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<document id="AC881FDA7DE2E98346F170B15FECE416" ID-CLB-Dataset="24795" ID-DOI="10.11646/zootaxa.4701.1.3" ID-GBIF-Dataset="1fb77063-018a-4222-8733-9450729ac222" ID-ISSN="1175-5326" ID-Zenodo-Dep="3557803" ID-ZooBank="668205B0-B6E9-48F3-9E21-87C534CDC837" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1575043522256" checkinUser="plazi" docAuthor="Huang, Sunbin, Tú, Výõng Tân, Phú, Ph ạm Văn, Tian, Mingyi &amp; Faille, Arnaud" docDate="2019" docId="0399DA2B7F38A81AFF33FD0DFAC0F728" docLanguage="en" docName="zootaxa.4701.1.3.pdf" docOrigin="Zootaxa 4701 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tonkinaphaenops impunctatus Huang &amp; Tú &amp; Phú &amp; Tian &amp; Faille 2019, sp. nov." docType="treatment" docVersion="8" lastPageNumber="42" masterDocId="FFA0A2537F3FA81DFFA4FF9CFFD8F02F" masterDocTitle="Contributions to the knowledge of the genus Tonkinaphaenops Deuve, 2013 from China and Vietnam (Coleoptera: Carabidae: Trechinae)" masterLastPageNumber="53" masterPageNumber="35" pageNumber="42" updateTime="1698769588363" updateUser="ExternalLinkService">
<mods:mods id="E13AF601490DDE9900298A73F3076093" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A55BEA3F88BE7A934EA8C5CD8C885C69">
<mods:title id="298F710A38DE85F3A8593311D39183BC">Contributions to the knowledge of the genus Tonkinaphaenops Deuve, 2013 from China and Vietnam (Coleoptera: Carabidae: Trechinae)</mods:title>
</mods:titleInfo>
<mods:name id="A224D40505FA42ED0CF115927FAC7A01" type="personal">
<mods:role id="8B00C57E10C387EA205755BA93B1FFC1">
<mods:roleTerm id="B349A1FDB008FE8990C3554BEF3A6856">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3420493B351F6DAE878F124E4F20CE63">Huang, Sunbin</mods:namePart>
</mods:name>
<mods:name id="C6F9901089DB678208FFE1559CE9CC0E" type="personal">
<mods:role id="29882D2A7A7D2E91B464AC05DC095A35">
<mods:roleTerm id="5E38AA9BA14E27614B728D271B6AE02C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A1AB569934C7B621C4A4D79990A46BBD">Tú, Výõng Tân</mods:namePart>
</mods:name>
<mods:name id="DEBF65FD6C74FA5671FC11EA69C27F68" type="personal">
<mods:role id="44047B51CBF974ADF2933480F192E33B">
<mods:roleTerm id="5EB68D8066E745493AB7848959D86AA3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C893B97B1B455B34FB84BA1DDDDFB015">Phú, Ph ạm Văn</mods:namePart>
</mods:name>
<mods:name id="0735E26DD4019FDB1846BC4DF73A5D84" type="personal">
<mods:role id="B4B0243C1FA6E3ACA50B22030DC1B6E1">
<mods:roleTerm id="6AA9047BCB0E13411F2CD2FC630FC239">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3230FE1EBAEE06B52DBF982FF909971A">Tian, Mingyi</mods:namePart>
</mods:name>
<mods:name id="6BFD7C72F2730F083D352777DA43155C" type="personal">
<mods:role id="3DE59A3E90483FAFB6923A4C7A438176">
<mods:roleTerm id="93A602178EC754BC68018BE2DAE94AF9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AA671D353F260033F1BF2EFEC3958FDD">Faille, Arnaud</mods:namePart>
</mods:name>
<mods:typeOfResource id="EF765661CADE5B76AF91884F9DC6C5B9">text</mods:typeOfResource>
<mods:relatedItem id="B43996CE81AB33EEA4465E2A7A46500C" type="host">
<mods:titleInfo id="DA4D4AE95DCEEC4561F6DAE75920B137">
<mods:title id="24442D7AF386B5E4C70313470E0071D2">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="4896CFA4F3E0F887AD0E9EA001AFD06E">
<mods:date id="3D9954908644D815545712B2A0C4C9FA">2019</mods:date>
<mods:detail id="769F1FE3FF7DEB1C6B53C16F7ED31156" type="pubDate">
<mods:number id="F3974644E80D5E860F5CEB0514961AD3">2019-11-26</mods:number>
</mods:detail>
<mods:detail id="15C1678DD70AF58408C5B30586F3769E" type="volume">
<mods:number id="255C3A7061E28EADD1C18B7213D15360">4701</mods:number>
</mods:detail>
<mods:detail id="555C79AA44B819DCACE2750A9913288B" type="issue">
<mods:number id="DB8B1315F24334432D22682329403625">1</mods:number>
</mods:detail>
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<mods:start id="C56777ABA953BC786D07B5A03CDD98A5">35</mods:start>
<mods:end id="4A5D54E232D97A1BD7EADEE8506D0E99">53</mods:end>
</mods:extent>
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<mods:classification id="7243982D8B68C88E466F84310A4DAC74">journal article</mods:classification>
<mods:identifier id="2F23180478644FC0DC263F4B21E41F12" type="CLB-Dataset">24795</mods:identifier>
<mods:identifier id="044110EA7F14D8D5C1EDC44F660602FD" type="DOI">10.11646/zootaxa.4701.1.3</mods:identifier>
<mods:identifier id="77E98F86EB94410D692BBC49E570F89A" type="GBIF-Dataset">1fb77063-018a-4222-8733-9450729ac222</mods:identifier>
<mods:identifier id="596EA81E0B9DF10A2D5E7D1266393CDC" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="4D013619393B1250054B300921157C78" type="Zenodo-Dep">3557803</mods:identifier>
<mods:identifier id="E82BDE4CECF257222231BDA1C45D95EA" type="ZooBank">668205B0-B6E9-48F3-9E21-87C534CDC837</mods:identifier>
</mods:mods>
<treatment id="0399DA2B7F38A81AFF33FD0DFAC0F728" ID-DOI="http://doi.org/10.5281/zenodo.5929260" ID-GBIF-Taxon="160605349" ID-Zenodo-Dep="5929260" LSID="urn:lsid:plazi:treatment:0399DA2B7F38A81AFF33FD0DFAC0F728" httpUri="http://treatment.plazi.org/id/0399DA2B7F38A81AFF33FD0DFAC0F728" lastPageNumber="42" pageId="7" pageNumber="42">
<subSubSection id="C32A38B67F38A81AFF33FD0DFC9AF284" box="[151,834,657,684]" pageId="7" pageNumber="42" type="nomenclature">
<paragraph id="8B8F6B3D7F38A81AFF33FD0DFC9AF284" blockId="7.[151,834,657,719]" box="[151,834,657,684]" pageId="7" pageNumber="42">
<heading id="D0C7DC517F38A81AFF33FD0DFC9AF284" bold="true" box="[151,834,657,684]" fontSize="11" level="1" pageId="7" pageNumber="42" reason="1">
<emphasis id="B944B72F7F38A81AFF33FD0DFC9AF284" bold="true" box="[151,834,657,684]" pageId="7" pageNumber="42">
<taxonomicName id="4C3010BE7F38A81AFF33FD0DFD00F284" ID-CoL="8MBHV" authority="Faille &amp; Huang" authorityName="Huang &amp; Tú &amp; Phú &amp; Tian &amp; Faille" authorityYear="2019" box="[151,728,657,684]" class="Insecta" family="Carabidae" genus="Tonkinaphaenops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="42" phylum="Arthropoda" rank="species" species="impunctatus" status="sp. nov.">
<emphasis id="B944B72F7F38A81AFF33FD0DFDD4F284" bold="true" box="[151,524,657,684]" italics="true" pageId="7" pageNumber="42">Tonkinaphaenops impunctatus</emphasis>
Faille &amp; Huang
</taxonomicName>
,
<taxonomicNameLabel id="A2770A547F38A81AFD40FD0DFC9AF284" box="[740,834,657,683]" pageId="7" pageNumber="42" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32A38B67F38A81AFF33FD29FEB4F2E0" box="[151,364,693,719]" pageId="7" pageNumber="42" type="description">
<paragraph id="8B8F6B3D7F38A81AFF33FD29FEB4F2E0" blockId="7.[151,834,657,719]" box="[151,364,693,719]" pageId="7" pageNumber="42">
(
<figureCitation id="130B77B87F38A81AFF3BFD29FF35F2E0" box="[159,237,693,719]" captionStart="FIGURE 1" captionStartId="1.[151,250,1801,1826]" captionTargetBox="[159,1428,877,1779]" captionTargetId="figure@1.[151,1436,877,1781]" captionTargetPageId="1" captionText="FIGURE 1. Distribution map of Tonkinaphaenops Deuve, 2013" figureDoi="http://doi.org/10.5281/zenodo.3557805" httpUri="https://zenodo.org/record/3557805/files/figure.png" pageId="7" pageNumber="42">Figs. 1</figureCitation>
,
<figureCitation id="130B77B87F38A81AFF5FFD29FEFFF2E0" box="[251,295,693,719]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStart-2="FIGURE 8" captionStartId-0="8.[151,250,2009,2034]" captionStartId-1="9.[151,250,1267,1292]" captionStartId-2="10.[151,250,1964,1989]" captionTargetBox-1="[169,1424,191,1220]" captionTargetBox-2="[168,1420,174,1945]" captionTargetId-0="figure@8.[386,1201,174,1990]" captionTargetId-1="figure@9.[151,1436,171,1245]" captionTargetId-2="figure@10.[168,1420,174,1945]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionTargetPageId-2="10" captionText-0="FIGURE 6. Tonkinaphaenops impunctatus sp. nov., habitus of holotype, male." captionText-1="FIGURE 7. Male genitalia of Tonkinaphaenops impunctatus sp. nov. (median lobe and parameres, lateral view)." captionText-2="FIGURE 8. Cave Hang Rắn, type locality of Tonkinaphaenops impunctatus sp. nov. (A. entrance; B. the wettest part of the cave where the beetles were collected)." figureDoi-0="http://doi.org/10.5281/zenodo.3614842" figureDoi-1="http://doi.org/10.5281/zenodo.3557813" figureDoi-2="http://doi.org/10.5281/zenodo.3557815" httpUri-0="https://zenodo.org/record/3614842/files/figure.png" httpUri-1="https://zenodo.org/record/3557813/files/figure.png" httpUri-2="https://zenodo.org/record/3557815/files/figure.png" pageId="7" pageNumber="42">6–8</figureCitation>
,
<figureCitation id="130B77B87F38A81AFE97FD29FEBCF2E0" box="[307,356,693,719]" captionStart="FIGURE 14" captionStartId="16.[151,250,1912,1937]" captionTargetBox="[214,1391,181,1869]" captionTargetId="figure@16.[189,1398,181,1887]" captionTargetPageId="16" captionText="FIGURE 14. Chaetotaxal pattern of Tonkinaphaenops (A. Tonkinaphaenops marinae Deuve, 2013, holotype, male from cave TH20; B, C. Tonkinaphaenops yinquanicus sp. nov.: B. male from Yinquan Dong, C. female from Yinquan Dong; D. Tonkinaphaenops impunctatus sp. nov., male from Hang Rắn cave; E. Tonkinaphaenops jingxicus sp. nov., female from Nianluo Dong; F. Tonkinaphaenops anthonyi sp. nov., male from Hang Dõi cave)." figureDoi="http://doi.org/10.5281/zenodo.3557823" httpUri="https://zenodo.org/record/3557823/files/figure.png" pageId="7" pageNumber="42">14D</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C32A38B67F38A81AFF33FD61FEEAF3E4" pageId="7" pageNumber="42" type="materials_examined">
<paragraph id="8B8F6B3D7F38A81AFF33FD61FA45F314" blockId="7.[151,1437,764,1799]" pageId="7" pageNumber="42">
<materialsCitation id="3B5861607F38A81AFF33FD61FA45F314" ID-GBIF-Occurrence="2460169385" collectingDate="2018-10-17" collectionCode="MNHN" collectorName="Cao B &amp; Hang R &amp; A. Faille &amp; Faille, P. V &amp; Phu, V. T" country="Vietnam" county="Danh" elevation="642" latitude="22.778416" location="Tra Linh district" longLatPrecision="1" longitude="106.313835" municipality="To" pageId="7" pageNumber="42" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B944B72F7F38A81AFF33FD61FED3F338" bold="true" box="[151,267,765,791]" pageId="7" pageNumber="42">
<typeStatus id="548BD59F7F38A81AFF33FD61FEDEF338" box="[151,262,765,791]" pageId="7" pageNumber="42" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCount id="9D36A0B47F38A81AFEB7FD61FEF3F338" box="[275,299,765,791]" pageId="7" pageNumber="42" type="male">♂</specimenCount>
;
<collectingCountry id="F3272BAD7F38A81AFE93FD61FE42F338" box="[311,410,765,791]" name="Vietnam" pageId="7" pageNumber="42">Vietnam</collectingCountry>
:
<collectorName id="26C50EEB7F38A81AFE02FD61FE35F338" box="[422,493,764,791]" pageId="7" pageNumber="42">Cao B</collectorName>
ằng province:
<location id="8EEF3DE67F38A81AFD2BFD61FC93F338" LSID="urn:lsid:plazi:treatment:0399DA2B7F38A81AFF33FD0DFAC0F728:8EEF3DE67F38A81AFD2BFD61FC93F338" box="[655,843,765,791]" country="Vietnam" county="Danh" latitude="22.778416" longLatPrecision="1" longitude="106.313835" municipality="To" name="Tra Linh district" pageId="7" pageNumber="42">Trà Lĩnh district</location>
,
<location id="8EEF3DE67F38A81AFCF3FD60FCA2F338" LSID="urn:lsid:plazi:treatment:0399DA2B7F38A81AFF33FD0DFAC0F728:8EEF3DE67F38A81AFCF3FD60FCA2F338" box="[855,890,764,791]" country="Vietnam" county="Danh" latitude="22.778416" longLatPrecision="1" longitude="106.313835" municipality="To" name="Qu" pageId="7" pageNumber="42">Qu</location>
ốc
<collectingMunicipality id="6BEBF1477F38A81AFC3FFD60FC61F338" box="[923,953,764,791]" pageId="7" pageNumber="42">To</collectingMunicipality>
ản commune, Bản
<collectingCounty id="62EE13B17F38A81AFB29FD61FB16F338" box="[1165,1230,765,791]" pageId="7" pageNumber="42">Danh</collectingCounty>
:
<collectorName id="26C50EEB7F38A81AFB7DFD61FAE9F338" box="[1241,1329,764,791]" pageId="7" pageNumber="42">Hang R</collectorName>
ắn (snake cave) (=TH06);
<geoCoordinate id="EE040DFA7F38A81AFEEAFCBDFE28F314" box="[334,496,801,827]" degrees="22" direction="north" minutes="46" orientation="latitude" pageId="7" pageNumber="42" precision="1" seconds="42.3" value="22.778416">22°46’42.3’’N</geoCoordinate>
<geoCoordinate id="EE040DFA7F38A81AFE53FCBDFD7EF314" box="[503,678,801,827]" degrees="106" direction="east" minutes="18" orientation="longitude" pageId="7" pageNumber="42" precision="1" seconds="49.8" value="106.313835">106°18’49.8’’E</geoCoordinate>
,
<quantity id="4CC8C6D87F38A81AFD16FCBDFD2DF314" box="[690,757,801,827]" metricMagnitude="2" metricUnit="m" metricValue="6.42" pageId="7" pageNumber="42" unit="m" value="642.0">
<elevation id="001D8C0E7F38A81AFD16FCBDFD2DF314" box="[690,757,801,827]" metricMagnitude="2" metricUnit="m" metricValue="6.42" pageId="7" pageNumber="42" unit="m" value="642.0">642m</elevation>
</quantity>
; coll.
<collectorName id="26C50EEB7F38A81AFC9CFCBDFC45F314" box="[824,925,801,827]" pageId="7" pageNumber="42">A. Faille</collectorName>
, P.V.
<collectorName id="26C50EEB7F38A81AFC7EFCBDFBE6F314" box="[986,1086,801,827]" pageId="7" pageNumber="42">Phu, V.T</collectorName>
. Tu;
<date id="FF8E4DFD7F38A81AFBD0FCBDFAECF314" box="[1140,1332,801,827]" pageId="7" pageNumber="42" value="2018-10-17">
<collectingDate id="EFCAB4157F38A81AFBD0FCBDFAECF314" box="[1140,1332,801,827]" pageId="7" pageNumber="42" value="2018-10-17">17 October 2018</collectingDate>
</date>
;
<collectionCode id="ED21F3F87F38A81AFAE4FCBDFA41F314" box="[1344,1433,801,827]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="7" pageNumber="42">MNHN</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8B8F6B3D7F38A81AFF33FCD9FEEAF3E4" blockId="7.[151,1437,764,1799]" pageId="7" pageNumber="42">
<materialsCitation id="3B5861607F38A81AFF33FCD9FCC1F370" ID-GBIF-Occurrence="2460169378" box="[151,793,837,863]" collectingDate="2018-10-17" collectionCode="SMNS" collectorName="Cao B &amp; Hang R &amp; A. Faille &amp; Faille, P. V &amp; Phu, V. T" country="Vietnam" county="Danh" elevation="642" latitude="22.778416" location="Tra Linh district" longLatPrecision="1" longitude="106.313835" municipality="To" pageId="7" pageNumber="42" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
<emphasis id="B944B72F7F38A81AFF33FCD9FECEF370" bold="true" box="[151,278,837,863]" pageId="7" pageNumber="42">
<typeStatus id="548BD59F7F38A81AFF33FCD9FEC9F370" box="[151,273,837,863]" pageId="7" pageNumber="42" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<specimenCount id="9D36A0B47F38A81AFEBAFCD9FE92F370" box="[286,330,837,863]" pageId="7" pageNumber="42" type="male">1 ♂</specimenCount>
and
<specimenCount id="9D36A0B47F38A81AFE27FCD9FE77F370" box="[387,431,837,863]" pageId="7" pageNumber="42" type="female">1 ♀</specimenCount>
; same data as holotype;
<collectionCode id="ED21F3F87F38A81AFD6DFCD9FCC1F370" box="[713,793,837,863]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="7" pageNumber="42">SMNS</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B5861607F38A81AFC81FCD9FB1CF370" ID-GBIF-Occurrence="2460169384" box="[805,1220,837,863]" collectingDate="2018-10-17" collectionCode="MNHN" collectorName="Cao B &amp; Hang R &amp; A. Faille &amp; Faille, P. V &amp; Phu, V. T" country="Vietnam" county="Danh" elevation="642" latitude="22.778416" location="Tra Linh district" longLatPrecision="1" longitude="106.313835" municipality="To" pageId="7" pageNumber="42" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="9D36A0B47F38A81AFC81FCD9FC89F370" box="[805,849,837,863]" pageId="7" pageNumber="42" type="female">1 ♀</specimenCount>
; same data as holotype;
<collectionCode id="ED21F3F87F38A81AFBCFFCD9FB1CF370" box="[1131,1220,837,863]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="7" pageNumber="42">MNHN</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B5861607F38A81AFB74FCD9FE95F3AC" ID-GBIF-Occurrence="2460169375" collectingDate="2018-10-17" collectionCode="IEBR" collectorName="Cao B &amp; Hang R &amp; A. Faille &amp; Faille, P. V &amp; Phu, V. T" country="Vietnam" county="Danh" elevation="642" latitude="22.778416" location="Tra Linh district" longLatPrecision="1" longitude="106.313835" municipality="To" pageId="7" pageNumber="42" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="9D36A0B47F38A81AFB74FCD9FB25F370" box="[1232,1277,837,863]" pageId="7" pageNumber="42" type="female">1 ♀</specimenCount>
; same data as holotype;
<collectionCode id="ED21F3F87F38A81AFEAEFCF5FE95F3AC" box="[266,333,873,899]" pageId="7" pageNumber="42">IEBR</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B5861607F38A81AFEFEFCF5FC81F3AC" ID-GBIF-Occurrence="2460169383" box="[346,857,873,899]" collectingDate="2018-10-17" collectionCode="SCAU" collectorName="Cao B &amp; Hang R &amp; A. Faille &amp; Faille, P. V &amp; Phu, V. T" country="Vietnam" county="Danh" elevation="642" latitude="22.778416" location="Tra Linh district" longLatPrecision="1" longitude="106.313835" municipality="To" pageId="7" pageNumber="42" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D36A0B47F38A81AFEFEFCF5FE5EF3AC" box="[346,390,873,899]" pageId="7" pageNumber="42" type="male">1 ♂</specimenCount>
and
<specimenCount id="9D36A0B47F38A81AFE64FCF5FE35F3AC" box="[448,493,873,899]" pageId="7" pageNumber="42" type="female">1 ♀</specimenCount>
; same data as holotype;
<collectionCode id="ED21F3F87F38A81AFCAFFCF5FC81F3AC" box="[779,857,873,899]" pageId="7" pageNumber="42">SCAU</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B5861607F38A81AFCC1FCF5FF28F388" ID-GBIF-Occurrence="2460169381" collectingDate="2018-10-18" collectionCode="MNHN" collectorName="Cao B &amp; Hang R &amp; A. Faille &amp; Faille, P. V &amp; Phu, V. T" country="Vietnam" county="Danh" elevation="642" latitude="22.778416" location="Tra Linh district" longLatPrecision="1" longitude="106.313835" municipality="To" pageId="7" pageNumber="42" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D36A0B47F38A81AFCC1FCF5FC4BF3AC" box="[869,915,873,899]" pageId="7" pageNumber="42" type="male">1 ♂</specimenCount>
, same locality as holotype;
<date id="FF8E4DFD7F38A81AFB70FCF5FA4FF3AC" box="[1236,1431,873,899]" pageId="7" pageNumber="42" value="2018-10-18">
<collectingDate id="EFCAB4157F38A81AFB70FCF5FA4FF3AC" box="[1236,1431,873,899]" pageId="7" pageNumber="42" value="2018-10-18">18 October 2018</collectingDate>
</date>
;
<collectionCode id="ED21F3F87F38A81AFF33FC11FF28F388" box="[151,240,909,935]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="7" pageNumber="42">MNHN</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B5861607F38A81AFF5FFC11FC6BF388" ID-GBIF-Occurrence="2460169373" box="[251,947,909,935]" collectingDate="2018-10-17" collectionCode="SMNS" collectorName="Cao B &amp; Hang R &amp; A. Faille &amp; Faille, P. V &amp; Phu, V. T" country="Vietnam" county="Danh" elevation="642" latitude="22.778416" location="Tra Linh district" longLatPrecision="1" longitude="106.313835" municipality="To" pageId="7" pageNumber="42" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D36A0B47F38A81AFF5FFC11FEFDF388" box="[251,293,909,935]" pageId="7" pageNumber="42" type="male">1 ♂</specimenCount>
and
<specimenCount id="9D36A0B47F38A81AFEFFFC11FE5EF388" box="[347,390,909,935]" pageId="7" pageNumber="42" type="female">1 ♀</specimenCount>
; spares in ethanol; same data as holotype;
<collectionCode id="ED21F3F87F38A81AFCC7FC11FC6BF388" box="[867,947,909,935]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="7" pageNumber="42">SMNS</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B5861607F38A81AFC1AFC11FEF6F3E4" ID-GBIF-Occurrence="2460169382" collectingDate="2018-10-18" collectionCode="SMNS" collectorName="Cao B &amp; Hang R &amp; A. Faille &amp; Faille, P. V &amp; Phu, V. T" country="Vietnam" county="Danh" elevation="642" latitude="22.778416" location="Tra Linh district" longLatPrecision="1" longitude="106.313835" municipality="To" pageId="7" pageNumber="42" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="9D36A0B47F38A81AFC1AFC11FC30F388" box="[958,1000,909,935]" pageId="7" pageNumber="42" type="female">1 ♀</specimenCount>
, same locality as holotype,
<date id="FF8E4DFD7F38A81AFABBFC11FF0AF3E4" pageId="7" pageNumber="42" value="2018-10-18">
<collectingDate id="EFCAB4157F38A81AFABBFC11FF0AF3E4" pageId="7" pageNumber="42" value="2018-10-18">18 October 2018</collectingDate>
</date>
;
<collectionCode id="ED21F3F87F38A81AFF7AFC2DFEF6F3E4" box="[222,302,945,971]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="7" pageNumber="42">SMNS</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C32A38B67F38A81AFF63FC48FE40F4E8" pageId="7" pageNumber="42" type="description">
<paragraph id="8B8F6B3D7F38A81AFF63FC48FE40F4E8" blockId="7.[151,1437,764,1799]" pageId="7" pageNumber="42">
<emphasis id="B944B72F7F38A81AFF63FC48FE81F3C0" bold="true" box="[199,345,980,1007]" pageId="7" pageNumber="42">Description.</emphasis>
Similar to
<taxonomicName id="4C3010BE7F38A81AFE7EFC4AFDA5F3C0" authorityName="Huang &amp; Tú &amp; Phú &amp; Tian &amp; Faille" authorityYear="2019" box="[474,637,982,1007]" class="Insecta" family="Carabidae" genus="Tonkinaphaenops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="42" phylum="Arthropoda" rank="species" species="yinquanicus">
<emphasis id="B944B72F7F38A81AFE7EFC4AFDA5F3C0" box="[474,637,982,1007]" italics="true" pageId="7" pageNumber="42">T. yinquanicus</emphasis>
</taxonomicName>
. Habitus as in
<figureCitation id="130B77B87F38A81AFC8EFC49FCA8F3C0" box="[810,880,981,1007]" captionStart="FIGURE 6" captionStartId="8.[151,250,2009,2034]" captionTargetId="figure@8.[386,1201,174,1990]" captionTargetPageId="8" captionText="FIGURE 6. Tonkinaphaenops impunctatus sp. nov., habitus of holotype, male." figureDoi="http://doi.org/10.5281/zenodo.3614842" httpUri="https://zenodo.org/record/3614842/files/figure.png" pageId="7" pageNumber="42">Fig. 6</figureCitation>
. Length:
<quantity id="4CC8C6D87F38A81AFC7FFC48FBEAF3C0" box="[987,1074,980,1007]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="7" pageNumber="42" unit="mm" value="5.6">5.6 mm</quantity>
(from the middle of the labrum to apex of the elytron). Body elongate, elytra wide and convex, amygdaloid. Narrow fore-body; elytra wider than pronotum, which is as wide as the head. EL:
<quantity id="4CC8C6D87F38A81AFD37FB81FD0EF418" box="[659,726,1053,1079]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="7" pageNumber="42" unit="mm" value="3.0">3 mm</quantity>
, EW:
<quantity id="4CC8C6D87F38A81AFCB8FB81FCADF418" box="[796,885,1053,1079]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="7" pageNumber="42" unit="mm" value="1.8">1.8 mm</quantity>
. PL:
<quantity id="4CC8C6D87F38A81AFC14FB81FBD1F418" box="[944,1033,1053,1079]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="7" pageNumber="42" unit="mm" value="1.3">1.3 mm</quantity>
, PW:
<quantity id="4CC8C6D87F38A81AFBE9FB81FB6DF418" box="[1101,1205,1053,1079]" metricMagnitude="-4" metricUnit="m" metricValue="9.8" pageId="7" pageNumber="42" unit="mm" value="0.98">0.98 mm</quantity>
. HL:
<quantity id="4CC8C6D87F38A81AFB50FB80FA83F418" box="[1268,1371,1052,1079]" metricMagnitude="-3" metricUnit="m" metricValue="1.05" pageId="7" pageNumber="42" unit="mm" value="1.05">1.05 mm</quantity>
, HW:
<quantity id="4CC8C6D87F38A81AFF33FBDDFF27F474" box="[151,255,1089,1115]" metricMagnitude="-4" metricUnit="m" metricValue="7.6" pageId="7" pageNumber="42" unit="mm" value="0.76">0.76 mm</quantity>
. AL:
<quantity id="4CC8C6D87F38A81AFE99FBDCFE7DF474" box="[317,421,1088,1115]" metricMagnitude="-3" metricUnit="m" metricValue="3.95" pageId="7" pageNumber="42" unit="mm" value="3.95">3.95 mm</quantity>
. Color: body reddish brown, concolor, shiny. Fore body long and narrow, hind body elongated obovate, strongly convex dorsally. Posterior pronotal setae lacking. Pronotum covered with a small, sparse and short pubescence. Microsculpture: transverse meshes, very fine on elytra and pronotum, more visible and less transverse on the head.
</paragraph>
</subSubSection>
<subSubSection id="C32A38B67F38A81AFF63FB4DFBC7F520" pageId="7" pageNumber="42" type="etymology">
<paragraph id="8B8F6B3D7F38A81AFF63FB4DFBC7F520" blockId="7.[151,1437,764,1799]" pageId="7" pageNumber="42">
<emphasis id="B944B72F7F38A81AFF63FB4DFE96F4C4" bold="true" box="[199,334,1233,1259]" pageId="7" pageNumber="42">Etymology.</emphasis>
The specific epithet is an adjective referring to one of the main character differing from the other species of the genus, the absence of punctuate striae in the last fifth of the elytra.
</paragraph>
</subSubSection>
<subSubSection id="C32A38B67F38A81AFF63FA85FDA9F6B4" pageId="7" pageNumber="42" type="discussion">
<paragraph id="8B8F6B3D7F38A81AFF63FA85FC90F600" blockId="7.[151,1437,764,1799]" pageId="7" pageNumber="42">
<emphasis id="B944B72F7F38A81AFF63FA85FEE3F51C" bold="true" box="[199,315,1305,1331]" pageId="7" pageNumber="42">Remarks.</emphasis>
The new species is close to
<taxonomicName id="4C3010BE7F38A81AFDD6FA85FC43F51C" authorityName="Deuve" authorityYear="2013" box="[626,923,1305,1331]" class="Insecta" family="Carabidae" genus="Tonkinaphaenops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="42" phylum="Arthropoda" rank="species" species="marinae">
<emphasis id="B944B72F7F38A81AFDD6FA85FC43F51C" box="[626,923,1305,1331]" italics="true" pageId="7" pageNumber="42">Tonkinaphaenops marinae</emphasis>
</taxonomicName>
(
<figureCitation id="130B77B87F38A81AFC0EFA85FC34F51C" box="[938,1004,1305,1331]" captionStart="FIGURE 2" captionStartId="3.[151,250,1997,2022]" captionTargetBox="[188,1398,175,1963]" captionTargetId="figure@3.[172,1415,174,1978]" captionTargetPageId="3" captionText="FIGURE 2. Tonkinaphaenops marinae Deuve, 2013, habitus of holotype, male." figureDoi="http://doi.org/10.5281/zenodo.3557807" httpUri="https://zenodo.org/record/3557807/files/figure.png" pageId="7" pageNumber="42">Fig. 2</figureCitation>
) and
<taxonomicName id="4C3010BE7F38A81AFB8DFA86FB12F51C" authorityName="Huang &amp; Tú &amp; Phú &amp; Tian &amp; Faille" authorityYear="2019" box="[1065,1226,1306,1331]" class="Insecta" family="Carabidae" genus="Tonkinaphaenops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="42" phylum="Arthropoda" rank="species" species="yinquanicus">
<emphasis id="B944B72F7F38A81AFB8DFA86FB12F51C" box="[1065,1226,1306,1331]" italics="true" pageId="7" pageNumber="42">T. yinquanicus</emphasis>
</taxonomicName>
(
<figureCitation id="130B77B87F38A81AFB7DFA85FAC2F51C" box="[1241,1306,1305,1331]" captionStart="FIGURE 3" captionStartId="4.[151,250,1997,2022]" captionTargetId="figure@4.[405,1182,174,1978]" captionTargetPageId="4" captionText="FIGURE 3. Tonkinaphaenops yinquanicus sp. nov., habitus of holotype, male." figureDoi="http://doi.org/10.5281/zenodo.3614840" httpUri="https://zenodo.org/record/3614840/files/figure.png" pageId="7" pageNumber="42">Fig. 3</figureCitation>
) but differs from the first in the punctuation of elytra, which is effaced well before the end of the apex whereas the dots reach the apex of the elytra in
<taxonomicName id="4C3010BE7F38A81AFE02FAFEFDC6F554" authorityName="Deuve" authorityYear="2013" box="[422,542,1378,1403]" class="Insecta" family="Carabidae" genus="Tonkinaphaenops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="42" phylum="Arthropoda" rank="species" species="marinae">
<emphasis id="B944B72F7F38A81AFE02FAFEFDC6F554" box="[422,542,1378,1403]" italics="true" pageId="7" pageNumber="42">T. marinae</emphasis>
</taxonomicName>
. It is also distinct by the shape of the male genitalia, whose apex is rounded and flattened, similar to the next described species, while it is hooked in
<taxonomicName id="4C3010BE7F38A81AFC22FA1AFC25F5B0" authorityName="Deuve" authorityYear="2013" box="[902,1021,1414,1439]" class="Insecta" family="Carabidae" genus="Tonkinaphaenops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="42" phylum="Arthropoda" rank="species" species="marinae">
<emphasis id="B944B72F7F38A81AFC22FA1AFC25F5B0" box="[902,1021,1414,1439]" italics="true" pageId="7" pageNumber="42">T. marinae</emphasis>
</taxonomicName>
. From
<taxonomicName id="4C3010BE7F38A81AFBE9FA1AFB36F5B0" authorityName="Huang &amp; Tú &amp; Phú &amp; Tian &amp; Faille" authorityYear="2019" box="[1101,1262,1414,1439]" class="Insecta" family="Carabidae" genus="Tonkinaphaenops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="42" phylum="Arthropoda" rank="species" species="yinquanicus">
<emphasis id="B944B72F7F38A81AFBE9FA1AFB36F5B0" box="[1101,1262,1414,1439]" italics="true" pageId="7" pageNumber="42">T. yinquanicus</emphasis>
</taxonomicName>
, it differs in the shape of male genitalia less rounded at apex and the position of the 3
<superScript id="7C45C6757F38A81AFC04FA34FC76F599" attach="left" box="[928,942,1448,1462]" fontSize="6" pageId="7" pageNumber="42">rd</superScript>
pore of marginal umbilicate series closer to the 2
<superScript id="7C45C6757F38A81AFF6AFA50FF06F5F5" attach="left" box="[206,222,1484,1498]" fontSize="6" pageId="7" pageNumber="42">nd</superScript>
than to the 4
<superScript id="7C45C6757F38A81AFECAFA50FEA3F5F5" attach="left" box="[366,379,1484,1498]" fontSize="6" pageId="7" pageNumber="42">th</superScript>
whereas it is equidistant in
<emphasis id="B944B72F7F38A81AFD14FA52FC81F5C8" box="[688,857,1486,1511]" italics="true" pageId="7" pageNumber="42">
<taxonomicName id="4C3010BE7F38A81AFD14FA52FC8AF5C8" authorityName="Huang &amp; Tú &amp; Phú &amp; Tian &amp; Faille" authorityYear="2019" box="[688,850,1486,1511]" class="Insecta" family="Carabidae" genus="Tonkinaphaenops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="42" phylum="Arthropoda" rank="species" species="yinquanicus">T. yinquanicus</taxonomicName>
.
</emphasis>
Furthermore, the species is clearly distinct from the other species of
<taxonomicName id="4C3010BE7F38A81AFEE8FA6DFDC9F624" authorityName="Deuve" authorityYear="2013" box="[332,529,1521,1547]" class="Insecta" family="Carabidae" genus="Tonkinaphaenops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="42" phylum="Arthropoda" rank="genus">
<emphasis id="B944B72F7F38A81AFEE8FA6DFDC9F624" box="[332,529,1521,1547]" italics="true" pageId="7" pageNumber="42">Tonkinaphaenops</emphasis>
</taxonomicName>
by the presence of a small, sparse and short pubescence on the pronotum (mainly visible in lateral view) and the lack of pronotal posterior seta.
</paragraph>
<paragraph id="8B8F6B3D7F38A81AFF63F9A5FDA9F6B4" blockId="7.[151,1437,764,1799]" pageId="7" pageNumber="42">
Nevertheless, and because the only known specimen of
<taxonomicName id="4C3010BE7F38A81AFC95F9A6FC71F67C" authorityName="Deuve" authorityYear="2013" box="[817,937,1594,1619]" class="Insecta" family="Carabidae" genus="Tonkinaphaenops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="42" phylum="Arthropoda" rank="species" species="marinae">
<emphasis id="B944B72F7F38A81AFC95F9A6FC71F67C" box="[817,937,1594,1619]" italics="true" pageId="7" pageNumber="42">T. marinae</emphasis>
</taxonomicName>
is immature with a genitalia weakly sclerotized, new exemplars of
<taxonomicName id="4C3010BE7F38A81AFE3DF9C2FDC9F658" authorityName="Deuve" authorityYear="2013" box="[409,529,1630,1655]" class="Insecta" family="Carabidae" genus="Tonkinaphaenops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="42" phylum="Arthropoda" rank="species" species="marinae">
<emphasis id="B944B72F7F38A81AFE3DF9C2FDC9F658" box="[409,529,1630,1655]" italics="true" pageId="7" pageNumber="42">T. marinae</emphasis>
</taxonomicName>
from the
<typeStatus id="548BD59F7F38A81AFDDAF9C1FD76F658" box="[638,686,1629,1655]" pageId="7" pageNumber="42">type</typeStatus>
locality would be necessary to confirm the differences in the shape of male genitalia between the two species.
</paragraph>
</subSubSection>
<subSubSection id="C32A38B67F38A81AFF63F938FAC0F728" pageId="7" pageNumber="42" type="distribution">
<paragraph id="8B8F6B3D7F38A81AFF63F938FC95F690" blockId="7.[151,1437,764,1799]" box="[199,845,1700,1727]" pageId="7" pageNumber="42">
<emphasis id="B944B72F7F38A81AFF63F938FEB9F690" bold="true" box="[199,353,1700,1727]" pageId="7" pageNumber="42">Distribution.</emphasis>
Known only from the
<typeStatus id="548BD59F7F38A81AFDC4F939FD48F690" box="[608,656,1701,1727]" pageId="7" pageNumber="42">type</typeStatus>
locality (
<figureCitation id="130B77B87F38A81AFD5FF939FCE4F690" box="[763,828,1701,1727]" captionStart="FIGURE 1" captionStartId="1.[151,250,1801,1826]" captionTargetBox="[159,1428,877,1779]" captionTargetId="figure@1.[151,1436,877,1781]" captionTargetPageId="1" captionText="FIGURE 1. Distribution map of Tonkinaphaenops Deuve, 2013" figureDoi="http://doi.org/10.5281/zenodo.3557805" httpUri="https://zenodo.org/record/3557805/files/figure.png" pageId="7" pageNumber="42">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="8B8F6B3D7F38A81AFF63F955FAC0F728" blockId="7.[151,1437,764,1799]" pageId="7" pageNumber="42">
This cave is a tunnel, with a development of more than
<quantity id="4CC8C6D87F38A81AFC82F955FCB4F6CC" box="[806,876,1737,1763]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="7" pageNumber="42" unit="m" value="800.0">800 m</quantity>
(R. Huttler pers. comm.), the main part of the cave is fossil and dry. The beetles were collected in the wettest part of the cave, running on the clay (
<figureCitation id="130B77B87F38A81AFB62F971FAD2F728" box="[1222,1290,1773,1799]" captionStart="FIGURE 8" captionStartId="10.[151,250,1964,1989]" captionTargetBox="[168,1420,174,1945]" captionTargetId="figure@10.[168,1420,174,1945]" captionTargetPageId="10" captionText="FIGURE 8. Cave Hang Rắn, type locality of Tonkinaphaenops impunctatus sp. nov. (A. entrance; B. the wettest part of the cave where the beetles were collected)." figureDoi="http://doi.org/10.5281/zenodo.3557815" httpUri="https://zenodo.org/record/3557815/files/figure.png" pageId="7" pageNumber="42">Fig. 8</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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