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<document id="5B1D2B9F89F6F71B451F6C6246A24087" ID-CLB-Dataset="26919" ID-DOI="10.11646/zootaxa.4598.1.1" ID-GBIF-Dataset="475fd53e-c5c6-4a99-bb2e-cc2382f0171f" ID-ISSN="1175-5326" ID-Zenodo-Dep="2668063" ID-ZooBank="6BA5F086-1014-46B5-B6D5-77FE0CD07AEC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1557127411994" checkinUser="plazi" docAuthor="Seldon, David S. &amp; Buckley, Thomas R." docDate="2019" docId="03BBD0260223B72690FBB3F3E873FBD8" docLanguage="en" docName="zootaxa.4598.1.1.pdf.imf" docOrigin="Zootaxa 4598 (1)" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Mecodema curvidens Broun 1915" docType="treatment" docVersion="11" lastPageNumber="41" masterDocId="FF82A85E0205B70E906CB267ED0BFFA4" masterDocTitle="The genus Mecodema Blanchard 1853 (Coleoptera: Carabidae: Broscini) from the North Island, New Zealand" masterLastPageNumber="148" masterPageNumber="1" pageNumber="39" updateTime="1698726038790" updateUser="ExternalLinkService">
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<mods:title id="D14CAD5E31C99896E26997E76FC73439">The genus Mecodema Blanchard 1853 (Coleoptera: Carabidae: Broscini) from the North Island, New Zealand</mods:title>
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<mods:namePart id="35221499666FA1185985DCC8385611BD">Seldon, David S.</mods:namePart>
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<mods:namePart id="FD131EF1D24BD49E9A1775DED53FEBB4">Buckley, Thomas R.</mods:namePart>
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<treatment id="03BBD0260223B72690FBB3F3E873FBD8" ID-DOI="http://doi.org/10.5281/zenodo.5678369" ID-GBIF-Taxon="157062665" ID-Zenodo-Dep="5678369" LSID="urn:lsid:plazi:treatment:03BBD0260223B72690FBB3F3E873FBD8" httpUri="http://treatment.plazi.org/id/03BBD0260223B72690FBB3F3E873FBD8" lastPageId="40" lastPageNumber="41" pageId="38" pageNumber="39">
<subSubSection id="C30832BB0223B72890FBB3F3EF35FE0A" box="[151,574,404,430]" pageId="38" pageNumber="39" type="nomenclature">
<paragraph id="8BAD61300223B72890FBB3F3EF35FE0A" blockId="38.[151,574,404,463]" box="[151,574,404,430]" pageId="38" pageNumber="39">
<heading id="D0E5D65C0223B72890FBB3F3EF35FE0A" bold="true" box="[151,574,404,430]" fontSize="11" level="1" pageId="38" pageNumber="39" reason="1">
<emphasis id="B966BD220223B72890FBB3F3EF35FE0A" bold="true" box="[151,574,404,430]" pageId="38" pageNumber="39">
<taxonomicName id="4C121AB30223B72890FBB3F3EF31FE0A" ID-CoL="3YKSD" authority="Broun, 1915" authorityName="Broun" authorityYear="1915" box="[151,570,404,430]" class="Insecta" family="Carabidae" genus="Mecodema" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="curvidens">
<emphasis id="B966BD220223B72890FBB3F3EC9CFE0A" bold="true" box="[151,407,404,430]" italics="true" pageId="38" pageNumber="39">Mecodema curvidens</emphasis>
<bibRefCitation id="EF831CC10223B72891F2B3F3EF31FE0A" author="Broun, T." box="[414,570,404,430]" pageId="38" pageNumber="39" pagination="267 - 346" refId="ref91488" refString="Broun, T. (1915) Descriptions of new genera and species of Coleoptera. [Part IV]. Bulletin of the New Zealand Institute, 1, 267 - 346." type="journal article" year="1915">Broun, 1915</bibRefCitation>
</taxonomicName>
.
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30832BB0223B72890FBB3D1EC77FE6B" box="[151,380,438,463]" pageId="38" pageNumber="39" type="description">
<paragraph id="8BAD61300223B72890FBB3D1EC77FE6B" blockId="38.[151,574,404,463]" box="[151,380,438,463]" pageId="38" pageNumber="39">
<figureCitation id="13297DB50223B72890FBB3D1EC29FE6B" box="[151,290,438,463]" captionStart="FIGURE 12" captionStartId="18.[151,250,1905,1927]" captionTargetBox="[311,1244,697,1878]" captionTargetId="figure@18.[310,1250,689,1889]" captionTargetPageId="18" captionText="FIGURE 12. Ventral view of the mentum with a number of different character states. Illustrations AD are the different forms of the indentation in the apex of the median process: (A) distinctly indentate (M. manaia); (B) moderately indentate (M. pluto); (C) slightly indentate (M. infimate); (D) indentation notched (M. parataiko). Illustrations EG are the three general forms (i.e., hashed lines) of the mentum lobes: (E) triangular; (F) rounded; (G) squared." httpUri="https://zenodo.org/record/2668087/files/figure.png" pageId="38" pageNumber="39">Figures 12G</figureCitation>
and
<figureCitation id="13297DB50223B7289134B3D0EC7CFE6B" box="[344,375,439,463]" captionStart="FIGURE 24" captionStartId="39.[151,250,1867,1889]" captionTargetBox="[251,1341,201,1832]" captionTargetId="figure@39.[238,1349,195,1846]" captionTargetPageId="39" captionText="FIGURE 24. Mecodema curvidens plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668111/files/figure.png" pageId="38" pageNumber="39">24</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30832BB0223B72890FBB399E919FDF6" pageId="38" pageNumber="39" type="reference_group">
<paragraph id="8BAD61300223B72890FBB399EED2FDB0" blockId="38.[151,1042,510,594]" box="[151,985,510,532]" pageId="38" pageNumber="39">
<taxonomicName id="4C121AB30223B72890FBB399EF02FDB0" ID-CoL="429YX" authority="Broun, 1915" authorityName="Broun" authorityYear="1915" box="[151,521,510,532]" class="Insecta" family="Carabidae" genus="Metaglymma" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="curvidens">
<emphasis id="B966BD220223B72890FBB399EC8FFDB0" box="[151,388,510,532]" italics="true" pageId="38" pageNumber="39">Metaglymma curvidens</emphasis>
<bibRefCitation id="EF831CC10223B72891E6B398EF02FDB0" author="Broun, T." box="[394,521,510,532]" pageId="38" pageNumber="39" pagination="267 - 346" refId="ref91488" refString="Broun, T. (1915) Descriptions of new genera and species of Coleoptera. [Part IV]. Bulletin of the New Zealand Institute, 1, 267 - 346." type="journal article" year="1915">Broun, 1915</bibRefCitation>
</taxonomicName>
(Opotiki, BP), synonymised by
<bibRefCitation id="EF831CC10223B728933DB399EEDEFDB0" author="Britton, E. B." box="[849,981,510,532]" pageId="38" pageNumber="39" pagination="533 - 581" refId="ref90973" refString="Britton, E. B. (1949) The Carabidae (Coleoptera) of New Zealand part 3: A revision of the tribe Broscini. Transactions of the Royal Society of New Zealand, 77, 533 - 581." type="journal article" year="1949">Britton 1949</bibRefCitation>
.
</paragraph>
<paragraph id="8BAD61300223B72890FBB07AEE4CFD97" blockId="38.[151,1042,510,594]" box="[151,839,541,563]" pageId="38" pageNumber="39">
<taxonomicName id="4C121AB30223B72890FBB07AECF0FD97" ID-CoL="8LJ36" authority="Broun, 1923" authorityName="Broun" authorityYear="1923" box="[151,507,541,563]" class="Insecta" family="Carabidae" genus="Mecodema" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="occiputale">
<emphasis id="B966BD220223B72890FBB07AEC7EFD97" box="[151,373,541,563]" italics="true" pageId="38" pageNumber="39">Mecodema occiputale</emphasis>
<bibRefCitation id="EF831CC10223B7289110B079ECF0FD97" author="Broun, T." box="[380,507,541,563]" pageId="38" pageNumber="39" pagination="667 - 708" refId="ref91555" refString="Broun, T. (1923) Descriptions of new genera and species of Coleoptera. [Part VIII]. Bulletin of the New Zealand Institute, 1, 667 - 708." type="journal article" year="1923">Broun, 1923</bibRefCitation>
</taxonomicName>
(Tokaanu, TO),
<emphasis id="B966BD220223B72892CFB07AEE4AFD97" bold="true" box="[675,833,541,563]" pageId="38" pageNumber="39">new synonymy</emphasis>
.
</paragraph>
<paragraph id="8BAD61300223B72890FBB05BE919FDF6" blockId="38.[151,1042,510,594]" box="[151,1042,572,594]" pageId="38" pageNumber="39">
<taxonomicName id="4C121AB30223B72890FBB05BECF4FDF6" authority="Brookes, 1926" authorityName="Brookes" authorityYear="1926" box="[151,511,572,594]" class="Insecta" family="Carabidae" genus="Mecodema" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="exitiosus">
<emphasis id="B966BD220223B72890FBB05BEC6EFDF6" box="[151,357,572,594]" italics="true" pageId="38" pageNumber="39">Mecodema exitiosus</emphasis>
<bibRefCitation id="EF831CC10223B7289107B05BECF4FDF6" author="Brookes, A. E." box="[363,511,572,594]" pageId="38" pageNumber="39" pagination="440 - 448" refId="ref91052" refString="Brookes, A. E. (1926) A new genus and eight new species of Coleoptera. Transactions and Proceedings of the New Zealand Institute, 56, 440 - 448." type="journal article" year="1926">Brookes, 1926</bibRefCitation>
</taxonomicName>
(nr. Matamata, WO), synonymised by
<bibRefCitation id="EF831CC10223B72893E6B05BE904FDF6" author="Britton, E. B." box="[906,1039,572,594]" pageId="38" pageNumber="39" pagination="533 - 581" refId="ref90973" refString="Britton, E. B. (1949) The Carabidae (Coleoptera) of New Zealand part 3: A revision of the tribe Broscini. Transactions of the Royal Society of New Zealand, 77, 533 - 581." type="journal article" year="1949">Britton 1949</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30832BB0223B72890FBB019E9C3FD7B" pageId="38" pageNumber="39" type="diagnosis">
<paragraph id="8BAD61300223B72890FBB019E9C3FD7B" blockId="38.[151,1437,638,1998]" pageId="38" pageNumber="39">
<emphasis id="B966BD220223B72890FBB019EC18FD33" bold="true" box="[151,275,638,663]" pageId="38" pageNumber="39">Diagnosis:</emphasis>
Distinguished from other North Island
<taxonomicName id="4C121AB30223B72892BEB019EE46FD32" authorityName="Blanchard" authorityYear="1853" box="[722,845,638,662]" class="Insecta" family="Carabidae" genus="Mecodema" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="B966BD220223B72892BEB019EE46FD32" box="[722,845,638,662]" italics="true" pageId="38" pageNumber="39">Mecodema</emphasis>
</taxonomicName>
species by having:
<emphasis id="B966BD220223B7289440B019E931FD33" bold="true" box="[1068,1082,638,663]" pageId="38" pageNumber="39">1</emphasis>
, pronotal carina with 46 setae each side (
<taxonomicName id="4C121AB30223B7289178B0C4EC8AFD1F" authorityName="Broun" authorityYear="1915" box="[276,385,675,699]" class="Insecta" family="Carabidae" genus="Metaglymma" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="curvidens">
<emphasis id="B966BD220223B7289178B0C4EC8AFD1F" box="[276,385,675,699]" italics="true" pageId="38" pageNumber="39">curvidens</emphasis>
</taxonomicName>
species group);
<emphasis id="B966BD220223B728922CB0C5EF45FD1F" bold="true" box="[576,590,674,699]" pageId="38" pageNumber="39">2</emphasis>
, a vertexal groove that is narrow and defined entire length;
<emphasis id="B966BD220223B7289565B0C5E81CFD1F" bold="true" box="[1289,1303,674,699]" pageId="38" pageNumber="39">3</emphasis>
, distinctive form of the basal lobe and setal distribution along ventral edge of the left paramere (
<figureCitation id="13297DB50223B728942BB0A1E992FD7B" box="[1095,1177,710,735]" captionStart="FIGURE 24" captionStartId="39.[151,250,1867,1889]" captionTargetBox="[251,1341,201,1832]" captionTargetId="figure@39.[238,1349,195,1846]" captionTargetPageId="39" captionText="FIGURE 24. Mecodema curvidens plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668111/files/figure.png" pageId="38" pageNumber="39">Fig. 24</figureCitation>
LP).
</paragraph>
</subSubSection>
<subSubSection id="C30832BB0223B72690ABB08DEC10FE23" lastPageId="40" lastPageNumber="41" pageId="38" pageNumber="39" type="description">
<paragraph id="8BAD61300223B72890ABB08DEEC2FC83" blockId="38.[151,1437,638,1998]" pageId="38" pageNumber="39">
<emphasis id="B966BD220223B72890ABB08DEC50FCA7" bold="true" box="[199,347,746,771]" pageId="38" pageNumber="39">Description:</emphasis>
Length
<quantity id="4CEACCD50223B72891D7B08DEF33FCA6" box="[443,568,746,771]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" metricValueMax="2.6" metricValueMin="1.8" pageId="38" pageNumber="39" unit="mm" value="22.0" valueMax="26.0" valueMin="18.0">1826 mm</quantity>
, pronotal width
<quantity id="4CEACCD50223B7289298B08DEE87FCA6" box="[756,908,746,771]" metricMagnitude="-3" metricUnit="m" metricValue="6.465" metricValueMax="7.53" metricValueMin="5.4" pageId="38" pageNumber="39" unit="mm" value="6.465" valueMax="7.53" valueMin="5.4">5.47.53 mm</quantity>
, elytral width
<quantity id="4CEACCD50223B728945EB08DE9D1FCA6" box="[1074,1242,746,771]" metricMagnitude="-3" metricUnit="m" metricValue="7.345" metricValueMax="8.56" metricValueMin="6.13" pageId="38" pageNumber="39" unit="mm" value="7.345" valueMax="8.56" valueMin="6.13">6.138.56 mm</quantity>
. Colour of entire body glossy to matte black, except coxae to tarsi reddish-brown to black.
</paragraph>
<paragraph id="8BAD61300223B72890ABB154EF4CFBCE" blockId="38.[151,1437,638,1998]" pageId="38" pageNumber="39">
<emphasis id="B966BD220223B72890ABB154EC06FCEF" box="[199,269,819,843]" italics="true" pageId="38" pageNumber="39">Head:</emphasis>
Broad and flat (
<figureCitation id="13297DB50223B72891A1B155EF3CFCEF" box="[461,567,818,843]" captionStart="FIGURE 14" captionStartId="20.[151,250,936,958]" captionTargetBox="[171,1412,204,911]" captionTargetId="figure@20.[160,1412,193,921]" captionTargetPageId="20" captionText="FIGURE 14. Diagrammatic cross sections that represent the width and convexity of head or deflection of pronotum (frontal view): (A) broad and flat; (B) broad and convex; (C) narrow and flat; (D) narrow and convex." httpUri="https://zenodo.org/record/2668091/files/figure.png" pageId="38" pageNumber="39">Fig. 14A</figureCitation>
). Vertex smooth (
<figureCitation id="13297DB50223B7289368B155EE41FCEF" box="[772,842,818,843]" captionStart="FIGURE 9" captionStartId="15.[151,250,1706,1728]" captionTargetBox="[157,1433,759,1683]" captionTargetId="figure@15.[151,1436,757,1683]" captionTargetPageId="15" captionText="FIGURE 9. Dorsal view of head with detail of the morphological structures used in the species descriptions. Ƒ = vertex; SPS = supraorbital puncture setae; SP = supraorbital puncture; F = frons area; FCS = frontoclypeal suture; C = clypeus; M = mandible; LS = labrum setae; L = labrum; ACM = anterior clypeal microsculpture; CSP = clypeal setose puncture; TP = tentorial pit; SG = supraorbital grooves; E = eye; ƑG = vertexal groove." httpUri="https://zenodo.org/record/2668081/files/figure.png" pageId="38" pageNumber="39">Fig. 9</figureCitation>
); vertexal groove narrow and defined entire length; large supraorbital puncture bearing 34 setae; 4 supraorbital grooves slightly impressed, extended onto lateral area of frons; frons smooth; frontoclypeal suture well-defined the entire length, tentorial pits small; anterior area of clypeus narrowly grooved (may be indistinct), 1 large, setose puncture each side bearing 2 setae. Labrum rounded, anterior edge straight with 3 evenly spaced setae each side. Mentum lobes squared (
<figureCitation id="13297DB50223B7289406B1A5E9DEFC7F" box="[1130,1237,962,987]" captionStart="FIGURE 12" captionStartId="18.[151,250,1905,1927]" captionTargetBox="[311,1244,697,1878]" captionTargetId="figure@18.[310,1250,689,1889]" captionTargetPageId="18" captionText="FIGURE 12. Ventral view of the mentum with a number of different character states. Illustrations AD are the different forms of the indentation in the apex of the median process: (A) distinctly indentate (M. manaia); (B) moderately indentate (M. pluto); (C) slightly indentate (M. infimate); (D) indentation notched (M. parataiko). Illustrations EG are the three general forms (i.e., hashed lines) of the mentum lobes: (E) triangular; (F) rounded; (G) squared." httpUri="https://zenodo.org/record/2668087/files/figure.png" pageId="38" pageNumber="39">Fig. 12G</figureCitation>
), median process broad at base narrowed (long) to apex, upward angle absent, distinctly indentate (
<figureCitation id="13297DB50223B728944AB181E985FC5B" box="[1062,1166,998,1023]" captionStart="FIGURE 12" captionStartId="18.[151,250,1905,1927]" captionTargetBox="[311,1244,697,1878]" captionTargetId="figure@18.[310,1250,689,1889]" captionTargetPageId="18" captionText="FIGURE 12. Ventral view of the mentum with a number of different character states. Illustrations AD are the different forms of the indentation in the apex of the median process: (A) distinctly indentate (M. manaia); (B) moderately indentate (M. pluto); (C) slightly indentate (M. infimate); (D) indentation notched (M. parataiko). Illustrations EG are the three general forms (i.e., hashed lines) of the mentum lobes: (E) triangular; (F) rounded; (G) squared." httpUri="https://zenodo.org/record/2668087/files/figure.png" pageId="38" pageNumber="39">Fig. 12A</figureCitation>
); mentum setae present. Submentum sclerite constriction (
<figureCitation id="13297DB50223B7289263B66DEF5AFB87" box="[527,593,1034,1059]" captionStart="FIGURE 8" captionStartId="13.[151,250,1097,1119]" captionTargetBox="[155,1432,309,1072]" captionTargetId="figure@13.[151,1436,303,1075]" captionTargetPageId="13" captionText="FIGURE 8. Detail of the ventral head with the taxonomic structures used in the species descriptions. SBMS = submentum setae; SBM = submentum sclerite; ML = mentum lobe; MP = mentum process (with indentation); MPW = mentum process width; SBS = stipes basal setae; MPL = mentum process length; MPS = mentum process setae; SBMC = submentum sclerite constriction; GP = gula pits; GS = gula suture." httpUri="https://zenodo.org/record/2668079/files/figure.png" pageId="38" pageNumber="39">Fig. 8</figureCitation>
) narrow with 3 setae clustered each side (large gap medially). Stipes with 2 basal setae. Gula (
<figureCitation id="13297DB50223B7289106B649ECA5FBE3" box="[362,430,1070,1095]" captionStart="FIGURE 8" captionStartId="13.[151,250,1097,1119]" captionTargetBox="[155,1432,309,1072]" captionTargetId="figure@13.[151,1436,303,1075]" captionTargetPageId="13" captionText="FIGURE 8. Detail of the ventral head with the taxonomic structures used in the species descriptions. SBMS = submentum setae; SBM = submentum sclerite; ML = mentum lobe; MP = mentum process (with indentation); MPW = mentum process width; SBS = stipes basal setae; MPL = mentum process length; MPS = mentum process setae; SBMC = submentum sclerite constriction; GP = gula pits; GS = gula suture." httpUri="https://zenodo.org/record/2668079/files/figure.png" pageId="38" pageNumber="39">Fig. 8</figureCitation>
) pits small, suture indistinct, gula flat and smooth. Gena with fine wrinkles forming an isodiametric pattern across entire area.
</paragraph>
<paragraph id="8BAD61300223B72890ABB611E9F0FAE7" blockId="38.[151,1437,638,1998]" pageId="38" pageNumber="39">
<emphasis id="B966BD220223B72890ABB611EC49FB2A" box="[199,322,1142,1166]" italics="true" pageId="38" pageNumber="39">Prothorax:</emphasis>
Prothoracic carina narrow the entire length, slightly crenulated with 47 setae each side (
<figureCitation id="13297DB50223B7289557B611E884FB2B" box="[1339,1423,1142,1167]" captionStart="FIGURE 24" captionStartId="39.[151,250,1867,1889]" captionTargetBox="[251,1341,201,1832]" captionTargetId="figure@39.[238,1349,195,1846]" captionTargetPageId="39" captionText="FIGURE 24. Mecodema curvidens plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668111/files/figure.png" pageId="38" pageNumber="39">Fig. 24</figureCitation>
), extended beyond anterior angle; posterior lateral sinuation indistinctly to evidently carinate, parallel; pronotum broad and flattened, overall shape squared (
<figureCitation id="13297DB50223B72892E7B6D9EFEAFB73" box="[651,737,1214,1239]" captionStart="FIGURE 24" captionStartId="39.[151,250,1867,1889]" captionTargetBox="[251,1341,201,1832]" captionTargetId="figure@39.[238,1349,195,1846]" captionTargetPageId="39" captionText="FIGURE 24. Mecodema curvidens plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668111/files/figure.png" pageId="38" pageNumber="39">Fig. 24</figureCitation>
); midline well-defined, medial impressions absent, very fine, transverse lines that may be present laterally on disc; pronotal foveae shallow and narrow; anterior edge inwardly curved and posterior edge relatively straight. Prosternum (
<figureCitation id="13297DB50223B7289324B761EE9BFABB" box="[840,912,1286,1311]" captionStart="FIGURE 2" captionStartId="6.[151,250,800,822]" captionTargetBox="[159,1432,197,773]" captionTargetId="figure@6.[151,1436,193,779]" captionTargetPageId="6" captionText="FIGURE 2. Ventral view of a Mecodema specimen showing specific morphological structures, excluding taxonomic structures indicated in detail figures, used in the species descriptions. ƑLF = ventrite lateral foveae; ƑSP = ventrite setose punctures; MTC = metacoxa; MTƑP = metaventrite process; MSC = mesocoxa; PC = procoxa; PS = prosternum; G = gena; PES = proepisternum; MSE = mesepisternum; MTE = metepisternum; Ƒ1Ƒ6 = ventrites 1-6 (ventrites 1-3 may be fused); M = midline (dashed line, not a taxonomic structure)." httpUri="https://zenodo.org/record/2668067/files/figure.png" pageId="38" pageNumber="39">Fig. 2</figureCitation>
) weakly convex and smooth; proepisternum without microsculpture. Procoxal setae absent; protibia distally expanded and shovel-like (
<figureCitation id="13297DB50223B72894E2B74DE9E5FAE7" box="[1166,1262,1322,1347]" captionStart="FIGURE 10" captionStartId="16.[151,250,1442,1464]" captionTargetBox="[152,1428,191,1421]" captionTargetId="figure@16.[151,1435,183,1427]" captionTargetPageId="16" captionText="FIGURE 10. The protibia form is a character used mainly to determine between genera, except for one species in Mecodema where the difference between (A) greatly distally expanded (M. antarctica) and (B) expanded distally (all other Mecodema species) is used." httpUri="https://zenodo.org/record/2668083/files/figure.png" pageId="38" pageNumber="39">Fig 10B</figureCitation>
).
</paragraph>
<paragraph id="8BAD61300223B72890ABB729E989F9E6" blockId="38.[151,1437,638,1998]" pageId="38" pageNumber="39">
<emphasis id="B966BD220223B72890ABB729EC1DFAC2" box="[199,278,1358,1382]" italics="true" pageId="38" pageNumber="39">Elytra:</emphasis>
Narrow and flat; humeral angle anteriorly convergent (making elytra look narrow at base) (
<figureCitation id="13297DB50223B728955FB729E885FAC3" box="[1331,1422,1358,1383]" captionStart="FIGURE 6" captionStartId="10.[151,250,519,541]" captionTargetBox="[156,1432,193,498]" captionTargetId="figure@10.[151,1436,193,498]" captionTargetPageId="10" captionText="FIGURE 6. Detail of the three different forms of the elytral humeral angle: (A) = anteriorly convergent (M. parataiko), (B) = evenly convex (M. oconnori), (C) = subangulate (M. atuanui)." httpUri="https://zenodo.org/record/2668075/files/figure.png" pageId="38" pageNumber="39">Fig. 6A</figureCitation>
); basal margin slightly curved (almost straight) and bevelled to base, interval 1 extended to scutellum, other intervals and striae terminated at margin; lateral carina narrow the entire length, extended to humeral angle; humerus with 2 setose punctures proximate angle; suture well-defined, slightly broadened at humeral angle; striae with small, regularly distributed, asetose punctures, slightly increased in size laterally; intervals 14 flat, intervals 59 weakly convex (convexity increased laterally); interval microsculpture absent; 7
<superScript id="7C67CC780223B72893B3B465EEE0F9B4" attach="left" box="[991,1003,1538,1552]" fontSize="6" pageId="38" pageNumber="39">th</superScript>
strial setal pattern (
<figureCitation id="13297DB50223B72894B5B462E815F9BA" box="[1241,1310,1541,1566]" captionStart="FIGURE 1" captionStartId="5.[151,250,1912,1934]" captionTargetBox="[159,1427,1029,1884]" captionTargetId="figure@5.[151,1436,1023,1890]" captionTargetPageId="5" captionText="FIGURE 1. Dorsal view of a Mecodema specimen showing taxonomic structures used in the species descriptions. H =head; P = pronotum; E = elytra; S = striae; I1 = interval 1; A = apex; 7S = 7 th strial setae; HSP = humerus setose puncture; HA = humeral angle; BM = basal margin; P = protibia." httpUri="https://zenodo.org/record/2668065/files/figure.png" pageId="38" pageNumber="39">Fig. 1</figureCitation>
) with 01 setose puncture in anterior ½, 34 setose punctures in posterior ½, setose punctures large.
</paragraph>
<paragraph id="8BAD61300223B72890ABB42AE861F952" blockId="38.[151,1437,638,1998]" pageId="38" pageNumber="39">
<emphasis id="B966BD220223B72890ABB42AEC71F9C2" box="[199,378,1613,1638]" italics="true" pageId="38" pageNumber="39">Ventral surface:</emphasis>
Mesepisternum and metepisternum with fine wrinkles; setose punctures present on mesocoxae (1) and metacoxae (0). All abdominal ventrites finely lineate laterally; ventrites 13 setose without punctures, ventrites 4 and 5 with 1 setose puncture each side of midline; ventrite 6 (
<figureCitation id="13297DB50223B72893A7B4F2E904F90A" box="[971,1039,1685,1710]" captionStart="FIGURE 2" captionStartId="6.[151,250,800,822]" captionTargetBox="[159,1432,197,773]" captionTargetId="figure@6.[151,1436,193,779]" captionTargetPageId="6" captionText="FIGURE 2. Ventral view of a Mecodema specimen showing specific morphological structures, excluding taxonomic structures indicated in detail figures, used in the species descriptions. ƑLF = ventrite lateral foveae; ƑSP = ventrite setose punctures; MTC = metacoxa; MTƑP = metaventrite process; MSC = mesocoxa; PC = procoxa; PS = prosternum; G = gena; PES = proepisternum; MSE = mesepisternum; MTE = metepisternum; Ƒ1Ƒ6 = ventrites 1-6 (ventrites 1-3 may be fused); M = midline (dashed line, not a taxonomic structure)." httpUri="https://zenodo.org/record/2668067/files/figure.png" pageId="38" pageNumber="39">Fig. 2</figureCitation>
) setae present:
<specimenCount id="9D14AAB90223B72894D2B4F1E9D8F909" box="[1214,1235,1686,1709]" pageId="38" pageNumber="39" type="male"></specimenCount>
with 2 setae each side at junction of the apical edge flattened curve and curve anterad,
<specimenCount id="9D14AAB90223B72893F8B4DDEEA2F975" box="[916,937,1722,1745]" pageId="38" pageNumber="39" type="female"></specimenCount>
with 4 setae each side, apical edge a pointed curve; lateral foveae absent. Anterior metaventrite process (
<figureCitation id="13297DB50223B7289358B4BAEE7CF952" box="[820,887,1757,1782]" captionStart="FIGURE 2" captionStartId="6.[151,250,800,822]" captionTargetBox="[159,1432,197,773]" captionTargetId="figure@6.[151,1436,193,779]" captionTargetPageId="6" captionText="FIGURE 2. Ventral view of a Mecodema specimen showing specific morphological structures, excluding taxonomic structures indicated in detail figures, used in the species descriptions. ƑLF = ventrite lateral foveae; ƑSP = ventrite setose punctures; MTC = metacoxa; MTƑP = metaventrite process; MSC = mesocoxa; PC = procoxa; PS = prosternum; G = gena; PES = proepisternum; MSE = mesepisternum; MTE = metepisternum; Ƒ1Ƒ6 = ventrites 1-6 (ventrites 1-3 may be fused); M = midline (dashed line, not a taxonomic structure)." httpUri="https://zenodo.org/record/2668067/files/figure.png" pageId="38" pageNumber="39">Fig. 2</figureCitation>
) a long and rounded triangle without carina.
</paragraph>
<paragraph id="8BAD61300223B72690ABB566E879FEB8" blockId="38.[151,1437,638,1998]" lastBlockId="40.[151,1437,151,1149]" lastPageId="40" lastPageNumber="41" pageId="38" pageNumber="39">
<emphasis id="B966BD220223B72890ABB566EC7EF8BD" box="[199,373,1793,1817]" italics="true" pageId="38" pageNumber="39">Male genitalia:</emphasis>
Apical portion of penis lobe symmetrically rounded (
<figureCitation id="13297DB50223B72893B5B566E920F8BE" box="[985,1067,1793,1818]" captionStart="FIGURE 24" captionStartId="39.[151,250,1867,1889]" captionTargetBox="[251,1341,201,1832]" captionTargetId="figure@39.[238,1349,195,1846]" captionTargetPageId="39" captionText="FIGURE 24. Mecodema curvidens plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668111/files/figure.png" pageId="38" pageNumber="39">Fig. 24</figureCitation>
PL) but with a flattened curve at the apex, slight deflection to right of vertical axis (VV) (
<figureCitation id="13297DB50223B728937CB542EE7DF89A" box="[784,886,1829,1854]" captionStart="FIGURE 15" captionStartId="20.[151,250,1943,1965]" captionTargetBox="[162,1424,1039,1914]" captionTargetId="figure@20.[162,1424,1038,1921]" captionTargetPageId="20" captionText="FIGURE 15. The penis lobe in ventral view showing the relative deflection to right of vertical: (A) distinctly deflected (M. regulus); (B) moderately deflected (M. haunoho); (C) slightly deflected (M. oregoides). Scale bars = 1 mm. Structures of penis lobe indicated are: base (B); lateral edge (LE); apical portion (AP); ventral edge (ƑE)." httpUri="https://zenodo.org/record/2668093/files/figure.png" pageId="38" pageNumber="39">Fig. 15C</figureCitation>
); penis lobe shaft straight the entire length (VV), shaft width equally narrow from apex to base, overall length of penis lobe distinctly curved ventrally (LV) (
<figureCitation id="13297DB50223B7289503B52EEDC2F822" captionStart="FIGURE 16" captionStartId="21.[151,250,897,919]" captionTargetBox="[151,1436,181,869]" captionTargetId="figure@21.[151,1436,181,876]" captionTargetPageId="21" captionText="FIGURE 16. Relative strength of the ventral curve over the length of the penis lobe (lateral view): (A) slightly curved (M. oblongum); (B) moderately curved (M. ponaiti); (C) distinctly curved (M. manaia). Scale bar = 1 mm." httpUri="https://zenodo.org/record/2668095/files/figure.png" pageId="38" pageNumber="39">Fig. 16C</figureCitation>
). Structures of the endophallus (
<figureCitation id="13297DB50223B7289258B50AEF71F822" box="[564,634,1901,1926]" captionStart="FIGURE 4" captionStartId="7.[151,250,973,995]" captionTargetBox="[354,1232,193,952]" captionTargetId="figure@7.[354,1232,193,952]" captionTargetPageId="7" captionText="FIGURE 4. An Environmental Scanning Electron Microscope (ESEM) photograph of the four main structures of the apical plate of the endophallus within the penis lobe. This is an example from Mecodema haunoho showing the central spicule (c.s), left (l.s.f) and the right setose flange (r.s.f), and microtrichial field (m.f). These structures are variable among species but within a species, they are very consistent, even over distant geographical ranges. The microtrichial field can be covered in flattened scales as in photo above, or with elongated setae-like scales." httpUri="https://zenodo.org/record/2668071/files/figure.png" pageId="38" pageNumber="39">Fig. 4</figureCitation>
): lateral form of the apex of the central spicule rounded; dorsal form of the apex of the central spicule short and narrow; setal (scales) coverage of the apical plate is sparse to moderate (125%); size of left setose flange small; size of right setose flange large. Left paramere basal lobe small (may be slightly larger in specimens of some populations), rectangular with a long and gentle slope (15°) to arm; arm relatively large (½ of whole paramere), gradually narrowed to terminal lobe (
<figureCitation id="13297DB5022DB7269395B2DBE947FF70" box="[1017,1100,188,213]" captionStart="FIGURE 24" captionStartId="39.[151,250,1867,1889]" captionTargetBox="[251,1341,201,1832]" captionTargetId="figure@39.[238,1349,195,1846]" captionTargetPageId="39" captionText="FIGURE 24. Mecodema curvidens plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668111/files/figure.png" pageId="40" pageNumber="41">Fig. 24</figureCitation>
LP); terminal lobe not distinct from arm, apical tuft of elongate setae, medium lengthed setae continue along apical ¾ of ventral edge. Right paramere narrowly triangular with medium to long setae extending along apical ¾ of ventral edge (
<figureCitation id="13297DB5022DB7269482B363E84BFEB8" box="[1262,1344,260,285]" captionStart="FIGURE 24" captionStartId="39.[151,250,1867,1889]" captionTargetBox="[251,1341,201,1832]" captionTargetId="figure@39.[238,1349,195,1846]" captionTargetPageId="39" captionText="FIGURE 24. Mecodema curvidens plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe." httpUri="https://zenodo.org/record/2668111/files/figure.png" pageId="40" pageNumber="41">Fig. 24</figureCitation>
RP).
</paragraph>
<caption id="DF6D31B80222B72990FBB52CE808F83B" httpUri="https://zenodo.org/record/2668111/files/figure.png" pageId="39" pageNumber="40" startId="39.[151,250,1867,1889]" targetBox="[251,1341,201,1832]" targetPageId="39">
<paragraph id="8BAD61300222B72990FBB52CE808F83B" blockId="39.[151,1436,1867,1951]" pageId="39" pageNumber="40">
<emphasis id="B966BD220222B72990FBB52CEC2AF8C5" bold="true" box="[151,289,1867,1889]" pageId="39" pageNumber="40">FIGURE 24.</emphasis>
<taxonomicName id="4C121AB30222B7299144B52CEF0AF8C5" authorityName="Broun" authorityYear="1915" box="[296,513,1867,1889]" class="Insecta" family="Carabidae" genus="Mecodema" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="curvidens">
<emphasis id="B966BD220222B7299144B52CEF0AF8C5" box="[296,513,1867,1889]" italics="true" pageId="39" pageNumber="40">Mecodema curvidens</emphasis>
</taxonomicName>
plate with half habitus (left) and the three (detached) structures of the male aedeagus: the right (RP) and left parameres (LP), plus the penis lobe (PL). The bottom scale bar is the width across the widest point of the pronotum of the specimen used for this habitus and top scale bar equates to the half the length of the penis lobe.
</paragraph>
</caption>
<paragraph id="8BAD6130022DB72690ABB34FEC10FE23" blockId="40.[151,1437,151,1149]" pageId="40" pageNumber="41">
<emphasis id="B966BD22022DB72690ABB34FEC85FEE4" box="[199,398,296,320]" italics="true" pageId="40" pageNumber="41">Female genitalia:</emphasis>
Basal gonocoxite 1 short and broad, flattened apically, large shallow groove on inward side, internal latero-dorsal carina without setae. Gonocoxite 2 (
<figureCitation id="13297DB5022DB726934EB32BEE6EFEC1" box="[802,869,332,357]" captionStart="FIGURE 5" captionStartId="9.[151,250,1714,1736]" captionTargetBox="[167,1436,790,1674]" captionTargetId="figure@9.[151,1436,790,1693]" captionTargetPageId="9" captionText="FIGURE 5. The entire female reproductive tract (ventral view) including taxonomic structures (for all other abbreviated structure names see Seldon et al. 2012). However, for this revision only the following structures are important: HS = helminthoid sclerite; G1 = gonocoxite 1; G2 = gonocoxite 2; TDC = transverse dorsal carina; R = ramus." httpUri="https://zenodo.org/record/2668073/files/figure.png" pageId="40" pageNumber="41">Fig. 5</figureCitation>
) bluntly rounded and triangular. Ramus very long and narrow.
</paragraph>
</subSubSection>
<subSubSection id="C30832BB022DB72690ABB3F3EED7FDC4" pageId="40" pageNumber="41" type="discussion">
<paragraph id="8BAD6130022DB72690ABB3F3EED7FDC4" blockId="40.[151,1437,151,1149]" pageId="40" pageNumber="41">
<emphasis id="B966BD22022DB72690ABB3F3EC44FE09" bold="true" box="[199,335,404,429]" pageId="40" pageNumber="41">Comments.</emphasis>
<taxonomicName id="4C121AB3022DB7269139B3F3EF4FFE08" authorityName="Broun" authorityYear="1915" box="[341,580,404,428]" class="Insecta" family="Carabidae" genus="Mecodema" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="curvidens">
<emphasis id="B966BD22022DB7269139B3F3EF4FFE08" box="[341,580,404,428]" italics="true" pageId="40" pageNumber="41">Mecodema curvidens</emphasis>
</taxonomicName>
and
<taxonomicName id="4C121AB3022DB7269216B3F2EE1FFE08" authorityName="Broun" authorityYear="1923" box="[634,788,404,428]" class="Insecta" family="Carabidae" genus="Mecodema" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="occiputale">
<emphasis id="B966BD22022DB7269216B3F2EE1FFE08" box="[634,788,404,428]" italics="true" pageId="40" pageNumber="41">M. occiputale</emphasis>
</taxonomicName>
were described by Broun from badly damaged specimens in
<date id="FFAC47F0022DB72690D5B3D0EC60FE74" box="[185,363,439,464]" pageId="40" pageNumber="41" value="1915" valueMax="1923">1915 and 1923</date>
respectively.
<bibRefCitation id="EF831CC1022DB726927CB3D0EE4BFE74" author="Seldon, D. S. &amp; Leschen, R. A. B." box="[528,832,439,464]" pageId="40" pageNumber="41" pagination="1 - 45" refId="ref93265" refString="Seldon, D. S. &amp; Leschen, R. A. B. (2011) Revision of the Mecodema curvidens species group (Coleoptera: Carabidae: Broscini). Zootaxa, 2829, 1 - 45. https: // doi. org / 10.11646 / zootaxa. 2829.1.1" type="journal article" year="2011">Seldon &amp; Leschen (2011)</bibRefCitation>
commented on the similarities in morphology and distribution, with
<taxonomicName id="4C121AB3022DB7269108B3BAECF2FE50" authorityName="Broun" authorityYear="1915" box="[356,505,476,500]" class="Insecta" family="Carabidae" genus="Mecodema" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="curvidens">
<emphasis id="B966BD22022DB7269108B3BAECF2FE50" box="[356,505,476,500]" italics="true" pageId="40" pageNumber="41">M. curvidens</emphasis>
</taxonomicName>
being found more often on the eastern slopes of the Kaimai Ranges and coastal forest remnants, whereas
<taxonomicName id="4C121AB3022DB72691BFB066EF73FDBC" authorityName="Broun" authorityYear="1923" box="[467,632,512,536]" class="Insecta" family="Carabidae" genus="Mecodema" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="occiputale">
<emphasis id="B966BD22022DB72691BFB066EF73FDBC" box="[467,632,512,536]" italics="true" pageId="40" pageNumber="41">M. occiputale</emphasis>
</taxonomicName>
is widespread across the Central Plateau. Further morphological investigations and the discovery of populations in northwestern
<collectingRegion id="49D6AFD2022DB726930FB043EEF3FD99" box="[867,1016,548,573]" country="New Zealand" name="Hawke's Bay" pageId="40" pageNumber="41">Hawkes Bay</collectingRegion>
and East Cape have shown that these two species are actually one widespread species, with distinctive ecotypes.
</paragraph>
</subSubSection>
<subSubSection id="C30832BB022DB72690ABB00BEC2BFD0C" pageId="40" pageNumber="41" type="distribution">
<paragraph id="8BAD6130022DB72690ABB00BEC2BFD0C" blockId="40.[151,1437,151,1149]" pageId="40" pageNumber="41">
<emphasis id="B966BD22022DB72690ABB00BEC69FD21" bold="true" box="[199,354,620,645]" pageId="40" pageNumber="41">Distribution:</emphasis>
<collectingCountry id="F30521A0022DB726911DB00AEF19FD21" box="[369,530,620,645]" name="New Zealand" pageId="40" pageNumber="41">New Zealand</collectingCountry>
, North Island, Coromandel, Bay of Plenty, Hawkes Bay, Taupo, Waikato, Whanganui.
</paragraph>
</subSubSection>
<subSubSection id="C30832BB022DB72690ABB0D3E9D6FC24" pageId="40" pageNumber="41" type="materials_examined">
<paragraph id="8BAD6130022DB72690ABB0D3EE2FFCB1" blockId="40.[151,1437,151,1149]" pageId="40" pageNumber="41">
<emphasis id="B966BD22022DB72690ABB0D3EC3FFD69" bold="true" box="[199,308,692,717]" pageId="40" pageNumber="41">
<typeStatus id="54A9DF92022DB72690ABB0D3EC3FFD69" box="[199,308,692,717]" pageId="40" pageNumber="41" type="holotype">Holotype</typeStatus>
</emphasis>
(
<taxonomicName id="4C121AB3022DB7269129B0D3EF46FD68" authorityName="Broun" authorityYear="1915" box="[325,589,692,716]" class="Insecta" family="Carabidae" genus="Metaglymma" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="curvidens">
<emphasis id="B966BD22022DB7269129B0D3EF46FD68" box="[325,589,692,716]" italics="true" pageId="40" pageNumber="41">Metaglymma curvidens</emphasis>
</taxonomicName>
): BMNH. 3683. [hw] / New Zeal. [red underlined] Broun Coll. Brit. Mus. 1923-482. / Opotiki. Decr. 1912. [hw] /
<taxonomicName id="4C121AB3022DB7269209B0B0EE7FFD54" authorityName="Broun" authorityYear="1915" box="[613,884,727,752]" class="Insecta" family="Carabidae" genus="Metaglymma" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="curvidens">Metaglymma curvidens</taxonomicName>
.
<specimenCount id="9D14AAB9022DB72693ECB0BFEE9DFD4B" box="[896,918,728,751]" pageId="40" pageNumber="41" type="female"></specimenCount>
. [hw] /
<typeStatus id="54A9DF92022DB7269398B0BFE983FD54" box="[1012,1160,728,752]" pageId="40" pageNumber="41" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4C121AB3022DB72694FDB0B0EDD6FCB0" authority="Broun" authorityName="Broun" authorityYear="1915" class="Insecta" family="Carabidae" genus="Metaglymma" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="curvidens">Metaglymma curvidens Broun</taxonomicName>
confirmed by Seldon and Leschen 2010 [red label].
</paragraph>
<paragraph id="8BAD6130022DB72690ABB178E9D6FC24" blockId="40.[151,1437,151,1149]" pageId="40" pageNumber="41">
<emphasis id="B966BD22022DB72690ABB178EC36FC9C" bold="true" box="[199,317,799,824]" pageId="40" pageNumber="41">
<typeStatus id="54A9DF92022DB72690ABB178EC36FC9C" box="[199,317,799,824]" pageId="40" pageNumber="41" type="lectotype">Lectotype</typeStatus>
</emphasis>
(
<taxonomicName id="4C121AB3022DB726913CB147EF42FC9C" authorityName="Broun" authorityYear="1923" box="[336,585,800,824]" class="Insecta" family="Carabidae" genus="Mecodema" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="occiputale">
<emphasis id="B966BD22022DB726913CB147EF42FC9C" box="[336,585,800,824]" italics="true" pageId="40" pageNumber="41">Mecodema occiputale</emphasis>
</taxonomicName>
): BMNH. Para-type [round label with yellow border] / 4256 [hw] / New Zeal. [red line beneath
<collectingCountry id="F30521A0022DB72691F7B122ECC8FCF8" box="[411,451,837,860]" name="New Zealand" pageId="40" pageNumber="41">NZ</collectingCountry>
] Broun Coll. Brit. Mus. 1922-482. / Tokaanau.
<date id="FFAC47F0022DB72693B7B123E94BFCF9" box="[987,1088,836,861]" pageId="40" pageNumber="41" value="1918-04-06">6.4.1918</date>
. [hw] /
<taxonomicName id="4C121AB3022DB72694FBB123E81CFCF9" authorityName="Blanchard" authorityYear="1853" box="[1175,1303,836,861]" class="Insecta" family="Carabidae" genus="Mecodema" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">Mecodema</taxonomicName>
, occiputale. [hw] /
<typeStatus id="54A9DF92022DB726908DB10FEC8AFC24" box="[225,385,872,896]" pageId="40" pageNumber="41" type="lectotype">LECTOTYPE</typeStatus>
<taxonomicName id="4C121AB3022DB72691E4B10FEFC0FCDB" authority="Broun" authorityName="Broun" authorityYear="1923" box="[392,715,872,896]" class="Insecta" family="Carabidae" genus="Mecodema" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="occiputale">
<emphasis id="B966BD22022DB72691E4B10FEF74FC24" box="[392,639,872,896]" italics="true" pageId="40" pageNumber="41">Mecodema occiputale</emphasis>
Broun
</taxonomicName>
design. by Seldon &amp; Leschen 2010 [red label].
</paragraph>
</subSubSection>
<subSubSection id="C30832BB022DB72690ABB1EBE912FC6C" pageId="40" pageNumber="41" type="description">
<paragraph id="8BAD6130022DB72690ABB1EBE912FC6C" blockId="40.[151,1437,151,1149]" pageId="40" pageNumber="41">
<materialsCitation id="3B7A6B6D022DB72690ABB1EBEEB9FC01" ID-GBIF-Occurrence="2237851751" box="[199,946,908,933]" collectingDate="21-01-16" collectionCode="NZAC" pageId="40" pageNumber="41" specimenCount="1" typeStatus="paratype">
<emphasis id="B966BD22022DB72690ABB1EBEC3FFC01" bold="true" box="[199,308,908,933]" pageId="40" pageNumber="41">
<typeStatus id="54A9DF92022DB72690ABB1EBEC3FFC01" box="[199,308,908,933]" pageId="40" pageNumber="41" type="paratype">Paratype</typeStatus>
</emphasis>
(
<taxonomicName id="4C121AB3022DB7269120B1EBEF33FC00" authorityName="Brookes" authorityYear="1926" box="[332,568,908,932]" class="Insecta" family="Carabidae" genus="Mecodema" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="exitiosus">
<emphasis id="B966BD22022DB7269120B1EBEF33FC00" box="[332,568,908,932]" italics="true" pageId="40" pageNumber="41">Mecodema exitiosus</emphasis>
</taxonomicName>
):
<collectionCode id="ED03F9F5022DB7269234B1EBEFA3FC00" box="[600,680,908,932]" country="New Zealand" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34070" lsid="urn:lsid:biocol.org:col:34070" name="New Zealand Arthropod Collection" pageId="40" pageNumber="41" type="Other institutional type">NZAC</collectionCode>
. Okauia
<date id="FFAC47F0022DB7269376B1EBEE65FC01" box="[794,878,908,933]" pageId="40" pageNumber="41" value="21-01-16">
<collectingDate id="EFE8BE18022DB7269376B1EBEE65FC01" box="[794,878,908,933]" pageId="40" pageNumber="41" value="21-01-16">16.1.21</collectingDate>
</date>
[hw]
</materialsCitation>
/
<materialsCitation id="3B7A6B6D022DB72693B4B1EBE91DFC6C" ID-GBIF-Occurrence="2237851757" collectionCode="NZAC" country="New Zealand" location="Arthropod Collection" pageId="40" pageNumber="41" specimenCode="NZAC04011128" specimenCount="1" typeStatus="holotype">
<typeStatus id="54A9DF92022DB72693B4B1EBE94BFC01" box="[984,1088,908,933]" pageId="40" pageNumber="41" type="holotype">Holotype</typeStatus>
(faded orange label) [hw] /
<taxonomicName id="4C121AB3022DB72690FBB1C8ECE9FC6C" authority="Brookes" authorityName="Brookes" authorityYear="1926" box="[151,482,943,968]" class="Insecta" family="Carabidae" genus="Mecodema" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="exitiosus">Mecodema exitiosus Brookes</taxonomicName>
[hw] /
<collectingCountry id="F30521A0022DB7269258B1D7EF52FC63" box="[564,601,944,967]" name="New Zealand" pageId="40" pageNumber="41">NZ</collectingCountry>
<location id="8ECD37EB022DB726920CB1C8EE5AFC6C" LSID="urn:lsid:plazi:treatment:03BBD0260223B72690FBB3F3E873FBD8:8ECD37EB022DB726920CB1C8EE5AFC6C" box="[608,849,943,968]" country="New Zealand" name="Arthropod Collection" pageId="40" pageNumber="41">Arthropod Collection</location>
<specimenCode id="DBB4C94B022DB7269334B1D7E91DFC6C" box="[856,1046,944,968]" collectionCode="NZAC" country="New Zealand" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34070" lsid="urn:lsid:biocol.org:col:34070" name="New Zealand Arthropod Collection" pageId="40" pageNumber="41" type="Other institutional type">
NZAC
<date id="FFAC47F0022DB72693C9B1D7E91DFC6C" box="[933,1046,944,968]" pageId="40" pageNumber="41" value="1128-04-01">04011128</date>
</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30832BB022DB72690ABB1B3E873FBD8" pageId="40" pageNumber="41" type="materials_examined">
<paragraph id="8BAD6130022DB72690ABB1B3E873FBD8" blockId="40.[151,1437,151,1149]" pageId="40" pageNumber="41">
<emphasis id="B966BD22022DB72690ABB1B3ECBEFC49" bold="true" box="[199,437,980,1005]" pageId="40" pageNumber="41">Material examined:</emphasis>
1,
<materialsCitation id="3B7A6B6D022DB72691B3B1B2E97DFBB4" ID-GBIF-Occurrence="2237851817" collectingDate="2011-01-04" collectionCode="NZAC" collectorName="D. S. Seldon &amp; Podocarp-kamahi" country="New Zealand" location="Boundary Stream Mainland Island" municipality="Kamahi Loop Tk" pageId="40" pageNumber="41" specimenCount="1">
<collectingCountry id="F30521A0022DB72691B3B1B2EFA6FC48" box="[479,685,980,1004]" name="New Zealand" pageId="40" pageNumber="41">NEW ZEALAND</collectingCountry>
HB
<location id="8ECD37EB022DB7269284B1B3E97DFC49" LSID="urn:lsid:plazi:treatment:03BBD0260223B72690FBB3F3E873FBD8:8ECD37EB022DB7269284B1B3E97DFC49" box="[744,1142,980,1005]" country="New Zealand" municipality="Kamahi Loop Tk" name="Boundary Stream Mainland Island" pageId="40" pageNumber="41">Boundary Stream Mainland Island</location>
,
<collectingMunicipality id="6BC9FB4A022DB72694E8B1B3E845FC49" box="[1156,1358,980,1005]" pageId="40" pageNumber="41">Kamahi Loop Tk</collectingMunicipality>
, under logs,
<date id="FFAC47F0022DB72690BFB19FEC46FBB4" box="[211,333,1016,1040]" pageId="40" pageNumber="41" value="2011-01-04">
<collectingDate id="EFE8BE18022DB72690BFB19FEC46FBB4" box="[211,333,1016,1040]" pageId="40" pageNumber="41" value="2011-01-04">4 Jan 2011</collectingDate>
</date>
,
<collectorName id="26E704E6022DB7269134B19FECD4FBB4" box="[344,479,1015,1040]" pageId="40" pageNumber="41">D.S. Seldon</collectorName>
/
<collectorName id="26E704E6022DB7269198B190EFB1FBB4" box="[500,698,1015,1040]" pageId="40" pageNumber="41">Podocarp-kamahi</collectorName>
canopy, E2839579 N6225723 (
<collectionCode id="ED03F9F5022DB7269477B19FE966FBB4" box="[1051,1133,1016,1040]" country="New Zealand" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34070" lsid="urn:lsid:biocol.org:col:34070" name="New Zealand Arthropod Collection" pageId="40" pageNumber="41" type="Other institutional type">NZAC</collectionCode>
)
</materialsCitation>
; 1,
<materialsCitation id="3B7A6B6D022DB72694F0B19FEF93FBFC" ID-GBIF-Occurrence="2237851784" collectingDate="2015-12-20" collectingDateMax="2015-12-28" collectingDateMin="2015-12-20" collectionCode="NZAC" collectorName="D. S. Seldon &amp; Podocarp-broadleaf" country="New Zealand" elevation="552" latitude="-38.982002" location="State Highway" longLatPrecision="1" longitude="176.53345" municipality="Oputo Falls" pageId="40" pageNumber="41" specimenCount="1">
<collectingCountry id="F30521A0022DB72694F0B19FE86CFBB4" box="[1180,1383,1016,1040]" name="New Zealand" pageId="40" pageNumber="41">NEW ZEALAND</collectingCountry>
HB/ TO
<location id="8ECD37EB022DB72690A8B67BEC60FB91" LSID="urn:lsid:plazi:treatment:03BBD0260223B72690FBB3F3E873FBD8:8ECD37EB022DB72690A8B67BEC60FB91" box="[196,363,1052,1077]" country="New Zealand" latitude="-38.982002" longLatPrecision="1" longitude="176.53345" municipality="Oputo Falls" name="State Highway" pageId="40" pageNumber="41">State Highway</location>
5,
<collectingMunicipality id="6BC9FB4A022DB72691FEB67BEF13FB91" box="[402,536,1052,1077]" pageId="40" pageNumber="41">Oputo Falls</collectingMunicipality>
S.R.,
<date id="FFAC47F0022DB726920CB67BEE2BFB91" box="[608,800,1052,1077]" pageId="40" pageNumber="41" value="2015-12-20" valueMax="2015-12-28" valueMin="2015-12-20">
<collectingDate id="EFE8BE18022DB726920CB67BEE2BFB91" box="[608,800,1052,1077]" pageId="40" pageNumber="41" value="2015-12-20" valueMax="2015-12-28" valueMin="2015-12-20">2028 Dec 2015</collectingDate>
</date>
,
<collectorName id="26E704E6022DB7269341B67BEEBEFB91" box="[813,949,1052,1077]" pageId="40" pageNumber="41">D.S. Seldon</collectorName>
/
<collectorName id="26E704E6022DB72693A3B67BE9A7FB91" box="[975,1196,1052,1077]" pageId="40" pageNumber="41">Podocarp-broadleaf</collectorName>
canopy,
<collectingMethod id="52531927022DB726957FB67BE893FB90" box="[1299,1432,1052,1077]" pageId="40" pageNumber="41">pitfall traps</collectingMethod>
,
<geoCoordinate id="EE2607F7022DB72690FBB658EC28FBFC" box="[151,291,1087,1112]" direction="south" orientation="latitude" pageId="40" pageNumber="41" precision="1" value="-38.982002">S38°5855.2</geoCoordinate>
<geoCoordinate id="EE2607F7022DB7269143B658ECC5FBFC" box="[303,462,1087,1112]" direction="east" orientation="longitude" pageId="40" pageNumber="41" precision="1" value="176.53345">E176°3200.4</geoCoordinate>
,
<quantity id="4CEACCD5022DB72691B2B658EF21FBF3" box="[478,554,1087,1112]" metricMagnitude="2" metricUnit="m" metricValue="5.52" pageId="40" pageNumber="41" unit="m" value="552.0">
<elevation id="003F8603022DB72691B2B658EF21FBF3" box="[478,554,1087,1112]" metricMagnitude="2" metricUnit="m" metricValue="5.52" pageId="40" pageNumber="41" unit="m" value="552.0">552 m</elevation>
</quantity>
(
<collectionCode id="ED03F9F5022DB7269251B627EF9BFBFC" box="[573,656,1088,1112]" country="New Zealand" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34070" lsid="urn:lsid:biocol.org:col:34070" name="New Zealand Arthropod Collection" pageId="40" pageNumber="41" type="Other institutional type">NZAC</collectionCode>
)
</materialsCitation>
; 1,
<materialsCitation id="3B7A6B6D022DB72692A4B627E87EFBD8" ID-GBIF-Occurrence="2237851610" collectingDate="2015-12-19" collectingDateMax="2015-12-30" collectingDateMin="2015-12-19" collectionCode="NZAC" collectorName="D. S. Seldon &amp; J. Tizard &amp; Podocarp-broadleaf" country="New Zealand" latitude="-39.271027" location="Puketitiri" longLatPrecision="1" longitude="176.49873" municipality="Ball's Clearing" pageId="40" pageNumber="41" specimenCount="1">
<collectingCountry id="F30521A0022DB72692A4B627EE91FBFC" box="[712,922,1088,1112]" name="New Zealand" pageId="40" pageNumber="41">NEW ZEALAND</collectingCountry>
, HB,
<location id="8ECD37EB022DB726938FB658E945FBFC" LSID="urn:lsid:plazi:treatment:03BBD0260223B72690FBB3F3E873FBD8:8ECD37EB022DB726938FB658E945FBFC" box="[995,1102,1087,1112]" country="New Zealand" latitude="-39.271027" longLatPrecision="1" longitude="176.49873" municipality="Ball's Clearing" name="Puketitiri" pageId="40" pageNumber="41">Puketitiri</location>
,
<collectingMunicipality id="6BC9FB4A022DB7269432B658E805FBFC" box="[1118,1294,1087,1112]" pageId="40" pageNumber="41">Balls Clearing</collectingMunicipality>
,
<date id="FFAC47F0022DB7269572B627EDD9FBD9" pageId="40" pageNumber="41" value="2015-12-19" valueMax="2015-12-30" valueMin="2015-12-19">
<collectingDate id="EFE8BE18022DB7269572B627EDD9FBD9" pageId="40" pageNumber="41" value="2015-12-19" valueMax="2015-12-30" valueMin="2015-12-19">1930 Dec 2015</collectingDate>
</date>
,
<collectorName id="26E704E6022DB72690B1B603EC6DFBD9" box="[221,358,1124,1149]" pageId="40" pageNumber="41">D.S. Seldon</collectorName>
,
<collectorName id="26E704E6022DB726911DB602ECD9FBD9" box="[369,466,1124,1149]" pageId="40" pageNumber="41">J. Tizard</collectorName>
/
<collectorName id="26E704E6022DB7269184B603EFCEFBD9" box="[488,709,1124,1149]" pageId="40" pageNumber="41">Podocarp-broadleaf</collectorName>
canopy, 30
<collectingMethod id="52531927022DB7269320B603EEC4FBD8" box="[844,975,1124,1149]" pageId="40" pageNumber="41">pitfall traps</collectingMethod>
,
<geoCoordinate id="EE2607F7022DB72693B6B603E963FBD9" box="[986,1128,1124,1149]" direction="south" orientation="latitude" pageId="40" pageNumber="41" precision="1" value="-39.271027">S39°1615.7</geoCoordinate>
<geoCoordinate id="EE2607F7022DB7269403B603E800FBD9" box="[1135,1291,1124,1149]" direction="east" orientation="longitude" pageId="40" pageNumber="41" precision="1" value="176.49873">E176°2955.4</geoCoordinate>
(
<collectionCode id="ED03F9F5022DB7269576B603E866FBD8" box="[1306,1389,1124,1148]" country="New Zealand" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34070" lsid="urn:lsid:biocol.org:col:34070" name="New Zealand Arthropod Collection" pageId="40" pageNumber="41" type="Other institutional type">NZAC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>