376 lines
41 KiB
XML
376 lines
41 KiB
XML
<document ID-DOI="10.11646/zootaxa.5278.2.4" ID-ISSN="1175-5326" ID-Zenodo-Dep="7906172" ID-ZooBank="A5A18196-D519-4D43-BF03-5C6D2DEAC842" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1683529272315" checkinUser="plazi" docAuthor="Archibald, S. Bruce, Ware, Jessica L., Rasmussen, Jan A., Sylvestersen, René L., Olsen, Kent & Simonsen, Thomas J." docDate="2023" docId="03A5BA5CFFB5FFB7FF40E9B3E8234ECC" docLanguage="en" docName="zootaxa.5278.2.4.pdf" docOrigin="Zootaxa 5278 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5278.2.4" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Furagrion ansorgei Archibald & Ware & Rasmussen & Sylvestersen & Olsen & Simonsen 2023, comb. nov." docType="treatment" docVersion="3" lastPageNumber="314" masterDocId="FF9CC224FFA2FFAEFFD7EA60E938481D" masterDocTitle="The damselfly genus Furagrion Petrulevičius et al. (Odonata, Zygoptera) from the early Eocene Fur Formation of Denmark and the dysagrionoid grade" masterLastPageNumber="317" masterPageNumber="289" pageNumber="312" updateTime="1683553539095" updateUser="juliana" zenodo-license-document="CLOSED">
|
||
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo>
|
||
<mods:title>The damselfly genus Furagrion Petrulevičius et al. (Odonata, Zygoptera) from the early Eocene Fur Formation of Denmark and the dysagrionoid grade</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Archibald, S. Bruce</mods:namePart>
|
||
<mods:nameIdentifier type="ORCID">0000-0002-4397-2497</mods:nameIdentifier>
|
||
<mods:affiliation>Beaty Biodiversity Museum, University of British Columbia, Vancouver, British Columbia, Canada. https: // orcid. org / 0000 - 0002 - 4397 - 2497 & Museum of Comparative Zoology, 26 Oxford Street, Cambridge, Massachusetts, 02138, United States of America</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Ware, Jessica L.</mods:namePart>
|
||
<mods:affiliation>Division of Invertebrate Zoology, American Museum of Natural History, New York, New York, 10024, United States of America.</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Rasmussen, Jan A.</mods:namePart>
|
||
<mods:nameIdentifier type="ORCID">0000-0003-0520-9148</mods:nameIdentifier>
|
||
<mods:affiliation>Museum Mors, Skarrehagevej 8, DK- 7950 NykØbing Mors, Denmark. https: // orcid. org / 0000 - 0003 - 0520 - 9148 & Natural History Museum of Denmark, Øster Voldgade 5 - 7, DK- 1350 Copenhagen K, Denmark. jan. rasmussen @ museummors. dk</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Sylvestersen, René L.</mods:namePart>
|
||
<mods:nameIdentifier type="ORCID">0000-0002-1946-4716</mods:nameIdentifier>
|
||
<mods:affiliation>Fur Museum, Nederby 28, DK- 7884 Fur, Denmark. RLSY @ museumsalling. dk; https: // orcid. org / 0000 - 0002 - 1946 - 4716</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Olsen, Kent</mods:namePart>
|
||
<mods:affiliation>Natural History Museum Aarhus, Wilhelm Meyers Allé 10, Aarhus, DK- 8000 Aarhus C, Denmark.</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Simonsen, Thomas J.</mods:namePart>
|
||
<mods:nameIdentifier type="ORCID">0000-0001-9857-9564</mods:nameIdentifier>
|
||
<mods:affiliation>Natural History Museum Aarhus, Wilhelm Meyers Allé 10, Aarhus, DK- 8000 Aarhus C, Denmark. https: // orcid. org / 0000 - 0001 - 9857 - 9564</mods:affiliation>
|
||
</mods:name>
|
||
<mods:typeOfResource>text</mods:typeOfResource>
|
||
<mods:relatedItem type="host">
|
||
<mods:titleInfo>
|
||
<mods:title>Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part>
|
||
<mods:date>2023</mods:date>
|
||
<mods:detail type="pubDate">
|
||
<mods:number>2023-05-05</mods:number>
|
||
</mods:detail>
|
||
<mods:detail type="volume">
|
||
<mods:number>5278</mods:number>
|
||
</mods:detail>
|
||
<mods:detail type="issue">
|
||
<mods:number>2</mods:number>
|
||
</mods:detail>
|
||
<mods:extent unit="page">
|
||
<mods:start>289</mods:start>
|
||
<mods:end>317</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location>
|
||
<mods:url>http://dx.doi.org/10.11646/zootaxa.5278.2.4</mods:url>
|
||
</mods:location>
|
||
<mods:classification>journal article</mods:classification>
|
||
<mods:identifier type="DOI">10.11646/zootaxa.5278.2.4</mods:identifier>
|
||
<mods:identifier type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier type="Zenodo-Dep">7906172</mods:identifier>
|
||
<mods:identifier type="ZooBank">A5A18196-D519-4D43-BF03-5C6D2DEAC842</mods:identifier>
|
||
</mods:mods>
|
||
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7906168" ID-Zenodo-Dep="7906168" LSID="urn:lsid:plazi:treatment:03A5BA5CFFB5FFB7FF40E9B3E8234ECC" httpUri="http://treatment.plazi.org/id/03A5BA5CFFB5FFB7FF40E9B3E8234ECC" lastPageId="25" lastPageNumber="314" pageId="23" pageNumber="312">
|
||
<subSubSection box="[151,710,978,1005]" pageId="23" pageNumber="312" type="nomenclature">
|
||
<paragraph blockId="23.[151,710,978,1041]" box="[151,710,978,1005]" pageId="23" pageNumber="312">
|
||
<heading bold="true" box="[151,710,978,1005]" fontSize="11" level="1" pageId="23" pageNumber="312" reason="1">
|
||
<emphasis bold="true" box="[151,710,978,1005]" pageId="23" pageNumber="312">
|
||
<taxonomicName authority="(Zessin, 2011)" authorityName="Archibald & Ware & Rasmussen & Sylvestersen & Olsen & Simonsen" authorityYear="2023" baseAuthorityName="Zessin" baseAuthorityYear="2011" box="[151,569,978,1005]" class="Insecta" family="Megapodagrionidae" genus="Furagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="23" pageNumber="312" phylum="Arthropoda" rank="species" species="ansorgei" status="comb. nov.">
|
||
<emphasis bold="true" box="[151,390,979,1005]" italics="true" pageId="23" pageNumber="312">Furagrion ansorgei</emphasis>
|
||
(
|
||
<bibRefCitation author="Zessin, W." box="[405,561,978,1004]" pageId="23" pageNumber="312" pagination="64 - 73" refId="ref14309" refString="Zessin, W. (2011) Neue Insekten aus dem Moler (Palaozan / Eozan) von Danemark Teil 1 (Odonata: Epallagidae, Megapodagrioniidae), Virgo, Mitteilungsblatt des Entomologischen Vereins Mecklenburg, 14, 64 - 73." type="journal article" year="2011">Zessin, 2011</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[576,710,979,1005]" pageId="23" pageNumber="312" rank="species">comb. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection box="[151,284,1015,1041]" pageId="23" pageNumber="312" type="description">
|
||
<paragraph blockId="23.[151,710,978,1041]" box="[151,284,1015,1041]" pageId="23" pageNumber="312">
|
||
<figureCitation box="[151,284,1015,1041]" captionStart-0="FIGURE 22" captionStart-1="FIGURE 23" captionStartId-0="24.[151,250,1877,1901]" captionStartId-1="25.[151,250,1216,1240]" captionTargetBox-0="[151,1431,849,1843]" captionTargetBox-1="[156,1434,422,1182]" captionTargetId-0="figure-365@24.[151,1436,849,1854]" captionTargetId-1="figure-150@25.[151,1436,417,1193]" captionTargetPageId-0="24" captionTargetPageId-1="25" captionText-0="FIGURE 22. MGUH 34113, a forewing: A, photograph of the part; B, photograph of the counterpart; C, drawing from both A and B. Note infuscation very light on the part as preserved, darker on the counterpart (see text). All to scale, 5 mm." captionText-1="FIGURE 23. A, Furagrion ansorgei MOA 770 redrawn from a higher resolution copy of Zessin’s (2011) figure 3 photograph provided by J. Ansorge; shape compared with that of the proposed neotype (blue) B, forewing; C, hind wing. A to scale, 5 mm; B, C wings to no scales, the sizes of proposed neotype wings are adjusted to match arculus to apex and anterior wing margin of MOA 770 to compare shape." figureDoi-0="http://doi.org/10.5281/zenodo.7906220" figureDoi-1="http://doi.org/10.5281/zenodo.7906222" httpUri-0="https://zenodo.org/record/7906220/files/figure.png" httpUri-1="https://zenodo.org/record/7906222/files/figure.png" pageId="23" pageNumber="312">Figs. 22–23</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="23" pageNumber="312" type="diagnosis">
|
||
<paragraph blockId="23.[151,1437,1086,1725]" pageId="23" pageNumber="312">
|
||
<emphasis bold="true" box="[151,391,1086,1113]" pageId="23" pageNumber="312">Emended diagnosis.</emphasis>
|
||
The wing of
|
||
<taxonomicName authorityName="Archibald & Ware & Rasmussen & Sylvestersen & Olsen & Simonsen" authorityYear="2023" baseAuthorityName="Zessin" baseAuthorityYear="2011" box="[546,768,1087,1112]" class="Insecta" family="Megapodagrionidae" genus="Furagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="23" pageNumber="312" phylum="Arthropoda" rank="species" species="ansorgei">
|
||
<emphasis box="[546,768,1087,1112]" italics="true" pageId="23" pageNumber="312">Furagrion ansorgei</emphasis>
|
||
</taxonomicName>
|
||
may be distinguished from those of
|
||
<taxonomicName baseAuthorityName="Henriksen" baseAuthorityYear="1922" box="[1187,1339,1087,1113]" class="Insecta" family="Megapodagrionidae" genus="Furagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="23" pageNumber="312" phylum="Arthropoda" rank="species" species="jutlandicus">
|
||
<emphasis box="[1187,1339,1087,1113]" italics="true" pageId="23" pageNumber="312">F. jutlandicus</emphasis>
|
||
</taxonomicName>
|
||
as in its diagnosis, above.
|
||
</paragraph>
|
||
<paragraph blockId="23.[151,1437,1086,1725]" pageId="23" pageNumber="312">
|
||
<emphasis bold="true" box="[199,462,1158,1185]" pageId="23" pageNumber="312">Emended description.</emphasis>
|
||
<typeStatus box="[469,578,1159,1185]" pageId="23" pageNumber="312">Holotype</typeStatus>
|
||
: see genus description and its distinctive character states in the diagnosis of
|
||
<taxonomicName baseAuthorityName="Henriksen" baseAuthorityYear="1922" box="[151,303,1195,1221]" class="Insecta" family="Megapodagrionidae" genus="Furagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="23" pageNumber="312" phylum="Arthropoda" rank="species" species="jutlandicus">
|
||
<emphasis box="[151,303,1195,1221]" italics="true" pageId="23" pageNumber="312">F. jutlandicus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="23" pageNumber="312" type="materials_examined">
|
||
<paragraph blockId="23.[151,1437,1086,1725]" pageId="23" pageNumber="312">
|
||
<emphasis bold="true" box="[199,376,1230,1257]" pageId="23" pageNumber="312">Type material.</emphasis>
|
||
<materialsCitation box="[387,790,1231,1257]" collectorName="Jorg Ansorge" country="Denmark" location="Denmark" pageId="23" pageNumber="312" specimenCount="1" typeStatus="holotype">
|
||
<typeStatus box="[387,491,1231,1257]" pageId="23" pageNumber="312">Holotype</typeStatus>
|
||
(
|
||
<figureCitation box="[510,594,1231,1257]" captionStart="FIGURE 23" captionStartId="25.[151,250,1216,1240]" captionTargetBox="[156,1434,422,1182]" captionTargetId="figure-150@25.[151,1436,417,1193]" captionTargetPageId="25" captionText="FIGURE 23. A, Furagrion ansorgei MOA 770 redrawn from a higher resolution copy of Zessin’s (2011) figure 3 photograph provided by J. Ansorge; shape compared with that of the proposed neotype (blue) B, forewing; C, hind wing. A to scale, 5 mm; B, C wings to no scales, the sizes of proposed neotype wings are adjusted to match arculus to apex and anterior wing margin of MOA 770 to compare shape." figureDoi="http://doi.org/10.5281/zenodo.7906222" httpUri="https://zenodo.org/record/7906222/files/figure.png" pageId="23" pageNumber="312">Fig. 23</figureCitation>
|
||
) MOA 770/1, 2:
|
||
</materialsCitation>
|
||
likely a forewing by its slender shape (
|
||
<figureCitation box="[1258,1363,1231,1257]" captionStart="FIGURE 23" captionStartId="25.[151,250,1216,1240]" captionTargetBox="[156,1434,422,1182]" captionTargetId="figure-150@25.[151,1436,417,1193]" captionTargetPageId="25" captionText="FIGURE 23. A, Furagrion ansorgei MOA 770 redrawn from a higher resolution copy of Zessin’s (2011) figure 3 photograph provided by J. Ansorge; shape compared with that of the proposed neotype (blue) B, forewing; C, hind wing. A to scale, 5 mm; B, C wings to no scales, the sizes of proposed neotype wings are adjusted to match arculus to apex and anterior wing margin of MOA 770 to compare shape." figureDoi="http://doi.org/10.5281/zenodo.7906222" httpUri="https://zenodo.org/record/7906222/files/figure.png" pageId="23" pageNumber="312">Fig. 23B</figureCitation>
|
||
), coll.
|
||
<collectorName box="[151,302,1267,1293]" pageId="23" pageNumber="312">Jörg Ansorge</collectorName>
|
||
, northern coast of Mors, Skaerbaek, Fur Formation, in a calcareous concretion, in the Natural History Museum, Natural Research Society Mecklenburg, Natureum at Ludwigslust Castle (part) and Natural History Museum of
|
||
<collectingCountry box="[287,394,1339,1365]" name="Denmark" pageId="23" pageNumber="312">Denmark</collectingCountry>
|
||
, Copenhagen,
|
||
<collectingCountry box="[559,663,1339,1365]" name="Denmark" pageId="23" pageNumber="312">Denmark</collectingCountry>
|
||
(counterpart). We were not able to examine the fossil as the part and counterpart could not be located at the institutions listed. Our discussion is based on
|
||
<bibRefCitation author="Zessin, W." box="[1087,1259,1375,1401]" pageId="23" pageNumber="312" pagination="64 - 73" refId="ref14309" refString="Zessin, W. (2011) Neue Insekten aus dem Moler (Palaozan / Eozan) von Danemark Teil 1 (Odonata: Epallagidae, Megapodagrioniidae), Virgo, Mitteilungsblatt des Entomologischen Vereins Mecklenburg, 14, 64 - 73." type="journal article" year="2011">Zessin’s (2011)</bibRefCitation>
|
||
description and illustrations, and a higher resolution copy of his figure 3 provided by Jörg Ansorge.
|
||
</paragraph>
|
||
<paragraph blockId="23.[151,1437,1086,1725]" pageId="23" pageNumber="312">
|
||
<emphasis bold="true" box="[199,384,1446,1472]" pageId="23" pageNumber="312">Other material.</emphasis>
|
||
<collectionCode box="[389,475,1447,1473]" country="Denmark" httpUri="http://grbio.org/cool/c1ae-v64p" name="Museum Geologicum Universitatis Hafniensis" pageId="23" pageNumber="312">MGUH</collectionCode>
|
||
34113 (
|
||
<figureCitation box="[563,643,1447,1473]" captionStart="FIGURE 22" captionStartId="24.[151,250,1877,1901]" captionTargetBox="[151,1431,849,1843]" captionTargetId="figure-365@24.[151,1436,849,1854]" captionTargetPageId="24" captionText="FIGURE 22. MGUH 34113, a forewing: A, photograph of the part; B, photograph of the counterpart; C, drawing from both A and B. Note infuscation very light on the part as preserved, darker on the counterpart (see text). All to scale, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7906220" httpUri="https://zenodo.org/record/7906220/files/figure.png" pageId="23" pageNumber="312">Fig. 22</figureCitation>
|
||
) tentatively belongs to the species: a complete forewing in a concretion, coll. Thomas Klode on Fur Island, Knudeklint Member, possibly between ash layers -11 and -13, Fur Formation, 1968, in the Natural History Museum of
|
||
<collectingCountry box="[608,715,1519,1545]" name="Denmark" pageId="23" pageNumber="312">Denmark</collectingCountry>
|
||
, University of Copenhagen.
|
||
</paragraph>
|
||
<paragraph blockId="23.[151,1437,1086,1725]" pageId="23" pageNumber="312">
|
||
<emphasis bold="true" box="[199,391,1555,1581]" pageId="23" pageNumber="312">Range and age.</emphasis>
|
||
<location LSID="urn:lsid:plazi:treatment:03A5BA5CFFB5FFB7FF40E9B3E8234ECC:8ED35D91FFB5FFB9FE44EC73EA344E30" box="[403,780,1555,1581]" country="Denmark" county="Range" municipality="Jutland" name="Earliest Ypresian Fur Formation" pageId="23" pageNumber="312">Earliest Ypresian Fur Formation</location>
|
||
,
|
||
<collectingMunicipality box="[795,879,1555,1581]" pageId="23" pageNumber="312">Jutland</collectingMunicipality>
|
||
,
|
||
<collectingCountry box="[895,1003,1555,1581]" name="Denmark" pageId="23" pageNumber="312">Denmark</collectingCountry>
|
||
.
|
||
<collectingCounty box="[1018,1094,1555,1581]" pageId="23" pageNumber="312">Range</collectingCounty>
|
||
: see
|
||
<typeStatus box="[1156,1204,1555,1581]" pageId="23" pageNumber="312">type</typeStatus>
|
||
and other material, above.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="25" lastPageNumber="314" pageId="23" pageNumber="312" type="discussion">
|
||
<paragraph blockId="23.[151,1437,1086,1725]" pageId="23" pageNumber="312">
|
||
<emphasis bold="true" box="[199,315,1627,1653]" pageId="23" pageNumber="312">Remarks.</emphasis>
|
||
With the ability to examine a larger sample of
|
||
<taxonomicName box="[862,979,1627,1652]" class="Insecta" family="Megapodagrionidae" genus="Furagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="23" pageNumber="312" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[862,979,1627,1652]" italics="true" pageId="23" pageNumber="312">Furagrion</emphasis>
|
||
</taxonomicName>
|
||
specimens than was available to
|
||
<bibRefCitation author="Zessin, W." pageId="23" pageNumber="312" pagination="64 - 73" refId="ref14309" refString="Zessin, W. (2011) Neue Insekten aus dem Moler (Palaozan / Eozan) von Danemark Teil 1 (Odonata: Epallagidae, Megapodagrioniidae), Virgo, Mitteilungsblatt des Entomologischen Vereins Mecklenburg, 14, 64 - 73." type="journal article" year="2011">Zessin (2011)</bibRefCitation>
|
||
, we find that the character states of
|
||
<taxonomicName authorityName="Zessin. As" authorityYear="2011" box="[638,772,1663,1688]" class="Insecta" family="Megapodagrionidae" genus="Morsagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="23" pageNumber="312" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[638,772,1663,1688]" italics="true" pageId="23" pageNumber="312">Morsagrion</emphasis>
|
||
</taxonomicName>
|
||
that he proposed would separate it from
|
||
<taxonomicName box="[1240,1357,1663,1688]" class="Insecta" family="Megapodagrionidae" genus="Furagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="23" pageNumber="312" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[1240,1357,1663,1688]" italics="true" pageId="23" pageNumber="312">Furagrion</emphasis>
|
||
</taxonomicName>
|
||
do not do so. These are:
|
||
</paragraph>
|
||
<paragraph blockId="23.[151,1437,1764,2043]" box="[151,1217,1764,1791]" pageId="23" pageNumber="312">1- The RA-RP1 space is two cells wide from about the middle of the level of the pterostigma.</paragraph>
|
||
<paragraph blockId="23.[151,1437,1764,2043]" box="[151,1394,1800,1827]" pageId="23" pageNumber="312">- We find this difficult to discern from his fig. 3 photograph and we are not confident that this is established.</paragraph>
|
||
<paragraph blockId="23.[151,1437,1764,2043]" box="[151,521,1836,1862]" pageId="23" pageNumber="312">2- The form of the pterostigma.</paragraph>
|
||
<paragraph blockId="23.[151,1437,1764,2043]" box="[151,681,1872,1899]" pageId="23" pageNumber="312">- We find that it does not significantly differ.</paragraph>
|
||
<paragraph blockId="23.[151,1437,1764,2043]" pageId="23" pageNumber="312">
|
||
3- The relationship between Zessin’s “premedial cell” (“prmc”) and the cells below it in the MA-MP space. The premedial cell is the space between RP to slightly past the base of RP3-4 and MA without crossveins from the arculus to the first crossvein between RP-3-4 and MA. He excludes the quadrangle and counts four cells in the MA-MP space subtending this in
|
||
<taxonomicName authorityName="Zessin. As" authorityYear="2011" box="[574,708,2017,2042]" class="Insecta" family="Megapodagrionidae" genus="Morsagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="23" pageNumber="312" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[574,708,2017,2042]" italics="true" pageId="23" pageNumber="312">Morsagrion</emphasis>
|
||
</taxonomicName>
|
||
and two cells in this space in
|
||
<taxonomicName box="[1044,1161,2017,2042]" class="Insecta" family="Megapodagrionidae" genus="Furagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="23" pageNumber="312" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[1044,1161,2017,2042]" italics="true" pageId="23" pageNumber="312">Furagrion</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph blockId="24.[151,1437,151,825]" pageId="24" pageNumber="313">
|
||
- By our count, this is about three and a half cells long in
|
||
<taxonomicName authorityName="Archibald & Ware & Rasmussen & Sylvestersen & Olsen & Simonsen" authorityYear="2023" baseAuthorityName="Zessin" baseAuthorityYear="2011" box="[850,986,151,176]" class="Insecta" family="Megapodagrionidae" genus="Furagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="313" phylum="Arthropoda" rank="species" species="ansorgei">
|
||
<emphasis box="[850,986,151,176]" italics="true" pageId="24" pageNumber="313">M. ansorgei</emphasis>
|
||
</taxonomicName>
|
||
, and in the larger sample of
|
||
<taxonomicName box="[1320,1437,151,176]" class="Insecta" family="Megapodagrionidae" genus="Furagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="313" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[1320,1437,151,176]" italics="true" pageId="24" pageNumber="313">Furagrion</emphasis>
|
||
</taxonomicName>
|
||
available to us, two (
|
||
<emphasis box="[441,474,187,212]" italics="true" pageId="24" pageNumber="313">e.g</emphasis>
|
||
., FUM-N 13856,
|
||
<figureCitation box="[683,752,187,213]" captionStart="FIGURE 2" captionStartId="2.[152,255,1807,1831]" captionTargetBox="[151,1421,1101,1773]" captionTargetId="figure-470@2.[151,1436,1101,1784]" captionTargetPageId="2" captionText="FIGURE 2. FUM-N 13856, with which Petrulevičius et al. (2008) defined Furagrion (as ERK-KL-T1): A, photograph; B, drawing. Both to scale, 5 mm. A forewing. Ax1, Ax2 = antenodal cross veins 1 and 2; arc = arculus; b = brace vein; CuA = anterior branch of the cubitus; IR1, IR2 = intercalar vein 1 and 2; MA, MP = the anterior and posterior branches of the media; nod = nodus; pt = pterostigma; Q = quadrangle; RA = radius anterior; RP1, RP2, RP3-4 = branches of the radius posterior; sn = subnodus; * = origin of RP2." figureDoi="http://doi.org/10.5281/zenodo.7906176" httpUri="https://zenodo.org/record/7906176/files/figure.png" pageId="24" pageNumber="313">Fig. 2</figureCitation>
|
||
) to four (FUM-N 14704,
|
||
<figureCitation box="[1052,1134,187,213]" captionStart="FIGURE 17" captionStartId="20.[151,250,913,937]" captionTargetBox="[151,1436,181,875]" captionTargetId="figure-28@20.[151,1436,181,889]" captionTargetPageId="20" captionText="FIGURE 17. FUM-N 14704: A, photograph; B, drawing. A forewing. Both to scale, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7906210" httpUri="https://zenodo.org/record/7906210/files/figure.png" pageId="24" pageNumber="313">Fig. 17</figureCitation>
|
||
) cells long and, therefore, consider this to be within intraspecific variation.
|
||
</paragraph>
|
||
<paragraph blockId="24.[151,1437,151,825]" pageId="24" pageNumber="313">
|
||
4- The number of cells in the precubital field (MP-CuA space) to the level of the origin of RP3-4 (three in
|
||
<taxonomicName authority="Archibald & Ware & Rasmussen & Sylvestersen & Olsen & Simonsen, 2023" authorityName="Archibald & Ware & Rasmussen & Sylvestersen & Olsen & Simonsen" authorityYear="2023" box="[198,332,295,320]" class="Insecta" family="Megapodagrionidae" genus="Morsagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="313" phylum="Arthropoda" rank="genus" status="n. gen.">
|
||
<emphasis box="[198,332,295,320]" italics="true" pageId="24" pageNumber="313">Morsagrion</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[339,417,295,321]" pageId="24" pageNumber="313" rank="genus">n. gen.</taxonomicNameLabel>
|
||
, four at
|
||
<taxonomicName box="[509,626,295,320]" class="Insecta" family="Megapodagrionidae" genus="Furagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="313" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[509,626,295,320]" italics="true" pageId="24" pageNumber="313">Furagrion</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="24.[151,1437,151,825]" pageId="24" pageNumber="313">
|
||
- Like Zessin, we count three in
|
||
<taxonomicName authorityName="Archibald & Ware & Rasmussen & Sylvestersen & Olsen & Simonsen" authorityYear="2023" baseAuthorityName="Zessin" baseAuthorityYear="2011" box="[535,668,331,356]" class="Insecta" family="Megapodagrionidae" genus="Furagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="313" phylum="Arthropoda" rank="species" species="ansorgei">
|
||
<emphasis box="[535,668,331,356]" italics="true" pageId="24" pageNumber="313">M. ansorgei</emphasis>
|
||
</taxonomicName>
|
||
, but with our larger sample we find between two (hind wings of MM-10752) to five (FUM-N 11146) in
|
||
<taxonomicName box="[583,700,367,392]" class="Insecta" family="Megapodagrionidae" genus="Furagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="313" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[583,700,367,392]" italics="true" pageId="24" pageNumber="313">Furagrion</emphasis>
|
||
</taxonomicName>
|
||
. We, therefore, also consider this to be intraspecific variation.
|
||
</paragraph>
|
||
<paragraph blockId="24.[151,1437,151,825]" pageId="24" pageNumber="313">
|
||
5- Eight cells between the beginning of IR2 and RP
|
||
<quantity box="[744,786,403,429]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="24" pageNumber="313" unit="in" value="2.0">2 in</quantity>
|
||
<taxonomicName authorityName="Archibald & Ware & Rasmussen & Sylvestersen & Olsen & Simonsen" authorityYear="2023" box="[793,927,403,428]" class="Insecta" family="Megapodagrionidae" genus="Morsagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="313" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[793,927,403,428]" italics="true" pageId="24" pageNumber="313">Morsagrion</emphasis>
|
||
</taxonomicName>
|
||
and five in
|
||
<taxonomicName box="[1059,1176,403,428]" class="Insecta" family="Megapodagrionidae" genus="Furagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="313" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[1059,1176,403,428]" italics="true" pageId="24" pageNumber="313">Furagrion</emphasis>
|
||
</taxonomicName>
|
||
. This varies from eight cells in Henriksen’s
|
||
<typeStatus box="[422,470,439,465]" pageId="24" pageNumber="313">type</typeStatus>
|
||
MGUH 1819 to five in FUM-N 13856, with fewer in hind wings,
|
||
<emphasis box="[1199,1232,439,464]" italics="true" pageId="24" pageNumber="313">e.g</emphasis>
|
||
., MM-10750 with 7.5 and 6.5 cells in the forewings and
|
||
<quantity box="[624,688,474,501]" metricMagnitude="-1" metricUnit="m" metricValue="1.143" pageId="24" pageNumber="313" unit="in" value="4.5">4.5 in</quantity>
|
||
the hind wing.
|
||
</paragraph>
|
||
<paragraph blockId="24.[151,1437,151,825]" pageId="24" pageNumber="313">- In all of these except MGUH 34113, which we tentatively associate with MOA 770, the IR1-RP2 space is more than one cells wide distad about the level of the pterostigma base (see below).</paragraph>
|
||
<paragraph blockId="24.[151,1437,151,825]" box="[151,741,583,609]" pageId="24" pageNumber="313">6- The shape of the discoidal and subdiscoidal cell.</paragraph>
|
||
<paragraph blockId="24.[151,1437,151,825]" box="[151,644,619,645]" pageId="24" pageNumber="313">- We find that this does not greatly differ.</paragraph>
|
||
<paragraph blockId="24.[151,1437,151,825]" box="[151,877,655,681]" pageId="24" pageNumber="313">7- The length of the double rows of cells in the IR1-RP2 space.</paragraph>
|
||
<paragraph blockId="24.[151,1437,151,825]" pageId="24" pageNumber="313">
|
||
- This space is more than one cell wide for only about three cells from ending on the wing margin, whereas in
|
||
<taxonomicName baseAuthorityName="Henriksen" baseAuthorityYear="1922" class="Insecta" family="Megapodagrionidae" genus="Furagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="313" phylum="Arthropoda" rank="species" species="jutlandicus">
|
||
<emphasis italics="true" pageId="24" pageNumber="313">F. jutlandicus</emphasis>
|
||
</taxonomicName>
|
||
, this space is more than one cell wide to about the level of or slightly more basal of the basal end of the pterostigma. This occurs in all
|
||
<taxonomicName box="[628,745,763,788]" class="Insecta" family="Megapodagrionidae" genus="Furagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="24" pageNumber="313" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[628,745,763,788]" italics="true" pageId="24" pageNumber="313">Furagrion</emphasis>
|
||
</taxonomicName>
|
||
specimens that we examined except possibly MGUH 34113 (below) and does appear distinctive.
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7906220" ID-Zenodo-Dep="7906220" httpUri="https://zenodo.org/record/7906220/files/figure.png" pageId="24" pageNumber="313" startId="24.[151,250,1877,1901]" targetBox="[151,1431,849,1843]" targetPageId="24" targetType="figure">
|
||
<paragraph blockId="24.[151,1436,1877,1938]" pageId="24" pageNumber="313">
|
||
<emphasis bold="true" box="[151,288,1877,1901]" pageId="24" pageNumber="313">FIGURE 22.</emphasis>
|
||
MGUH 34113, a forewing: A, photograph of the part; B, photograph of the counterpart; C, drawing from both A and B. Note infuscation very light on the part as preserved, darker on the counterpart (see text). All to scale, 5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="25.[151,1437,151,393]" pageId="25" pageNumber="314">
|
||
We, therefore, find that MOA 770/1and MGUH 34113 do not differ enough from the fossils assigned here to
|
||
<taxonomicName baseAuthorityName="Henriksen" baseAuthorityYear="1922" class="Insecta" family="Megapodagrionidae" genus="Furagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="314" phylum="Arthropoda" rank="species" species="jutlandicus">
|
||
<emphasis italics="true" pageId="25" pageNumber="314">F. jutlandicus</emphasis>
|
||
</taxonomicName>
|
||
to justify placing them in a separate genus, and so treat
|
||
<taxonomicName authorityName="Archibald & Ware & Rasmussen & Sylvestersen & Olsen & Simonsen" authorityYear="2023" box="[905,1039,187,212]" class="Insecta" family="Megapodagrionidae" genus="Morsagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="314" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[905,1039,187,212]" italics="true" pageId="25" pageNumber="314">Morsagrion</emphasis>
|
||
</taxonomicName>
|
||
as a junior synonym of
|
||
<taxonomicName box="[1309,1426,187,212]" class="Insecta" family="Megapodagrionidae" genus="Furagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="314" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[1309,1426,187,212]" italics="true" pageId="25" pageNumber="314">Furagrion</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph blockId="25.[151,1437,151,393]" pageId="25" pageNumber="314">
|
||
Character state 7 of MOA 770 does appear distinctive among the
|
||
<taxonomicName box="[971,1088,223,248]" class="Insecta" family="Megapodagrionidae" genus="Furagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="314" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[971,1088,223,248]" italics="true" pageId="25" pageNumber="314">Furagrion</emphasis>
|
||
</taxonomicName>
|
||
specimens examined (but see
|
||
<specimenCode box="[151,316,259,285]" collectionCode="MGUH" country="Denmark" httpUri="http://grbio.org/cool/c1ae-v64p" name="Museum Geologicum Universitatis Hafniensis" pageId="25" pageNumber="314">MGUH 34113</specimenCode>
|
||
, below), supporting its status as a separate species.
|
||
<collectorName box="[899,1003,259,285]" pageId="25" pageNumber="314">However</collectorName>
|
||
, while it is shown in
|
||
<collectorName box="[1245,1337,259,285]" pageId="25" pageNumber="314">Zessin’s</collectorName>
|
||
drawing (his figs. 1 and 4), this region appears unclear in his photograph (his fig. 3) and in a slightly higher resolution version of it provided by
|
||
<collectorName box="[345,462,331,357]" pageId="25" pageNumber="314">J. Ansorge</collectorName>
|
||
(see our drawing made from this,
|
||
<figureCitation box="[847,931,331,357]" captionStart="FIGURE 23" captionStartId="25.[151,250,1216,1240]" captionTargetBox="[156,1434,422,1182]" captionTargetId="figure-150@25.[151,1436,417,1193]" captionTargetPageId="25" captionText="FIGURE 23. A, Furagrion ansorgei MOA 770 redrawn from a higher resolution copy of Zessin’s (2011) figure 3 photograph provided by J. Ansorge; shape compared with that of the proposed neotype (blue) B, forewing; C, hind wing. A to scale, 5 mm; B, C wings to no scales, the sizes of proposed neotype wings are adjusted to match arculus to apex and anterior wing margin of MOA 770 to compare shape." figureDoi="http://doi.org/10.5281/zenodo.7906222" httpUri="https://zenodo.org/record/7906222/files/figure.png" pageId="25" pageNumber="314">Fig. 23</figureCitation>
|
||
).
|
||
<collectorName box="[949,986,331,357]" pageId="25" pageNumber="314">We</collectorName>
|
||
consider this character state as tentative until the
|
||
<typeStatus box="[251,299,367,393]" pageId="25" pageNumber="314">type</typeStatus>
|
||
specimen can be examined.
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7906222" ID-Zenodo-Dep="7906222" httpUri="https://zenodo.org/record/7906222/files/figure.png" pageId="25" pageNumber="314" startId="25.[151,250,1216,1240]" targetBox="[156,1434,422,1182]" targetPageId="25" targetType="figure">
|
||
<paragraph blockId="25.[151,1436,1216,1348]" pageId="25" pageNumber="314">
|
||
<emphasis bold="true" box="[151,289,1216,1240]" pageId="25" pageNumber="314">FIGURE 23.</emphasis>
|
||
A,
|
||
<taxonomicName authority="MOA" authorityName="MOA" box="[325,590,1216,1240]" class="Insecta" family="Megapodagrionidae" genus="Furagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="314" phylum="Arthropoda" rank="species" species="ansorgei">
|
||
<emphasis box="[325,526,1217,1240]" italics="true" pageId="25" pageNumber="314">Furagrion ansorgei</emphasis>
|
||
MOA
|
||
</taxonomicName>
|
||
770 redrawn from a higher resolution copy of
|
||
<bibRefCitation author="Zessin, W." box="[1070,1227,1216,1240]" pageId="25" pageNumber="314" pagination="64 - 73" refId="ref14309" refString="Zessin, W. (2011) Neue Insekten aus dem Moler (Palaozan / Eozan) von Danemark Teil 1 (Odonata: Epallagidae, Megapodagrioniidae), Virgo, Mitteilungsblatt des Entomologischen Vereins Mecklenburg, 14, 64 - 73." type="journal article" year="2011">Zessin’s (2011)</bibRefCitation>
|
||
figure 3 photograph provided by J. Ansorge; shape compared with that of the proposed neotype (blue) B, forewing; C, hind wing. A to scale, 5 mm; B, C wings to no scales, the sizes of proposed neotype wings are adjusted to match arculus to apex and anterior wing margin of MOA 770 to compare shape.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="25.[151,1436,1394,1745]" pageId="25" pageNumber="314">
|
||
The shape of wing MOA 770 provides stronger support for recognizing it as a separate species (
|
||
<figureCitation box="[1262,1424,1394,1420]" captionStart="FIGURE 23" captionStartId="25.[151,250,1216,1240]" captionTargetBox="[156,1434,422,1182]" captionTargetId="figure-150@25.[151,1436,417,1193]" captionTargetPageId="25" captionText="FIGURE 23. A, Furagrion ansorgei MOA 770 redrawn from a higher resolution copy of Zessin’s (2011) figure 3 photograph provided by J. Ansorge; shape compared with that of the proposed neotype (blue) B, forewing; C, hind wing. A to scale, 5 mm; B, C wings to no scales, the sizes of proposed neotype wings are adjusted to match arculus to apex and anterior wing margin of MOA 770 to compare shape." figureDoi="http://doi.org/10.5281/zenodo.7906222" httpUri="https://zenodo.org/record/7906222/files/figure.png" pageId="25" pageNumber="314">Fig. 23B, 23C</figureCitation>
|
||
). It has a greater length / width ratio than all
|
||
<taxonomicName box="[634,751,1431,1456]" class="Insecta" family="Megapodagrionidae" genus="Furagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="314" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[634,751,1431,1456]" italics="true" pageId="25" pageNumber="314">Furagrion</emphasis>
|
||
</taxonomicName>
|
||
wings, fore- or hind, by all length measurements except RP2 to apex / width (
|
||
<tableCitation box="[335,428,1466,1492]" captionStart="TABLE 2" captionStartId="5.[152,244,151,175]" captionTargetId="graphics-41@5.[151,1434,180,606]" captionTargetPageId="5" captionText="TABLE 2. Ratios of measurements in Table 1. Abbreviations as in Table 1." pageId="25" pageNumber="314">Tables 2</tableCitation>
|
||
and
|
||
<tableCitation box="[483,497,1466,1492]" captionStart="TABLE 3" captionStartId="6.[152,244,1052,1076]" captionTargetId="graphics-456@6.[151,1435,1117,1636]" captionTargetPageId="6" captionText="TABLE 3. Measurements in millimeters and ratios of these for wings not known to be fore- or hind wings. Abbreviations as in Table 1." pageId="25" pageNumber="314">3</tableCitation>
|
||
and see PCA, above).
|
||
</paragraph>
|
||
<paragraph blockId="25.[151,1436,1394,1745]" box="[199,1103,1502,1529]" pageId="25" pageNumber="314">
|
||
By these reasons, we treat MOA 770 as
|
||
<taxonomicName authority="(Zessin)" authorityName="Archibald & Ware & Rasmussen & Sylvestersen & Olsen & Simonsen" authorityYear="2023" baseAuthorityName="Zessin" baseAuthorityYear="2011" box="[648,968,1502,1528]" class="Insecta" family="Megapodagrionidae" genus="Furagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="314" phylum="Arthropoda" rank="species" species="ansorgei" status="comb. nov.">
|
||
<emphasis box="[648,869,1503,1528]" italics="true" pageId="25" pageNumber="314">Furagrion ansorgei</emphasis>
|
||
(Zessin)
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[974,1103,1502,1528]" pageId="25" pageNumber="314">
|
||
<taxonomicNameLabel box="[974,1103,1502,1528]" pageId="25" pageNumber="314" rank="species">comb. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph blockId="25.[151,1436,1394,1745]" pageId="25" pageNumber="314">
|
||
MGUH 34113 (
|
||
<figureCitation box="[378,460,1538,1564]" captionStart="FIGURE 22" captionStartId="24.[151,250,1877,1901]" captionTargetBox="[151,1431,849,1843]" captionTargetId="figure-365@24.[151,1436,849,1854]" captionTargetPageId="24" captionText="FIGURE 22. MGUH 34113, a forewing: A, photograph of the part; B, photograph of the counterpart; C, drawing from both A and B. Note infuscation very light on the part as preserved, darker on the counterpart (see text). All to scale, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7906220" httpUri="https://zenodo.org/record/7906220/files/figure.png" pageId="25" pageNumber="314">Fig. 22</figureCitation>
|
||
) might be conspecific with MOA 770: Its IR1-RP2 space becomes more than one cell wide closer to the level of the distal end of pterostigma than basal, suggesting an association with
|
||
<taxonomicName authority="MOA" authorityName="MOA" box="[1242,1436,1574,1600]" class="Insecta" family="Megapodagrionidae" genus="Furagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="314" phylum="Arthropoda" rank="species" species="ansorgei">
|
||
<emphasis box="[1242,1366,1575,1600]" italics="true" pageId="25" pageNumber="314">F. ansorgei</emphasis>
|
||
MOA
|
||
</taxonomicName>
|
||
770. This is, however, not as distinct as in Zessin’s drawing of the MOA 770 wing. More importantly, this wing is relatively narrow like MOA 770. Furthermore, it is smaller (see PCA and
|
||
<tableCitation box="[1023,1119,1646,1672]" captionStart="TABLE 3" captionStartId="6.[152,244,1052,1076]" captionTargetId="graphics-456@6.[151,1435,1117,1636]" captionTargetPageId="6" captionText="TABLE 3. Measurements in millimeters and ratios of these for wings not known to be fore- or hind wings. Abbreviations as in Table 1." pageId="25" pageNumber="314">Tables 3</tableCitation>
|
||
and
|
||
<tableCitation box="[1176,1192,1646,1672]" captionStart="TABLE 4" captionStartId="14.[151,243,553,579]" captionTargetId="graphics-88@14.[150,2037,618,1603]" captionTargetPageId="14" captionText="TABLE 4. Measurements in millimeters and ratios of these used in principle component analysis. Colours correspond to those in Fig. 10: known forewings (FW) in red and hindwings (HW) in blue, those unknown as fore- or hind wings are in green, and the F. ansorgei wing is in purple. Abbreviations as in Table 1." pageId="25" pageNumber="314">4</tableCitation>
|
||
), although this might represent intraspecific variation or sexual dimorphism. Given this uncertainty, we tentatively treat this specimen as
|
||
<taxonomicName box="[151,276,1719,1744]" class="Insecta" family="Megapodagrionidae" genus="Furagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="25" pageNumber="314" phylum="Arthropoda" rank="species" species="ansorgei">
|
||
<emphasis box="[151,276,1719,1744]" italics="true" pageId="25" pageNumber="314">F. ansorgei</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |