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<document id="27785DB6759F08B8CFC78A8DD4730B49" ID-CLB-Dataset="8257" ID-DOI="10.11646/zootaxa.4922.1.1" ID-GBIF-Dataset="188411de-569c-4408-a016-1a04b858c142" ID-ISSN="1175-5326" ID-Zenodo-Dep="4496316" ID-ZooBank="D7F9C6A3-9C28-4F4C-8E81-BF24849FDD8C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612331093960" checkinUser="plazi" docAuthor="Fabrizi, Silvia, Liu, Wan-Gang, Bai, Ming, Yang, Xing-Ke &amp; Ahrens, Dirk" docDate="2021" docId="03BD87E66A2DFE88AF89FB74FD103E11" docLanguage="en" docName="zootaxa.4922.1.1.pdf" docOrigin="Zootaxa 4922 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Maladera nanlingensis Ahrens, Fabrizi &amp; Liu 2021, sp. n." docType="treatment" docVersion="8" lastPageNumber="334" masterDocId="FF84FF9E6B61FFC5AF1EF86AFFDE3D4C" masterDocTitle="A monograph of the genus Maladera Mulsant &amp; Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini)" masterLastPageNumber="400" masterPageNumber="1" pageNumber="333" updateTime="1698926631736" updateUser="ExternalLinkService">
<mods:mods id="24DFCF4FD2FFC517E562EC844CC7086C" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="781A7DE907287F180F2312116FBEDBA1">A monograph of the genus Maladera Mulsant &amp; Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini)</mods:title>
</mods:titleInfo>
<mods:name id="02D2F627C719F22F5E76D54D87106129" type="personal">
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</mods:role>
<mods:namePart id="E8E45B252E1FEC669B0542969D69DBAE">Fabrizi, Silvia</mods:namePart>
<mods:nameIdentifier id="03FFFC67F73C77556CB203094399A429" type="ORCID">0000-0003-3524-7153</mods:nameIdentifier>
<mods:nameIdentifier id="5B5796BA4087787B5447169360384D28" type="email">ahrens.dirk_col@gmx.de</mods:nameIdentifier>
</mods:name>
<mods:name id="206FFA4831B59ACCCB647B75C08856FC" type="personal">
<mods:role id="693470C877C0D8C6142D78E684287DEB">
<mods:roleTerm id="E4D334D1B16A03880C737C2643A1B15F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CD528F0663BA86DD1500DC0FB624C0DE">Liu, Wan-Gang</mods:namePart>
<mods:nameIdentifier id="BD6FEAD4922786E06325EEE0D1B2D26A" type="ORCID">0000-0003-4788-7967</mods:nameIdentifier>
<mods:nameIdentifier id="B9A3EAC62F39B3FDEE05B88CCE1DBA42" type="email">liuwangang@ieecas.cn</mods:nameIdentifier>
</mods:name>
<mods:name id="0A38C2B19BBD3A632C3759C748A89841" type="personal">
<mods:role id="7BD2E43D53EB023E01B15296033AB336">
<mods:roleTerm id="3AEE5889BB170E4691289D2A5F5F66FF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C615F7F0AFC3267E1F4BA03BC2931E45">Bai, Ming</mods:namePart>
<mods:nameIdentifier id="61E90F26332C251ED97FFA34868EBD1E" type="ORCID">0000-0003-3676-6828</mods:nameIdentifier>
<mods:nameIdentifier id="1133E744F8B42C1E1D1030E2F58C95F4" type="email">liuwangang@ieecas.cn</mods:nameIdentifier>
</mods:name>
<mods:name id="529B147122B9C1D2CEFA637B17CAFE70" type="personal">
<mods:role id="CCBF6532CBBA537B6FB9372CC4B6DE58">
<mods:roleTerm id="A0C0CAA43BDEE4DED9BE587D7F8D1E8F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="52869820D43DC42B04861643244FB2AB">Yang, Xing-Ke</mods:namePart>
<mods:nameIdentifier id="154A070CCA81F1CFB96ECCF44F6613D7" type="ORCID">0000-0003-3676-6828</mods:nameIdentifier>
<mods:nameIdentifier id="D23101DAC8BD8BFEE2B01382B75067E9" type="email">liuwangang@ieecas.cn</mods:nameIdentifier>
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<mods:name id="6DFCD778668CF6090040DD1C652E3BE8" type="personal">
<mods:role id="AA1D9EB539DDA3CC59590D86D611AD6F">
<mods:roleTerm id="7CB9BF51EEFE69D1B5034FD194561AA2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E2624CD370BFF2F4BEE459A9C580F49F">Ahrens, Dirk</mods:namePart>
<mods:nameIdentifier id="FB8C9D48AAD28497C11527977E4A1CAD" type="ORCID">0000-0003-3524-7153</mods:nameIdentifier>
<mods:nameIdentifier id="BF15CC89D9B08AB806D328D36BF04887" type="email">ahrens.dirk_col@gmx.de</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="AA5CAAB5ACB47D880F53C38886E64685">text</mods:typeOfResource>
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<mods:title id="99E1A78621329C86E570B68EF1DCE91E">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="5326762619F4E6901D7A8FF27706346F">
<mods:date id="9915661E78F760D6976067E5C22B7564">2021</mods:date>
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<mods:number id="D5D37DBCF0DA0F423B068F3E53C85CF8">2021-02-03</mods:number>
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<mods:number id="FBDCB5C0C2BEB9BA4C9D18BF052014A3">4922</mods:number>
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<mods:number id="5FA5822326B18AF897EEFC6BB7ED611D">1</mods:number>
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<mods:start id="FF010FD83FE1C4263756F2B761103ACE">1</mods:start>
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<mods:classification id="4103725B7DF0BA4F9131B8827DFA6995">journal article</mods:classification>
<mods:identifier id="A9F60394EC2C68C76D6FA71419C64DE0" type="CLB-Dataset">8257</mods:identifier>
<mods:identifier id="AA0D83562F964C91D8CA8371FCB0E199" type="DOI">10.11646/zootaxa.4922.1.1</mods:identifier>
<mods:identifier id="22C42F17B0611D44BCEC9006CDA9F395" type="GBIF-Dataset">188411de-569c-4408-a016-1a04b858c142</mods:identifier>
<mods:identifier id="F918457443978C13D7AF87B463582A13" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="6B6E41909BC16969B4BF15B37804EEC8" type="Zenodo-Dep">4496316</mods:identifier>
<mods:identifier id="632DDB56750E8EC20485B02744685722" type="ZooBank">D7F9C6A3-9C28-4F4C-8E81-BF24849FDD8C</mods:identifier>
</mods:mods>
<treatment id="03BD87E66A2DFE88AF89FB74FD103E11" ID-DOI="http://doi.org/10.5281/zenodo.4534969" ID-GBIF-Taxon="177323740" ID-Zenodo-Dep="4534969" LSID="urn:lsid:plazi:treatment:03BD87E66A2DFE88AF89FB74FD103E11" httpUri="http://treatment.plazi.org/id/03BD87E66A2DFE88AF89FB74FD103E11" lastPageId="333" lastPageNumber="334" pageId="332" pageNumber="333">
<subSubSection id="C30E657B6A2DFE89AF89FB74FCC13E75" box="[151,799,798,825]" pageId="332" pageNumber="333" type="nomenclature">
<paragraph id="8BAB36F06A2DFE89AF89FB74FCC13E75" blockId="332.[151,799,798,861]" box="[151,799,798,825]" pageId="332" pageNumber="333">
<heading id="D0E3819C6A2DFE89AF89FB74FCC13E75" bold="true" box="[151,799,798,825]" fontSize="11" level="2" pageId="332" pageNumber="333" reason="4">
<emphasis id="B960EAE26A2DFE89AF89FB74FCC13E75" bold="true" box="[151,799,798,825]" pageId="332" pageNumber="333">
<taxonomicName id="4C144D736A2DFE89AF89FB74FD0E3E74" authority="Ahrens, Fabrizi &amp; Liu" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" box="[151,720,798,825]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="332" pageNumber="333" phylum="Arthropoda" rank="species" species="nanlingensis" status="sp. nov.">
<emphasis id="B960EAE26A2DFE89AF89FB74FE713E75" bold="true" box="[151,431,798,825]" italics="true" pageId="332" pageNumber="333">Maladera nanlingensis</emphasis>
Ahrens, Fabrizi &amp; Liu
</taxonomicName>
,
<taxonomicNameLabel id="A25357996A2DFE89ADC3FB75FCC13E75" box="[733,799,799,825]" pageId="332" pageNumber="333" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6A2DFE89AF89FB29FEB13E11" box="[151,367,835,861]" pageId="332" pageNumber="333" type="description">
<paragraph id="8BAB36F06A2DFE89AF89FB29FEB13E11" blockId="332.[151,799,798,861]" box="[151,367,835,861]" pageId="332" pageNumber="333">
<figureCitation id="132F2A756A2DFE89AF89FB29FED03E11" box="[151,270,835,861]" captionStart="FIGURE 79" captionStartId="330.[151,250,1943,1967]" captionTargetBox="[172,1417,198,1910]" captionTargetId="figure-19@330.[156,1431,181,1920]" captionTargetPageId="330" captionText="FIGURE 79. A–D: Maladera wandingana Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); E–H: M. trifidiforceps Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); I–L: M. nanlingensis Ahrens, Fabrizi &amp; Liu, sp. n. (holotype). A, E, I: aedeagus, left side lateral view; C, G, K: aedeagus, right side lateral view; B, F, J: parameres, dorsal view; D, H, L: habitus. Scale: 0.5 mm. Habitus not to scale" figureDoi="http://doi.org/10.5281/zenodo.4496528" httpUri="https://zenodo.org/record/4496528/files/figure.png" pageId="332" pageNumber="333">Figures 79</figureCitation>
I–L, 129
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6A2DFE89AF89FBE1FA8A395D" pageId="332" pageNumber="333" type="materials_examined">
<paragraph id="8BAB36F06A2DFE89AF89FBE1FA8A395D" blockId="332.[151,1437,906,2013]" pageId="332" pageNumber="333">
<emphasis id="B960EAE26A2DFE89AF89FBE1FE1F3EE8" bold="true" box="[151,449,906,933]" pageId="332" pageNumber="333">Type material examined.</emphasis>
<materialsCitation id="3B7C3CAD6A2DFE89AED7FBE1FE5A3EA1" ID-GBIF-Occurrence="3034555609" collectingDate="2013-04-18" collectingDateMax="2013-04-21" collectingDateMin="2013-04-18" collectionCode="NMPC" collectorName="J. Hajek &amp; J. Ruzicka" country="China" elevation="690" latitude="24.933332" location="Nanling National Nature Reserve" longLatPrecision="124" longitude="112.715" municipality="Nanling National Nature Reserve Dadongshan" pageId="332" pageNumber="333" specimenCount="1" specimenCount-male="1" stateProvince="Guangdong" typeStatus="holotype">
<typeStatus id="54AF88526A2DFE89AED7FBE1FDE83EE9" box="[457,566,907,933]" pageId="332" pageNumber="333" type="holotype">Holotype</typeStatus>
:
<specimenCount id="9D12FD796A2DFE89AD5FFBE1FD883EE9" box="[577,598,907,933]" pageId="332" pageNumber="333" type="male">♂</specimenCount>
“
<collectingCountry id="F30376606A2DFE89AD76FBE1FD6F3EE9" box="[616,689,907,933]" name="China" pageId="332" pageNumber="333">China</collectingCountry>
:
<collectingRegion id="49D0F8126A2DFE89ADA3FBE1FC5F3EE9" box="[701,897,907,933]" country="China" name="Guangdong" pageId="332" pageNumber="333">Guangdong prov.</collectingRegion>
<collectingMunicipality id="6BCFAC8A6A2DFE89AC96FBE1FA463EE9" box="[904,1432,907,933]" pageId="332" pageNumber="333">
<location id="8ECB602B6A2DFE89AC96FBE1FADF3EE9" LSID="urn:lsid:plazi:treatment:03BD87E66A2DFE88AF89FB74FD103E11:8ECB602B6A2DFE89AC96FBE1FADF3EE9" box="[904,1281,907,933]" country="China" latitude="24.933332" longLatPrecision="124" longitude="112.715" municipality="Nanling National Nature Reserve Dadongshan" name="Nanling National Nature Reserve" pageId="332" pageNumber="333" stateProvince="Guangdong">Nanling National Nature Reserve</location>
Dadongshan
</collectingMunicipality>
,
<date id="FFAA10306A2DFE89AF89FBC5FEE43E85" box="[151,314,943,969]" pageId="332" pageNumber="333" value="2013-04-18" valueMax="2013-04-21" valueMin="2013-04-18">
<collectingDate id="EFEEE9D86A2DFE89AF89FBC5FEE43E85" box="[151,314,943,969]" pageId="332" pageNumber="333" value="2013-04-18" valueMax="2013-04-21" valueMin="2013-04-18">18.-21.iv.2013</collectingDate>
</date>
(border of mixed forest, at light)
<geoCoordinate id="EE2050376A2DFE89ADB2FBC4FCFA3E85" box="[684,804,942,969]" degrees="24" direction="north" minutes="56.0" orientation="latitude" pageId="332" pageNumber="333" precision="92" value="24.933332">24°56.0’N</geoCoordinate>
,
<geoCoordinate id="EE2050376A2DFE89AC31FBC5FC6C3E85" box="[815,946,943,969]" degrees="112" direction="east" minutes="42.9" orientation="longitude" pageId="332" pageNumber="333" precision="92" value="112.715">112°42.9’E</geoCoordinate>
,
<quantity id="4CEC9B156A2DFE89ACA2FBC5FBDC3E85" box="[956,1026,943,969]" metricMagnitude="2" metricUnit="m" metricValue="6.9" pageId="332" pageNumber="333" unit="m" value="690.0">
<elevation id="0039D1C36A2DFE89ACA2FBC5FBDC3E85" box="[956,1026,943,969]" metricMagnitude="2" metricUnit="m" metricValue="6.9" pageId="332" pageNumber="333" unit="m" value="690.0">690 m</elevation>
</quantity>
<collectorName id="26E153266A2DFE89AB16FBC5FBBD3E85" box="[1032,1123,943,969]" pageId="332" pageNumber="333">J. Hájek</collectorName>
&amp;
<collectorName id="26E153266A2DFE89AB9BFBC5FB263E85" box="[1157,1272,942,969]" pageId="332" pageNumber="333">J. Růžicka</collectorName>
/ 1077
<taxonomicName id="4C144D736A2DFE89AA5BFBC5FA423E85" box="[1349,1436,943,969]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="332" pageNumber="380" phylum="Arthropoda" rank="tribe" tribe="Sericini">Sericini</taxonomicName>
Asia spec.” (
<collectionCode id="ED05AE356A2DFE89AE38FBB9FEA23EA1" box="[294,380,979,1005]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="332" pageNumber="333">NMPC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B7C3CAD6A2DFE89AE90FBB9FA8E395D" ID-GBIF-Occurrence="3034556779" collectingDate="2013-04-18" collectingDateMax="2013-04-21" collectingDateMin="2013-04-18" collectionCode="ZFMK" collectorName="J. Hajek &amp; J. Ruzicka" country="China" elevation="690" latitude="24.933332" location="Nanling National Nature Reserve" longLatPrecision="124" longitude="112.715" municipality="Nanling National Nature Reserve Dadongshan" pageId="332" pageNumber="333" specimenCount="1" specimenCount-female="1" stateProvince="Guangdong" typeStatus="paratype">
<typeStatus id="54AF88526A2DFE89AE90FBB9FE2A3EA1" box="[398,500,979,1005]" pageId="332" pageNumber="333" type="paratype">Paratype</typeStatus>
:
<specimenCount id="9D12FD796A2DFE89AEE1FBB9FDF73EA1" box="[511,553,979,1005]" pageId="332" pageNumber="333" type="female">1 ♀</specimenCount>
“
<collectingCountry id="F30376606A2DFE89AD25FBB9FD5A3EA1" box="[571,644,979,1005]" name="China" pageId="332" pageNumber="333">China</collectingCountry>
:
<collectingRegion id="49D0F8126A2DFE89AD91FBB9FC8C3EA1" box="[655,850,979,1005]" country="China" name="Guangdong" pageId="332" pageNumber="333">Guangdong prov.</collectingRegion>
<collectingMunicipality id="6BCFAC8A6A2DFE89AC46FBB9FABB3EA1" box="[856,1381,979,1005]" pageId="332" pageNumber="333">
<location id="8ECB602B6A2DFE89AC46FBB9FB103EA1" LSID="urn:lsid:plazi:treatment:03BD87E66A2DFE88AF89FB74FD103E11:8ECB602B6A2DFE89AC46FBB9FB103EA1" box="[856,1230,979,1005]" country="China" latitude="24.933332" longLatPrecision="124" longitude="112.715" municipality="Nanling National Nature Reserve Dadongshan" name="Nanling National Nature Reserve" pageId="332" pageNumber="333" stateProvince="Guangdong">Nanling National Nature Reserve</location>
Dadongshan
</collectingMunicipality>
,
<date id="FFAA10306A2DFE89AA6EFBB9FED3395D" pageId="332" pageNumber="333" value="2013-04-18" valueMax="2013-04-21" valueMin="2013-04-18">
<collectingDate id="EFEEE9D86A2DFE89AA6EFBB9FED3395D" pageId="332" pageNumber="333" value="2013-04-18" valueMax="2013-04-21" valueMin="2013-04-18">18.- 21.iv.2013</collectingDate>
</date>
(border of mixed forest, at light)
<geoCoordinate id="EE2050376A2DFE89AD98FB9CFD20395D" box="[646,766,1014,1041]" degrees="24" direction="north" minutes="56.0" orientation="latitude" pageId="332" pageNumber="333" precision="92" value="24.933332">24°56.0’N</geoCoordinate>
,
<geoCoordinate id="EE2050376A2DFE89AC17FB9DFC52395D" box="[777,908,1015,1041]" degrees="112" direction="east" minutes="42.9" orientation="longitude" pageId="332" pageNumber="333" precision="92" value="112.715">112°42.9’E</geoCoordinate>
,
<quantity id="4CEC9B156A2DFE89AC89FB9DFC00395D" box="[919,990,1015,1041]" metricMagnitude="2" metricUnit="m" metricValue="6.9" pageId="332" pageNumber="333" unit="m" value="690.0">
<elevation id="0039D1C36A2DFE89AC89FB9DFC00395D" box="[919,990,1015,1041]" metricMagnitude="2" metricUnit="m" metricValue="6.9" pageId="332" pageNumber="333" unit="m" value="690.0">690 m</elevation>
</quantity>
<collectorName id="26E153266A2DFE89ACFBFB9DFB9E395D" box="[997,1088,1015,1041]" pageId="332" pageNumber="333">J. Hájek</collectorName>
&amp;
<collectorName id="26E153266A2DFE89AB7AFB9DFB02395D" box="[1124,1244,1014,1041]" pageId="332" pageNumber="333">J. Růžicka</collectorName>
” (
<collectionCode id="ED05AE356A2DFE89ABEAFB9DFA96395D" box="[1268,1352,1015,1041]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="332" pageNumber="333" type="Museum">ZFMK</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6A2DFE88AFD9FC70FD453F2D" lastPageId="333" lastPageNumber="334" pageId="332" pageNumber="333" type="description">
<paragraph id="8BAB36F06A2DFE89AFD9FC70FAAA3915" blockId="332.[151,1437,906,2013]" pageId="332" pageNumber="333">
<emphasis id="B960EAE26A2DFE89AFD9FC70FE873978" bold="true" box="[199,345,1050,1076]" pageId="332" pageNumber="333">Description.</emphasis>
Length:
<quantity id="4CEC9B156A2DFE89AEA2FC71FDCA3979" box="[444,532,1051,1077]" metricMagnitude="-3" metricUnit="m" metricValue="7.8" pageId="332" pageNumber="333" unit="mm" value="7.8">7.8 mm</quantity>
, length of elytra:
<quantity id="4CEC9B156A2DFE89ADC8FC70FCF33979" box="[726,813,1050,1077]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="332" pageNumber="333" unit="mm" value="5.6">5.6 mm</quantity>
, width:
<quantity id="4CEC9B156A2DFE89AC98FC71FC013979" box="[902,991,1051,1077]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="332" pageNumber="333" unit="mm" value="4.6">4.6 mm</quantity>
. Body oblong, dark brown, antenna yellowish, surface dull, labroclypeus and anterior frons narrowly shiny, except a few long setae on head, glabrous.
</paragraph>
<paragraph id="8BAB36F06A2DFE89AFD9FC09FA9838D1" blockId="332.[151,1437,906,2013]" pageId="332" pageNumber="333">
Labroclypeus wide and trapezoidal, widest at base, lateral margins straight and strongly convergent anteriorly, anterior angles moderately rounded; anterior margin weakly emarginate medially; margins moderately reflexed; lateral margin and ocular canthus produce an indistinct angle; surface flat, very coarsely, densely punctate, with numerous erect setae; frontoclypeal suture finely incised, angled medially; smooth area anterior to eye convex, 2.5 times as wide as long; ocular canthus short and narrow (1/3 of ocular diameter), finely densely punctate, with a long terminal seta. Frons in posterior part dull, anteriorly tomentum lacking and shiny, with coarse, moderately dense, irregular, punctures, with a few single setae beside eyes and along a transversal line on disc. Eyes moderately large, ratio diameter/ interocular width: 0.61. Antenna with ten antennomeres; club with three antennomeres and straight, slightly shorter than remaining antennomeres combined.
<collectingRegion id="49D0F8126A2DFE89AC08FDE9FCAB38D1" box="[790,885,1411,1437]" country="Myanmar" name="Chin" pageId="332" pageNumber="333">Mentum</collectingRegion>
elevated and slightly flattened anteriorly.
</paragraph>
<paragraph id="8BAB36F06A2DFE89AFD9FDCDFCFA3B1D" blockId="332.[151,1437,906,2013]" pageId="332" pageNumber="333">Pronotum moderately wide, widest at base, lateral margins in evenly convex, moderately convergent anteriorly and posteriorly, anterior angles moderately produced and slightly rounded at tip, posterior angles blunt; anterior margin weakly convex, marginal line absent, base without marginal line; surface finely and densely punctate, with minute setae in punctures; anterior and lateral margin robustly setose; hypomeron carinate, not produced ventrally. Scutellum wide, triangular, with fine and dense punctures.</paragraph>
<paragraph id="8BAB36F06A2DFE89AFD9FE31FF203A4A" blockId="332.[151,1437,906,2013]" pageId="332" pageNumber="333">Elytra oblong, widest at posterior third, striae finely impressed, finely and densely punctate, intervals weakly convex, with fine, moderately dense punctures, odd intervals with a few single short, erect setae, otherwise with minute setae in punctures; epipleural margin robust, ending at convex external apical angle of elytra, epipleura sparsely setose; apical border of elytra membraneous, with a fine rim of microtrichomes (visible at ca 100x magnification).</paragraph>
<paragraph id="8BAB36F06A2DFE89AFD9FF65FEE63A91" blockId="332.[151,1437,906,2013]" pageId="332" pageNumber="333">Ventral surface dull, finely and densely punctate, with dense and fine, adpressed setae, metasternal disc sparsely covered with longer setae; metacoxa with a few longer setae laterally. Abdominal sternites finely and densely punctate, glabrous, each sternite with a transverse row of punctures each bearing a fine seta. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.42. Pygidium moderately convex, dull, finely and densely punctate, without smooth midline, with short setae throughout and a few long setae along apical margin.</paragraph>
<paragraph id="8BAB36F06A2CFE88AFD9F8FDFE8E3F71" blockId="333.[151,1437,151,861]" pageId="333" pageNumber="334">Legs long and moderately wide; femora shiny, with two longitudinal rows of setae, finely and sparsely punctate. Anterior margin of metafemur acute, without adjacent serrated line; posterior ventral margin smooth, not widened at apex, dorsal posterior margin smooth, neither serrate, glabrous. Metatibia long and moderately wide, widest at middle, ratio of width/length: 1/3.6, weakly carinate dorsally, with two groups of spines, basal group at one third, apical group at three quarters of metatibial length, in basal half a few short single spines; lateral face longitudinally convex, finely sparsely punctate on sides, with a few minute setae in punctures; ventral margin finely serrate, with five robust equidistant setae; medial face smooth and glabrous; apex finely serrate, shallowly emarginate interiorly near tarsal articulation. Tarsomeres dorsally impunctate, glabrous, neither laterally nor dorsally carinate, moderately setose ventrally; metatarsomeres glabrous and with a strongly serrated ridge ventrally and a smooth subventral longitudinal carina; first metatarsomere as long as following two tarsomeres combined and distinctly longer than dorsal tibial spur. Protibia moderately long, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex.</paragraph>
<paragraph id="8BAB36F06A2CFE88AFD9FA2DFD453F2D" blockId="333.[151,1437,151,861]" box="[199,667,583,609]" pageId="333" pageNumber="334">
Aedeagus:
<figureCitation id="132F2A756A2CFE88AE5AFA2DFE4B3F2D" box="[324,405,583,609]" captionStart="FIGURE 79" captionStartId="330.[151,250,1943,1967]" captionTargetBox="[172,1417,198,1910]" captionTargetId="figure-19@330.[156,1431,181,1920]" captionTargetPageId="330" captionText="FIGURE 79. A–D: Maladera wandingana Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); E–H: M. trifidiforceps Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); I–L: M. nanlingensis Ahrens, Fabrizi &amp; Liu, sp. n. (holotype). A, E, I: aedeagus, left side lateral view; C, G, K: aedeagus, right side lateral view; B, F, J: parameres, dorsal view; D, H, L: habitus. Scale: 0.5 mm. Habitus not to scale" figureDoi="http://doi.org/10.5281/zenodo.4496528" httpUri="https://zenodo.org/record/4496528/files/figure.png" pageId="333" pageNumber="334">Fig. 79</figureCitation>
I–K. Habitus:
<figureCitation id="132F2A756A2CFE88AD2DFA2DFD493F2D" box="[563,663,583,609]" captionStart="FIGURE 79" captionStartId="330.[151,250,1943,1967]" captionTargetBox="[172,1417,198,1910]" captionTargetId="figure-19@330.[156,1431,181,1920]" captionTargetPageId="330" captionText="FIGURE 79. A–D: Maladera wandingana Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); E–H: M. trifidiforceps Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); I–L: M. nanlingensis Ahrens, Fabrizi &amp; Liu, sp. n. (holotype). A, E, I: aedeagus, left side lateral view; C, G, K: aedeagus, right side lateral view; B, F, J: parameres, dorsal view; D, H, L: habitus. Scale: 0.5 mm. Habitus not to scale" figureDoi="http://doi.org/10.5281/zenodo.4496528" httpUri="https://zenodo.org/record/4496528/files/figure.png" pageId="333" pageNumber="334">
Fig.
<specimenCount id="9D12FD796A2CFE88AD79FA2DFD493F2D" box="[615,663,583,609]" pageId="333" pageNumber="334" type="generic">79L</specimenCount>
</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6A2CFE88AFD9FA00FC0B3FE5" pageId="333" pageNumber="334" type="diagnosis">
<paragraph id="8BAB36F06A2CFE88AFD9FA00FC0B3FE5" blockId="333.[151,1437,151,861]" pageId="333" pageNumber="334">
<emphasis id="B960EAE26A2CFE88AFD9FA00FE9C3FC8" bold="true" box="[199,322,618,644]" pageId="333" pageNumber="334">Diagnosis.</emphasis>
This new species resembles somewhat members of the subgenus
<taxonomicName id="4C144D736A2CFE88AB06FA01FB583FC9" authorityName="Mulsant &amp; Rey" authorityYear="1871" box="[1048,1158,619,645]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="333" pageNumber="334" phylum="Arthropoda" rank="genus">
<emphasis id="B960EAE26A2CFE88AB06FA01FB583FC9" box="[1048,1158,619,645]" italics="true" pageId="333" pageNumber="334">Maladera</emphasis>
</taxonomicName>
; however, the parameres are sysmmetrical and the anterior serrated line of the metafemur is absent.
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6A2CFE88AFD9FAD9FD9D3FBE" pageId="333" pageNumber="334" type="etymology">
<paragraph id="8BAB36F06A2CFE88AFD9FAD9FD9D3FBE" blockId="333.[151,1437,151,861]" pageId="333" pageNumber="334">
<emphasis id="B960EAE26A2CFE88AFD9FAD9FE903F81" bold="true" box="[199,334,691,717]" pageId="333" pageNumber="334">Etymology.</emphasis>
The name of the new species is derived from its occurrence in the Nanling National Nature Reserve (adjective in the nominative singular).
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6A2CFE88AFD9FA90FAF03E75" pageId="333" pageNumber="334" type="description">
<paragraph id="8BAB36F06A2CFE88AFD9FA90FAF03E75" blockId="333.[151,1437,151,861]" pageId="333" pageNumber="334">
<emphasis id="B960EAE26A2CFE88AFD9FA90FEE13E58" bold="true" box="[199,319,762,788]" pageId="333" pageNumber="334">Variation.</emphasis>
Length:
<quantity id="4CEC9B156A2CFE88AEBBFA91FDF03E59" box="[421,558,763,789]" metricMagnitude="-3" metricUnit="m" metricValue="7.699999999999999" metricValueMax="7.8" metricValueMin="7.6" pageId="333" pageNumber="334" unit="mm" value="7.699999999999999" valueMax="7.8" valueMin="7.6">7.6–7.8 mm</quantity>
, length of elytra:
<quantity id="4CEC9B156A2CFE88ADEBFA90FCA13E59" box="[757,895,762,789]" metricMagnitude="-3" metricUnit="m" metricValue="5.55" metricValueMax="5.6" metricValueMin="5.5" pageId="333" pageNumber="334" unit="mm" value="5.55" valueMax="5.6" valueMin="5.5">5.5–5.6 mm</quantity>
, width:
<quantity id="4CEC9B156A2CFE88ACC7FA91FBBA3E59" box="[985,1124,763,789]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="4.6" metricValueMin="4.4" pageId="333" pageNumber="334" unit="mm" value="4.5" valueMax="4.6" valueMin="4.4">4.4–4.6 mm</quantity>
. Female: Pygidium flat, antennal club as long as in male; Eyes slightly smaller than in male, ratio diameter/ interocular width: 0.53.
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6A2CFE88AFD9FB28FD103E11" box="[199,718,834,861]" pageId="333" pageNumber="334" type="distribution">
<paragraph id="8BAB36F06A2CFE88AFD9FB28FD103E11" blockId="333.[151,1437,151,861]" box="[199,718,834,861]" pageId="333" pageNumber="334">
<emphasis id="B960EAE26A2CFE88AFD9FB28FEBF3E10" bold="true" box="[199,353,834,860]" pageId="333" pageNumber="334">Distribution.</emphasis>
See map (
<figureCitation id="132F2A756A2CFE88AEC8FB29FDE83E11" box="[470,566,835,861]" captionStart="FIGURE 129" captionStartId="366.[151,250,1001,1025]" captionTargetBox="[231,1354,187,971]" captionTargetId="figure-19@366.[226,1362,181,977]" captionTargetPageId="366" captionText="FIGURE 129. Distribution of Maladera species with uncertain systematic status: Maladera baishaoensis sp. n., M. hutiaoensis sp. n., M. jinggangshanica sp. n., M. lukjanovitschi (Medvedev, 1966), M. luoxiangensis sp. n., M. nabanensis sp. n., M. nanlingensis sp. n.." figureDoi="http://doi.org/10.5281/zenodo.4496668" httpUri="https://zenodo.org/record/4496668/files/figure.png" pageId="333" pageNumber="334">Fig. 129</figureCitation>
) and
<tableCitation id="C696034B6A2CFE88AD6AFB29FD163E11" box="[628,712,835,861]" captionStart="TABLE 1" captionStartId="368.[151,239,544,570]" captionText="TABLE 1. Overview on the distribuition of the Chinese mainland species of Maladera Mulsant &amp; Rey, 1871 (x – records from material examined herein, o – records from literature, with no material of this study)." pageId="333" pageNumber="334">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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