427 lines
62 KiB
XML
427 lines
62 KiB
XML
<document id="9D05B38981D39C2E1B1B7C9C9DE0B132" ID-CLB-Dataset="25493" ID-DOI="10.11646/zootaxa.4668.1.6" ID-GBIF-Dataset="3996d4b6-345e-4db4-a214-4729d5844552" ID-ISSN="1175-5326" ID-Zenodo-Dep="3449013" ID-ZooBank="4F3088B0-9803-45D3-8ECD-ED88C8141C57" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1568897344918" checkinUser="plazi" docAuthor="Chen, Mengzhen, Ma, Zijun & Tian, Mingyi" docDate="2019" docId="03C187C8FF8D8C78B5B2FDFDFBAEFC74" docLanguage="en" docName="zootaxa.4668.1.6.pdf" docOrigin="Zootaxa 4668 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Qianaphaenops (Sanwangius) rowselli Tian, n. sp." docType="treatment" docVersion="9" lastPageNumber="112" masterDocId="FFF8FFB0FF8F8C7FB525FF90FF8BFFBF" masterDocTitle="Discovery of a cavernicolous trechine species from the Sanwang Dong-Erwang Dong cave system, Wulong, Chongqing, southwestern China (Coleoptera Carabidae: Trechinae)" masterLastPageNumber="114" masterPageNumber="105" pageNumber="107" updateTime="1698755406474" updateUser="ExternalLinkService">
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<mods:title id="FA892CC3CECDF93E5BBBEEA42B27E1B6">Discovery of a cavernicolous trechine species from the Sanwang Dong-Erwang Dong cave system, Wulong, Chongqing, southwestern China (Coleoptera Carabidae: Trechinae)</mods:title>
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<mods:namePart id="0EB4F388F1BD3660035881EEC64C55CF">Chen, Mengzhen</mods:namePart>
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<mods:namePart id="C9BEFCC42FE627F4B2AA551F5DE397D1">Ma, Zijun</mods:namePart>
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<mods:namePart id="ADADA16CF55F66CDDC4563D5F1A84E6D">Tian, Mingyi</mods:namePart>
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<taxonomicName id="4C684D5DFF8D8C7DB5B2FDFDFD27FD38" ID-CoL="8M678" authority="Tian" authorityName="Tian" box="[151,684,621,648]" class="Insecta" family="Carabidae" genus="Qianaphaenops" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="107" phylum="Arthropoda" rank="species" species="rowselli" subGenus="Sanwangius">
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<emphasis id="B91CEACCFF8D8C7DB5B2FDFDFDE0FD37" bold="true" box="[151,619,621,648]" italics="true" pageId="2" pageNumber="107">Qianaphaenops (Sanwangius) rowselli</emphasis>
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Tian
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et Chen,
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<taxonomicNameLabel id="A22F57B7FF8D8C7DB606FDFDFCEEFD38" box="[803,869,621,647]" pageId="2" pageNumber="107" rank="species">n. sp.</taxonomicNameLabel>
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<paragraph id="8BD736DEFF8D8C7DB5B2FD01FE99FD14" blockId="2.[151,869,621,683]" box="[151,274,657,683]" pageId="2" pageNumber="107">
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(
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<figureCitation id="13532A5BFF8D8C7DB5BAFD01FE82FD14" box="[159,265,657,683]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStart-6="FIGURE 7" captionStartId-0="3.[151,250,1946,1971]" captionStartId-1="4.[151,250,1303,1328]" captionStartId-2="4.[151,250,1957,1982]" captionStartId-3="5.[151,250,1911,1936]" captionStartId-4="6.[151,250,1543,1568]" captionStartId-5="7.[151,250,1925,1950]" captionStartId-6="8.[151,250,1886,1911]" captionTargetBox-0="[323,1251,179,1918]" captionTargetBox-1="[345,1243,181,1261]" captionTargetBox-2="[321,1265,1386,1932]" captionTargetBox-3="[239,1357,209,1877]" captionTargetBox-4="[165,1425,193,1511]" captionTargetBox-5="[157,1430,1001,1894]" captionTargetBox-6="[151,1435,181,1850]" captionTargetId-0="figure@3.[323,1263,179,1919]" captionTargetId-1="figure@4.[345,1243,181,1277]" captionTargetId-2="figure@4.[321,1265,1386,1932]" captionTargetId-3="figure@5.[221,1367,181,1885]" captionTargetId-4="figure@6.[151,1435,181,1517]" captionTargetId-5="figure@7.[151,1435,993,1900]" captionTargetId-6="figure@8.[151,1435,181,1860]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="4" captionTargetPageId-3="5" captionTargetPageId-4="6" captionTargetPageId-5="7" captionTargetPageId-6="8" captionText-0="FIGURE 1 Habitus of Qianaphaenops (Sanwangius) rowselli Tian et Chen, n. subgen., n. sp., holotype, male." captionText-1="FIGURE 2 Qianaphaenops (Sanwangius) rowselli Tian et Chen, n. subgen., n. sp., A head (ventral), male, paratype B left elytron (chaetotaxy indicated by white points), male, paratype C ventrite VII, male, paratype." captionText-2="FIGURE 3 Male genitalia of Qianaphaenops (Sanwangius) rowselli Tian et Chen, n. subgen., n. sp., holotype A median lobe and parameres, lateral view B apical lobe, dorsal view." captionText-3="FIGURE 4 Larva of Qianaphaenops (Sanwangius) rowselli Tian et Chen, n. subgen., n. sp. A habitus, ventral view B head (cephalic capsule) and thoraces, dorsal view (FA: frontale, PA: parietal, PN: pronotum, MsN: mesonotum, MtN: metanotum) C head (cephalic capsule), ventral view D urogomphi and pygidium, dorsolateral view E right antenna, dorsal view F left maxilla, ventral view G tarsal claw of left middle leg, dorsal view, arrowhead indicating the small seta near base." captionText-4="FIGURE 5 Geographical positions of Sanwang Dong and Erwang Dong, the localities of Qianaphaenops (Sanwangius) rowselli Tian et Chen, n. subgen., n. sp. A map of China to show the site of Chongqing B map of Chonqing to show the site of Wulong Qu C map of Wulong Qu, to show the site of Sanwang Dong-Erwang Dong cave system D sites of Sanwang Dong (a) and Erwang Dong (b)" captionText-5="FIGURE 6 Simplified maps of Sanwang Dong and Erwang Dong, to show the site where the cave beetles were found and collected (in courtesy of Dr. Philip John Rowsell, Hong Meigui Cave Exploration Society)." captionText-6="FIGURE 7 Sanwang Dong-Erwang Dong cave system, type locality of Qianaphaenops (Sanwangius) rowselli Tian et Chen, n. subgen., n. sp. A entrance of Sanwang Dong B Qingkou Tiankeng C “Xiong Da” (right) and Tian MY in front of Erwang Dong D an adult in Erwang Dong E a larva in Erwang Dong." figureDoi-0="http://doi.org/10.5281/zenodo.3449017" figureDoi-1="http://doi.org/10.5281/zenodo.3449019" figureDoi-2="http://doi.org/10.5281/zenodo.3449023" figureDoi-3="http://doi.org/10.5281/zenodo.3449027" figureDoi-4="http://doi.org/10.5281/zenodo.3449031" figureDoi-5="http://doi.org/10.5281/zenodo.3449035" figureDoi-6="http://doi.org/10.5281/zenodo.3449037" httpUri-0="https://zenodo.org/record/3449017/files/figure.png" httpUri-1="https://zenodo.org/record/3449019/files/figure.png" httpUri-2="https://zenodo.org/record/3449023/files/figure.png" httpUri-3="https://zenodo.org/record/3449027/files/figure.png" httpUri-4="https://zenodo.org/record/3449031/files/figure.png" httpUri-5="https://zenodo.org/record/3449035/files/figure.png" httpUri-6="https://zenodo.org/record/3449037/files/figure.png" pageId="2" pageNumber="107">Figs. 1–7</figureCitation>
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)
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<emphasis id="B91CEACCFF8D8C7DB5B2FD49FEC3FD4C" bold="true" box="[151,328,728,755]" pageId="2" pageNumber="107">Type material.</emphasis>
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<materialsCitation id="3B003C83FF8D8C7DB477FD49FBD1FC84" ID-GBIF-Occurrence="2418078732" collectingDate="2018-08-12" collectingDateMax="2018-08-13" collectingDateMin="2018-08-12" collectionCode="SCAU" country="China" county="Cave Erwang Dong" elevation="930" latitude="29.588057" location="Cave Sanwang Dong" longLatPrecision="20" longitude="107.99638" municipality="Cave Erwang Dong" pageId="2" pageNumber="107" specimenCount="1" specimenCount-female="39" specimenCount-male="51" stateProvince="Chongqing Shi" typeStatus="holotype">
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male, investigation site 1,
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<location id="8EB76005FF8D8C7DB7D3FD49FC60FD4C" LSID="urn:lsid:plazi:treatment:03C187C8FF8D8C78B5B2FDFDFBAEFC74:8EB76005FF8D8C7DB7D3FD49FC60FD4C" box="[758,1003,729,755]" country="China" county="Cave Erwang Dong" latitude="29.588057" longLatPrecision="20" longitude="107.99638" municipality="Cave Erwang Dong" name="Cave Sanwang Dong" pageId="2" pageNumber="107" stateProvince="Chongqing Shi">Cave Sanwang Dong</location>
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,
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<location id="8EB76005FF8D8C7DB6DCFD49FB5CFD4C" LSID="urn:lsid:plazi:treatment:03C187C8FF8D8C78B5B2FDFDFBAEFC74:8EB76005FF8D8C7DB6DCFD49FB5CFD4C" box="[1017,1239,729,755]" country="China" county="Cave Erwang Dong" latitude="29.588057" longLatPrecision="20" longitude="107.99638" municipality="Cave Erwang Dong" name="Erlongdong village" pageId="2" pageNumber="107" stateProvince="Chongqing Shi">Erlongdong village</location>
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,
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<location id="8EB76005FF8D8C7DB1C0FD49FA13FD4C" LSID="urn:lsid:plazi:treatment:03C187C8FF8D8C78B5B2FDFDFBAEFC74:8EB76005FF8D8C7DB1C0FD49FA13FD4C" box="[1253,1432,729,755]" country="China" county="Cave Erwang Dong" latitude="29.588057" longLatPrecision="20" longitude="107.99638" municipality="Cave Erwang Dong" name="Houping Xiang" pageId="2" pageNumber="107" stateProvince="Chongqing Shi">Houping Xiang</location>
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,
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<location id="8EB76005FF8D8C7DB5B2FD6DFEABFCA8" LSID="urn:lsid:plazi:treatment:03C187C8FF8D8C78B5B2FDFDFBAEFC74:8EB76005FF8D8C7DB5B2FD6DFEABFCA8" box="[151,288,765,791]" country="China" county="Cave Erwang Dong" latitude="29.588057" longLatPrecision="20" longitude="107.99638" municipality="Cave Erwang Dong" name="Wulong Qu" pageId="2" pageNumber="107" stateProvince="Chongqing Shi">Wulong Qu</location>
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,
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<collectingRegion id="49ACF83CFF8D8C7DB40AFD6DFE54FCA8" box="[303,479,765,791]" country="China" name="Chongqing" pageId="2" pageNumber="107">Chongqing Shi</collectingRegion>
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, southwestern
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<collectingCountry id="F37F764EFF8D8C7DB7A8FD6DFD58FCA8" box="[653,723,765,791]" name="China" pageId="2" pageNumber="107">China</collectingCountry>
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,
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/
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,
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</quantity>
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,
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<date id="FFD6101EFF8D8C7DB148FD6DFA8DFCA8" box="[1133,1286,765,791]" pageId="2" pageNumber="107" value="2018-08-12">
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<collectingDate id="EF92E9F6FF8D8C7DB148FD6DFA8DFCA8" box="[1133,1286,765,791]" pageId="2" pageNumber="107" value="2018-08-12">VIII-12-2018</collectingDate>
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</date>
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, leg. “Xiong Da” (Philip John Rowsell),
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<location id="8EB76005FF8D8C7DB4F7FCB1FDEAFC84" LSID="urn:lsid:plazi:treatment:03C187C8FF8D8C78B5B2FDFDFBAEFC74:8EB76005FF8D8C7DB4F7FCB1FDEAFC84" box="[466,609,801,827]" country="China" county="Cave Erwang Dong" latitude="29.588057" longLatPrecision="20" longitude="107.99638" municipality="Cave Erwang Dong" name="Mingyi Tian" pageId="2" pageNumber="107" stateProvince="Chongqing Shi">Mingyi Tian</location>
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,
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<location id="8EB76005FF8D8C7DB74BFCB1FC8BFC84" LSID="urn:lsid:plazi:treatment:03C187C8FF8D8C78B5B2FDFDFBAEFC74:8EB76005FF8D8C7DB74BFCB1FC8BFC84" box="[622,768,801,827]" country="China" county="Cave Erwang Dong" latitude="29.588057" longLatPrecision="20" longitude="107.99638" municipality="Cave Erwang Dong" name="Jingli Cheng" pageId="2" pageNumber="107" stateProvince="Chongqing Shi">Jingli Cheng</location>
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,
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<location id="8EB76005FF8D8C7DB628FCB1FC08FC84" LSID="urn:lsid:plazi:treatment:03C187C8FF8D8C78B5B2FDFDFBAEFC74:8EB76005FF8D8C7DB628FCB1FC08FC84" box="[781,899,801,827]" country="China" county="Cave Erwang Dong" latitude="29.588057" longLatPrecision="20" longitude="107.99638" municipality="Cave Erwang Dong" name="Mengzhen" pageId="2" pageNumber="107" stateProvince="Chongqing Shi">Mengzhen</location>
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<taxonomicName id="4C684D5DFF8D8C7DB6AEFCB1FC4DFC84" authority="Chen & Ma & Tian, 2019" authorityName="Chen & Ma & Tian" authorityYear="2019" box="[907,966,801,827]" pageId="2" pageNumber="113" rank="subGenus" status="subgen. nov." subGenus="Chen">Chen</taxonomicName>
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&
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<location id="8EB76005FF8D8C7DB6C9FCB1FBD1FC84" LSID="urn:lsid:plazi:treatment:03C187C8FF8D8C78B5B2FDFDFBAEFC74:8EB76005FF8D8C7DB6C9FCB1FBD1FC84" box="[1004,1114,801,827]" country="China" county="Cave Erwang Dong" latitude="29.588057" longLatPrecision="20" longitude="107.99638" municipality="Cave Erwang Dong" name="Zijun Ma" pageId="2" pageNumber="107" stateProvince="Chongqing Shi">Zijun Ma</location>
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</materialsCitation>
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;
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:
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and
|
||
<specimenCount id="9D6EFD57FF8D8C7DB0A5FCB1FF7AFCE0" pageId="2" pageNumber="107" type="female">31 females</specimenCount>
|
||
, idem;
|
||
<specimenCount id="9D6EFD57FF8D8C7DB466FCD4FE2CFCE0" box="[323,423,836,863]" pageId="2" pageNumber="107" type="male">15 males</specimenCount>
|
||
and
|
||
<specimenCount id="9D6EFD57FF8D8C7DB4F8FCD5FDC7FCE0" box="[477,588,837,863]" pageId="2" pageNumber="107" type="female">3 females</specimenCount>
|
||
, investigation site 1,
|
||
<collectingMunicipality id="6BB3ACA4FF8D8C7DB61CFCD5FB93FCE0" box="[825,1048,837,863]" pageId="2" pageNumber="107">Cave Erwang Dong</collectingMunicipality>
|
||
,
|
||
<geoCoordinate id="EE5C5019FF8D8C7DB106FCD4FB23FCE0" box="[1059,1192,836,863]" degrees="29" direction="north" minutes="35" orientation="latitude" pageId="2" pageNumber="107" precision="15" seconds="17" value="29.588057">29°35′17″N</geoCoordinate>
|
||
/
|
||
<geoCoordinate id="EE5C5019FF8D8C7DB194FCD4FAC8FCE0" box="[1201,1347,836,863]" degrees="107" direction="east" minutes="59" orientation="longitude" pageId="2" pageNumber="107" precision="15" seconds="47" value="107.99638">107°59′47″E</geoCoordinate>
|
||
,
|
||
<quantity id="4C909B3BFF8D8C7DB06BFCD5FA1CFCE0" box="[1358,1431,837,863]" metricMagnitude="2" metricUnit="m" metricValue="8.8" pageId="2" pageNumber="107" unit="m" value="880.0">
|
||
<elevation id="0045D1EDFF8D8C7DB06BFCD5FA1CFCE0" box="[1358,1431,837,863]" metricMagnitude="2" metricUnit="m" metricValue="8.8" pageId="2" pageNumber="107" unit="m" value="880.0">880 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="FFD6101EFF8D8C7DB5B2FCF9FEBBFC3C" box="[151,304,873,899]" pageId="2" pageNumber="107" value="2018-08-13">
|
||
<collectingDate id="EF92E9F6FF8D8C7DB5B2FCF9FEBBFC3C" box="[151,304,873,899]" pageId="2" pageNumber="107" value="2018-08-13">VIII-13-2018</collectingDate>
|
||
</date>
|
||
, same collectors
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B003C83FF8D8C7DB4DAFCF9FE5FFC18" ID-GBIF-Occurrence="2418078731" collectingDate="2018-08-12" collectingDateMax="2018-08-13" collectingDateMin="2018-08-12" collectionCode="SCAU" country="China" county="Cave Erwang Dong" elevation="930" latitude="29.588057" location="Cave Erwang Dong" longLatPrecision="20" longitude="107.99638" municipality="Cave Erwang Dong" pageId="2" pageNumber="107" specimenCount="21" specimenCount-female="5" specimenCount-male="16" stateProvince="Chongqing" typeStatus="holotype">
|
||
<specimenCount id="9D6EFD57FF8D8C7DB4DAFCF9FDEEFC3C" box="[511,613,873,899]" pageId="2" pageNumber="107" type="male">16 males</specimenCount>
|
||
and
|
||
<specimenCount id="9D6EFD57FF8D8C7DB784FCF8FC99FC3C" box="[673,786,872,899]" pageId="2" pageNumber="107" type="female">5 females</specimenCount>
|
||
, investigation site 2,
|
||
<collectingCounty id="62B64E52FF8D8C7DB122FCF9FB67FC3C" box="[1031,1260,873,899]" pageId="2" pageNumber="107">Cave Erwang Dong</collectingCounty>
|
||
, same date and collectors. All are in
|
||
<collectionCode id="ED79AE1BFF8D8C7DB4A4FC1DFE44FC18" box="[385,463,909,935]" pageId="2" pageNumber="107">SCAU</collectionCode>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3726555FF8D8C78B5E2FC20FF41FD80" lastPageId="7" lastPageNumber="112" pageId="2" pageNumber="107" type="description">
|
||
<paragraph id="8BD736DEFF8D8C7DB5E2FC20FD5AFC50" blockId="2.[151,1437,728,2015]" pageId="2" pageNumber="107">
|
||
<emphasis id="B91CEACCFF8D8C7DB5E2FC20FED2FC74" bold="true" box="[199,345,944,971]" pageId="2" pageNumber="107">Description.</emphasis>
|
||
Length:
|
||
<quantity id="4C909B3BFF8D8C7DB49AFC20FDCDFC74" box="[447,582,944,971]" metricMagnitude="-3" metricUnit="m" metricValue="6.05" metricValueMax="6.3" metricValueMin="5.8" pageId="2" pageNumber="107" unit="mm" value="6.05" valueMax="6.3" valueMin="5.8">5.8–6.3 mm</quantity>
|
||
(from apex of right mandibles to elytral apex) or
|
||
<quantity id="4C909B3BFF8D8C7DB152FC20FB74FC74" box="[1143,1279,944,971]" metricMagnitude="-3" metricUnit="m" metricValue="5.35" metricValueMax="5.5" metricValueMin="5.2" pageId="2" pageNumber="107" unit="mm" value="5.35" valueMax="5.5" valueMin="5.2">5.2–5.5 mm</quantity>
|
||
(from labrum to elytra); width:
|
||
<quantity id="4C909B3BFF8D8C7DB47FFC44FE6FFC50" box="[346,484,980,1007]" metricMagnitude="-3" metricUnit="m" metricValue="1.55" metricValueMax="1.6" metricValueMin="1.5" pageId="2" pageNumber="107" unit="mm" value="1.55" valueMax="1.6" valueMin="1.5">1.5–1.6 mm</quantity>
|
||
. Habitus as in
|
||
<figureCitation id="13532A5BFF8D8C7DB7ADFC45FD40FC50" box="[648,715,981,1007]" captionStart="FIGURE 1" captionStartId="3.[151,250,1946,1971]" captionTargetBox="[323,1251,179,1918]" captionTargetId="figure@3.[323,1263,179,1919]" captionTargetPageId="3" captionText="FIGURE 1 Habitus of Qianaphaenops (Sanwangius) rowselli Tian et Chen, n. subgen., n. sp., holotype, male." figureDoi="http://doi.org/10.5281/zenodo.3449017" httpUri="https://zenodo.org/record/3449017/files/figure.png" pageId="2" pageNumber="107">Fig. 1</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BD736DEFF8D8C7DB5E2FC69FF7EFB1C" blockId="2.[151,1437,728,2015]" pageId="2" pageNumber="107">Body yellowish brown, but pale on apical joint of antennae, mouthparts palps and tarsi; moderately shiny and whole body including abdominal ventrites glabrous though head with several short hairs on genae, and legs pubescent; microsculptural engraved meshes moderately transverse near base of head, transverse striate on frons and vertex of head, pronotum and elytra; fore body much shorter than elytra, (HmL+PL) / EL = 0.808, (HlL+PrL) / EL = 0.693.</paragraph>
|
||
<paragraph id="8BD736DEFF8D8C7DB5E2FB3DFAA8F9EC" blockId="2.[151,1437,728,2015]" pageId="2" pageNumber="107">
|
||
Head elongate, much longer than wide, HLm/HW = 1.96–1.97, HLl/HW = 1.70–1.71; genae fairly developed, nearly parallel-sided, widest at about middle of head from neck to apical margin of labrum; frons and vertex convex, frontal furrows well-marked, very long, widely and strongly divergent posteriad, ended behind posterior supraorbital pores; clypeus transverse, quadrisetose; labrum transverse, frontal margin slightly emarginate, 6-setose; two pairs of supraorbital setiferous pores present at about 3/7 and 3/4 of head from labrum; labial suture clearly visible, mentum bisetose on either side of tooth, mental tooth short and blunt at apex, basal fovea widely concave; submentum 6-setose; both maxillary and labial palps moderately subcylindrical, glabrous except for the 2
|
||
<superScript id="7C1D9B96FF8D8C7DB15BFA14FB05FA2D" attach="left" box="[1150,1166,1412,1426]" fontSize="6" pageId="2" pageNumber="107">nd</superScript>
|
||
labial palpomere which is bisetose on inner margin, and with an additional one at subapex; the 2
|
||
<superScript id="7C1D9B96FF8D8C7DB6E5FA38FC5BFA09" attach="left" box="[960,976,1448,1462]" fontSize="6" pageId="2" pageNumber="107">nd</superScript>
|
||
labial palpomere slightly longer than the 3
|
||
<superScript id="7C1D9B96FF8D8C7DB580FA5CFF38FA65" attach="none" box="[165,179,1484,1498]" fontSize="6" pageId="2" pageNumber="107">rd</superScript>
|
||
; the 3
|
||
<superScript id="7C1D9B96FF8D8C7DB5DCFA5CFE8CFA65" attach="left" box="[249,263,1484,1498]" fontSize="6" pageId="2" pageNumber="107">rd</superScript>
|
||
maxillary palpomere slightly longer than the 4
|
||
<superScript id="7C1D9B96FF8D8C7DB630FA5CFCA9FA65" attach="none" box="[789,802,1484,1498]" fontSize="6" pageId="2" pageNumber="107">th</superScript>
|
||
; suborbital pore just between base and submentum (
|
||
<figureCitation id="13532A5BFF8D8C7DB04AFA5DFF36F9B4" captionStart="FIGURE 2" captionStartId="4.[151,250,1303,1328]" captionTargetBox="[345,1243,181,1261]" captionTargetId="figure@4.[345,1243,181,1277]" captionTargetPageId="4" captionText="FIGURE 2 Qianaphaenops (Sanwangius) rowselli Tian et Chen, n. subgen., n. sp., A head (ventral), male, paratype B left elytron (chaetotaxy indicated by white points), male, paratype C ventrite VII, male, paratype." figureDoi="http://doi.org/10.5281/zenodo.3449019" httpUri="https://zenodo.org/record/3449019/files/figure.png" pageId="2" pageNumber="107">Fig. 2A</figureCitation>
|
||
). Antennae thin and rather long, but extending only to the site of preapical pores of elytra; the 1
|
||
<superScript id="7C1D9B96FF8D8C7DB1DCFA60FA8FFA41" attach="left" box="[1273,1284,1520,1534]" fontSize="6" pageId="2" pageNumber="107">st</superScript>
|
||
antennomere stouter, as long as the 2
|
||
<superScript id="7C1D9B96FF8D8C7DB4B8F984FE26F99D" attach="none" box="[413,429,1556,1570]" fontSize="6" pageId="2" pageNumber="107">nd</superScript>
|
||
, the 5
|
||
<superScript id="7C1D9B96FF8D8C7DB4D6F984FD8BF99D" attach="left" box="[499,512,1556,1570]" fontSize="6" pageId="2" pageNumber="107">th</superScript>
|
||
longest, the 11
|
||
<superScript id="7C1D9B96FF8D8C7DB78FF984FD3CF99D" attach="left" box="[682,695,1556,1570]" fontSize="6" pageId="2" pageNumber="107">th</superScript>
|
||
longer than 10
|
||
<superScript id="7C1D9B96FF8D8C7DB645F984FCE6F99D" attach="none" box="[864,877,1556,1570]" fontSize="6" pageId="2" pageNumber="107">th</superScript>
|
||
, as long as the 7
|
||
<superScript id="7C1D9B96FF8D8C7DB10DF984FBBEF99D" attach="none" box="[1064,1077,1556,1570]" fontSize="6" pageId="2" pageNumber="107">th</superScript>
|
||
, the comparative length of each antennomere from the 1
|
||
<superScript id="7C1D9B96FF8D8C7DB486F9A8FE25F9F9" attach="left" box="[419,430,1592,1606]" fontSize="6" pageId="2" pageNumber="107">st</superScript>
|
||
to 11
|
||
<superScript id="7C1D9B96FF8D8C7DB4C8F9A8FE71F9F9" attach="left" box="[493,506,1592,1606]" fontSize="6" pageId="2" pageNumber="107">th</superScript>
|
||
as: 1.00 / 1.00 / 1.54 / 1.73 / 1.85 / 1.73 / 1.58 / 1.46 / 1.35 / 1.19 / 1.50.
|
||
</paragraph>
|
||
<paragraph id="8BD736DEFF8D8C7DB5E2F9CDFDB8F8B8" blockId="2.[151,1437,728,2015]" pageId="2" pageNumber="107">Pronotum subquadrate, as long as wide, wider than head, PW/HW = 1.16–1.21, slightly shorter than head excluding mandibles, widest a little before middle, gently narrowed anteriorly and posteriorly, latero-margins finely bordered; front and base slightly concave and convex respectively, both unbordered; fore latero-marginal setae at about apical 1/8, hind ones on hind angles; disc moderately convex; median line clear, basal transversal impression well-marked. Scutellum fairly small.</paragraph>
|
||
<paragraph id="8BD736DEFF8D8C7DB5E2F881FAE5F860" blockId="2.[151,1437,728,2015]" pageId="2" pageNumber="107">
|
||
Elytra (
|
||
<figureCitation id="13532A5BFF8D8C7DB43CF881FED4F894" box="[281,351,1809,1835]" captionStart="FIGURE 1" captionStartId="3.[151,250,1946,1971]" captionTargetBox="[323,1251,179,1918]" captionTargetId="figure@3.[323,1263,179,1919]" captionTargetPageId="3" captionText="FIGURE 1 Habitus of Qianaphaenops (Sanwangius) rowselli Tian et Chen, n. subgen., n. sp., holotype, male." figureDoi="http://doi.org/10.5281/zenodo.3449017" httpUri="https://zenodo.org/record/3449017/files/figure.png" pageId="2" pageNumber="107">Figs 1</figureCitation>
|
||
,
|
||
<figureCitation id="13532A5BFF8D8C7DB44EF881FE04F894" box="[363,399,1809,1835]" captionStart="FIGURE 2" captionStartId="4.[151,250,1303,1328]" captionTargetBox="[345,1243,181,1261]" captionTargetId="figure@4.[345,1243,181,1277]" captionTargetPageId="4" captionText="FIGURE 2 Qianaphaenops (Sanwangius) rowselli Tian et Chen, n. subgen., n. sp., A head (ventral), male, paratype B left elytron (chaetotaxy indicated by white points), male, paratype C ventrite VII, male, paratype." figureDoi="http://doi.org/10.5281/zenodo.3449019" httpUri="https://zenodo.org/record/3449019/files/figure.png" pageId="2" pageNumber="107">2B</figureCitation>
|
||
), elongated ovate, much longer than wide, EL/EW = 1.61–1.65; longer than prothorax, EL/PL = 3.17–3.38, much wider than prothorax, EW/PW = 2.04–2.12; widest at about 3/7 of elytra from base, lateral sides smooth, finely bordered throughout, marginal gutters well-marked; slightly depressed near base, disc moderately convex; striae well-defined and punctate, intervals moderately convex. Chaetotaxy (
|
||
<figureCitation id="13532A5BFF8D8C7DB173F8EDFB3AF828" box="[1110,1201,1917,1943]" captionStart="FIGURE 2" captionStartId="4.[151,250,1303,1328]" captionTargetBox="[345,1243,181,1261]" captionTargetId="figure@4.[345,1243,181,1277]" captionTargetPageId="4" captionText="FIGURE 2 Qianaphaenops (Sanwangius) rowselli Tian et Chen, n. subgen., n. sp., A head (ventral), male, paratype B left elytron (chaetotaxy indicated by white points), male, paratype C ventrite VII, male, paratype." figureDoi="http://doi.org/10.5281/zenodo.3449019" httpUri="https://zenodo.org/record/3449019/files/figure.png" pageId="2" pageNumber="107">Fig. 2B</figureCitation>
|
||
): dorsal pores of the 3
|
||
<superScript id="7C1D9B96FF8D8C7DB580F830FF38F811" attach="left" box="[165,179,1952,1966]" fontSize="6" pageId="2" pageNumber="107">rd</superScript>
|
||
stria at about 1/5 and 3/7 of elytra from base respectively; the preapical pore at about apical 1/8 of elytra, much closer to elytral suture than to apical margin; basal pore present; the marginal umbilicate pores not aggregated.
|
||
</paragraph>
|
||
<caption id="DF176656FF8C8C7CB5B2F80AFB74F80C" ID-DOI="http://doi.org/10.5281/zenodo.3449017" ID-Zenodo-Dep="3449017" box="[151,1279,1946,1971]" httpUri="https://zenodo.org/record/3449017/files/figure.png" pageId="3" pageNumber="108" startId="3.[151,250,1946,1971]" targetBox="[323,1251,179,1918]" targetPageId="3">
|
||
<paragraph id="8BD736DEFF8C8C7CB5B2F80AFB74F80C" blockId="3.[151,1279,1946,1971]" box="[151,1279,1946,1971]" pageId="3" pageNumber="108">
|
||
<emphasis id="B91CEACCFF8C8C7CB5B2F80AFE86F80C" bold="true" box="[151,269,1946,1971]" pageId="3" pageNumber="108">FIGURE 1</emphasis>
|
||
Habitus of
|
||
<taxonomicName id="4C684D5DFF8C8C7CB4A1F80AFCCAF80C" authority="Tian" authorityName="Tian" box="[388,833,1946,1971]" class="Insecta" family="Carabidae" genus="Qianaphaenops" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="108" phylum="Arthropoda" rank="species" species="rowselli" subGenus="Sanwangius">
|
||
<emphasis id="B91CEACCFF8C8C7CB4A1F80AFC86F80D" box="[388,781,1946,1970]" italics="true" pageId="3" pageNumber="108">Qianaphaenops (Sanwangius) rowselli</emphasis>
|
||
Tian
|
||
</taxonomicName>
|
||
et
|
||
<taxonomicName id="4C684D5DFF8C8C7CB67AF80AFC1CF80C" authority="Chen & Ma & Tian, 2019" authorityName="Chen & Ma & Tian" authorityYear="2019" box="[863,919,1946,1971]" pageId="3" pageNumber="108" rank="subGenus" status="subgen. nov." subGenus="Chen">Chen</taxonomicName>
|
||
,
|
||
<emphasis id="B91CEACCFF8C8C7CB684F80AFBDFF80D" bold="true" box="[929,1108,1946,1970]" pageId="3" pageNumber="108">
|
||
<taxonomicNameLabel id="A22F57B7FF8C8C7CB684F80AFBDFF80D" box="[929,1108,1946,1970]" pageId="3" pageNumber="108" rank="species">n. subgen., n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype, male.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF176656FF8B8C7BB5B2FA87FBC1FAEB" ID-DOI="http://doi.org/10.5281/zenodo.3449019" ID-Zenodo-Dep="3449019" httpUri="https://zenodo.org/record/3449019/files/figure.png" pageId="4" pageNumber="109" startId="4.[151,250,1303,1328]" targetBox="[345,1243,181,1261]" targetPageId="4">
|
||
<paragraph id="8BD736DEFF8B8C7BB5B2FA87FBC1FAEB" blockId="4.[151,1436,1303,1364]" pageId="4" pageNumber="109">
|
||
<emphasis id="B91CEACCFF8B8C7BB5B2FA87FE84FA8F" bold="true" box="[151,271,1303,1328]" pageId="4" pageNumber="109">FIGURE 2</emphasis>
|
||
<taxonomicName id="4C684D5DFF8B8C7BB432FA87FD50FA8F" authority="Tian" authorityName="Tian" box="[279,731,1303,1328]" class="Insecta" family="Carabidae" genus="Qianaphaenops" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="109" phylum="Arthropoda" rank="species" species="rowselli" subGenus="Sanwangius">
|
||
<emphasis id="B91CEACCFF8B8C7BB432FA87FD2EFA90" box="[279,677,1303,1327]" italics="true" pageId="4" pageNumber="109">Qianaphaenops (Sanwangius) rowselli</emphasis>
|
||
Tian
|
||
</taxonomicName>
|
||
et
|
||
<taxonomicName id="4C684D5DFF8B8C7BB7D8FA87FCBEFA8F" authority="Chen & Ma & Tian, 2019" authorityName="Chen & Ma & Tian" authorityYear="2019" box="[765,821,1303,1328]" pageId="4" pageNumber="109" rank="subGenus" status="subgen. nov." subGenus="Chen">Chen</taxonomicName>
|
||
,
|
||
<emphasis id="B91CEACCFF8B8C7BB667FA88FC70FA8F" bold="true" box="[834,1019,1304,1328]" pageId="4" pageNumber="109">
|
||
<taxonomicNameLabel id="A22F57B7FF8B8C7BB667FA88FC70FA8F" box="[834,1019,1304,1328]" pageId="4" pageNumber="109" rank="species">n. subgen., n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<emphasis id="B91CEACCFF8B8C7BB12FFA87FB90FA8F" bold="true" box="[1034,1051,1303,1328]" pageId="4" pageNumber="109">A</emphasis>
|
||
head (ventral), male, paratype
|
||
<emphasis id="B91CEACCFF8B8C7BB044FA88FAF9FA8F" bold="true" box="[1377,1394,1304,1328]" pageId="4" pageNumber="109">B</emphasis>
|
||
left elytron (chaetotaxy indicated by white points), male, paratype
|
||
<emphasis id="B91CEACCFF8B8C7BB637FAABFCAFFAEB" bold="true" box="[786,804,1339,1364]" pageId="4" pageNumber="109">C</emphasis>
|
||
ventrite VII, male, paratype.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF176656FF8B8C7BB5B2F835FD49F85D" ID-DOI="http://doi.org/10.5281/zenodo.3449023" ID-Zenodo-Dep="3449023" httpUri="https://zenodo.org/record/3449023/files/figure.png" pageId="4" pageNumber="109" startId="4.[151,250,1957,1982]" targetBox="[321,1265,1386,1932]" targetPageId="4">
|
||
<paragraph id="8BD736DEFF8B8C7BB5B2F835FD49F85D" blockId="4.[151,1437,1957,2018]" pageId="4" pageNumber="109">
|
||
<emphasis id="B91CEACCFF8B8C7BB5B2F835FE86F801" bold="true" box="[151,269,1957,1982]" pageId="4" pageNumber="109">FIGURE 3</emphasis>
|
||
Male genitalia of
|
||
<taxonomicName id="4C684D5DFF8B8C7BB4EDF836FC0EF801" authority="Tian" authorityName="Tian" box="[456,901,1957,1982]" class="Insecta" family="Carabidae" genus="Qianaphaenops" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="109" phylum="Arthropoda" rank="species" species="rowselli" subGenus="Sanwangius">
|
||
<emphasis id="B91CEACCFF8B8C7BB4EDF836FCD9F801" box="[456,850,1958,1982]" italics="true" pageId="4" pageNumber="109">Qianaphaenops (Sanwangius) rowselli</emphasis>
|
||
Tian
|
||
</taxonomicName>
|
||
et
|
||
<taxonomicName id="4C684D5DFF8B8C7BB681F835FC57F801" authority="Chen & Ma & Tian, 2019" authorityName="Chen & Ma & Tian" authorityYear="2019" box="[932,988,1957,1982]" pageId="4" pageNumber="109" rank="subGenus" status="subgen. nov." subGenus="Chen">Chen</taxonomicName>
|
||
,
|
||
<emphasis id="B91CEACCFF8B8C7BB6C2F836FB10F801" bold="true" box="[999,1179,1958,1982]" pageId="4" pageNumber="109">
|
||
<taxonomicNameLabel id="A22F57B7FF8B8C7BB6C2F836FB10F801" box="[999,1179,1958,1982]" pageId="4" pageNumber="109" rank="species">n. subgen., n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype
|
||
<emphasis id="B91CEACCFF8B8C7BB022F835FA92F801" bold="true" box="[1287,1305,1957,1982]" pageId="4" pageNumber="109">A</emphasis>
|
||
median lobe and parameres, lateral view
|
||
<emphasis id="B91CEACCFF8B8C7BB491F85AFE4EF85D" bold="true" box="[436,453,1994,2018]" pageId="4" pageNumber="109">B</emphasis>
|
||
apical lobe, dorsal view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF176656FF8A8C7AB5B2F8E7FB33F843" ID-DOI="http://doi.org/10.5281/zenodo.3449027" ID-Zenodo-Dep="3449027" httpUri="https://zenodo.org/record/3449027/files/figure.png" pageId="5" pageNumber="110" startId="5.[151,250,1911,1936]" targetBox="[239,1357,209,1877]" targetPageId="5">
|
||
<paragraph id="8BD736DEFF8A8C7AB5B2F8E7FB33F843" blockId="5.[151,1436,1911,2044]" pageId="5" pageNumber="110">
|
||
<emphasis id="B91CEACCFF8A8C7AB5B2F8E7FE84F82F" bold="true" box="[151,271,1911,1936]" pageId="5" pageNumber="110">FIGURE 4</emphasis>
|
||
Larva of
|
||
<taxonomicName id="4C684D5DFF8A8C7AB453F8E7FCB3F82F" authority="Tian" authorityName="Tian" box="[374,824,1911,1936]" class="Insecta" family="Carabidae" genus="Qianaphaenops" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="110" phylum="Arthropoda" rank="species" species="rowselli" subGenus="Sanwangius">
|
||
<emphasis id="B91CEACCFF8A8C7AB453F8E7FC88F830" box="[374,771,1911,1935]" italics="true" pageId="5" pageNumber="110">Qianaphaenops (Sanwangius) rowselli</emphasis>
|
||
Tian
|
||
</taxonomicName>
|
||
et
|
||
<taxonomicName id="4C684D5DFF8A8C7AB67CF8E7FC1AF82F" authority="Chen & Ma & Tian, 2019" authorityName="Chen & Ma & Tian" authorityYear="2019" box="[857,913,1911,1936]" pageId="5" pageNumber="110" rank="subGenus" status="subgen. nov." subGenus="Chen">Chen</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A22F57B7FF8A8C7AB6B8F8E8FBDFF82F" box="[925,1108,1912,1936]" pageId="5" pageNumber="110" rank="species">
|
||
<emphasis id="B91CEACCFF8A8C7AB6B8F8E8FB80F82F" bold="true" box="[925,1035,1912,1936]" pageId="5" pageNumber="110">n. subgen.</emphasis>
|
||
,
|
||
<emphasis id="B91CEACCFF8A8C7AB13FF8E8FBDFF82F" bold="true" box="[1050,1108,1912,1936]" pageId="5" pageNumber="110">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
<emphasis id="B91CEACCFF8A8C7AB179F8E7FBE6F82F" bold="true" box="[1116,1133,1911,1936]" pageId="5" pageNumber="110">A</emphasis>
|
||
habitus, ventral view
|
||
<emphasis id="B91CEACCFF8A8C7AB071F8E8FAEEF82F" bold="true" box="[1364,1381,1912,1936]" pageId="5" pageNumber="110">B</emphasis>
|
||
head (cephalic capsule) and thoraces, dorsal view (FA: frontale, PA: parietal, PN: pronotum, MsN: mesonotum, MtN: metanotum)
|
||
<emphasis id="B91CEACCFF8A8C7AB0AFF80BFA17F80B" bold="true" box="[1418,1436,1947,1972]" pageId="5" pageNumber="110">C</emphasis>
|
||
head (cephalic capsule), ventral view
|
||
<emphasis id="B91CEACCFF8A8C7AB736F850FDAEF867" bold="true" box="[531,549,1984,2008]" pageId="5" pageNumber="110">D</emphasis>
|
||
urogomphi and pygidium, dorsolateral view
|
||
<emphasis id="B91CEACCFF8A8C7AB6C9F850FC76F867" bold="true" box="[1004,1021,1984,2008]" pageId="5" pageNumber="110">E</emphasis>
|
||
right antenna, dorsal view
|
||
<emphasis id="B91CEACCFF8A8C7AB028F850FA97F867" bold="true" box="[1293,1308,1984,2008]" pageId="5" pageNumber="110">F</emphasis>
|
||
left maxilla, ventral view
|
||
<emphasis id="B91CEACCFF8A8C7AB43EF873FEA4F843" bold="true" box="[283,303,2019,2044]" pageId="5" pageNumber="110">G</emphasis>
|
||
tarsal claw of left middle leg, dorsal view, arrowhead indicating the small seta near base.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF176656FF898C79B5B2F997FEC2F933" ID-DOI="http://doi.org/10.5281/zenodo.3449031" ID-Zenodo-Dep="3449031" httpUri="https://zenodo.org/record/3449031/files/figure.png" pageId="6" pageNumber="111" startId="6.[151,250,1543,1568]" targetBox="[165,1425,193,1511]" targetPageId="6">
|
||
<paragraph id="8BD736DEFF898C79B5B2F997FEC2F933" blockId="6.[151,1437,1543,1677]" pageId="6" pageNumber="111">
|
||
<emphasis id="B91CEACCFF898C79B5B2F997FE87F99F" bold="true" box="[151,268,1543,1568]" pageId="6" pageNumber="111">FIGURE 5</emphasis>
|
||
Geographical positions of Sanwang Dong and Erwang Dong, the localities of
|
||
<taxonomicName id="4C684D5DFF898C79B13EF998FF53F9FB" authority="Tian" authorityName="Tian" class="Insecta" family="Carabidae" genus="Qianaphaenops" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="111" phylum="Arthropoda" rank="species" species="rowselli" subGenus="Sanwangius">
|
||
<emphasis id="B91CEACCFF898C79B13EF998FAC7F9A0" box="[1051,1356,1544,1568]" italics="true" pageId="6" pageNumber="111">Qianaphaenops (Sanwangius)</emphasis>
|
||
<emphasis id="B91CEACCFF898C79B074F998FF2EF9FB" italics="true" pageId="6" pageNumber="111">rowselli</emphasis>
|
||
Tian
|
||
</taxonomicName>
|
||
et
|
||
<taxonomicName id="4C684D5DFF898C79B5D0F9BBFEA6F9FB" authority="Chen & Ma & Tian, 2019" authorityName="Chen & Ma & Tian" authorityYear="2019" box="[245,301,1579,1604]" pageId="6" pageNumber="111" rank="subGenus" status="subgen. nov." subGenus="Chen">Chen</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A22F57B7FF898C79B412F9BCFE6CF9FB" box="[311,487,1580,1604]" pageId="6" pageNumber="111" rank="species">
|
||
<emphasis id="B91CEACCFF898C79B412F9BCFE29F9FB" bold="true" box="[311,418,1580,1604]" pageId="6" pageNumber="111">
|
||
<taxonomicNameLabel id="A22F57B7FF898C79B412F9BCFE29F9FB" box="[311,418,1580,1604]" pageId="6" pageNumber="111" rank="subGenus">n. subgen.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<emphasis id="B91CEACCFF898C79B48AF9BCFE6CF9FB" bold="true" box="[431,487,1580,1604]" pageId="6" pageNumber="111">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
<emphasis id="B91CEACCFF898C79B4C9F9BBFE75F9FB" bold="true" box="[492,510,1579,1604]" pageId="6" pageNumber="111">A</emphasis>
|
||
map of China to show the site of Chongqing
|
||
<emphasis id="B91CEACCFF898C79B6E1F9BCFC5EF9FB" bold="true" box="[964,981,1580,1604]" pageId="6" pageNumber="111">B</emphasis>
|
||
map of Chonqing to show the site of Wulong Qu
|
||
<emphasis id="B91CEACCFF898C79B59AF9DFFF5AF9D7" bold="true" box="[191,209,1615,1640]" pageId="6" pageNumber="111">C</emphasis>
|
||
map of Wulong Qu, to show the site of Sanwang Dong-Erwang Dong cave system
|
||
<emphasis id="B91CEACCFF898C79B11AF9C0FBDAF9D7" bold="true" box="[1087,1105,1616,1640]" pageId="6" pageNumber="111">D</emphasis>
|
||
sites of Sanwang Dong (a) and Erwang Dong (b)
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD736DEFF898C79B5E2F92BFDF0F8A3" blockId="6.[151,1436,1722,2037]" pageId="6" pageNumber="111">
|
||
Legs thin and elongate, bearing short pubescence; femora sparsely setose; fore tibia with a longitudinal furrow or sulcus dorsally; the 1
|
||
<superScript id="7C1D9B96FF898C79B486F94EFE25F953" attach="left" box="[419,430,1758,1772]" fontSize="6" pageId="6" pageNumber="111">st</superScript>
|
||
tarsomere shorter than, as long as, and slightly longer than the 2
|
||
<superScript id="7C1D9B96FF898C79B1A5F94DFB2BF946" attach="none" box="[1152,1184,1757,1785]" fontSize="6" pageId="6" pageNumber="111">nd–</superScript>
|
||
4
|
||
<superScript id="7C1D9B96FF898C79B188F94DFB31F954" attach="left" box="[1197,1210,1757,1771]" fontSize="6" pageId="6" pageNumber="111">th</superScript>
|
||
tarsomeres together in fore, middle, and hind legs, respectively.
|
||
</paragraph>
|
||
<paragraph id="8BD736DEFF898C79B5E2F8B6FCC0F8FE" blockId="6.[151,1436,1722,2037]" box="[199,843,1830,1857]" pageId="6" pageNumber="111">Ventrites IV–VI each with two pairs of paramedian setae.</paragraph>
|
||
<paragraph id="8BD736DEFF898C79B5E2F8DAFD72F86E" blockId="6.[151,1436,1722,2037]" pageId="6" pageNumber="111">
|
||
Male genitalia (
|
||
<figureCitation id="13532A5BFF898C79B450F8DAFE33F8DA" box="[373,440,1866,1893]" captionStart="FIGURE 3" captionStartId="4.[151,250,1957,1982]" captionTargetBox="[321,1265,1386,1932]" captionTargetId="figure@4.[321,1265,1386,1932]" captionTargetPageId="4" captionText="FIGURE 3 Male genitalia of Qianaphaenops (Sanwangius) rowselli Tian et Chen, n. subgen., n. sp., holotype A median lobe and parameres, lateral view B apical lobe, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3449023" httpUri="https://zenodo.org/record/3449023/files/figure.png" pageId="6" pageNumber="111">Fig. 3</figureCitation>
|
||
): The median lobe of aedeagus quite small but stout, arcuate medially; endophallus armed with a large copulatory piece which is about 1/4 as long as the median lobe; base large; in dorsal view the apical lobe quite elongate, rounded at apex; each parameres bearing six long setae at apex (except only two on right one and four on the left in one male from Sanwang Dong).
|
||
</paragraph>
|
||
<paragraph id="8BD736DEFF898C78B5E2F84BFF41FD80" blockId="6.[151,1436,1722,2037]" lastBlockId="7.[151,1437,153,971]" lastPageId="7" lastPageNumber="112" pageId="6" pageNumber="111">
|
||
<emphasis id="B91CEACCFF898C79B5E2F84BFE47F84A" bold="true" box="[199,460,2010,2037]" pageId="6" pageNumber="111">Larval characteristics</emphasis>
|
||
(
|
||
<figureCitation id="13532A5BFF898C79B4FEF84AFD94F84A" box="[475,543,2010,2037]" captionStart="FIGURE 4" captionStartId="5.[151,250,1911,1936]" captionTargetBox="[239,1357,209,1877]" captionTargetId="figure@5.[221,1367,181,1885]" captionTargetPageId="5" captionText="FIGURE 4 Larva of Qianaphaenops (Sanwangius) rowselli Tian et Chen, n. subgen., n. sp. A habitus, ventral view B head (cephalic capsule) and thoraces, dorsal view (FA: frontale, PA: parietal, PN: pronotum, MsN: mesonotum, MtN: metanotum) C head (cephalic capsule), ventral view D urogomphi and pygidium, dorsolateral view E right antenna, dorsal view F left maxilla, ventral view G tarsal claw of left middle leg, dorsal view, arrowhead indicating the small seta near base." figureDoi="http://doi.org/10.5281/zenodo.3449027" httpUri="https://zenodo.org/record/3449027/files/figure.png" pageId="6" pageNumber="111">Fig. 4</figureCitation>
|
||
): Habitus as in
|
||
<figureCitation id="13532A5BFF898C79B7EFF84AFC95F84A" box="[714,798,2010,2037]" captionStart="FIGURE 4" captionStartId="5.[151,250,1911,1936]" captionTargetBox="[239,1357,209,1877]" captionTargetId="figure@5.[221,1367,181,1885]" captionTargetPageId="5" captionText="FIGURE 4 Larva of Qianaphaenops (Sanwangius) rowselli Tian et Chen, n. subgen., n. sp. A habitus, ventral view B head (cephalic capsule) and thoraces, dorsal view (FA: frontale, PA: parietal, PN: pronotum, MsN: mesonotum, MtN: metanotum) C head (cephalic capsule), ventral view D urogomphi and pygidium, dorsolateral view E right antenna, dorsal view F left maxilla, ventral view G tarsal claw of left middle leg, dorsal view, arrowhead indicating the small seta near base." figureDoi="http://doi.org/10.5281/zenodo.3449027" httpUri="https://zenodo.org/record/3449027/files/figure.png" pageId="6" pageNumber="111">Fig. 4A</figureCitation>
|
||
and
|
||
<figureCitation id="13532A5BFF898C79B677F84AFC2CF84A" box="[850,935,2010,2037]" captionStart="FIGURE 7" captionStartId="8.[151,250,1886,1911]" captionTargetBox="[151,1435,181,1850]" captionTargetId="figure@8.[151,1435,181,1860]" captionTargetPageId="8" captionText="FIGURE 7 Sanwang Dong-Erwang Dong cave system, type locality of Qianaphaenops (Sanwangius) rowselli Tian et Chen, n. subgen., n. sp. A entrance of Sanwang Dong B Qingkou Tiankeng C “Xiong Da” (right) and Tian MY in front of Erwang Dong D an adult in Erwang Dong E a larva in Erwang Dong." figureDoi="http://doi.org/10.5281/zenodo.3449037" httpUri="https://zenodo.org/record/3449037/files/figure.png" pageId="6" pageNumber="111">Fig. 7E</figureCitation>
|
||
. Body campodeiform, long and slender, head width subequal to thoraces, abdominal segment I width equal to metathorax, then gradually narrowed towards apex. Length (from apex of mandibles to apex of urogomphi)
|
||
<quantity id="4C909B3BFF888C78B605FF2CFC1DFF68" box="[800,918,188,215]" metricMagnitude="-2" metricUnit="m" metricValue="1.125" pageId="7" pageNumber="112" unit="mm" value="11.25">11.25 mm</quantity>
|
||
; moderately sclerotized, head light yellowish brown, other parts lighter; body covered with some secondary setae. Head width subequal to thorax; abdominal segments bead-like. Cephalic capsule subquadrate, longer than wide (HL/HW = 1.10), parallel-sided, slightly sinuated before base; anterior margin of nasale triangular, protruding at middle, strongly serrated on either side which are symmetrical; frontale moderately curved, U-shaped at base; stemmata, ocular groove and egg-brusters absent; frontale (FR) with setae FR1–4 and FR7; parietal (PA) with setae PA1–10 and several secondary setae on each side (
|
||
<figureCitation id="13532A5BFF888C78B5BAFE05FF67FE10" box="[159,236,405,431]" captionStart="FIGURE 4" captionStartId="5.[151,250,1911,1936]" captionTargetBox="[239,1357,209,1877]" captionTargetId="figure@5.[221,1367,181,1885]" captionTargetPageId="5" captionText="FIGURE 4 Larva of Qianaphaenops (Sanwangius) rowselli Tian et Chen, n. subgen., n. sp. A habitus, ventral view B head (cephalic capsule) and thoraces, dorsal view (FA: frontale, PA: parietal, PN: pronotum, MsN: mesonotum, MtN: metanotum) C head (cephalic capsule), ventral view D urogomphi and pygidium, dorsolateral view E right antenna, dorsal view F left maxilla, ventral view G tarsal claw of left middle leg, dorsal view, arrowhead indicating the small seta near base." figureDoi="http://doi.org/10.5281/zenodo.3449027" httpUri="https://zenodo.org/record/3449027/files/figure.png" pageId="7" pageNumber="112">Figs. 4</figureCitation>
|
||
B–C); mandibles falciform; maxilla slender (
|
||
<figureCitation id="13532A5BFF888C78B7F9FE05FCBAFE10" box="[732,817,405,431]" captionStart="FIGURE 4" captionStartId="5.[151,250,1911,1936]" captionTargetBox="[239,1357,209,1877]" captionTargetId="figure@5.[221,1367,181,1885]" captionTargetPageId="5" captionText="FIGURE 4 Larva of Qianaphaenops (Sanwangius) rowselli Tian et Chen, n. subgen., n. sp. A habitus, ventral view B head (cephalic capsule) and thoraces, dorsal view (FA: frontale, PA: parietal, PN: pronotum, MsN: mesonotum, MtN: metanotum) C head (cephalic capsule), ventral view D urogomphi and pygidium, dorsolateral view E right antenna, dorsal view F left maxilla, ventral view G tarsal claw of left middle leg, dorsal view, arrowhead indicating the small seta near base." figureDoi="http://doi.org/10.5281/zenodo.3449027" httpUri="https://zenodo.org/record/3449027/files/figure.png" pageId="7" pageNumber="112">Fig. 4F</figureCitation>
|
||
); labium shorter than maxilla; antenna slender and long, 4 segmented, each much longer than wide, separated by weak membrane (
|
||
<figureCitation id="13532A5BFF888C78B6F7FE29FBA2FE6C" box="[978,1065,441,467]" captionStart="FIGURE 4" captionStartId="5.[151,250,1911,1936]" captionTargetBox="[239,1357,209,1877]" captionTargetId="figure@5.[221,1367,181,1885]" captionTargetPageId="5" captionText="FIGURE 4 Larva of Qianaphaenops (Sanwangius) rowselli Tian et Chen, n. subgen., n. sp. A habitus, ventral view B head (cephalic capsule) and thoraces, dorsal view (FA: frontale, PA: parietal, PN: pronotum, MsN: mesonotum, MtN: metanotum) C head (cephalic capsule), ventral view D urogomphi and pygidium, dorsolateral view E right antenna, dorsal view F left maxilla, ventral view G tarsal claw of left middle leg, dorsal view, arrowhead indicating the small seta near base." figureDoi="http://doi.org/10.5281/zenodo.3449027" httpUri="https://zenodo.org/record/3449027/files/figure.png" pageId="7" pageNumber="112">Fig. 4E</figureCitation>
|
||
). Thoraces transverse: pronotum, mesonotum and metanotum transverse with many secondary setae. Legs with only one claw (
|
||
<figureCitation id="13532A5BFF888C78B180FE4DFB74FE48" box="[1189,1279,477,503]" captionStart="FIGURE 4" captionStartId="5.[151,250,1911,1936]" captionTargetBox="[239,1357,209,1877]" captionTargetId="figure@5.[221,1367,181,1885]" captionTargetPageId="5" captionText="FIGURE 4 Larva of Qianaphaenops (Sanwangius) rowselli Tian et Chen, n. subgen., n. sp. A habitus, ventral view B head (cephalic capsule) and thoraces, dorsal view (FA: frontale, PA: parietal, PN: pronotum, MsN: mesonotum, MtN: metanotum) C head (cephalic capsule), ventral view D urogomphi and pygidium, dorsolateral view E right antenna, dorsal view F left maxilla, ventral view G tarsal claw of left middle leg, dorsal view, arrowhead indicating the small seta near base." figureDoi="http://doi.org/10.5281/zenodo.3449027" httpUri="https://zenodo.org/record/3449027/files/figure.png" pageId="7" pageNumber="112">Fig. 4G</figureCitation>
|
||
). Abdomen 9- segmented; urogomphi long and slender, each with setae UR2–10; pygidium long, but shorter than urogomphi (
|
||
<figureCitation id="13532A5BFF888C78B04AFD91FF35FD80" captionStart="FIGURE 4" captionStartId="5.[151,250,1911,1936]" captionTargetBox="[239,1357,209,1877]" captionTargetId="figure@5.[221,1367,181,1885]" captionTargetPageId="5" captionText="FIGURE 4 Larva of Qianaphaenops (Sanwangius) rowselli Tian et Chen, n. subgen., n. sp. A habitus, ventral view B head (cephalic capsule) and thoraces, dorsal view (FA: frontale, PA: parietal, PN: pronotum, MsN: mesonotum, MtN: metanotum) C head (cephalic capsule), ventral view D urogomphi and pygidium, dorsolateral view E right antenna, dorsal view F left maxilla, ventral view G tarsal claw of left middle leg, dorsal view, arrowhead indicating the small seta near base." figureDoi="http://doi.org/10.5281/zenodo.3449027" httpUri="https://zenodo.org/record/3449027/files/figure.png" pageId="7" pageNumber="112">Fig. 4D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3726555FF888C78B5E2FDD9FF41FD14" pageId="7" pageNumber="112" type="discussion">
|
||
<paragraph id="8BD736DEFF888C78B5E2FDD9FF41FD14" blockId="7.[151,1437,153,971]" pageId="7" pageNumber="112">
|
||
<emphasis id="B91CEACCFF888C78B5E2FDD9FEB0FDDC" bold="true" box="[199,315,585,611]" pageId="7" pageNumber="112">Remarks.</emphasis>
|
||
Larva of
|
||
<taxonomicName id="4C684D5DFF888C78B489FDD9FDD2FDDC" box="[428,601,585,611]" class="Insecta" family="Carabidae" genus="Qianaphaenops" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="112" phylum="Arthropoda" rank="species" species="rowselli" subGenus="Sanwangius">
|
||
<emphasis id="B91CEACCFF888C78B489FDD9FDD2FDDC" box="[428,601,585,611]" italics="true" pageId="7" pageNumber="112">Q. (S.) rowselli</emphasis>
|
||
</taxonomicName>
|
||
is a semi-aphaenopsian
|
||
<typeStatus id="54D3887CFF888C78B64AFDD9FC29FDDC" box="[879,930,585,611]" pageId="7" pageNumber="112">type</typeStatus>
|
||
, similar to that of
|
||
<taxonomicName id="4C684D5DFF888C78B151FDD9FEA3FD38" authority="Deuve, 1999" authorityName="Deuve" authorityYear="1999" class="Insecta" family="Carabidae" genus="Guizhaphaenops" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="112" phylum="Arthropoda" rank="species" species="lipsorum">
|
||
<emphasis id="B91CEACCFF888C78B151FDD9FA16FDDC" box="[1140,1437,585,611]" italics="true" pageId="7" pageNumber="112">Guizhaphaenops lipsorum</emphasis>
|
||
Deuve, 1999
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFF94B2FFF888C78B41CFDFDFE7BFD38" author="Luo, X. Z. & Yin, H. M. & Huang, S. B. & Tian, M. Y." box="[313,496,621,647]" pageId="7" pageNumber="112" pagination="97 - 109" refId="ref3994" refString="Luo, X. Z., Yin, H. M., Huang, S. B. & Tian, M. Y. (2017) Differences between anophthalmic and aphenopsian beetles: evidences from larvae of Chinese species. Tropical Zoology, 30 (3), 97 - 109. http: // doi. org / 10.1080 / 03946975.2017.1301629" type="journal article" year="2017">
|
||
Luo
|
||
<emphasis id="B91CEACCFF888C78B455FDFEFE21FD38" box="[368,426,621,647]" italics="true" pageId="7" pageNumber="112">et al.</emphasis>
|
||
2017
|
||
</bibRefCitation>
|
||
). But nasale is evenly serrated at front margin instead of trilobed in the latter species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3726555FF888C78B5E2FD25FEA2FD4C" pageId="7" pageNumber="112" type="etymology">
|
||
<paragraph id="8BD736DEFF888C78B5E2FD25FEA2FD4C" blockId="7.[151,1437,153,971]" pageId="7" pageNumber="112">
|
||
<emphasis id="B91CEACCFF888C78B5E2FD25FEC5FD70" bold="true" box="[199,334,693,719]" pageId="7" pageNumber="112">Etymology.</emphasis>
|
||
Dedicated to Dr. Philip John Rowsell (Hong Meigui Cave Exploration Society), a well-known Speleologist.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3726555FF888C78B5E2FD6CFBAEFC74" pageId="7" pageNumber="112" type="distribution">
|
||
<paragraph id="8BD736DEFF888C78B5E2FD6CFF59FC84" blockId="7.[151,1437,153,971]" pageId="7" pageNumber="112">
|
||
<emphasis id="B91CEACCFF888C78B5E2FD6CFEEAFCA8" bold="true" box="[199,353,764,791]" pageId="7" pageNumber="112">Distribution.</emphasis>
|
||
<collectingCountry id="F37F764EFF888C78B442FD6DFE21FCA8" box="[359,426,765,791]" name="China" pageId="7" pageNumber="112">China</collectingCountry>
|
||
(
|
||
<collectingRegion id="49ACF83CFF888C78B49CFD6DFDB0FCA8" box="[441,571,765,791]" country="China" name="Chongqing" pageId="7" pageNumber="112">Chongqing</collectingRegion>
|
||
: Wulong). Known from the Sanwang Dong-Erwang Dong cave system (
|
||
<figureCitation id="13532A5BFF888C78B041FD6DFF4EFC84" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStart-2="FIGURE 7" captionStartId-0="6.[151,250,1543,1568]" captionStartId-1="7.[151,250,1925,1950]" captionStartId-2="8.[151,250,1886,1911]" captionTargetBox-0="[165,1425,193,1511]" captionTargetBox-1="[157,1430,1001,1894]" captionTargetBox-2="[151,1435,181,1850]" captionTargetId-0="figure@6.[151,1435,181,1517]" captionTargetId-1="figure@7.[151,1435,993,1900]" captionTargetId-2="figure@8.[151,1435,181,1860]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="8" captionText-0="FIGURE 5 Geographical positions of Sanwang Dong and Erwang Dong, the localities of Qianaphaenops (Sanwangius) rowselli Tian et Chen, n. subgen., n. sp. A map of China to show the site of Chongqing B map of Chonqing to show the site of Wulong Qu C map of Wulong Qu, to show the site of Sanwang Dong-Erwang Dong cave system D sites of Sanwang Dong (a) and Erwang Dong (b)" captionText-1="FIGURE 6 Simplified maps of Sanwang Dong and Erwang Dong, to show the site where the cave beetles were found and collected (in courtesy of Dr. Philip John Rowsell, Hong Meigui Cave Exploration Society)." captionText-2="FIGURE 7 Sanwang Dong-Erwang Dong cave system, type locality of Qianaphaenops (Sanwangius) rowselli Tian et Chen, n. subgen., n. sp. A entrance of Sanwang Dong B Qingkou Tiankeng C “Xiong Da” (right) and Tian MY in front of Erwang Dong D an adult in Erwang Dong E a larva in Erwang Dong." figureDoi-0="http://doi.org/10.5281/zenodo.3449031" figureDoi-1="http://doi.org/10.5281/zenodo.3449035" figureDoi-2="http://doi.org/10.5281/zenodo.3449037" httpUri-0="https://zenodo.org/record/3449031/files/figure.png" httpUri-1="https://zenodo.org/record/3449035/files/figure.png" httpUri-2="https://zenodo.org/record/3449037/files/figure.png" pageId="7" pageNumber="112">Figs. 5–7</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BD736DEFF888C78B5E2FCD5FBAEFC74" blockId="7.[151,1437,153,971]" pageId="7" pageNumber="112">
|
||
Entrances of the both caves are about
|
||
<quantity id="4C909B3BFF888C78B753FCD4FD35FCE0" box="[630,702,836,863]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="7" pageNumber="112" unit="m" value="850.0">850 m</quantity>
|
||
from each other (
|
||
<figureCitation id="13532A5BFF888C78B6ADFCD5FC69FCE0" box="[904,994,836,863]" captionStart="FIGURE 5" captionStartId="6.[151,250,1543,1568]" captionTargetBox="[165,1425,193,1511]" captionTargetId="figure@6.[151,1435,181,1517]" captionTargetPageId="6" captionText="FIGURE 5 Geographical positions of Sanwang Dong and Erwang Dong, the localities of Qianaphaenops (Sanwangius) rowselli Tian et Chen, n. subgen., n. sp. A map of China to show the site of Chongqing B map of Chonqing to show the site of Wulong Qu C map of Wulong Qu, to show the site of Sanwang Dong-Erwang Dong cave system D sites of Sanwang Dong (a) and Erwang Dong (b)" figureDoi="http://doi.org/10.5281/zenodo.3449031" httpUri="https://zenodo.org/record/3449031/files/figure.png" pageId="7" pageNumber="112">Fig. 5D</figureCitation>
|
||
). The beetles were found and collected from three sites: one site in Sanwang Dong and two in Erwang Dong. In Sanwang Dong the beetles were collected in a twilight zone just close to the entrance (
|
||
<figureCitation id="13532A5BFF888C78B74CFC1DFD33FC18" box="[617,696,909,935]" captionStart="FIGURE 6" captionStartId="7.[151,250,1925,1950]" captionTargetBox="[157,1430,1001,1894]" captionTargetId="figure@7.[151,1435,993,1900]" captionTargetPageId="7" captionText="FIGURE 6 Simplified maps of Sanwang Dong and Erwang Dong, to show the site where the cave beetles were found and collected (in courtesy of Dr. Philip John Rowsell, Hong Meigui Cave Exploration Society)." figureDoi="http://doi.org/10.5281/zenodo.3449035" httpUri="https://zenodo.org/record/3449035/files/figure.png" pageId="7" pageNumber="112">Figs. 6</figureCitation>
|
||
,
|
||
<figureCitation id="13532A5BFF888C78B7E1FC1DFD62FC18" box="[708,745,909,935]" captionStart="FIGURE 7" captionStartId="8.[151,250,1886,1911]" captionTargetBox="[151,1435,181,1850]" captionTargetId="figure@8.[151,1435,181,1860]" captionTargetPageId="8" captionText="FIGURE 7 Sanwang Dong-Erwang Dong cave system, type locality of Qianaphaenops (Sanwangius) rowselli Tian et Chen, n. subgen., n. sp. A entrance of Sanwang Dong B Qingkou Tiankeng C “Xiong Da” (right) and Tian MY in front of Erwang Dong D an adult in Erwang Dong E a larva in Erwang Dong." figureDoi="http://doi.org/10.5281/zenodo.3449037" httpUri="https://zenodo.org/record/3449037/files/figure.png" pageId="7" pageNumber="112">7A</figureCitation>
|
||
). In Erwang Dong they were collected in dark zones, at about 100 metres from entrance and from Qingkou Tiankeng respectively (
|
||
<figureCitation id="13532A5BFF888C78B6BCFC21FC63FC74" box="[921,1000,945,971]" captionStart="FIGURE 6" captionStartId="7.[151,250,1925,1950]" captionTargetBox="[157,1430,1001,1894]" captionTargetId="figure@7.[151,1435,993,1900]" captionTargetPageId="7" captionText="FIGURE 6 Simplified maps of Sanwang Dong and Erwang Dong, to show the site where the cave beetles were found and collected (in courtesy of Dr. Philip John Rowsell, Hong Meigui Cave Exploration Society)." figureDoi="http://doi.org/10.5281/zenodo.3449035" httpUri="https://zenodo.org/record/3449035/files/figure.png" pageId="7" pageNumber="112">Figs. 6</figureCitation>
|
||
,
|
||
<figureCitation id="13532A5BFF888C78B6D1FC21FB93FC74" box="[1012,1048,945,971]" captionStart="FIGURE 7" captionStartId="8.[151,250,1886,1911]" captionTargetBox="[151,1435,181,1850]" captionTargetId="figure@8.[151,1435,181,1860]" captionTargetPageId="8" captionText="FIGURE 7 Sanwang Dong-Erwang Dong cave system, type locality of Qianaphaenops (Sanwangius) rowselli Tian et Chen, n. subgen., n. sp. A entrance of Sanwang Dong B Qingkou Tiankeng C “Xiong Da” (right) and Tian MY in front of Erwang Dong D an adult in Erwang Dong E a larva in Erwang Dong." figureDoi="http://doi.org/10.5281/zenodo.3449037" httpUri="https://zenodo.org/record/3449037/files/figure.png" pageId="7" pageNumber="112">7B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |