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<document id="6A448F05B7788B692C17C91CE7804795" ID-CLB-Dataset="29039" ID-DOI="10.11646/zootaxa.4459.1.1" ID-GBIF-Dataset="9acf71bc-6414-4878-8af0-a64d3ec1fea6" ID-ISSN="1175-5326" ID-Zenodo-Dep="1458302" ID-ZooBank="633A9498-FCC6-4439-BE33-A45B9E51B550" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1539287574730" checkinUser="plazi" docAuthor="Grandinete, Yuri Campanholo, Noll, Fernando Barbosa &amp; Carpenter, James" docDate="2018" docId="03C5878FFFEDF321FF3096C3D788FCE7" docLanguage="en" docName="zootaxa.4459.1.1.pdf" docOrigin="Zootaxa 4459 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Eumenes verticalis Say 1824" docType="treatment" docVersion="9" lastPageNumber="34" masterDocId="FFFCFFF7FFF3F300FFA79155D339FFDF" masterDocTitle="Taxonomic Review of Eumenes Latreille, 1802 (Hymenoptera, Vespidae, Eumeninae) from the New World" masterLastPageNumber="52" masterPageNumber="1" pageNumber="31" updateTime="1698577660185" updateUser="ExternalLinkService">
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<mods:title id="0CEEA0652883FB9CF64D16D73C923D5B">Taxonomic Review of Eumenes Latreille, 1802 (Hymenoptera, Vespidae, Eumeninae) from the New World</mods:title>
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<mods:namePart id="5DF085B6D9D1DA99EEBBC5426DB397B5">Grandinete, Yuri Campanholo</mods:namePart>
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<mods:namePart id="212EEB7CCED6CB35BB7AC81D6038B731">Noll, Fernando Barbosa</mods:namePart>
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<treatment id="03C5878FFFEDF321FF3096C3D788FCE7" ID-DOI="http://doi.org/10.5281/zenodo.5986392" ID-GBIF-Taxon="148692483" ID-Zenodo-Dep="5986392" LSID="urn:lsid:plazi:treatment:03C5878FFFEDF321FF3096C3D788FCE7" httpUri="http://treatment.plazi.org/id/03C5878FFFEDF321FF3096C3D788FCE7" lastPageId="33" lastPageNumber="34" pageId="30" pageNumber="31">
<subSubSection id="C3766512FFEDF31EFF3096C3D168F871" box="[151,593,1941,1966]" pageId="30" pageNumber="31" type="nomenclature">
<paragraph id="8BD33699FFEDF31EFF3096C3D168F871" blockId="30.[151,1437,1941,2038]" box="[151,593,1941,1966]" pageId="30" pageNumber="31">
<taxonomicName id="4C6C4D1AFFEDF31EFF3096C3D2D1F871" ID-CoL="3CDQ8" authority="Say, 1824" authorityName="Say" authorityYear="1824" box="[151,488,1941,1966]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="verticalis">
<emphasis id="B918EA8BFFEDF31EFF3096C3D2D1F871" bold="true" box="[151,488,1941,1966]" pageId="30" pageNumber="31">
<emphasis id="B918EA8BFFEDF31EFF3096C3D248F871" bold="true" box="[151,369,1941,1966]" italics="true" pageId="30" pageNumber="31">Eumenes verticalis</emphasis>
Say, 1824
</emphasis>
</taxonomicName>
(
<figureCitation id="13572A1CFFEDF31EFE5F96C0D170F871" box="[504,585,1941,1966]" captionStart="FIGURES 1326" captionStartId="42.[151,264,1649,1671]" captionTargetBox="[152,1436,193,1628]" captionTargetId="figure@42.[151,1436,193,1628]" captionTargetPageId="42" captionText="FIGURES 1326. Eumenes species, lateral view. 13) E. cubensis Cresson, 1865 (female); 14) E. flavitinctus Bohart, 1950 (Holotype, female); 15) E. fervens Say, 1824 (Lectotype, female); 16) E. pictus (Smith, 1857) (female); 17) E. sculleni Bohart, 1950 (Holotype, male); 18) E. smithii de Saussure, 1852 (female); 19) E. belfragei Cresson, 1872 (Lectotype, female); 20) E. verticalis Say, 1824 (female); 21) Eumenes verticalis coloradensis Cresson, 1875 (lectotype, female); 22) E. verticalis neoboreus Bequaert, 1944 (Holotype, male); 23) E. verticalis tricinctus Isely, 1917 (Holotype, female); 24) E. filiformis de Saussure, 1855 (famele); 25) E. rufomaculata Fox, 1899 (Lectotype, female); 26) Eumenes versicolor de Saussure, 1852 (Lectotype, female). Scale bar = 1mm." httpUri="https://zenodo.org/record/1458306/files/figure.png" pageId="30" pageNumber="31">Fig. 20</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3766512FFEDF320FF6096ECD28EF9B5" lastPageId="32" lastPageNumber="33" pageId="30" pageNumber="31" type="description">
<paragraph id="8BD33699FFEDF31EFF6096ECD6A4F829" blockId="30.[151,1437,1941,2038]" pageId="30" pageNumber="31">
<emphasis id="B918EA8BFFEDF31EFF6096ECD2F9F80D" bold="true" box="[199,448,1977,2002]" pageId="30" pageNumber="31">Female. Color, head:</emphasis>
Black, with yellow marks as follows: wide spot on basal half of clypeus; inter-antennal region and a narrow stripe on basal half of scape and on upper half of gena, adjacent to the compound eyes.
</paragraph>
<paragraph id="8BD33699FFECF31FFF3091C2D179FDC7" blockId="31.[151,1437,151,2013]" pageId="31" pageNumber="32">
Brownish marks: on the two-third apical region of the mandibles; labrum; apex of the clypeus (on the apical projections) and marks on flagellum, mainly the first three and last two flagellomeres.
<emphasis id="B918EA8BFFECF31FFB2A91E9D62DFF0A" bold="true" box="[1165,1300,188,213]" pageId="31" pageNumber="32">Mesosoma:</emphasis>
Black with yellow marks as follows: a broad band on the anterior region on the dorsal face of the pronotum; small spot on upper half of mesepisternum, adjacent to the mesepimeron; tegula (with a brownish translucent mark on the center region of it); apical two-third of metanotum; small spot on the side of the upper half of the posterior face of propodeum and one wide mark on the basal half of it. Marks on the apical region of femora; almost entirely tibiae; first two tarsi yellow-brownish and the last three dark brownish.
<emphasis id="B918EA8BFFECF31FFC2D903AD736FE57" bold="true" box="[906,1039,367,392]" pageId="31" pageNumber="32">Metasoma:</emphasis>
predominantly black, with yellow marks as follows: two small spots on each side of the middle region of T1; a transverse band on apex of T1; a relatively broad band on each side of T2 and transverse band on apex of T2T4 and S2S5 (narrower on those sclerites and almost absent on S5). Wings hyaline, with the costal, medial and submedial cells darkened yellow. Veins and pterostigma dark brownish.
</paragraph>
<paragraph id="8BD33699FFECF31FFF609371D13CFCCB" blockId="31.[151,1437,151,2013]" pageId="31" pageNumber="32">
<emphasis id="B918EA8BFFECF31FFF609371D260FDE2" bold="true" box="[199,345,548,573]" pageId="31" pageNumber="32">Pubescence:</emphasis>
Golden pubescence covering the body.
<emphasis id="B918EA8BFFECF31FFC8B9371D04CFDE2" bold="true" box="[812,885,548,573]" pageId="31" pageNumber="32">Head:</emphasis>
long pubescence on frons, vertex and occipital region. Basal and lateral regions of clypeus with slightly shorter pubescence, while on center and apical regions it is very short. In oblique view, the pubescence on clypeus generally appears to be whitish. Pubescence on scape moderately long (shorter than on frons).
<emphasis id="B918EA8BFFECF31FFDDD93DAD03BFD77" bold="true" box="[634,770,655,680]" pageId="31" pageNumber="32">Mesosoma:</emphasis>
covered with long pubescence (as long as on frons and vertex).
<emphasis id="B918EA8BFFECF31FFF5093E1D245FD12" bold="true" box="[247,380,692,717]" pageId="31" pageNumber="32">Metasoma:</emphasis>
T1 with moderately long pubescence (shorter than on mesosoma); basal region of T2 with pubescence as long as on T1 and becoming shorter towards apex. The other sclerites have very short pubescence with some slightly long on apex.
</paragraph>
<paragraph id="8BD33699FFECF31FFF60924AD257FB87" blockId="31.[151,1437,151,2013]" pageId="31" pageNumber="32">
<emphasis id="B918EA8BFFECF31FFF60924AD111FCE7" bold="true" box="[199,552,799,824]" pageId="31" pageNumber="32">Surface of integument, head:</emphasis>
Clypeus with coarse punctation homogenously distributed. Frons and vertex with coarse punctation, denser than on clypeus. Behind the lateral ocelli the punctation becomes sparser. Gena with weak punctation and very sparse distributed.
<emphasis id="B918EA8BFFECF31FFD359232D023FC5F" bold="true" box="[658,794,871,896]" pageId="31" pageNumber="32">Mesosoma:</emphasis>
Coarse and dense (as on frons) punctation, except on the upper half of mesepisternum which are slightly sparser and on anterior half of the lateral face of the propodeum which is reduced.
<emphasis id="B918EA8BFFECF31FFECC92FAD2C8FC17" bold="true" box="[363,497,943,968]" pageId="31" pageNumber="32">Metasoma:</emphasis>
absent on the basal third of T1 and coarse on the two-third of the apical region, sparser than on mesosoma, but becoming denser on apex region. T2 with coarse and dense punctation, denser on apex. On lateral margins of T2 the punctation becomes sparser than on dorsal face. S2 with coarse and sparse punctation mainly on lateral margins. T3T5 and S3S5 with weak and very sparse punctation. T6 and S6 with no evident punctation.
</paragraph>
<paragraph id="8BD33699FFECF31FFF609531D6A4FB36" blockId="31.[151,1437,151,2013]" pageId="31" pageNumber="32">
<emphasis id="B918EA8BFFECF31FFF609531D2B0FBA2" bold="true" box="[199,393,1124,1149]" pageId="31" pageNumber="32">Structure, head:</emphasis>
<collectingRegion id="49A8F87BFFECF31FFE379531D2E2FBA3" box="[400,475,1124,1148]" country="France" name="Centre" pageId="31" pageNumber="32">Center</collectingRegion>
of clypeus strongly convex; apical region of the clypeus concave, forming two apical and rounded projections with very weak carinae on apex (
<figureCitation id="13572A1CFFECF31FFC8795D2D04CFB7F" box="[800,885,1159,1184]" captionStart="FIGURES 6676" captionStartId="47.[151,264,1772,1794]" captionTargetBox="[151,1436,193,1750]" captionTargetId="figure@47.[151,1436,193,1750]" captionTargetPageId="47" captionText="FIGURES 6676. Clypeus. 66) Eumenes americanus (female); 67) E. smithii (female); 68) E. crucifera (female); 69) E. crucifera (male); 70) E. verticalis (female); 71) E. fraternus (female); 72) E. fraternus (male); 73) E. sculleni (female); 74) E. filiformis (female); 75) E. rufomaculatus (female); 76) E. versicolor (female). Scale bar = 1mm." httpUri="https://zenodo.org/record/1458316/files/figure.png" pageId="31" pageNumber="32">Fig. 70</figureCitation>
). Inter-antennal region longitudinally cariniform, slightly wider than the antennal socket; lateral ocelli closer to each other than to the compound eyes; occipital carina very close to the compound eye next to the mandibles and weakly angled on middle region of gena.
</paragraph>
<paragraph id="8BD33699FFECF31FFF3095A1D743FA3A" blockId="31.[151,1437,151,2013]" pageId="31" pageNumber="32">
<emphasis id="B918EA8BFFECF31FFF3095A1D227FAD2" bold="true" box="[151,286,1268,1293]" pageId="31" pageNumber="32">Mesosoma:</emphasis>
carina on dorsal face of pronotum weakly developed, while on lateral face it is more developed. Pronotal fovea present; mesepimeron slightly elevated regarding to the mesepisternum level; tegula with posterior projection weakly developed, rounded; posterior face of strongly concave on basal half; carina on upper half of external side of fore coxa.
<emphasis id="B918EA8BFFECF31FFE63940AD170FAA7" bold="true" box="[452,585,1375,1400]" pageId="31" pageNumber="32">Metasoma:</emphasis>
T1 not more than 2.5 times longer than wide; moderately swollen in dorsal view (
<figureCitation id="13572A1CFFECF31FFF7A94D1D209FA43" box="[221,304,1412,1437]" captionStart="FIGURES 2742" captionStartId="43.[151,264,1902,1924]" captionTargetBox="[151,1435,193,1881]" captionTargetId="figure@43.[151,1436,193,1881]" captionTargetPageId="43" captionText="FIGURES 2742. First tergum, dorsal view. 27) Eumenes americanus; 28) E. bequaerti; 29) E. bollii; 30) E. consobrinus; 31) E. coyotae; 32) E. crucifera; 33) E. cubensis; 34) E. flavitinctus; 35) E. fraternus; 36) E. pictus; 37) E. sculleni; 38) E. smithii; 39) E. verticalis; 40) E. filiformis; 41) E. rufomaculactus; 42) E. versicolor. Scale bar = 1mm." httpUri="https://zenodo.org/record/1458308/files/figure.png" pageId="31" pageNumber="32">Fig. 38</figureCitation>
), with the lateral margins on apical half concave and lateral margins on apex divergent. T2, in dorsal view, longer than wide, with the lateral margins weakly concave (
<figureCitation id="13572A1CFFECF31FFCD694F2D0FCFA1F" box="[881,965,1447,1472]" captionStart="FIGURES 4358" captionStartId="44.[151,264,1946,1968]" captionTargetBox="[202,1388,195,1934]" captionTargetId="figure@44.[201,1388,193,1934]" captionTargetPageId="44" captionText="FIGURES 4358. Second tergum, dorsal view. 43) Eumenes americanus; 44) E. bequaerti; 45) E. bollii; 46) E. consobrinus; 47) E. coyotae; 48) E. crucifera; 49) E. cubensis; 50) E. flavitinctus; 51) E. fraternus; 52) E. pictus; 53) E. sculleni; 54) E. smithii; 55) E. verticalis; 56) E. filiformis; 57) E. rufomaculactus; 58) E. versicolor. Scale bar = 1mm." httpUri="https://zenodo.org/record/1458310/files/figure.png" pageId="31" pageNumber="32">Fig. 54</figureCitation>
). Apex of T2 with lamella which becomes reduced until completely disappear. Basal angle of T2, in lateral view, rising up abruptly.
</paragraph>
<paragraph id="8BD33699FFECF31FFF6094BAD0C7F98F" blockId="31.[151,1437,151,2013]" pageId="31" pageNumber="32">
<emphasis id="B918EA8BFFECF31FFF6094BAD234F9D7" bold="true" box="[199,269,1519,1544]" pageId="31" pageNumber="32">Male:</emphasis>
Clypeus narrower than females; F11 long, narrow basally, with the apex pointed and surpassing the apical margin of F8 (
<figureCitation id="13572A1CFFECF31FFE289741D2DFF9F3" box="[399,486,1556,1581]" captionStart="FIGURES 7792" captionStartId="48.[151,264,1946,1968]" captionTargetBox="[151,1436,193,1925]" captionTargetId="figure@48.[151,1436,193,1925]" captionTargetPageId="48" captionText="FIGURES 7792. First tergum, dorsal view. 77) Eumenes americanus; 78) E. bequaerti; 79) E. bollii; 80) E. consobrinus; 81) E. coyotae; 82) E. crucifera; 83) E. cubensis; 84) E. flavitinctus; 85) E. fraternus; 86) E. pictus; 87) E. sculleni; 88) E. smithii; 89) E. verticalis; 90) E. filiformis; 91) E. rufomaculactus; 92) E. versicolor. Scale bar = 1mm." httpUri="https://zenodo.org/record/1458318/files/figure.png" pageId="31" pageNumber="32">Fig. 89</figureCitation>
). Dorsal surface without longitudinal carina. Microscopic erect bristles on ventral surface of F11. S7 flattened apically, with some bristles scattered on the apex.
</paragraph>
<paragraph id="8BD33699FFECF31FFF609709D0A6F93F" blockId="31.[151,1437,151,2013]" pageId="31" pageNumber="32">
<emphasis id="B918EA8BFFECF31FFF609709D245F9AA" bold="true" box="[199,380,1628,1653]" pageId="31" pageNumber="32">Male genitalia:</emphasis>
Aedeagus as in figure 105a, b. Paramere (
<figureCitation id="13572A1CFFECF31FFCCE9709D0EEF9AA" box="[873,983,1628,1653]" captionStart="FIGURES 102109" captionStartId="50.[151,264,1926,1948]" captionTargetBox="[151,1436,193,1903]" captionTargetId="figure@50.[151,1436,193,1904]" captionTargetPageId="50" captionText="FIGURES 102109. Male genitalia: a, b) aedeagus (lateral and ventral view, respectively); c) paramere, internal view. 102) E. pictus; 103) E. sculleni; 104) E. smithii; 105) E. verticalis; 106) E. filiformis; 107) E. rufomaculatus; 108) E. versicolor; 109) General scheme of the apical projection of the aedeagus: a) weakly concave/truncate; b) strongly concave. Scale bar = 1mm." httpUri="https://zenodo.org/record/1458322/files/figure.png" pageId="31" pageNumber="32">Fig. 105c</figureCitation>
) with long bristles on middle region of gonostyle; digitus broad, not reaching the middle region of the gonostyle, covered with short bristles (
<figureCitation id="13572A1CFFECF31FFA8C972AD6B6F947" box="[1323,1423,1663,1688]" captionStart="FIGURES 110125" captionStartId="51.[151,264,1899,1921]" captionTargetBox="[163,1425,209,1857]" captionTargetId="figure@51.[150,1437,193,1878]" captionTargetPageId="51" captionText="FIGURES 110125. Male genitalia: digitus. 110) Eumenes americanus; 111) E. bequaerti; 112) E. bollii; 113) E. consobrinus; 114) E. coyotae; 115) E. crucifera; 116) E. cubensis; 117) E. flavitinctus; 118) E. fraternus; 119) E. pictus; 120) E. sculleni; 121) E. smithii; 122) E. verticalis; 123) E. filiformis; 124) E. rufomaculatus; 125) E. versicolor." httpUri="https://zenodo.org/record/1458324/files/figure.png" pageId="31" pageNumber="32">Fig. 122</figureCitation>
); volsella with long bristles on ventral margin and short scattered bristles; cuspis with moderately long bristles (longer than on digitus); distal lobe truncate, without evident bristles.
</paragraph>
<paragraph id="8BD33699FFECF320FF6097B9D2F6FF0B" blockId="31.[151,1437,151,2013]" lastBlockId="32.[151,1437,151,2035]" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32">
<emphasis id="B918EA8BFFECF31FFF6097B9D278F8DA" bold="true" box="[199,321,1772,1797]" pageId="31" pageNumber="32">Variation:</emphasis>
The specimens from
<collectingCountry id="F37B7609FFECF31FFD9197B9D1B3F8DA" box="[566,650,1772,1797]" name="Canada" pageId="31" pageNumber="32">Canada</collectingCountry>
to the central west of the
<collectingCountry id="F37B7609FFECF31FFC1697B9D0D0F8DB" box="[945,1001,1772,1796]" name="United States of America" pageId="31" pageNumber="32">USA</collectingCountry>
have the yellow marks less developed than those from the central east. Color variation is as follows: apex of clypeus with spots on each side or almost entirely yellow, with just a longitudinal black band on the apex; marks on mesepisternum, upper half of posterior face of propodeum and T
<date id="FFD21059FFECF31FFE18960DD13AF8AE" box="[447,515,1880,1905]" pageId="31" pageNumber="32">1 may</date>
be reduced or even absent; small spot on each side of scutellum (one specimen presented a spot only in one side of the scutellum); marks on lateral margins of T
<date id="FFD21059FFECF31FFBF19629D7A3F84A" box="[1110,1178,1916,1941]" pageId="31" pageNumber="32">1 may</date>
be reduced; brownish marks on the flagellum may be absent. The integument may be violaceous mixed with blackish, better observed on laterally on the mesosoma and metasoma. Some specimens have the pubescence of clypeus whitish in oblique view. The posterior face of the propodeum may have the concavity weaker; and the punctation of T
<date id="FFD21059FFD3F320FA8991CDD64EFF6F" box="[1326,1399,152,176]" pageId="32" pageNumber="33">2 may</date>
be slightly weaker and sparser.
</paragraph>
<paragraph id="8BD33699FFD3F320FF6091B5D053FD7A" blockId="32.[151,1437,151,2035]" pageId="32" pageNumber="33">
We examined all the
<typeStatus id="54D7883BFFD3F320FE1A91B5D2D4FF27" box="[445,493,224,248]" pageId="32" pageNumber="33">type</typeStatus>
series of
<taxonomicName id="4C6C4D1AFFD3F320FDC691B4D087FF27" authority="Cresson, 1875" authorityName="Cresson" authorityYear="1875" box="[609,958,223,248]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="coloradensis">
<emphasis id="B918EA8BFFD3F320FDC691B4D02BFF27" box="[609,786,224,248]" italics="true" pageId="32" pageNumber="33">E. coloradensis</emphasis>
Cresson, 1875
</taxonomicName>
(
<figureCitation id="13572A1CFFD3F320FC76918AD71FFF27" box="[977,1062,223,248]" captionStart="FIGURES 1326" captionStartId="42.[151,264,1649,1671]" captionTargetBox="[152,1436,193,1628]" captionTargetId="figure@42.[151,1436,193,1628]" captionTargetPageId="42" captionText="FIGURES 1326. Eumenes species, lateral view. 13) E. cubensis Cresson, 1865 (female); 14) E. flavitinctus Bohart, 1950 (Holotype, female); 15) E. fervens Say, 1824 (Lectotype, female); 16) E. pictus (Smith, 1857) (female); 17) E. sculleni Bohart, 1950 (Holotype, male); 18) E. smithii de Saussure, 1852 (female); 19) E. belfragei Cresson, 1872 (Lectotype, female); 20) E. verticalis Say, 1824 (female); 21) Eumenes verticalis coloradensis Cresson, 1875 (lectotype, female); 22) E. verticalis neoboreus Bequaert, 1944 (Holotype, male); 23) E. verticalis tricinctus Isely, 1917 (Holotype, female); 24) E. filiformis de Saussure, 1855 (famele); 25) E. rufomaculata Fox, 1899 (Lectotype, female); 26) Eumenes versicolor de Saussure, 1852 (Lectotype, female). Scale bar = 1mm." httpUri="https://zenodo.org/record/1458306/files/figure.png" pageId="32" pageNumber="33">Fig. 21</figureCitation>
);
<taxonomicName id="4C6C4D1AFFD3F320FB9991B4D6A5FF27" authority="Viereck, 1908" authorityName="Viereck" authorityYear="1908" box="[1086,1436,223,248]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="cruciferorum">
<emphasis id="B918EA8BFFD3F320FB9991B4D7CAFF27" box="[1086,1267,224,248]" italics="true" pageId="32" pageNumber="33">E. cruciferorum</emphasis>
Viereck, 1908
</taxonomicName>
(
<typeStatus id="54D7883BFFD3F320FF399056D23EFEC3" box="[158,263,259,284]" pageId="32" pageNumber="33" type="lectotype">lectotype</typeStatus>
female and
<typeStatus id="54D7883BFFD3F320FE3D9056D108FEC3" box="[410,561,259,284]" pageId="32" pageNumber="33" type="paralectotype">paralectotype</typeStatus>
male by present designation);
<taxonomicName id="4C6C4D1AFFD3F320FC3D9051D7D6FEC3" authority="Viereck, 1908" authorityName="Viereck" authorityYear="1908" box="[922,1263,259,284]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="enigmaticus">
<emphasis id="B918EA8BFFD3F320FC3D9051D77AFEC4" box="[922,1091,260,283]" italics="true" pageId="32" pageNumber="33">E. enigmaticus</emphasis>
Viereck, 1908
</taxonomicName>
(
<typeStatus id="54D7883BFFD3F320FAA69056D650FEC3" box="[1281,1385,259,284]" pageId="32" pageNumber="33" type="lectotype">lectotype</typeStatus>
and
<typeStatus id="54D7883BFFD3F320FF309072D217FE9F" box="[151,302,295,320]" pageId="32" pageNumber="33" type="paralectotype">paralectotype</typeStatus>
male by present designation);
<taxonomicName id="4C6C4D1AFFD3F320FD2C907DD08AFE9F" authority="Isely, 1917" authorityName="Isely" authorityYear="1917" box="[651,947,295,320]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="stenogaster">
<emphasis id="B918EA8BFFD3F320FD2C907DD015FEE0" box="[651,812,296,319]" italics="true" pageId="32" pageNumber="33">E. stenogaster</emphasis>
<bibRefCitation id="EFFD4B68FFD3F320FC939072D08AFE9F" box="[820,947,295,320]" pageId="32" pageNumber="33" refId="ref43377" refString="Isely, D. (1917) A synopsis of the petiolate wasps of the family Eumenidae (Hymenoptera), found in America North of Mexico. Annals of the Entomological Society of America, 10 (4), 345 - 366. https: // doi. org / 10.1093 / aesa / 10.4.345" type="journal article">Isely, 1917</bibRefCitation>
</taxonomicName>
; and most of the specimens of
<taxonomicName id="4C6C4D1AFFD3F320FAB3907DD21FFEBC" authorityName="Cresson" authorityYear="1875" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="subSpecies" species="verticalis" subSpecies="coloradensis">
<emphasis id="B918EA8BFFD3F320FAB3907DD21FFEBC" italics="true" pageId="32" pageNumber="33">E. verticalis coloradensis</emphasis>
</taxonomicName>
. All of them presented the same features described for
<taxonomicName id="4C6C4D1AFFD3F320FC389019D71EFEBC" authorityName="Say" authorityYear="1824" box="[927,1063,331,355]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="verticalis">
<emphasis id="B918EA8BFFD3F320FC389019D71EFEBC" box="[927,1063,331,355]" italics="true" pageId="32" pageNumber="33">E. verticalis</emphasis>
</taxonomicName>
, showing only the yellow marks frequently more developed as follows: clypeus often yellow with a black mark that goes from the base to the middle area and sometime on the lateral margins (or only with the black on center region of it); mark on the scape longer; spots on each side of the scutellum, which vary in size (and sometimes absent); the spots on the sides of the posterior face of the propodeum may be wider and often contiguous (in some species totally absent); the band on the lateral margins of T2 broader and sometimes contiguous laterally with the apical band, which it is also clearly broader, as long as the apical band on the T3T5 and S2S6. Although the differences relating to the yellow marks, the pubescence, punctation, and external morphology (and male genitalia) were the same as typical
<taxonomicName id="4C6C4D1AFFD3F320FAAF9310D649FD82" authorityName="Say" authorityYear="1824" box="[1288,1392,581,605]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="verticalis">
<emphasis id="B918EA8BFFD3F320FAAF9310D649FD82" box="[1288,1392,581,605]" italics="true" pageId="32" pageNumber="33">verticalis</emphasis>
</taxonomicName>
, all included in the range of variation listed. One specimen from
<collectingRegion id="49A8F87BFFD3F320FC9C933FD0A6FD5E" box="[827,927,618,641]" country="United States of America" name="Montana" pageId="32" pageNumber="33">Montana</collectingRegion>
showed the punctation of the basal half of T2 sparser, as seen in
<taxonomicName id="4C6C4D1AFFD3F320FEC093D8D2D7FD7A" authorityName="Say" authorityYear="1824" box="[359,494,653,677]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="verticalis">
<emphasis id="B918EA8BFFD3F320FEC093D8D2D7FD7A" box="[359,494,653,677]" italics="true" pageId="32" pageNumber="33">E. verticalis</emphasis>
</taxonomicName>
from
<collectingRegion id="49A8F87BFFD3F320FD9493D9D14FFD7A" box="[563,630,652,677]" country="United States of America" name="Idaho" pageId="32" pageNumber="33">Idaho</collectingRegion>
,
<collectingRegion id="49A8F87BFFD3F320FD2793D9D1C8FD7A" box="[640,753,652,677]" country="United States of America" name="Wyoming" pageId="32" pageNumber="33">Wyoming</collectingRegion>
, and
<collectingRegion id="49A8F87BFFD3F320FC8B93D9D05FFD7A" box="[812,870,652,677]" country="United States of America" name="Utah" pageId="32" pageNumber="33">Utah</collectingRegion>
.
</paragraph>
<paragraph id="8BD33699FFD3F320FF6093E4D14FFB46" blockId="32.[151,1437,151,2035]" pageId="32" pageNumber="33">
<bibRefCitation id="EFFD4B68FFD3F320FF6093E4D2BBFD16" box="[199,386,689,713]" pageId="32" pageNumber="33" refId="ref42355" refString="Bequaert, J. (1944) The genus Eumenes in the Dominion of Canada. Le Naturaliste Canadien, 71, 75 - 88." type="journal article">Bequaert (1944)</bibRefCitation>
described
<taxonomicName id="4C6C4D1AFFD3F320FDA393E4D07DFD17" authorityName="Bequaert" authorityYear="1944" box="[516,836,688,712]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="variety" species="verticalis" variety="neoboreus">
<emphasis id="B918EA8BFFD3F320FDA393E4D1A9FD17" box="[516,656,688,712]" italics="true" pageId="32" pageNumber="33">E. verticalis</emphasis>
var.
<emphasis id="B918EA8BFFD3F320FD7793E5D07DFD17" box="[720,836,688,712]" italics="true" pageId="32" pageNumber="33">neoboreus</emphasis>
</taxonomicName>
(
<figureCitation id="13572A1CFFD3F320FCF093E5D094FD16" box="[855,941,688,713]" captionStart="FIGURES 1326" captionStartId="42.[151,264,1649,1671]" captionTargetBox="[152,1436,193,1628]" captionTargetId="figure@42.[151,1436,193,1628]" captionTargetPageId="42" captionText="FIGURES 1326. Eumenes species, lateral view. 13) E. cubensis Cresson, 1865 (female); 14) E. flavitinctus Bohart, 1950 (Holotype, female); 15) E. fervens Say, 1824 (Lectotype, female); 16) E. pictus (Smith, 1857) (female); 17) E. sculleni Bohart, 1950 (Holotype, male); 18) E. smithii de Saussure, 1852 (female); 19) E. belfragei Cresson, 1872 (Lectotype, female); 20) E. verticalis Say, 1824 (female); 21) Eumenes verticalis coloradensis Cresson, 1875 (lectotype, female); 22) E. verticalis neoboreus Bequaert, 1944 (Holotype, male); 23) E. verticalis tricinctus Isely, 1917 (Holotype, female); 24) E. filiformis de Saussure, 1855 (famele); 25) E. rufomaculata Fox, 1899 (Lectotype, female); 26) Eumenes versicolor de Saussure, 1852 (Lectotype, female). Scale bar = 1mm." httpUri="https://zenodo.org/record/1458306/files/figure.png" pageId="32" pageNumber="33">Fig. 22</figureCitation>
) and stated that this variety has the same markings as
<taxonomicName id="4C6C4D1AFFD3F320FE819380D297FD33" authorityName="Say" authorityYear="1824" box="[294,430,724,748]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="verticalis">
<emphasis id="B918EA8BFFD3F320FE819380D297FD33" box="[294,430,724,748]" italics="true" pageId="32" pageNumber="33">E. verticalis</emphasis>
</taxonomicName>
, but with “white to creamy-white instead of yellow” and “the more northern form of the species and is known only from the
<collectingCountry id="F37B7609FFD3F320FD9393A2D01FFCCF" box="[564,806,759,784]" name="Canada" pageId="32" pageNumber="33">Dominion of Canada</collectingCountry>
… In the western provinces, it occurs in some localities together with the typical form and there are also a transitional species” (pag. 87). We examined just one male in addition to the
<typeStatus id="54D7883BFFD3F320FEE29215D24CFC87" box="[325,373,832,856]" pageId="32" pageNumber="33">type</typeStatus>
specimens (for which we did not have authorization to extract the genitalia), and we did not find evident differences in pubescence and morphological structure, including the morphology of the male genitalia. One difference that we did find was a male with the punctation of T2 weaker and sparser than that on the
<typeStatus id="54D7883BFFD3F320FF3092FED3FEFC1C" box="[151,199,939,963]" pageId="32" pageNumber="33">type</typeStatus>
species, a pattern commonly seen in the East coast of the
<collectingCountry id="F37B7609FFD3F320FCFB92FED0ADFC1C" box="[860,916,939,963]" name="United States of America" pageId="32" pageNumber="33">USA</collectingCountry>
(see below), although this species is recorded from
<collectingRegion id="49A8F87BFFD3F320FF73929BD27BFC38" box="[212,322,974,999]" country="United States of America" name="Wyoming" pageId="32" pageNumber="33">Wyoming</collectingRegion>
state (species of
<taxonomicName id="4C6C4D1AFFD3F320FDA6929AD190FC39" authorityName="de Saussure" authorityYear="1855" box="[513,681,974,998]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="consobrinus">
<emphasis id="B918EA8BFFD3F320FDA6929AD190FC39" box="[513,681,974,998]" italics="true" pageId="32" pageNumber="33">E. consobrinus</emphasis>
</taxonomicName>
also presented this pattern on the same region). The morphology of the male genitalia was similar to other genitalia examined (see
<emphasis id="B918EA8BFFD3F320FCFD92A7D0C2FBD5" box="[858,1019,1010,1034]" italics="true" pageId="32" pageNumber="33">male genitalia</emphasis>
section), which supported placing
<taxonomicName id="4C6C4D1AFFD3F320FA2392A6D242FBF1" authorityName="Bequaert" authorityYear="1944" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="subSpecies" species="verticalis" subSpecies="neoboreus">
<emphasis id="B918EA8BFFD3F320FA2392A6D242FBF1" italics="true" pageId="32" pageNumber="33">E. verticalis neoboreus</emphasis>
</taxonomicName>
as a synonym of
<taxonomicName id="4C6C4D1AFFD3F320FDE39542D1F4FBF1" authorityName="Say" authorityYear="1824" box="[580,717,1046,1070]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="verticalis">
<emphasis id="B918EA8BFFD3F320FDE39542D1F4FBF1" box="[580,717,1046,1070]" italics="true" pageId="32" pageNumber="33">E. verticalis</emphasis>
</taxonomicName>
with a color variation possibly due the longitudinal differences between populations. We decided to maintain the term
<taxonomicName id="4C6C4D1AFFD3F320FD5D956CD057FB8E" authorityName="Bequaert" authorityYear="1944" box="[762,878,1081,1105]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="subSpecies" species="verticalis" subSpecies="neoboreus">
<emphasis id="B918EA8BFFD3F320FD5D956CD057FB8E" box="[762,878,1081,1105]" italics="true" pageId="32" pageNumber="33">neoboreus</emphasis>
</taxonomicName>
to facilitate the recognition of the variation, as we proposed for the other variants below. Among all
<typeStatus id="54D7883BFFD3F320FD6B950BD00EFBA9" box="[716,823,1118,1142]" pageId="32" pageNumber="33" type="paratype">paratypes</typeStatus>
of
<taxonomicName id="4C6C4D1AFFD3F320FCF9950BD75BFBAA" authorityName="Bequaert" authorityYear="1944" box="[862,1122,1117,1141]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="subSpecies" species="verticalis" subSpecies="neoboreus">
<emphasis id="B918EA8BFFD3F320FCF9950BD75BFBAA" box="[862,1122,1117,1141]" italics="true" pageId="32" pageNumber="33">E. verticalis neoboreus</emphasis>
</taxonomicName>
deposited in the ROM, one specimen (male) was actually
<taxonomicName id="4C6C4D1AFFD3F320FE4D95D7D156FB46" authorityName="Say" authorityYear="1824" box="[490,623,1153,1177]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="fraternus">
<emphasis id="B918EA8BFFD3F320FE4D95D7D156FB46" box="[490,623,1153,1177]" italics="true" pageId="32" pageNumber="33">E. fraternus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BD33699FFD3F320FF6095F0D28EF9B5" blockId="32.[151,1437,151,2035]" pageId="32" pageNumber="33">
As we observed for
<taxonomicName id="4C6C4D1AFFD3F320FE0A95F0D16FFB62" authorityName="de Saussure" authorityYear="1855" box="[429,598,1189,1213]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="consobrinus">
<emphasis id="B918EA8BFFD3F320FE0A95F0D16FFB62" box="[429,598,1189,1213]" italics="true" pageId="32" pageNumber="33">E. consobrinus</emphasis>
</taxonomicName>
, the West coast of the
<collectingCountry id="F37B7609FFD3F320FCFC95F0D0AAFB62" box="[859,915,1189,1213]" name="United States of America" pageId="32" pageNumber="33">USA</collectingCountry>
has populations with different punctation and color than the typical eastern forms. We observed the same in the material examined of
<taxonomicName id="4C6C4D1AFFD3F320FBC4959CD3F6FADA" authority="Isely, 1917" authorityName="Isely" authorityYear="1917" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="subSpecies" species="verticalis" subSpecies="tricinctus">
<emphasis id="B918EA8BFFD3F320FBC4959CD663FB3F" box="[1123,1370,1224,1248]" italics="true" pageId="32" pageNumber="33">E. verticalis tricinctus</emphasis>
<bibRefCitation id="EFFD4B68FFD3F320FAC7959DD3F6FADA" pageId="32" pageNumber="33" refId="ref43377" refString="Isely, D. (1917) A synopsis of the petiolate wasps of the family Eumenidae (Hymenoptera), found in America North of Mexico. Annals of the Entomological Society of America, 10 (4), 345 - 366. https: // doi. org / 10.1093 / aesa / 10.4.345" type="journal article">Isely, 1917</bibRefCitation>
</taxonomicName>
(
<figureCitation id="13572A1CFFD3F320FF7995B9D20BFADA" box="[222,306,1260,1285]" captionStart="FIGURES 1326" captionStartId="42.[151,264,1649,1671]" captionTargetBox="[152,1436,193,1628]" captionTargetId="figure@42.[151,1436,193,1628]" captionTargetPageId="42" captionText="FIGURES 1326. Eumenes species, lateral view. 13) E. cubensis Cresson, 1865 (female); 14) E. flavitinctus Bohart, 1950 (Holotype, female); 15) E. fervens Say, 1824 (Lectotype, female); 16) E. pictus (Smith, 1857) (female); 17) E. sculleni Bohart, 1950 (Holotype, male); 18) E. smithii de Saussure, 1852 (female); 19) E. belfragei Cresson, 1872 (Lectotype, female); 20) E. verticalis Say, 1824 (female); 21) Eumenes verticalis coloradensis Cresson, 1875 (lectotype, female); 22) E. verticalis neoboreus Bequaert, 1944 (Holotype, male); 23) E. verticalis tricinctus Isely, 1917 (Holotype, female); 24) E. filiformis de Saussure, 1855 (famele); 25) E. rufomaculata Fox, 1899 (Lectotype, female); 26) Eumenes versicolor de Saussure, 1852 (Lectotype, female). Scale bar = 1mm." httpUri="https://zenodo.org/record/1458306/files/figure.png" pageId="32" pageNumber="33">Fig. 23</figureCitation>
), including the following yellow marks: wider marks on mesepisternum, including on the basal half; marks on scutellum wider, often contiguous, occupying the anterior half; marks on the lateral margins of the posterior face of the propodeum more developed, marks on the middle area of T1 more developed, often contiguous laterally with an apical band; lateral and apical bands of T2 more developed, occupying almost the entire sclerite, except for black marks on the basal half (and sometimes also laterally) and a transverse band on the middle area (sometimes the transverse band is contiguous with the basal mark). This pattern of marks was observed mostly in females, while the males generally had the same pattern that we described for the typical species and/or
<taxonomicName id="4C6C4D1AFFD3F320FF3094B3D21FFA21" authorityName="Cresson" authorityYear="1875" box="[151,294,1510,1534]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="coloradensis">
<emphasis id="B918EA8BFFD3F320FF3094B3D21FFA21" box="[151,294,1510,1534]" italics="true" pageId="32" pageNumber="33">coloradensis</emphasis>
</taxonomicName>
variants and the genitalia were not different from the variants of
<taxonomicName id="4C6C4D1AFFD3F320FBB194B2D7A6FA21" authorityName="Say" authorityYear="1824" box="[1046,1183,1510,1534]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="verticalis">
<emphasis id="B918EA8BFFD3F320FBB194B2D7A6FA21" box="[1046,1183,1510,1534]" italics="true" pageId="32" pageNumber="33">E. verticalis</emphasis>
</taxonomicName>
. All the specimens of
<taxonomicName id="4C6C4D1AFFD3F320FF30975FD3C6F9FD" authorityName="Say" authorityYear="1824" box="[151,255,1546,1570]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="verticalis">
<emphasis id="B918EA8BFFD3F320FF30975FD3C6F9FD" box="[151,255,1546,1570]" italics="true" pageId="32" pageNumber="33">verticalis</emphasis>
</taxonomicName>
variety presented the punctation of T2 clearly sparser than the typical form, mainly on the basal half. The
<typeStatus id="54D7883BFFD3F320FF30977AD3EBF998" box="[151,210,1583,1607]" pageId="32" pageNumber="33">types</typeStatus>
of
<taxonomicName id="4C6C4D1AFFD3F320FEA4977AD13EF999" authorityName="Isely" authorityYear="1917" box="[259,519,1582,1606]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="subSpecies" species="verticalis" subSpecies="tricinctus">
<emphasis id="B918EA8BFFD3F320FEA4977AD13EF999" box="[259,519,1582,1606]" italics="true" pageId="32" pageNumber="33">E. verticalis tricinctus</emphasis>
</taxonomicName>
also presented the mesosomal punctation slightly weak, but most specimens matched the typical form.
</paragraph>
</subSubSection>
<subSubSection id="C3766512FFD3F321FF609720D788FCE7" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33" type="materials_examined">
<paragraph id="8BD33699FFD3F320FF609720D75AF82D" blockId="32.[151,1437,151,2035]" pageId="32" pageNumber="33">
<emphasis id="B918EA8BFFD3F320FF609720D24CF951" bold="true" box="[199,373,1653,1678]" pageId="32" pageNumber="33">
<typeStatus id="54D7883BFFD3F320FF609720D23BF951" box="[199,258,1653,1678]" pageId="32" pageNumber="33">Type</typeStatus>
material.
</emphasis>
The
<typeStatus id="54D7883BFFD3F320FE099720D129F951" box="[430,528,1653,1678]" pageId="32" pageNumber="33" type="holotype">holotype</typeStatus>
male of
<taxonomicName id="4C6C4D1AFFD3F320FDD49723D79CF951" authority="Bequaert, 1944" authorityName="Bequaert" authorityYear="1944" box="[627,1189,1653,1678]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="variety" species="verticalis" variety="neoboreus">
<emphasis id="B918EA8BFFD3F320FDD49723D07EF952" box="[627,839,1653,1677]" italics="true" pageId="32" pageNumber="33">Eumenes verticalis</emphasis>
var.
<emphasis id="B918EA8BFFD3F320FCD89720D0CAF952" box="[895,1011,1653,1677]" italics="true" pageId="32" pageNumber="33">neoboreus</emphasis>
<bibRefCitation id="EFFD4B68FFD3F320FC5D9723D79CF951" box="[1018,1189,1654,1678]" pageId="32" pageNumber="33" refId="ref42355" refString="Bequaert, J. (1944) The genus Eumenes in the Dominion of Canada. Le Naturaliste Canadien, 71, 75 - 88." type="journal article">Bequaert, 1944</bibRefCitation>
</taxonomicName>
is excellent condition and bears the labels: 8 La Treppe \
<date id="FFD21059FFD3F320FD9897CFD1B5F96E" box="[575,652,1689,1713]" pageId="32" pageNumber="33" value="1942-07">VII 42</date>
Qué. \ J. Ouellet [Red Label] M.C.Z. \
<typeStatus id="54D7883BFFD3F320FBD697CCD7EAF96D" box="[1137,1235,1689,1714]" pageId="32" pageNumber="33" type="holotype">holotype</typeStatus>
\
<taxonomicName id="4C6C4D1AFFD3F320FB4997CCD65BF96E" authorityName="Bequaert" authorityYear="1944" box="[1262,1378,1689,1713]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="subSpecies" species="verticalis" subSpecies="neoboreus">
<emphasis id="B918EA8BFFD3F320FB4997CCD65BF96E" box="[1262,1378,1689,1713]" italics="true" pageId="32" pageNumber="33">neoboreus</emphasis>
</taxonomicName>
\
<specimenCount id="9D6AFD10FFD3F320FADA97CCD6AFF96E" box="[1405,1430,1689,1713]" pageId="32" pageNumber="33" type="male"></specimenCount>
<taxonomicName id="4C6C4D1AFFD3F320FF0797E8D23CF90B" box="[160,261,1725,1748]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="B918EA8BFFD3F320FF0797E8D23CF90B" box="[160,261,1725,1748]" italics="true" pageId="32" pageNumber="33">Eumenes</emphasis>
</taxonomicName>
\
<taxonomicName id="4C6C4D1AFFD3F320FE8297E8D2B4F90A" authorityName="Say" authorityYear="1824" box="[293,397,1725,1749]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="verticalis">
<emphasis id="B918EA8BFFD3F320FE8297E8D2B4F90A" box="[293,397,1725,1749]" italics="true" pageId="32" pageNumber="33">verticalis</emphasis>
</taxonomicName>
\ var. \
<taxonomicName id="4C6C4D1AFFD3F320FE5097E8D152F90A" authorityName="Bequaert" authorityYear="1944" box="[503,619,1725,1749]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="subSpecies" species="verticalis" subSpecies="neoboreus">
<emphasis id="B918EA8BFFD3F320FE5097E8D152F90A" box="[503,619,1725,1749]" italics="true" pageId="32" pageNumber="33">neoboreus</emphasis>
</taxonomicName>
\ J. Beq [Red Label] M.C.Z. \
<typeStatus id="54D7883BFFD3F320FC5A97E9D75CF90A" box="[1021,1125,1724,1749]" pageId="32" pageNumber="33" type="holotype">Holotype</typeStatus>
\ 33185. The four males
<typeStatus id="54D7883BFFD3F320FF3097B4D23BF926" box="[151,258,1761,1785]" pageId="32" pageNumber="33" type="paratype">paratypes</typeStatus>
deposited at
<collectionCode id="ED7DAE5CFFD3F320FE3C97B4D2E2F926" box="[411,475,1761,1785]" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="32" pageNumber="33">ROM</collectionCode>
are in good condition and bear the labels: Wm. Brodie \ Collection
<typeStatus id="54D7883BFFD3F320FB5C97B4D6B5F926" box="[1275,1420,1761,1785]" pageId="32" pageNumber="33" type="paratype">PARATYPE</typeStatus>
\
<taxonomicName id="4C6C4D1AFFD3F320FF309650D254F8C3" authorityName="Say" authorityYear="1824" box="[151,365,1796,1820]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="verticalis">
<emphasis id="B918EA8BFFD3F320FF309650D254F8C3" box="[151,365,1796,1820]" italics="true" pageId="32" pageNumber="33">Eumenes verticalis</emphasis>
</taxonomicName>
\
<taxonomicName id="4C6C4D1AFFD3F320FE209651D150F8C3" authority="Bequaert" authorityName="Bequaert" authorityYear="1944" box="[391,617,1796,1820]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="subSpecies" species="verticalis" subSpecies="neoboreus">
<emphasis id="B918EA8BFFD3F320FE209651D2C2F8C3" box="[391,507,1796,1820]" italics="true" pageId="32" pageNumber="33">neoboreus</emphasis>
Bequaert
</taxonomicName>
\ Det.
<bibRefCitation id="EFFD4B68FFD3F320FD1C9650D05BF8C2" box="[699,866,1797,1821]" pageId="32" pageNumber="33" refId="ref42355" refString="Bequaert, J. (1944) The genus Eumenes in the Dominion of Canada. Le Naturaliste Canadien, 71, 75 - 88." type="journal article">Bequaert 1944</bibRefCitation>
\ ROMEnt Spec.
<collectingRegion id="49A8F87BFFD3F320FB959650D761F8C3" box="[1074,1112,1797,1820]" country="Haiti" name="Nord" pageId="32" pageNumber="33">No</collectingRegion>
. 81631 \
<collectionCode id="ED7DAE5CFFD3F320FB639650D63DF8C2" box="[1220,1284,1797,1821]" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="32" pageNumber="33">ROM</collectionCode>
003001683; Toronto, Ont. \
<date id="FFD21059FFD3F320FEFA967DD28FF89F" box="[349,438,1832,1856]" pageId="32" pageNumber="33" value="94-07-19">19.7.94</date>
Wm. Brodie \ Collection [red square label]
<typeStatus id="54D7883BFFD3F320FC5A967DD7B7F89F" box="[1021,1166,1832,1856]" pageId="32" pageNumber="33" type="paratype">PARATYPE</typeStatus>
\
<taxonomicName id="4C6C4D1AFFD3F320FB09967CD6BEF89F" authorityName="Say" authorityYear="1824" box="[1198,1415,1832,1856]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="verticalis">
<emphasis id="B918EA8BFFD3F320FB09967CD6BEF89F" box="[1198,1415,1832,1856]" italics="true" pageId="32" pageNumber="33">Eumenes verticalis</emphasis>
</taxonomicName>
\
<taxonomicName id="4C6C4D1AFFD3F320FF309619D245F8BC" authority="Bequaert" authorityName="Bequaert" authorityYear="1944" box="[151,380,1868,1892]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="subSpecies" species="verticalis" subSpecies="neoboreus">
<emphasis id="B918EA8BFFD3F320FF309619D232F8BB" box="[151,267,1868,1892]" italics="true" pageId="32" pageNumber="33">neoboreus</emphasis>
Bequaert
</taxonomicName>
\ Det.
<bibRefCitation id="EFFD4B68FFD3F320FE7F9619D1B8F8BB" box="[472,641,1868,1892]" pageId="32" pageNumber="33" refId="ref42355" refString="Bequaert, J. (1944) The genus Eumenes in the Dominion of Canada. Le Naturaliste Canadien, 71, 75 - 88." type="journal article">Bequaert 1944</bibRefCitation>
\ ROMEnt Spec.
<collectingRegion id="49A8F87BFFD3F320FCF89619D0BCF8BC" box="[863,901,1868,1891]" country="Haiti" name="Nord" pageId="32" pageNumber="33">No</collectingRegion>
. 81633 \
<collectionCode id="ED7DAE5CFFD3F320FC5D9619D703F8BB" box="[1018,1082,1868,1892]" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="32" pageNumber="33">ROM</collectionCode>
004000144; Toronto, Ont. \
<date id="FFD21059FFD3F320FF309625D3C9F857" box="[151,240,1904,1928]" pageId="32" pageNumber="33" value="91-01-23">23.1.91</date>
Wm. Brodie \ Collection [red square label]
<typeStatus id="54D7883BFFD3F320FCB29625D09CF857" box="[789,933,1904,1928]" pageId="32" pageNumber="33" type="paratype">PARATYPE</typeStatus>
\
<taxonomicName id="4C6C4D1AFFD3F320FC1A9625D7ABF858" authorityName="Say" authorityYear="1824" box="[957,1170,1903,1927]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="verticalis">
<emphasis id="B918EA8BFFD3F320FC1A9625D7ABF858" box="[957,1170,1903,1927]" italics="true" pageId="32" pageNumber="33">Eumenes verticalis</emphasis>
</taxonomicName>
\
<taxonomicName id="4C6C4D1AFFD3F320FB0C963AD6B2F858" authority="Bequaert" authorityName="Bequaert" authorityYear="1944" box="[1195,1419,1903,1927]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="subSpecies" species="verticalis" subSpecies="neoboreus">
<emphasis id="B918EA8BFFD3F320FB0C963AD626F858" box="[1195,1311,1903,1927]" italics="true" pageId="32" pageNumber="33">neoboreus</emphasis>
Bequaert
</taxonomicName>
\ Det.
<bibRefCitation id="EFFD4B68FFD3F320FF7596C1D243F874" box="[210,378,1939,1963]" pageId="32" pageNumber="33" refId="ref42355" refString="Bequaert, J. (1944) The genus Eumenes in the Dominion of Canada. Le Naturaliste Canadien, 71, 75 - 88." type="journal article">Bequaert 1944</bibRefCitation>
\ ROMEnt Spec.
<collectingRegion id="49A8F87BFFD3F320FDF396C1D143F874" box="[596,634,1940,1963]" country="Haiti" name="Nord" pageId="32" pageNumber="33">No</collectingRegion>
. 81634 \
<collectionCode id="ED7DAE5CFFD3F320FD4A96C6D014F874" box="[749,813,1939,1963]" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="32" pageNumber="33">ROM</collectionCode>
004000145; Toronto, Ont. \
<date id="FFD21059FFD3F320FB3196C6D7D8F874" box="[1174,1249,1939,1963]" pageId="32" pageNumber="33" value="94-07-02">2.7.94</date>
Wm. Brodie \ Collection [red square label]
<typeStatus id="54D7883BFFD3F320FDAD96E2D1A3F810" box="[522,666,1975,1999]" pageId="32" pageNumber="33" type="paratype">PARATYPE</typeStatus>
\
<taxonomicName id="4C6C4D1AFFD3F320FD1196EDD0B7F810" authorityName="Say" authorityYear="1824" box="[694,910,1975,1999]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="verticalis">
<emphasis id="B918EA8BFFD3F320FD1196EDD0B7F810" box="[694,910,1975,1999]" italics="true" pageId="32" pageNumber="33">Eumenes verticalis</emphasis>
</taxonomicName>
\
<taxonomicName id="4C6C4D1AFFD3F320FC0D96E2D7B4F810" authority="Bequaert" authorityName="Bequaert" authorityYear="1944" box="[938,1165,1975,1999]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="subSpecies" species="verticalis" subSpecies="neoboreus">
<emphasis id="B918EA8BFFD3F320FC0D96E2D727F810" box="[938,1054,1975,1999]" italics="true" pageId="32" pageNumber="33">neoboreus</emphasis>
Bequaert
</taxonomicName>
\ Det.
<bibRefCitation id="EFFD4B68FFD3F320FB4496EDD6B3F810" box="[1251,1418,1975,1999]" pageId="32" pageNumber="33" refId="ref42355" refString="Bequaert, J. (1944) The genus Eumenes in the Dominion of Canada. Le Naturaliste Canadien, 71, 75 - 88." type="journal article">Bequaert 1944</bibRefCitation>
\ ROMEnt Spec.
<collectingRegion id="49A8F87BFFD3F320FEEE968ED257F82D" box="[329,366,2011,2034]" country="Haiti" name="Nord" pageId="32" pageNumber="33">No</collectingRegion>
. 81632 \
<collectionCode id="ED7DAE5CFFD3F320FE72968ED12CF82C" box="[469,533,2011,2035]" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="32" pageNumber="33">ROM</collectionCode>
004000143 [this specimen is actually
<taxonomicName id="4C6C4D1AFFD3F320FC6A968ED76AF82C" authorityName="Say" authorityYear="1824" box="[973,1107,2011,2035]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="fraternus">
<emphasis id="B918EA8BFFD3F320FC6A968ED76AF82C" box="[973,1107,2011,2035]" italics="true" pageId="32" pageNumber="33">E. fraternus</emphasis>
</taxonomicName>
].
</paragraph>
<paragraph id="8BD33699FFD2F321FF6091C2D06AFD77" blockId="33.[151,1437,151,824]" pageId="33" pageNumber="34">
The holotype female of
<taxonomicName id="4C6C4D1AFFD2F321FE7391CCD041FF6F" authority="Cresson, 1875" authorityName="Cresson" authorityYear="1875" box="[468,888,151,176]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="coloradensis">
<emphasis id="B918EA8BFFD2F321FE7391CCD1E9FF6F" box="[468,720,152,176]" italics="true" pageId="33" pageNumber="34">Eumenes coloradensis</emphasis>
Cresson, 1875
</taxonomicName>
is in good condition but it is lacking the F3F10 of the right antenna and bears the labels: Col. [Red Label] LectoTYPE \ 2728
<taxonomicName id="4C6C4D1AFFD2F321FB8E91E8D7B7FF0B" box="[1065,1166,189,212]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="B918EA8BFFD2F321FB8E91E8D7B7FF0B" box="[1065,1166,189,212]" italics="true" pageId="33" pageNumber="34">Eumenes</emphasis>
</taxonomicName>
\
<taxonomicName id="4C6C4D1AFFD2F321FB0491E9D60BFF0B" authorityName="Cresson" authorityYear="1875" box="[1187,1330,188,212]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="coloradensis">
<emphasis id="B918EA8BFFD2F321FB0491E9D60BFF0B" box="[1187,1330,188,212]" italics="true" pageId="33" pageNumber="34">coloradensis</emphasis>
</taxonomicName>
\ Cress.. The Lectotype female of
<taxonomicName id="4C6C4D1AFFD2F321FE1091B4D067FF27" authority="Viereck, 1908" authorityName="Viereck" authorityYear="1908" box="[439,862,223,248]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="cruciferorum">
<emphasis id="B918EA8BFFD2F321FE1091B4D18EFF27" box="[439,695,224,248]" italics="true" pageId="33" pageNumber="34">Eumenes cruciferorum</emphasis>
Viereck, 1908
</taxonomicName>
is lacking the fore and mid right legs, and the last two tarsi of the left fore leg. It bears the labels: Oak Creek Canon \ Ariz.
<quantity id="4C949B7CFFD2F321FC689051D718FEC2" box="[975,1057,260,285]" metricMagnitude="3" metricUnit="m" metricValue="1.8288" pageId="33" pageNumber="34" unit="ft" value="6000.0">6000 ft</quantity>
. Aug \ F. H. Snow 1708a [Red Label]
<taxonomicName id="4C6C4D1AFFD2F321FF40907CD275FE9F" box="[231,332,297,320]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="B918EA8BFFD2F321FF40907CD275FE9F" box="[231,332,297,320]" italics="true" pageId="33" pageNumber="34">Eumenes</emphasis>
</taxonomicName>
\
<taxonomicName id="4C6C4D1AFFD2F321FEC5907DD2CCFE9F" authorityName="Viereck" authorityYear="1908" box="[354,501,296,320]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="cruciferorum">
<emphasis id="B918EA8BFFD2F321FEC5907DD2CCFE9F" box="[354,501,296,320]" italics="true" pageId="33" pageNumber="34">cruciferorum</emphasis>
</taxonomicName>
\ Type Vier.. The paralectotype male of
<taxonomicName id="4C6C4D1AFFD2F321FC65907CD65CFE9F" authority="Viereck, 1908" authorityName="Viereck" authorityYear="1908" box="[962,1381,295,320]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="cruciferorum">
<emphasis id="B918EA8BFFD2F321FC65907CD7F8FE9F" box="[962,1217,296,320]" italics="true" pageId="33" pageNumber="34">Eumenes cruciferorum</emphasis>
Viereck, 1908
</taxonomicName>
is in bad condition, lacking: left antenna; left fore and hind legs (except for the coxa); left mid leg from tibia and beyond; right leg; right mid and hind legs from tibia and beyond; metasoma broken but still pinned with the specimen. Bears the labels: Oak Creek Canon \ Ariz.
<quantity id="4C949B7CFFD2F321FD5C90C1D077FE72" box="[763,846,404,429]" metricMagnitude="3" metricUnit="m" metricValue="1.8288" pageId="33" pageNumber="34" unit="ft" value="6000.0">6000 ft</quantity>
. Aug \ F. H. Snow 1708b [Red Label]
<taxonomicName id="4C6C4D1AFFD2F321FA8090C0D6B5FE73" box="[1319,1420,405,428]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="B918EA8BFFD2F321FA8090C0D6B5FE73" box="[1319,1420,405,428]" italics="true" pageId="33" pageNumber="34">Eumenes</emphasis>
</taxonomicName>
\
<taxonomicName id="4C6C4D1AFFD2F321FF3090EDD213FE0F" authorityName="Viereck" authorityYear="1908" box="[151,298,440,464]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="cruciferorum">
<emphasis id="B918EA8BFFD2F321FF3090EDD213FE0F" box="[151,298,440,464]" italics="true" pageId="33" pageNumber="34">cruciferorum</emphasis>
</taxonomicName>
\ Type Vier.. The lectotype male of
<taxonomicName id="4C6C4D1AFFD2F321FD6090ECD767FE0F" authority="Viereck, 1908" authorityName="Viereck" authorityYear="1908" box="[711,1118,439,464]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="enigmaticus">
<emphasis id="B918EA8BFFD2F321FD6090ECD083FE10" box="[711,954,440,464]" italics="true" pageId="33" pageNumber="34">Eumenes enigmaticus</emphasis>
Viereck, 1908
</taxonomicName>
is in good condition, except for the lacking of the third tarsus and beyond of the hind righ leg. Bears the labels: Oak Creek Canon \ Ariz.
<quantity id="4C949B7CFFD2F321FAC39089D395FDC7" metricMagnitude="3" metricUnit="m" metricValue="1.8288" pageId="33" pageNumber="34" unit="ft" value="6000.0">6000 ft</quantity>
. Aug \ F. H. Snow 1708c [Red Label]
<taxonomicName id="4C6C4D1AFFD2F321FD339354D1C0FDC7" box="[660,761,513,536]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="B918EA8BFFD2F321FD339354D1C0FDC7" box="[660,761,513,536]" italics="true" pageId="33" pageNumber="34">Eumenes</emphasis>
</taxonomicName>
\
<taxonomicName id="4C6C4D1AFFD2F321FCB49355D0A3FDC8" authorityName="Viereck" authorityYear="1908" box="[787,922,512,535]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="enigmaticus">
<emphasis id="B918EA8BFFD2F321FCB49355D0A3FDC8" box="[787,922,512,535]" italics="true" pageId="33" pageNumber="34">enigmaticus</emphasis>
</taxonomicName>
\ Paratype Vier.. The paralectotype male of
<taxonomicName id="4C6C4D1AFFD2F321FF309370D116FDE3" authority="Viereck, 1908" authorityName="Viereck" authorityYear="1908" box="[151,559,548,573]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="enigmaticus">
<emphasis id="B918EA8BFFD2F321FF309370D2B3FDE3" box="[151,394,549,572]" italics="true" pageId="33" pageNumber="34">Eumenes enigmaticus</emphasis>
Viereck, 1908
</taxonomicName>
is in excellent condition and bears the same labels as the lectotype, except for one: 1708 instead of 1708c. The holotype of
<taxonomicName id="4C6C4D1AFFD2F321FD09931CD722FDBF" authority="Isely, 1917" authorityName="Isely" authorityYear="1917" box="[686,1051,583,608]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="stenogaster">
<emphasis id="B918EA8BFFD2F321FD09931CD0A2FDBF" box="[686,923,585,608]" italics="true" pageId="33" pageNumber="34">Eumenes stenogaster</emphasis>
<bibRefCitation id="EFFD4B68FFD2F321FC069312D722FDBF" box="[929,1051,583,608]" pageId="33" pageNumber="34" refId="ref43377" refString="Isely, D. (1917) A synopsis of the petiolate wasps of the family Eumenidae (Hymenoptera), found in America North of Mexico. Annals of the Entomological Society of America, 10 (4), 345 - 366. https: // doi. org / 10.1093 / aesa / 10.4.345" type="journal article">Isely, 1917</bibRefCitation>
</taxonomicName>
is in excellent condition and bears the labels: Coll. \ Townsend White Mts \ 8-1 NM Rio Ruidoso \ Abt
<quantity id="4C949B7CFFD2F321FC6E9339D719FD5A" box="[969,1056,620,645]" metricMagnitude="3" metricUnit="m" metricValue="1.9812" pageId="33" pageNumber="34" unit="ft" value="6500.0">6500 ft</quantity>
. On fls
<taxonomicName id="4C6C4D1AFFD2F321FB209339D7D6FD5B" box="[1159,1263,620,644]" class="Insecta" family="Sphingidae" genus="Monarda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="B918EA8BFFD2F321FB209339D7D6FD5B" box="[1159,1263,620,644]" italics="true" pageId="33" pageNumber="34">Monarda</emphasis>
</taxonomicName>
\
<emphasis id="B918EA8BFFD2F321FAA39338D675FD5B" box="[1284,1356,621,644]" italics="true" pageId="33" pageNumber="34">stricta</emphasis>
[Red Label] Type \ No. 21379 \ U.S.N.M.
<emphasis id="B918EA8BFFD2F321FDE093C4D166FD77" box="[583,607,657,680]" italics="true" pageId="33" pageNumber="34">E.</emphasis>
\
<taxonomicName id="4C6C4D1AFFD2F321FDD393C4D1CCFD77" authorityName="Isely" authorityYear="1917" box="[628,757,657,680]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="stenogaster">
<emphasis id="B918EA8BFFD2F321FDD393C4D1CCFD77" box="[628,757,657,680]" italics="true" pageId="33" pageNumber="34">stenogaster</emphasis>
</taxonomicName>
\ Isely.
</paragraph>
<paragraph id="8BD33699FFD2F321FF6093E1D0C1FD2F" blockId="33.[151,1437,151,824]" pageId="33" pageNumber="34">
The holotype female of
<taxonomicName id="4C6C4D1AFFD2F321FE7493E0D191FD13" authorityName="Isely" authorityYear="1917" box="[467,680,693,716]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="tricinctus">
<emphasis id="B918EA8BFFD2F321FE7493E0D191FD13" box="[467,680,693,716]" italics="true" pageId="33" pageNumber="34">Eumenes tricinctus</emphasis>
</taxonomicName>
is in good condition and bear the labels: Oreg \ 2511 Collection \ CFBaker [Red Label] Type \ No. 21383 \ U.S.N.M.
<emphasis id="B918EA8BFFD2F321FCA3938CD024FD2F" box="[772,797,729,752]" italics="true" pageId="33" pageNumber="34">E.</emphasis>
\
<taxonomicName id="4C6C4D1AFFD2F321FC95938DD0A5FD30" authorityName="Isely" authorityYear="1917" box="[818,924,728,751]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="tricinctus">
<emphasis id="B918EA8BFFD2F321FC95938DD0A5FD30" box="[818,924,728,751]" italics="true" pageId="33" pageNumber="34">tricinctus</emphasis>
</taxonomicName>
\ Isely.
</paragraph>
<paragraph id="8BD33699FFD2F321FF6093A9D788FCE7" blockId="33.[151,1437,151,824]" pageId="33" pageNumber="34">
The holotype female of
<taxonomicName id="4C6C4D1AFFD2F321FE7493A8D1E9FCCB" authorityName="Giordani Soika" authorityYear="1978" box="[467,720,764,788]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="cruciferoides">
<emphasis id="B918EA8BFFD2F321FE7493A8D1E9FCCB" box="[467,720,764,788]" italics="true" pageId="33" pageNumber="34">Eumenes cruciferoides</emphasis>
</taxonomicName>
is in good condition and bears the labels: Nogares \ (Orizaba) \
<quantity id="4C949B7CFFD2F321FF30924AD3D6FCE8" box="[151,239,799,824]" metricMagnitude="3" metricUnit="m" metricValue="1.54" pageId="33" pageNumber="34" unit="m" value="1540.0">1540 m</quantity>
.
<date id="FFD21059FFD2F321FF5D924AD276FCE7" box="[250,335,799,824]" pageId="33" pageNumber="34" value="1975-09-01">1.IX.75</date>
\ Giordani Soika [Red Label] HOLOTYPE \
<taxonomicName id="4C6C4D1AFFD2F321FCC79274D0FCFCE7" box="[864,965,801,824]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="B918EA8BFFD2F321FCC79274D0FCFCE7" box="[864,965,801,824]" italics="true" pageId="33" pageNumber="34">Eumenes</emphasis>
</taxonomicName>
\
<taxonomicName id="4C6C4D1AFFD2F321FC7C9275D79EFCE7" authority="Sss." authorityName="Sss." box="[987,1191,800,824]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="cruciferoides">
<emphasis id="B918EA8BFFD2F321FC7C9275D757FCE7" box="[987,1134,800,824]" italics="true" pageId="33" pageNumber="34">cruciferoides</emphasis>
Sss.
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>