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<document id="5C86264B19D114830C09F9D5C78401CE" ID-CLB-Dataset="3383" ID-DOI="10.1093/zoolinnean/zly037" ID-GBIF-Dataset="d90d12b0-99f7-4d30-8eea-1f5364bf490c" ID-ISSN="0024-4082" ID-Zenodo-Dep="5715063" ID-ZooBank="03DAADC2-E954-42C5-B44D-55927DF55837" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1619628054066" checkinUser="carolina" docAuthor="Geroto, Caio Fabricio Cezar &amp; Bertini, Reinaldo J." docDate="2019" docId="03C35358FFC1936AFE8B3B89FD9E92B4" docLanguage="en" docName="ZoolJLinnSoc.185.312-334.pdf.imf" docOrigin="Zoological Journal of the Linnean Society 185" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Peirosaurinae Geroto &amp; Bertini 2019, SUBFAM. NOV." docType="treatment" docVersion="7" lastPageNumber="329" masterDocId="FFFA2B20FFD09378FF873E3FFFBE9013" masterDocTitle="New material of Pepesuchus (Crocodyliformes; Mesoeucrocodylia) from the Bauru Group: implications about its phylogeny and the age of the Adamantina Formation" masterLastPageNumber="334" masterPageNumber="312" pageNumber="329" updateTime="1698937884311" updateUser="ExternalLinkService">
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<mods:title id="F19B0A5DB719A74E58B65FCFD1E9F923">New material of Pepesuchus (Crocodyliformes; Mesoeucrocodylia) from the Bauru Group: implications about its phylogeny and the age of the Adamantina Formation</mods:title>
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<mods:name id="8B1D5FCB9141A9BDF9953E2291B34E06" type="personal">
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<mods:namePart id="0428E7CF36FF12ADA8B08DA9212A1DD7">Geroto, Caio Fabricio Cezar</mods:namePart>
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<mods:namePart id="FDED13135F1BA11796E4A8D84FCD4F61">Bertini, Reinaldo J.</mods:namePart>
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<mods:typeOfResource id="43B1ABF4F848E39104D0BBAFA492E8E9">text</mods:typeOfResource>
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<mods:title id="8E8E9F2013D3CAFAEC9D4C4A43ACAED6">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="74FF84D9307A78111A87F9C335EB4E09">2019</mods:date>
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<mods:number id="A0E08AD6FEFC6EA13A11B913485E6B87">2018-07-02</mods:number>
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<mods:detail id="72F19DAA592FCFD51E28DF176132DE89" type="volume">
<mods:number id="04B4165B2F6DD1FD1584D94858AE6AC1">185</mods:number>
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<mods:classification id="FD17866B088ED15F812BAB08E14009A8">journal article</mods:classification>
<mods:identifier id="08588B9E303314C17BBC63F1362C3C28" type="CLB-Dataset">3383</mods:identifier>
<mods:identifier id="FE2979DB4BB713EE07B59FF3ED96362E" type="DOI">10.1093/zoolinnean/zly037</mods:identifier>
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<mods:identifier id="37C758F30656042302CF01FC93FBB8D9" type="ISSN">0024-4082</mods:identifier>
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<treatment id="03C35358FFC1936AFE8B3B89FD9E92B4" ID-DOI="http://doi.org/10.5281/zenodo.5715079" ID-GBIF-Taxon="190310838" ID-Zenodo-Dep="5715079" LSID="urn:lsid:plazi:treatment:03C35358FFC1936AFE8B3B89FD9E92B4" httpUri="http://treatment.plazi.org/id/03C35358FFC1936AFE8B3B89FD9E92B4" lastPageId="18" lastPageNumber="329" pageId="17" pageNumber="329">
<subSubSection id="C370B1C5FFC19369FE8B3B89FEFA9601" pageId="17" pageNumber="329" type="nomenclature">
<paragraph id="8BD5E24EFFC19369FE8B3B89FDC395DD" blockId="17.[268,637,1462,1486]" box="[268,637,1462,1486]" pageId="17" pageNumber="329">
<heading id="D09D5522FFC19369FE8B3B89FDC395DD" box="[268,637,1462,1486]" centered="true" fontSize="9" level="2" pageId="17" pageNumber="329" reason="3">
<taxonomicName id="4C6A99CDFFC19369FE8B3B89FE7195DF" authority="Geroto &amp; Bertini, 2019" authorityName="Geroto &amp; Bertini" authorityYear="2019" box="[268,463,1462,1486]" class="Reptilia" family="Peirosauridae" kingdom="Animalia" order="Crocodylia" pageId="17" pageNumber="329" phylum="Chordata" rank="subFamily" status="subfam. nov." subFamily="Peirosaurinae">PEIROSAURINAE</taxonomicName>
<emphasis id="B91E3E5CFFC19369FE503B86FDC395DD" bold="true" box="[471,637,1462,1486]" pageId="17" pageNumber="329">
<taxonomicNameLabel id="A22D8327FFC19369FE503B86FDC395DD" box="[471,637,1462,1486]" pageId="17" pageNumber="329" rank="subFamily">SUBFAM. NOV.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BD5E24EFFC19369FF163BE1FEFA9601" blockId="17.[145,761,1502,1554]" pageId="17" pageNumber="329">
<uri id="FFFBEE4CFFC19369FF163BE1FEFA9601" pageId="17" pageNumber="329">
urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
</subSubSection>
<subSubSection id="C370B1C5FFC19369FF163817FBFF9278" pageId="17" pageNumber="329" type="diagnosis">
<paragraph id="8BD5E24EFFC19369FF163817FEAE962C" blockId="17.[145,762,1576,1906]" box="[145,272,1576,1599]" pageId="17" pageNumber="329">
<emphasis id="B91E3E5CFFC19369FF163817FEAE962C" box="[145,272,1576,1599]" italics="true" pageId="17" pageNumber="329">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BD5E24EFFC19369FF163877FACB91A2" blockId="17.[145,762,1576,1906]" lastBlockId="17.[809,1425,197,434]" pageId="17" pageNumber="329">
Crocodyliformes with a heavily sculpted mandible and skull (
<bibRefCitation id="EFFB9FBFFFC19369FE933859FE6A966F" author="Gasparini Z" box="[276,468,1638,1660]" pageId="17" pageNumber="329" pagination="317 - 329" refId="ref14720" refString="Gasparini Z. 1982. Una nueva familia de cocodrilos zifodontes cretacicos de America del Sur. Quinto Congresso Latinoamericano de Geologia, Actas IV. Buenos Aires, Argentina, 317 - 329." type="book chapter" year="1982">Gasparini, 1982</bibRefCitation>
;
<bibRefCitation id="EFFB9FBFFFC19369FE633859FD57966F" author="Gasparini Z &amp; Chiappe LM &amp; Fernandez M" box="[484,745,1638,1660]" pageId="17" pageNumber="329" pagination="316 - 333" refId="ref14755" refString="Gasparini Z, Chiappe LM, Fernandez M. 1991. A new Senonian peirosaurid (Crocodylomorpha) from Argentina and a synopsis of the South American Cretaceous crocodilians. Journal of Vertebrate Paleontology 11: 316 - 333." type="journal article" year="1991">
Gasparini
<emphasis id="B91E3E5CFFC19369FDE33858FD1F9668" box="[612,673,1638,1660]" italics="true" pageId="17" pageNumber="329">et al.</emphasis>
, 1991
</bibRefCitation>
); moderately high snout and a large wedged maxillary process inserted dorsolaterally between the premaxilla; a high maxillary notch; anterolaterally oriented external nares separated by the nasals; nasals lacking contact with the lacrimals; large and paired supraorbitals; broad and partly divided internal nares located between the palatine and pterygoid; a broad and anteroventrally inclined basisphenoid that is exposed only in occipital view; a supraoccipital that does not take up part of the cranial roof; the basioccipital has an elongated central crest and heavy lateral bumps; two to three teeth posterior to the anterior margin of the suborbital fenestra; subcircular teeth that are moderately compressed, with anterior and posterior serrated edges. Additionally,
<taxonomicName id="4C6A99CDFFC19369FBFE3F41FB449180" authorityName="Price" authorityYear="1955" box="[1145,1274,382,403]" class="Reptilia" family="Peirosauridae" genus="Peirosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="17" pageNumber="329" phylum="Chordata" rank="species" species="torminni">
<emphasis id="B91E3E5CFFC19369FBFE3F41FB449180" box="[1145,1274,382,403]" italics="true" pageId="17" pageNumber="329">P. torminni</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6A99CDFFC19369FA8F3F41FA2F9181" box="[1288,1425,381,403]" class="Reptilia" family="Peirosauridae" genus="Uberabasuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="17" pageNumber="329" phylum="Chordata" rank="species" species="terrificus">
<emphasis id="B91E3E5CFFC19369FA8F3F41FA2F9181" box="[1288,1425,381,403]" italics="true" pageId="17" pageNumber="329">U. terrificus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6A99CDFFC19369FCDC3FA3FB9291A2" authorityName="Carvalho, Vasconcellos &amp; Tavares" authorityYear="2007" box="[859,1068,412,433]" class="Reptilia" family="Peirosauridae" genus="Montealtosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="17" pageNumber="329" phylum="Chordata" rank="species" species="arrudacamposi">
<emphasis id="B91E3E5CFFC19369FCDC3FA3FB9291A2" box="[859,1068,412,433]" italics="true" pageId="17" pageNumber="329">M. arrudacamposi</emphasis>
</taxonomicName>
also have this nasal process.
</paragraph>
<paragraph id="8BD5E24EFFC19369FCAE3FF6FC1491F2" blockId="17.[809,1424,457,572]" box="[809,938,457,481]" pageId="17" pageNumber="329">
<emphasis id="B91E3E5CFFC19369FCAE3FF6FC1491F2" box="[809,938,457,481]" italics="true" pageId="17" pageNumber="329">Etymology</emphasis>
</paragraph>
<paragraph id="8BD5E24EFFC19369FCAE3FD6FCD1922F" blockId="17.[809,1424,457,572]" pageId="17" pageNumber="329">
<taxonomicName id="4C6A99CDFFC19369FCAE3FD6FC7691EC" authority="Geroto &amp; Bertini, 2019" authorityName="Geroto &amp; Bertini" authorityYear="2019" box="[809,968,489,511]" class="Reptilia" family="Peirosauridae" kingdom="Animalia" order="Crocodylia" pageId="17" pageNumber="329" phylum="Chordata" rank="subFamily" status="subfam. nov." subFamily="Peirosaurinae">Peirosaurinae</taxonomicName>
was established as the name of the subfamily based on the family
<taxonomicName id="4C6A99CDFFC19369FBE13C37FCD5922F" authority="(Gasparini, 1982)" authorityName="GASPARINI" authorityYear="1982" baseAuthorityName="Gasparini" baseAuthorityYear="1982" class="Reptilia" family="Peirosauridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="17" pageNumber="329" phylum="Chordata" rank="family">
Peirosauridae (
<bibRefCitation id="EFFB9FBFFFC19369FA903C37FCDE922F" author="Gasparini Z" pageId="17" pageNumber="329" pagination="317 - 329" refId="ref14720" refString="Gasparini Z. 1982. Una nueva familia de cocodrilos zifodontes cretacicos de America del Sur. Quinto Congresso Latinoamericano de Geologia, Actas IV. Buenos Aires, Argentina, 317 - 329." type="book chapter" year="1982">Gasparini, 1982</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="8BD5E24EFFC19369FCAE3C6CFBFF9278" blockId="17.[809,1425,595,895]" box="[809,1089,595,619]" pageId="17" pageNumber="329">
<emphasis id="B91E3E5CFFC19369FCAE3C6CFBFF9278" box="[809,1089,595,619]" italics="true" pageId="17" pageNumber="329">Phylogenetic definition</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C370B1C5FFC19369FCAE3C4AFC6F936C" pageId="17" pageNumber="329" type="reference_group">
<paragraph id="8BD5E24EFFC19369FCAE3C4AFC6F936C" blockId="17.[809,1425,595,895]" pageId="17" pageNumber="329">
<taxonomicName id="4C6A99CDFFC19369FCAE3C4AFAA89299" authority="Price, 1955" authorityName="Price" authorityYear="1955" box="[809,1302,628,650]" class="Reptilia" family="Peirosauridae" genus="Peirosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="17" pageNumber="329" phylum="Chordata" rank="species" species="torminni">
<emphasis id="B91E3E5CFFC19369FCAE3C4AFBE0929A" box="[809,1118,628,650]" italics="true" pageId="17" pageNumber="329">Peirosaurus torminni</emphasis>
<bibRefCitation id="EFFB9FBFFFC19369FBF63C4BFAA89299" author="Price LI" box="[1137,1302,628,650]" pageId="17" pageNumber="329" refId="ref15723" refString="Price LI. 1955. Novos Crocodilideos dos Arenitos da Serie Bauru, Cretaceo do Estado de Minas Gerasi. Anais da Academia Brasileira de Ciencias 27: 487 - 453." type="journal volume" year="1955">Price, 1955</bibRefCitation>
</taxonomicName>
and all Crocodyliformes that share a more recent common ancestor with
<taxonomicName id="4C6A99CDFFC19369FC593C8DFBD492D4" authorityName="Price" authorityYear="1955" box="[990,1130,690,711]" class="Reptilia" family="Peirosauridae" genus="Peirosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="17" pageNumber="329" phylum="Chordata" rank="species" species="torminni">
<emphasis id="B91E3E5CFFC19369FC593C8DFBD492D4" box="[990,1130,690,711]" italics="true" pageId="17" pageNumber="329">P. torminni</emphasis>
</taxonomicName>
than with
<taxonomicName id="4C6A99CDFFC19369FB7C3C8DFC4E92F5" authority="Woodward, 1896" authorityName="Woodward" authorityYear="1896" class="Reptilia" genus="Notosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="17" pageNumber="329" phylum="Chordata" rank="species" species="terrestris">
<emphasis id="B91E3E5CFFC19369FB7C3C8DFA2E92D4" box="[1275,1424,690,711]" italics="true" pageId="17" pageNumber="329">N. terrestris</emphasis>
<bibRefCitation id="EFFB9FBFFFC19369FCAE3CEFFC4E92F5" author="Woodward AS" box="[809,1008,720,742]" pageId="17" pageNumber="329" pagination="1 - 20" refId="ref16486" refString="Woodward AS. 1896. On two Mesozoic crocodilians, Notosuchus (genus novum) and Cynodontosuchus (genum novum) from the red sandstones of the Territory of Neuquen (Argentine Republic). Anales del Museo de La Plata, La Plata, Serie Paleontologia Argentina 4: 1 - 20." type="journal article" year="1896">Woodward, 1896</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C6A99CDFFC19369FC793CEFFAAB92F5" authority="Price, 1945" authorityName="Price" authorityYear="1945" box="[1022,1301,720,742]" class="Reptilia" family="Baurusuchidae" genus="Baurusuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="17" pageNumber="329" phylum="Chordata" rank="species" species="pachecoi">
<emphasis id="B91E3E5CFFC19369FC793CEFFB3D92F6" box="[1022,1155,720,741]" italics="true" pageId="17" pageNumber="329">B. pachecoi</emphasis>
<bibRefCitation id="EFFB9FBFFFC19369FB0A3CEFFAAB92F5" author="Price LI" box="[1165,1301,720,742]" pageId="17" pageNumber="329" pagination="1 - 9" refId="ref15697" refString="Price LI. 1945. A new reptil from the Cretaceous Brazil. Divisao de Geologia e Mineralogia Notas Preliminares e estudos 1 - 9." type="book chapter" year="1945">Price, 1945</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C6A99CDFFC19369FAA43CEFFC159317" authority="Price, 1950" authorityName="Price" authorityYear="1950" class="Reptilia" family="Sphagesauridae" genus="Sphagesaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="17" pageNumber="329" phylum="Chordata" rank="species" species="huenei">
<emphasis id="B91E3E5CFFC19369FAA43CEFFA2F92F6" box="[1315,1425,720,741]" italics="true" pageId="17" pageNumber="329">S. huenei</emphasis>
Price, 1950
</taxonomicName>
,
<taxonomicName id="4C6A99CDFFC19369FC303CD0FB0C9317" authority="Price, 1955" authorityName="Price" authorityYear="1955" box="[951,1202,750,772]" class="Reptilia" family="Itasuchidae" genus="Itasuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="17" pageNumber="329" phylum="Chordata" rank="species" species="jesuinoi">
<emphasis id="B91E3E5CFFC19369FC303CD0FB979317" box="[951,1065,751,772]" italics="true" pageId="17" pageNumber="329">I. jesuinoi</emphasis>
<bibRefCitation id="EFFB9FBFFFC19369FBB63CD1FB0C9317" author="Price LI" box="[1073,1202,750,772]" pageId="17" pageNumber="329" refId="ref15723" refString="Price LI. 1955. Novos Crocodilideos dos Arenitos da Serie Bauru, Cretaceo do Estado de Minas Gerasi. Anais da Academia Brasileira de Ciencias 27: 487 - 453." type="journal volume" year="1955">Price, 1955</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C6A99CDFFC19369FB383CD0FC079331" authority="Campos et al., 2011" authorityName="Campos" authorityYear="2011" class="Reptilia" family="Peirosauridae" genus="Pepesuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="17" pageNumber="329" phylum="Chordata" rank="species" species="deiseae">
<emphasis id="B91E3E5CFFC19369FB383CD0FA939310" box="[1215,1325,750,772]" italics="true" pageId="17" pageNumber="329">P. deiseae</emphasis>
<bibRefCitation id="EFFB9FBFFFC19369FAB33CD0FC079331" author="Campos DA &amp; Oliveira GR &amp; Figueiredo RG &amp; Riff D &amp; Azevedo SAK &amp; Carvalho LB &amp; Kellner AWA" pageId="17" pageNumber="329" pagination="317 - 327" refId="ref14304" refString="Campos DA, Oliveira GR, Figueiredo RG, Riff D, Azevedo SAK, Carvalho LB, Kellner AWA. 2011. On a new peirosaurid crocodyliform from the Upper Cretaceous, Bauru Group, southeastern Brazil. Anais da Academia Brasileira de Ciencias 83: 317 - 327." type="journal article" year="2011">
Campos
<emphasis id="B91E3E5CFFC19369FCAE3D31FCDF9331" box="[809,865,781,803]" italics="true" pageId="17" pageNumber="329">et al</emphasis>
., 2011
</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C6A99CDFFC19369FC4D3D31FA359331" authority="Iori &amp; Carvalho, 2012" authorityName="Iori &amp; Carvalho" authorityYear="2012" box="[970,1419,781,803]" class="Reptilia" family="Trematochampsidae" genus="Barreirosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="17" pageNumber="329" phylum="Chordata" rank="species" species="franciscoi">
<emphasis id="B91E3E5CFFC19369FC4D3D31FBD99331" box="[970,1127,781,803]" italics="true" pageId="17" pageNumber="329">B. franciscoi</emphasis>
Iori &amp; Carvalho, 2012
</taxonomicName>
,
<taxonomicName id="4C6A99CDFFC19369FCAE3D13FB979351" authority="Price, 1959" authorityName="Price" authorityYear="1959" box="[809,1065,812,834]" class="Reptilia" genus="Araripesuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="17" pageNumber="329" phylum="Chordata" rank="species" species="gomesii">
<emphasis id="B91E3E5CFFC19369FCAE3D13FC219352" box="[809,927,812,833]" italics="true" pageId="17" pageNumber="329">A. gomesii</emphasis>
<bibRefCitation id="EFFB9FBFFFC19369FC203D13FB979351" author="Price LI" box="[935,1065,812,834]" pageId="17" pageNumber="329" pagination="7 - 55" refId="ref15755" refString="Price LI. 1959. Sobre um novo crocodilideo notossuquido do Cretacico brasileiro. Boletim do Departamento Nacional de Producao Mineral, Divisao de Geologia e Mineralogia 188: 7 - 55." type="journal article" year="1959">Price, 1959</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C6A99CDFFC19369FBB13D13FA329352" authority="Simpson, 1937" authorityName="Simpson" authorityYear="1937" box="[1078,1420,812,834]" class="Reptilia" family="Sebecidae" genus="Sebecus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="17" pageNumber="329" phylum="Chordata" rank="species" species="icaeorhinus">
<emphasis id="B91E3E5CFFC19369FBB13D13FB679352" box="[1078,1241,812,833]" italics="true" pageId="17" pageNumber="329">S. icaeorhinus</emphasis>
<bibRefCitation id="EFFB9FBFFFC19369FB663D13FA329352" author="Simpson GG" box="[1249,1420,812,834]" pageId="17" pageNumber="329" pagination="1 - 20" refId="ref16226" refString="Simpson GG. 1937. An ancient eusuchian crocodile from Patagonia. American Museum Novitates. Published by the American Museum of Natural History of New York. 1 - 20." type="book chapter" year="1937">Simpson, 1937</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C6A99CDFFC19369FCAE3D74FB6A9373" authority="Carvalho &amp; Bertini, 1999" authorityName="Carvalho &amp; Bertini" authorityYear="1999" box="[809,1236,842,864]" class="Reptilia" family="Candidodontidae" genus="Mariliasuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="17" pageNumber="329" phylum="Chordata" rank="species" species="amarali">
<emphasis id="B91E3E5CFFC19369FCAE3D74FC17934C" box="[809,937,842,864]" italics="true" pageId="17" pageNumber="329">M. amarali</emphasis>
<bibRefCitation id="EFFB9FBFFFC19369FC373D75FB6A9373" author="Carvalho IS &amp; Bertini RJ" box="[944,1236,842,864]" pageId="17" pageNumber="329" pagination="83 - 105" refId="ref14355" refString="Carvalho IS, Bertini RJ. 1999. Mariliasuchus: um novo Crocodylomorpha (Notosuchia) do Cretaceo da Bacia Bauru, Brasil. Geologia Colombiana 24: 83 - 105." type="journal article" year="1999">Carvalho &amp; Bertini, 1999</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C6A99CDFFC19369FA963D74FC73936C" authority="Laurent, 1768" authorityName="Laurent" authorityYear="1768" class="Reptilia" family="Crocodylidae" genus="Crocodylus" kingdom="Animalia" order="Crocodylia" pageId="17" pageNumber="329" phylum="Chordata" rank="species" species="niloticus">
<emphasis id="B91E3E5CFFC19369FA963D74FA2F934C" box="[1297,1425,842,864]" italics="true" pageId="17" pageNumber="329">C. niloticus</emphasis>
Laurent, 1768
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C370B1C5FFC1936AFCAE3D9BFD9E92B4" lastPageId="18" lastPageNumber="330" pageId="17" pageNumber="329" type="discussion">
<paragraph id="8BD5E24EFFC19369FCAE3D9BFC1193A8" blockId="17.[809,1426,932,1906]" box="[809,943,932,955]" pageId="17" pageNumber="329">
<emphasis id="B91E3E5CFFC19369FCAE3D9BFC1193A8" box="[809,943,932,955]" italics="true" pageId="17" pageNumber="329">Discussion</emphasis>
</paragraph>
<paragraph id="8BD5E24EFFC19369FCAE3DFBFADC96E4" blockId="17.[809,1426,932,1906]" pageId="17" pageNumber="329">
<taxonomicName id="4C6A99CDFFC19369FCAE3DFBFC5C93CA" authority="Geroto &amp; Bertini, 2019" authorityName="Geroto &amp; Bertini" authorityYear="2019" box="[809,994,964,986]" class="Reptilia" family="Peirosauridae" kingdom="Animalia" order="Crocodylia" pageId="17" pageNumber="329" phylum="Chordata" rank="subFamily" status="subfam. nov." subFamily="Peirosaurinae">Peirosaurinae</taxonomicName>
(
<figureCitation id="1351FECBFFC19369FC7B3DFBFBF493CA" box="[1020,1098,964,986]" captionStart="Figure 9" captionStartId="16.[164,247,811,833]" captionTargetBox="[164,1442,197,771]" captionTargetId="figure-492@16.[163,1443,195,772]" captionTargetPageId="16" captionText="Figure 9. Calibrated cladogram for the consensus strictus for the three most parsimonious trees, depicting only Mesoeucrocodylia. For the complete cladogram, refer to Supplementary Information Data S3. Mesoeucrocodylia. For the complete cladogram, refer." figureDoi="http://doi.org/10.5281/zenodo.5715085" httpUri="https://zenodo.org/record/5715085/files/figure.png" pageId="17" pageNumber="329">Fig. 9</figureCitation>
; node
<figureCitation id="1351FECBFFC19369FB2A3DFBFB6493C9" box="[1197,1242,964,986]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStart-3="Figure 4" captionStart-4="Figure 5" captionStart-5="Figure 6" captionStartId-0="2.[164,242,1733,1755]" captionStartId-1="6.[162,241,1141,1163]" captionStartId-2="7.[146,224,1098,1120]" captionStartId-3="8.[164,242,1037,1059]" captionStartId-4="9.[145,225,1007,1029]" captionStartId-5="11.[145,225,1852,1874]" captionTargetBox-0="[228,1377,199,1691]" captionTargetBox-1="[164,1443,196,1099]" captionTargetBox-2="[147,1426,197,1056]" captionTargetBox-3="[164,1444,197,995]" captionTargetBox-4="[305,1265,198,966]" captionTargetBox-5="[305,1264,1004,1811]" captionTargetId-0="figure-97@2.[225,1380,195,1693]" captionTargetId-1="figure-384@6.[163,1443,195,1102]" captionTargetId-2="figure-379@7.[145,1428,195,1058]" captionTargetId-3="figure-425@8.[163,1446,195,997]" captionTargetId-4="figure-445@9.[303,1267,196,968]" captionTargetId-5="figure-432@11.[303,1266,1002,1813]" captionTargetPageId-0="2" captionTargetPageId-1="6" captionTargetPageId-2="7" captionTargetPageId-3="8" captionTargetPageId-4="9" captionTargetPageId-5="11" captionText-0="Figure 1. Map of the Bauru Group (modified from Fernandes &amp; Coimbra, 1996)." captionText-1="Figure 2. MCT 1723-R, anterior portion of the rostrum. A, D, dorsal view. B, E, ventral view. C, F, left lateral view. Grey areas are associated sediments; hatched areas are broken parts. Scale bars represent 5 cm." captionText-2="Figure 3. MCT 1723-R, rostrum.A, B, dorsal views. C, D, ventral views. Grey areas are associated sediments; hatched areas are broken parts. Scale bars represent 5 cm." captionText-3="Figure 4. MCT 1723-R, rostrum in lateral views. A, B, left lateral views. C, D, right lateral views. Grey areas are associated sediments; hatched areas are broken parts. Scale bars represent 5 cm." captionText-4="Figure 5. Palatal surfaces of the maxillary from A, Pepesuchus deiseae (MCT 1788-R), with an anterior portion of the palatines showing a tipping point, compared with B, MCT 1723-R, with a broad and round anterior portion. Grey areas are associated sediments; hatched areas are broken parts. Scale bars represent 5 cm." captionText-5="Figure 6. MCT 1723-R, mandible. A, B, distal views. C, D, ventral views. Grey areas are associated sediments; hatched areas are broken parts. Scale bars represent 5 cm." figureDoi-0="http://doi.org/10.5281/zenodo.5715067" figureDoi-1="http://doi.org/10.5281/zenodo.5715069" figureDoi-2="http://doi.org/10.5281/zenodo.5715071" figureDoi-3="http://doi.org/10.5281/zenodo.5715073" figureDoi-4="http://doi.org/10.5281/zenodo.5715075" figureDoi-5="http://doi.org/10.5281/zenodo.5715077" httpUri-0="https://zenodo.org/record/5715067/files/figure.png" httpUri-1="https://zenodo.org/record/5715069/files/figure.png" httpUri-2="https://zenodo.org/record/5715071/files/figure.png" httpUri-3="https://zenodo.org/record/5715073/files/figure.png" httpUri-4="https://zenodo.org/record/5715075/files/figure.png" httpUri-5="https://zenodo.org/record/5715077/files/figure.png" pageId="17" pageNumber="329">156</figureCitation>
; refer to the Supplementary Information Data S3 for the consensus tree with the numbered node) consists of
<taxonomicName id="4C6A99CDFFC19369FA923A3DFA349405" authorityName="Buffetaut" authorityYear="1994" box="[1301,1418,1025,1047]" class="Reptilia" family="Peirosauridae" genus="Hamadasuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="17" pageNumber="329" phylum="Chordata" rank="species" species="rebouli">
<emphasis id="B91E3E5CFFC19369FA923A3DFA349405" box="[1301,1418,1025,1047]" italics="true" pageId="17" pageNumber="329">H. rebouli</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6A99CDFFC19369FCAE3A1FFC609426" authorityName="Gasparini, Chiappe &amp; Fernandez" authorityYear="1991" box="[809,990,1056,1077]" class="Reptilia" family="Peirosauridae" genus="Lomasuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="17" pageNumber="329" phylum="Chordata" rank="species" species="palpebrosus">
<emphasis id="B91E3E5CFFC19369FCAE3A1FFC609426" box="[809,990,1056,1077]" italics="true" pageId="17" pageNumber="329">L. palpebrosus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6A99CDFFC19369FC763A1FFB779426" authorityName="Martinelli, Sertich, Garrido &amp; Praderio" authorityYear="2012" box="[1009,1225,1056,1077]" class="Reptilia" family="Peirosauridae" genus="Gasparinisuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="17" pageNumber="329" phylum="Chordata" rank="species" species="peirosauroides">
<emphasis id="B91E3E5CFFC19369FC763A1FFB779426" box="[1009,1225,1056,1077]" italics="true" pageId="17" pageNumber="329">G. peirosauroides</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6A99CDFFC19369FB5B3A1FFC3D9440" authorityName="Leardi &amp; Pol" authorityYear="2009" class="Reptilia" family="Peirosauridae" genus="Barcinosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="17" pageNumber="329" phylum="Chordata" rank="species" species="gradilis">
<emphasis id="B91E3E5CFFC19369FB5B3A1FFC3D9440" italics="true" pageId="17" pageNumber="329">Barcinosuchus gradilis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6A99CDFFC19369FC163A00FBAC9447" authorityName="Price" authorityYear="1955" box="[913,1042,1087,1108]" class="Reptilia" family="Peirosauridae" genus="Peirosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="17" pageNumber="329" phylum="Chordata" rank="species" species="torminni">
<emphasis id="B91E3E5CFFC19369FC163A00FBAC9447" box="[913,1042,1087,1108]" italics="true" pageId="17" pageNumber="329">P. torminni</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6A99CDFFC19369FBA73A00FB169447" box="[1056,1192,1087,1108]" class="Reptilia" family="Peirosauridae" genus="Uberabasuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="17" pageNumber="329" phylum="Chordata" rank="species" species="terrificus">
<emphasis id="B91E3E5CFFC19369FBA73A00FB169447" box="[1056,1192,1087,1108]" italics="true" pageId="17" pageNumber="329">U. terrificus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6A99CDFFC19369FB653A00FCE69460" authorityName="Carvalho, Vasconcellos &amp; Tavares" authorityYear="2007" class="Reptilia" family="Peirosauridae" genus="Montealtosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="17" pageNumber="329" phylum="Chordata" rank="species" species="arrudacamposi">
<emphasis id="B91E3E5CFFC19369FB653A00FCE69460" italics="true" pageId="17" pageNumber="329">M. arrudacamposi</emphasis>
</taxonomicName>
. A clade supported by 13 unambiguous synapomorphies was recovered based on: skull height in posterior view higher than wide (10.0); anteroposterior length of the premaxilla in relationship to the rostrum long (&gt; 30% of the total length of the rostrum) (11.1); a premaxilla with a wedge-like process of the maxilla on the lateral surface of the premaxillary–maxillary suture (13.1); perinarial fossa facing anterolaterally (20.2); a foramen on the posterior surface of the base of the postorbital bar (67.3); posterodorsal inclination of the external surface of the occipital portion of the squamosal (110.1); shape of the dentary near the mandibular symphysis, forming an acute angle (183.0); dorsal edge of the dentary in the lateral view with two concave festooned regions (195.2); ziphodont teeth (219.0); lateral or anterolateral external nares (248); a foramen in the perinarial depression of the premaxilla (249.1); a notch for hypertrophied mandibular caniniform teeth on the premaxilla–maxilla suture, with a medial bent border of the premaxilla and maxilla (252.1); and reduced participation of the maxilla at the anterior margin of the suborbital fenestra (264.1).
</paragraph>
<paragraph id="8BD5E24EFFC1936AFCC6393FFD9E92B4" blockId="17.[809,1426,932,1906]" lastBlockId="18.[163,779,197,679]" lastPageId="18" lastPageNumber="330" pageId="17" pageNumber="329">
The aim of this analysis was to provide some internal stability to
<taxonomicName id="4C6A99CDFFC19369FC363921FBEB9727" authorityName="GASPARINI" authorityYear="1982" box="[945,1109,1822,1844]" class="Reptilia" family="Peirosauridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="17" pageNumber="329" phylum="Chordata" rank="family">Peirosauridae</taxonomicName>
, thus both
<taxonomicName id="4C6A99CDFFC19369FB5C3921FA2F9720" authorityName="Buffetaut" authorityYear="1994" box="[1243,1425,1822,1843]" class="Reptilia" family="Peirosauridae" genus="Hamadasuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="17" pageNumber="329" phylum="Chordata" rank="genus">
<emphasis id="B91E3E5CFFC19369FB5C3921FA2F9720" box="[1243,1425,1822,1843]" italics="true" pageId="17" pageNumber="329">Hamadasuchus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6A99CDFFC19369FCE53902FB9F9741" box="[866,1057,1853,1874]" class="Reptilia" family="Peirosauridae" genus="Uberabasuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="17" pageNumber="329" phylum="Chordata" rank="genus">
<emphasis id="B91E3E5CFFC19369FCE53902FB9F9741" box="[866,1057,1853,1874]" italics="true" pageId="17" pageNumber="329">Uberabasuchus</emphasis>
</taxonomicName>
were not chosen as internal specifiers of the subfamily
<taxonomicName id="4C6A99CDFFC19369FBEA3963FAAA9761" authorityName="Geroto &amp; Bertini" authorityYear="2019" box="[1133,1300,1884,1906]" class="Reptilia" family="Peirosauridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="17" pageNumber="329" phylum="Chordata" rank="subFamily" subFamily="Peirosaurinae">Peirosaurinae</taxonomicName>
, given the problems related to specimens of these taxa. The
<typeStatus id="54D15CECFFC2936AFD543EFAFF6A90EA" pageId="18" pageNumber="330" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C6A99CDFFC2936AFF723EDBFE1990EA" authorityName="Buffetaut" authorityYear="1994" box="[245,423,228,249]" class="Reptilia" family="Peirosauridae" genus="Hamadasuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="18" pageNumber="330" phylum="Chordata" rank="genus">
<emphasis id="B91E3E5CFFC2936AFF723EDBFE1990EA" box="[245,423,228,249]" italics="true" pageId="18" pageNumber="330">Hamadasuchus</emphasis>
</taxonomicName>
has some differences compared with a more complete specimen described by
<bibRefCitation id="EFFB9FBFFFC2936AFD293F3CFEF89124" author="Larsson HCE &amp; Sues HD" pageId="18" pageNumber="330" pagination="533 - 567" refId="ref15191" refString="Larsson HCE, Sues HD. 2007. Cranial osteology and phylogenetic relationships of Hamadasuchus rebouli (Crocodyliformes: Mesoeucrocodylia) from the Cretaceous of Morocco. Zoological Journal of Linnean Society 149: 533 - 567." type="journal article" year="2007">Larsson &amp; Sues (2007)</bibRefCitation>
and the MNHN specimen (MRS 3101). A polytomy between
<taxonomicName id="4C6A99CDFFC2936AFE1A3F7FFD9F9146" authorityName="Price" authorityYear="1955" box="[413,545,320,341]" class="Reptilia" family="Peirosauridae" genus="Peirosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="18" pageNumber="330" phylum="Chordata" rank="species" species="torminni">
<emphasis id="B91E3E5CFFC2936AFE1A3F7FFD9F9146" box="[413,545,320,341]" italics="true" pageId="18" pageNumber="330">P. torminni</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6A99CDFFC2936AFDB63F7FFCB49146" authorityName="Carvalho, Vasconcellos &amp; Tavares" authorityYear="2007" box="[561,778,320,341]" class="Reptilia" family="Peirosauridae" genus="Montealtosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="18" pageNumber="330" phylum="Chordata" rank="species" species="arrudacamposi">
<emphasis id="B91E3E5CFFC2936AFDB63F7FFCB49146" box="[561,778,320,341]" italics="true" pageId="18" pageNumber="330">M. arrudacamposi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6A99CDFFC2936AFF533F60FEE69167" box="[212,344,351,372]" class="Reptilia" family="Peirosauridae" genus="Uberabasuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="18" pageNumber="330" phylum="Chordata" rank="species" species="terrificus">
<emphasis id="B91E3E5CFFC2936AFF533F60FEE69167" box="[212,344,351,372]" italics="true" pageId="18" pageNumber="330">U. terrificus</emphasis>
</taxonomicName>
is caused by an overlap of characteristics with
<taxonomicName id="4C6A99CDFFC2936AFE8B3F42FE059181" box="[268,443,381,402]" class="Reptilia" family="Peirosauridae" genus="Uberabasuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="18" pageNumber="330" phylum="Chordata" rank="genus">
<emphasis id="B91E3E5CFFC2936AFE8B3F42FE059181" box="[268,443,381,402]" italics="true" pageId="18" pageNumber="330">Uberabasuchus</emphasis>
</taxonomicName>
recovered in a different position within node 153.
<bibRefCitation id="EFFB9FBFFFC2936AFE253FA3FD1591A1" author="Martinelli AG &amp; Sertich JJW &amp; Garrido AC &amp; Praderio AM" box="[418,683,412,434]" pageId="18" pageNumber="330" pagination="191 - 200" refId="ref15371" refString="Martinelli AG, Sertich JJW, Garrido AC, Praderio AM. 2012. A new peirosaurid from the Upper Cretaceous of Argentina: implications for specimens referred to Peirosaurus torminni Price (Crocodyliformes: Peirosauridae). Cretaceous Research 37: 191 - 200." type="journal article" year="2012">
Martinelli
<emphasis id="B91E3E5CFFC2936AFDA73FA3FDE491A2" box="[544,602,412,433]" italics="true" pageId="18" pageNumber="330">et al.</emphasis>
(2012)
</bibRefCitation>
and
<bibRefCitation id="EFFB9FBFFFC2936AFD613FA3FE9691C3" author="Lio G &amp; Agnolin FL &amp; Valieri RJ &amp; Filippi L" pageId="18" pageNumber="330" pagination="835 - 841" refId="ref15290" refString="Lio G, Agnolin FL, Valieri RJ, Filippi L. 2015. A new peirosaurid (Crocodilyformes) from the Late Cretaceous (Turonian - Coniacian) of Patagonia, Argentina. Historical Biology 28: 835 - 841." type="journal article" year="2015">
Lio
<emphasis id="B91E3E5CFFC2936AFF243F84FF6491DC" box="[163,218,442,464]" italics="true" pageId="18" pageNumber="330">et al.</emphasis>
(2015)
</bibRefCitation>
proposed that
<taxonomicName id="4C6A99CDFFC2936AFE573F84FDF091C3" authorityName="Price" authorityYear="1955" box="[464,590,443,464]" class="Reptilia" family="Peirosauridae" genus="Peirosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="18" pageNumber="330" phylum="Chordata" rank="species" species="torminni">
<emphasis id="B91E3E5CFFC2936AFE573F84FDF091C3" box="[464,590,443,464]" italics="true" pageId="18" pageNumber="330">P. torminni</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6A99CDFFC2936AFD013F84FCB491C3" box="[646,778,443,464]" class="Reptilia" family="Peirosauridae" genus="Uberabasuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="18" pageNumber="330" phylum="Chordata" rank="species" species="terrificus">
<emphasis id="B91E3E5CFFC2936AFD013F84FCB491C3" box="[646,778,443,464]" italics="true" pageId="18" pageNumber="330">U. terrificus</emphasis>
</taxonomicName>
are synonymous based on morphological similarities, such as the premaxillary length relative to the rest of the rostrum. The clade is sustained by six non-ambiguous synapomorphies (node 153, Supplementary Information Data S3), hinting that the genus might be the same as the three taxa. That revision, however, is not the scope of this contribution.
</paragraph>
</subSubSection>
</treatment>
</document>
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