276 lines
32 KiB
XML
276 lines
32 KiB
XML
<document id="0A6700154CFE7B96E3B2D05C9056D7FC" ID-CLB-Dataset="29024" ID-DOI="10.11646/zootaxa.4459.2.2" ID-GBIF-Dataset="634e23da-84c4-46a4-8bb8-94fcf1637160" ID-ISSN="1175-5326" ID-Zenodo-Dep="1458548" ID-ZooBank="73510C3B-6482-4760-8173-B6E701839940" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1539288809625" checkinUser="plazi" docAuthor="Sheth, Sayali D., Ghate, Hemant V. & Hájek, Jiří" docDate="2018" docId="03BC3747FF92FF99FF5E1396FBC7FAD0" docLanguage="en" docName="zootaxa.4459.2.2.pdf" docOrigin="Zootaxa 4459 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Copelatus maushomi Sheth & Ghate & Hájek 2018" docType="treatment" docVersion="7" lastPageNumber="245" masterDocId="FF854F3FFF9BFF93FFC91143FFF8FFA8" masterDocTitle="Copelatus Erichson, 1832 from Maharashtra, India, with description of three new species and notes on other taxa of the genus (Coleoptera: Dytiscidae: Copelatinae)" masterLastPageNumber="260" masterPageNumber="235" pageNumber="244" updateTime="1698578320975" updateUser="ExternalLinkService">
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<mods:mods id="83F13E8B17A7F97DEFB8BDA6CB0C13BF" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="81C37BF209EA05C9C5D376DE6AE46CD4">
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<mods:title id="23BCDB50B54F3BDAE1E8537244EFC28C">Copelatus Erichson, 1832 from Maharashtra, India, with description of three new species and notes on other taxa of the genus (Coleoptera: Dytiscidae: Copelatinae)</mods:title>
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</mods:titleInfo>
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<mods:name id="73619FAF864F6A2168B5863C20547F6D" type="personal">
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<mods:role id="9F1F6D24AC9F7CB4CE1FE5242498C273">
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<mods:roleTerm id="6C03B921CA4629A0154C55DCCB487386">Author</mods:roleTerm>
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<mods:namePart id="6C4B4948F70337DB75784EF600217922">Sheth, Sayali D.</mods:namePart>
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</mods:name>
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<mods:name id="2C7EA2111AADA623C5E8C2E6A40C3597" type="personal">
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<mods:role id="53F150F17E98AB15527338A118962A5A">
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<mods:roleTerm id="A8EB0FD5BB28845AA173CBECD2A32053">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="03CBDA2CD4194EB343294D8834937C29">Ghate, Hemant V.</mods:namePart>
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</mods:name>
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<mods:name id="7FD62DC26B32AD39F9DBC1CE192375E1" type="personal">
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<mods:role id="A3A5DC2A6E58DA2769340A7C89A4C36B">
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<mods:roleTerm id="CE5303898B43D7F302B89EAA136AE5E0">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="F1D2FFE8004DDDA1144AD2321187B191">Hájek, Jiří</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="861A6874170BC57BB51E54080927D92F">text</mods:typeOfResource>
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<mods:titleInfo id="AA81848DE1BD967E95AD21138F2E7C76">
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<mods:title id="4A09B46F4E6FFDE1D197BB1A8FE3DA9E">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="197F20A3C172E4F051CC4209752EC274">
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<mods:date id="4DE677CBA3FA776BA32347E6378568E6">2018</mods:date>
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<mods:detail id="C594463396A18A12691989C4BCB0D714" type="pubDate">
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<mods:number id="B0D4B223BE2178D868885DFF61915937">2018-08-15</mods:number>
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</mods:detail>
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<mods:detail id="85A9F03A5CA73A005814B66B966EBC41" type="volume">
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<mods:number id="2FA33F4FABCD23B64BD29E558742FCB2">4459</mods:number>
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</mods:detail>
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<mods:detail id="DFECCDE7B0635DC2F8F79244A2FF0984" type="issue">
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<mods:number id="7CFB8537EE8EBD4B1D980DC095D60639">2</mods:number>
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<mods:start id="9DFB5F3DDABD5749D68673E962949D12">235</mods:start>
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<mods:classification id="F656F60C7626558ECB9488DA72AB98CA">journal article</mods:classification>
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<mods:identifier id="9877553A1E7BF2189ED786F74CABDA53" type="CLB-Dataset">29024</mods:identifier>
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<mods:identifier id="C8DC64C4A25219E1A9FCE22EB3E2BF5D" type="DOI">10.11646/zootaxa.4459.2.2</mods:identifier>
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<mods:identifier id="0EC0C99B681313475D64B1C0BD63E460" type="GBIF-Dataset">634e23da-84c4-46a4-8bb8-94fcf1637160</mods:identifier>
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<mods:identifier id="31774C979F585EB68FDDD0EB031C1EA7" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="92C0ED509D8865D334F661D9D81ED201" type="Zenodo-Dep">1458548</mods:identifier>
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</mods:mods>
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<treatment id="03BC3747FF92FF99FF5E1396FBC7FAD0" ID-DOI="http://doi.org/10.5281/zenodo.5966529" ID-GBIF-Taxon="148692679" ID-Zenodo-Dep="5966529" LSID="urn:lsid:plazi:treatment:03BC3747FF92FF99FF5E1396FBC7FAD0" httpUri="http://treatment.plazi.org/id/03BC3747FF92FF99FF5E1396FBC7FAD0" lastPageId="10" lastPageNumber="245" pageId="9" pageNumber="244">
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<subSubSection id="C30FD5DAFF92FF9AFF5E1396FEBBFCB8" pageId="9" pageNumber="244" type="nomenclature">
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||
<paragraph id="8BAA8651FF92FF9AFF5E1396FE02FD47" blockId="9.[151,506,725,784]" box="[151,506,725,751]" pageId="9" pageNumber="244">
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<heading id="D0E2313DFF92FF9AFF5E1396FE02FD47" bold="true" box="[151,506,725,751]" fontSize="11" level="1" pageId="9" pageNumber="244" reason="1">
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||
<emphasis id="B9615A43FF92FF9AFF5E1396FE02FD47" bold="true" box="[151,506,725,751]" pageId="9" pageNumber="244">
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||
<emphasis id="B9615A43FF92FF9AFF5E1396FE50FD47" bold="true" box="[151,424,725,751]" italics="true" pageId="9" pageNumber="244">
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||
<taxonomicName id="4C15FDD2FF92FF9AFF5E1396FE6EFD47" ID-CoL="8LZDW" authorityName="Sheth & Ghate & Hájek" authorityYear="2018" box="[151,406,725,751]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="244" phylum="Arthropoda" rank="species" species="maushomi">Copelatus maushomi</taxonomicName>
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s
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</emphasis>
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p. nov.
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BAA8651FF92FF9AFF5E13B4FEBBFCB8" blockId="9.[151,506,725,784]" box="[151,323,759,784]" pageId="9" pageNumber="244">
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||
(
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<figureCitation id="132E9AD4FF92FF9AFF5613B4FF1FFCB8" box="[159,231,759,784]" captionStart="FIGURES 1–4" captionStartId="3.[151,263,1966,1989]" captionTargetBox="[252,1350,214,1928]" captionTargetId="figure@3.[221,1371,195,1943]" captionTargetPageId="3" captionText="FIGURES 1–4. Habitus of Copelatus. 1–2. C. deccanensis sp. nov.: 1. male holotype; 2. male paratype with striolate elytra (Harishchandragad); 3. C. schuhi, male (Lonavala); 4. C. maushomi sp. nov., male holotype." httpUri="https://zenodo.org/record/1458550/files/figure.png" pageId="9" pageNumber="244">Figs 4</figureCitation>
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,
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<figureCitation id="132E9AD4FF92FF9AFF3A13BBFEC2FCB8" box="[243,314,760,784]" captionStart="FIGURES 17–22" captionStartId="7.[151,263,1689,1712]" captionTargetBox="[175,1410,219,1644]" captionTargetId="figure@7.[151,1436,193,1668]" captionTargetPageId="7" captionText="FIGURES 17–22. Male genitalia of Copelatus. 17–18. C. deccanensis sp. nov.; 19–20. C. schuhi; 21–22. C. maushomi sp. nov. 17, 19, 21. median lobe in lateral view; 18, 20, 22. paramere. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/1458558/files/figure.png" pageId="9" pageNumber="244">21–22</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C30FD5DAFF92FF9AFF5E127CFCE4FB98" pageId="9" pageNumber="244" type="materials_examined">
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<paragraph id="8BAA8651FF92FF9AFF5E127CFF33FCD5" blockId="9.[151,1437,831,2009]" pageId="9" pageNumber="244">
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<materialsCitation id="3B7D8C0CFF92FF9AFF5E127CFF33FCD5" ID-GBIF-Occurrence="1917355458" country="India" elevation="600" latitude="19.705" location="Igatpuri" longLatPrecision="127" longitude="73.55167" pageId="9" pageNumber="244" specimenCount="1" stateProvince="Maharashtra" typeStatus="holotype">
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<emphasis id="B9615A43FF92FF9AFF5E127CFEC0FCF0" bold="true" box="[151,312,831,856]" pageId="9" pageNumber="244">
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<typeStatus id="54AE38F3FF92FF9AFF5E127CFF2AFCF0" box="[151,210,831,856]" pageId="9" pageNumber="244">Type</typeStatus>
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locality.
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</emphasis>
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<collectingCountry id="F302C6C1FF92FF9AFE8B127CFE86FCF0" box="[322,382,831,856]" name="India" pageId="9" pageNumber="244">India</collectingCountry>
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,
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<collectingRegion id="49D148B3FF92FF9AFE44127CFDE4FCF0" box="[397,540,831,856]" country="India" name="Maharashtra" pageId="9" pageNumber="244">Maharashtra</collectingRegion>
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,
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||
<locationDeviation id="9A189F73FF92FF9AFDE31203FCC7FCF0" box="[554,831,831,856]" pageId="9" pageNumber="244">
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||
<quantity id="4CED2BB4FF92FF9AFDE31203FD7AFCF0" box="[554,642,831,856]" metricMagnitude="5" metricUnit="m" metricValue="1.2" pageId="9" pageNumber="244" unit="km" value="120.0">120 km</quantity>
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NE of Mumbai
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</locationDeviation>
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,
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<location id="8ECAD08AFF92FF9AFC84127CFC5EFCF0" LSID="urn:lsid:plazi:treatment:03BC3747FF92FF99FF5E1396FBC7FAD0:8ECAD08AFF92FF9AFC84127CFC5EFCF0" box="[845,934,831,856]" country="India" latitude="19.705" longLatPrecision="127" longitude="73.55167" name="Igatpuri" pageId="9" pageNumber="244" stateProvince="Maharashtra">Igatpuri</location>
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environment,
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||
<geoCoordinate id="EE21E096FF92FF9AFB87127CFB3BFCF0" box="[1102,1219,831,856]" direction="north" orientation="latitude" pageId="9" pageNumber="244" precision="92" value="19.705">19°42.3′N</geoCoordinate>
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,
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||
<geoCoordinate id="EE21E096FF92FF9AFB1B127CFABCFCF0" box="[1234,1348,831,856]" direction="east" orientation="longitude" pageId="9" pageNumber="244" precision="92" value="73.55167">73°33.1′E</geoCoordinate>
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,
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<elevation id="00386162FF92FF9AFA9B127CFF33FCD5" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="9" pageNumber="244" unit="m" value="600.0">
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||
<quantity id="4CED2BB4FF92FF9AFA9B127CFA64FCFF" box="[1362,1436,831,856]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="9" pageNumber="244" unit="m" value="600.0">600 m</quantity>
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a.s.l.
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||
</elevation>
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</materialsCitation>
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||
</paragraph>
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<paragraph id="8BAA8651FF92FF9AFF0E12C4FCE4FB98" blockId="9.[151,1437,831,2009]" pageId="9" pageNumber="244">
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||
<emphasis id="B9615A43FF92FF9AFF0E12C4FE8DFC08" bold="true" box="[199,373,903,928]" pageId="9" pageNumber="244">Type material.</emphasis>
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Holotype ♂ (NMPC), labelled: "INDIA occ. centr. / MAHARASHTRA prov. /
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<quantity id="4CED2BB4FF92FF9AFB3212CBFAA9FC08" box="[1275,1361,903,928]" metricMagnitude="5" metricUnit="m" metricValue="1.2" pageId="9" pageNumber="244" unit="km" value="120.0">120 km</quantity>
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NE of MUMBAI / IGATPURI env.,
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||
<quantity id="4CED2BB4FF92FF9AFE2312EFFDD2FC6D" box="[490,554,940,965]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="9" pageNumber="244" unit="m" value="600.0">600m</quantity>
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[printed] // INDIA 2002 Expedition / 19°42.17′N, 73°33.06′E /
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<date id="FFABA091FF92FF9AFACF12EFFF37FC40" pageId="9" pageNumber="244" value="2002-08-01" valueMax="2002-08-12" valueMin="2002-08-01">1. – 12. VIII. 2002</date>
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/ P.Šípek & M.Fikáček leg. [printed] // HOLOTYPE /
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<emphasis id="B9615A43FF92FF9AFCFE1293FBA2FC40" box="[823,1114,975,1000]" italics="true" pageId="9" pageNumber="244">
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<taxonomicName id="4C15FDD2FF92FF9AFCFE1293FC2AFC4F" authority="Sheth & Ghate & Hájek, 2018" authorityName="Sheth & Ghate & Hájek" authorityYear="2018" box="[823,978,976,999]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="244" phylum="Arthropoda" rank="genus" status="sp. nov.">COPELATUS</taxonomicName>
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/
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<taxonomicName id="4C15FDD2FF92FF9AFC2E128CFBA2FC40" authority="Sheth & Ghate & Hájek, 2018" authorityName="Sheth & Ghate & Hájek" authorityYear="2018" box="[999,1114,975,1000]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="244" phylum="Arthropoda" rank="species" species="maushomi" status="sp. nov.">maushomi</taxonomicName>
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</emphasis>
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<taxonomicNameLabel id="A252E738FF92FF9AFBAB1293FB41FC4F" box="[1122,1209,976,999]" pageId="9" pageNumber="244" rank="species">sp. nov.</taxonomicNameLabel>
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/ S. Sheth
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<emphasis id="B9615A43FF92FF9AFAFA1292FA9DFC40" box="[1331,1381,975,1000]" italics="true" pageId="9" pageNumber="244">et al</emphasis>
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. det. 2016 [red label, printed]" (NMPC). Paratypes: 4♂, 1♀ same data as holotype (LHCM, NMPC, ZSMG). Each paratype is provided with the respective red printed label.
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</paragraph>
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</subSubSection>
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<subSubSection id="C30FD5DAFF92FF99FF0E157FFE2CFC28" lastPageId="10" lastPageNumber="245" pageId="9" pageNumber="244" type="description">
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<paragraph id="8BAA8651FF92FF9AFF0E157FFDCCFB35" blockId="9.[151,1437,831,2009]" pageId="9" pageNumber="244">
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<emphasis id="B9615A43FF92FF9AFF0E157FFD66FBFC" bold="true" box="[199,670,1083,1109]" pageId="9" pageNumber="244">
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Description of male
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<typeStatus id="54AE38F3FF92FF9AFE01157FFDC9FBFD" box="[456,561,1084,1109]" pageId="9" pageNumber="244" type="holotype">holotype</typeStatus>
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.
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<emphasis id="B9615A43FF92FF9AFD8B1578FD66FBFC" bold="true" box="[578,670,1083,1108]" italics="true" pageId="9" pageNumber="244">Habitus</emphasis>
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||
</emphasis>
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||
(
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<figureCitation id="132E9AD4FF92FF9AFD7A157FFD04FBFC" box="[691,764,1084,1109]" captionStart="FIGURES 1–4" captionStartId="3.[151,263,1966,1989]" captionTargetBox="[252,1350,214,1928]" captionTargetId="figure@3.[221,1371,195,1943]" captionTargetPageId="3" captionText="FIGURES 1–4. Habitus of Copelatus. 1–2. C. deccanensis sp. nov.: 1. male holotype; 2. male paratype with striolate elytra (Harishchandragad); 3. C. schuhi, male (Lonavala); 4. C. maushomi sp. nov., male holotype." httpUri="https://zenodo.org/record/1458550/files/figure.png" pageId="9" pageNumber="244">Fig. 4</figureCitation>
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) elongate oblong oval, nearly parallel sided; outline not continuous as pronotal posterior corners protrude; broadest in basal third of pronotum; very slightly convex. Dorsal surface matt due to dense striolation.
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</paragraph>
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<paragraph id="8BAA8651FF92FF9AFF0E15E4FD51FAA0" blockId="9.[151,1437,831,2009]" pageId="9" pageNumber="244">
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||
<emphasis id="B9615A43FF92FF9AFF0E15E4FEB2FB68" bold="true" box="[199,330,1191,1216]" italics="true" pageId="9" pageNumber="244">Coloration.</emphasis>
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||
Dorsally almost uniformly testaceous; head slightly darker than pronotum and elytra, infuscate posterior to eyes; pronotum indistinctly infuscate on disc; elytra laterally and apically somewhat paler; appendages testaceous. Ventral part testaceous to brownish.
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||
</paragraph>
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||
<paragraph id="8BAA8651FF92FF9AFF0E1450FE9EF980" blockId="9.[151,1437,831,2009]" pageId="9" pageNumber="244">
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||
<emphasis id="B9615A43FF92FF9AFF0E1450FEF4FA85" bold="true" box="[199,268,1299,1325]" pageId="9" pageNumber="244">
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||
<emphasis id="B9615A43FF92FF9AFF0E1450FEFDFA84" bold="true" box="[199,261,1299,1324]" italics="true" pageId="9" pageNumber="244">Head</emphasis>
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.
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</emphasis>
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Moderately broad, ca. 0.6× width of pronotum, almost semicircular. Labrum medially emarginate. Anterior margin of clypeus slightly concave. Antennae with antennomeres slender, club-shaped, antennomere I longest. Eyes emarginate anterolaterally, small, eye width only ca. 0.1× width of head. Reticulation consisting of well impressed polygonal meshes; meshes slightly larger in anterior region. Rather long, longitudinal or oblique strioles present between eyes and on vertex. Punctation double; several large setigerous punctures present in fronto-clypeal depressions, frontal depressions at level of anterior margin of eyes, and in depressions along inner margin of eyes; very fine and sparsely distributed punctures placed among meshes of microreticulation, punctures denser posteriorly.
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</paragraph>
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<paragraph id="8BAA8651FF92FF9AFF0E1777FC99F88D" blockId="9.[151,1437,831,2009]" pageId="9" pageNumber="244">
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<emphasis id="B9615A43FF92FF9AFF0E1777FEBBF9E5" bold="true" box="[199,323,1588,1613]" pageId="9" pageNumber="244">
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<emphasis id="B9615A43FF92FF9AFF0E1777FEC4F9E3" bold="true" box="[199,316,1588,1611]" italics="true" pageId="9" pageNumber="244">Pronotum</emphasis>
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.
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</emphasis>
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Transverse, broadest in basal third. Anterior angles acute, posterior angles rectangular. Sides largely and evenly curved, with lateral beading very thin and indistinct. Anterior margin straight, posterior margin sinuate. Surface reticulation consisting of polygonal meshes, similar to that of head, but slightly less impressed. Disc of pronotum completely longitudinally striolate; strioles mostly long, well impressed, rarely confluent; few short, shallow strioles present between long strioles. Punctation double; row of coarse setigerous punctures present along anterior margin, basal margin (except medially), and laterally close to sides; fine punctures placed among meshes of microreticulation. Scutellar shield broadly triangular.
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</paragraph>
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<paragraph id="8BAA8651FF92FF9AFF0E166CFDFDF871" blockId="9.[151,1437,831,2009]" pageId="9" pageNumber="244">
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<emphasis id="B9615A43FF92FF9AFF0E166CFEF7F8E0" bold="true" box="[199,271,1839,1864]" italics="true" pageId="9" pageNumber="244">Elytra</emphasis>
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. Elytral striation consisting of nine complete shallow discal striae; striae almost imperceptible due to dense striolation of elytra. Strioles very long, rarely confluent. Surface reticulation consisting of fine, shallowly impressed isodiametric polygonal meshes. Punctation consisting of setigerous punctures only, few punctures present along elytral striae, but predominantly apically and along lateral margin of elytra; fine punctures, due to dense striolation not perceptible.
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</paragraph>
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<paragraph id="8BAA8651FF91FF99FF0E11DBFB36FF7D" blockId="10.[151,1437,151,1401]" pageId="10" pageNumber="245">
|
||
<emphasis id="B9615A43FF91FF99FF0E11DBFEFDFF07" bold="true" box="[199,261,152,175]" italics="true" pageId="10" pageNumber="245">Legs.</emphasis>
|
||
Protibia modified, angled near base, distinctly broadened anteriorly, club shaped. Pro- and mesotarsomeres 1–3 distinctly broadened, with four rows of adhesive setae on their ventral side.
|
||
</paragraph>
|
||
<paragraph id="8BAA8651FF91FF99FF0E119CFDCDFD95" blockId="10.[151,1437,151,1401]" pageId="10" pageNumber="245">
|
||
<emphasis id="B9615A43FF91FF99FF0E119CFEAEFF50" bold="true" box="[199,342,223,248]" italics="true" pageId="10" pageNumber="245">Ventral side.</emphasis>
|
||
Prosternum sinuate anteriorly, obtusely keeled medially. Prosternal process shortly lanceolate, in cross-section convex, apex rounded; distinctly bordered laterally; reticulation or punctation absent. Metaventrite with microsculpture consisting of polygonal meshes; punctation imperceptible. Lateral parts of metaventrite ('metasternal wings') tongue-shaped, slender. Metacoxal lines well impressed, incomplete—absent in anterior fourth. Metacoxal plates covered with deep, longitudinal or oblique strioles; reticulation consisting of elongate, longitudinal polygonal meshes. Punctation on metacoxae absent. Metacoxal processes rounded and incised at posterior margin. Abdominal ventrites I–II with longitudinal strioles; ventrites III–IV with oblique strioles laterally, absent medially. Abdominal reticulation consisting of elongate polygonal meshes, longitudinal on ventrites I–II, oblique on ventrite III and transverse on ventrites IV–VI. Punctation consisting of fine punctures medially, and larger and deeper punctures laterally.
|
||
</paragraph>
|
||
<paragraph id="8BAA8651FF91FF99FF0E1304FB99FD00" blockId="10.[151,1437,151,1401]" pageId="10" pageNumber="245">
|
||
<emphasis id="B9615A43FF91FF99FF0E1304FE80FDC8" bold="true" box="[199,376,583,608]" italics="true" pageId="10" pageNumber="245">
|
||
<collectingRegion id="49D148B3FF91FF99FF0E1304FEFAFDC8" box="[199,258,583,608]" country="Maldives" name="Maale" pageId="10" pageNumber="245">Male</collectingRegion>
|
||
genitalia.
|
||
</emphasis>
|
||
Median lobe in lateral aspect almost evenly curved; narrowing from base to pointed apex; broadest in middle (
|
||
<figureCitation id="132E9AD4FF91FF99FE4D132FFE24FD2C" box="[388,476,620,645]" captionStart="FIGURES 17–22" captionStartId="7.[151,263,1689,1712]" captionTargetBox="[175,1410,219,1644]" captionTargetId="figure@7.[151,1436,193,1668]" captionTargetPageId="7" captionText="FIGURES 17–22. Male genitalia of Copelatus. 17–18. C. deccanensis sp. nov.; 19–20. C. schuhi; 21–22. C. maushomi sp. nov. 17, 19, 21. median lobe in lateral view; 18, 20, 22. paramere. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/1458558/files/figure.png" pageId="10" pageNumber="245">Fig. 21</figureCitation>
|
||
). A fold present till subapical region. Parameres more or less 'D'-shaped, slightly sinuate on outer margin, apex very narrow and long; apical lobe club-shaped (
|
||
<figureCitation id="132E9AD4FF91FF99FBC913CCFBACFD00" box="[1024,1108,655,680]" captionStart="FIGURES 17–22" captionStartId="7.[151,263,1689,1712]" captionTargetBox="[175,1410,219,1644]" captionTargetId="figure@7.[151,1436,193,1668]" captionTargetPageId="7" captionText="FIGURES 17–22. Male genitalia of Copelatus. 17–18. C. deccanensis sp. nov.; 19–20. C. schuhi; 21–22. C. maushomi sp. nov. 17, 19, 21. median lobe in lateral view; 18, 20, 22. paramere. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/1458558/files/figure.png" pageId="10" pageNumber="245">Fig. 22</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BAA8651FF91FF99FF0E13F7FC23FD47" blockId="10.[151,1437,151,1401]" pageId="10" pageNumber="245">
|
||
<emphasis id="B9615A43FF91FF99FF0E13F7FEDDFD65" bold="true" box="[199,293,692,717]" pageId="10" pageNumber="245">Female.</emphasis>
|
||
Females do not differ in external morphology from male except for nearly straight, apically less broadened protibia, and slender pro- and mesotarsi without adhesive setae.
|
||
</paragraph>
|
||
<paragraph id="8BAA8651FF91FF99FF0E13BFFEC3FC9F" blockId="10.[151,1437,151,1401]" pageId="10" pageNumber="245">
|
||
<emphasis id="B9615A43FF91FF99FF0E13BFFEB5FCBD" bold="true" box="[199,333,764,789]" pageId="10" pageNumber="245">Variability.</emphasis>
|
||
All specimens of the
|
||
<typeStatus id="54AE38F3FF91FF99FD8C13BEFD8DFCBD" box="[581,629,765,789]" pageId="10" pageNumber="245">type</typeStatus>
|
||
series are rather uniform and vary only in extent of infuscation of head and pronotum.
|
||
</paragraph>
|
||
<paragraph id="8BAA8651FF91FF99FF0E1207FE2CFC28" blockId="10.[151,1437,151,1401]" pageId="10" pageNumber="245">
|
||
<emphasis id="B9615A43FF91FF99FF0E1207FE8EFCF5" bold="true" box="[199,374,836,861]" pageId="10" pageNumber="245">Measurements</emphasis>
|
||
(N=5). TL: 4.6–5.0 mm (
|
||
<typeStatus id="54AE38F3FF91FF99FD5F1207FD06FCF5" box="[662,766,836,861]" pageId="10" pageNumber="245" type="holotype">holotype</typeStatus>
|
||
:
|
||
<quantity id="4CED2BB4FF91FF99FCC11207FC9AFCF4" box="[776,866,836,860]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="10" pageNumber="245" unit="mm" value="4.8">4.8 mm</quantity>
|
||
); Tl-h:
|
||
<quantity id="4CED2BB4FF91FF99FC7C1207FBC3FCF4" box="[949,1083,836,861]" metricMagnitude="-3" metricUnit="m" metricValue="4.35" metricValueMax="4.5" metricValueMin="4.2" pageId="10" pageNumber="245" unit="mm" value="4.35" valueMax="4.5" valueMin="4.2">4.2–4.5 mm</quantity>
|
||
(
|
||
<typeStatus id="54AE38F3FF91FF99FB831207FB4AFCF5" box="[1098,1202,836,861]" pageId="10" pageNumber="245" type="holotype">holotype</typeStatus>
|
||
:
|
||
<quantity id="4CED2BB4FF91FF99FB751207FAEEFCF4" box="[1212,1302,836,860]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="10" pageNumber="245" unit="mm" value="4.4">4.4 mm</quantity>
|
||
); MW: 2.0–
|
||
<quantity id="4CED2BB4FF91FF99FF5E122BFF15FCD7" box="[151,237,872,896]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="10" pageNumber="245" unit="mm" value="2.1">2.1 mm</quantity>
|
||
(
|
||
<typeStatus id="54AE38F3FF91FF99FF321224FE9BFC28" box="[251,355,871,896]" pageId="10" pageNumber="245" type="holotype">holotype</typeStatus>
|
||
: 2.0 mm).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30FD5DAFF91FF99FF0E12CFFB27FBD4" pageId="10" pageNumber="245" type="diagnosis">
|
||
<paragraph id="8BAA8651FF91FF99FF0E12CFFB24FC44" blockId="10.[151,1437,151,1401]" pageId="10" pageNumber="245">
|
||
<emphasis id="B9615A43FF91FF99FF0E12CFFE2AFC0D" bold="true" box="[199,466,908,933]" pageId="10" pageNumber="245">Differential diagnosis.</emphasis>
|
||
Based on the presence of nine dorsal striae on the elytra, the new species can be tentatively classified within the
|
||
<taxonomicName id="4C15FDD2FF91FF99FE3412ECFD35FC60" authorityName="Sharp" authorityYear="1882" box="[509,717,943,968]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="245" phylum="Arthropoda" rank="species" species="consors">
|
||
<emphasis id="B9615A43FF91FF99FE3412ECFD35FC60" box="[509,717,943,968]" italics="true" pageId="10" pageNumber="245">Copelatus consors</emphasis>
|
||
</taxonomicName>
|
||
species group sensu
|
||
<bibRefCitation id="EF84FBA0FF91FF99FC7212ECFB94FC60" box="[955,1132,943,968]" pageId="10" pageNumber="245" refId="ref15617" refString="Guignot, F. (1961) Revision des hydrocanthares d'Afrique (Coleoptera Dityscoidea [sic!]). Troisieme partie. Annales du Musee Royal du Congo Belge, Serie 8 vo (Sciences Zoologiques), 90, 659 - 995." type="journal article">Guignot (1961)</bibRefCitation>
|
||
. This group so far contains eighteen species:
|
||
<quantity id="4CED2BB4FF91FF99FE951297FE6CFC45" box="[348,404,980,1005]" metricMagnitude="-1" metricUnit="m" metricValue="2.794" pageId="10" pageNumber="245" unit="in" value="11.0">11 in</quantity>
|
||
the Afrotropical and seven in the Nearctic region (
|
||
<bibRefCitation id="EF84FBA0FF91FF99FC071297FB28FC44" box="[974,1232,980,1005]" pageId="10" pageNumber="245" refId="ref16087" refString="Nilsson, A. N. & Hajek, J. (2018) A world catalogue of the family Dytiscidae, or the diving beetles (Coleoptera, Adephaga). Uersion 1. I. 2018. Distributed as a PDF file via Internet. Available from: http: // www. waterbeetles. eu (accessed 10 May 2018)" type="url">Nilsson & Hájek 2018</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BAA8651FF91FF99FF0E12B4FB27FBD4" blockId="10.[151,1437,151,1401]" pageId="10" pageNumber="245">
|
||
<taxonomicName id="4C15FDD2FF91FF99FF0E12B4FE4CFBB8" authority="Sheth & Ghate & Hájek, 2018" authorityName="Sheth & Ghate & Hájek" authorityYear="2018" box="[199,436,1015,1040]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="245" phylum="Arthropoda" rank="species" species="maushomi" status="sp. nov.">
|
||
<emphasis id="B9615A43FF91FF99FF0E12B4FE4CFBB8" box="[199,436,1015,1040]" italics="true" pageId="10" pageNumber="245">Copelatus maushomi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A252E738FF91FF99FE7512B4FDEEFBB8" box="[444,534,1015,1040]" pageId="10" pageNumber="245" rank="species">
|
||
<emphasis id="B9615A43FF91FF99FE7512B4FDEEFBB8" bold="true" box="[444,534,1015,1040]" pageId="10" pageNumber="245">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
does not seem to be related to any species of the
|
||
<taxonomicName id="4C15FDD2FF91FF99FB8712BBFB3FFBB8" authorityName="Sharp" authorityYear="1882" box="[1102,1223,1016,1040]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="245" phylum="Arthropoda" rank="species" species="consors">
|
||
<emphasis id="B9615A43FF91FF99FB8712BBFB3FFBB8" box="[1102,1223,1016,1040]" italics="true" pageId="10" pageNumber="245">C. consors</emphasis>
|
||
</taxonomicName>
|
||
group. With small eyes, pronotum distinctly broader than elytra, and elytra with dense striolation, the new species has very unique appearance within all known
|
||
<taxonomicName id="4C15FDD2FF91FF99FE2D157CFDAEFBF0" authorityName="Sheth & Ghate & Hájek" authorityYear="2018" box="[484,598,1087,1112]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="245" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9615A43FF91FF99FE2D157CFDAEFBF0" box="[484,598,1087,1112]" italics="true" pageId="10" pageNumber="245">Copelatus</emphasis>
|
||
</taxonomicName>
|
||
species. The shape of the male median lobe suggests that the species may be related to Indian species of the
|
||
<taxonomicName id="4C15FDD2FF91FF99FDDF1526FD34FBD5" authorityName="Sharp" authorityYear="1882" box="[534,716,1124,1149]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="245" phylum="Arthropoda" rank="species" species="nigrolineatus">
|
||
<emphasis id="B9615A43FF91FF99FDDF1526FD34FBD5" box="[534,716,1124,1149]" italics="true" pageId="10" pageNumber="245">C. nigrolineatus</emphasis>
|
||
</taxonomicName>
|
||
group—
|
||
<taxonomicName id="4C15FDD2FF91FF99FCE61526FC2FFBD5" authority="Sheth & Ghate & Hájek, 2018" authorityName="Sheth & Ghate & Hájek" authorityYear="2018" box="[815,983,1124,1149]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="245" phylum="Arthropoda" rank="species" species="deccanensis" status="sp. nov.">
|
||
<emphasis id="B9615A43FF91FF99FCE61526FC2FFBD5" box="[815,983,1124,1149]" italics="true" pageId="10" pageNumber="245">C. deccanensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A252E738FF91FF99FC171527FBC0FBD5" box="[990,1080,1124,1149]" pageId="10" pageNumber="245" rank="species">
|
||
<emphasis id="B9615A43FF91FF99FC171527FBC0FBD5" bold="true" box="[990,1080,1124,1149]" pageId="10" pageNumber="245">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C15FDD2FF91FF99FBA71526FB20FBD5" authorityName="Hendrich & Balke" authorityYear="1998" box="[1134,1240,1124,1149]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="245" phylum="Arthropoda" rank="species" species="schuhi">
|
||
<emphasis id="B9615A43FF91FF99FBA71526FB20FBD5" box="[1134,1240,1124,1149]" italics="true" pageId="10" pageNumber="245">C. schuhi</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30FD5DAFF91FF99FF0E15C4FC20FAFC" pageId="10" pageNumber="245" type="etymology">
|
||
<paragraph id="8BAA8651FF91FF99FF0E15C4FABFFB6C" blockId="10.[151,1437,151,1401]" pageId="10" pageNumber="245">
|
||
<emphasis id="B9615A43FF91FF99FF0E15C4FEB6FB08" bold="true" box="[199,334,1159,1184]" pageId="10" pageNumber="245">Etymology.</emphasis>
|
||
The species is named after the 'maushom'—a local name for the monsoon, indicating that the specimens were collected at the beginning of the monsoon season. The name is a noun in the genitive case.
|
||
</paragraph>
|
||
<paragraph id="8BAA8651FF91FF99FF0E158CFC20FAFC" blockId="10.[151,1437,151,1401]" pageId="10" pageNumber="245">
|
||
<emphasis id="B9615A43FF91FF99FF0E158CFE00FB40" bold="true" box="[199,504,1231,1256]" pageId="10" pageNumber="245">Collecting circumstances.</emphasis>
|
||
The specimens were collected in small deep pools in a stony stream below a table mountain (
|
||
<figureCitation id="132E9AD4FF91FF99FEDB15B7FE90FAA4" box="[274,360,1268,1293]" captionStart="FIGURES 41–44" captionStartId="19.[151,263,1551,1574]" captionTargetBox="[151,1436,195,1529]" captionTargetId="figure@19.[150,1436,195,1530]" captionTargetPageId="19" captionText="FIGURES 41–44. Habitat of Copelatus. 41. ephemeral puddle (Tamhini); 42. rock puddle (Harishchandragad); 43. puddle (Tamhini); 44. mountain stream (Igatpuri area; photo M. Fikáček)." httpUri="https://zenodo.org/record/1458566/files/figure.png" pageId="10" pageNumber="245">Fig. 44</figureCitation>
|
||
). The place was visited at the beginning of the monsoon. Sudden large amount of water could have brought the specimens to the normal stream from less accessible habitat, e.g. wet gravels on the stream bottom or other interstitial water habitats (M. Fikáček, pers. comm. 2017).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30FD5DAFF91FF99FF0E141CFBC7FAD0" box="[199,1087,1375,1401]" pageId="10" pageNumber="245" type="distribution">
|
||
<paragraph id="8BAA8651FF91FF99FF0E141CFBC7FAD0" blockId="10.[151,1437,151,1401]" box="[199,1087,1375,1401]" pageId="10" pageNumber="245">
|
||
<emphasis id="B9615A43FF91FF99FF0E141CFE98FAD0" bold="true" box="[199,352,1375,1400]" pageId="10" pageNumber="245">Distribution.</emphasis>
|
||
The species is so far known only from the
|
||
<typeStatus id="54AE38F3FF91FF99FC8D1422FC8CFAD1" box="[836,884,1377,1401]" pageId="10" pageNumber="245">type</typeStatus>
|
||
locality (
|
||
<figureCitation id="132E9AD4FF91FF99FC17141CFBCBFAD0" box="[990,1075,1375,1400]" captionStart="FIGURE 45" captionStartId="21.[151,249,1915,1937]" captionTargetBox="[190,1401,196,1869]" captionTargetId="figure@21.[153,1434,158,1905]" captionTargetPageId="21" captionText="FIGURE 45. Distributional map of Copelatus species in Maharashtra." httpUri="https://zenodo.org/record/1458568/files/figure.png" pageId="10" pageNumber="245">Fig. 45</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |