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<document id="F7FF4284106178E9EBB69BB9590FD5F7" ID-DOI="10.5281/zenodo.189550" ID-GBIF-Dataset="61a00500-771b-4656-83c9-eff9b3627d9f" ID-ISSN="1175-5326" ID-Zenodo-Dep="189550" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461103111820" checkinUser="plazi" docAuthor="Rowley, Jodi J. L. &amp; Trung, Cao Tien" docDate="2009" docId="03B1B8600A3EFB05E38EFA8E4CCDED26" docLanguage="en" docName="zt02198p060.pdf" docOrigin="Zootaxa 2198" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Leptolalax applebyi Rowley &amp; Trung, 2009, sp. nov." docType="treatment" docVersion="10" lastPageNumber="59" masterDocId="FF88C0180A3FFB0DE319FFC74E18EB04" masterDocTitle="A new species of Leptolalax (Anura: Megophryidae) from central Vietnam" masterLastPageNumber="60" masterPageNumber="51" pageNumber="52" updateTime="1698592069855" updateUser="plazi">
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<mods:title id="43C491678CD5BB8F2B0EE41F1188F319">A new species of Leptolalax (Anura: Megophryidae) from central Vietnam</mods:title>
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<mods:namePart id="CFC9A8AE8007B2C5AF90CA249BFDFB3A">Rowley, Jodi J. L.</mods:namePart>
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<mods:namePart id="1BD3FB259654992C16072DF36FD48F04">Trung, Cao Tien</mods:namePart>
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<mods:date id="AD4D375332ED208441837DB75DDAAA6E">2009</mods:date>
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<treatment id="03B1B8600A3EFB05E38EFA8E4CCDED26" ID-DOI="http://doi.org/10.5281/zenodo.5679225" ID-GBIF-Taxon="119622377" ID-Zenodo-Dep="5679225" LSID="urn:lsid:plazi:treatment:03B1B8600A3EFB05E38EFA8E4CCDED26" httpUri="http://treatment.plazi.org/id/03B1B8600A3EFB05E38EFA8E4CCDED26" lastPageId="8" lastPageNumber="59" pageId="1" pageNumber="52">
<subSubSection id="C3025AFD0A3EFB0CE38EFA8E4FF0EE67" box="[151,488,1353,1379]" pageId="1" pageNumber="52" type="nomenclature">
<paragraph id="8BA709760A3EFB0CE38EFA8E4FF0EE67" blockId="1.[151,488,1353,1379]" box="[151,488,1353,1379]" pageId="1" pageNumber="52">
<heading id="D0EFBE1A0A3EFB0CE38EFA8E4FF0EE67" bold="true" box="[151,488,1353,1379]" fontSize="11" level="1" pageId="1" pageNumber="52" reason="1">
<emphasis id="B96CD5640A3EFB0CE38EFA8E4FF0EE67" bold="true" box="[151,488,1353,1379]" pageId="1" pageNumber="52">
<taxonomicName id="4C1872F50A3EFB0CE38EFA8E4F9CEE67" ID-CoL="3TQPD" box="[151,388,1353,1379]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="1" pageNumber="52" phylum="Chordata" rank="species" species="applebyi" status="sp. nov.">
<emphasis id="B96CD5640A3EFB0CE38EFA8E4F9CEE67" bold="true" box="[151,388,1353,1379]" italics="true" pageId="1" pageNumber="52">Leptolalax applebyi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F681F0A3EFB0CE292FA8E4FF0EE67" box="[395,488,1353,1379]" pageId="1" pageNumber="52" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3025AFD0A3EFB0CE38EFA554C3CED90" pageId="1" pageNumber="52" type="materials_examined">
<paragraph id="8BA709760A3EFB0CE38EFA554A81EEFE" blockId="1.[151,1437,1426,2033]" pageId="1" pageNumber="52">
<typeStatus id="54A3B7D40A3EFB0CE38EFA554F11EEA8" box="[151,265,1426,1452]" pageId="1" pageNumber="52" type="holotype">
<emphasis id="B96CD5640A3EFB0CE38EFA554F11EEA8" bold="true" box="[151,265,1426,1452]" pageId="1" pageNumber="52">Holotype</emphasis>
</typeStatus>
: AMS R171703, an adult male, calling in leaf litter,
<quantity id="4CE0A4930A3EFB0CE065FA554DB2EEA8" box="[892,938,1426,1452]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="52" unit="m" value="2.0">2 m</quantity>
from steep, narrow, rocky stream in Song Thanh Proposed Nature Reserve, Phouc Son district, Quang Nam Province,
<collectingCountry id="F30F49E60A3EFB0CE772FA7E4ACEEED7" box="[1131,1238,1465,1491]" name="Vietnam" pageId="1" pageNumber="52">Vietnam</collectingCountry>
(
<geoCoordinate id="EE2C6FB10A3EFB0CE7F1FA7E4B8FEED7" box="[1256,1431,1465,1491]" direction="north" orientation="latitude" pageId="1" pageNumber="52" precision="1" value="15.27394">15.27394 º N</geoCoordinate>
,
<geoCoordinate id="EE2C6FB10A3EFB0CE38EFA274F27EEFE" box="[151,319,1504,1530]" direction="east" orientation="longitude" pageId="1" pageNumber="52" precision="1" value="107.76015">107.76015 º E</geoCoordinate>
,
<quantity id="4CE0A4930A3EFB0CE252FA274FBFEEFE" box="[331,423,1504,1530]" metricMagnitude="3" metricUnit="m" metricValue="1.402" pageId="1" pageNumber="52" unit="m" value="1402.0">1402 m</quantity>
;
<figureCitation id="132315F30A3EFB0CE2ADFA274C0EEEFE" box="[436,534,1504,1530]" captionStart="FIGURE 1" captionStartId="2.[151,255,1469,1493]" captionTargetBox="[367,1217,274,1445]" captionTargetId="figure@2.[365,1221,271,1445]" captionTargetPageId="2" captionText="FIGURE 1. Type locality of Leptolalax applebyi sp. nov. (black circle), central Vietnam." httpUri="https://zenodo.org/record/189551/files/figure.png" pageId="1" pageNumber="52">Figure 1</figureCitation>
). Collected at 18:15 h on
<date id="FFA62FB60A3EFB0CE05EFA274D86EEFE" box="[839,926,1504,1530]" pageId="1" pageNumber="52" value="1923-07">23 July</date>
, 2007 by the authors.
</paragraph>
<paragraph id="8BA709760A3EFB0CE3DCF9C14C3CED90" blockId="1.[151,1437,1426,2033]" pageId="1" pageNumber="52">
<emphasis id="B96CD5640A3EFB0CE3DCF9C14F5AED24" bold="true" box="[197,322,1542,1568]" pageId="1" pageNumber="52">
<typeStatus id="54A3B7D40A3EFB0CE3DCF9C14F5AED24" box="[197,322,1542,1568]" pageId="1" pageNumber="52" type="paratype">Paratypes</typeStatus>
</emphasis>
: AMS R 171704171706 adult males from
<typeStatus id="54A3B7D40A3EFB0CE054F9C04D98ED25" box="[845,896,1543,1569]" pageId="1" pageNumber="52">type</typeStatus>
locality (
<figureCitation id="132315F30A3EFB0CE0E9F9C04A4BED25" box="[1008,1107,1543,1569]" captionStart="FIGURE 1" captionStartId="2.[151,255,1469,1493]" captionTargetBox="[367,1217,274,1445]" captionTargetId="figure@2.[365,1221,271,1445]" captionTargetPageId="2" captionText="FIGURE 1. Type locality of Leptolalax applebyi sp. nov. (black circle), central Vietnam." httpUri="https://zenodo.org/record/189551/files/figure.png" pageId="1" pageNumber="52">Figure 1</figureCitation>
), collected between 18:16 18:30 h on
<date id="FFA62FB60A3EFB0CE203F9EA4F68ED43" box="[282,368,1581,1607]" pageId="1" pageNumber="52" value="1923-07">23 July</date>
, 2007. All males were calling. AMS R 171707, one adult female collected at 19:00 h, on
<date id="FFA62FB60A3EFB0CE666F9EA4ED1ED6A" pageId="1" pageNumber="52" value="1924-07">24 July</date>
, 2007, from
<geoCoordinate id="EE2C6FB10A3EFB0CE246F9934FE6ED6A" box="[351,510,1620,1646]" direction="north" orientation="latitude" pageId="1" pageNumber="52" precision="1" value="15.26253">15.26253 º N</geoCoordinate>
,
<geoCoordinate id="EE2C6FB10A3EFB0CE113F9934CACED6A" box="[522,692,1620,1646]" direction="east" orientation="longitude" pageId="1" pageNumber="52" precision="1" value="107.75759">107.75759 º E</geoCoordinate>
,
<quantity id="4CE0A4930A3EFB0CE1D8F9934D05ED6A" box="[705,797,1620,1646]" metricMagnitude="3" metricUnit="m" metricValue="1.3119999999999998" pageId="1" pageNumber="52" unit="m" value="1312.0">1312 m</quantity>
. All specimens were found
<quantity id="4CE0A4930A3EFB0CE77CF9934AA8ED6A" box="[1125,1200,1620,1646]" metricMagnitude="0" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="1" pageNumber="52" unit="m" value="1.5" valueMax="2.0" valueMin="1.0">12 m</quantity>
from steep, narrow, rocky streams in evergreen forest.
</paragraph>
</subSubSection>
<subSubSection id="C3025AFD0A3EFB0CE3DCF9664D1EEDE6" pageId="1" pageNumber="52" type="etymology">
<paragraph id="8BA709760A3EFB0CE3DCF9664D1EEDE6" blockId="1.[151,1437,1426,2033]" pageId="1" pageNumber="52">
<emphasis id="B96CD5640A3EFB0CE3DCF9664F4DEDBF" bold="true" box="[197,341,1697,1723]" pageId="1" pageNumber="52">Etymology</emphasis>
: specific epithet is a patronym honouring Robert Appleby, an investor in biodiversity conservation and scientific capacity building in Asia.
</paragraph>
</subSubSection>
<subSubSection id="C3025AFD0A3EFB0FE3DCF9294A83EBDC" lastPageId="2" lastPageNumber="53" pageId="1" pageNumber="52" type="diagnosis">
<paragraph id="8BA709760A3EFB0FE3DCF9294A83EBDC" blockId="1.[151,1437,1426,2033]" lastBlockId="2.[151,1436,152,216]" lastPageId="2" lastPageNumber="53" pageId="1" pageNumber="52">
<emphasis id="B96CD5640A3EFB0CE3DCF9294F25EC0C" bold="true" box="[197,317,1774,1800]" pageId="1" pageNumber="52">Diagnosis</emphasis>
: Assigned to the genus
<taxonomicName id="4C1872F50A3EFB0CE143F9284CCFEC0C" box="[602,727,1775,1800]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="1" pageNumber="52" phylum="Chordata" rank="genus">
<emphasis id="B96CD5640A3EFB0CE143F9284CCFEC0C" box="[602,727,1775,1800]" italics="true" pageId="1" pageNumber="52">Leptolalax</emphasis>
</taxonomicName>
on the basis of the following: small size, the presence of an elevated thenar tubercle not continuous to the thumb, chest glands present but not forming teats, vomerine teeth absent, anterior tip of snout with vertical white bar (Dubois 1980;
<bibRefCitation id="EF8974870A3EFB0CE708F8FB4B1CEC52" author="Lathrop" box="[1041,1284,1852,1878]" pageId="1" pageNumber="60" refString="Lathrop, A., Murphy, R. W., Orlov, N. &amp; Ho, C. T. (1998) Two new species of Leptolalax (Anura: Megophryidae) from northern Vietnam. Amphibia-Reptilia, 19, 253 - 267." type="journal article" year="1998">
Lathrop
<emphasis id="B96CD5640A3EFB0CE762F8FB4AAAEC51" box="[1147,1202,1852,1877]" italics="true" pageId="1" pageNumber="52">et al</emphasis>
. 1998
</bibRefCitation>
).
<taxonomicName id="4C1872F50A3EFB0CE603F8FB4EE4EC78" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="1" pageNumber="52" phylum="Chordata" rank="species" species="applebyi">
<emphasis id="B96CD5640A3EFB0CE603F8FB4EE4EC78" italics="true" pageId="1" pageNumber="52">Leptolalax applebyi</emphasis>
</taxonomicName>
is distinguished from its congeners by a combination of (1) body size (
<quantity id="4CE0A4930A3EFB0CE755F8A54AE1EC78" box="[1100,1273,1890,1916]" metricMagnitude="-2" metricUnit="m" metricValue="2.02" metricValueMax="2.08" metricValueMin="1.96" pageId="1" pageNumber="52" unit="mm" value="20.2" valueMax="20.8" valueMin="19.6">19.620.8 mm</quantity>
for five adult males;
<quantity id="4CE0A4930A3EFB0CE3F7F84E4F43ECA7" box="[238,347,1929,1955]" metricMagnitude="-2" metricUnit="m" metricValue="2.17" pageId="1" pageNumber="52" unit="mm" value="21.7">21.7 mm</quantity>
for single adult female), (2) uniformly smooth dorsum lacking tubercles, (3) dark brown dorsal surface lacking distinct patterns and dark brownish pink ventral surface with white speckling (4) an absence of webbing and dermal fringes on fingers, (5) slight basal webbing and no dermal fringes on toes (6) short tibia (TIB:SVL 0.4660.480), and (7) unique advertisement call consisting of 45 notes with a dominant frequency of 3962.14306.6 Hz, repeated at a rate of approximately 9 notes per second.
</paragraph>
</subSubSection>
<caption id="DF6759FE0A3DFB0FE38EFA7A4A4AEED0" box="[151,1106,1469,1493]" httpUri="https://zenodo.org/record/189551/files/figure.png" pageId="2" pageNumber="53" targetBox="[367,1217,274,1445]" targetPageId="2">
<paragraph id="8BA709760A3DFB0FE38EFA7A4A4AEED0" blockId="2.[151,1106,1469,1493]" box="[151,1106,1469,1493]" pageId="2" pageNumber="53">
<emphasis id="B96CD5640A3DFB0FE38EFA7A4F01EED1" bold="true" box="[151,281,1469,1493]" pageId="2" pageNumber="53">FIGURE 1.</emphasis>
Type locality of
<taxonomicName id="4C1872F50A3DFB0FE2D6FA794CBBEED1" box="[463,675,1470,1493]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="2" pageNumber="53" phylum="Chordata" rank="species" species="applebyi" status="sp. nov.">
<emphasis id="B96CD5640A3DFB0FE2D6FA794CBBEED1" box="[463,675,1470,1493]" italics="true" pageId="2" pageNumber="53">Leptolalax applebyi</emphasis>
</taxonomicName>
<emphasis id="B96CD5640A3DFB0FE1B3FA7A4D18EED1" bold="true" box="[682,768,1469,1493]" pageId="2" pageNumber="53">
<taxonomicNameLabel id="A25F681F0A3DFB0FE1B3FA7A4D18EED1" box="[682,768,1469,1493]" pageId="2" pageNumber="53" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(black circle), central Vietnam.
</paragraph>
</caption>
<subSubSection id="C3025AFD0A3DFB05E3DCF9C54CCDED26" lastPageId="8" lastPageNumber="59" pageId="2" pageNumber="53" type="description">
<paragraph id="8BA709760A3DFB0EE3DCF9C54F49EBFB" blockId="2.[151,1437,1538,2029]" lastBlockId="3.[151,1437,152,255]" lastPageId="3" lastPageNumber="54" pageId="2" pageNumber="53">
<emphasis id="B96CD5640A3DFB0FE3DCF9C54FF0ED18" bold="true" box="[197,488,1538,1564]" pageId="2" pageNumber="53">
Description of
<typeStatus id="54A3B7D40A3DFB0FE264F9C54FF0ED18" box="[381,488,1538,1564]" pageId="2" pageNumber="53" type="holotype">holotype</typeStatus>
</emphasis>
: Head longer than wide; snout rounded or truncate in profile, projecting slightly over lower jaw; nostril closer to tip of snout than eye; canthi rostralis rounded, constricted; lores sloping, concave; vertical pupil; diameter of eye less than length of snout; tympanum distinct, round, diameter smaller than that of the eye; anterioventral three-quaters of tympanic rim slightly elevated relative to skin of temporal region; vomerine teeth absent; pineal ocellus absent; vocal sac openings absent; tongue large, broad, and weakly notched posteriorly; weak supratympanic ridge running from eye towards axilla. Tips of fingers rounded, slightly enlarged; relative finger lengths I&lt;II=IV&lt;III; nuptial pad absent; subarticular tubercles absent; a large, round inner palmar tubercle distinctly separated from small, laterally compressed outer palmar tubercle; no finger webbing or lateral fringes. Tips of toes like fingers; relative toe length I&lt;II&lt;V&lt;III&lt;IV; subarticular tubercles absent, replaced by dermal ridges, distinct on second, third and fourth toes; small, oval inner metatarsal tubercle pronounced, outer metatarsal tubercle absent; webbing basal, confined to very base of toes; no lateral fringes. Tibia short and stout, width approximately one-third of length; tibiotarsal articulation reaches anterior corner of eye. Skin on dorsal and ventral surfaces smooth, only eyelid with low, indistinct tubercles; pectoral gland small, indistinct, circular; femoral gland distinct, small white, on posteroventral surface of thigh, closer to knee than to vent; small gland above axilla; no ventrolateral glandular ridge.
</paragraph>
<caption id="DF6759FE0A3CFB0EE38EF9B64DC6EDAF" httpUri="https://zenodo.org/record/189552/files/figure.png" pageId="3" pageNumber="54" targetBox="[151,1436,313,1615]" targetPageId="3">
<paragraph id="8BA709760A3CFB0EE38EF9B64DC6EDAF" blockId="3.[151,1435,1649,1708]" pageId="3" pageNumber="54">
<emphasis id="B96CD5640A3CFB0EE38EF9B64F02ED8D" bold="true" box="[151,282,1649,1673]" pageId="3" pageNumber="54">FIGURE 2.</emphasis>
(A) Male holotype of
<taxonomicName id="4C1872F50A3CFB0EE116F9B54C90ED8D" box="[527,648,1650,1673]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="3" pageNumber="54" phylum="Chordata" rank="species" species="applebyi" status="sp. nov.">
<emphasis id="B96CD5640A3CFB0EE116F9B54C90ED8D" box="[527,648,1650,1673]" italics="true" pageId="3" pageNumber="54">L. applebyi</emphasis>
</taxonomicName>
<emphasis id="B96CD5640A3CFB0EE189F9B64CFEED8D" bold="true" box="[656,742,1649,1673]" pageId="3" pageNumber="54">
<taxonomicNameLabel id="A25F681F0A3CFB0EE189F9B64CFEED8D" box="[656,742,1649,1673]" pageId="3" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in life (AMS R 171703), and (B) ventral view of male holotype and paratypes of
<taxonomicName id="4C1872F50A3CFB0EE254F9534FDDEDAF" box="[333,453,1684,1707]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="3" pageNumber="54" phylum="Chordata" rank="species" species="applebyi" status="sp. nov.">
<emphasis id="B96CD5640A3CFB0EE254F9534FDDEDAF" box="[333,453,1684,1707]" italics="true" pageId="3" pageNumber="54">L. applebyi</emphasis>
</taxonomicName>
<emphasis id="B96CD5640A3CFB0EE2D5F9534C3AEDA8" bold="true" box="[460,546,1684,1708]" pageId="3" pageNumber="54">
<taxonomicNameLabel id="A25F681F0A3CFB0EE2D5F9534C3AEDA8" box="[460,546,1684,1708]" pageId="3" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in life (AMS R 171703171706, L to R).
</paragraph>
</caption>
<paragraph id="8BA709760A3CFB0BE3DCF91E4DC0EA22" blockId="3.[151,1437,1753,2012]" lastBlockId="6.[151,1437,152,1183]" lastPageId="6" lastPageNumber="57" pageId="3" pageNumber="54">
<emphasis id="B96CD5640A3CFB0EE3DCF91E4C39EDF7" bold="true" box="[197,545,1753,1779]" pageId="3" pageNumber="54">
Colour of
<typeStatus id="54A3B7D40A3CFB0EE257F91E4FD8EDF7" box="[334,448,1753,1779]" pageId="3" pageNumber="54" type="holotype">holotype</typeStatus>
in life:
</emphasis>
Dorsal surface dark brown with indistinct, diffuse darker brown patch between axillae; vertical darker brown bars on upper lip, indistinct lighter vertical stripes at tip of snout and under eyes; no interorbital bar; black line along canthus rostralis, through eye, and continuing along supratympanic ridge, encompassing most of tympanum, terminating above axilla; transverse darker bars on dorsal surface of limbs; large, black blotch on posterior flank anterior to sacrum and two smaller black spots on lateral flank; ventral surface of elbow and upper arm without dark bars, indistinct paler colouration on elbow; fingers with indistinct transverse barring; ventral surface dark brownish pink, with white speckling concentrated on belly, but also on throat, ventral surfaces of arms, tibio-tarsus and feet; fine white speckling extends to ventrolateral surface of belly and on lower margins of head, with a single distinct and slightly larger white spot just posterior to tympanum; ventral margin of throat bearing row of larger white spots. Macroglands white. Iris coppery gold, with minute, black reticulations.
</paragraph>
<caption id="DF6759FE0A3BFB09E38EF84A4B28ECEE" httpUri="https://zenodo.org/record/189553/files/figure.png" pageId="4" pageNumber="55" targetBox="[219,1360,260,1888]" targetPageId="4">
<paragraph id="8BA709760A3BFB09E38EF84A4B28ECEE" blockId="4.[151,1436,1933,2026]" pageId="4" pageNumber="55">
<emphasis id="B96CD5640A3BFB09E38EF84A4F00ECA1" bold="true" box="[151,280,1933,1957]" pageId="4" pageNumber="55">FIGURE 3.</emphasis>
(A) Dorsal view, (B) ventral view, and (C) lateral view of head of preserved holotype (AMS R 171703) of
<taxonomicName id="4C1872F50A3BFB09E69FF84A4EEBECC3" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="4" pageNumber="55" phylum="Chordata" rank="species" species="applebyi" status="sp. nov.">
<emphasis id="B96CD5640A3BFB09E69FF84A4EEBECC3" italics="true" pageId="4" pageNumber="55">L. applebyi</emphasis>
</taxonomicName>
<emphasis id="B96CD5640A3BFB09E3E3F8684F57ECC3" bold="true" box="[250,335,1967,1991]" pageId="4" pageNumber="55">
<taxonomicNameLabel id="A25F681F0A3BFB09E3E3F8684F57ECC3" box="[250,335,1967,1991]" pageId="4" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, (D) ventral surface of right hand of preserved male paratype of
<taxonomicName id="4C1872F50A3BFB09E71CF8774A65ECC3" box="[1029,1149,1968,1991]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="4" pageNumber="55" phylum="Chordata" rank="species" species="applebyi" status="sp. nov.">
<emphasis id="B96CD5640A3BFB09E71CF8774A65ECC3" box="[1029,1149,1968,1991]" italics="true" pageId="4" pageNumber="55">L. applebyi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F681F0A3BFB09E79DF8684AC2ECC3" box="[1156,1242,1967,1991]" pageId="4" pageNumber="55" rank="species">
<emphasis id="B96CD5640A3BFB09E79DF8684AC2ECC3" bold="true" box="[1156,1242,1967,1991]" pageId="4" pageNumber="55">sp. nov.</emphasis>
</taxonomicNameLabel>
(AMS R171705), and (E) lateral view of preserved female paratype of
<taxonomicName id="4C1872F50A3BFB09E1D3F8144D5AECEE" box="[714,834,2003,2026]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="4" pageNumber="55" phylum="Chordata" rank="species" species="applebyi" status="sp. nov.">
<emphasis id="B96CD5640A3BFB09E1D3F8144D5AECEE" box="[714,834,2003,2026]" italics="true" pageId="4" pageNumber="55">L. applebyi</emphasis>
</taxonomicName>
<emphasis id="B96CD5640A3BFB09E050F8154D86ECEE" bold="true" box="[841,926,2002,2026]" pageId="4" pageNumber="55">
<taxonomicNameLabel id="A25F681F0A3BFB09E050F8154D86ECEE" box="[841,926,2002,2026]" pageId="4" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(AMS R 171707). Scale bar = 2 mm.
</paragraph>
</caption>
<caption id="DF6759FE0A3AFB08E38EF84D4CB2ECE3" httpUri="https://zenodo.org/record/189554/files/figure.png" pageId="5" pageNumber="56" targetBox="[180,1414,200,1892]" targetPageId="5">
<paragraph id="8BA709760A3AFB08E38EF84D4CB2ECE3" blockId="5.[151,1436,1930,2023]" pageId="5" pageNumber="56">
<emphasis id="B96CD5640A3AFB08E38EF84D4F00ECA6" bold="true" box="[151,280,1930,1954]" pageId="5" pageNumber="56">FIGURE 4.</emphasis>
Advertisement call of
<taxonomicName id="4C1872F50A3AFB08E112F84D4C9AECA5" box="[523,642,1930,1953]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="5" pageNumber="56" phylum="Chordata" rank="species" species="applebyi" status="sp. nov.">
<emphasis id="B96CD5640A3AFB08E112F84D4C9AECA5" box="[523,642,1930,1953]" italics="true" pageId="5" pageNumber="56">L. applebyi</emphasis>
</taxonomicName>
<emphasis id="B96CD5640A3AFB08E191F84D4CC5ECA6" bold="true" box="[648,733,1930,1954]" pageId="5" pageNumber="56">
<taxonomicNameLabel id="A25F681F0A3AFB08E191F84D4CC5ECA6" box="[648,733,1930,1954]" pageId="5" pageNumber="56" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(holotype AMS R 171703) recorded at ambient air temperature of 21.5º C. (A) a 3 s waveform of relative amplitude, (B) corresponding 3 s spectrogram, (C) expanded, 0.27 s waveform and (D) corresponding spectrogram for same call.
</paragraph>
</caption>
<paragraph id="8BA709760A39FB0BE3DCFEF54CC2EA77" blockId="6.[151,1437,152,1183]" pageId="6" pageNumber="57">
<emphasis id="B96CD5640A39FB0BE3DCFEF54C99EA48" bold="true" box="[197,641,306,332]" pageId="6" pageNumber="57">
Colour of
<typeStatus id="54A3B7D40A39FB0BE25FFEF54FAAEA48" box="[326,434,306,332]" pageId="6" pageNumber="57" type="holotype">holotype</typeStatus>
in preservative:
</emphasis>
Dark brown dorsal surface. Ventral surface pale brown with white speckling and white macroglands. Colours faded.
</paragraph>
<paragraph id="8BA709760A39FB0BE3DCFEB84FA6EAC4" blockId="6.[151,1437,152,1183]" pageId="6" pageNumber="57">
<emphasis id="B96CD5640A39FB0BE3DCFEB84F65EA9D" bold="true" box="[197,381,383,409]" pageId="6" pageNumber="57">Measurements</emphasis>
:
<typeStatus id="54A3B7D40A39FB0BE295FE474FE6EA9E" box="[396,510,384,410]" pageId="6" pageNumber="57" type="holotype">Holotype</typeStatus>
: SVL 19.6, HDL 7.4, HDW 6.6, SNT 3.0, EYE 2.0, IOD 2.6, TMP: 1.1, TEY 0.7, TIB 9.2, weight
<quantity id="4CE0A4930A39FB0BE292FE614FA6EAC4" box="[395,446,422,448]" metricMagnitude="-4" metricUnit="kg" metricValue="8.0" pageId="6" pageNumber="57" unit="g" value="0.8">0.8g</quantity>
</paragraph>
<paragraph id="8BA709760A39FB0BE3DCFE0A4D58E894" blockId="6.[151,1437,152,1183]" pageId="6" pageNumber="57">
<emphasis id="B96CD5640A39FB0BE3DCFE0A4F20EAE3" bold="true" box="[197,312,461,487]" pageId="6" pageNumber="57">Variation</emphasis>
: Specimens vary in dorsal colouration, from uniformly coloured dark brown (AMS R 171706) to weakly mottled (AMS R 1717045). Dark barring on limbs may be indistinct and diffuse (AMS R 171706). Female (AMS R 171707) pale brown with diffuse darker chevrons on dorsum, above axilla. Paler vertical stripes at tip of snout and under eyes vary from indistinct (AMS R 171706) to distinct (AMS R 1717045), most distinct in the single female (AMS R 171707). Ventral surface of chest and belly may be white with dark speckling in places, as opposed to all dark with white speckling. Female (AMS R 171707) has slightly paler throat. Elbows of female (AMS R 171707) and one male (AMS R 171705) are distinctly paler. Colouration on dorsal surface of head between the snout and eyes may be slightly paler. Iris colour varies from uniform gold to uniform reddish copper. Black spots on lateral flank variable in number (14 spots), size and shape. Female (AMS R 171707) has black spot on left knee. Tibiotarsal articulation reaches between anterior corner of eye and nostril. In life, males weighed
<quantity id="4CE0A4930A39FB0BE128FC974C83E86E" box="[561,667,848,874]" metricMagnitude="-4" metricUnit="kg" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="6" pageNumber="57" unit="g" value="0.85" valueMax="0.9" valueMin="0.8">0.80.9g</quantity>
. Female, with distinct follicles present in ovary, weighed 1.0 g in life. Measurements of the
<typeStatus id="54A3B7D40A39FB0BE2D3FCB14FE4E894" box="[458,508,886,912]" pageId="6" pageNumber="57">type</typeStatus>
series are shown in table 1.
</paragraph>
<paragraph id="8BA709760A39FB0BE3DCFC5A4B94EF9B" blockId="6.[151,1437,152,1183]" pageId="6" pageNumber="57">
<emphasis id="B96CD5640A39FB0BE3DCFC5A4FD8E8B3" bold="true" box="[197,448,925,951]" pageId="6" pageNumber="57">Advertisement call</emphasis>
: Call descriptions are based on the calls of the
<typeStatus id="54A3B7D40A39FB0BE707FC5A4A94E8B3" box="[1054,1164,925,951]" pageId="6" pageNumber="57" type="holotype">holotype</typeStatus>
and a non-vouchered individual, both taken at 21.5ºC ambient temperature. Calls consisted of 45 notes, repeated at a rate of approximately 9 per second, with each note consisting of between one and five distinct pulses (Table 2,
<figureCitation id="132315F30A39FB0BE38EFBD64F1EEF2F" box="[151,262,1041,1067]" captionStart="FIGURE 4" captionStartId="5.[151,255,1930,1954]" captionTargetBox="[180,1414,200,1892]" captionTargetId="figure@5.[174,1418,194,1895]" captionTargetPageId="5" captionText="FIGURE 4. Advertisement call of L. applebyi sp. nov. (holotype AMS R 171703) recorded at ambient air temperature of 21.5 º C. (A) a 3 s waveform of relative amplitude, (B) corresponding 3 s spectrogram, (C) expanded, 0.27 s waveform and (D) corresponding spectrogram for same call." httpUri="https://zenodo.org/record/189554/files/figure.png" pageId="6" pageNumber="57">Figures 4</figureCitation>
A, C). There was little frequency modulation, with the dominant frequency between 3962.14306.6 Hz (
<figureCitation id="132315F30A39FB0BE3D3FBFF4F27EF56" box="[202,319,1080,1106]" captionStart="FIGURE 4" captionStartId="5.[151,255,1930,1954]" captionTargetBox="[180,1414,200,1892]" captionTargetId="figure@5.[174,1418,194,1895]" captionTargetPageId="5" captionText="FIGURE 4. Advertisement call of L. applebyi sp. nov. (holotype AMS R 171703) recorded at ambient air temperature of 21.5 º C. (A) a 3 s waveform of relative amplitude, (B) corresponding 3 s spectrogram, (C) expanded, 0.27 s waveform and (D) corresponding spectrogram for same call." httpUri="https://zenodo.org/record/189554/files/figure.png" pageId="6" pageNumber="57">Figures 4</figureCitation>
B, D), and harmonics at approximately 7752 Hz. Pulse duration was between 27 ms. To the human ear, the call of
<taxonomicName id="4C1872F50A39FB0BE286FB984C3CEF7C" box="[415,548,1119,1144]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="6" pageNumber="57" phylum="Chordata" rank="species" species="applebyi">
<emphasis id="B96CD5640A39FB0BE286FB984C3CEF7C" box="[415,548,1119,1144]" italics="true" pageId="6" pageNumber="57">L. applebyi</emphasis>
</taxonomicName>
sounds like a faint, rapid rasping, similar to an orthopteran. The calls of
<taxonomicName id="4C1872F50A39FB0BE69DFB984EE4EF9A" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="6" pageNumber="57" phylum="Chordata" rank="species" species="applebyi">
<emphasis id="B96CD5640A39FB0BE69DFB984EE4EF9A" italics="true" pageId="6" pageNumber="57">L. applebyi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1872F50A39FB0BE22DFB424FDCEF9A" box="[308,452,1157,1182]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="6" pageNumber="57" phylum="Chordata" rank="species" species="tuberosus">
<emphasis id="B96CD5640A39FB0BE22DFB424FDCEF9A" box="[308,452,1157,1182]" italics="true" pageId="6" pageNumber="57">L. tuberosus</emphasis>
</taxonomicName>
differ considerably in structure and frequency (Rowley &amp; Cao, unpublished data).
</paragraph>
<caption id="DF6759FE0A39FB0BE38EFB154AD2EFED" ID-Table-UUID="DF6759FE0A39FB0BE38EFB154AD2EFED" box="[151,1226,1234,1258]" httpUri="http://table.plazi.org/id/DF6759FE0A39FB0BE38EFB154AD2EFED" pageId="6" pageNumber="57" targetBox="[190,1041,1420,1904]" targetIsTable="true" targetPageId="6">
<paragraph id="8BA709760A39FB0BE38EFB154AD2EFED" blockId="6.[151,1226,1234,1258]" box="[151,1226,1234,1258]" pageId="6" pageNumber="57">
<emphasis id="B96CD5640A39FB0BE38EFB154F1AEFEE" bold="true" box="[151,258,1234,1258]" pageId="6" pageNumber="57">TABLE 1</emphasis>
. Measurements (mm) of adult
<taxonomicName id="4C1872F50A39FB0BE154FB154D3AEFED" box="[589,802,1234,1257]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="6" pageNumber="57" phylum="Chordata" rank="species" species="applebyi" status="sp. nov.">
<emphasis id="B96CD5640A39FB0BE154FB154D3AEFED" box="[589,802,1234,1257]" italics="true" pageId="6" pageNumber="57">Leptolalax applebyi</emphasis>
</taxonomicName>
,
<emphasis id="B96CD5640A39FB0BE036FB154D9DEFEE" bold="true" box="[815,901,1234,1258]" pageId="6" pageNumber="57">
<taxonomicNameLabel id="A25F681F0A39FB0BE036FB154D9DEFEE" box="[815,901,1234,1258]" pageId="6" pageNumber="57" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Abbreviations defined in text.
</paragraph>
</caption>
<paragraph id="8BA709760A39FB0AE27CFAF04C3BEA77" blockId="6.[357,1041,1335,1400]" lastBlockId="7.[151,1437,152,371]" lastPageId="7" lastPageNumber="58" pageId="6" pageNumber="57">
Males Female R171703 R171704 R171705 R171706 R171707
<emphasis id="B96CD5640A39FB0BE3DCF86A4F21ECC3" bold="true" box="[197,313,1965,1991]" pageId="6" pageNumber="57">Ecology:</emphasis>
All specimens were found at the headwaters of rocky streams in medium montane forest (evergreen forest between
<quantity id="4CE0A4930A39FB0BE2CFF8134C6FECEA" box="[470,631,2004,2030]" metricMagnitude="3" metricUnit="m" metricValue="1.35" metricValueMax="1.5" metricValueMin="1.2" pageId="6" pageNumber="57" unit="m" value="1350.0" valueMax="1500.0" valueMin="1200.0">12001500m</quantity>
;
<bibRefCitation id="EF8974870A39FB0BE19DF8134D74ECEA" author="Tordoff" box="[644,876,2004,2030]" pageId="6" pageNumber="60" refString="Tordoff, A., Timmins, R., Smith, R. &amp; Mai Ky Vinh (2003) A Biological Assessment of the Central Truong Son Landscape. Central Truong Son Initiative Report No. 1. WWF Indochina, Hanoi, Vietnam. 135 pp." type="book" year="2003">
Tordoff
<emphasis id="B96CD5640A39FB0BE1FFF8134D3BECE9" box="[742,803,2004,2029]" italics="true" pageId="6" pageNumber="57">et al.</emphasis>
2003
</bibRefCitation>
). Streams were steep, narrow (&lt;
<quantity id="4CE0A4930A39FB0BE7F7F8134B05ECEA" box="[1262,1309,2004,2030]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="57" unit="m" value="2.0">2 m</quantity>
wide) and had little water flow. Despite intensive surveys,
<taxonomicName id="4C1872F50A38FB0AE1F4FF5F4D61EBB5" box="[749,889,152,177]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="58" phylum="Chordata" rank="species" species="applebyi">
<emphasis id="B96CD5640A38FB0AE1F4FF5F4D61EBB5" box="[749,889,152,177]" italics="true" pageId="7" pageNumber="58">L. applebyi</emphasis>
</taxonomicName>
was never heard or observed below
<quantity id="4CE0A4930A38FB0AE626FF5F4B84EBB6" box="[1343,1436,152,178]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="7" pageNumber="58" unit="m" value="1300.0">1300 m</quantity>
elevation. Male
<taxonomicName id="4C1872F50A38FB0AE245FF784FFDEBDC" box="[348,485,191,216]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="58" phylum="Chordata" rank="species" species="applebyi">
<emphasis id="B96CD5640A38FB0AE245FF784FFDEBDC" box="[348,485,191,216]" italics="true" pageId="7" pageNumber="58">L. applebyi</emphasis>
</taxonomicName>
were found calling from under leaf litter, always approximately
<quantity id="4CE0A4930A38FB0AE7E4FF794B33EBDC" box="[1277,1323,190,216]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="58" unit="m" value="2.0">2 m</quantity>
from the stream.
<taxonomicName id="4C1872F50A38FB0AE3EDFF224FC6EBFA" box="[244,478,229,254]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="58" phylum="Chordata" rank="species" species="applebyi">
<emphasis id="B96CD5640A38FB0AE3EDFF224FC6EBFA" box="[244,478,229,254]" italics="true" pageId="7" pageNumber="58">Leptolalax applebyi</emphasis>
</taxonomicName>
was found in sympatry with
<taxonomicName id="4C1872F50A38FB0AE02FFF224DDFEBFA" box="[822,967,229,254]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="58" phylum="Chordata" rank="species" species="tuberosus">
<emphasis id="B96CD5640A38FB0AE02FFF224DDFEBFA" box="[822,967,229,254]" italics="true" pageId="7" pageNumber="58">L. tuberosus</emphasis>
</taxonomicName>
.
<taxonomicName id="4C1872F50A38FB0AE0CCFF224AD3EBFA" box="[981,1227,229,254]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="58" phylum="Chordata" rank="species" species="tuberosus">
<emphasis id="B96CD5640A38FB0AE0CCFF224AD3EBFA" box="[981,1227,229,254]" italics="true" pageId="7" pageNumber="58">Leptolalax tuberosus</emphasis>
</taxonomicName>
was more widely distributed in terms of elevation, with individuals collected between
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, and was almost always found greater than
<quantity id="4CE0A4930A38FB0AE26FFEF54FBBEA48" box="[374,419,306,332]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="58" unit="m" value="5.0">5 m</quantity>
distance from streams. The stomach contents of AMS R171707 included remains of a small cockroach (order Blattaria).
</paragraph>
<paragraph id="8BA709760A39FB0BE3FDFA4B4DC6EC74" pageId="6" pageNumber="57">
<table id="F918FBD60A3904F2E3A7FA4B4A09EC74" box="[190,1041,1420,1904]" gridcols="5" gridrows="12" pageId="6" pageNumber="57">
<tr id="35280B340A3904F2E3A7FA4B4A09EEA7" box="[190,1041,1420,1443]" gridrow="0" pageId="6" pageNumber="57">
<th id="76F962480A3904F2E3A7FA4B4F8DEEA7" box="[190,405,1420,1443]" gridcol="0" gridrow="0" pageId="6" pageNumber="57">SVL 19.6</th>
<th id="76F962480A3904F2E2E1FA4B4C3FEEA7" box="[504,551,1420,1443]" gridcol="1" gridrow="0" pageId="6" pageNumber="57">20.7</th>
<th id="76F962480A3904F2E193FA4B4CA1EEA7" box="[650,697,1420,1443]" gridcol="2" gridrow="0" pageId="6" pageNumber="57">20.8</th>
<th id="76F962480A3904F2E005FA4B4D54EEA7" box="[796,844,1420,1443]" gridcol="3" gridrow="0" pageId="6" pageNumber="57">20.8</th>
<th id="76F962480A3904F2E0B6FA4B4A09EEA7" box="[943,1041,1420,1443]" gridcol="4" gridrow="0" pageId="6" pageNumber="57">21.7</th>
</tr>
<tr id="35280B340A3904F2E3A7FA724A09EEC8" box="[190,1041,1461,1484]" gridrow="1" pageId="6" pageNumber="57">
<th id="76F962480A3904F2E3A7FA724F8DEEC8" box="[190,405,1461,1484]" gridcol="0" gridrow="1" pageId="6" pageNumber="57">HDL 7.4</th>
<td id="76F962480A3904F2E2E1FA724C3FEEC8" box="[504,551,1461,1484]" gridcol="1" gridrow="1" pageId="6" pageNumber="57">7.8</td>
<td id="76F962480A3904F2E193FA724CA1EEC8" box="[650,697,1461,1484]" gridcol="2" gridrow="1" pageId="6" pageNumber="57">7.7</td>
<td id="76F962480A3904F2E005FA724D54EEC8" box="[796,844,1461,1484]" gridcol="3" gridrow="1" pageId="6" pageNumber="57">7.6</td>
<td id="76F962480A3904F2E0B6FA724A09EEC8" box="[943,1041,1461,1484]" gridcol="4" gridrow="1" pageId="6" pageNumber="57">8.2</td>
</tr>
<tr id="35280B340A3904F2E3A7FA184A09EEF2" box="[190,1041,1503,1526]" gridrow="2" pageId="6" pageNumber="57">
<th id="76F962480A3904F2E3A7FA184F8DEEF2" box="[190,405,1503,1526]" gridcol="0" gridrow="2" pageId="6" pageNumber="57">HDW 6.6</th>
<td id="76F962480A3904F2E2E1FA184C3FEEF2" box="[504,551,1503,1526]" gridcol="1" gridrow="2" pageId="6" pageNumber="57">7.3</td>
<td id="76F962480A3904F2E193FA184CA1EEF2" box="[650,697,1503,1526]" gridcol="2" gridrow="2" pageId="6" pageNumber="57">7.3</td>
<td id="76F962480A3904F2E005FA184D54EEF2" box="[796,844,1503,1526]" gridcol="3" gridrow="2" pageId="6" pageNumber="57">7.2</td>
<td id="76F962480A3904F2E0B6FA184A09EEF2" box="[943,1041,1503,1526]" gridcol="4" gridrow="2" pageId="6" pageNumber="57">7.4</td>
</tr>
<tr id="35280B340A3904F2E3A7F9CE4A09ED24" box="[190,1041,1545,1568]" gridrow="3" pageId="6" pageNumber="57">
<th id="76F962480A3904F2E3A7F9CE4F8DED24" box="[190,405,1545,1568]" gridcol="0" gridrow="3" pageId="6" pageNumber="57">SNT 3.0</th>
<td id="76F962480A3904F2E2E1F9CE4C3FED24" box="[504,551,1545,1568]" gridcol="1" gridrow="3" pageId="6" pageNumber="57">3.4</td>
<td id="76F962480A3904F2E193F9CE4CA1ED24" box="[650,697,1545,1568]" gridcol="2" gridrow="3" pageId="6" pageNumber="57">3.0</td>
<td id="76F962480A3904F2E005F9CE4D54ED24" box="[796,844,1545,1568]" gridcol="3" gridrow="3" pageId="6" pageNumber="57">3.2</td>
<td id="76F962480A3904F2E0B6F9CE4A09ED24" box="[943,1041,1545,1568]" gridcol="4" gridrow="3" pageId="6" pageNumber="57">3.8</td>
</tr>
<tr id="35280B340A3904F2E3A7F9F34A09ED4F" box="[190,1041,1588,1611]" gridrow="4" pageId="6" pageNumber="57">
<th id="76F962480A3904F2E3A7F9F34F8DED4F" box="[190,405,1588,1611]" gridcol="0" gridrow="4" pageId="6" pageNumber="57">EYE 2.0</th>
<td id="76F962480A3904F2E2E1F9F34C3FED4F" box="[504,551,1588,1611]" gridcol="1" gridrow="4" pageId="6" pageNumber="57">2.4</td>
<td id="76F962480A3904F2E193F9F34CA1ED4F" box="[650,697,1588,1611]" gridcol="2" gridrow="4" pageId="6" pageNumber="57">2.3</td>
<td id="76F962480A3904F2E005F9F34D54ED4F" box="[796,844,1588,1611]" gridcol="3" gridrow="4" pageId="6" pageNumber="57">2.4</td>
<td id="76F962480A3904F2E0B6F9F34A09ED4F" box="[943,1041,1588,1611]" gridcol="4" gridrow="4" pageId="6" pageNumber="57">2.7</td>
</tr>
<tr id="35280B340A3904F2E3A7F99A4A09ED70" box="[190,1041,1629,1652]" gridrow="5" pageId="6" pageNumber="57">
<th id="76F962480A3904F2E3A7F99A4F8DED70" box="[190,405,1629,1652]" gridcol="0" gridrow="5" pageId="6" pageNumber="57">IOD 2.6</th>
<td id="76F962480A3904F2E2E1F99A4C3FED70" box="[504,551,1629,1652]" gridcol="1" gridrow="5" pageId="6" pageNumber="57">2.8</td>
<td id="76F962480A3904F2E193F99A4CA1ED70" box="[650,697,1629,1652]" gridcol="2" gridrow="5" pageId="6" pageNumber="57">3.1</td>
<td id="76F962480A3904F2E005F99A4D54ED70" box="[796,844,1629,1652]" gridcol="3" gridrow="5" pageId="6" pageNumber="57">2.8</td>
<td id="76F962480A3904F2E0B6F99A4A09ED70" box="[943,1041,1629,1652]" gridcol="4" gridrow="5" pageId="6" pageNumber="57">2.9</td>
</tr>
<tr id="35280B340A3904F2E3A7F9404A09ED9A" box="[190,1041,1671,1694]" gridrow="6" pageId="6" pageNumber="57">
<th id="76F962480A3904F2E3A7F9404F8DED9A" box="[190,405,1671,1694]" gridcol="0" gridrow="6" pageId="6" pageNumber="57">TMP 1.1</th>
<td id="76F962480A3904F2E2E1F9404C3FED9A" box="[504,551,1671,1694]" gridcol="1" gridrow="6" pageId="6" pageNumber="57">1.4</td>
<td id="76F962480A3904F2E193F9404CA1ED9A" box="[650,697,1671,1694]" gridcol="2" gridrow="6" pageId="6" pageNumber="57">1.6</td>
<td id="76F962480A3904F2E005F9404D54ED9A" box="[796,844,1671,1694]" gridcol="3" gridrow="6" pageId="6" pageNumber="57">1.5</td>
<td id="76F962480A3904F2E0B6F9404A09ED9A" box="[943,1041,1671,1694]" gridcol="4" gridrow="6" pageId="6" pageNumber="57">1.6</td>
</tr>
<tr id="35280B340A3904F2E3A7F9764A09EDCC" box="[190,1041,1713,1736]" gridrow="7" pageId="6" pageNumber="57">
<th id="76F962480A3904F2E3A7F9764F8DEDCC" box="[190,405,1713,1736]" gridcol="0" gridrow="7" pageId="6" pageNumber="57">TEY 0.7</th>
<td id="76F962480A3904F2E2E1F9764C3FEDCC" box="[504,551,1713,1736]" gridcol="1" gridrow="7" pageId="6" pageNumber="57">1.0</td>
<td id="76F962480A3904F2E193F9764CA1EDCC" box="[650,697,1713,1736]" gridcol="2" gridrow="7" pageId="6" pageNumber="57">0.9</td>
<td id="76F962480A3904F2E005F9764D54EDCC" box="[796,844,1713,1736]" gridcol="3" gridrow="7" pageId="6" pageNumber="57">0.6</td>
<td id="76F962480A3904F2E0B6F9764A09EDCC" box="[943,1041,1713,1736]" gridcol="4" gridrow="7" pageId="6" pageNumber="57">0.9</td>
</tr>
<tr id="35280B340A3904F2E3A7F91B4A09EDF7" box="[190,1041,1756,1779]" gridrow="8" pageId="6" pageNumber="57">
<th id="76F962480A3904F2E3A7F91B4F8DEDF7" box="[190,405,1756,1779]" gridcol="0" gridrow="8" pageId="6" pageNumber="57">TIB 9.2</th>
<td id="76F962480A3904F2E2E1F91B4C3FEDF7" box="[504,551,1756,1779]" gridcol="1" gridrow="8" pageId="6" pageNumber="57">9.7</td>
<td id="76F962480A3904F2E193F91B4CA1EDF7" box="[650,697,1756,1779]" gridcol="2" gridrow="8" pageId="6" pageNumber="57">10.0</td>
<td id="76F962480A3904F2E005F91B4D54EDF7" box="[796,844,1756,1779]" gridcol="3" gridrow="8" pageId="6" pageNumber="57">9.7</td>
<td id="76F962480A3904F2E0B6F91B4A09EDF7" box="[943,1041,1756,1779]" gridcol="4" gridrow="8" pageId="6" pageNumber="57">10.4</td>
</tr>
<tr id="35280B340A3904F2E3A7F8C24A09EC18" box="[190,1041,1797,1820]" gridrow="9" pageId="6" pageNumber="57">
<th id="76F962480A3904F2E3A7F8C24F8DEC18" box="[190,405,1797,1820]" gridcol="0" gridrow="9" pageId="6" pageNumber="57">HDL:HDW 1.11</th>
<td id="76F962480A3904F2E2E1F8C24C3FEC18" box="[504,551,1797,1820]" gridcol="1" gridrow="9" pageId="6" pageNumber="57">1.06</td>
<td id="76F962480A3904F2E193F8C24CA1EC18" box="[650,697,1797,1820]" gridcol="2" gridrow="9" pageId="6" pageNumber="57">1.05</td>
<td id="76F962480A3904F2E005F8C24D54EC18" box="[796,844,1797,1820]" gridcol="3" gridrow="9" pageId="6" pageNumber="57">1.05</td>
<td id="76F962480A3904F2E0B6F8C24A09EC18" box="[943,1041,1797,1820]" gridcol="4" gridrow="9" pageId="6" pageNumber="57">1.1</td>
</tr>
<tr id="35280B340A3904F2E3A7F8E84A09EC42" box="[190,1041,1839,1862]" gridrow="10" pageId="6" pageNumber="57">
<th id="76F962480A3904F2E3A7F8E84F8DEC42" box="[190,405,1839,1862]" gridcol="0" gridrow="10" pageId="6" pageNumber="57">HDL:SVL 0.38</th>
<td id="76F962480A3904F2E2E1F8E84C3FEC42" box="[504,551,1839,1862]" gridcol="1" gridrow="10" pageId="6" pageNumber="57">0.38</td>
<td id="76F962480A3904F2E193F8E84CA1EC42" box="[650,697,1839,1862]" gridcol="2" gridrow="10" pageId="6" pageNumber="57">0.37</td>
<td id="76F962480A3904F2E005F8E84D54EC42" box="[796,844,1839,1862]" gridcol="3" gridrow="10" pageId="6" pageNumber="57">0.36</td>
<td id="76F962480A3904F2E0B6F8E84A09EC42" box="[943,1041,1839,1862]" gridcol="4" gridrow="10" pageId="6" pageNumber="57">0.38</td>
</tr>
<tr id="35280B340A3904F2E3A7F89E4A09EC74" box="[190,1041,1881,1904]" gridrow="11" pageId="6" pageNumber="57">
<th id="76F962480A3904F2E3A7F89E4F8DEC74" box="[190,405,1881,1904]" gridcol="0" gridrow="11" pageId="6" pageNumber="57">TIB:SVL 0.47</th>
<td id="76F962480A3904F2E2E1F89E4C3FEC74" box="[504,551,1881,1904]" gridcol="1" gridrow="11" pageId="6" pageNumber="57">0.47</td>
<td id="76F962480A3904F2E193F89E4CA1EC74" box="[650,697,1881,1904]" gridcol="2" gridrow="11" pageId="6" pageNumber="57">0.48</td>
<td id="76F962480A3904F2E005F89E4D54EC74" box="[796,844,1881,1904]" gridcol="3" gridrow="11" pageId="6" pageNumber="57">0.47</td>
<td id="76F962480A3904F2E0B6F89E4A09EC74" box="[943,1041,1881,1904]" gridcol="4" gridrow="11" pageId="6" pageNumber="57">0.48</td>
</tr>
</table>
</paragraph>
<paragraph id="8BA709760A38FB0AE38EFE614ADEE8B9" pageId="7" pageNumber="58">
<table id="F918FBD60A3804F2E38EFE614B84E8B9" box="[151,1436,422,957]" gridcols="5" gridrows="12" pageId="7" pageNumber="58">
<tr id="35280B340A3804F2E38EFE614B84EAE4" box="[151,1436,422,480]" gridrow="0" pageId="7" pageNumber="58">
<th id="76F962480A3804F2E38EFE614C7DEAE4" box="[151,613,422,480]" gridcol="0" gridrow="0" pageId="7" pageNumber="58">
<emphasis id="B96CD5640A38FB0AE38EFE614F11EABA" bold="true" box="[151,265,422,446]" pageId="7" pageNumber="58">TABLE 2.</emphasis>
Measurements of advertisement are given as means (and ranges).
</th>
<th id="76F962480A3804F2E174FE614DB6EAE4" box="[621,942,422,480]" gridcol="1" gridrow="0" pageId="7" pageNumber="58">
call parameters for
<taxonomicName id="4C1872F50A38FB0AE025FE614DB6EAB9" box="[828,942,422,445]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="58" phylum="Chordata" rank="genus">
<emphasis id="B96CD5640A38FB0AE025FE614DB6EAB9" box="[828,942,422,445]" italics="true" pageId="7" pageNumber="58">Leptolalax</emphasis>
</taxonomicName>
</th>
<th id="76F962480A3804F2E0ACFE614AC6EAE4" box="[949,1246,422,480]" gridcol="2" gridrow="0" pageId="7" pageNumber="58">
<taxonomicName id="4C1872F50A38FB0AE0ACFE614A09EAB9" box="[949,1041,422,445]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="58" phylum="Chordata" rank="species" species="applebyi" status="sp. nov.">
<emphasis id="B96CD5640A38FB0AE0ACFE614A09EAB9" box="[949,1041,422,445]" italics="true" pageId="7" pageNumber="58">applebyi</emphasis>
</taxonomicName>
<emphasis id="B96CD5640A38FB0AE701FE614A76EABA" bold="true" box="[1048,1134,422,446]" pageId="7" pageNumber="58">
<taxonomicNameLabel id="A25F681F0A38FB0AE701FE614A76EABA" box="[1048,1134,422,446]" pageId="7" pageNumber="58" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
at 21.5ºC.
</th>
<th id="76F962480A3804F2E7FCFE614B49EAE4" box="[1253,1361,422,480]" gridcol="3" gridrow="0" pageId="7" pageNumber="58">Parameter</th>
<th id="76F962480A3804F2E641FE614B84EAE4" box="[1368,1436,422,480]" gridcol="4" gridrow="0" pageId="7" pageNumber="58">values</th>
</tr>
<tr id="35280B340A3804F2E38EFE3B4B84E917" box="[151,1436,508,531]" gridrow="1" pageId="7" pageNumber="58" rowspan-3="1" rowspan-4="1">
<th id="76F962480A3804F2E38EFE3B4C7DE917" box="[151,613,508,531]" gridcol="0" gridrow="1" pageId="7" pageNumber="58">Recording</th>
<td id="76F962480A3804F2E174FE3B4DB6E917" box="[621,942,508,531]" gridcol="1" gridrow="1" pageId="7" pageNumber="58">AMS R 171003*</td>
<td id="76F962480A3804F2E0ACFE3B4AC6E917" box="[949,1246,508,531]" gridcol="2" gridrow="1" pageId="7" pageNumber="58">non-vouchered</td>
</tr>
<tr id="35280B340A3804F2E38EFDE14B84E939" box="[151,1436,550,573]" gridrow="2" pageId="7" pageNumber="58" rowspan-3="1" rowspan-4="1">
<th id="76F962480A3804F2E38EFDE14C7DE939" box="[151,613,550,573]" gridcol="0" gridrow="2" pageId="7" pageNumber="58">Number of calls</th>
<td id="76F962480A3804F2E174FDE14DB6E939" box="[621,942,550,573]" gridcol="1" gridrow="2" pageId="7" pageNumber="58">6</td>
<td id="76F962480A3804F2E0ACFDE14AC6E939" box="[949,1246,550,573]" gridcol="2" gridrow="2" pageId="7" pageNumber="58">7</td>
</tr>
<tr id="35280B340A3804F2E38EFD964B84E96C" box="[151,1436,593,616]" gridrow="3" pageId="7" pageNumber="58" rowspan-3="1" rowspan-4="1">
<th id="76F962480A3804F2E38EFD964C7DE96C" box="[151,613,593,616]" gridcol="0" gridrow="3" pageId="7" pageNumber="58">Number of notes</th>
<td id="76F962480A3804F2E174FD964DB6E96C" box="[621,942,593,616]" gridcol="1" gridrow="3" pageId="7" pageNumber="58">24</td>
<td id="76F962480A3804F2E0ACFD964AC6E96C" box="[949,1246,593,616]" gridcol="2" gridrow="3" pageId="7" pageNumber="58">34</td>
</tr>
<tr id="35280B340A3804F2E38EFDBB4B84E997" box="[151,1436,636,659]" gridrow="4" pageId="7" pageNumber="58" rowspan-3="1" rowspan-4="1">
<th id="76F962480A3804F2E38EFDBB4C7DE997" box="[151,613,636,659]" gridcol="0" gridrow="4" pageId="7" pageNumber="58">Call duration (ms)</th>
<td id="76F962480A3804F2E174FDBB4DB6E997" box="[621,942,636,659]" gridcol="1" gridrow="4" pageId="7" pageNumber="58">332 (280358)</td>
<td id="76F962480A3804F2E0ACFDBB4AC6E997" box="[949,1246,636,659]" gridcol="2" gridrow="4" pageId="7" pageNumber="58">405 (340441)</td>
</tr>
<tr id="35280B340A3804F2E38EFD614B84E9B9" box="[151,1436,678,701]" gridrow="5" pageId="7" pageNumber="58" rowspan-3="1" rowspan-4="1">
<th id="76F962480A3804F2E38EFD614C7DE9B9" box="[151,613,678,701]" gridcol="0" gridrow="5" pageId="7" pageNumber="58">Intercall interval (ms)</th>
<td id="76F962480A3804F2E174FD614DB6E9B9" box="[621,942,678,701]" gridcol="1" gridrow="5" pageId="7" pageNumber="58">245 (226287)</td>
<td id="76F962480A3804F2E0ACFD614AC6E9B9" box="[949,1246,678,701]" gridcol="2" gridrow="5" pageId="7" pageNumber="58">287 (234413)</td>
</tr>
<tr id="35280B340A3804F2E38EFD164B84E9EC" box="[151,1436,721,744]" gridrow="6" pageId="7" pageNumber="58" rowspan-3="1" rowspan-4="1">
<th id="76F962480A3804F2E38EFD164C7DE9EC" box="[151,613,721,744]" gridcol="0" gridrow="6" pageId="7" pageNumber="58">Notes/call</th>
<td id="76F962480A3804F2E174FD164DB6E9EC" box="[621,942,721,744]" gridcol="1" gridrow="6" pageId="7" pageNumber="58">4</td>
<td id="76F962480A3804F2E0ACFD164AC6E9EC" box="[949,1246,721,744]" gridcol="2" gridrow="6" pageId="7" pageNumber="58">4.8 (45)</td>
</tr>
<tr id="35280B340A3804F2E38EFD3B4B84E817" box="[151,1436,764,787]" gridrow="7" pageId="7" pageNumber="58" rowspan-3="1" rowspan-4="1">
<th id="76F962480A3804F2E38EFD3B4C7DE817" box="[151,613,764,787]" gridcol="0" gridrow="7" pageId="7" pageNumber="58">Note duration (ms)</th>
<td id="76F962480A3804F2E174FD3B4DB6E817" box="[621,942,764,787]" gridcol="1" gridrow="7" pageId="7" pageNumber="58">16.6 (832)</td>
<td id="76F962480A3804F2E0ACFD3B4AC6E817" box="[949,1246,764,787]" gridcol="2" gridrow="7" pageId="7" pageNumber="58">14.3 (431)</td>
</tr>
<tr id="35280B340A3804F2E38EFCE14B84E839" box="[151,1436,806,829]" gridrow="8" pageId="7" pageNumber="58" rowspan-3="1" rowspan-4="1">
<th id="76F962480A3804F2E38EFCE14C7DE839" box="[151,613,806,829]" gridcol="0" gridrow="8" pageId="7" pageNumber="58">Internote interval (ms)</th>
<td id="76F962480A3804F2E174FCE14DB6E839" box="[621,942,806,829]" gridcol="1" gridrow="8" pageId="7" pageNumber="58">86.7 (48140)</td>
<td id="76F962480A3804F2E0ACFCE14AC6E839" box="[949,1246,806,829]" gridcol="2" gridrow="8" pageId="7" pageNumber="58">83.7 (38113)</td>
</tr>
<tr id="35280B340A3804F2E38EFC964B84E86C" box="[151,1436,849,872]" gridrow="9" pageId="7" pageNumber="58" rowspan-3="1" rowspan-4="1">
<th id="76F962480A3804F2E38EFC964C7DE86C" box="[151,613,849,872]" gridcol="0" gridrow="9" pageId="7" pageNumber="58">Pulses/note</th>
<td id="76F962480A3804F2E174FC964DB6E86C" box="[621,942,849,872]" gridcol="1" gridrow="9" pageId="7" pageNumber="58">3.29 (24)</td>
<td id="76F962480A3804F2E0ACFC964AC6E86C" box="[949,1246,849,872]" gridcol="2" gridrow="9" pageId="7" pageNumber="58">2.52 (15)</td>
</tr>
<tr id="35280B340A3804F2E38EFCBB4B84E897" box="[151,1436,892,915]" gridrow="10" pageId="7" pageNumber="58" rowspan-3="1" rowspan-4="1">
<th id="76F962480A3804F2E38EFCBB4C7DE897" box="[151,613,892,915]" gridcol="0" gridrow="10" pageId="7" pageNumber="58">Note repetition rate (notes/s)</th>
<td id="76F962480A3804F2E174FCBB4DB6E897" box="[621,942,892,915]" gridcol="1" gridrow="10" pageId="7" pageNumber="58">9.0 (8.410.7)</td>
<td id="76F962480A3804F2E0ACFCBB4AC6E897" box="[949,1246,892,915]" gridcol="2" gridrow="10" pageId="7" pageNumber="58">9.3 (8.611.2)</td>
</tr>
<tr id="35280B340A3804F2E38EFC614B84E8B9" box="[151,1436,934,957]" gridrow="11" pageId="7" pageNumber="58" rowspan-3="1" rowspan-4="1">
<th id="76F962480A3804F2E38EFC614C7DE8B9" box="[151,613,934,957]" gridcol="0" gridrow="11" pageId="7" pageNumber="58">Dominant frequency (Hz)</th>
<td id="76F962480A3804F2E174FC614DB6E8B9" box="[621,942,934,957]" gridcol="1" gridrow="11" pageId="7" pageNumber="58">3962.1</td>
<td id="76F962480A3804F2E0ACFC614AC6E8B9" box="[949,1246,934,957]" gridcol="2" gridrow="11" pageId="7" pageNumber="58">4222.9 (3962.14306.6)</td>
</tr>
</table>
</paragraph>
<paragraph id="8BA709760A38FB0AE38EFC264F19E8FC" blockId="7.[151,257,993,1016]" box="[151,257,993,1016]" pageId="7" pageNumber="58">
*
<typeStatus id="54A3B7D40A38FB0AE3B8FC264F19E8FC" box="[161,257,993,1016]" pageId="7" pageNumber="58" type="holotype">holotype</typeStatus>
</paragraph>
<paragraph id="8BA709760A38FB0AE3DCFBED4CBCEE7E" blockId="7.[151,1437,1066,2021]" pageId="7" pageNumber="58">
<emphasis id="B96CD5640A38FB0AE3DCFBED4FDCEF40" bold="true" box="[197,452,1066,1092]" pageId="7" pageNumber="58">Conservation status</emphasis>
: The five
<typeStatus id="54A3B7D40A38FB0AE15DFBED4C60EF40" box="[580,632,1066,1092]" pageId="7" pageNumber="58">type</typeStatus>
specimens are the only known representatives of the new species. Given the available information, we suggest the species should be considered Data Deficient following IUCNs Red List categories (
<bibRefCitation id="EF8974870A38FB0AE2F3FBBF4C62EF96" author="IUCN" box="[490,634,1144,1170]" pageId="7" pageNumber="60" refString="IUCN. (2001) IUCN Red List Categories and Criteria: Version 3.1. IUCN Species Survival Commission. IUCN, Gland, Switzerland and Cambridge, UK. 30 pp." type="book" year="2001">IUCN 2001</bibRefCitation>
). Being known from only two localities approximately
<quantity id="4CE0A4930A38FB0AE618FBBF4B4BEF96" box="[1281,1363,1144,1170]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="7" pageNumber="58" unit="km" value="1.3">1.3 km</quantity>
apart, its extent of occurrence is unknown but probably more widespread. The known habitat of
<taxonomicName id="4C1872F50A38FB0AE7C1FB584B78EFBC" box="[1240,1376,1183,1208]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="58" phylum="Chordata" rank="species" species="applebyi">
<emphasis id="B96CD5640A38FB0AE7C1FB584B78EFBC" box="[1240,1376,1183,1208]" italics="true" pageId="7" pageNumber="58">L. applebyi</emphasis>
</taxonomicName>
falls within a protected area and is remote and steep enough that immediate deforestation risk is likely low. The habitat of
<taxonomicName id="4C1872F50A38FB0AE20AFB2B4F84EE01" box="[275,412,1260,1285]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="58" phylum="Chordata" rank="species" species="applebyi">
<emphasis id="B96CD5640A38FB0AE20AFB2B4F84EE01" box="[275,412,1260,1285]" italics="true" pageId="7" pageNumber="58">L. applebyi</emphasis>
</taxonomicName>
is also contiguous with similar areas of habitat in the central Truong Son landscape (
<bibRefCitation id="EF8974870A38FB0AE387FAD54F99EE28" author="Tordoff" box="[158,385,1298,1324]" pageId="7" pageNumber="60" refString="Tordoff, A., Timmins, R., Smith, R. &amp; Mai Ky Vinh (2003) A Biological Assessment of the Central Truong Son Landscape. Central Truong Son Initiative Report No. 1. WWF Indochina, Hanoi, Vietnam. 135 pp." type="book" year="2003">
Tordoff
<emphasis id="B96CD5640A38FB0AE219FAD44F2CEE28" box="[256,308,1299,1324]" italics="true" pageId="7" pageNumber="58">et al</emphasis>
. 2003
</bibRefCitation>
), including Ngoc Linh Nature Reserve in Kon Tum Province to the south, and
<collectingCountry id="F30F49E60A38FB0AE603FAD54B4BEE28" box="[1306,1363,1298,1324]" name="Laos" pageId="7" pageNumber="58">Laos</collectingCountry>
in the west, but areas above
<quantity id="4CE0A4930A38FB0AE28FFAFE4FF6EE57" box="[406,494,1337,1363]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="7" pageNumber="58" unit="m" value="1200.0">1200 m</quantity>
elevation compose only 8.5% of existing forest in the Truong Son landscape and are patchily distributed (
<bibRefCitation id="EF8974870A38FB0AE2AAFAA74C8FEE7E" author="Tordoff" box="[435,663,1376,1402]" pageId="7" pageNumber="60" refString="Tordoff, A., Timmins, R., Smith, R. &amp; Mai Ky Vinh (2003) A Biological Assessment of the Central Truong Son Landscape. Central Truong Son Initiative Report No. 1. WWF Indochina, Hanoi, Vietnam. 135 pp." type="book" year="2003">
Tordoff
<emphasis id="B96CD5640A38FB0AE10FFAA74C51EE7D" box="[534,585,1376,1401]" italics="true" pageId="7" pageNumber="58">et al</emphasis>
. 2003
</bibRefCitation>
).
</paragraph>
<paragraph id="8BA709760A38FB0AE3DCFA414C50ED8C" blockId="7.[151,1437,1066,2021]" pageId="7" pageNumber="58">
<emphasis id="B96CD5640A38FB0AE3DCFA414F69EEA4" bold="true" box="[197,369,1414,1440]" pageId="7" pageNumber="58">Comparisons</emphasis>
:
<taxonomicName id="4C1872F50A38FB0AE29DFA404C60EEA4" box="[388,632,1415,1440]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="58" phylum="Chordata" rank="species" species="applebyi">
<emphasis id="B96CD5640A38FB0AE29DFA404C60EEA4" box="[388,632,1415,1440]" italics="true" pageId="7" pageNumber="58">Leptolalax applebyi</emphasis>
</taxonomicName>
is distinguished from all other described
<taxonomicName id="4C1872F50A38FB0AE762FA404AE6EEA4" box="[1147,1278,1415,1440]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="58" phylum="Chordata" rank="genus">
<emphasis id="B96CD5640A38FB0AE762FA404AE6EEA4" box="[1147,1278,1415,1440]" italics="true" pageId="7" pageNumber="58">Leptolalax</emphasis>
</taxonomicName>
species by a combination of (1) body size (
<quantity id="4CE0A4930A38FB0AE117FA6A4CDBEEC3" box="[526,707,1453,1479]" metricMagnitude="-2" metricUnit="m" metricValue="2.02" metricValueMax="2.08" metricValueMin="1.96" pageId="7" pageNumber="58" unit="mm" value="20.2" valueMax="20.8" valueMin="19.6">19.620.8 mm</quantity>
for five adult males;
<quantity id="4CE0A4930A38FB0AE0D6FA6A4A24EEC3" box="[975,1084,1453,1479]" metricMagnitude="-2" metricUnit="m" metricValue="2.17" pageId="7" pageNumber="58" unit="mm" value="21.7">21.7 mm</quantity>
for single adult female), (2) uniformly smooth dorsum lacking tubercles, (3) dark brown dorsal surface lacking distinct patterns and dark brownish pink ventral surface with white speckling (4) an absence of webbing and dermal fringes on fingers, (5) slight basal webbing and no dermal fringes on toes, (6) short tibia (TIB:SVL 0.4660.480), and (7) unique advertisement call consisting of 45 notes with a dominant frequency of 3962.14306.6 Hz, repeated at a rate of approximately 9 notes per second.
</paragraph>
<paragraph id="8BA709760A38FB0AE3DCF9524A63ECE1" blockId="7.[151,1437,1066,2021]" pageId="7" pageNumber="58">
<taxonomicName id="4C1872F50A38FB0AE3DCF9524FDEEDAA" box="[197,454,1685,1710]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="58" phylum="Chordata" rank="species" species="applebyi">
<emphasis id="B96CD5640A38FB0AE3DCF9524FDEEDAA" box="[197,454,1685,1710]" italics="true" pageId="7" pageNumber="58">Leptolalax applebyi</emphasis>
</taxonomicName>
is one of the smallest species in the genus. Two species of
<taxonomicName id="4C1872F50A38FB0AE7D7F9524B40EDAA" box="[1230,1368,1685,1710]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="58" phylum="Chordata" rank="genus">
<emphasis id="B96CD5640A38FB0AE7D7F9524B40EDAA" box="[1230,1368,1685,1710]" italics="true" pageId="7" pageNumber="58">Leptolalax</emphasis>
</taxonomicName>
have overlapping male adult body sizes;
<taxonomicName id="4C1872F50A38FB0AE12EF97B4CA2EDD1" box="[567,698,1724,1749]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="58" phylum="Chordata" rank="species" species="pluvialis">
<emphasis id="B96CD5640A38FB0AE12EF97B4CA2EDD1" box="[567,698,1724,1749]" italics="true" pageId="7" pageNumber="58">L. pluvialis</emphasis>
</taxonomicName>
, known only from northern
<collectingCountry id="F30F49E60A38FB0AE71EF97B4A72EDD2" box="[1031,1130,1724,1750]" name="Vietnam" pageId="7" pageNumber="58">Vietnam</collectingCountry>
at
<quantity id="4CE0A4930A38FB0AE797F97B4B23EDD2" box="[1166,1339,1724,1750]" metricMagnitude="-2" metricUnit="m" metricValue="2.18" metricValueMax="2.23" metricValueMin="2.13" pageId="7" pageNumber="58" unit="mm" value="21.8" valueMax="22.3" valueMin="21.3">21.322.3 mm</quantity>
, and the recently described
<taxonomicName id="4C1872F50A38FB0AE26DF9244C37EDF8" box="[372,559,1763,1788]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="58" phylum="Chordata" rank="species" species="kecil">
<emphasis id="B96CD5640A38FB0AE26DF9244C37EDF8" box="[372,559,1763,1788]" italics="true" pageId="7" pageNumber="58">Leptolalax kecil</emphasis>
</taxonomicName>
from the Malay Peninsula, at
<quantity id="4CE0A4930A38FB0AE096F9254A25EDF8" box="[911,1085,1762,1788]" metricMagnitude="-2" metricUnit="m" metricValue="1.9899999999999998" metricValueMax="2.05" metricValueMin="1.93" pageId="7" pageNumber="58" unit="mm" value="19.9" valueMax="20.5" valueMin="19.3">19.320.5 mm</quantity>
. Similar body sizes have been reported for
<taxonomicName id="4C1872F50A38FB0AE236F8CE4FBAEC26" box="[303,418,1801,1826]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="58" phylum="Chordata" rank="species" species="alpinis">
<emphasis id="B96CD5640A38FB0AE236F8CE4FBAEC26" box="[303,418,1801,1826]" italics="true" pageId="7" pageNumber="58">L. alpinis</emphasis>
</taxonomicName>
(24.026.4),
<taxonomicName id="4C1872F50A38FB0AE151F8CE4CC6EC26" box="[584,734,1801,1826]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="58" phylum="Chordata" rank="species" species="heteropus">
<emphasis id="B96CD5640A38FB0AE151F8CE4CC6EC26" box="[584,734,1801,1826]" italics="true" pageId="7" pageNumber="58">L. heteropus</emphasis>
</taxonomicName>
(24.0
<quantity id="4CE0A4930A38FB0AE02FF8CE4D96EC27" box="[822,910,1801,1827]" metricMagnitude="-2" metricUnit="m" metricValue="3.3" pageId="7" pageNumber="58" unit="mm" value="33.0">33 mm</quantity>
),
<taxonomicName id="4C1872F50A38FB0AE0BDF8CE4DF5EC26" box="[932,1005,1801,1826]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="58" phylum="Chordata" rank="species" species="liui">
<emphasis id="B96CD5640A38FB0AE0BDF8CE4DF5EC26" box="[932,1005,1801,1826]" italics="true" pageId="7" pageNumber="58">L. liui</emphasis>
</taxonomicName>
(23.0
<quantity id="4CE0A4930A38FB0AE75AF8CE4AA9EC27" box="[1091,1201,1801,1827]" metricMagnitude="-2" metricUnit="m" metricValue="2.87" pageId="7" pageNumber="58" unit="mm" value="28.7">28.7 mm</quantity>
) and
<taxonomicName id="4C1872F50A38FB0AE7ECF8CE4B92EC26" box="[1269,1418,1801,1826]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="58" phylum="Chordata" rank="species" species="tuberosus">
<emphasis id="B96CD5640A38FB0AE7ECF8CE4B92EC26" box="[1269,1418,1801,1826]" italics="true" pageId="7" pageNumber="58">L. tuberosus</emphasis>
</taxonomicName>
(
<quantity id="4CE0A4930A38FB0AE38EF8F74F51EC4E" box="[151,329,1840,1866]" metricMagnitude="-2" metricUnit="m" metricValue="2.695" metricValueMax="2.95" metricValueMin="2.44" pageId="7" pageNumber="58" unit="mm" value="26.95" valueMax="29.5" valueMin="24.4">24.429.5 mm</quantity>
). All other species have considerably larger male body sizes (
<taxonomicName id="4C1872F50A38FB0AE737F8F74A84EC4D" box="[1070,1180,1840,1865]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="58" phylum="Chordata" rank="species" species="arayai">
<emphasis id="B96CD5640A38FB0AE737F8F74A84EC4D" box="[1070,1180,1840,1865]" italics="true" pageId="7" pageNumber="58">L. arayai</emphasis>
</taxonomicName>
<quantity id="4CE0A4930A38FB0AE7BDF8F74B0AEC4E" box="[1188,1298,1840,1866]" metricMagnitude="-2" metricUnit="m" metricValue="2.96" pageId="7" pageNumber="58" unit="mm" value="29.6">29.6 mm</quantity>
;
<taxonomicName id="4C1872F50A38FB0AE606F8F74B84EC4D" box="[1311,1436,1840,1865]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="58" phylum="Chordata" rank="species" species="bouretti">
<emphasis id="B96CD5640A38FB0AE606F8F74B84EC4D" box="[1311,1436,1840,1865]" italics="true" pageId="7" pageNumber="58">L. bouretti</emphasis>
</taxonomicName>
<quantity id="4CE0A4930A38FB0AE38EF8914F5CEC74" box="[151,324,1878,1904]" metricMagnitude="-2" metricUnit="m" metricValue="3.17" metricValueMax="3.62" metricValueMin="2.7199999999999998" pageId="7" pageNumber="58" unit="mm" value="31.7" valueMax="36.2" valueMin="27.2">27.236.2 mm</quantity>
;
<taxonomicName id="4C1872F50A38FB0AE248F8904FA0EC74" box="[337,440,1879,1904]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="58" phylum="Chordata" rank="species" species="dringi">
<emphasis id="B96CD5640A38FB0AE248F8904FA0EC74" box="[337,440,1879,1904]" italics="true" pageId="7" pageNumber="58">L. dringi</emphasis>
</taxonomicName>
<quantity id="4CE0A4930A38FB0AE2A6F8914C75EC74" box="[447,621,1878,1904]" metricMagnitude="-2" metricUnit="m" metricValue="2.95" metricValueMax="3.03" metricValueMin="2.87" pageId="7" pageNumber="58" unit="mm" value="29.5" valueMax="30.3" valueMin="28.7">28.730.3 mm</quantity>
;
<taxonomicName id="4C1872F50A38FB0AE163F8904D09EC74" box="[634,785,1879,1904]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="58" phylum="Chordata" rank="species" species="fulignosus">
<emphasis id="B96CD5640A38FB0AE163F8904D09EC74" box="[634,785,1879,1904]" italics="true" pageId="7" pageNumber="58">L. fulignosus</emphasis>
</taxonomicName>
28.230.0 mm;
<taxonomicName id="4C1872F50A38FB0AE0CAF8904A54EC74" box="[979,1100,1879,1904]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="58" phylum="Chordata" rank="species" species="gracilis">
<emphasis id="B96CD5640A38FB0AE0CAF8904A54EC74" box="[979,1100,1879,1904]" italics="true" pageId="7" pageNumber="58">L. gracilis</emphasis>
</taxonomicName>
<quantity id="4CE0A4930A38FB0AE74DF8914ACDEC74" box="[1108,1237,1878,1904]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" metricValueMax="3.9" metricValueMin="3.1" pageId="7" pageNumber="58" unit="mm" value="35.0" valueMax="39.0" valueMin="31.0">3139 mm</quantity>
;
<taxonomicName id="4C1872F50A38FB0AE7FBF8904B4AEC74" box="[1250,1362,1879,1904]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="58" phylum="Chordata" rank="species" species="hamidi">
<emphasis id="B96CD5640A38FB0AE7FBF8904B4AEC74" box="[1250,1362,1879,1904]" italics="true" pageId="7" pageNumber="58">L. hamidi</emphasis>
</taxonomicName>
<quantity id="4CE0A4930A38FB0AE643F8914F1DEC93" metricMagnitude="-2" metricUnit="m" metricValue="3.0" metricValueMax="3.13" metricValueMin="2.87" pageId="7" pageNumber="58" unit="mm" value="30.0" valueMax="31.3" valueMin="28.7">28.7 31.3 mm</quantity>
;
<taxonomicName id="4C1872F50A38FB0AE208F8BA4FA5EC92" box="[273,445,1917,1942]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="58" phylum="Chordata" rank="species" species="kajangensis">
<emphasis id="B96CD5640A38FB0AE208F8BA4FA5EC92" box="[273,445,1917,1942]" italics="true" pageId="7" pageNumber="58">L. kajangensis</emphasis>
</taxonomicName>
<quantity id="4CE0A4930A38FB0AE2DDF8BA4C50EC93" box="[452,584,1917,1943]" metricMagnitude="-2" metricUnit="m" metricValue="3.45" metricValueMax="3.5" metricValueMin="3.4" pageId="7" pageNumber="58" unit="mm" value="34.5" valueMax="35.0" valueMin="34.0">3435 mm</quantity>
;
<taxonomicName id="4C1872F50A38FB0AE14DF8BA4CCEEC92" box="[596,726,1917,1942]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="58" phylum="Chordata" rank="species" species="lateralis">
<emphasis id="B96CD5640A38FB0AE14DF8BA4CCEEC92" box="[596,726,1917,1942]" italics="true" pageId="7" pageNumber="58">L. lateralis</emphasis>
</taxonomicName>
<quantity id="4CE0A4930A38FB0AE1C6F8BA4D97EC93" box="[735,911,1917,1943]" metricMagnitude="-2" metricUnit="m" metricValue="2.76" metricValueMax="2.83" metricValueMin="2.69" pageId="7" pageNumber="58" unit="mm" value="27.6" valueMax="28.3" valueMin="26.9">26.928.3 mm</quantity>
;
<taxonomicName id="4C1872F50A38FB0AE082F8BA4A0BEC92" box="[923,1043,1917,1942]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="58" phylum="Chordata" rank="species" species="maurus">
<emphasis id="B96CD5640A38FB0AE082F8BA4A0BEC92" box="[923,1043,1917,1942]" italics="true" pageId="7" pageNumber="58">L. maurus</emphasis>
</taxonomicName>
<quantity id="4CE0A4930A38FB0AE705F8BA4A91EC93" box="[1052,1161,1917,1943]" metricMagnitude="-2" metricUnit="m" metricValue="2.61" pageId="7" pageNumber="58" unit="mm" value="26.1">26.1 mm</quantity>
;
<taxonomicName id="4C1872F50A38FB0AE78FF8BA4B4AEC92" box="[1174,1362,1917,1942]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="58" phylum="Chordata" rank="species" species="melanolectus">
<emphasis id="B96CD5640A38FB0AE78FF8BA4B4AEC92" box="[1174,1362,1917,1942]" italics="true" pageId="7" pageNumber="58">L. melanolectus</emphasis>
</taxonomicName>
<quantity id="4CE0A4930A38FB0AE643F8BA4F1BECBA" metricMagnitude="-2" metricUnit="m" metricValue="2.77" metricValueMax="2.88" metricValueMin="2.66" pageId="7" pageNumber="58" unit="mm" value="27.7" valueMax="28.8" valueMin="26.6">26.6 28.8 mm</quantity>
;
<taxonomicName id="4C1872F50A38FB0AE208F8634FDBECB9" box="[273,451,1956,1981]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="58" phylum="Chordata" rank="species" species="nahangensis">
<emphasis id="B96CD5640A38FB0AE208F8634FDBECB9" box="[273,451,1956,1981]" italics="true" pageId="7" pageNumber="58">L. nahangensis</emphasis>
</taxonomicName>
<quantity id="4CE0A4930A38FB0AE2D5F8634C21ECBA" box="[460,569,1956,1982]" metricMagnitude="-2" metricUnit="m" metricValue="4.08" pageId="7" pageNumber="58" unit="mm" value="40.8">40.8 mm</quantity>
;
<taxonomicName id="4C1872F50A38FB0AE15CF8634CFEECB9" box="[581,742,1956,1981]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="58" phylum="Chordata" rank="species" species="oshanensis">
<emphasis id="B96CD5640A38FB0AE15CF8634CFEECB9" box="[581,742,1956,1981]" italics="true" pageId="7" pageNumber="58">L. oshanensis</emphasis>
</taxonomicName>
<quantity id="4CE0A4930A38FB0AE1F6F8634D87ECBA" box="[751,927,1956,1982]" metricMagnitude="-2" metricUnit="m" metricValue="2.8649999999999998" metricValueMax="3.07" metricValueMin="2.66" pageId="7" pageNumber="58" unit="mm" value="28.65" valueMax="30.7" valueMin="26.6">26.630.7 mm</quantity>
;
<taxonomicName id="4C1872F50A38FB0AE0B2F8634A78ECB9" box="[939,1120,1956,1981]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="58" phylum="Chordata" rank="species" species="pelodytoides">
<emphasis id="B96CD5640A38FB0AE0B2F8634A78ECB9" box="[939,1120,1956,1981]" italics="true" pageId="7" pageNumber="58">L. pelodytoides</emphasis>
</taxonomicName>
<quantity id="4CE0A4930A38FB0AE771F8634AA6ECBA" box="[1128,1214,1956,1982]" metricMagnitude="-2" metricUnit="m" metricValue="3.7" pageId="7" pageNumber="58" unit="mm" value="37.0">37 mm</quantity>
;
<taxonomicName id="4C1872F50A38FB0AE7D5F8634B2AECB9" box="[1228,1330,1956,1981]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="58" phylum="Chordata" rank="species" species="pictus">
<emphasis id="B96CD5640A38FB0AE7D5F8634B2AECB9" box="[1228,1330,1956,1981]" italics="true" pageId="7" pageNumber="58">L. pictus</emphasis>
</taxonomicName>
up to
<quantity id="4CE0A4930A38FB0AE667F8634ED0ECE1" metricMagnitude="-2" metricUnit="m" metricValue="3.6" pageId="7" pageNumber="58" unit="mm" value="36.0">36 mm</quantity>
;
<taxonomicName id="4C1872F50A38FB0AE3CDF80C4F37ECE0" box="[212,303,1995,2020]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="58" phylum="Chordata" rank="species" species="solus">
<emphasis id="B96CD5640A38FB0AE3CDF80C4F37ECE0" box="[212,303,1995,2020]" italics="true" pageId="7" pageNumber="58">L. solus</emphasis>
</taxonomicName>
<quantity id="4CE0A4930A38FB0AE22FF80C4FA0ECE1" box="[310,440,1995,2021]" metricMagnitude="-2" metricUnit="m" metricValue="3.25" metricValueMax="3.5" metricValueMin="3.0" pageId="7" pageNumber="58" unit="mm" value="32.5" valueMax="35.0" valueMin="30.0">3035 mm</quantity>
;
<taxonomicName id="4C1872F50A38FB0AE2DDF80C4C3BECE0" box="[452,547,1995,2020]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="58" phylum="Chordata" rank="species" species="sungi">
<emphasis id="B96CD5640A38FB0AE2DDF80C4C3BECE0" box="[452,547,1995,2020]" italics="true" pageId="7" pageNumber="58">L. sungi</emphasis>
</taxonomicName>
<quantity id="4CE0A4930A38FB0AE133F80C4CCFECE1" box="[554,727,1995,2021]" metricMagnitude="-2" metricUnit="m" metricValue="5.05" metricValueMax="5.27" metricValueMin="4.83" pageId="7" pageNumber="58" unit="mm" value="50.5" valueMax="52.7" valueMin="48.3">48.352.7 mm</quantity>
;
<taxonomicName id="4C1872F50A38FB0AE1FAF80C4DA0ECE0" box="[739,952,1995,2020]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="58" phylum="Chordata" rank="species" species="ventripunctatus">
<emphasis id="B96CD5640A38FB0AE1FAF80C4DA0ECE0" box="[739,952,1995,2020]" italics="true" pageId="7" pageNumber="58">L. ventripunctatus</emphasis>
</taxonomicName>
25.528.0 mm).
</paragraph>
<paragraph id="8BA709760A37FB05E3DCFF5F4F91EA48" blockId="8.[151,1437,152,1570]" pageId="8" pageNumber="59">
The single female specimen of
<taxonomicName id="4C1872F50A37FB05E12EFF5F4CA3EBB5" box="[567,699,152,177]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="applebyi">
<emphasis id="B96CD5640A37FB05E12EFF5F4CA3EBB5" box="[567,699,152,177]" italics="true" pageId="8" pageNumber="59">L. applebyi</emphasis>
</taxonomicName>
(
<quantity id="4CE0A4930A37FB05E1D2FF5F4D20EBB6" box="[715,824,152,178]" metricMagnitude="-2" metricUnit="m" metricValue="2.17" pageId="8" pageNumber="59" unit="mm" value="21.7">21.7 mm</quantity>
) does not fall within the range of female body sizes reported in the genus to date (
<taxonomicName id="4C1872F50A37FB05E2E7FF784C77EBDC" box="[510,623,191,216]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="alpinis">
<emphasis id="B96CD5640A37FB05E2E7FF784C77EBDC" box="[510,623,191,216]" italics="true" pageId="8" pageNumber="59">L. alpinis</emphasis>
</taxonomicName>
<quantity id="4CE0A4930A37FB05E161FF794CFEEBDC" box="[632,742,190,216]" metricMagnitude="-2" metricUnit="m" metricValue="3.21" pageId="8" pageNumber="59" unit="mm" value="32.1">32.1 mm</quantity>
;
<taxonomicName id="4C1872F50A37FB05E1EBFF784D68EBDC" box="[754,880,191,216]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="bouretti">
<emphasis id="B96CD5640A37FB05E1EBFF784D68EBDC" box="[754,880,191,216]" italics="true" pageId="8" pageNumber="59">L. bouretti</emphasis>
</taxonomicName>
42.045.0 mm;
<taxonomicName id="4C1872F50A37FB05E72CFF784A86EBDC" box="[1077,1182,191,216]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="dringi">
<emphasis id="B96CD5640A37FB05E72CFF784A86EBDC" box="[1077,1182,191,216]" italics="true" pageId="8" pageNumber="59">L. dringi</emphasis>
</taxonomicName>
<quantity id="4CE0A4930A37FB05E7BFFF794B0DEBDC" box="[1190,1301,190,216]" metricMagnitude="-2" metricUnit="m" metricValue="3.75" pageId="8" pageNumber="59" unit="mm" value="37.5">37.5 mm</quantity>
;
<taxonomicName id="4C1872F50A37FB05E638FF784B83EBDC" box="[1313,1435,191,216]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="gracilis">
<emphasis id="B96CD5640A37FB05E638FF784B83EBDC" box="[1313,1435,191,216]" italics="true" pageId="8" pageNumber="59">L. gracilis</emphasis>
</taxonomicName>
<quantity id="4CE0A4930A37FB05E38EFF224F01EBFB" box="[151,281,229,255]" metricMagnitude="-2" metricUnit="m" metricValue="4.05" metricValueMax="4.1" metricValueMin="4.0" pageId="8" pageNumber="59" unit="mm" value="40.5" valueMax="41.0" valueMin="40.0">4041 mm</quantity>
;
<taxonomicName id="4C1872F50A37FB05E23CFF224F8DEBFA" box="[293,405,229,254]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="hamidi">
<emphasis id="B96CD5640A37FB05E23CFF224F8DEBFA" box="[293,405,229,254]" italics="true" pageId="8" pageNumber="59">L. hamidi</emphasis>
</taxonomicName>
36.142.8;
<taxonomicName id="4C1872F50A37FB05E13BFF224C60EBFA" box="[546,632,229,254]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="kecil">
<emphasis id="B96CD5640A37FB05E13BFF224C60EBFA" box="[546,632,229,254]" italics="true" pageId="8" pageNumber="59">L. kecil</emphasis>
</taxonomicName>
25.0 mm,
<taxonomicName id="4C1872F50A37FB05E1EEFF224D61EBFA" box="[759,889,229,254]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="lateralis">
<emphasis id="B96CD5640A37FB05E1EEFF224D61EBFA" box="[759,889,229,254]" italics="true" pageId="8" pageNumber="59">L. lateralis</emphasis>
</taxonomicName>
<quantity id="4CE0A4930A37FB05E099FF224DCDEBFB" box="[896,981,229,255]" metricMagnitude="-2" metricUnit="m" metricValue="3.6" pageId="8" pageNumber="59" unit="mm" value="36.0">36 mm</quantity>
;
<taxonomicName id="4C1872F50A37FB05E0FBFF224A30EBFA" box="[994,1064,229,254]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="liui">
<emphasis id="B96CD5640A37FB05E0FBFF224A30EBFA" box="[994,1064,229,254]" italics="true" pageId="8" pageNumber="59">L. liui</emphasis>
</taxonomicName>
<quantity id="4CE0A4930A37FB05E729FF224AC6EBFB" box="[1072,1246,229,255]" metricMagnitude="-2" metricUnit="m" metricValue="2.56" metricValueMax="2.81" metricValueMin="2.31" pageId="8" pageNumber="59" unit="mm" value="25.6" valueMax="28.1" valueMin="23.1">23.128.1 mm</quantity>
;
<taxonomicName id="4C1872F50A37FB05E7F3FF224B79EBFA" box="[1258,1377,229,254]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="maurus">
<emphasis id="B96CD5640A37FB05E7F3FF224B79EBFA" box="[1258,1377,229,254]" italics="true" pageId="8" pageNumber="59">L. maurus</emphasis>
</taxonomicName>
<quantity id="4CE0A4930A37FB05E670FF224ED1EA22" metricMagnitude="-2" metricUnit="m" metricValue="3.18" pageId="8" pageNumber="59" unit="mm" value="31.8">31.8 mm</quantity>
;
<taxonomicName id="4C1872F50A37FB05E3CFFECB4F8CEA21" box="[214,404,268,293]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="melanolectus">
<emphasis id="B96CD5640A37FB05E3CFFECB4F8CEA21" box="[214,404,268,293]" italics="true" pageId="8" pageNumber="59">L. melanolectus</emphasis>
</taxonomicName>
<quantity id="4CE0A4930A37FB05E285FECB4C13EA22" box="[412,523,268,294]" metricMagnitude="-2" metricUnit="m" metricValue="3.27" pageId="8" pageNumber="59" unit="mm" value="32.7">32.7 mm</quantity>
;
<taxonomicName id="4C1872F50A37FB05E101FECB4CA4EA21" box="[536,700,268,293]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="oshanensis">
<emphasis id="B96CD5640A37FB05E101FECB4CA4EA21" box="[536,700,268,293]" italics="true" pageId="8" pageNumber="59">L. oshanensis</emphasis>
</taxonomicName>
<quantity id="4CE0A4930A37FB05E1DDFECB4D2BEA22" box="[708,819,268,294]" metricMagnitude="-2" metricUnit="m" metricValue="3.16" pageId="8" pageNumber="59" unit="mm" value="31.6">31.6 mm</quantity>
;
<taxonomicName id="4C1872F50A37FB05E059FECB4DB0EA21" box="[832,936,268,293]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="pictus">
<emphasis id="B96CD5640A37FB05E059FECB4DB0EA21" box="[832,936,268,293]" italics="true" pageId="8" pageNumber="59">L. pictus</emphasis>
</taxonomicName>
up to
<quantity id="4CE0A4930A37FB05E0EFFECB4A56EA22" box="[1014,1102,268,294]" metricMagnitude="-2" metricUnit="m" metricValue="4.7" pageId="8" pageNumber="59" unit="mm" value="47.0">47 mm</quantity>
;
<taxonomicName id="4C1872F50A37FB05E742FECB4AA5EA21" box="[1115,1213,268,293]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="sungi">
<emphasis id="B96CD5640A37FB05E742FECB4AA5EA21" box="[1115,1213,268,293]" italics="true" pageId="8" pageNumber="59">L. sungi</emphasis>
</taxonomicName>
<quantity id="4CE0A4930A37FB05E7DCFECB4B6FEA22" box="[1221,1399,268,294]" metricMagnitude="-2" metricUnit="m" metricValue="5.779999999999999" metricValueMax="5.89" metricValueMin="5.67" pageId="8" pageNumber="59" unit="mm" value="57.8" valueMax="58.9" valueMin="56.7">56.758.9 mm</quantity>
;
<taxonomicName id="4C1872F50A37FB05E69DFECB4F10EA4F" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="tuberosus">
<emphasis id="B96CD5640A37FB05E69DFECB4F10EA4F" italics="true" pageId="8" pageNumber="59">L. tuberosus</emphasis>
</taxonomicName>
<quantity id="4CE0A4930A37FB05E216FEF54F65EA48" box="[271,381,306,332]" metricMagnitude="-2" metricUnit="m" metricValue="3.02" pageId="8" pageNumber="59" unit="mm" value="30.2">30.2 mm</quantity>
).
</paragraph>
<paragraph id="8BA709760A37FB05E3DCFE9E4A16E9CB" blockId="8.[151,1437,152,1570]" pageId="8" pageNumber="59">
The skin texture and colouration of
<taxonomicName id="4C1872F50A37FB05E168FE9E4CEFEA76" box="[625,759,345,370]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="applebyi">
<emphasis id="B96CD5640A37FB05E168FE9E4CEFEA76" box="[625,759,345,370]" italics="true" pageId="8" pageNumber="59">L. applebyi</emphasis>
</taxonomicName>
differs from all previously described
<taxonomicName id="4C1872F50A37FB05E7A0FE9E4B2FEA76" box="[1209,1335,345,370]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="genus">
<emphasis id="B96CD5640A37FB05E7A0FE9E4B2FEA76" box="[1209,1335,345,370]" italics="true" pageId="8" pageNumber="59">Leptolalax</emphasis>
</taxonomicName>
species. In terms of skin texture, only
<emphasis id="B96CD5640A37FB05E110FE474D37EA9D" box="[521,815,384,409]" italics="true" pageId="8" pageNumber="59">
<taxonomicName id="4C1872F50A37FB05E110FE474C8FEA9D" box="[521,663,384,409]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="applebyi">L. applebyi</taxonomicName>
,
<taxonomicName id="4C1872F50A37FB05E1BDFE474D37EA9D" box="[676,815,384,409]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="pluvialis">L. pluvialis</taxonomicName>
</emphasis>
,
<taxonomicName id="4C1872F50A37FB05E059FE474DC1EA9D" box="[832,985,384,409]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="heteropus">
<emphasis id="B96CD5640A37FB05E059FE474DC1EA9D" box="[832,985,384,409]" italics="true" pageId="8" pageNumber="59">L. heteropus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1872F50A37FB05E0F3FE474A4DEA9D" box="[1002,1109,384,409]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="pictus">
<emphasis id="B96CD5640A37FB05E0F3FE474A4DEA9D" box="[1002,1109,384,409]" italics="true" pageId="8" pageNumber="59">L. pictus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1872F50A37FB05E78BFE474B5CEA9D" box="[1170,1348,384,409]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="kajangensis">
<emphasis id="B96CD5640A37FB05E78BFE474B5CEA9D" box="[1170,1348,384,409]" italics="true" pageId="8" pageNumber="59">L. kajangensis</emphasis>
</taxonomicName>
have a smooth dorsum lacking tubercles or dorsal ridges. Of these species,
<taxonomicName id="4C1872F50A37FB05E0ABFE604A2DEAC4" box="[946,1077,423,448]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="applebyi">
<emphasis id="B96CD5640A37FB05E0ABFE604A2DEAC4" box="[946,1077,423,448]" italics="true" pageId="8" pageNumber="59">L. applebyi</emphasis>
</taxonomicName>
is the only species with a dark brown dorsal surface lacking distinct dorsal patterns (
<taxonomicName id="4C1872F50A37FB05E011FE0A4D80EAE2" box="[776,920,461,486]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="heteropus">
<emphasis id="B96CD5640A37FB05E011FE0A4D80EAE2" box="[776,920,461,486]" italics="true" pageId="8" pageNumber="59">L. heteropus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1872F50A37FB05E0C8FE0A4A2DEAE2" box="[977,1077,461,486]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="pictus">
<emphasis id="B96CD5640A37FB05E0C8FE0A4A2DEAE2" box="[977,1077,461,486]" italics="true" pageId="8" pageNumber="59">L. pictus</emphasis>
</taxonomicName>
have extensive, distinct dorsal markings or patches,
<taxonomicName id="4C1872F50A37FB05E28AFE334C0FE909" box="[403,535,500,525]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="pluvialis">
<emphasis id="B96CD5640A37FB05E28AFE334C0FE909" box="[403,535,500,525]" italics="true" pageId="8" pageNumber="59">L. pluvialis</emphasis>
</taxonomicName>
has distinct black interorbital markings and
<taxonomicName id="4C1872F50A37FB05E73CFE334AD7E909" box="[1061,1231,500,525]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="kajangensis">
<emphasis id="B96CD5640A37FB05E73CFE334AD7E909" box="[1061,1231,500,525]" italics="true" pageId="8" pageNumber="59">L. kajangensis</emphasis>
</taxonomicName>
has almost black dorsal surfaces with minute pale spots). Additionally, most
<taxonomicName id="4C1872F50A37FB05E04EFDDD4DCDE937" box="[855,981,538,563]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="genus">
<emphasis id="B96CD5640A37FB05E04EFDDD4DCDE937" box="[855,981,538,563]" italics="true" pageId="8" pageNumber="59">Leptolalax</emphasis>
</taxonomicName>
species lacking a smooth dorsum also differ from
<taxonomicName id="4C1872F50A37FB05E206FD864FBBE95E" box="[287,419,577,602]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="applebyi">
<emphasis id="B96CD5640A37FB05E206FD864FBBE95E" box="[287,419,577,602]" italics="true" pageId="8" pageNumber="59">L. applebyi</emphasis>
</taxonomicName>
in terms of dorsal colour and pattern (
<taxonomicName id="4C1872F50A37FB05E07EFD864DC1E95E" box="[871,985,577,602]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="gracilis">
<emphasis id="B96CD5640A37FB05E07EFD864DC1E95E" box="[871,985,577,602]" italics="true" pageId="8" pageNumber="59">L gracilis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1872F50A37FB05E70BFD864A9AE95E" box="[1042,1154,577,602]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="hamidi">
<emphasis id="B96CD5640A37FB05E70BFD864A9AE95E" box="[1042,1154,577,602]" italics="true" pageId="8" pageNumber="59">L. hamidi</emphasis>
</taxonomicName>
have extensive, distinct dorsal markings or patches,
<emphasis id="B96CD5640A37FB05E2C6FDAF4D42E985" box="[479,858,616,641]" italics="true" pageId="8" pageNumber="59">
<taxonomicName id="4C1872F50A37FB05E2C6FDAF4C56E985" box="[479,590,616,641]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="arayai">L. arayai</taxonomicName>
,
<taxonomicName id="4C1872F50A37FB05E141FDAF4CEAE985" box="[600,754,616,641]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="fulignosus">L. fulignosus</taxonomicName>
,
<taxonomicName id="4C1872F50A37FB05E1E7FDAF4D42E985" box="[766,858,616,641]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="kecil">L. kecil</taxonomicName>
</emphasis>
,
<taxonomicName id="4C1872F50A37FB05E071FDAF4DF1E985" box="[872,1001,616,641]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="lateralis">
<emphasis id="B96CD5640A37FB05E071FDAF4DF1E985" box="[872,1001,616,641]" italics="true" pageId="8" pageNumber="59">L. lateralis</emphasis>
</taxonomicName>
,
<emphasis id="B96CD5640A37FB05E0EEFDAF4B6EE985" box="[1015,1398,616,641]" italics="true" pageId="8" pageNumber="59">
<taxonomicName id="4C1872F50A37FB05E0EEFDAF4AA2E985" box="[1015,1210,616,641]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="melanoleucus">L. melanoleucus</taxonomicName>
,
<taxonomicName id="4C1872F50A37FB05E7DCFDAF4B6EE985" box="[1221,1398,616,641]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="nahangensis">L. nahangensis</taxonomicName>
</emphasis>
,
<taxonomicName id="4C1872F50A37FB05E69DFDAF4F00E9AC" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="oshanensis">
<emphasis id="B96CD5640A37FB05E69DFDAF4F00E9AC" italics="true" pageId="8" pageNumber="59">L. oshanensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1872F50A37FB05E248FD484FA8E9AC" box="[337,432,655,680]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="sungi">
<emphasis id="B96CD5640A37FB05E248FD484FA8E9AC" box="[337,432,655,680]" italics="true" pageId="8" pageNumber="59">L. sungi</emphasis>
</taxonomicName>
have distinct black interorbital markings,
<taxonomicName id="4C1872F50A37FB05E0BBFD484A01E9AC" box="[930,1049,655,680]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="maurus">
<emphasis id="B96CD5640A37FB05E0BBFD484A01E9AC" box="[930,1049,655,680]" italics="true" pageId="8" pageNumber="59">L. maurus</emphasis>
</taxonomicName>
has almost black dorsal surfaces and
<taxonomicName id="4C1872F50A37FB05E3D1FD724F40E9CA" box="[200,344,693,718]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="tuberosus">
<emphasis id="B96CD5640A37FB05E3D1FD724F40E9CA" box="[200,344,693,718]" italics="true" pageId="8" pageNumber="59">L. tuberosus</emphasis>
</taxonomicName>
has a sandy dorsum with an olive brown hatching pattern).
</paragraph>
<paragraph id="8BA709760A37FB05E3DCFD1B4ACBEF00" blockId="8.[151,1437,152,1570]" pageId="8" pageNumber="59">
The dark brownish pink ventral surface with white speckling of
<taxonomicName id="4C1872F50A37FB05E0CDFD1B4A45E9F1" box="[980,1117,732,757]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="applebyi">
<emphasis id="B96CD5640A37FB05E0CDFD1B4A45E9F1" box="[980,1117,732,757]" italics="true" pageId="8" pageNumber="59">L. applebyi</emphasis>
</taxonomicName>
is also distinctly different from other species in the genus (
<emphasis id="B96CD5640A37FB05E135FCC54A2BE81F" box="[556,1075,770,795]" italics="true" pageId="8" pageNumber="59">
<taxonomicName id="4C1872F50A37FB05E135FCC54CBDE81F" box="[556,677,770,795]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="alpinis">L. alpinis</taxonomicName>
,
<taxonomicName id="4C1872F50A37FB05E1ABFCC54D3FE81F" box="[690,807,770,795]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="arayai">L. arayai</taxonomicName>
,
<taxonomicName id="4C1872F50A37FB05E02DFCC54DA1E81F" box="[820,953,770,795]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="bouretti">L. bouretti</taxonomicName>
,
<taxonomicName id="4C1872F50A37FB05E0DFFCC54A2BE81F" box="[966,1075,770,795]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="dringi">L. dringi</taxonomicName>
</emphasis>
,
<emphasis id="B96CD5640A37FB05E75DFCC54F6FE846" italics="true" pageId="8" pageNumber="59">
<taxonomicName id="4C1872F50A37FB05E75DFCC54AFFE81F" box="[1092,1255,770,795]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="fulignosus">L. fulignosus</taxonomicName>
,
<taxonomicName id="4C1872F50A37FB05E7EDFCC54B6EE81F" box="[1268,1398,770,795]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="gracilis">L. gracilis</taxonomicName>
,
<taxonomicName id="4C1872F50A37FB05E69DFCC54EF4E846" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="hamidi">L. hamidi</taxonomicName>
,
<taxonomicName id="4C1872F50A37FB05E3EEFCEE4F6FE846" box="[247,375,809,834]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="lateralis">L. lateralis</taxonomicName>
</emphasis>
,
<emphasis id="B96CD5640A37FB05E29EFCEE4DE7E846" box="[391,1023,809,834]" italics="true" pageId="8" pageNumber="59">
<taxonomicName id="4C1872F50A37FB05E29EFCEE4FD7E846" box="[391,463,809,834]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="liui">L. liui</taxonomicName>
,
<taxonomicName id="4C1872F50A37FB05E2C5FCEE4C89E846" box="[476,657,809,834]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="nahangensis">L. nahangensis</taxonomicName>
,
<taxonomicName id="4C1872F50A37FB05E184FCEE4D58E846" box="[669,832,809,834]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="oshanensis">L. oshanensis</taxonomicName>
,
<taxonomicName id="4C1872F50A37FB05E055FCEE4DE7E846" box="[844,1023,809,834]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="pelodytoides">L. pelodytoides</taxonomicName>
</emphasis>
,
<emphasis id="B96CD5640A37FB05E714FCEE4B53E846" box="[1037,1355,809,834]" italics="true" pageId="8" pageNumber="59">
<taxonomicName id="4C1872F50A37FB05E714FCEE4A6DE846" box="[1037,1141,809,834]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="pictus">L. pictus</taxonomicName>
,
<taxonomicName id="4C1872F50A37FB05E798FCEE4AF8E846" box="[1153,1248,809,834]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="solus">L. solus</taxonomicName>
,
<taxonomicName id="4C1872F50A37FB05E7F5FCEE4B53E846" box="[1260,1355,809,834]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="sungi">L. sungi</taxonomicName>
</emphasis>
and
<taxonomicName id="4C1872F50A37FB05E69DFCEE4F13E86D" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="tuberosus">
<emphasis id="B96CD5640A37FB05E69DFCEE4F13E86D" italics="true" pageId="8" pageNumber="59">L. tuberosus</emphasis>
</taxonomicName>
have entirely or mostly white or pale grey venters;
<taxonomicName id="4C1872F50A37FB05E09BFC974A13E86D" box="[898,1035,848,873]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="pluvialis">
<emphasis id="B96CD5640A37FB05E09BFC974A13E86D" box="[898,1035,848,873]" italics="true" pageId="8" pageNumber="59">L. pluvialis</emphasis>
</taxonomicName>
has a grey venter with dark grey marbling, uniform pale grey throat with speckling around the border and large whitish grey pectoral glands;
<taxonomicName id="4C1872F50A37FB05E69DFCB04F2AE8B2" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="melanolectus">
<emphasis id="B96CD5640A37FB05E69DFCB04F2AE8B2" italics="true" pageId="8" pageNumber="59">L. melanolectus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1872F50A37FB05E274FC5A4C5CE8B2" box="[365,580,925,950]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="ventripunctatus">
<emphasis id="B96CD5640A37FB05E274FC5A4C5CE8B2" box="[365,580,925,950]" italics="true" pageId="8" pageNumber="59">L. ventripunctatus</emphasis>
</taxonomicName>
display large patches of distinct black and white marbling,
<taxonomicName id="4C1872F50A37FB05E610FC5A4B84E8B2" box="[1289,1436,925,950]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="heteropus">
<emphasis id="B96CD5640A37FB05E610FC5A4B84E8B2" box="[1289,1436,925,950]" italics="true" pageId="8" pageNumber="59">L. heteropus</emphasis>
</taxonomicName>
has a grey venter, speckled with black;
<taxonomicName id="4C1872F50A37FB05E17DFC034CC4E8D9" box="[612,732,964,989]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="maurus">
<emphasis id="B96CD5640A37FB05E17DFC034CC4E8D9" box="[612,732,964,989]" italics="true" pageId="8" pageNumber="59">L. maurus</emphasis>
</taxonomicName>
has a black or dark grey brown venter, with indistinct small light areas, and
<taxonomicName id="4C1872F50A37FB05E249FC2D4FBEEF07" box="[336,422,1002,1027]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="kecil">
<emphasis id="B96CD5640A37FB05E249FC2D4FBEEF07" box="[336,422,1002,1027]" italics="true" pageId="8" pageNumber="59">L. kecil</emphasis>
</taxonomicName>
has a uniformly dark venter with large, dark orange pectoral glands).
</paragraph>
<paragraph id="8BA709760A37FB05E3DCFBD64C53EF9B" blockId="8.[151,1437,152,1570]" pageId="8" pageNumber="59">
In lacking webbing and dermal fringes on fingers and having only slight basal webbing and no dermal fringes on toes,
<taxonomicName id="4C1872F50A37FB05E243FBFF4FF9EF55" box="[346,481,1080,1105]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="applebyi">
<emphasis id="B96CD5640A37FB05E243FBFF4FF9EF55" box="[346,481,1080,1105]" italics="true" pageId="8" pageNumber="59">L. applebyi</emphasis>
</taxonomicName>
is distinguished further from
<emphasis id="B96CD5640A37FB05E055FBFF4D04EF7C" italics="true" pageId="8" pageNumber="59">
<taxonomicName id="4C1872F50A37FB05E055FBFF4DDAEF55" box="[844,962,1080,1105]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="alpinis">L. alpinis</taxonomicName>
,
<taxonomicName id="4C1872F50A37FB05E0D6FBFF4A48EF55" box="[975,1104,1080,1105]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="bouretti">L. bouretti</taxonomicName>
,
<taxonomicName id="4C1872F50A37FB05E744FBFF4AD3EF55" box="[1117,1227,1080,1105]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="dringi">L. dringi</taxonomicName>
,
<taxonomicName id="4C1872F50A37FB05E7C1FBFF4B6FEF55" box="[1240,1399,1080,1105]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="fulignosus">L. fulignosus</taxonomicName>
,
<taxonomicName id="4C1872F50A37FB05E69DFBFF4EF6EF7C" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="hamidi">L. hamidi</taxonomicName>
,
<taxonomicName id="4C1872F50A37FB05E3E3FB984F8CEF7C" box="[250,404,1119,1144]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="heteropus">L. heteropus</taxonomicName>
,
<taxonomicName id="4C1872F50A37FB05E2B8FB984FE5EF7C" box="[417,509,1119,1144]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="kecil">L. kecil</taxonomicName>
,
<taxonomicName id="4C1872F50A37FB05E113FB984C4EEF7C" box="[522,598,1119,1144]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="liui">L. liui</taxonomicName>
,
<taxonomicName id="4C1872F50A37FB05E17DFB984D04EF7C" box="[612,796,1119,1144]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="pelodytoides">L. pelodytoides</taxonomicName>
</emphasis>
,
<emphasis id="B96CD5640A37FB05E035FB984A04EF7C" box="[812,1052,1119,1144]" italics="true" pageId="8" pageNumber="59">
<taxonomicName id="4C1872F50A37FB05E035FB984DA8EF7C" box="[812,944,1119,1144]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="pluvalis">L. pluvalis</taxonomicName>
,
<taxonomicName id="4C1872F50A37FB05E0A4FB984A04EF7C" box="[957,1052,1119,1144]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="solus">L. solus</taxonomicName>
</emphasis>
and
<taxonomicName id="4C1872F50A37FB05E741FB984AA1EF7C" box="[1112,1209,1119,1144]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="sungi">
<emphasis id="B96CD5640A37FB05E741FB984AA1EF7C" box="[1112,1209,1119,1144]" italics="true" pageId="8" pageNumber="59">L. sungi</emphasis>
</taxonomicName>
, all of which have dermal fringes or extensive webbing.
</paragraph>
<paragraph id="8BA709760A37FB05E3DCFB6B4AE7EFE8" blockId="8.[151,1437,152,1570]" pageId="8" pageNumber="59">
The relative tibia length of
<taxonomicName id="4C1872F50A37FB05E115FB6B4C8AEFC1" box="[524,658,1196,1221]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="applebyi">
<emphasis id="B96CD5640A37FB05E115FB6B4C8AEFC1" box="[524,658,1196,1221]" italics="true" pageId="8" pageNumber="59">L. applebyi</emphasis>
</taxonomicName>
(TIB:SVL 0.470.48) distinguishes it from
<taxonomicName id="4C1872F50A37FB05E7B8FB6B4AE3EFC1" box="[1185,1275,1196,1221]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="kucil">
<emphasis id="B96CD5640A37FB05E7B8FB6B4AE3EFC1" box="[1185,1275,1196,1221]" italics="true" pageId="8" pageNumber="59">L. kucil</emphasis>
</taxonomicName>
(males 0.56 0.58)
<taxonomicName id="4C1872F50A37FB05E3C2FB144F47EFE8" box="[219,351,1235,1260]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="pluvialis">
<emphasis id="B96CD5640A37FB05E3C2FB144F47EFE8" box="[219,351,1235,1260]" italics="true" pageId="8" pageNumber="59">L. pluvialis</emphasis>
</taxonomicName>
(0.520.56)
<taxonomicName id="4C1872F50A37FB05E2EFFB144CB1EFE8" box="[502,681,1235,1260]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="pelodytoides">
<emphasis id="B96CD5640A37FB05E2EFFB144CB1EFE8" box="[502,681,1235,1260]" italics="true" pageId="8" pageNumber="59">L. pelodytoides</emphasis>
</taxonomicName>
(0.56),
<taxonomicName id="4C1872F50A37FB05E01FFB144D6AEFE8" box="[774,882,1235,1260]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="arayai">
<emphasis id="B96CD5640A37FB05E01FFB144D6AEFE8" box="[774,882,1235,1260]" italics="true" pageId="8" pageNumber="59">L. arayai</emphasis>
</taxonomicName>
(0.62), and
<taxonomicName id="4C1872F50A37FB05E719FB144AB1EFE8" box="[1024,1193,1235,1260]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="kajangensis">
<emphasis id="B96CD5640A37FB05E719FB144AB1EFE8" box="[1024,1193,1235,1260]" italics="true" pageId="8" pageNumber="59">L. kajangensis</emphasis>
</taxonomicName>
(0.42).
</paragraph>
<paragraph id="8BA709760A37FB05E3DCFB3E4CCDED26" blockId="8.[151,1437,152,1570]" pageId="8" pageNumber="59">
The advertisement call of
<taxonomicName id="4C1872F50A37FB05E2EFFB3E4C62EE16" box="[502,634,1273,1298]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="applebyi">
<emphasis id="B96CD5640A37FB05E2EFFB3E4C62EE16" box="[502,634,1273,1298]" italics="true" pageId="8" pageNumber="59">L. applebyi</emphasis>
</taxonomicName>
also differs from the twelve
<taxonomicName id="4C1872F50A37FB05E0D2FB3E4A50EE16" box="[971,1096,1273,1298]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="genus">
<emphasis id="B96CD5640A37FB05E0D2FB3E4A50EE16" box="[971,1096,1273,1298]" italics="true" pageId="8" pageNumber="59">Leptolalax</emphasis>
</taxonomicName>
species with described calls;
<emphasis id="B96CD5640A37FB05E38EFAE74FAEEE64" italics="true" pageId="8" pageNumber="59">
<taxonomicName id="4C1872F50A37FB05E38EFAE74F09EE3D" box="[151,273,1312,1337]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="arayai">L. arayai</taxonomicName>
,
<taxonomicName id="4C1872F50A37FB05E206FAE74F8CEE3D" box="[287,404,1312,1337]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="dringi">L. dringi</taxonomicName>
,
<taxonomicName id="4C1872F50A37FB05E2BBFAE74C51EE3D" box="[418,585,1312,1337]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="fuliginous">L. fuliginous</taxonomicName>
,
<taxonomicName id="4C1872F50A37FB05E14EFAE74CF9EE3D" box="[599,737,1312,1337]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="gracilis">L. gracilis</taxonomicName>
,
<taxonomicName id="4C1872F50A37FB05E1F6FAE74D76EE3D" box="[751,878,1312,1337]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="hamidi">L. hamidi</taxonomicName>
,
<taxonomicName id="4C1872F50A37FB05E065FAE74A07EE3D" box="[892,1055,1312,1337]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="heteropus">L. heteropus</taxonomicName>
,
<taxonomicName id="4C1872F50A37FB05E737FAE74A88EE3D" box="[1070,1168,1312,1337]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="kecil">L. kecil</taxonomicName>
,
<taxonomicName id="4C1872F50A37FB05E786FAE74B6DEE3D" box="[1183,1397,1312,1337]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="melanoleucus">L. melanoleucus</taxonomicName>
,
<taxonomicName id="4C1872F50A37FB05E69AFAE74F22EE64" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="pelodytoides">L. pelodytoides</taxonomicName>
,
<taxonomicName id="4C1872F50A37FB05E251FA804FAEEE64" box="[328,438,1351,1376]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="pictus">L. pictus</taxonomicName>
</emphasis>
and
<emphasis id="B96CD5640A37FB05E2ECFA804D0FEE64" box="[501,791,1351,1376]" italics="true" pageId="8" pageNumber="59">
<taxonomicName id="4C1872F50A37FB05E2ECFA804CBDEE64" box="[501,677,1351,1376]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="oshanensis">L. oshanensis</taxonomicName>
,
<taxonomicName id="4C1872F50A37FB05E1AAFA804D0FEE64" box="[691,791,1351,1376]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="solus">L. solus</taxonomicName>
</emphasis>
. Whereas the advertisement calls of
<emphasis id="B96CD5640A37FB05E7F7FA804C2DEE82" italics="true" pageId="8" pageNumber="59">
<taxonomicName id="4C1872F50A37FB05E7F7FA804B6DEE64" box="[1262,1397,1351,1376]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="gracilis">L. gracilis</taxonomicName>
,
<taxonomicName id="4C1872F50A37FB05E69AFA804F15EE82" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="heteropus">L. heteropus</taxonomicName>
,
<taxonomicName id="4C1872F50A37FB05E201FAAA4F68EE82" box="[280,368,1389,1414]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="kecil">L. kecil</taxonomicName>
,
<taxonomicName id="4C1872F50A37FB05E265FAAA4C29EE82" box="[380,561,1389,1414]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="pelodytoides">L. pelodytoides</taxonomicName>
,
</emphasis>
and
<taxonomicName id="4C1872F50A37FB05E177FAAA4CD0EE82" box="[622,712,1389,1414]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="solus">
<emphasis id="B96CD5640A37FB05E177FAAA4CD0EE82" box="[622,712,1389,1414]" italics="true" pageId="8" pageNumber="59">L. solus</emphasis>
</taxonomicName>
overlap with
<taxonomicName id="4C1872F50A37FB05E073FAAA4DF5EE82" box="[874,1005,1389,1414]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="applebyi">
<emphasis id="B96CD5640A37FB05E073FAAA4DF5EE82" box="[874,1005,1389,1414]" italics="true" pageId="8" pageNumber="59">L. applebyi</emphasis>
</taxonomicName>
in terms of number of notes per call, compared to
<taxonomicName id="4C1872F50A37FB05E222FA534FD0EEA9" box="[315,456,1428,1453]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="applebyi">
<emphasis id="B96CD5640A37FB05E222FA534FD0EEA9" box="[315,456,1428,1453]" italics="true" pageId="8" pageNumber="59">L. applebyi</emphasis>
</taxonomicName>
, the calls of
<emphasis id="B96CD5640A37FB05E177FA534D86EEA9" box="[622,926,1428,1453]" italics="true" pageId="8" pageNumber="59">
<taxonomicName id="4C1872F50A37FB05E177FA534CEBEEA9" box="[622,755,1428,1453]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="gracilis">L. gracilis</taxonomicName>
,
<taxonomicName id="4C1872F50A37FB05E018FA534D86EEA9" box="[769,926,1428,1453]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="heteropus">L. heteropus</taxonomicName>
</emphasis>
and
<taxonomicName id="4C1872F50A37FB05E0C5FA534A26EEA9" box="[988,1086,1428,1453]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="solus">
<emphasis id="B96CD5640A37FB05E0C5FA534A26EEA9" box="[988,1086,1428,1453]" italics="true" pageId="8" pageNumber="59">L. solus</emphasis>
</taxonomicName>
have much lower dominant frequencies (25403200Hz) and
<taxonomicName id="4C1872F50A37FB05E129FA7C4CF7EED0" box="[560,751,1467,1492]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="pelodytoides">
<emphasis id="B96CD5640A37FB05E129FA7C4CF7EED0" box="[560,751,1467,1492]" italics="true" pageId="8" pageNumber="59">L. pelodytoides</emphasis>
</taxonomicName>
has higher dominant frequencies (63508100Hz). The advertisement call of
<taxonomicName id="4C1872F50A37FB05E282FA264FEBEEFE" box="[411,499,1505,1530]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="kecil">
<emphasis id="B96CD5640A37FB05E282FA264FEBEEFE" box="[411,499,1505,1530]" italics="true" pageId="8" pageNumber="59">L. kecil</emphasis>
</taxonomicName>
has both a lower dominant frequency, a lower number of pulses per note, and higher intercall interval compared to
<taxonomicName id="4C1872F50A37FB05E153F9CF4CD6ED25" box="[586,718,1544,1569]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="applebyi">
<emphasis id="B96CD5640A37FB05E153F9CF4CD6ED25" box="[586,718,1544,1569]" italics="true" pageId="8" pageNumber="59">L. applebyi</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>