treatments-xml/data/03/87/76/03877623623DFF982DEFB2ECFAEE59C6.xml

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<mods:title id="493EB779BABF08D48CF0789E1B9DF341">Taxonomic revision of the genus Parena Motschulsky, 1860 (Coleoptera, Carabidae, Lebiini, Metallicina)</mods:title>
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<mods:namePart id="2DD4827C6BDB48725281C426DA7F2EBA">Shi, Hongliang</mods:namePart>
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<paragraph id="8B91C735623DFF982DEFB2ECFD275DEB" blockId="123.[151,704,151,213]" box="[151,704,151,178]" pageId="123" pageNumber="124">
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[38] 
<taxonomicName id="4C2EBCB6623DFF982DB7B2ECFD275DEB" ID-CoL="4DKP8" authority="Basilewsky, 1955" authorityName="Basilewsky. C" authorityYear="1955" box="[207,704,151,178]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="dorae" subGenus="Parena">
<emphasis id="B95A1B27623DFF982DB7B2ECFEC05DEB" bold="true" box="[207,295,151,177]" italics="true" pageId="123" pageNumber="124">Parena</emphasis>
(
<emphasis id="B95A1B27623DFF982C40B2ECFE775DEB" bold="true" box="[312,400,151,177]" italics="true" pageId="123" pageNumber="124">Parena</emphasis>
) 
<emphasis id="B95A1B27623DFF982CDAB2ECFE015DE8" bold="true" box="[418,486,151,178]" italics="true" pageId="123" pageNumber="124">dorae</emphasis>
<bibRefCitation id="EFBFBAC4623DFF982C95B2ECFD275DEB" author="Basilewsky, P." box="[493,704,151,177]" pageId="123" pageNumber="124" pagination="93 - 137" refId="ref109889" refString="Basilewsky, P. (1955) Carabidae (Coleoptera) de l'Angola. Publicacoes Culturais da Companhia de Diamantes de Angola, 27, 93 - 137." type="journal article" year="1955">Basilewsky, 1955</bibRefCitation>
</taxonomicName>
</emphasis>
</paragraph>
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<subSubSection id="C33494BE623DFF982DEFB2C0FE815D8F" box="[151,358,187,213]" pageId="123" pageNumber="124" type="description">
<paragraph id="8B91C735623DFF982DEFB2C0FE815D8F" blockId="123.[151,704,151,213]" box="[151,358,187,213]" pageId="123" pageNumber="124">
Habitus: 
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.
</paragraph>
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<paragraph id="8B91C735623DFF982DEFB37BFC745C42" blockId="123.[151,915,255,280]" box="[151,915,255,280]" pageId="123" pageNumber="124">
<materialsCitation id="3B46CD68623DFF982DEFB37BFC685C42" ID-GBIF-Occurrence="4115871460" box="[151,911,255,280]" country="Angola" location="Dundo" pageId="123" pageNumber="124" specimenCount="1" typeStatus="holotype">
<bibRefCitation id="EFBFBAC4623DFF982DEFB37BFE9A5C42" author="Basilewsky, P." box="[151,381,255,280]" pageId="123" pageNumber="124" pagination="93 - 137" refId="ref109889" refString="Basilewsky, P. (1955) Carabidae (Coleoptera) de l'Angola. Publicacoes Culturais da Companhia de Diamantes de Angola, 27, 93 - 137." type="journal article" year="1955">Basilewsky, 1955: 129</bibRefCitation>
(type locality: 
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, 
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; 
<typeStatus id="54957997623DFF982FC7B37BFCFF5C42" box="[703,792,256,280]" pageId="123" pageNumber="124" type="holotype">holotype</typeStatus>
in MRAC)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33494BE623DFF982DEFB33DFDB25CF2" pageId="123" pageNumber="124" type="materials_examined">
<paragraph id="8B91C735623DFF982DEFB33DFDB25CF2" blockId="123.[151,1437,325,1180]" pageId="123" pageNumber="124">
<emphasis id="B95A1B27623DFF982DEFB33DFE525C05" bold="true" box="[151,437,325,352]" pageId="123" pageNumber="124">Type material examined</emphasis>
. 
<emphasis id="B95A1B27623DFF982CBAB33DFC845C3A" bold="true" box="[450,867,325,352]" pageId="123" pageNumber="124">
<taxonomicName id="4C2EBCB6623DFF982CBAB33DFD0B5C05" authority="Basilewsky" authorityName="Basilewsky. C" authorityYear="1955" box="[450,748,325,352]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="dorae">
<emphasis id="B95A1B27623DFF982CBAB33DFDBA5C3A" bold="true" box="[450,605,325,352]" italics="true" pageId="123" pageNumber="124">Parena dorae</emphasis>
Basilewsky
</taxonomicName>
: 
<typeStatus id="54957997623DFF982F8EB33DFC845C3A" box="[758,867,326,352]" pageId="123" pageNumber="124" type="holotype">Holotype</typeStatus>
</emphasis>
(MRAC, 
<figureCitation id="1315DBB0623DFF982EA9B33DFBD05C3A" box="[977,1079,326,352]" captionStart="FIGURE 73" captionStartId="121.[151,250,1902,1926]" captionTargetBox="[167,1409,298,1862]" captionTargetId="figure-87@121.[151,1436,294,1878]" captionTargetPageId="121" captionText="FIGURE 73. Habitus of Parena (Parena) stigmatica species group. A Lectotype of Parena stigmatica (Fairmaire). B Holotype of Parena dorae Basilewsky. C Holotype of Parena fulva sp. n. D Paratype of Parena fulva sp. n., female from Zimbabwe. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7959090" httpUri="https://zenodo.org/record/7959090/files/figure.png" pageId="123" pageNumber="124">Fig. 73B</figureCitation>
): female, body length = 
<quantity id="4CD66AD0623DFF982839B33DFA7F5C3A" box="[1345,1432,326,352]" metricMagnitude="-3" metricUnit="m" metricValue="9.1" pageId="123" pageNumber="124" unit="mm" value="9.1">9.1 mm</quantity>
, pin mounted, &quot; 
<typeStatus id="54957997623DFF982C42B311FE0F5CDE" box="[314,488,362,388]" pageId="123" pageNumber="124" type="holotype">HOLOTYPUS</typeStatus>
&quot; [red label]; &quot; 
<collectingCountry id="F33987A5623DFF982FFAB311FD3B5CDE" box="[642,732,362,388]" name="Angola" pageId="123" pageNumber="124">Angola</collectingCountry>
: Dundo / 
<emphasis id="B95A1B27623DFF982E31B310FC455CDE" box="[841,930,362,388]" italics="true" pageId="123" pageNumber="124">
<date id="FF90E1F5623DFF982E31B310FC455CDE" box="[841,930,362,388]" pageId="123" pageNumber="124" value="1948-02">II. 1948</date>
</emphasis>
/ A. de Barros Machado&quot;, &quot; 
<emphasis id="B95A1B27623DFF9829A2B311FAF05CDE" box="[1242,1303,362,388]" italics="true" pageId="123" pageNumber="124">411.6</emphasis>
&quot;, &quot; 
<emphasis id="B95A1B27623DFF982844B310FECD5CF2" italics="true" pageId="123" pageNumber="124">
<taxonomicName id="4C2EBCB6623DFF982844B310FA6A5CDE" box="[1340,1421,363,388]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="genus">Parena</taxonomicName>
/ 
<taxonomicName id="4C2EBCB6623DFF982DEFB3F4FF385CF2" authority="Shi &amp; Liang, 2023" authorityName="Shi &amp; Liang" authorityYear="2023" box="[151,223,399,424]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="dorae" status="sp. nov.">Dorae</taxonomicName>
<taxonomicNameLabel id="A269A65C623DFF982D9EB3F4FEFC5CF2" box="[230,283,399,424]" pageId="123" pageNumber="124" rank="species">n.sp.</taxonomicNameLabel>
/
</emphasis>
P. Basilewsky det., 19 
<emphasis id="B95A1B27623DFF982F5EB3F5FDA55CF2" box="[550,578,398,424]" italics="true" pageId="123" pageNumber="124">54</emphasis>
&quot;.
</paragraph>
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<paragraph id="8B91C735623DFF982DBFB3CAFB955F06" blockId="123.[151,1437,325,1180]" pageId="123" pageNumber="124">
<emphasis id="B95A1B27623DFF982DBFB3CAFE815C91" bold="true" box="[199,358,433,459]" pageId="123" pageNumber="124">Comparisons</emphasis>
. This species differs from all other species of subgenus 
<taxonomicName id="4C2EBCB6623DFF982E92B3C8FBDC5C96" box="[1002,1083,435,460]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="genus">
<emphasis id="B95A1B27623DFF982E92B3C8FBDC5C96" box="[1002,1083,435,460]" italics="true" pageId="123" pageNumber="124">Parena</emphasis>
</taxonomicName>
by the combination of dorsum reddish brown with distinct lustre, elytra without pattern, antennae distinctly bicolor; pronotum with lateral margins strongly sinuate before posterior angles; postgenae with suborbital setae; elytra disc shallowly depressed. Among all African species, this species is most similar to 
<taxonomicName id="4C2EBCB6623DFF982FB0B064FCBE5F62" baseAuthorityName="Fairmaire" baseAuthorityYear="1899" box="[712,857,543,568]" class="Insecta" family="Carabidae" genus="Parena" kingdom="Animalia" order="Coleoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="stigmatica">
<emphasis id="B95A1B27623DFF982FB0B064FCBE5F62" box="[712,857,543,568]" italics="true" pageId="123" pageNumber="124">P. stigmatica</emphasis>
</taxonomicName>
, but can be distinguished from the latter species by its smaller size, lighter color, elytra with shallower depressions, and antennae bicolored.
</paragraph>
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<subSubSection id="C33494BE623DFF982DBFB01EFCA05922" pageId="123" pageNumber="124" type="description">
<paragraph id="8B91C735623DFF982DBFB01EFCA05922" blockId="123.[151,1437,325,1180]" pageId="123" pageNumber="124">
<emphasis id="B95A1B27623DFF982DBFB01EFEB65F25" bold="true" box="[199,337,613,639]" pageId="123" pageNumber="124">Description</emphasis>
. Body length 
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. Dorsum reddish brown, strongly polished; basal four antennomeres yellowish brown, apical ones nearly black; pronotal disc dark brown, distinctly darker than the anterior, posterior and lateral regions; elytra largely brown, base of inner intervals darker, but not forming distinct pattern; venter and legs yellowish brown, tarsi slightly darker. Vertex with sparse fine punctures; postgenae with suborbital setae; mentum with a pair of median setae which are slightly shorter than the terminal labial palpomere. Pronotum sub-quadrate, PW/PL = 1.49, slightly wider than head, PW/HW = 1.07; lateral margins indistinctly angulate at anterior third, strongly sinuate before posterior angles; posterior angles nearly rectangular, slightly pointed outwards; lateral explanations wide; disc with very sparse fine punctures and transverse wrinkles. Elytra slightly dilated to apex; with very shallow isodiametric microsculpture, visible only near base; striae shallowly incised, with dense and fine puncture rows, punctures absent near elytral apices; intervals distinctly punctate, similar to punctures in striae, even intervals evidently convex near base; discal depression large and very shallow, nearly round, at anterior third of intervals 3 to 5; apical truncation rounded, outer apical angles completely rounded; sutural angles not pointed. Apex of abdominal sternite VII straight in females, with two setae on each side. Gonocoxite II of ovipositor nearly quadrate, length slightly greater than basal width, apex concave, inner apical angle strongly pointed; with two ensiform setae on outer and inner apical angles respectively. Male unknown.
</paragraph>
</subSubSection>
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<paragraph id="8B91C735623DFF982DBFB6FAFAEE59C6" blockId="123.[151,1437,325,1180]" box="[199,1289,1153,1180]" pageId="123" pageNumber="124">
<emphasis id="B95A1B27623DFF982DBFB6FAFEBE59C1" bold="true" box="[199,345,1153,1179]" pageId="123" pageNumber="124">Distribution</emphasis>
(
<figureCitation id="1315DBB0623DFF982C10B6F9FE2559C6" box="[360,450,1154,1180]" captionStart="MAP 12" captionStartId="120.[152,211,1374,1398]" captionTargetBox="[151,1436,700,1351]" captionTargetId="figure-240@120.[151,1436,700,1351]" captionText="MAP 12. Distributions for species in the Parena (Parena) stigmatica group" figureDoi="http://doi.org/10.5281/zenodo.7963697" httpUri="https://zenodo.org/record/7963697/files/figure.png" pageId="123" pageNumber="124">Map 12</figureCitation>
, green). Only known from the 
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locality Dundo in northeastern 
<collectingCountry id="F33987A5623DFF9829D6B6F9FAE359C6" box="[1198,1284,1154,1180]" name="Angola" pageId="123" pageNumber="124">Angola</collectingCountry>
.
</paragraph>
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</treatment>
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