551 lines
66 KiB
XML
551 lines
66 KiB
XML
<document id="0ECAC2304EF6B25AE75FE0B630381046" ID-CLB-Dataset="263342" ID-DOI="10.11646/zootaxa.5286.1.1" ID-GBIF-Dataset="67e7a105-1f62-414a-83d9-f758039eb726" ID-ISSN="1175-5326" ID-Zenodo-Dep="7958937" ID-ZooBank="F9834684-24D3-4795-B5EB-77B451DF856D" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="diego" checkinTime="1684793492880" checkinUser="plazi" docAuthor="Shi, Hongliang & Liang, Hongbin" docDate="2023" docId="03877623625EFFF92DEFB341FA7B5FA2" docLanguage="en" docName="zootaxa.5286.1.1.pdf" docOrigin="Zootaxa 5286 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5286.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Crossoglossa Chaudoir 1872" docType="treatment" docVersion="8" lastPageNumber="27" masterDocId="FFBE0E5B6246FFE32D78B27BFFE75D5A" masterDocTitle="Taxonomic revision of the genus Parena Motschulsky, 1860 (Coleoptera, Carabidae, Lebiini, Metallicina)" masterLastPageNumber="144" masterPageNumber="1" pageNumber="25" updateTime="1699506634024" updateUser="plazi">
|
||
<mods:mods id="8371ACC5F6261CCD2273541BED1B9C25" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="6A7B2E3E279862465D93A79D9F76B5A8">
|
||
<mods:title id="62012D7D651B22C6A3E76B3F6061D82A">Taxonomic revision of the genus Parena Motschulsky, 1860 (Coleoptera, Carabidae, Lebiini, Metallicina)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="545900009E4E5A7C332B66BF9A6C326F" type="personal">
|
||
<mods:role id="01BDB03E93873066DBFF5991BCF5B0BD">
|
||
<mods:roleTerm id="B7CE16623F41CAA74B793EFE81C9D253">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="99BCBA2D2A53184A3D64867CEC9F4B4B">Shi, Hongliang</mods:namePart>
|
||
<mods:nameIdentifier id="0F935F4E7A338256D3B79D1A36EC5994" type="ORCID">0000-0002-9989-5830</mods:nameIdentifier>
|
||
<mods:nameIdentifier id="817F2EDF60E5096D823C0A53F6C33AA8" type="email">shihl@bjfu.edu.cn</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:name id="99F569CB3666104E3774AFBE33AA5F2C" type="personal">
|
||
<mods:role id="E815714BEB9EF6F5AE2C7044C89DD0A6">
|
||
<mods:roleTerm id="E1B9BBC4B72BB5351D5FA0EC66B1BCE4">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="F2F6FDA5D4A6D6FDCE3A2A69C80341DE">Liang, Hongbin</mods:namePart>
|
||
<mods:nameIdentifier id="7F1F6727EF2297B37A812FEC26EC6413" type="ORCID">0000-0002-9989-5830</mods:nameIdentifier>
|
||
<mods:nameIdentifier id="757258FB57FC5D4BB79364EAD9A890B2" type="email">shihl@bjfu.edu.cn</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:typeOfResource id="620A5B4F7B537F051CD2F8DE06E62C1A">text</mods:typeOfResource>
|
||
<mods:relatedItem id="B1CD73B21BBD16FE46372532C3EA7479" type="host">
|
||
<mods:titleInfo id="364E2FC9BEF998FF4E3BF6D7913D6F8F">
|
||
<mods:title id="D500DBD25DC30C2EDC87E239171BB1EB">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="B31D656CD91A109170F10D95370B8D1E">
|
||
<mods:date id="1E185C26EF774C8724BD02E1045799BA">2023</mods:date>
|
||
<mods:detail id="E69B5AC06CE579ADB63EA7C8F95BA11E" type="pubDate">
|
||
<mods:number id="E23EE53018EF0BC9553AEF6BCBD4A25E">2023-05-18</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="CE9A70C275CE57B050AAB6ED96A40CD5" type="volume">
|
||
<mods:number id="DE5EDCF7BC51334D78C9A2F8ABC2521A">5286</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="CF731CF3A06F4337C285A1BD4541A07A" type="issue">
|
||
<mods:number id="8778329C52748D757BBBEA2678A5EE30">1</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="DA06E5BEE134746B29DDEE3343F27CE4" unit="page">
|
||
<mods:start id="4A92D2D199CAF00DFEEABBBF4834A44D">1</mods:start>
|
||
<mods:end id="51DF863841BDB6C1AACFB423C95E01E9">144</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location id="ED4C5A1B108F0AA931A05876D7C52327">
|
||
<mods:url id="047DB62A1E170B2C594025631B9E5E7F">http://dx.doi.org/10.11646/zootaxa.5286.1.1</mods:url>
|
||
</mods:location>
|
||
<mods:classification id="801C21EC9334BDF233631EC29732D74F">journal article</mods:classification>
|
||
<mods:identifier id="C2C053744AA47DC442FFB88ED8C39780" type="CLB-Dataset">263342</mods:identifier>
|
||
<mods:identifier id="932B058EB81920E4BEA33FC581474138" type="DOI">10.11646/zootaxa.5286.1.1</mods:identifier>
|
||
<mods:identifier id="6D76E9C6DBDAF3E27B3FC358B96E1A31" type="GBIF-Dataset">67e7a105-1f62-414a-83d9-f758039eb726</mods:identifier>
|
||
<mods:identifier id="9ED87BE5C18399051D4EC2482AB41971" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="8615D5AC88B9F7B24881917556592F72" type="Zenodo-Dep">7958937</mods:identifier>
|
||
<mods:identifier id="D0DCE3063EDAAA90A84780D4B0833810" type="ZooBank">F9834684-24D3-4795-B5EB-77B451DF856D</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03877623625EFFF92DEFB341FA7B5FA2" ID-DOI="http://doi.org/10.5281/zenodo.7963713" ID-GBIF-Taxon="209629033" ID-Zenodo-Dep="7963713" LSID="urn:lsid:plazi:treatment:03877623625EFFF92DEFB341FA7B5FA2" httpUri="http://treatment.plazi.org/id/03877623625EFFF92DEFB341FA7B5FA2" lastPageId="26" lastPageNumber="27" pageId="24" pageNumber="25">
|
||
<subSubSection id="C33494BE625EFFFB2DEFB341FD9B5C0E" box="[151,636,314,341]" pageId="24" pageNumber="25" type="nomenclature">
|
||
<paragraph id="8B91C735625EFFFB2DEFB341FD9B5C0E" blockId="24.[151,636,314,341]" box="[151,636,314,341]" pageId="24" pageNumber="25">
|
||
<emphasis id="B95A1B27625EFFFB2DEFB341FD9B5C0E" bold="true" box="[151,636,314,341]" pageId="24" pageNumber="25">
|
||
Subgenus
|
||
<taxonomicName id="4C2EBCB6625EFFFB2C6FB341FD9B5C0E" ID-CoL="8KWFM" ID-ENA="796493" authority="Chaudoir, 1872" authorityName="Chaudoir" authorityYear="1872" box="[279,636,314,341]" class="Insecta" family="Carabidae" genus="Crossoglossa" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95A1B27625EFFFB2C6FB341FE525C0F" bold="true" box="[279,437,314,341]" italics="true" pageId="24" pageNumber="25">Crossoglossa</emphasis>
|
||
<bibRefCitation id="EFBFBAC4625EFFFB2CC5B341FD9B5C0E" author="Chaudoir, B. M." box="[445,636,314,340]" pageId="24" pageNumber="25" pagination="97 - 204" refId="ref110142" refString="Chaudoir, B. M. (1872) Monographie des Callidides. Annales de Societe Entomologique de Belgique, 16, 97 - 204." type="journal article" year="1872">Chaudoir, 1872</bibRefCitation>
|
||
</taxonomicName>
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33494BE625EFFFB2DEFB304FB465CCD" box="[151,1185,383,407]" pageId="24" pageNumber="25" type="type_taxon">
|
||
<paragraph id="8B91C735625EFFFB2DEFB304FB465CCD" blockId="24.[151,1436,383,468]" box="[151,1185,383,407]" pageId="24" pageNumber="25">
|
||
Type-species:
|
||
<taxonomicName id="4C2EBCB6625EFFFB2C50B304FD565CCD" ID-CoL="ZMF7" authority="Chaudoir, 1872" authorityName="Chaudoir" authorityYear="1872" box="[296,689,383,407]" class="Insecta" family="Carabidae" genus="Crossoglossa" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="testacea">
|
||
<emphasis id="B95A1B27625EFFFB2C50B304FDED5CCC" bold="true" box="[296,522,383,407]" italics="true" pageId="24" pageNumber="25">Crossoglossa testacea</emphasis>
|
||
<bibRefCitation id="EFBFBAC4625EFFFB2F69B304FD565CCD" author="Chaudoir, B. M." box="[529,689,383,407]" pageId="24" pageNumber="25" pagination="97 - 204" refId="ref110142" refString="Chaudoir, B. M. (1872) Monographie des Callidides. Annales de Societe Entomologique de Belgique, 16, 97 - 204." type="journal article" year="1872">Chaudoir, 1872</bibRefCitation>
|
||
</taxonomicName>
|
||
. Subsequently designated by
|
||
<bibRefCitation id="EFBFBAC4625EFFFB2EA3B304FB7A5CCD" author="Jeannel, R." box="[987,1181,383,407]" pageId="24" pageNumber="25" pagination="767 - 1146" refId="ref111235" refString="Jeannel, R. (1949) Faune de l'Empire francais. XI. Coleopteres carabiques de la region malgache (troisieme partie). Paris: Librairie Larose, 767 - 1146." type="book chapter" year="1949">Jeannel, 1949: 971</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33494BE625EFFFB2DEFB3E6FB5E5C8E" pageId="24" pageNumber="25" type="reference_group">
|
||
<paragraph id="8B91C735625EFFFB2DEFB3E6FB5E5C8E" blockId="24.[151,1436,383,468]" pageId="24" pageNumber="25">
|
||
<taxonomicName id="4C2EBCB6625EFFFB2DEFB3E6FE1C5CEC" ID-CoL="8KWFM" authority="Chaudoir, 1872: 177" authorityName="Chaudoir" authorityPageNumber="177" authorityYear="1872" box="[151,507,413,438]" class="Insecta" family="Carabidae" genus="Crossoglossa" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95A1B27625EFFFB2DEFB3E6FEC75CEF" bold="true" box="[151,288,413,437]" italics="true" pageId="24" pageNumber="25">Crossoglossa</emphasis>
|
||
<bibRefCitation id="EFBFBAC4625EFFFB2C5FB3E5FE1C5CEC" author="Chaudoir, B. M." box="[295,507,414,438]" pageId="24" pageNumber="25" pagination="97 - 204" refId="ref110142" refString="Chaudoir, B. M. (1872) Monographie des Callidides. Annales de Societe Entomologique de Belgique, 16, 97 - 204." type="journal article" year="1872">Chaudoir, 1872: 177</bibRefCitation>
|
||
</taxonomicName>
|
||
;
|
||
<bibRefCitation id="EFBFBAC4625EFFFB2F7EB3E5FD305CEF" author="Chaudoir, B. M." box="[518,727,413,438]" pageId="24" pageNumber="25" pagination="145 - 161" refId="ref110207" refString="Chaudoir, B. M. (1878) Enumeration des cincindeletes et des carabiques recueillis par M. A. Raffray, dans les iles de Zanzibar et de Pemba, ainsi qu'a Bagamoyo, Mombaze et sur les montagnes de Schimba, avec la description des especes nouvelles. 2. Revue et Magasin de Zoologie Pure et Appliquee, (3) 6, 145 - 161." type="journal article" year="1878">Chaudoir, 1878: 151</bibRefCitation>
|
||
(misspelled as
|
||
<taxonomicName id="4C2EBCB6625EFFFB2E0DB3E5FC005CEC" ID-CoL="8LRC5" authority="Chaudoir, 1878" box="[885,999,414,438]" class="Insecta" family="Carabidae" genus="Ceroglossa" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95A1B27625EFFFB2E0DB3E5FC005CEC" box="[885,999,414,438]" italics="true" pageId="24" pageNumber="25">Ceroglossa</emphasis>
|
||
</taxonomicName>
|
||
);
|
||
<bibRefCitation id="EFBFBAC4625EFFFB2E86B3E5FB345CEF" author="Jakobson, G. G." box="[1022,1235,413,438]" pageId="24" pageNumber="25" pagination="321 - 400" refId="ref111151" refString="Jakobson, G. G. (1907) Fasc. 5: pp. 321 - 400. In: Zhuki Rossii i Zapadnoi Evropy. Sankt-Petersburg: A. F. Devrien, 1024 pp., 83 pls." type="book chapter" year="1907">Jakobson, 1907: 395</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFBFBAC4625EFFFB29A4B3E5FA7F5CEF" author="Alluaud, C." box="[1244,1432,413,438]" pageId="24" pageNumber="25" pagination="85 - 88" refId="ref109310" refString="Alluaud, C. (1917) Contributions a l'etude des Carabiques d'Afrique et de Madagascar [Col.] XIV- Sur le genre Crossoglossa Chaudoir et descriptions de duex especes nouvelles. Bulletin de la Societe Entomologique de France, 1917, 85 - 88. https: // doi. org / 10.3406 / bsef. 1917.26042" type="journal article" year="1917">Alluaud, 1917: 85</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFBFBAC4625EFFFB2DBFB3C7FE465C8E" author="Andrewes, H. E." box="[199,417,444,468]" pageId="24" pageNumber="25" pagination="469 - 483" refId="ref109369" refString="Andrewes, H. E. (1919) Papers on Oriental Carabidae. - I. The Annals and Magazine of Natural History, (9) 3, 469 - 483. https: // doi. org / 10.1080 / 00222931908673846" type="journal article" year="1919">Andrewes, 1919: 483</bibRefCitation>
|
||
(synonymized with
|
||
<taxonomicName id="4C2EBCB6625EFFFB2F08B3C7FCED5C8E" ID-CoL="8LSMH" authorityName="Macleay" authorityYear="1871" box="[624,778,444,468]" class="Insecta" family="Carabidae" genus="Phloeodromius" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95A1B27625EFFFB2F08B3C7FCED5C8E" box="[624,778,444,468]" italics="true" pageId="24" pageNumber="25">Phloeodromius</emphasis>
|
||
</taxonomicName>
|
||
);
|
||
<bibRefCitation id="EFBFBAC4625EFFFB2E58B3C7FC055C8E" author="Jeannel, R." box="[800,994,444,468]" pageId="24" pageNumber="25" pagination="767 - 1146" refId="ref111235" refString="Jeannel, R. (1949) Faune de l'Empire francais. XI. Coleopteres carabiques de la region malgache (troisieme partie). Paris: Librairie Larose, 767 - 1146." type="book chapter" year="1949">Jeannel, 1949: 972</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFBFBAC4625EFFFB2E94B3C7FB525C8E" author="Jedlicka, A." box="[1004,1205,444,468]" pageId="24" pageNumber="25" pagination="207 - 211" refId="ref111420" refString="Jedlicka, A. (1952) Les Carabides nouveaux de la zone palaearctique (Col.). Sbornik Entomologickeho Oddeleni Narodniho Musea v Praze, 27 [1951], 207 - 211." type="journal article" year="1952">Jedlička, 1952: 210</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33494BE625EFFFB2DEFB079FA7A59FF" pageId="24" pageNumber="25" type="description">
|
||
<paragraph id="8B91C735625EFFFB2DEFB079FE775FAF" blockId="24.[151,1437,514,1801]" pageId="24" pageNumber="25">
|
||
<taxonomicName id="4C2EBCB6625EFFFB2DEFB079FECA5F46" authority="Chaudoir, 1873" box="[151,301,514,540]" class="Insecta" family="Carabidae" genus="Crossoglossa" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95A1B27625EFFFB2DEFB079FECA5F46" box="[151,301,514,540]" italics="true" pageId="24" pageNumber="25">Crossoglossa</emphasis>
|
||
</taxonomicName>
|
||
was erected by
|
||
<bibRefCitation id="EFBFBAC4625EFFFB2C94B079FD4E5F47" author="Chaudoir, B. M." box="[492,681,514,541]" pageId="24" pageNumber="25" pagination="97 - 204" refId="ref110142" refString="Chaudoir, B. M. (1872) Monographie des Callidides. Annales de Societe Entomologique de Belgique, 16, 97 - 204." type="journal article" year="1872">Chaudoir (1872)</bibRefCitation>
|
||
to accommodate his three species without considering the earlier name
|
||
<taxonomicName id="4C2EBCB6625EFFFB2DA2B05CFDEF5F1A" authority="Motschulsky, 1860" authorityName="Motschulsky" authorityYear="1860" box="[218,520,550,576]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95A1B27625EFFFB2DA2B05CFECC5F1A" box="[218,299,551,576]" italics="true" pageId="24" pageNumber="25">Parena</emphasis>
|
||
<bibRefCitation id="EFBFBAC4625EFFFB2C49B05DFDEF5F1A" author="Motschulsky, V." box="[305,520,550,576]" pageId="24" pageNumber="25" pagination="25 - 118" refId="ref112373" refString="Motschulsky, V. (1860) Entomologie speciale. Insectes des Indes orientales, et de contrees analogues. 2: de serie. Etudes Entomologiques, 8 [1859], 25 - 118." type="journal article" year="1860">Motschulsky, 1860</bibRefCitation>
|
||
</taxonomicName>
|
||
. The name
|
||
<taxonomicName id="4C2EBCB6625EFFFB2FFDB05DFCFC5F1A" authority="Chaudoir, 1873" box="[645,795,550,576]" class="Insecta" family="Carabidae" genus="Crossoglossa" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95A1B27625EFFFB2FFDB05DFCFC5F1A" box="[645,795,550,576]" italics="true" pageId="24" pageNumber="25">Crossoglossa</emphasis>
|
||
</taxonomicName>
|
||
was then widely used, and several species were described under that name (e.g.,
|
||
<bibRefCitation id="EFBFBAC4625EFFFB2CE0B030FDC75F3F" author="Bates, H. W." box="[408,544,587,613]" pageId="24" pageNumber="25" pagination="229 - 322" refId="ref109990" refString="Bates, H. W. (1873) On the Geodephagous Coleoptera of Japan. Fam. Carabidae. The Transactions of the Entomological Society of London, 1873 (2), 229 - 322." type="journal article" year="1873">Bates, 1873</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFBFBAC4625EFFFB2F54B031FD365F3F" author="Alluaud, C." box="[556,721,586,613]" pageId="24" pageNumber="25" pagination="85 - 88" refId="ref109310" refString="Alluaud, C. (1917) Contributions a l'etude des Carabiques d'Afrique et de Madagascar [Col.] XIV- Sur le genre Crossoglossa Chaudoir et descriptions de duex especes nouvelles. Bulletin de la Societe Entomologique de France, 1917, 85 - 88. https: // doi. org / 10.3406 / bsef. 1917.26042" type="journal article" year="1917">Alluaud, 1917</bibRefCitation>
|
||
). However, the
|
||
<typeStatus id="54957997625EFFFB2EFDB030FC525F3F" box="[901,949,587,613]" pageId="24" pageNumber="25">type</typeStatus>
|
||
species of
|
||
<taxonomicName id="4C2EBCB6625EFFFB294FB031FB2A5F3E" authority="Chaudoir, 1873" box="[1079,1229,586,612]" class="Insecta" family="Carabidae" genus="Crossoglossa" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95A1B27625EFFFB294FB031FB2A5F3E" box="[1079,1229,586,612]" italics="true" pageId="24" pageNumber="25">Crossoglossa</emphasis>
|
||
</taxonomicName>
|
||
was not fixed until
|
||
<bibRefCitation id="EFBFBAC4625EFFFB2DC0B015FE865FD3" author="Jeannel, R." box="[184,353,622,649]" pageId="24" pageNumber="25" pagination="767 - 1146" refId="ref111235" refString="Jeannel, R. (1949) Faune de l'Empire francais. XI. Coleopteres carabiques de la region malgache (troisieme partie). Paris: Librairie Larose, 767 - 1146." type="book chapter" year="1949">Jeannel (1949)</bibRefCitation>
|
||
designated
|
||
<taxonomicName id="4C2EBCB6625EFFFB2C94B015FCBD5FD2" authority="Chaudoir" authorityName="Chaudoir" authorityYear="1872" box="[492,858,622,648]" class="Insecta" family="Carabidae" genus="Crossoglossa" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="testacea">
|
||
<emphasis id="B95A1B27625EFFFB2C94B015FD0F5FD2" box="[492,744,622,648]" italics="true" pageId="24" pageNumber="25">Crossoglossa testacea</emphasis>
|
||
Chaudoir
|
||
</taxonomicName>
|
||
for it. The concept of
|
||
<taxonomicName id="4C2EBCB6625EFFFB291EB015FB1B5FD2" authority="Chaudoir, 1873" box="[1126,1276,622,648]" class="Insecta" family="Carabidae" genus="Crossoglossa" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95A1B27625EFFFB291EB015FB1B5FD2" box="[1126,1276,622,648]" italics="true" pageId="24" pageNumber="25">Crossoglossa</emphasis>
|
||
</taxonomicName>
|
||
(sensu
|
||
<bibRefCitation id="EFBFBAC4625EFFFB282FB014FF355FF7" author="Bates, H. W." pageId="24" pageNumber="25" pagination="229 - 322" refId="ref109990" refString="Bates, H. W. (1873) On the Geodephagous Coleoptera of Japan. Fam. Carabidae. The Transactions of the Entomological Society of London, 1873 (2), 229 - 322." type="journal article" year="1873">Bates, 1873</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFBFBAC4625EFFFB2DA4B0E9FE995FF7" author="Alluaud, C." box="[220,382,658,685]" pageId="24" pageNumber="25" pagination="85 - 88" refId="ref109310" refString="Alluaud, C. (1917) Contributions a l'etude des Carabiques d'Afrique et de Madagascar [Col.] XIV- Sur le genre Crossoglossa Chaudoir et descriptions de duex especes nouvelles. Bulletin de la Societe Entomologique de France, 1917, 85 - 88. https: // doi. org / 10.3406 / bsef. 1917.26042" type="journal article" year="1917">Alluaud, 1917</bibRefCitation>
|
||
) was similar to the genus
|
||
<taxonomicName id="4C2EBCB6625EFFFB2FDBB0E8FD135FF6" box="[675,756,659,684]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95A1B27625EFFFB2FDBB0E8FD135FF6" box="[675,756,659,684]" italics="true" pageId="24" pageNumber="25">Parena</emphasis>
|
||
</taxonomicName>
|
||
at present until
|
||
<bibRefCitation id="EFBFBAC4625EFFFB2EDFB0E9FB835FF7" author="Andrewes, H. E." box="[935,1124,658,685]" pageId="24" pageNumber="25" pagination="469 - 483" refId="ref109369" refString="Andrewes, H. E. (1919) Papers on Oriental Carabidae. - I. The Annals and Magazine of Natural History, (9) 3, 469 - 483. https: // doi. org / 10.1080 / 00222931908673846" type="journal article" year="1919">Andrewes (1919</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFBFBAC4625EFFFB2917B0E8FB4F5FF7" author="Andrewes, H. E." box="[1135,1192,659,685]" pageId="24" pageNumber="25" pagination="1 - 24" refId="ref109566" refString="Andrewes, H. E. (1928) On the types of Oriental Carabidae described by V. de Motschulsky. The Transactions of the Entomological Society of London, 76, 1 - 24. https: // doi. org / 10.1111 / j. 1365 - 2311.1928. tb 01185. x" type="journal article" year="1928">1928</bibRefCitation>
|
||
) synonymized it with
|
||
<taxonomicName id="4C2EBCB6625EFFFB2DEFB0CDFEA65F8A" authorityName="Macleay" authorityYear="1871" box="[151,321,694,720]" class="Insecta" family="Carabidae" genus="Phloeodromius" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95A1B27625EFFFB2DEFB0CDFEA65F8A" box="[151,321,694,720]" italics="true" pageId="24" pageNumber="25">Phloeodromius</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C2EBCB6625EFFFB2C00B0CCFE2E5F8A" box="[376,457,695,720]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95A1B27625EFFFB2C00B0CCFE2E5F8A" box="[376,457,695,720]" italics="true" pageId="24" pageNumber="25">Parena</emphasis>
|
||
</taxonomicName>
|
||
. Thereafter,
|
||
<taxonomicName id="4C2EBCB6625EFFFB2F21B0CDFD085F8A" authority="Chaudoir, 1873" box="[601,751,694,720]" class="Insecta" family="Carabidae" genus="Crossoglossa" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95A1B27625EFFFB2F21B0CDFD085F8A" box="[601,751,694,720]" italics="true" pageId="24" pageNumber="25">Crossoglossa</emphasis>
|
||
</taxonomicName>
|
||
was used as a subgenus name in different concepts (
|
||
<bibRefCitation id="EFBFBAC4625EFFFB2838B0CDFF355FAF" author="Jeannel, R." pageId="24" pageNumber="25" pagination="767 - 1146" refId="ref111235" refString="Jeannel, R. (1949) Faune de l'Empire francais. XI. Coleopteres carabiques de la region malgache (troisieme partie). Paris: Librairie Larose, 767 - 1146." type="book chapter" year="1949">Jeannel, 1949</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFBFBAC4625EFFFB2DA6B0A1FE635FAF" author="Jedlicka, A." box="[222,388,730,757]" pageId="24" pageNumber="25" pagination="207 - 211" refId="ref111420" refString="Jedlicka, A. (1952) Les Carabides nouveaux de la zone palaearctique (Col.). Sbornik Entomologickeho Oddeleni Narodniho Musea v Praze, 27 [1951], 207 - 211." type="journal article" year="1952">Jedlička, 1952</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B91C735625EFFFB2DBFB084FACF5EF2" blockId="24.[151,1437,514,1801]" pageId="24" pageNumber="25">
|
||
In the present study,
|
||
<taxonomicName id="4C2EBCB6625EFFFB2CC8B085FDA15E42" authority="Chaudoir, 1873" box="[432,582,766,792]" class="Insecta" family="Carabidae" genus="Crossoglossa" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95A1B27625EFFFB2CC8B085FDA15E42" box="[432,582,766,792]" italics="true" pageId="24" pageNumber="25">Crossoglossa</emphasis>
|
||
</taxonomicName>
|
||
is treated as a subgenus of
|
||
<taxonomicName id="4C2EBCB6625EFFFB2E01B084FC2D5E42" box="[889,970,767,792]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95A1B27625EFFFB2E01B084FC2D5E42" box="[889,970,767,792]" italics="true" pageId="24" pageNumber="25">Parena</emphasis>
|
||
</taxonomicName>
|
||
to accommodate species related to
|
||
<taxonomicName id="4C2EBCB6625EFFFB2820B084FF335E66" authorityName="Chaudoir" authorityYear="1872" class="Insecta" family="Carabidae" genus="Crossoglossa" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="testacea">
|
||
<emphasis id="B95A1B27625EFFFB2820B084FF335E66" italics="true" pageId="24" pageNumber="25">P. testacea</emphasis>
|
||
</taxonomicName>
|
||
, which are different from members of the other two subgenera of the genus in the following unique character states: (1) pronotum much wider (PW/PL 1.50–1.82) than other species of the genus; (2) mentum without setae, lateral lobes extremely large, inner margin nearly straight, epilobes very narrow (
|
||
<figureCitation id="1315DBB0625EFFFB2959B111FB9C5EDF" box="[1057,1147,874,901]" captionStart="FIGURE 3" captionStartId="10.[151,250,881,905]" captionTargetBox="[161,1430,189,857]" captionTargetId="figure-28@10.[151,1436,181,857]" captionTargetPageId="10" captionText="FIGURE 3. A–D Labia of Parena spp.: A P. (Crossoglossa) cavipennis (Bates), Beijing. B P. (Parena) latecincta (Bates), Zhejiang. C P. (Bothynoptera) dorsigera (Schaum), Guangxi. D P. (Bothynoptera) tripunctata (Bates), Beijing. E–F Heads of Parena spp. in lateral view: E P. (Bothynoptera) taiwana Hua, Taiwan, suborbital setae present. F P. (Parena) latecincta (Bates), Yunnan, suborbital seta absent. Scale bars = 0.5mm." figureDoi="http://doi.org/10.5281/zenodo.7958943" httpUri="https://zenodo.org/record/7958943/files/figure.png" pageId="24" pageNumber="25">Fig. 3A</figureCitation>
|
||
); (3) male genitalia rather slender, endophallus with the primary sclerite rudimentary; (4) gonocoxite II of ovipositor dichotomous.
|
||
</paragraph>
|
||
<paragraph id="8B91C735625EFFFB2DBFB1C9FCF25962" blockId="24.[151,1437,514,1801]" pageId="24" pageNumber="25">
|
||
The present concept of the subgenus
|
||
<taxonomicName id="4C2EBCB6625EFFFB2F1EB1C9FD1B5E96" authority="Chaudoir, 1873" box="[614,764,946,972]" class="Insecta" family="Carabidae" genus="Crossoglossa" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95A1B27625EFFFB2F1EB1C9FD1B5E96" box="[614,764,946,972]" italics="true" pageId="24" pageNumber="25">Crossoglossa</emphasis>
|
||
</taxonomicName>
|
||
is similar to that of
|
||
<bibRefCitation id="EFBFBAC4625EFFFB2EA4B1C9FB6B5E97" author="Jedlicka, A." box="[988,1164,946,973]" pageId="24" pageNumber="25" pagination="207 - 211" refId="ref111420" refString="Jedlicka, A. (1952) Les Carabides nouveaux de la zone palaearctique (Col.). Sbornik Entomologickeho Oddeleni Narodniho Musea v Praze, 27 [1951], 207 - 211." type="journal article" year="1952">Jedlička (1952)</bibRefCitation>
|
||
, containing nine species distributed through
|
||
<collectingCountry id="F33987A5625EFFFB2C0CB1ACFE075EAA" box="[372,480,982,1009]" name="China" pageId="24" pageNumber="25">East Asia</collectingCountry>
|
||
, all parts of the Oriental Realm,
|
||
<collectingCountry id="F33987A5625EFFFB2E36B1ACFBCD5EAA" box="[846,1066,982,1009]" name="Papua New Guinea" pageId="24" pageNumber="25">Papua New Guinea</collectingCountry>
|
||
, and easternmost to the
|
||
<collectingCountry id="F33987A5625EFFFB284FB1ADFF0D594E" name="Solomon Islands" pageId="24" pageNumber="25">Solomon Islands</collectingCountry>
|
||
. They are not known from Africa or
|
||
<collectingCountry id="F33987A5625EFFFB2FF8B181FD0D594E" box="[640,746,1018,1044]" name="Australia" pageId="24" pageNumber="25">Australia</collectingCountry>
|
||
. Three species groups are recognized in the subgenus, each of them containing one or several strictly allopatric species.
|
||
</paragraph>
|
||
<paragraph id="8B91C735625EFFFB2DBFB639FA7A59FF" blockId="24.[151,1437,514,1801]" pageId="24" pageNumber="25">
|
||
The exceptional male and female genital characters suggest
|
||
<taxonomicName id="4C2EBCB6625EFFFB2E08B639FBE15906" authority="Chaudoir, 1873" box="[880,1030,1090,1116]" class="Insecta" family="Carabidae" genus="Crossoglossa" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95A1B27625EFFFB2E08B639FBE15906" box="[880,1030,1090,1116]" italics="true" pageId="24" pageNumber="25">Crossoglossa</emphasis>
|
||
</taxonomicName>
|
||
is monophyletic. However, its relationship in the genus
|
||
<taxonomicName id="4C2EBCB6625EFFFB2CFFB61CFE3F59DA" box="[391,472,1127,1152]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95A1B27625EFFFB2CFFB61CFE3F59DA" box="[391,472,1127,1152]" italics="true" pageId="24" pageNumber="25">Parena</emphasis>
|
||
</taxonomicName>
|
||
is difficult to infer before a comprehensive phylogenetic analysis is conducted. Based on the modified male and female genitalia,
|
||
<taxonomicName id="4C2EBCB6625EFFFB2F0FB6F1FCEA59FE" authority="Chaudoir, 1873" box="[631,781,1162,1188]" class="Insecta" family="Carabidae" genus="Crossoglossa" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95A1B27625EFFFB2F0FB6F1FCEA59FE" box="[631,781,1162,1188]" italics="true" pageId="24" pageNumber="25">Crossoglossa</emphasis>
|
||
</taxonomicName>
|
||
is considered to be a highly specialized group in the genus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33494BE625EFFFB2DBFB6D5FD5E5A52" pageId="24" pageNumber="25" type="diagnosis">
|
||
<paragraph id="8B91C735625EFFFB2DBFB6D5FD5E5A52" blockId="24.[151,1437,514,1801]" pageId="24" pageNumber="25">
|
||
<emphasis id="B95A1B27625EFFFB2DBFB6D5FE325993" bold="true" box="[199,469,1198,1225]" pageId="24" pageNumber="25">Diagnostic characters.</emphasis>
|
||
Dorsal surface generally unicolor (except
|
||
<taxonomicName id="4C2EBCB6625EFFFB2EBEB6D4FBD45992" authority="Shi & Liang, 2023" authorityName="Shi & Liang" authorityYear="2023" box="[966,1075,1198,1224]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="sciakyi" status="sp. nov.">
|
||
<emphasis id="B95A1B27625EFFFB2EBEB6D4FBD45992" box="[966,1075,1198,1224]" italics="true" pageId="24" pageNumber="25">P. sciakyi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95A1B27625EFFFB2945B6D4FB675993" bold="true" box="[1085,1152,1199,1225]" pageId="24" pageNumber="25">
|
||
<taxonomicNameLabel id="A269A65C625EFFFB2945B6D4FB675993" box="[1085,1152,1199,1225]" pageId="24" pageNumber="25" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
), pale yellow to reddish brown. Tempora very short, abruptly narrowed behind eyes, length of tempora plus neck-constriction approximately one-third of diameter of the eye; accessory setae on antennomeres 1–3 absent or very short and sparse; postgenae with a pair of suborbital setae; mandibles extremely widened, semicircular, retinacular ridge of right mandible reduced to an elongate retinacular tooth; mentum without a pair of setae, lateral lobes extremely large, inner margins nearly straight, outer and apical margins widely arcuate, epilobes very narrow, apex much shorter than lateral lobes. Pronotum strongly transverse, PW/PL 1.50–1.82, generally greater than 1.60; much wider than head, PW/HW = 1.05–1.26, generally greater than 1.10; lateral explanations very wide; lateral margins usually slightly sinuate before posterior angles; posterior angles obtuse-rounded. Elytral interval 3 with three setigerous pores, first one very close to elytra base, near level of scutellar apex; disc with large depressions near middle of intervals 3 to 6 (except for
|
||
<taxonomicName id="4C2EBCB6625EFFFB28F0B788FF3C5B6A" authorityName="Andrewes, Borneo. D" authorityYear="1931" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="levata">
|
||
<emphasis id="B95A1B27625EFFFB28F0B788FF3C5B6A" italics="true" pageId="24" pageNumber="25">P. levata</emphasis>
|
||
</taxonomicName>
|
||
), lateral sides depressed near anterior third. Apex of abdominal sternite VII with two or three setae on each side, straight or slightly notched in males, straight or projected at middle in females. Male mesotarsi with biseriate ventral adhesive setae on basal three tarsomeres. Male genitalia with median lobe slender, AL/AW 5.4–6.8; apical lamella thin and wide, apical orifice opened to left; endophallus with primary sclerite reduced to a small triangular piece near base, apical sclerite ribbon-form, squamate sheath large and heavily scaled, squamate sac simple or divided. Female ovipositor with dichotomous gonocoxite II, inner branch with two ensiform setae apically, outer branch with two or three ensiform setae apically.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF5197BD625FFFFA2DEFB44BFC455B82" ID-DOI="http://doi.org/10.5281/zenodo.7958963" ID-Zenodo-Dep="7958963" httpUri="https://zenodo.org/record/7958963/files/figure.png" pageId="25" pageNumber="26" startId="25.[151,250,1584,1608]" targetBox="[151,1436,181,1561]" targetPageId="25" targetType="figure">
|
||
<paragraph id="8B91C735625FFFFA2DEFB44BFC455B82" blockId="25.[151,1436,1584,1753]" pageId="25" pageNumber="26">
|
||
<emphasis id="B95A1B27625FFFFA2DEFB44BFEFD5B12" bold="true" box="[151,282,1584,1608]" pageId="25" pageNumber="26">FIGURE 14</emphasis>
|
||
. Elytra apex of
|
||
<taxonomicName id="4C2EBCB6625FFFFA2CB9B44AFD4D5B13" box="[449,682,1584,1609]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="subGenus" subGenus="Crossoglossa">
|
||
<emphasis id="B95A1B27625FFFFA2CB9B44AFDEC5B12" box="[449,523,1585,1608]" italics="true" pageId="25" pageNumber="26">Parena</emphasis>
|
||
(
|
||
<emphasis id="B95A1B27625FFFFA2F62B44BFD455B12" box="[538,674,1584,1608]" italics="true" pageId="25" pageNumber="26">Crossoglossa</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
spp.
|
||
<emphasis id="B95A1B27625FFFFA2F9BB44BFCF25B12" bold="true" box="[739,789,1584,1608]" pageId="25" pageNumber="26">A–H</emphasis>
|
||
Elytra sutural angle (left), scale bar = 0.2mm.
|
||
<emphasis id="B95A1B27625FFFFA2981B44BFAEC5B12" bold="true" box="[1273,1291,1584,1608]" pageId="25" pageNumber="26">A</emphasis>
|
||
<taxonomicName id="4C2EBCB6625FFFFA286BB44AFF055B37" authority="(Bates)" baseAuthorityName="Bates" baseAuthorityYear="1873" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="cavipennis">
|
||
<emphasis id="B95A1B27625FFFFA286BB44AFAC55B12" box="[1299,1314,1585,1608]" italics="true" pageId="25" pageNumber="26">P</emphasis>
|
||
.
|
||
<emphasis id="B95A1B27625FFFFA2848B44AFA7B5B12" box="[1328,1436,1585,1608]" italics="true" pageId="25" pageNumber="26">cavipennis</emphasis>
|
||
(Bates)
|
||
</taxonomicName>
|
||
, Shaanxi.
|
||
<emphasis id="B95A1B27625FFFFA2C35B42EFEB95B37" bold="true" box="[333,350,1621,1645]" pageId="25" pageNumber="26">B</emphasis>
|
||
<taxonomicName id="4C2EBCB6625FFFFA2C1DB42EFDA15B37" authority="(Bates)" baseAuthorityName="Bates" baseAuthorityYear="1873" box="[357,582,1620,1645]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="laesipennis">
|
||
<emphasis id="B95A1B27625FFFFA2C1DB42EFE935B36" box="[357,372,1621,1644]" italics="true" pageId="25" pageNumber="26">P</emphasis>
|
||
.
|
||
<emphasis id="B95A1B27625FFFFA2CFAB42FFE135B36" box="[386,500,1620,1644]" italics="true" pageId="25" pageNumber="26">laesipennis</emphasis>
|
||
(Bates)
|
||
</taxonomicName>
|
||
, Vietnam.
|
||
<emphasis id="B95A1B27625FFFFA2FCCB42FFD215B36" bold="true" box="[692,710,1620,1644]" pageId="25" pageNumber="26">C</emphasis>
|
||
<taxonomicName id="4C2EBCB6625FFFFA2FB5B42EFBEA5B37" authority="Andrewes, Borneo. D" authorityName="Andrewes, Borneo. D" authorityYear="1931" box="[717,1037,1620,1645]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="levata">
|
||
<emphasis id="B95A1B27625FFFFA2FB5B42EFD3B5B36" box="[717,732,1621,1644]" italics="true" pageId="25" pageNumber="26">P</emphasis>
|
||
.
|
||
<emphasis id="B95A1B27625FFFFA2F92B42FFCCF5B36" box="[746,808,1620,1644]" italics="true" pageId="25" pageNumber="26">levata</emphasis>
|
||
Andrewes, Borneo.
|
||
<emphasis id="B95A1B27625FFFFA2E83B42EFBEA5B37" bold="true" box="[1019,1037,1621,1645]" pageId="25" pageNumber="26">D</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicName id="4C2EBCB6625FFFFA296CB42EFB1A5B36" authority="(Chaudoir)" baseAuthorityName="Chaudoir" box="[1044,1277,1620,1644]" class="Insecta" family="Carabidae" genus="Crossoglossa" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="testacea">
|
||
<emphasis id="B95A1B27625FFFFA296CB42EFBC45B36" box="[1044,1059,1621,1644]" italics="true" pageId="25" pageNumber="26">P</emphasis>
|
||
.
|
||
<emphasis id="B95A1B27625FFFFA2949B42EFB635B36" box="[1073,1156,1621,1644]" italics="true" pageId="25" pageNumber="26">testacea</emphasis>
|
||
(Chaudoir)
|
||
</taxonomicName>
|
||
, Guangdong.
|
||
<emphasis id="B95A1B27625FFFFA28F3B42EFA7B5B37" bold="true" box="[1419,1436,1621,1645]" pageId="25" pageNumber="26">E</emphasis>
|
||
<taxonomicName id="4C2EBCB6625FFFFA2DEFB402FE325BCA" authority="Kirschenhofer" authorityName="Kirschenhofer. D" authorityYear="2006" box="[151,469,1656,1680]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="sulawesiensis">
|
||
<emphasis id="B95A1B27625FFFFA2DEFB402FF415BCA" box="[151,166,1657,1680]" italics="true" pageId="25" pageNumber="26">P</emphasis>
|
||
.
|
||
<emphasis id="B95A1B27625FFFFA2DCAB403FEDB5BCA" box="[178,316,1656,1680]" italics="true" pageId="25" pageNumber="26">sulawesiensis</emphasis>
|
||
Kirschenhofer
|
||
</taxonomicName>
|
||
, Sulawesi.
|
||
<emphasis id="B95A1B27625FFFFA2F3DB402FDB35BCB" bold="true" box="[581,596,1657,1681]" pageId="25" pageNumber="26">F</emphasis>
|
||
<taxonomicName id="4C2EBCB6625FFFFA2F22B402FC415BCA" authority="Andrewes, Borneo. G" authorityName="Andrewes, Borneo. G" authorityYear="1935" box="[602,934,1656,1680]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="cruralis">
|
||
<emphasis id="B95A1B27625FFFFA2F22B402FD8E5BCA" box="[602,617,1657,1680]" italics="true" pageId="25" pageNumber="26">P</emphasis>
|
||
.
|
||
<emphasis id="B95A1B27625FFFFA2F0DB403FD225BCA" box="[629,709,1656,1680]" italics="true" pageId="25" pageNumber="26">cruralis</emphasis>
|
||
Andrewes, Borneo.
|
||
<emphasis id="B95A1B27625FFFFA2EEAB403FC415BCA" bold="true" box="[914,934,1656,1680]" pageId="25" pageNumber="26">G</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicName id="4C2EBCB6625FFFFA2ED3B402FB665BCA" authority="(Chaudoir)" baseAuthorityName="Chaudoir" baseAuthorityYear="1872" box="[939,1153,1656,1680]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="mellea">
|
||
<emphasis id="B95A1B27625FFFFA2ED3B402FC5D5BCA" box="[939,954,1657,1680]" italics="true" pageId="25" pageNumber="26">P</emphasis>
|
||
.
|
||
<emphasis id="B95A1B27625FFFFA2EBEB403FBED5BCA" box="[966,1034,1656,1680]" italics="true" pageId="25" pageNumber="26">mellea</emphasis>
|
||
(Chaudoir)
|
||
</taxonomicName>
|
||
, Philippine.
|
||
<emphasis id="B95A1B27625FFFFA2986B402FAF55BCB" bold="true" box="[1278,1298,1657,1681]" pageId="25" pageNumber="26">H</emphasis>
|
||
<taxonomicName id="4C2EBCB6625FFFFA286FB402FA9F5BCA" authority="Shi & Liang, 2023" authorityName="Shi & Liang" authorityYear="2023" box="[1303,1400,1656,1680]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="sciakyi" status="sp. nov.">
|
||
<emphasis id="B95A1B27625FFFFA286FB402FAC15BCA" box="[1303,1318,1657,1680]" italics="true" pageId="25" pageNumber="26">P</emphasis>
|
||
.
|
||
<emphasis id="B95A1B27625FFFFA284AB403FA9F5BCA" box="[1330,1400,1656,1680]" italics="true" pageId="25" pageNumber="26">sciakyi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95A1B27625FFFFA2806B402FF4B5BEF" bold="true" pageId="25" pageNumber="26">
|
||
<taxonomicNameLabel id="A269A65C625FFFFA2806B402FF4B5BEF" pageId="25" pageNumber="26" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype.
|
||
<emphasis id="B95A1B27625FFFFA2C66B4E6FEA25BEF" bold="true" box="[286,325,1693,1717]" pageId="25" pageNumber="26">I–L</emphasis>
|
||
Elytra outer apical angle (left), scale bar = 0.5mm.
|
||
<emphasis id="B95A1B27625FFFFA2E2AB4E6FCBB5BEF" bold="true" box="[850,860,1693,1717]" pageId="25" pageNumber="26">I</emphasis>
|
||
<taxonomicName id="4C2EBCB6625FFFFA2E1AB4E6FBDB5BEF" authority="(Bates)" baseAuthorityName="Bates" baseAuthorityYear="1873" box="[866,1084,1693,1717]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="cavipennis">
|
||
<emphasis id="B95A1B27625FFFFA2E1AB4E6FC965BEE" box="[866,881,1693,1716]" italics="true" pageId="25" pageNumber="26">P</emphasis>
|
||
.
|
||
<emphasis id="B95A1B27625FFFFA2E06B4E6FC0D5BEE" box="[894,1002,1693,1716]" italics="true" pageId="25" pageNumber="26">cavipennis</emphasis>
|
||
(Bates)
|
||
</taxonomicName>
|
||
, Shaanxi.
|
||
<emphasis id="B95A1B27625FFFFA29DDB4E6FB555BEF" bold="true" box="[1189,1202,1693,1717]" pageId="25" pageNumber="26">J</emphasis>
|
||
<taxonomicName id="4C2EBCB6625FFFFA29C0B4E6FA7F5BEF" authority="(Bates)" baseAuthorityName="Bates" baseAuthorityYear="1873" box="[1208,1432,1692,1717]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="laesipennis">
|
||
<emphasis id="B95A1B27625FFFFA29C0B4E6FB205BEE" box="[1208,1223,1693,1716]" italics="true" pageId="25" pageNumber="26">P</emphasis>
|
||
.
|
||
<emphasis id="B95A1B27625FFFFA29ACB4E7FAA15BEE" box="[1236,1350,1692,1716]" italics="true" pageId="25" pageNumber="26">laesipennis</emphasis>
|
||
(Bates)
|
||
</taxonomicName>
|
||
, Taiwan.
|
||
<emphasis id="B95A1B27625FFFFA2D96B4BAFEE55B83" bold="true" box="[238,258,1729,1753]" pageId="25" pageNumber="26">K</emphasis>
|
||
<taxonomicName id="4C2EBCB6625FFFFA2C70B4BAFE8D5B82" authority="Shi & Liang, 2023" authorityName="Shi & Liang" authorityYear="2023" box="[264,362,1728,1752]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="sciakyi" status="sp. nov.">
|
||
<emphasis id="B95A1B27625FFFFA2C70B4BAFEF05B82" box="[264,279,1729,1752]" italics="true" pageId="25" pageNumber="26">P</emphasis>
|
||
.
|
||
<emphasis id="B95A1B27625FFFFA2C5CB4BBFE8D5B82" box="[292,362,1728,1752]" italics="true" pageId="25" pageNumber="26">sciakyi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95A1B27625FFFFA2C09B4BAFE4D5B83" bold="true" box="[369,426,1729,1753]" pageId="25" pageNumber="26">
|
||
<taxonomicNameLabel id="A269A65C625FFFFA2C09B4BAFE4D5B83" box="[369,426,1729,1753]" pageId="25" pageNumber="26" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype.
|
||
<emphasis id="B95A1B27625FFFFA2F64B4BAFDCA5B83" bold="true" box="[540,557,1729,1753]" pageId="25" pageNumber="26">L</emphasis>
|
||
<taxonomicName id="4C2EBCB6625FFFFA2F4BB4BAFCFD5B82" authority="(Chaudoir)" baseAuthorityName="Chaudoir" box="[563,794,1728,1752]" class="Insecta" family="Carabidae" genus="Crossoglossa" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="testacea">
|
||
<emphasis id="B95A1B27625FFFFA2F4BB4BAFDA55B82" box="[563,578,1729,1752]" italics="true" pageId="25" pageNumber="26">P</emphasis>
|
||
.
|
||
<emphasis id="B95A1B27625FFFFA2F37B4BAFD455B82" box="[591,674,1729,1752]" italics="true" pageId="25" pageNumber="26">testacea</emphasis>
|
||
(Chaudoir)
|
||
</taxonomicName>
|
||
, Guangdong.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C33494BE625FFFF92DEFB550FA7B5FA2" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26" type="key">
|
||
<paragraph id="8B91C735625FFFFA2DEFB550FCEF5A1E" blockId="25.[151,776,1834,1861]" box="[151,776,1834,1861]" pageId="25" pageNumber="26">
|
||
<emphasis id="B95A1B27625FFFFA2DEFB550FCEF5A1E" bold="true" box="[151,776,1834,1861]" pageId="25" pageNumber="26">
|
||
Key to species of subgenus
|
||
<taxonomicName id="4C2EBCB6625FFFFA2C91B551FCEF5A1E" ID-CoL="8KWFM" authority="Chaudoir" authorityName="Chaudoir" box="[489,776,1834,1861]" class="Insecta" family="Carabidae" genus="Crossoglossa" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95A1B27625FFFFA2C91B551FD605A1F" bold="true" box="[489,647,1834,1861]" italics="true" pageId="25" pageNumber="26">Crossoglossa</emphasis>
|
||
Chaudoir
|
||
</taxonomicName>
|
||
</emphasis>
|
||
</paragraph>
|
||
<key id="FFBF9004625FFFF92DEFB50BFA7B5FA2" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26">
|
||
<keyStep id="30DADF90625FFFFA2DEFB50BFA7B5AF9" pageId="25" pageNumber="26">
|
||
<paragraph id="8B91C735625FFFFA2DEFB50BFA7B5ADF" blockId="25.[151,1437,1904,2014]" box="[151,1436,1904,1926]" pageId="25" pageNumber="26">
|
||
<keyLead id="30DF6400625FFFFA2DEFB50BFA7B5ADF" box="[151,1436,1904,1926]" pageId="25" pageNumber="26">1. Dorsum reddish yellow to reddish brown, tarsi same as or slightly darker than dorsum............................... 2</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B91C735625FFFFA2DEFB5F6FA7B5AF9" blockId="25.[151,1437,1904,2014]" box="[151,1436,1933,1955]" pageId="25" pageNumber="26">
|
||
<keyLead id="30DF6400625FFFFA2DEFB5F6FA7B5AF9" box="[151,1436,1933,1955]" pageId="25" pageNumber="26">
|
||
- Dorsum pale yellow, tarsi black, distinctly darker than dorsum. (3.
|
||
<taxonomicName id="4C2EBCB6625FFFFA2E1CB5F5FC2C5AF9" ID-CoL="ZMF7" box="[868,971,1934,1955]" class="Insecta" family="Carabidae" genus="Crossoglossa" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="testacea">
|
||
<emphasis id="B95A1B27625FFFFA2E1CB5F5FC2C5AF9" box="[868,971,1934,1955]" italics="true" pageId="25" pageNumber="26">P. testacea</emphasis>
|
||
</taxonomicName>
|
||
group)............................... 5
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30DADF90625FFFF92DEFB5D1FA7B5D90" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26">
|
||
<paragraph id="8B91C735625FFFFA2DEFB5D1FA7B5A87" blockId="25.[151,1437,1904,2014]" pageId="25" pageNumber="26">
|
||
<keyLead id="30DF6400625FFFFA2DEFB5D1FA7B5A87" pageId="25" pageNumber="26">
|
||
2. Elytral sutural angles only faintly pointed (
|
||
<figureCitation id="1315DBB0625FFFFA2FFFB5D1FD055AE5" box="[647,738,1962,1984]" captionStart="FIGURE 14" captionStartId="25.[151,250,1584,1608]" captionTargetBox="[151,1436,181,1561]" captionTargetId="figure-19@25.[151,1436,181,1561]" captionTargetPageId="25" captionText="FIGURE 14. Elytra apex of Parena (Crossoglossa) spp. A–H Elytra sutural angle (left), scale bar = 0.2mm. A P. cavipennis (Bates), Shaanxi. B P. laesipennis (Bates), Vietnam. C P. levata Andrewes, Borneo. D P. testacea (Chaudoir), Guangdong. E P. sulawesiensis Kirschenhofer, Sulawesi. F P. cruralis Andrewes, Borneo. G P. mellea (Chaudoir), Philippine. H P. sciakyi sp. n., holotype. I–L Elytra outer apical angle (left), scale bar = 0.5mm. I P. cavipennis (Bates), Shaanxi. J P. laesipennis (Bates), Taiwan. K P. sciakyi sp. n., holotype. L P. testacea (Chaudoir), Guangdong." figureDoi="http://doi.org/10.5281/zenodo.7958963" httpUri="https://zenodo.org/record/7958963/files/figure.png" pageId="25" pageNumber="26">Fig. 14A</figureCitation>
|
||
), outer apical angles completely rounded (
|
||
<figureCitation id="1315DBB0625FFFFA29FAB5D1FB345AE5" box="[1154,1235,1962,1984]" captionStart="FIGURE 14" captionStartId="25.[151,250,1584,1608]" captionTargetBox="[151,1436,181,1561]" captionTargetId="figure-19@25.[151,1436,181,1561]" captionTargetPageId="25" captionText="FIGURE 14. Elytra apex of Parena (Crossoglossa) spp. A–H Elytra sutural angle (left), scale bar = 0.2mm. A P. cavipennis (Bates), Shaanxi. B P. laesipennis (Bates), Vietnam. C P. levata Andrewes, Borneo. D P. testacea (Chaudoir), Guangdong. E P. sulawesiensis Kirschenhofer, Sulawesi. F P. cruralis Andrewes, Borneo. G P. mellea (Chaudoir), Philippine. H P. sciakyi sp. n., holotype. I–L Elytra outer apical angle (left), scale bar = 0.5mm. I P. cavipennis (Bates), Shaanxi. J P. laesipennis (Bates), Taiwan. K P. sciakyi sp. n., holotype. L P. testacea (Chaudoir), Guangdong." figureDoi="http://doi.org/10.5281/zenodo.7958963" httpUri="https://zenodo.org/record/7958963/files/figure.png" pageId="25" pageNumber="26">Fig. 14I</figureCitation>
|
||
); striae well incised, finely punctate; intervals slightly convex (1.
|
||
<taxonomicName id="4C2EBCB6625FFFFA2FE8B5B3FCE95A87" ID-CoL="4DKP6" baseAuthorityName="Bates" baseAuthorityYear="1873" box="[656,782,1992,2013]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="cavipennis">
|
||
<emphasis id="B95A1B27625FFFFA2FE8B5B3FCE95A87" box="[656,782,1992,2013]" italics="true" pageId="25" pageNumber="26">P. cavipennis</emphasis>
|
||
</taxonomicName>
|
||
group)............................ [1]
|
||
<taxonomicName id="4C2EBCB6625FFFFA29A8B5B3FA7B5A87" ID-CoL="4DKP6" authority="(Bates)" baseAuthorityName="Bates" baseAuthorityYear="1873" box="[1232,1436,1992,2013]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="cavipennis">
|
||
<emphasis id="B95A1B27625FFFFA29A8B5B3FAB65A87" bold="true" box="[1232,1361,1992,2013]" italics="true" pageId="25" pageNumber="26">P. cavipennis</emphasis>
|
||
(Bates)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B91C735625CFFF92DEFB2E3FA7B5D90" blockId="26.[151,1437,151,760]" pageId="26" pageNumber="27">
|
||
<keyLead id="30DF6400625CFFF92DEFB2E3FA7B5D90" pageId="26" pageNumber="27">
|
||
- Elytral sutural angles distinctly pointed, extended as short spines (
|
||
<figureCitation id="1315DBB0625CFFF92E1EB2ECFC1F5DF7" box="[870,1016,151,173]" captionStart="FIGURE 14" captionStartId="25.[151,250,1584,1608]" captionTargetBox="[151,1436,181,1561]" captionTargetId="figure-19@25.[151,1436,181,1561]" captionTargetPageId="25" captionText="FIGURE 14. Elytra apex of Parena (Crossoglossa) spp. A–H Elytra sutural angle (left), scale bar = 0.2mm. A P. cavipennis (Bates), Shaanxi. B P. laesipennis (Bates), Vietnam. C P. levata Andrewes, Borneo. D P. testacea (Chaudoir), Guangdong. E P. sulawesiensis Kirschenhofer, Sulawesi. F P. cruralis Andrewes, Borneo. G P. mellea (Chaudoir), Philippine. H P. sciakyi sp. n., holotype. I–L Elytra outer apical angle (left), scale bar = 0.5mm. I P. cavipennis (Bates), Shaanxi. J P. laesipennis (Bates), Taiwan. K P. sciakyi sp. n., holotype. L P. testacea (Chaudoir), Guangdong." figureDoi="http://doi.org/10.5281/zenodo.7958963" httpUri="https://zenodo.org/record/7958963/files/figure.png" pageId="26" pageNumber="27">Figs 14B, 14C</figureCitation>
|
||
), outer apical angles more or less rounded-angulate (
|
||
<figureCitation id="1315DBB0625CFFF92C38B2CEFE755D90" box="[320,402,181,203]" captionStart="FIGURE 14" captionStartId="25.[151,250,1584,1608]" captionTargetBox="[151,1436,181,1561]" captionTargetId="figure-19@25.[151,1436,181,1561]" captionTargetPageId="25" captionText="FIGURE 14. Elytra apex of Parena (Crossoglossa) spp. A–H Elytra sutural angle (left), scale bar = 0.2mm. A P. cavipennis (Bates), Shaanxi. B P. laesipennis (Bates), Vietnam. C P. levata Andrewes, Borneo. D P. testacea (Chaudoir), Guangdong. E P. sulawesiensis Kirschenhofer, Sulawesi. F P. cruralis Andrewes, Borneo. G P. mellea (Chaudoir), Philippine. H P. sciakyi sp. n., holotype. I–L Elytra outer apical angle (left), scale bar = 0.5mm. I P. cavipennis (Bates), Shaanxi. J P. laesipennis (Bates), Taiwan. K P. sciakyi sp. n., holotype. L P. testacea (Chaudoir), Guangdong." figureDoi="http://doi.org/10.5281/zenodo.7958963" httpUri="https://zenodo.org/record/7958963/files/figure.png" pageId="26" pageNumber="27">Fig. 14J</figureCitation>
|
||
); striae not incised, replaced by rows of fine punctures; intervals flat (2.
|
||
<taxonomicName id="4C2EBCB6625CFFF92932B2CEFB2A5D90" ID-CoL="4DKPN" baseAuthorityName="Bates" baseAuthorityYear="1873" box="[1098,1229,181,202]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="laesipennis">
|
||
<emphasis id="B95A1B27625CFFF92932B2CEFB2A5D90" box="[1098,1229,181,202]" italics="true" pageId="26" pageNumber="27">P. laesipennis</emphasis>
|
||
</taxonomicName>
|
||
group).......... 3
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30DADF90625CFFF92DEFB2A9FA7B5C5F" pageId="26" pageNumber="27">
|
||
<paragraph id="8B91C735625CFFF92DEFB2A9FA7B5DB2" blockId="26.[151,1437,151,760]" box="[151,1436,210,232]" pageId="26" pageNumber="27">
|
||
<keyLead id="30DF6400625CFFF92DEFB2A9FA7B5DB2" box="[151,1436,210,232]" pageId="26" pageNumber="27">
|
||
3. Elytra not depressed on intervals 3 to 6;
|
||
<collectingRegion id="49EA09D7625CFFF92F1EB2A9FD5F5DBD" box="[614,696,210,231]" country="Indonesia" name="Sumatera Utara" pageId="26" pageNumber="27">Sumatra</collectingRegion>
|
||
, Borneo.................................... [3]
|
||
<taxonomicName id="4C2EBCB6625CFFF9299CB2A9FA7B5DB2" ID-CoL="4DKPQ" authority="Andrewes" authorityName="Andrewes, Borneo. D" authorityYear="1931" box="[1252,1436,210,232]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="levata">
|
||
<emphasis id="B95A1B27625CFFF9299CB2A9FAD05DB2" bold="true" box="[1252,1335,210,232]" italics="true" pageId="26" pageNumber="27">P. levata</emphasis>
|
||
Andrewes
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B91C735625CFFF92DEFB28BFA7B5C5F" blockId="26.[151,1437,151,760]" box="[151,1436,239,261]" pageId="26" pageNumber="27">
|
||
<keyLead id="30DF6400625CFFF92DEFB28BFA7B5C5F" box="[151,1436,239,261]" pageId="26" pageNumber="27">- Elytra more or less depressed near middle on intervals 3 to 6; other localities...................................... 4</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30DADF90625CFFF92DEFB376FA7A5C21" pageId="26" pageNumber="27">
|
||
<paragraph id="8B91C735625CFFF92DEFB376FA7B5C65" blockId="26.[151,1437,151,760]" pageId="26" pageNumber="27">
|
||
<keyLead id="30DF6400625CFFF92DEFB376FA7B5C65" pageId="26" pageNumber="27">
|
||
4. Elytral outer apical angles only weakly angulate, forming rounded obtuse angles; apical lamella of aedeagus wider, LL/LW 1.50–1.65; widely in
|
||
<collectingCountry id="F33987A5625CFFF92CD7B350FDEE5C1A" box="[431,521,298,320]" name="China" pageId="26" pageNumber="27">East Asia</collectingCountry>
|
||
and Southeast Asia, except the Malay Archipelago............... [2]
|
||
<taxonomicName id="4C2EBCB6625CFFF929B3B351FA7B5C65" ID-CoL="4DKPN" authority="(Bates)" baseAuthorityName="Bates" baseAuthorityYear="1873" box="[1227,1436,298,320]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="laesipennis">
|
||
<emphasis id="B95A1B27625CFFF929B3B351FAB65C1A" bold="true" box="[1227,1361,298,320]" italics="true" pageId="26" pageNumber="27">P. laesipennis</emphasis>
|
||
(Bates)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B91C735625CFFF92DEFB333FA7A5C21" blockId="26.[151,1437,151,760]" pageId="26" pageNumber="27">
|
||
<keyLead id="30DF6400625CFFF92DEFB333FA7A5C21" pageId="26" pageNumber="27">
|
||
- Elytral outer apical angles distinctly angulate, forming sharper obtuse angles; apical lamella of aedeagus narrower, LL/LW approximately 1.75;
|
||
<collectingRegion id="49EA09D7625CFFF92CEBB31EFE595C20" box="[403,446,357,378]" country="Indonesia" name="Jawa Timur" pageId="26" pageNumber="27">Java</collectingRegion>
|
||
............................................................ [4]
|
||
<taxonomicName id="4C2EBCB6625CFFF929C1B31EFA7A5C21" ID-CoL="4DKPZ" authority="Jedlicka" authorityName="Jedlicka" authorityYear="1952" box="[1209,1437,356,379]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="obenbergeri">
|
||
<emphasis id="B95A1B27625CFFF929C1B31EFAAF5C20" bold="true" box="[1209,1352,356,378]" italics="true" pageId="26" pageNumber="27">P. obenbergeri</emphasis>
|
||
Jedlička
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30DADF90625CFFF92DEFB3F9FA7B5CEF" pageId="26" pageNumber="27">
|
||
<paragraph id="8B91C735625CFFF92DEFB3F9FA7B5CCD" blockId="26.[151,1437,151,760]" box="[151,1436,386,408]" pageId="26" pageNumber="27">
|
||
<keyLead id="30DF6400625CFFF92DEFB3F9FA7B5CCD" box="[151,1436,386,408]" pageId="26" pageNumber="27">
|
||
5. Elytral disc with large black patch; sutural angles not pointed (
|
||
<figureCitation id="1315DBB0625CFFF92E43B3F9FC725CCD" box="[827,917,386,408]" captionStart="FIGURE 14" captionStartId="25.[151,250,1584,1608]" captionTargetBox="[151,1436,181,1561]" captionTargetId="figure-19@25.[151,1436,181,1561]" captionTargetPageId="25" captionText="FIGURE 14. Elytra apex of Parena (Crossoglossa) spp. A–H Elytra sutural angle (left), scale bar = 0.2mm. A P. cavipennis (Bates), Shaanxi. B P. laesipennis (Bates), Vietnam. C P. levata Andrewes, Borneo. D P. testacea (Chaudoir), Guangdong. E P. sulawesiensis Kirschenhofer, Sulawesi. F P. cruralis Andrewes, Borneo. G P. mellea (Chaudoir), Philippine. H P. sciakyi sp. n., holotype. I–L Elytra outer apical angle (left), scale bar = 0.5mm. I P. cavipennis (Bates), Shaanxi. J P. laesipennis (Bates), Taiwan. K P. sciakyi sp. n., holotype. L P. testacea (Chaudoir), Guangdong." figureDoi="http://doi.org/10.5281/zenodo.7958963" httpUri="https://zenodo.org/record/7958963/files/figure.png" pageId="26" pageNumber="27">Fig. 14H</figureCitation>
|
||
);
|
||
<collectingCountry id="F33987A5625CFFF92EDEB3F9FBA05CC2" box="[934,1095,386,408]" name="Solomon Islands" pageId="26" pageNumber="27">Solomon Islands</collectingCountry>
|
||
........... [9]
|
||
<emphasis id="B95A1B27625CFFF92996B3F9FA7B5CCD" bold="true" box="[1262,1436,386,408]" pageId="26" pageNumber="27">
|
||
<taxonomicName id="4C2EBCB6625CFFF92996B3F9FAAD5CC2" authority="Shi & Liang, 2023" authorityName="Shi & Liang" authorityYear="2023" box="[1262,1354,386,408]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="sciakyi" status="sp. nov.">
|
||
<emphasis id="B95A1B27625CFFF92996B3F9FAAD5CC2" bold="true" box="[1262,1354,386,408]" italics="true" pageId="26" pageNumber="27">P. sciakyi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A269A65C625CFFF92828B3F9FA7B5CCD" box="[1360,1436,386,407]" pageId="26" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B91C735625CFFF92DEFB3DBFA7B5CEF" blockId="26.[151,1437,151,760]" box="[151,1436,415,437]" pageId="26" pageNumber="27">
|
||
<keyLead id="30DF6400625CFFF92DEFB3DBFA7B5CEF" box="[151,1436,415,437]" pageId="26" pageNumber="27">- Elytra uniformly yellow; sutural angles more or less pointed; other localities...................................... 6</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30DADF90625CFFF92DEFB3C6FA7B5CB5" pageId="26" pageNumber="27">
|
||
<paragraph id="8B91C735625CFFF92DEFB3C6FA7B5C88" blockId="26.[151,1437,151,760]" box="[151,1436,445,467]" pageId="26" pageNumber="27">
|
||
<keyLead id="30DF6400625CFFF92DEFB3C6FA7B5C88" box="[151,1436,445,467]" pageId="26" pageNumber="27">
|
||
6. Elytral sutural angles sharply pointed, forming short spines (
|
||
<figureCitation id="1315DBB0625CFFF92E5FB3C6FC535C88" box="[807,948,445,467]" captionStart="FIGURE 14" captionStartId="25.[151,250,1584,1608]" captionTargetBox="[151,1436,181,1561]" captionTargetId="figure-19@25.[151,1436,181,1561]" captionTargetPageId="25" captionText="FIGURE 14. Elytra apex of Parena (Crossoglossa) spp. A–H Elytra sutural angle (left), scale bar = 0.2mm. A P. cavipennis (Bates), Shaanxi. B P. laesipennis (Bates), Vietnam. C P. levata Andrewes, Borneo. D P. testacea (Chaudoir), Guangdong. E P. sulawesiensis Kirschenhofer, Sulawesi. F P. cruralis Andrewes, Borneo. G P. mellea (Chaudoir), Philippine. H P. sciakyi sp. n., holotype. I–L Elytra outer apical angle (left), scale bar = 0.5mm. I P. cavipennis (Bates), Shaanxi. J P. laesipennis (Bates), Taiwan. K P. sciakyi sp. n., holotype. L P. testacea (Chaudoir), Guangdong." figureDoi="http://doi.org/10.5281/zenodo.7958963" httpUri="https://zenodo.org/record/7958963/files/figure.png" pageId="26" pageNumber="27">Figs 14D, 14E</figureCitation>
|
||
)....................................... 7
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B91C735625CFFF92DEFB3A0FA7B5CB5" blockId="26.[151,1437,151,760]" box="[151,1436,474,496]" pageId="26" pageNumber="27">
|
||
<keyLead id="30DF6400625CFFF92DEFB3A0FA7B5CB5" box="[151,1436,474,496]" pageId="26" pageNumber="27">
|
||
- Elytral sutural angles gently pointed, forming small denticles (
|
||
<figureCitation id="1315DBB0625CFFF92E41B3A1FC255CB5" box="[825,962,474,496]" captionStart="FIGURE 14" captionStartId="25.[151,250,1584,1608]" captionTargetBox="[151,1436,181,1561]" captionTargetId="figure-19@25.[151,1436,181,1561]" captionTargetPageId="25" captionText="FIGURE 14. Elytra apex of Parena (Crossoglossa) spp. A–H Elytra sutural angle (left), scale bar = 0.2mm. A P. cavipennis (Bates), Shaanxi. B P. laesipennis (Bates), Vietnam. C P. levata Andrewes, Borneo. D P. testacea (Chaudoir), Guangdong. E P. sulawesiensis Kirschenhofer, Sulawesi. F P. cruralis Andrewes, Borneo. G P. mellea (Chaudoir), Philippine. H P. sciakyi sp. n., holotype. I–L Elytra outer apical angle (left), scale bar = 0.5mm. I P. cavipennis (Bates), Shaanxi. J P. laesipennis (Bates), Taiwan. K P. sciakyi sp. n., holotype. L P. testacea (Chaudoir), Guangdong." figureDoi="http://doi.org/10.5281/zenodo.7958963" httpUri="https://zenodo.org/record/7958963/files/figure.png" pageId="26" pageNumber="27">Figs 14F, 14G</figureCitation>
|
||
)..................................... 8
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30DADF90625CFFF92DEFB383FA7B5FFA" pageId="26" pageNumber="27">
|
||
<paragraph id="8B91C735625CFFF92DEFB383FA7B5F12" blockId="26.[151,1437,151,760]" pageId="26" pageNumber="27">
|
||
<keyLead id="30DF6400625CFFF92DEFB383FA7B5F12" pageId="26" pageNumber="27">
|
||
7. Antennomere 1 yellow, other ten antennomeres black; elytral intervals with dense punctures, similar density as punctures in striae, so that striae poorly defined; apical lamella of aedeagus shorter, LL/LW = 1.0–1.1;
|
||
<collectingCountry id="F33987A5625CFFF92939B06EFB925F71" box="[1089,1141,533,555]" name="India" pageId="26" pageNumber="27">India</collectingCountry>
|
||
,
|
||
<collectingCountry id="F33987A5625CFFF92907B06EFB5C5F71" box="[1151,1211,533,555]" name="Nepal" pageId="26" pageNumber="27">Nepal</collectingCountry>
|
||
,
|
||
<collectingCountry id="F33987A5625CFFF929BBB06EFAF05F71" box="[1219,1303,533,555]" name="Vietnam" pageId="26" pageNumber="27">Vietnam</collectingCountry>
|
||
,
|
||
<collectingCountry id="F33987A5625CFFF92859B06EFABD5F71" box="[1313,1370,533,555]" name="China" pageId="26" pageNumber="27">China</collectingCountry>
|
||
.................................................................................... [5]
|
||
<taxonomicName id="4C2EBCB6625CFFF929BFB049FA7B5F12" ID-CoL="ZMF7" authority="(Chaudoir)" baseAuthorityName="Chaudoir" box="[1223,1436,562,584]" class="Insecta" family="Carabidae" genus="Crossoglossa" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="testacea">
|
||
<emphasis id="B95A1B27625CFFF929BFB049FACA5F1D" bold="true" box="[1223,1325,562,583]" italics="true" pageId="26" pageNumber="27">P. testacea</emphasis>
|
||
(Chaudoir)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B91C735625CFFF92DEFB02BFA7B5FFA" blockId="26.[151,1437,151,760]" pageId="26" pageNumber="27">
|
||
<keyLead id="30DF6400625CFFF92DEFB02BFA7B5FFA" pageId="26" pageNumber="27">
|
||
- Antennomeres 1 to 3 yellow, other eight antennomeres black; elytral intervals with sparse punctures, much sparser than punctures in striae, so that striae clearly defined; apical lamella of aedeagus longer, LL/LW approximately 1.3;
|
||
<collectingRegion id="49EA09D7625CFFF929A7B016FAD15FD9" box="[1247,1334,621,643]" country="Indonesia" name="Sulawesi Utara" pageId="26" pageNumber="27">Sulawesi</collectingRegion>
|
||
................................................................................ [6]
|
||
<taxonomicName id="4C2EBCB6625CFFF92908B0F1FA7B5FFA" ID-CoL="4DKQN" authority="Kirschenhofer" authorityName="Kirschenhofer. D" authorityYear="2006" box="[1136,1436,650,672]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="sulawesiensis">
|
||
<emphasis id="B95A1B27625CFFF92908B0F1FAEA5FFA" bold="true" box="[1136,1293,650,672]" italics="true" pageId="26" pageNumber="27">P. sulawesiensis</emphasis>
|
||
Kirschenhofer
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30DADF90625CFFF92DEFB0D3FA7B5FA2" pageId="26" pageNumber="27">
|
||
<paragraph id="8B91C735625CFFF92DEFB0D3FA7B5F81" blockId="26.[151,1437,151,760]" pageId="26" pageNumber="27">
|
||
<keyLead id="30DF6400625CFFF92DEFB0D3FA7B5F81" pageId="26" pageNumber="27">
|
||
8. Tibiae yellow; elytra disc with depressions distinct;
|
||
<collectingCountry id="F33987A5625CFFF92FAAB0DCFC8E5FE7" box="[722,873,679,701]" name="Indonesia" pageId="26" pageNumber="27">Maluku Islands</collectingCountry>
|
||
, Adonara Island,
|
||
<collectingCountry id="F33987A5625CFFF92977B0D3FB2D5FE7" box="[1039,1226,679,701]" name="Papua New Guinea" pageId="26" pageNumber="27">Papua New Guinea</collectingCountry>
|
||
, E. Java, Sumatra, the
|
||
<collectingCountry id="F33987A5625CFFF92D9AB0BEFEA95F81" box="[226,334,709,731]" name="Philippines" pageId="26" pageNumber="27">Philippines</collectingCountry>
|
||
........................................................................[7]
|
||
<taxonomicName id="4C2EBCB6625CFFF929ACB0BEFA7B5F81" ID-CoL="4DKPT" authority="(Chaudoir)" baseAuthorityName="Chaudoir" baseAuthorityYear="1872" box="[1236,1436,708,731]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="mellea">
|
||
<emphasis id="B95A1B27625CFFF929ACB0BEFACA5F80" bold="true" box="[1236,1325,708,730]" italics="true" pageId="26" pageNumber="27">P. mellea</emphasis>
|
||
(Chaudoir)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B91C735625CFFF92DEFB098FA7B5FA2" blockId="26.[151,1437,151,760]" box="[151,1436,738,760]" pageId="26" pageNumber="27">
|
||
<keyLead id="30DF6400625CFFF92DEFB098FA7B5FA2" box="[151,1436,738,760]" pageId="26" pageNumber="27">
|
||
- Tibiae black; elytra disc with depressions absent or very shallow; W.
|
||
<collectingRegion id="49EA09D7625CFFF92E02B098FC405FA2" box="[890,935,739,760]" country="Indonesia" name="Jawa Timur" pageId="26" pageNumber="27">Java</collectingRegion>
|
||
, Borneo.............. [8]
|
||
<taxonomicName id="4C2EBCB6625CFFF929A8B099FA7B5FA2" authority="Andrewes" authorityName="Andrewes, Borneo. E" authorityYear="1933" box="[1232,1436,738,760]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="cruralis">
|
||
<emphasis id="B95A1B27625CFFF929A8B099FAD25FA2" bold="true" box="[1232,1333,738,760]" italics="true" pageId="26" pageNumber="27">P. cruralis</emphasis>
|
||
Andrewes
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</key>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |