149 lines
19 KiB
XML
149 lines
19 KiB
XML
<document id="7D06C088957C51160E1E9FEB606AC2D3" ID-DOI="10.2478/if-2019-0022" ID-ISSN="2533-4069" ID-Zenodo-Dep="5381828" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630294418819" checkinUser="diego" docAuthor="Syromyatnikova, Elena, Tesakov, Alexey, Mayda, Serdar, Kaya, Tanju & Saraç, Gerçek" docDate="2019" docId="03B387BBDF60FFA1FC7CF83B0A34FE72" docLanguage="en" docName="05_Syromyatnikova.pdf" docOrigin="Fossil Imprint 75 (3 - 4)" docSource="http://dx.doi.org/10.2478/if-2019-0022" docStyle="DocumentStyle:0EC54BF35E227CA4627F894F6BA30634.4:FossilImprint.2016-.journal_article" docStyleId="0EC54BF35E227CA4627F894F6BA30634" docStyleName="FossilImprint.2016-.journal_article" docStyleVersion="4" docTitle="Amphisbaenia GRAY 1844" docType="treatment" docVersion="4" lastPageNumber="352" masterDocId="FF8AFFC3DF68FFABFF82FF900860FFA2" masterDocTitle="Plio-Pleistocene Amphibians And Reptiles From Central Turkey: New Faunas And Faunal Records With Comments On Their Biochronological Position Based On Small Mammals" masterLastPageNumber="358" masterPageNumber="343" pageNumber="351" updateTime="1699217866505" updateUser="plazi" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title id="5E8A6F289E460D92165B534D0A5A4538">Plio-Pleistocene Amphibians And Reptiles From Central Turkey: New Faunas And Faunal Records With Comments On Their Biochronological Position Based On Small Mammals</mods:title>
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<mods:namePart id="FC3A30B75F7DC71CEA2582382C2F4073">Syromyatnikova, Elena</mods:namePart>
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<mods:affiliation id="B2805CF07A42B08C7BDD80C959D15CF5">A. A. Borissiak Paleontological Institute, Russian Academy of Sciences, Profsoyuznaya ul. 123, 117647 Moscow, Russia; e-mails: esyromyatnikova @ gmail. com, sev @ paleo. ru. & Zoological Institute, Russian Academy of Sciences, Universitetskaya emb. 1, 199034 St. Petersburg, Russia.</mods:affiliation>
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<mods:namePart id="6A2DBE2FCA5868719BD06EB00F82007D">Tesakov, Alexey</mods:namePart>
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<mods:affiliation id="4529B48116A22CF17F80E898562A196F">Geological Institute, Russian Academy of Sciences, Pyzhevsky per. 7, 119017 Moscow, Russia; e-mail: tesak @ ginras. ru.</mods:affiliation>
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<mods:nameIdentifier id="E2B062CB29A2DEA10F003E1D20A8D221" type="email">tesak@ginras.ru.</mods:nameIdentifier>
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<mods:namePart id="912FB47604E1AA2D7856B8754A70F970">Mayda, Serdar</mods:namePart>
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<mods:affiliation id="CD985D460B169E5C320967600B3848C7">Department of Biology, Faculty of Science, Ege University, Bornova, 35100 Izmir, Turkey; e-mail: serdar. mayda @ ege. edu. tr. & Ege University, Natural History Museum, 35100 Bornova-Izmir, Turkey.</mods:affiliation>
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<mods:nameIdentifier id="80EE3BD0737BC8F0075480018E34483D" type="email">serdar.mayda@ege.edu.tr.</mods:nameIdentifier>
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<mods:namePart id="E393EB9F3BC455A3514457697FE83C41">Kaya, Tanju</mods:namePart>
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<mods:affiliation id="B83F0779349F007D0CEA6A029ABDED58">Ege University, Natural History Museum, 35100 Bornova-Izmir, Turkey.</mods:affiliation>
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<mods:namePart id="C253D5EBC9F9041DA2E9BEDB20E2AA76">Saraç, Gerçek</mods:namePart>
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<mods:affiliation id="2F5845567CE8F7215469A71B152B2298">Sunar Sokak No: 3 / 3 Yenimahalle-Ankara, Turkey.</mods:affiliation>
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<mods:title id="E389E5A1D79E27F681E5501A72BA6222">Fossil Imprint</mods:title>
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<mods:date id="46F3EC02B58B7BBDE9B643376957E190">2019</mods:date>
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<mods:number id="B29C6B73F5EEB2CA64023D26BD8CA80A">2019-12-30</mods:number>
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<mods:number id="650CD1E7CBDE4BE6277B615A490C272B">75</mods:number>
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<mods:number id="70E9A50A95CC40AC6A29E3C561A3A595">3 - 4</mods:number>
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<mods:url id="E87E2D546E388A0C7039752E587C85B8">http://dx.doi.org/10.2478/if-2019-0022</mods:url>
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<mods:identifier id="58F9F712238E1B1C29023973B552885E" type="DOI">10.2478/if-2019-0022</mods:identifier>
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<mods:identifier id="FB4A7DDA99A90FE7075032C5BA755668" type="ISSN">2533-4069</mods:identifier>
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<subSubSection id="C3006526DF60FFA3FC7CF83B0CD1F846" pageId="8" pageNumber="351" type="nomenclature">
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<paragraph id="8BA536ADDF60FFA3FC7CF83B0C8AF861" blockId="8.[1022,1258,1963,1987]" box="[1022,1258,1963,1987]" pageId="8" pageNumber="351">
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<heading id="D0ED81C1DF60FFA3FC7CF83B0C8AF861" box="[1022,1258,1963,1987]" centered="true" fontSize="8" level="2" pageId="8" pageNumber="351" reason="2">
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<taxonomicName id="4C1A4D2EDF60FFA3FC7CF83B0CC2F861" authorityName="GRAY" authorityYear="1844" box="[1022,1186,1963,1987]" class="Reptilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphisbaenia" pageId="8" pageNumber="351" phylum="Chordata" rank="order">Amphisbaenia</taxonomicName>
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indet.
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</emphasis>
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</paragraph>
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<paragraph id="8BA536ADDF60FFA3FBB4F85C0CD1F846" blockId="8.[1078,1201,1996,2020]" box="[1078,1201,1996,2020]" pageId="8" pageNumber="351">
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<figureCitation id="13212A28DF60FFA3FBB4F85C0CD1F846" box="[1078,1201,1996,2020]" captionStart="Text-fig" captionStartId="9.[147,228,1484,1505]" captionTargetBox="[159,1458,164,1402]" captionTargetId="figure-210@9.[147,1462,151,1431]" captionTargetPageId="9" captionText="Text-fig. 6. Squamates: a – vertebra of Amphisbaenia indet. from Nasrettinhoca 1, in dorsal (a1), ventral (a2), lateral (a3), anterior (a4), and posterior (a5) views, EUNHM PV-13238; b – right maxilla of Lacertidae indet. from Nasrettinhoca 1, in lateral (b1) and medial (b2) views, EUNHM PV-13239; c – cervical vertebra of Natrix sp. from Hamamkarahisar B, in dorsal (c1), ventral (c2), lateral (c3), anterior (c4), andposterior (c5) views, EUNHM PV-13240; d – trunk vertebra of Natricinae indet. from Nasrettinhoca 2, in dorsal (d1), ventral (d2), lateral (d3), anterior (d4), and posterior (d5) views, EUNHM PV-13241; e – vertebra of cf. Colubrinae indet. from Yenişarbademli, in ventral (e1), dorsal (e2), and anterior (e3) views, EUNHM PV-13262. Scale equals 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5381840" httpUri="https://zenodo.org/record/5381840/files/figure.png" pageId="8" pageNumber="351">Text-fig. 6a</figureCitation>
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<paragraph id="8BA536ADDF60FFA3FCD8F86C0DD3F7B1" blockId="8.[820,1459,2043,2100]" box="[858,1459,2043,2067]" pageId="8" pageNumber="351">M a t e r i a l. One vertebra, EUNHM PV-13238,</paragraph>
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<paragraph id="8BA536ADDF60FFA3FCB6F78C0C82F796" blockId="8.[820,1459,2043,2100]" box="[820,1250,2076,2100]" pageId="8" pageNumber="351">Nasrettinhoca 1, early Pliocene, MN 14.</paragraph>
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<paragraph id="8BA536ADDF61FFA2FF7EFA190D2BFA0E" blockId="9.[252,561,1416,1452]" box="[252,1355,1416,1452]" lastBlockId="9.[792,1355,1416,1452]" pageId="9" pageNumber="352">d1 d2 d3 d4 d5</paragraph>
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<caption id="DF656625DF61FFA2FF11FA5C0D35F9D1" ID-DOI="http://doi.org/10.5281/zenodo.5381840" ID-Zenodo-Dep="5381840" httpUri="https://zenodo.org/record/5381840/files/figure.png" pageId="9" pageNumber="352" startId="9.[147,228,1484,1505]" targetBox="[159,1458,164,1402]" targetPageId="9">
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<paragraph id="8BA536ADDF61FFA2FF11FA5C0D35F9D1" blockId="9.[147,1463,1484,1651]" pageId="9" pageNumber="352">
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Text-fig. 6. Squamates: a – vertebra of
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||
<taxonomicName id="4C1A4D2EDF61FFA2FD98FA5C0ACEFA43" authorityName="GRAY" authorityYear="1844" box="[538,686,1484,1505]" class="Reptilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphisbaenia" pageId="9" pageNumber="352" phylum="Chordata" rank="order">Amphisbaenia</taxonomicName>
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indet. from Nasrettinhoca 1, in dorsal (a1), ventral (a2), lateral (a3), anterior (a4), and posterior (a5) views, EUNHM PV-13238; b – right maxilla of
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<taxonomicName id="4C1A4D2EDF61FFA2FCE6FA790BB3FA5C" authorityName="BONAPARTE" authorityYear="1831" box="[868,979,1513,1534]" class="Reptilia" family="Lacertidae" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="352" phylum="Chordata" rank="family">Lacertidae</taxonomicName>
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indet. from Nasrettinhoca 1, in lateral (b1) and medial (b2) views, EUNHM PV-13239; c – cervical vertebra of
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<taxonomicName id="4C1A4D2EDF61FFA2FCA1F9960BE7F9B9" box="[803,903,1542,1563]" class="Reptilia" family="Natricidae" genus="Natrix" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="352" phylum="Chordata" rank="species" species="undetermined">
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<emphasis id="B96EEABFDF61FFA2FCA1F9960B03F9B9" bold="true" box="[803,867,1542,1563]" italics="true" pageId="9" pageNumber="352">Natrix</emphasis>
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sp.
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</taxonomicName>
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from Hamamkarahisar B, in dorsal (c1), ventral (c2), lateral (c3), anterior (c4), and posterior (c5) views, EUNHM PV-13240; d – trunk vertebra of
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<taxonomicName id="4C1A4D2EDF61FFA2FBAFF9B40CF9F99B" baseAuthorityName="sensu Szyndlar" baseAuthorityYear="1991" box="[1069,1177,1572,1593]" class="Reptilia" family="Natricinae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="352" phylum="Chordata" rank="family">Natricinae</taxonomicName>
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indet. from Nasrettinhoca 2, in dorsal (d1), ventral (d2), lateral (d3), anterior (d4), and posterior (d5) views, EUNHM PV-13241; e – vertebra of cf. Colubrinae indet. from Yenişarbademli, in ventral (e1), dorsal (e2), and anterior (e3) views, EUNHM PV-13262. Scale equals 1 mm.
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</paragraph>
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</caption>
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<paragraph id="8BA536ADDF61FFA2FF24F92E0C1DF8B7" blockId="9.[129,768,1725,2085]" lastBlockId="9.[843,1482,1725,1813]" pageId="9" pageNumber="352">
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D e s c r i p t i o n. The presacral vertebra is small (centrum length about
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) with a centrum that is almost compressed dorsoventrally. The lateral margins of the centrum are almost parallel and divergent anteriorly. The ventral surface of the centrum bears two distinct subcentral foramina. The cotyles and condyles are oval in shape. The neural canal is semicircle in anterior and posterior views and its height is slightly lower than the height of the cotyle. The neural arch is almost smooth dorsally without a neural crest. The arch has a sinusoid shape in lateral aspect, gradually rising towards the posterior end. The anterior edge of the neural arch is deeply convex. The pre- and postzygapophyses are missing (except for a small fragment of the left prezygapophysis).
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</paragraph>
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<paragraph id="8BA536ADDF61FFA1FCF3F8BD0A34FE72" blockId="9.[843,1482,1837,2085]" lastBlockId="10.[106,745,151,464]" lastPageId="10" lastPageNumber="353" pageId="9" pageNumber="352">
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R e m a r k s. The vertebra is too fragmentary to provide any details of their morphology (i.e. morphology of prezygapophyses and synapophyses). The visible features of the specimen generally correspond to those of
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<taxonomicName id="4C1A4D2EDF61FFA2FAE0F81D0DAAF807" authorityName="Gray" authorityYear="1825" box="[1378,1482,1933,1957]" class="Reptilia" family="Anguidae" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="352" phylum="Chordata" rank="family">Anguidae</taxonomicName>
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(and
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<taxonomicName id="4C1A4D2EDF61FFA2FC02F83D0BAAF866" box="[896,970,1965,1988]" class="Reptilia" family="Anguidae" genus="Anguis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="352" phylum="Chordata" rank="genus">
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<emphasis id="B96EEABFDF61FFA2FC02F83D0BAAF866" box="[896,970,1965,1988]" italics="true" pageId="9" pageNumber="352">Anguis</emphasis>
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</taxonomicName>
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) based on a dorsoventrally compressed centrum and lateral margin of the centrum, which are almost parallel in ventral view (
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<bibRefCitation id="EF8B4B5CDF61FFA2FB84F87D0CD4F7A7" author="Klembara, J." box="[1030,1204,2029,2053]" pageId="9" pageNumber="352" pagination="121 - 168" refId="ref12448" refString="Klembara, J. (1982): Beitrag zur Kenntnis der Subfamilie Anguinae. - Acta Universitatis Carolinae, Geologica, 1981 (2): 121 - 168." type="journal article" year="1982">Klembara 1982</bibRefCitation>
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,
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<bibRefCitation id="EF8B4B5CDF61FFA2FB46F87D0DDFF7A7" author="Cernansky, A. & Yaryhin, O. & Cicekova, J. & Werneburg, I. & Hain, M. & Klembara, J." box="[1220,1471,2029,2055]" pageId="9" pageNumber="352" pagination="232 - 257" refId="ref11417" refString="Cernansky, A., Yaryhin, O., Cicekova, J., Werneburg, I., Hain, M., Klembara, J. (2019): Vertebral comparative anatomy and morphological differences in anguine lizards with a special reference to Pseudopus apodus. - The Anatomical Record, 302 (2): 232 - 257. https: // doi. org / 10.1002 / ar. 23944" type="journal article" year="2019">Čerňanský et al. 2019</bibRefCitation>
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||
). However,
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<taxonomicName id="4C1A4D2EDF61FFA2FC43F79D0C49F787" authorityName="Gray" authorityYear="1825" box="[961,1065,2061,2085]" class="Reptilia" family="Anguidae" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="352" phylum="Chordata" rank="family">Anguidae</taxonomicName>
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are characterized by the vertebrae having a clear neural spine and generally a larger size. The specimen from Nasrettinhoca 1 lacks a neural spine, its arch has a sinusoidal shape in lateral view, with a convex anterior edge in dorsal view, which support referral to an amphisbaenian (
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<bibRefCitation id="EF8B4B5CDF62FFA1FEA7FE8809CDFE92" author="Estes, R." box="[293,429,280,304]" pageId="10" pageNumber="353" refId="ref11659" refString="Estes, R. (1983): Sauria terrestria, Amphisbaenia (Handbuch der Palaoherpetologie, Part 10 A). - Gustav Fischer Verlag, Stuttgart, 249 pp." type="book" year="1983">Estes 1983</bibRefCitation>
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||
).
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<taxonomicName id="4C1A4D2EDF62FFA1FE49FE880A05FE92" authorityName="GRAY" authorityYear="1844" box="[459,613,280,304]" class="Reptilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphisbaenia" pageId="10" pageNumber="353" phylum="Chordata" rank="order">Amphisbaenia</taxonomicName>
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is widely distributed in modern day
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<collectingCountry id="F30D763DDF62FFA1FE03FEA809ADFEF2" box="[385,461,312,336]" name="Turkey" pageId="10" pageNumber="353">Turkey</collectingCountry>
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where the
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<taxonomicName id="4C1A4D2EDF62FFA1FDC3FEA80A88FEF2" baseAuthorityName="Bedriaga" baseAuthorityYear="1884" box="[577,744,312,336]" class="Reptilia" family="Blanidae" genus="Blanus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="353" phylum="Chordata" rank="species" species="strauchi">
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<emphasis id="B96EEABFDF62FFA1FDC3FEA80A88FEF2" box="[577,744,312,336]" italics="true" pageId="10" pageNumber="353">Blanus strauchi</emphasis>
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</taxonomicName>
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species complex is known (
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<bibRefCitation id="EF8B4B5CDF62FFA1FE0FFEC80A3EFED2" author="Sindaco, R. & Kornilios, P. & Sacchi, R. & Lymberakis, P." box="[397,606,344,368]" pageId="10" pageNumber="353" pagination="311 - 326" refId="ref13869" refString="Sindaco, R., Kornilios, P., Sacchi, R., Lymberakis, P. (2014): Taxonomic reassessment of Blanus strauchi (BEDRIAGA, 1884) (Squamata: Amphisbaenia: Blanidae), with the description of a new species from south-east Anatolia (Turkey). - Zootaxa, 3795 (5): 311 - 326. https: // doi. org / 10.11646 / zootaxa. 3795.3.6" type="journal article" year="2014">Sindaco et al. 2014</bibRefCitation>
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). Though the record from Nasrettinhoca 1 probably belongs to
|
||
<taxonomicName id="4C1A4D2EDF62FFA1FDE8FEE80AD4FE32" authorityName="Wagler" authorityYear="1830" box="[618,692,376,400]" class="Reptilia" family="Blanidae" genus="Blanus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="353" phylum="Chordata" rank="genus">
|
||
<emphasis id="B96EEABFDF62FFA1FDE8FEE80AD4FE32" box="[618,692,376,400]" italics="true" pageId="10" pageNumber="353">Blanus</emphasis>
|
||
</taxonomicName>
|
||
(and to
|
||
<taxonomicName id="4C1A4D2EDF62FFA1FF05FE080950FE12" baseAuthorityName="Bedriaga" baseAuthorityYear="1884" box="[135,304,408,432]" class="Reptilia" family="Blanidae" genus="Blanus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="353" phylum="Chordata" rank="species" species="strauchi">
|
||
<emphasis id="B96EEABFDF62FFA1FF05FE080950FE12" box="[135,304,408,432]" italics="true" pageId="10" pageNumber="353">Blanus strauchi</emphasis>
|
||
</taxonomicName>
|
||
species complex), the poor preservation of the material prevents precise identification.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |