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<mods:title id="9CEC9EE0D7D229971056A6EEC5AA72B4">Taxonomic revision of the subterranean genus Virpazaria Gittenberger, 1969 (Gastropoda, Spelaeodiscidae)</mods:title>
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<paragraph id="8BA7CB4CF710FC5FFDF0F1EF6991FC35" blockId="2.[575,1012,958,984]" box="[575,1012,958,984]" pageId="2" pageNumber="3">
<heading id="D0EF7C20F710FC5FFDF0F1EF6991FC35" box="[575,1012,958,984]" centered="true" fontSize="11" level="2" pageId="2" pageNumber="3" reason="2">
Genus
<taxonomicName id="4C18B0CFF710FC5FFD5DF1EF6991FC35" ID-CoL="7PKV4" authority="Gittenberger, 1969" authorityName="Gittenberger" authorityYear="1969" box="[658,1012,958,984]" class="Gastropoda" family="Spelaeodiscidae" genus="Virpazaria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="3" phylum="Mollusca" rank="genus">
<emphasis id="B96C175EF710FC5FFD5DF1EF6974FC35" bold="true" box="[658,785,958,984]" italics="true" pageId="2" pageNumber="3">Virpazaria</emphasis>
<bibRefCitation id="EF89B6BDF710FC5FFCD7F1EF6991FC35" author="Gittenberger E." box="[792,1012,958,984]" firstAuthor="Gittenberger" journalOrPublisher="Zoologische Mededelingen" pageId="2" pageNumber="3" pagination="287 - 306" part="43" refId="ref12709" refString="Gittenberger E. 1969. Beitrage zur Kenntnis der Pupillacea. I. Die Spelaeodiscinae. Zoologische Mededelingen 43 (22): 287 - 306." title="Beitrage zur Kenntnis der Pupillacea. I. Die Spelaeodiscinae" type="journal article" year="1969">Gittenberger, 1969</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C30298C7F710FC5FFF72F65169D6FBA5" pageId="2" pageNumber="3" type="type_taxon">
<paragraph id="8BA7CB4CF710FC5FFF72F6516B3DFBF4" blockId="2.[189,344,1023,1050]" box="[189,344,1023,1050]" pageId="2" pageNumber="3">
<heading id="D0EF7C20F710FC5FFF72F6516B3DFBF4" bold="true" box="[189,344,1023,1050]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
<emphasis id="B96C175EF710FC5FFF72F6516B3DFBF4" bold="true" box="[189,344,1023,1050]" pageId="2" pageNumber="3">
<typeStatus id="54A375EEF710FC5FFF72F6516A9FFBF7" box="[189,250,1024,1050]" pageId="2" pageNumber="3">Type</typeStatus>
species
</emphasis>
</heading>
</paragraph>
<paragraph id="8BA7CB4CF710FC5FFF72F67E69D6FBA5" blockId="2.[189,947,1070,1096]" box="[189,947,1070,1096]" pageId="2" pageNumber="3">
<taxonomicName id="4C18B0CFF710FC5FFF72F67E68F4FBA5" ID-CoL="5BK6Y" authority="Gittenberger" authorityName="Gittenberger" authorityYear="1969" box="[189,657,1070,1096]" class="Gastropoda" family="Spelaeodiscidae" genus="Virpazaria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="3" phylum="Mollusca" rank="species" species="adrianae">
<emphasis id="B96C175EF710FC5FFF72F67E6BCEFBA5" box="[189,427,1070,1096]" italics="true" pageId="2" pageNumber="3">Virpazaria adrianae</emphasis>
<bibRefCitation id="EF89B6BDF710FC5FFE7CF67F68F4FBA5" author="Gittenberger E." box="[435,657,1070,1096]" firstAuthor="Gittenberger" journalOrPublisher="Zoologische Mededelingen" pageId="2" pageNumber="3" pagination="287 - 306" part="43" refId="ref12709" refString="Gittenberger E. 1969. Beitrage zur Kenntnis der Pupillacea. I. Die Spelaeodiscinae. Zoologische Mededelingen 43 (22): 287 - 306." title="Beitrage zur Kenntnis der Pupillacea. I. Die Spelaeodiscinae" type="journal article" year="1969">Gittenberger, 1969</bibRefCitation>
</taxonomicName>
, by original designation.
</paragraph>
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<subSubSection id="C30298C7F710FC5FFF72F63E6B8AFAA8" pageId="2" pageNumber="3" type="diagnosis">
<paragraph id="8BA7CB4CF710FC5FFF72F63E6B53FB64" blockId="2.[189,310,1135,1161]" box="[189,310,1135,1161]" pageId="2" pageNumber="3">
<heading id="D0EF7C20F710FC5FFF72F63E6B53FB64" bold="true" box="[189,310,1135,1161]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
<emphasis id="B96C175EF710FC5FFF72F63E6B53FB64" bold="true" box="[189,310,1135,1161]" pageId="2" pageNumber="3">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BA7CB4CF710FC5FFF72F6CF6B8AFAA8" blockId="2.[189,1399,1182,1349]" pageId="2" pageNumber="3">Shell discoid, dextral, ribbed, colourless (weathered specimens are opaque white), peristome attached, thickened, usually with knob- or tooth-like prominences on palatal and/or basal sides. Peristome insertions are connected; this parietal connection varies from a swollen bulge to a lamelliform flap; in some species this flap might cover a significant part of the aperture in frontal view. Aperture somewhat narrow and moon-shaped.</paragraph>
</subSubSection>
<subSubSection id="C30298C7F710FC5FFF72F73D6951F80E" pageId="2" pageNumber="3" type="discussion">
<paragraph id="8BA7CB4CF710FC5FFF72F73D6B4AFA6B" blockId="2.[189,303,1388,1414]" box="[189,303,1388,1414]" pageId="2" pageNumber="3">
<heading id="D0EF7C20F710FC5FFF72F73D6B4AFA6B" bold="true" box="[189,303,1388,1414]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
<emphasis id="B96C175EF710FC5FFF72F73D6B4AFA6B" bold="true" box="[189,303,1388,1414]" pageId="2" pageNumber="3">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BA7CB4CF710FC5FFF72F7CB6933F988" blockId="2.[189,1399,1434,1637]" pageId="2" pageNumber="3">
According to
<bibRefCitation id="EF89B6BDF710FC5FFEA3F7CB6830FA59" author="Gittenberger E." box="[364,597,1434,1460]" firstAuthor="Gittenberger" journalOrPublisher="Zoologische Mededelingen" pageId="2" pageNumber="3" pagination="287 - 306" part="43" refId="ref12709" refString="Gittenberger E. 1969. Beitrage zur Kenntnis der Pupillacea. I. Die Spelaeodiscinae. Zoologische Mededelingen 43 (22): 287 - 306." title="Beitrage zur Kenntnis der Pupillacea. I. Die Spelaeodiscinae" type="journal article" year="1969">Gittenberger (1969</bibRefCitation>
,
<bibRefCitation id="EF89B6BDF710FC5FFDA6F7CB68C0FA58" author="Gittenberger E." box="[617,677,1434,1461]" firstAuthor="Gittenberger" journalOrPublisher="Zoologische Mededelingen" pageId="2" pageNumber="3" pagination="263 - 277" part="48" refId="ref12736" refString="Gittenberger E. 1975. Beitrage zur Kenntnis der Pupillacea. V. Die Spelaeodiscinae, erster Nachtrag. Zoologische Mededelingen 48 (23): 263 - 277." title="Beitrage zur Kenntnis der Pupillacea. V. Die Spelaeodiscinae, erster Nachtrag" type="journal article" year="1975">1975</bibRefCitation>
) as well
<bibRefCitation id="EF89B6BDF710FC5FFCECF7CA6EA1FA58" author="Reischutz A. &amp; Reischutz P. L." box="[803,1220,1434,1461]" firstAuthor="as Reischutz" pageId="2" pageNumber="3" pagination="51 - 60" refId="ref13167" refString="Reischutz A. &amp; Reischutz P. L. 2009. Ein Beitrag zur Kenntnis der Molluskenfauna von Montenegro, nebst Beschreibung zweier neuer Arten der Gattung Virpazaria Gittenberger 1969. Nachrichtenblatt der Ersten Vorarlberger Malakologischen Gesellschaft 16: 51 - 60." type="journal article" year="2009">as Reischütz &amp; Reischütz (2009)</bibRefCitation>
,
<taxonomicName id="4C18B0CFF710FC5FFB18F7CA6F31FA59" authorityName="Gittenberger" authorityYear="1969" box="[1239,1364,1435,1460]" class="Gastropoda" family="Spelaeodiscidae" genus="Virpazaria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="3" phylum="Mollusca" rank="genus">
<emphasis id="B96C175EF710FC5FFB18F7CA6F31FA59" box="[1239,1364,1435,1460]" italics="true" pageId="2" pageNumber="3">Virpazaria</emphasis>
</taxonomicName>
is distinguished from the conchologically similar genus
<taxonomicName id="4C18B0CFF710FC5FFC94F7EF6E64FA35" authorityName="Brusina" authorityYear="1886" box="[859,1025,1470,1496]" class="Gastropoda" family="Spelaeodiscidae" genus="Spelaeodiscus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="3" phylum="Mollusca" rank="genus">
<emphasis id="B96C175EF710FC5FFC94F7EF6E64FA35" box="[859,1025,1470,1496]" italics="true" pageId="2" pageNumber="3">Spelaeodiscus</emphasis>
</taxonomicName>
by the continuous peristome, i.e., the presence of a parietal bulge or flap that connects the peristome insertions.
<taxonomicName id="4C18B0CFF710FC5FFB1FF7B068A5F9F3" authority="Pall-Gergely" authorityName="Pall-Gergely" authorityYear="2018" class="Gastropoda" family="Spelaeodiscidae" genus="Spelaeodiscus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="3" phylum="Mollusca" rank="species" species="virpazarioides">
<emphasis id="B96C175EF710FC5FFB1FF7B06B03F9F3" italics="true" pageId="2" pageNumber="3">Spelaeodiscus virpazarioides</emphasis>
<bibRefCitation id="EF89B6BDF710FC5FFEBEF45568A5F9F3" author="Pall-Gergely B. &amp; Deli T. &amp; Eross Z. P. &amp; Reischutz P. L. &amp; Reischutz A. &amp; Feher Z." box="[369,704,1540,1566]" firstAuthor="Pall-Gergely" journalOrPublisher="ZooKeys" pageId="2" pageNumber="3" pagination="13 - 48" part="769" refId="ref13102" refString="Pall-Gergely B., Deli T., Eross Z. P., Reischutz P. L., Reischutz A. &amp; Feher Z. 2018. Revision of the subterranean genus Spelaeodiscus Brusina, 1886 (Gastropoda, Pulmonata, Spelaeodiscidae). ZooKeys 769: 13 - 48. https: // doi. org / 10.3897 / zookeys. 769.25258" title="Revision of the subterranean genus Spelaeodiscus Brusina, 1886 (Gastropoda, Pulmonata, Spelaeodiscidae)" type="journal article" year="2018">Páll-Gergely &amp; Fehér, 2018</bibRefCitation>
</taxonomicName>
also possesses a callous parietal structure, but it is much smaller in proportion to the size of the whole aperture. Also, the narrower, crescent-shaped aperture is a good feature to tell
<taxonomicName id="4C18B0CFF710FC5FFE6CF41A6845F989" authorityName="Gittenberger" authorityYear="1969" box="[419,544,1611,1636]" class="Gastropoda" family="Spelaeodiscidae" genus="Virpazaria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="3" phylum="Mollusca" rank="genus">
<emphasis id="B96C175EF710FC5FFE6CF41A6845F989" box="[419,544,1611,1636]" italics="true" pageId="2" pageNumber="3">Virpazaria</emphasis>
</taxonomicName>
apart from related genera.
</paragraph>
<paragraph id="8BA7CB4CF710FC5FFF72F4DC6951F80E" blockId="2.[189,1399,1676,2019]" pageId="2" pageNumber="3">
Due to their hidden life, in most of the cases only empty shells are found. So far, there is one reported case when
<taxonomicName id="4C18B0CFF710FC5FFECCF4E16BE5F924" authorityName="Gittenberger" authorityYear="1969" box="[259,384,1712,1737]" class="Gastropoda" family="Spelaeodiscidae" genus="Virpazaria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="3" phylum="Mollusca" rank="genus">
<emphasis id="B96C175EF710FC5FFECCF4E16BE5F924" box="[259,384,1712,1737]" italics="true" pageId="2" pageNumber="3">Virpazaria</emphasis>
</taxonomicName>
, more specifically two individuals of
<taxonomicName id="4C18B0CFF710FC5FFCF8F4E16E64F924" authorityName="Gittenberger" authorityYear="1969" box="[823,1025,1711,1737]" class="Gastropoda" family="Spelaeodiscidae" genus="Virpazaria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="3" phylum="Mollusca" rank="species" species="deelemanorum">
<emphasis id="B96C175EF710FC5FFCF8F4E16E64F924" box="[823,1025,1711,1737]" italics="true" pageId="2" pageNumber="3">V. deelemanorum</emphasis>
</taxonomicName>
, were found alive (
<bibRefCitation id="EF89B6BDF710FC5FFB2EF4FE6A99F900" author="Gittenberger E." firstAuthor="Gittenberger" journalOrPublisher="Zoologische Mededelingen" pageId="2" pageNumber="3" pagination="263 - 277" part="48" refId="ref12736" refString="Gittenberger E. 1975. Beitrage zur Kenntnis der Pupillacea. V. Die Spelaeodiscinae, erster Nachtrag. Zoologische Mededelingen 48 (23): 263 - 277." title="Beitrage zur Kenntnis der Pupillacea. V. Die Spelaeodiscinae, erster Nachtrag" type="journal article" year="1975">Gittenberger 1975</bibRefCitation>
). Having no pigmented eyes, they are presumably blind, as is usual for subterranean animals. Comparing their genitals to those of
<taxonomicName id="4C18B0CFF710FC5FFD4BF4A7694FF8FD" authorityName="Brusina" authorityYear="1886" box="[644,810,1782,1808]" class="Gastropoda" family="Spelaeodiscidae" genus="Spelaeodiscus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="3" phylum="Mollusca" rank="genus">
<emphasis id="B96C175EF710FC5FFD4BF4A7694FF8FD" box="[644,810,1782,1808]" italics="true" pageId="2" pageNumber="3">Spelaeodiscus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C18B0CFF710FC5FFCF2F4A769C2F8FD" authorityName="Gittenberger" authorityYear="1969" box="[829,935,1782,1808]" class="Gastropoda" family="Spelaeodiscidae" genus="Klemmia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="3" phylum="Mollusca" rank="genus">
<emphasis id="B96C175EF710FC5FFCF2F4A769C2F8FD" box="[829,935,1782,1808]" italics="true" pageId="2" pageNumber="3">Klemmia</emphasis>
</taxonomicName>
and
<taxonomicName id="4C18B0CFF710FC5FFC27F4A76F4FF8FD" authority="Westerlund, 1889" authorityName="Westerlund" authorityYear="1889" box="[1000,1322,1782,1808]" class="Gastropoda" family="Spelaeodiscidae" genus="Aspasita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="3" phylum="Mollusca" rank="genus">
<emphasis id="B96C175EF710FC5FFC27F4A76E28F8FD" box="[1000,1101,1782,1808]" italics="true" pageId="2" pageNumber="3">Aspasita</emphasis>
Westerlund, 1889
</taxonomicName>
(
<bibRefCitation id="EF89B6BDF710FC5FFAF2F4A76A9EF8D9" author="Bole J." firstAuthor="Bole" journalOrPublisher="International Journal of Speleology" pageId="2" pageNumber="3" pagination="349 - 356" part="1" refId="ref12540" refString="Bole J. 1965. Die Vertreter der Gattung Spelaeodiscus Brusina 1886 (Gastropoda, Pulmonata) in Jugoslawien. International Journal of Speleology 1 (3): 349 - 356. https: // doi. org / 10.5038 / 1827 - 806 X. 1.3.9" title="Die Vertreter der Gattung Spelaeodiscus Brusina 1886 (Gastropoda, Pulmonata) in Jugoslawien" type="journal article" year="1965">Bole 1965</bibRefCitation>
;
<bibRefCitation id="EF89B6BDF710FC5FFEC7F5486BFEF8D9" author="Hudec V." box="[264,411,1817,1844]" firstAuthor="Hudec" journalOrPublisher="Archiv fur Molluskenkunde" pageId="2" pageNumber="3" pagination="157 - 165" part="94" refId="ref12837" refString="Hudec V. 1965. Neue Erkenntnisse uber den Geschlechtsapparat von Argna bielzi (Rossmassler) und Bemerkungen zur systematischen Stellung der Gattung Argna. Archiv fur Molluskenkunde 94: 157 - 165." title="Neue Erkenntnisse uber den Geschlechtsapparat von Argna bielzi (Rossmassler) und Bemerkungen zur systematischen Stellung der Gattung Argna" type="journal article" year="1965">Hudec 1965</bibRefCitation>
;
<bibRefCitation id="EF89B6BDF710FC5FFE68F54868E4F8D9" author="Gittenberger E." box="[423,641,1817,1844]" firstAuthor="Gittenberger" journalOrPublisher="Zoologische Mededelingen" pageId="2" pageNumber="3" pagination="263 - 277" part="48" refId="ref12736" refString="Gittenberger E. 1975. Beitrage zur Kenntnis der Pupillacea. V. Die Spelaeodiscinae, erster Nachtrag. Zoologische Mededelingen 48 (23): 263 - 277." title="Beitrage zur Kenntnis der Pupillacea. V. Die Spelaeodiscinae, erster Nachtrag" type="journal article" year="1975">Gittenberger 1975</bibRefCitation>
;
<bibRefCitation id="EF89B6BDF710FC5FFD41F54869E8F8DE" author="Subai P. &amp; Dedov I." box="[654,909,1817,1843]" firstAuthor="Subai" journalOrPublisher="Basteria" pageId="2" pageNumber="3" pagination="111 - 118" part="72" refId="ref13604" refString="Subai P. &amp; Dedov I. 2008. A review of the Bulgarian species of Aspasita Westerlund, 1889 (Gastropoda; Pulmonata; Spelaeodiscidae), with description of A. bulgarica spec. nov. Basteria 72: 111 - 118." title="A review of the Bulgarian species of Aspasita Westerlund, 1889 (Gastropoda; Pulmonata; Spelaeodiscidae), with description of A. bulgarica spec. nov" type="journal article" year="2008">Subai &amp; Dedov 2008</bibRefCitation>
), one might discover differences in terms of proportional lengths of certain reproductive organs, and there are certain structures that are called differently by different authors (e.g., the same short protrusion at the border of penis and epiphallus is referred to as flagellum by
<bibRefCitation id="EF89B6BDF710FC5FFE33F5D36883F870" author="Gittenberger E." box="[508,742,1922,1949]" firstAuthor="Gittenberger" journalOrPublisher="Zoologische Mededelingen" pageId="2" pageNumber="3" pagination="263 - 277" part="48" refId="ref12736" refString="Gittenberger E. 1975. Beitrage zur Kenntnis der Pupillacea. V. Die Spelaeodiscinae, erster Nachtrag. Zoologische Mededelingen 48 (23): 263 - 277." title="Beitrage zur Kenntnis der Pupillacea. V. Die Spelaeodiscinae, erster Nachtrag" type="journal article" year="1975">Gittenberger (1975)</bibRefCitation>
and penial caecum by
<bibRefCitation id="EF89B6BDF710FC5FFC3AF5D36EABF870" author="Schileyko A. A." box="[1013,1230,1922,1949]" firstAuthor="Schileyko" journalOrPublisher="Ruthenica, Supplement" pageId="2" pageNumber="3" pagination="1 - 127" part="2" refId="ref13455" refString="Schileyko A. A. 1998. Treatise on Recent Terrestrial Pulmonate Molluscs, Part 1. Achatinellidae, Amastridae, Orculidae, Strobilopsidae, Spelaeodiscidae, Valloniidae, Cochlicopidae, Pupillidae, Chondrinidae, Pyramidulidae. Ruthenica, Supplement 2 (1): 1 - 127." title="Treatise on Recent Terrestrial Pulmonate Molluscs, Part 1. Achatinellidae, Amastridae, Orculidae, Strobilopsidae, Spelaeodiscidae, Valloniidae, Cochlicopidae, Pupillidae, Chondrinidae, Pyramidulidae" type="journal article" year="1998">Schileyko (1998))</bibRefCitation>
. However, the overall arrangement of the genitalia is similar in these four genera, and no genus-specific differences could be found on the basis of the available literature.
</paragraph>
</subSubSection>
<subSubSection id="C30298C7F711FC5EFF72F35A6F12FA7D" pageId="3" pageNumber="4" type="key">
<paragraph id="8BA7CB4CF711FC5EFF72F35A6842FEC8" blockId="3.[189,551,267,293]" box="[189,551,267,293]" pageId="3" pageNumber="4">
<heading id="D0EF7C20F711FC5EFF72F35A6842FEC8" bold="true" box="[189,551,267,293]" fontSize="11" level="3" pageId="3" pageNumber="4" reason="3">
<emphasis id="B96C175EF711FC5EFF72F35A6842FEC8" bold="true" box="[189,551,267,293]" pageId="3" pageNumber="4">
A key to species of
<taxonomicName id="4C18B0CFF711FC5EFE67F35A6842FEC8" ID-CoL="7PKV4" authorityName="Gittenberger" authorityYear="1969" box="[424,551,267,293]" class="Gastropoda" family="Spelaeodiscidae" genus="Virpazaria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="4" phylum="Mollusca" rank="genus">
<emphasis id="B96C175EF711FC5EFE67F35A6842FEC8" bold="true" box="[424,551,267,293]" italics="true" pageId="3" pageNumber="4">Virpazaria</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<key id="FF899C7DF711FC5EFF72F36B6F12FA7D" pageId="3" pageNumber="4">
<keyStep id="30ECD3E9F711FC5EFF72F36B6F13FE50" pageId="3" pageNumber="4">
<paragraph id="8BA7CB4CF711FC5EFF72F36B6F12FE9B" blockId="3.[189,1399,313,1424]" pageId="3" pageNumber="4">
<keyLead id="30E96879F711FC5EFF72F36B6F12FE9B" pageId="3" pageNumber="4">
1. Two lamellae, one palatal and one basal, reach deeply (more than 1/10 of whorl) into the aperture ... ..........................................................................................
<taxonomicName id="4C18B0CFF711FC5EFCB1F30C6F12FE9B" ID-CoL="7f998fcf-4d8d-487e-9ce9-203fa65dca0b" authority="Gittenberger" authorityName="Gittenberger" authorityYear="1969" box="[894,1399,348,375]" class="Gastropoda" family="Spelaeodiscidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="ripkeni" subGenus="Aemiliella">
<emphasis id="B96C175EF711FC5EFCB1F30C6EEBFE9A" bold="true" box="[894,1166,348,375]" italics="true" pageId="3" pageNumber="4">V. (Aemiliella) ripkeni</emphasis>
<bibRefCitation id="EF89B6BDF711FC5EFB57F30D6F12FE9B" author="Gittenberger E." box="[1176,1399,348,374]" firstAuthor="Gittenberger" journalOrPublisher="Zoologische Mededelingen" pageId="3" pageNumber="4" pagination="287 - 306" part="43" refId="ref12709" refString="Gittenberger E. 1969. Beitrage zur Kenntnis der Pupillacea. I. Die Spelaeodiscinae. Zoologische Mededelingen 43 (22): 287 - 306." title="Beitrage zur Kenntnis der Pupillacea. I. Die Spelaeodiscinae" type="journal article" year="1969">Gittenberger, 1969</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BA7CB4CF711FC5EFF0EF3D16F13FE50" blockId="3.[189,1399,313,1424]" pageId="3" pageNumber="4">
<keyLead id="30E96879F711FC5EFF0EF3D16F13FE50" pageId="3" pageNumber="4"> No deeply reaching lamellae in the aperture, just knob(s) or on elongated thickening parallel with the peristome......................................................................................................................................2</keyLead>
</paragraph>
</keyStep>
<keyStep id="30ECD3E9F711FC5EFF72F38C6F12FDF6" pageId="3" pageNumber="4">
<paragraph id="8BA7CB4CF711FC5EFF72F38C6F12FE1A" blockId="3.[189,1399,313,1424]" box="[189,1399,477,503]" pageId="3" pageNumber="4">
<keyLead id="30E96879F711FC5EFF72F38C6F12FE1A" box="[189,1399,477,503]" pageId="3" pageNumber="4">2. In frontal view, the crescent-shaped hole of the aperture is well visible..........................................3</keyLead>
</paragraph>
<paragraph id="8BA7CB4CF711FC5EFF0EF0506F12FDF6" blockId="3.[189,1399,313,1424]" box="[193,1399,512,539]" pageId="3" pageNumber="4">
<keyLead id="30E96879F711FC5EFF0EF0506F12FDF6" box="[193,1399,512,539]" pageId="3" pageNumber="4"> In frontal view, the hole of the aperture is almost or entirely covered.............................................5</keyLead>
</paragraph>
</keyStep>
<keyStep id="30ECD3E9F711FC5EFF72F06B6F12FD9A" pageId="3" pageNumber="4">
<paragraph id="8BA7CB4CF711FC5EFF72F06B6F12FDB9" blockId="3.[189,1399,313,1424]" box="[189,1399,570,597]" pageId="3" pageNumber="4">
<keyLead id="30E96879F711FC5EFF72F06B6F12FDB9" box="[189,1399,570,597]" pageId="3" pageNumber="4">
3. One knob and a basal thickening on the peristome ...........
<taxonomicName id="4C18B0CFF711FC5EFC4DF06B6F72FDB9" ID-CoL="7TM29" authority="Feher &amp; Eross" authorityName="Fehér &amp; Deli &amp; Erőss &amp; Lika" authorityYear="2019" box="[898,1303,570,597]" class="Gastropoda" family="Spelaeodiscidae" genus="Virpazaria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="gittenbergeri" status="sp. nov." subGenus="Virpazaria">
<emphasis id="B96C175EF711FC5EFC4DF06B6E0EFDB8" bold="true" box="[898,1131,570,597]" italics="true" pageId="3" pageNumber="4">V. (V.) gittenbergeri</emphasis>
Fehér &amp; Erőss
</taxonomicName>
<taxonomicNameLabel id="A25FAA25F711FC5EFAD2F06B6F12FDB9" box="[1309,1399,570,596]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BA7CB4CF711FC5EFF0EF00F6F12FD9A" blockId="3.[189,1399,313,1424]" box="[193,1399,605,632]" pageId="3" pageNumber="4">
<keyLead id="30E96879F711FC5EFF0EF00F6F12FD9A" box="[193,1399,605,632]" pageId="3" pageNumber="4"> Two knobs on the peristome...............................................................................................................4</keyLead>
</paragraph>
</keyStep>
<keyStep id="30ECD3E9F711FC5EFF72F0C66F12FD39" pageId="3" pageNumber="4">
<paragraph id="8BA7CB4CF711FC5EFF72F0C66F12FD5F" blockId="3.[189,1399,313,1424]" box="[189,1399,663,690]" pageId="3" pageNumber="4">
<keyLead id="30E96879F711FC5EFF72F0C66F12FD5F" box="[189,1399,663,690]" pageId="3" pageNumber="4">
4. Shell width greater than
<quantity id="4CE066A9F711FC5EFDDFF0C66833FD5F" box="[528,598,663,690]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="4" unit="mm" value="5.0">5 mm</quantity>
...................................................
<taxonomicName id="4C18B0CFF711FC5EFC1BF0C66F76FD5C" ID-CoL="7TM2B" authority="Feher &amp; Deli" authorityName="Fehér &amp; Deli &amp; Erőss &amp; Lika" authorityYear="2019" box="[980,1299,663,689]" class="Gastropoda" family="Spelaeodiscidae" genus="Virpazaria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="pesici" status="sp. nov." subGenus="Virpazaria">
<emphasis id="B96C175EF711FC5EFC1BF0C66E0EFD5C" bold="true" box="[980,1131,663,689]" italics="true" pageId="3" pageNumber="4">V. (V.) pesici</emphasis>
Fehér &amp; Deli
</taxonomicName>
<taxonomicNameLabel id="A25FAA25F711FC5EFAD4F0C96F12FD5F" box="[1307,1399,664,690]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BA7CB4CF711FC5EFF0EF0EA6F12FD39" blockId="3.[189,1399,313,1424]" box="[193,1399,698,725]" pageId="3" pageNumber="4">
<keyLead id="30E96879F711FC5EFF0EF0EA6F12FD39" box="[193,1399,698,725]" pageId="3" pageNumber="4">
Shell width less than
<quantity id="4CE066A9F711FC5EFE2BF0EB684DFD38" box="[484,552,698,725]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="4" unit="mm" value="5.0">5 mm</quantity>
................................................
<taxonomicName id="4C18B0CFF711FC5EFC40F0EA6F12FD39" ID-CoL="bb15b6e9-660d-4ab0-a359-56a8ba8e87db" authority="Gittenberger" authorityName="Gittenberger" authorityYear="1969" box="[911,1399,698,725]" class="Gastropoda" family="Spelaeodiscidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="deelemanorum" subGenus="V.">
<emphasis id="B96C175EF711FC5EFC40F0EA6EF1FD38" bold="true" box="[911,1172,698,725]" italics="true" pageId="3" pageNumber="4">V. (V.) deelemanorum</emphasis>
<bibRefCitation id="EF89B6BDF711FC5EFB54F0EB6F12FD39" author="Gittenberger E." box="[1179,1399,698,724]" firstAuthor="Gittenberger" journalOrPublisher="Zoologische Mededelingen" pageId="3" pageNumber="4" pagination="287 - 306" part="43" refId="ref12709" refString="Gittenberger E. 1969. Beitrage zur Kenntnis der Pupillacea. I. Die Spelaeodiscinae. Zoologische Mededelingen 43 (22): 287 - 306." title="Beitrage zur Kenntnis der Pupillacea. I. Die Spelaeodiscinae" type="journal article" year="1969">Gittenberger, 1969</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30ECD3E9F711FC5EFF72F0A56F13FCDC" pageId="3" pageNumber="4">
<paragraph id="8BA7CB4CF711FC5EFF72F0A56F12FCE3" blockId="3.[189,1399,313,1424]" box="[189,1399,756,783]" pageId="3" pageNumber="4">
<keyLead id="30E96879F711FC5EFF72F0A56F12FCE3" box="[189,1399,756,783]" pageId="3" pageNumber="4">
5. Upper side of the shell is entirely flat in frontal view ........
<taxonomicName id="4C18B0CFF711FC5EFC6BF0A46F12FCE3" ID-CoL="7fdca4a2-f809-4644-86f2-3dbedace63f7" authority="Reischutz" authorityName="Reischutz" authorityYear="2009" box="[932,1399,756,783]" class="Gastropoda" family="Spelaeodiscidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="stojaspali" subGenus="V.">
<emphasis id="B96C175EF711FC5EFC6BF0A46E0FFCE2" bold="true" box="[932,1130,756,783]" italics="true" pageId="3" pageNumber="4">V. (V.) stojaspali</emphasis>
<bibRefCitation id="EF89B6BDF711FC5EFBBCF0A56F12FCE3" author="Reischutz A. &amp; Reischutz P. L." box="[1139,1399,756,783]" etAl="et al." firstAuthor="Reischutz" journalOrPublisher="Nachrichtenblatt der Ersten Vorarlberger Malakologischen Gesellschaft" pageId="3" pageNumber="4" pagination="51 - 60" part="16" refId="ref13167" refString="Reischutz A. &amp; Reischutz P. L. 2009. Ein Beitrag zur Kenntnis der Molluskenfauna von Montenegro, nebst Beschreibung zweier neuer Arten der Gattung Virpazaria Gittenberger 1969. Nachrichtenblatt der Ersten Vorarlberger Malakologischen Gesellschaft 16: 51 - 60." title="Ein Beitrag zur Kenntnis der Molluskenfauna von Montenegro, nebst Beschreibung zweier neuer Arten der Gattung Virpazaria Gittenberger 1969" type="journal article" year="2009">
Reischütz
<emphasis id="B96C175EF711FC5EFB20F0A46F49FCE3" box="[1263,1324,756,782]" italics="true" pageId="3" pageNumber="4">et al.</emphasis>
, 2009
</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BA7CB4CF711FC5EFF0EF1496F13FCDC" blockId="3.[189,1399,313,1424]" box="[193,1398,791,818]" pageId="3" pageNumber="4">
<keyLead id="30E96879F711FC5EFF0EF1496F13FCDC" box="[193,1398,791,818]" pageId="3" pageNumber="4"> Shell is conical in frontal view...........................................................................................................6</keyLead>
</paragraph>
</keyStep>
<keyStep id="30ECD3E9F711FC5EFF72F1006F13FC63" pageId="3" pageNumber="4">
<paragraph id="8BA7CB4CF711FC5EFF72F1006F13FC81" blockId="3.[189,1399,313,1424]" box="[189,1398,849,876]" pageId="3" pageNumber="4">
<keyLead id="30E96879F711FC5EFF72F1006F13FC81" box="[189,1398,849,876]" pageId="3" pageNumber="4">6. No knobs on the peristome................................................................................................................7</keyLead>
</paragraph>
<paragraph id="8BA7CB4CF711FC5EFF0EF1246F13FC63" blockId="3.[189,1399,313,1424]" box="[193,1398,884,911]" pageId="3" pageNumber="4">
<keyLead id="30E96879F711FC5EFF0EF1246F13FC63" box="[193,1398,884,911]" pageId="3" pageNumber="4"> At least one knob on the peristome..................................................................................................8</keyLead>
</paragraph>
</keyStep>
<keyStep id="30ECD3E9F711FC5EFF72F1FE6F12FC06" pageId="3" pageNumber="4">
<paragraph id="8BA7CB4CF711FC5EFF72F1FE6F12FC25" blockId="3.[189,1399,313,1424]" box="[189,1399,942,969]" pageId="3" pageNumber="4">
<keyLead id="30E96879F711FC5EFF72F1FE6F12FC25" box="[189,1399,942,969]" pageId="3" pageNumber="4">
7. Very narrow umbilicus .....
<taxonomicName id="4C18B0CFF711FC5EFDEDF1FE6F12FC25" ID-CoL="b6da985b-a8af-468c-a0a2-7fe270816198" authority="Reischutz et al. in Reischutz" authorityName="Reischutz et al. in Reischutz &amp; Reischutz" authorityYear="2009" box="[546,1399,942,969]" class="Gastropoda" family="Spelaeodiscidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="aspectulabeatidis" subGenus="V.">
<emphasis id="B96C175EF711FC5EFDEDF1FE6924FC24" bold="true" box="[546,833,942,969]" italics="true" pageId="3" pageNumber="4">V. (V.) aspectulabeatidis</emphasis>
Reischütz
<emphasis id="B96C175EF711FC5EFC0DF1FE6998FC25" box="[962,1021,942,968]" italics="true" pageId="3" pageNumber="4">et al.</emphasis>
in
<bibRefCitation id="EF89B6BDF711FC5EFBEEF1FF6F12FC25" author="Reischutz A. &amp; Reischutz P. L." box="[1057,1399,942,968]" firstAuthor="Reischutz" journalOrPublisher="Nachrichtenblatt der Ersten Vorarlberger Malakologischen Gesellschaft" pageId="3" pageNumber="4" pagination="51 - 60" part="16" refId="ref13167" refString="Reischutz A. &amp; Reischutz P. L. 2009. Ein Beitrag zur Kenntnis der Molluskenfauna von Montenegro, nebst Beschreibung zweier neuer Arten der Gattung Virpazaria Gittenberger 1969. Nachrichtenblatt der Ersten Vorarlberger Malakologischen Gesellschaft 16: 51 - 60." title="Ein Beitrag zur Kenntnis der Molluskenfauna von Montenegro, nebst Beschreibung zweier neuer Arten der Gattung Virpazaria Gittenberger 1969" type="journal article" year="2009">Reischütz &amp; Reischütz, 2009</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BA7CB4CF711FC5EFF0EF1836F12FC06" blockId="3.[189,1399,313,1424]" box="[193,1399,977,1004]" pageId="3" pageNumber="4">
<keyLead id="30E96879F711FC5EFF0EF1836F12FC06" box="[193,1399,977,1004]" pageId="3" pageNumber="4">
Wide, perspectivical umbilicus .......................................................
<taxonomicName id="4C18B0CFF711FC5EFC3BF1836F12FC06" ID-CoL="8576217c-0585-4821-82fc-5dbbfa10fe7e" authority="Gittenberger" authorityName="Gittenberger" authorityYear="1969" box="[1012,1399,977,1004]" class="Gastropoda" family="Spelaeodiscidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="pageti" subGenus="V.">
<emphasis id="B96C175EF711FC5EFC3BF1836EF5FC06" bold="true" box="[1012,1168,977,1004]" italics="true" pageId="3" pageNumber="4">V. (V.) pageti</emphasis>
<bibRefCitation id="EF89B6BDF711FC5EFB56F1806F12FC06" author="Gittenberger E." box="[1177,1399,977,1003]" firstAuthor="Gittenberger" journalOrPublisher="Zoologische Mededelingen" pageId="3" pageNumber="4" pagination="287 - 306" part="43" refId="ref12709" refString="Gittenberger E. 1969. Beitrage zur Kenntnis der Pupillacea. I. Die Spelaeodiscinae. Zoologische Mededelingen 43 (22): 287 - 306." title="Beitrage zur Kenntnis der Pupillacea. I. Die Spelaeodiscinae" type="journal article" year="1969">Gittenberger, 1969</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30ECD3E9F711FC5EFF72F65A6F13FBA4" pageId="3" pageNumber="4">
<paragraph id="8BA7CB4CF711FC5EFF72F65A6F12FBC8" blockId="3.[189,1399,313,1424]" box="[189,1399,1035,1062]" pageId="3" pageNumber="4">
<keyLead id="30E96879F711FC5EFF72F65A6F12FBC8" box="[189,1399,1035,1062]" pageId="3" pageNumber="4">8. Less than three knobs on the peristome............................................................................................9</keyLead>
</paragraph>
<paragraph id="8BA7CB4CF711FC5EFF0EF67E6F13FBA4" blockId="3.[189,1399,313,1424]" box="[193,1398,1070,1097]" pageId="3" pageNumber="4">
<keyLead id="30E96879F711FC5EFF0EF67E6F13FBA4" box="[193,1398,1070,1097]" pageId="3" pageNumber="4"> Three knobs on the peristome..........................................................................................................10</keyLead>
</paragraph>
</keyStep>
<keyStep id="30ECD3E9F711FC5EFF72F6386F12FB00" pageId="3" pageNumber="4">
<paragraph id="8BA7CB4CF711FC5EFF72F6386F12FB48" blockId="3.[189,1399,313,1424]" pageId="3" pageNumber="4">
<keyLead id="30E96879F711FC5EFF72F6386F12FB48" pageId="3" pageNumber="4">
9. An elongated basal thickening and a detached palatal knob on the peristome ............................... ..................................................................................................
<taxonomicName id="4C18B0CFF711FC5EFC76F6DD6F12FB48" ID-CoL="9f62cb2c-3bdd-434e-b467-fd6e27ad1c07" authority="Gittenberger" authorityName="Gittenberger" authorityYear="1969" box="[953,1399,1163,1190]" class="Gastropoda" family="Spelaeodiscidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="backhuysi" subGenus="V.">
<emphasis id="B96C175EF711FC5EFC76F6DD6EEEFB4B" bold="true" box="[953,1163,1163,1190]" italics="true" pageId="3" pageNumber="4">V. (V.) backhuysi</emphasis>
<bibRefCitation id="EF89B6BDF711FC5EFB58F6DD6F12FB48" author="Gittenberger E." box="[1175,1399,1163,1190]" firstAuthor="Gittenberger" journalOrPublisher="Zoologische Mededelingen" pageId="3" pageNumber="4" pagination="287 - 306" part="43" refId="ref12709" refString="Gittenberger E. 1969. Beitrage zur Kenntnis der Pupillacea. I. Die Spelaeodiscinae. Zoologische Mededelingen 43 (22): 287 - 306." title="Beitrage zur Kenntnis der Pupillacea. I. Die Spelaeodiscinae" type="journal article" year="1969">Gittenberger, 1969</bibRefCitation>
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Palatal knob is fused to the basal thickening of the peristome ......................................................... ......................................................................................
<taxonomicName id="4C18B0CFF711FC5EFCAEF6836F6BFB01" ID-CoL="7TM29" authority="Feher &amp; Eross" authorityName="Fehér &amp; Deli &amp; Erőss &amp; Lika" authorityYear="2019" box="[865,1294,1234,1261]" class="Gastropoda" family="Spelaeodiscidae" genus="Virpazaria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="gittenbergeri" status="sp. nov." subGenus="Virpazaria">
<emphasis id="B96C175EF711FC5EFCAEF6836E36FB00" bold="true" box="[865,1107,1234,1261]" italics="true" pageId="3" pageNumber="4">V. (V.) gittenbergeri</emphasis>
Fehér &amp; Erőss
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<taxonomicNameLabel id="A25FAA25F711FC5EFAD6F6826F12FB00" box="[1305,1399,1235,1261]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="8BA7CB4CF711FC5EFF72F75D6F12FAA4" blockId="3.[189,1399,313,1424]" pageId="3" pageNumber="4">
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10. In bottom-view the umbilicus seems wider than ¼ of the shell width, in frontal view the shell is flat conical (H:W &lt;2:3) ........................................
<taxonomicName id="4C18B0CFF711FC5EFC74F7616F12FAA4" ID-CoL="b8a6163b-bea2-440f-91ba-aaa51fd94d47" authority="Gittenberger" authorityName="Gittenberger" authorityYear="1969" box="[955,1399,1327,1354]" class="Gastropoda" family="Spelaeodiscidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="adrianae" subGenus="V.">
<emphasis id="B96C175EF711FC5EFC74F7616EE3FAA7" bold="true" box="[955,1158,1327,1354]" italics="true" pageId="3" pageNumber="4">V. (V.) adrianae</emphasis>
<bibRefCitation id="EF89B6BDF711FC5EFB5BF77E6F12FAA4" author="Gittenberger E." box="[1172,1399,1327,1353]" firstAuthor="Gittenberger" journalOrPublisher="Zoologische Mededelingen" pageId="3" pageNumber="4" pagination="287 - 306" part="43" refId="ref12709" refString="Gittenberger E. 1969. Beitrage zur Kenntnis der Pupillacea. I. Die Spelaeodiscinae. Zoologische Mededelingen 43 (22): 287 - 306." title="Beitrage zur Kenntnis der Pupillacea. I. Die Spelaeodiscinae" type="journal article" year="1969">Gittenberger, 1969</bibRefCitation>
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<keyLead id="30E96879F711FC5EFF0EF7026F12FA7D" pageId="3" pageNumber="4">
In bottom-view the umbilicus seems narrower than ¼ of the shell width, in frontal view the shell is conical (H:W ≈ 2:3) ...................................
<taxonomicName id="4C18B0CFF711FC5EFC9CF7276F12FA7D" ID-CoL="a4a493ce-07b1-4981-ac82-ff6fa482b5cc" authority="Reischutz" authorityName="Reischutz" authorityYear="2012" box="[851,1399,1397,1424]" class="Gastropoda" family="Spelaeodiscidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="nicoleae" subGenus="V.">
<emphasis id="B96C175EF711FC5EFC9CF7276E6BFA7D" bold="true" box="[851,1038,1397,1424]" italics="true" pageId="3" pageNumber="4">V. (V.) nicoleae</emphasis>
<bibRefCitation id="EF89B6BDF711FC5EFBD7F7246F12FA7D" author="Reischutz A. &amp; Reischutz P. L." box="[1048,1399,1397,1424]" firstAuthor="Reischutz" journalOrPublisher="Nachrichtenblatt der Ersten Vorarlberger Malakologischen Gesellschaft" pageId="3" pageNumber="4" pagination="17 - 18" part="19" refId="ref13210" refString="Reischutz A. &amp; Reischutz P. L. 2012. Virpazaria nicoleae nov. spec. (Spelaeodiscidae: Pulmonata: Gastropoda) aus Montenegro. Nachrichtenblatt der Ersten Vorarlberger Malakologischen Gesellschaft 19: 17 - 18." title="Virpazaria nicoleae nov. spec. (Spelaeodiscidae: Pulmonata: Gastropoda) aus Montenegro" type="journal article" year="2012">
Reischütz &amp; Reischütz,
<treatmentHeading id="AD54A448F711FC5EFAF3F7276F12FA7D" box="[1340,1399,1398,1424]" pageId="3" pageNumber="4">2012</treatmentHeading>
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