treatments-xml/data/03/59/11/035911EC1AA158DBB7F7943D70F22D47.xml

<document ID-DOI="http://dx.doi.org/10.3897/asp.81.e81961" ID-Pensoft-Pub="1864-8312-81-79" ID-Pensoft-UUID="CA16EA0C5A33560FBC955981DFC9D502" ID-ZooBank="5C1FDC828F9F4869ADDD83FA96E507ED" ModsDocID="1864-8312-81-79" checkinTime="1674309819629" checkinUser="pensoft" docAuthor="Domahovski, Alexandre Cruz &amp; Cavichioli, Rodney Ramiro" docDate="2023" docId="035911EC1AA158DBB7F7943D70F22D47" docLanguage="en" docName="ArthropodSystPhyl 81: 79-164" docOrigin="Arthropod Systematics &amp; amp; Phylogeny 81" docPubDate="2023-01-20" docSource="http://dx.doi.org/10.3897/asp.81.e81961" docTitle="Coarctana striata Domahovski &amp; Cavichioli 2023, sp. nov." docType="treatment" docUuid="B32E1D93-4E16-4DC8-BBDE-6835F171450C" docUuidSource="ZooBank" docVersion="1" id="CA16EA0C5A33560FBC955981DFC9D502" lastPageNumber="79" masterDocId="CA16EA0C5A33560FBC955981DFC9D502" masterDocTitle="Phylogenetic analysis and revision of the leafhopper genus Acuera DeLong &amp; Freytag (Hemiptera: Cicadellidae: Gyponini) based on morphological data" masterLastPageNumber="164" masterPageNumber="79" pageNumber="79" updateTime="1674309819629" updateUser="pensoft">
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<mods:title>Phylogenetic analysis and revision of the leafhopper genus Acuera DeLong &amp; Freytag (Hemiptera: Cicadellidae: Gyponini) based on morphological data</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Domahovski, Alexandre Cruz</mods:namePart>
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<mods:affiliation>Departamento de Zoologia, Setor de Ciencias Biologicas, Universidade Federal do Parana, Av. Cel. Francisco H. dos Santos, 100, Caixa Postal 19020, 81531 - 980, Curitiba, PR, Brasil</mods:affiliation>
<mods:nameIdentifier type="email">domahovskiac@yahoo.com.br</mods:nameIdentifier>
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<mods:namePart>Cavichioli, Rodney Ramiro</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-5996-0629</mods:nameIdentifier>
<mods:affiliation>Departamento de Zoologia, Setor de Ciencias Biologicas, Universidade Federal do Parana, Av. Cel. Francisco H. dos Santos, 100, Caixa Postal 19020, 81531 - 980, Curitiba, PR, Brasil</mods:affiliation>
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<mods:title>Arthropod Systematics &amp; amp; Phylogeny</mods:title>
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<mods:date>2023</mods:date>
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<mods:number>2023-01-20</mods:number>
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<mods:number>81</mods:number>
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<mods:url>http://dx.doi.org/10.3897/asp.81.e81961</mods:url>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/asp.81.e81961</mods:identifier>
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<paragraph pageId="0" pageNumber="79">
<taxonomicName LSID="https://zoobank.org/B32E1D93-4E16-4DC8-BBDE-6835F171450C" authority="Domahovski &amp; Cavichioli, 2023" authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" family="Cicadellidae" genus="Coarctana" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Coarctana striata" order="Hemiptera" pageId="0" pageNumber="79" rank="species" species="striata" status="sp. nov.">Coarctana striata</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="79">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="description">
<paragraph pageId="0" pageNumber="79">
<figureCitation captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Coarctana striata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure26" httpUri="https://binary.pensoft.net/fig/798179" pageId="0" pageNumber="79">Figs 26</figureCitation>
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Coarctana striata sp. nov., paratype female. A distal portion of abdomen, ventral view. B distal portion of abdomen, lateral view. C first valvifer and first valvula, lateral view. D apical portion of first valvula. E second valvula, lateral view. F apical portion of second valvula. G second valvifer and gonoplac, lateral view. H apical portion of gonoplac. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure27" httpUri="https://binary.pensoft.net/fig/798180" pageId="0" pageNumber="79">, 27</figureCitation>
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Species of Coarctana gen. nov., dorsal (left) and lateral (right) views. A, B C. punctata comb. nov., holotype female, NHRS GULI 000069783. C, D C. rana comb. nov., holotype male, OSU. E, F C. serpenta comb. nov., holotype male, AMNH. G, H C. striata sp. nov., holotype male, DZUP. I, J C. taurina sp. nov., holotype male, DZUP. K, L C. vilavelha sp. nov., holotype male, DZUP." figureDoi="10.3897/asp.81.e81961.figure32" httpUri="https://binary.pensoft.net/fig/798185" pageId="0" pageNumber="79">, 32G, H</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="diagnosis">
<paragraph pageId="0" pageNumber="79">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="79">
Pygofer (Fig. 
<figureCitation captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Coarctana striata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure26" httpUri="https://binary.pensoft.net/fig/798179" pageId="0" pageNumber="79">26G</figureCitation>
) with apex truncate. Style, in lateral view (Fig. 
<figureCitation captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Coarctana striata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure26" httpUri="https://binary.pensoft.net/fig/798179" pageId="0" pageNumber="79">26K</figureCitation>
), blade curved dorsally, ventral margin not serrated and with lateral carina. Aedeagus (Fig. 
<figureCitation captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Coarctana striata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure26" httpUri="https://binary.pensoft.net/fig/798179" pageId="0" pageNumber="79">26L, M</figureCitation>
) apodemal processes narrow basally, curved in right angle dorsally; ventral margin forming triangular lobe produced ventrally; posterior margin with transverse and parallel striae; shaft apex with pair of short processes, curved laterally and crossed over the posterior surface.
</paragraph>
<caption doi="10.3897/asp.81.e81961.figure26" httpUri="https://binary.pensoft.net/fig/798179" pageId="0" pageNumber="79" start="Figure 26" startId="F26">
<paragraph pageId="0" pageNumber="79">
<emphasis bold="true" pageId="0" pageNumber="79">Figure 26.</emphasis>
<taxonomicName authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" family="Cicadellidae" genus="Coarctana" kingdom="Animalia" lsidName="Coarctana striata" order="Hemiptera" pageId="0" pageNumber="79" rank="species" species="striata">
<emphasis italics="true" pageId="0" pageNumber="79">Coarctana striata</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="79">sp. nov.</emphasis>
, holotype male. 
<emphasis bold="true" pageId="0" pageNumber="79">A</emphasis>
head and thorax, dorsal view. 
<emphasis bold="true" pageId="0" pageNumber="79">B</emphasis>
head, ventral view. 
<emphasis bold="true" pageId="0" pageNumber="79">C</emphasis>
head and thorax, lateral view. 
<emphasis bold="true" pageId="0" pageNumber="79">D</emphasis>
forewing. 
<emphasis bold="true" pageId="0" pageNumber="79">E</emphasis>
sternite VIII, ventral view. 
<emphasis bold="true" pageId="0" pageNumber="79">F</emphasis>
valve, ventral view. 
<emphasis bold="true" pageId="0" pageNumber="79">G</emphasis>
pygofer, valve and subgenital plate, lateral view. 
<emphasis bold="true" pageId="0" pageNumber="79">H</emphasis>
subgenital plate, ventral view. 
<emphasis bold="true" pageId="0" pageNumber="79">I</emphasis>
connective, dorsal view. 
<emphasis bold="true" pageId="0" pageNumber="79">J</emphasis>
style, dorsal view. 
<emphasis bold="true" pageId="0" pageNumber="79">K</emphasis>
style, lateral view. 
<emphasis bold="true" pageId="0" pageNumber="79">L</emphasis>
aedeagus, lateral view. 
<emphasis bold="true" pageId="0" pageNumber="79">M</emphasis>
aedeagus, posterior view. Scale bars in mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="description">
<paragraph pageId="0" pageNumber="79">Measurements.</paragraph>
<paragraph pageId="0" pageNumber="79">Total length: holotype male 8.5 mm; paratypes, females (n = 3) 9.6-10.4 mm.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="description">
<paragraph pageId="0" pageNumber="79">Description.</paragraph>
<paragraph pageId="0" pageNumber="79">
Head, in frontal view (Fig. 
<figureCitation captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Coarctana striata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure26" httpUri="https://binary.pensoft.net/fig/798179" pageId="0" pageNumber="79">26B</figureCitation>
), frons 1.2 
<normalizedToken originalValue="×">x</normalizedToken>
wider than long; frontogenal suture distant from eye margin by less than half maximum width of clypeus; clypeus 1.4 
<normalizedToken originalValue="×">x</normalizedToken>
wider than long; lateral margins parallel; apex straight. Head, in lateral view (Fig. 
<figureCitation captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Coarctana striata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure26" httpUri="https://binary.pensoft.net/fig/798179" pageId="0" pageNumber="79">26C</figureCitation>
), crown-face transition thin with 4-5 transverse carinae; clypeus not inflated. Profemur AV row with 5-6 setae; PV row with 1 seta near apex. Protibia PD row with 4 setae and undifferentiated intercalary setae; PV row with 4 setae and undifferentiated intercalary setae. Metatibia PD, AD and AV rows with 23, 12 and 14-16 macrosetae respectively. Metatarsomere I 4 
<normalizedToken originalValue="×">x</normalizedToken>
longer than apical width; inner row of the ventral surface with 7-8 setae. Other characteristics as in generic description.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="coloration">
<paragraph pageId="0" pageNumber="79">Coloration.</paragraph>
<paragraph pageId="0" pageNumber="79">
Head and thorax (Figs 
<figureCitation captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Coarctana striata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure26" httpUri="https://binary.pensoft.net/fig/798179" pageId="0" pageNumber="79">26A, B</figureCitation>
, 
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Species of Coarctana gen. nov., dorsal (left) and lateral (right) views. A, B C. punctata comb. nov., holotype female, NHRS GULI 000069783. C, D C. rana comb. nov., holotype male, OSU. E, F C. serpenta comb. nov., holotype male, AMNH. G, H C. striata sp. nov., holotype male, DZUP. I, J C. taurina sp. nov., holotype male, DZUP. K, L C. vilavelha sp. nov., holotype male, DZUP." figureDoi="10.3897/asp.81.e81961.figure32" httpUri="https://binary.pensoft.net/fig/798185" pageId="0" pageNumber="79">32G, H</figureCitation>
) brown. Head (Fig. 
<figureCitation captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Coarctana striata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure26" httpUri="https://binary.pensoft.net/fig/798179" pageId="0" pageNumber="79">26A</figureCitation>
) with pair of small circular black spots, near posterior margin, behind ocellus; coronal suture black. Face (Fig. 
<figureCitation captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Coarctana striata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure26" httpUri="https://binary.pensoft.net/fig/798179" pageId="0" pageNumber="79">26B</figureCitation>
) yellowish; frons reddish-brown dorsally with pair of transverse black maculae below crown-face transition; black macula adjacent to antennal base. Pronotum (Fig. 
<figureCitation captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Coarctana striata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure26" httpUri="https://binary.pensoft.net/fig/798179" pageId="0" pageNumber="79">26A</figureCitation>
) with black punctures, irregular black maculae near anterior margin; proepimeron (Fig. 
<figureCitation captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Coarctana striata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure26" httpUri="https://binary.pensoft.net/fig/798179" pageId="0" pageNumber="79">26C</figureCitation>
) with black band below pronotal carina; lateral carina yellow. Mesonotum (Fig. 
<figureCitation captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Coarctana striata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure26" httpUri="https://binary.pensoft.net/fig/798179" pageId="0" pageNumber="79">26A</figureCitation>
) with pair of large black maculae near lateral angles and pair of small median black spots near scutoscutellar suture. Forewing (Fig. 
<figureCitation captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Coarctana striata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure26" httpUri="https://binary.pensoft.net/fig/798179" pageId="0" pageNumber="79">26D</figureCitation>
) with scattered dark-brown mottling, larger black maculae on apex of anal veins, cross veins of inner discal cell and base of first apical cell; veins brown, margined by black punctures on basal half of wing. Metatibia (Fig. 
<figureCitation captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Coarctana striata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure26" httpUri="https://binary.pensoft.net/fig/798179" pageId="0" pageNumber="79">26C</figureCitation>
) with cucullate bases of setae blacks.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="male terminalia">
<paragraph pageId="0" pageNumber="79">Male terminalia.</paragraph>
<paragraph pageId="0" pageNumber="79">
Sternite VIII (Fig. 
<figureCitation captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Coarctana striata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure26" httpUri="https://binary.pensoft.net/fig/798179" pageId="0" pageNumber="79">26E</figureCitation>
) 1.4 
<normalizedToken originalValue="×">x</normalizedToken>
wider than long; lateral margins parallel; posterior margin slightly sinuous. Valve (Fig. 
<figureCitation captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Coarctana striata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure26" httpUri="https://binary.pensoft.net/fig/798179" pageId="0" pageNumber="79">26F</figureCitation>
) 1.6 
<normalizedToken originalValue="×">x</normalizedToken>
wider than long; posterior margin rounded. Pygofer, in lateral view (Fig. 
<figureCitation captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Coarctana striata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure26" httpUri="https://binary.pensoft.net/fig/798179" pageId="0" pageNumber="79">26G</figureCitation>
), 1.7 
<normalizedToken originalValue="×">x</normalizedToken>
longer than maximum height; basodorsal process present but reduced; anteroventral margin broadly rounded; posterodorsal and posteroventral margins straight; apex truncate. Subgenital plate, in lateral view (Fig. 
<figureCitation captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Coarctana striata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure26" httpUri="https://binary.pensoft.net/fig/798179" pageId="0" pageNumber="79">26G</figureCitation>
), short, not reaching pygofer apex; in ventral view (Fig. 
<figureCitation captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Coarctana striata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure26" httpUri="https://binary.pensoft.net/fig/798179" pageId="0" pageNumber="79">26H</figureCitation>
), elongated, 3.8 
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide, maximum width near half length; dorsal surface with filiform setae near outer margin; lateral margins slightly rounded; apex weakly tapered and rounded. Connective (Fig. 
<figureCitation captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Coarctana striata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure26" httpUri="https://binary.pensoft.net/fig/798179" pageId="0" pageNumber="79">26I</figureCitation>
) almost as wide as long; anterior margin excavated; dorsal keel and stem reduced; stem wide. Style, in dorsal view (Fig. 
<figureCitation captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Coarctana striata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure26" httpUri="https://binary.pensoft.net/fig/798179" pageId="0" pageNumber="79">26J</figureCitation>
), with outer lobe truncated; in lateral view (Fig. 
<figureCitation captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Coarctana striata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure26" httpUri="https://binary.pensoft.net/fig/798179" pageId="0" pageNumber="79">26K</figureCitation>
), blade curved dorsally, wider near base and tapered toward apex; ventral margin not serrated, with a longitudinal carina; apex tapered, forming a small dorsal process. Aedeagus (Fig. 
<figureCitation captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Coarctana striata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure26" httpUri="https://binary.pensoft.net/fig/798179" pageId="0" pageNumber="79">26L, M</figureCitation>
) preatrium weakly developed; dorsal apodeme with dorsal margin straight, lateral margins slightly produced laterally; apodemal processes narrow basally, curved in right angle dorsally at basal third; ventral margin forming triangular lobe produced ventrally; posterior margin with transverse and parallel striae; apex tapered, subacute; shaft approximately cylindrical, symmetrical, strongly curved dorsally near base, with a basolateral expansion that surrounds the previous portion of the shaft, basal third slightly curved posterad; apical third straight; apex with pair of short processes, curved laterally and crossed over the posterior surface. Other characteristics as in generic description.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="female terminalia">
<paragraph pageId="0" pageNumber="79">Female terminalia.</paragraph>
<paragraph pageId="0" pageNumber="79">
Sternite VII (Fig. 
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Coarctana striata sp. nov., paratype female. A distal portion of abdomen, ventral view. B distal portion of abdomen, lateral view. C first valvifer and first valvula, lateral view. D apical portion of first valvula. E second valvula, lateral view. F apical portion of second valvula. G second valvifer and gonoplac, lateral view. H apical portion of gonoplac. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure27" httpUri="https://binary.pensoft.net/fig/798180" pageId="0" pageNumber="79">27A</figureCitation>
) 1.5 
<normalizedToken originalValue="×">x</normalizedToken>
wider than long; posterolateral angles produced as far as posterior margin; posterior margin excavated each side of emarginated median lobe which occupies the median third. Internal sternite VIII membranous. Pygofer (Fig. 
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Coarctana striata sp. nov., paratype female. A distal portion of abdomen, ventral view. B distal portion of abdomen, lateral view. C first valvifer and first valvula, lateral view. D apical portion of first valvula. E second valvula, lateral view. F apical portion of second valvula. G second valvifer and gonoplac, lateral view. H apical portion of gonoplac. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure27" httpUri="https://binary.pensoft.net/fig/798180" pageId="0" pageNumber="79">27A, B</figureCitation>
) 1.9 
<normalizedToken originalValue="×">x</normalizedToken>
longer than maximum height; ventral margin slightly rounded; dorsal margin straight; apex obliquely truncate. First valvifer (Fig. 
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Coarctana striata sp. nov., paratype female. A distal portion of abdomen, ventral view. B distal portion of abdomen, lateral view. C first valvifer and first valvula, lateral view. D apical portion of first valvula. E second valvula, lateral view. F apical portion of second valvula. G second valvifer and gonoplac, lateral view. H apical portion of gonoplac. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure27" httpUri="https://binary.pensoft.net/fig/798180" pageId="0" pageNumber="79">27C</figureCitation>
) triangular, 1.4 
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide; lateral and dorsal margins straight. First valvula (Fig. 
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Coarctana striata sp. nov., paratype female. A distal portion of abdomen, ventral view. B distal portion of abdomen, lateral view. C first valvifer and first valvula, lateral view. D apical portion of first valvula. E second valvula, lateral view. F apical portion of second valvula. G second valvifer and gonoplac, lateral view. H apical portion of gonoplac. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure27" httpUri="https://binary.pensoft.net/fig/798180" pageId="0" pageNumber="79">27C</figureCitation>
) 5.2 
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide; apex (Fig. 
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Coarctana striata sp. nov., paratype female. A distal portion of abdomen, ventral view. B distal portion of abdomen, lateral view. C first valvifer and first valvula, lateral view. D apical portion of first valvula. E second valvula, lateral view. F apical portion of second valvula. G second valvifer and gonoplac, lateral view. H apical portion of gonoplac. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure27" httpUri="https://binary.pensoft.net/fig/798180" pageId="0" pageNumber="79">27D</figureCitation>
) abruptly tapered and acute. Second valvula (Fig. 
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Coarctana striata sp. nov., paratype female. A distal portion of abdomen, ventral view. B distal portion of abdomen, lateral view. C first valvifer and first valvula, lateral view. D apical portion of first valvula. E second valvula, lateral view. F apical portion of second valvula. G second valvifer and gonoplac, lateral view. H apical portion of gonoplac. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure27" httpUri="https://binary.pensoft.net/fig/798180" pageId="0" pageNumber="79">27E</figureCitation>
) 3.8 
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide; strongly expanded near mid-length; dorsal protuberance weakly developed; dorsal margin (Fig. 
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Coarctana striata sp. nov., paratype female. A distal portion of abdomen, ventral view. B distal portion of abdomen, lateral view. C first valvifer and first valvula, lateral view. D apical portion of first valvula. E second valvula, lateral view. F apical portion of second valvula. G second valvifer and gonoplac, lateral view. H apical portion of gonoplac. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure27" httpUri="https://binary.pensoft.net/fig/798180" pageId="0" pageNumber="79">27F</figureCitation>
) with few minute rounded teeth; apex gradually tapered and triangular. Second valvifer (Fig. 
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Coarctana striata sp. nov., paratype female. A distal portion of abdomen, ventral view. B distal portion of abdomen, lateral view. C first valvifer and first valvula, lateral view. D apical portion of first valvula. E second valvula, lateral view. F apical portion of second valvula. G second valvifer and gonoplac, lateral view. H apical portion of gonoplac. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure27" httpUri="https://binary.pensoft.net/fig/798180" pageId="0" pageNumber="79">27G</figureCitation>
) 2.5 
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide. Gonoplac (Fig. 
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Coarctana striata sp. nov., paratype female. A distal portion of abdomen, ventral view. B distal portion of abdomen, lateral view. C first valvifer and first valvula, lateral view. D apical portion of first valvula. E second valvula, lateral view. F apical portion of second valvula. G second valvifer and gonoplac, lateral view. H apical portion of gonoplac. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure27" httpUri="https://binary.pensoft.net/fig/798180" pageId="0" pageNumber="79">27G</figureCitation>
) 3.7 
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide; apex (Fig. 
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Coarctana striata sp. nov., paratype female. A distal portion of abdomen, ventral view. B distal portion of abdomen, lateral view. C first valvifer and first valvula, lateral view. D apical portion of first valvula. E second valvula, lateral view. F apical portion of second valvula. G second valvifer and gonoplac, lateral view. H apical portion of gonoplac. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure27" httpUri="https://binary.pensoft.net/fig/798180" pageId="0" pageNumber="79">27H</figureCitation>
) rounded. Other characteristics as in generic description.
</paragraph>
<caption doi="10.3897/asp.81.e81961.figure27" httpUri="https://binary.pensoft.net/fig/798180" pageId="0" pageNumber="79" start="Figure 27" startId="F27">
<paragraph pageId="0" pageNumber="79">
<emphasis bold="true" pageId="0" pageNumber="79">Figure 27.</emphasis>
<taxonomicName authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" family="Cicadellidae" genus="Coarctana" kingdom="Animalia" lsidName="Coarctana striata" order="Hemiptera" pageId="0" pageNumber="79" rank="species" species="striata">
<emphasis italics="true" pageId="0" pageNumber="79">Coarctana striata</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="79">sp. nov.</emphasis>
, paratype female. 
<emphasis bold="true" pageId="0" pageNumber="79">A</emphasis>
distal portion of abdomen, ventral view. 
<emphasis bold="true" pageId="0" pageNumber="79">B</emphasis>
distal portion of abdomen, lateral view. 
<emphasis bold="true" pageId="0" pageNumber="79">C</emphasis>
first valvifer and first valvula, lateral view. 
<emphasis bold="true" pageId="0" pageNumber="79">D</emphasis>
apical portion of first valvula. 
<emphasis bold="true" pageId="0" pageNumber="79">E</emphasis>
second valvula, lateral view. 
<emphasis bold="true" pageId="0" pageNumber="79">F</emphasis>
apical portion of second valvula. 
<emphasis bold="true" pageId="0" pageNumber="79">G</emphasis>
second valvifer and gonoplac, lateral view. 
<emphasis bold="true" pageId="0" pageNumber="79">H</emphasis>
apical portion of gonoplac. Scale bars in mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="materials_examined">
<paragraph pageId="0" pageNumber="79">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="79">
<materialsCitation collectingDate="2012-08-10" collectingDateMax="2012-08-17" collectingDateMin="2012-08-10" collectorName="Lima, H. I. L." country="Brazil" latitude="-10.2491665" location="Tocantins" longLatPrecision="15" longitude="-48.1265" specimenCount="1" stateProvince="Tocantins" typeStatus="Holotype">
<emphasis bold="true" pageId="0" pageNumber="79">
<typeStatus>Holotype</typeStatus>
male: 
<collectingCountry name="Brazil">Brazil</collectingCountry>
: 
<collectingRegion country="Brazil" name="Tocantins">Tocantins</collectingRegion>
</emphasis>
: &quot;Brasil, Tocantins, Palmas\ 
<normalizedToken originalValue="Taquaruçu">Taquarucu</normalizedToken>
, Fazenda\ Encantada 
<geoCoordinate degrees="10" direction="south" minutes="14" orientation="latitude" precision="15" seconds="57" value="-10.2491665">10°14′57″S</geoCoordinate>
<geoCoordinate degrees="48" direction="west" minutes="07" orientation="longitude" precision="1" seconds="35.4" value="-48.1265">48°07′35.4″W</geoCoordinate>
Malaise\ 
<collectingDate value="2012-08-10" valueMax="2012-08-17" valueMin="2012-08-10">10-17.viii.2012</collectingDate>
Krolow,\ T.K. &amp; 
<collectorName>Lima, H.I.L.</collectorName>
leg.&quot; (DZUP)
</materialsCitation>
. 
<materialsCitation collectingDate="2012-06-15" collectingDateMax="2012-06-22" collectingDateMin="2012-06-15" collectorName="Lima, H. I. L." country="Brazil" latitude="-10.2491665" location="Tocantins" longLatPrecision="15" longitude="-48.1265" specimenCount="1" specimenCount-female="1" stateProvince="Paratypes" typeStatus="Paratypes">
<emphasis bold="true" pageId="0" pageNumber="79">
<typeStatus>Paratypes</typeStatus>
</emphasis>
: 
<specimenCount type="female">1♀</specimenCount>
, same data as holotype except 
<collectingDate value="2012-06-15" valueMax="2012-06-22" valueMin="2012-06-15">15-22.vi.2012</collectingDate>
(DZUP)
</materialsCitation>
. - 
<materialsCitation collectingDate="2010-11-01" collectingDateMax="2010-11-06" collectingDateMin="2010-11-01" collectorName="F. Limei, D. W. A. Mar, E. A. S. Barbosa" country="Brazil" location="Maranhao" specimenCount="1" specimenCount-female="1" stateProvince="Maranhao" typeStatus="paratype">
<emphasis bold="true" pageId="0" pageNumber="79">
<normalizedToken originalValue="Maranhão">
<collectingRegion country="Brazil" name="Maranhao">Maranhao</collectingRegion>
</normalizedToken>
</emphasis>
: 
<specimenCount type="female">1♀</specimenCount>
, &quot;Brasil (MA), Bom Jardim\ REBIO-Res. Biol Gurupi\ Armadilha Malaise&quot;, &quot; 
<collectingDate value="2010-11-01" valueMax="2010-11-06" valueMin="2010-11-01">01-06.xi.2010</collectingDate>
, 
<collectorName>F.Limei</collectorName>
\ ra-de-Oliveira, 
<collectorName>D.W.A. Mar</collectorName>
\ ques &amp; 
<collectorName>E.A. S. Barbosa</collectorName>
&quot; (CZMA)
</materialsCitation>
; 
<materialsCitation collectingDate="2010-09-02" collectingDateMax="2010-09-11" collectingDateMin="2010-09-02" collectorName="E. A. S. Barbosa, J. A. Silva, M. M. Abreu" country="Brazil" specimenCount="1" specimenCount-female="1" stateProvince="Maranhao" typeStatus="paratype">
<specimenCount type="female">1♀</specimenCount>
, &quot;Brasil (MA), Bom Jardim\ REBIO-Res. Biol Gurupi\ Armadilha Malaise&quot;, &quot; 
<collectingDate value="2010-09-02" valueMax="2010-09-11" valueMin="2010-09-02">02-11.ix.2010</collectingDate>
, D.W.A.\ Marques, 
<collectorName>E.A. S. Barbosa</collectorName>
\ 
<collectorName>J.A. Silva</collectorName>
&amp; 
<collectorName>M.M. Abreu</collectorName>
&quot; (CZMA)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="etymology">
<paragraph pageId="0" pageNumber="79">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="79">The new species name refers to the aedeagus with apodemal processes bearing many transverse striae.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="remarks">
<paragraph pageId="0" pageNumber="79">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="79">
<taxonomicName authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" family="Cicadellidae" genus="Coarctana" kingdom="Animalia" lsidName="Coarctana striata" order="Hemiptera" pageId="0" pageNumber="79" rank="species" species="striata">
<emphasis italics="true" pageId="0" pageNumber="79">Coarctana striata</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="79">sp. nov.</emphasis>
(Fig. 
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Phylogenetic relationships of Acuera found in the implied weighting analysis for the two equally most parsimonious trees, K = 3.700 and 4.284, length = 514, consistency index = 0.362, retention index = 0.771. Symmetric resampling values (&lt;50) are marked next to branches; red stars represent the type species of the three subgenera of Acuera. Synapomorphies are marked with black circles, while homoplasies are marked with empty circles." figureDoi="10.3897/asp.81.e81961.figure7" httpUri="https://binary.pensoft.net/fig/798160" pageId="0" pageNumber="79">7</figureCitation>
) was recovered with low support (SR = 54) as sister to the clade 
<taxonomicName authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" family="Cicadellidae" kingdom="Animalia" lsidName="C. taurina" order="Hemiptera" pageId="0" pageNumber="79" rank="species" species="taurina">
<emphasis italics="true" pageId="0" pageNumber="79">C. taurina</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="79">sp. nov.</emphasis>
+ 
<taxonomicName authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" family="Cicadellidae" kingdom="Animalia" lsidName="C. oricula" order="Hemiptera" pageId="0" pageNumber="79" rank="species" species="oricula">
<emphasis italics="true" pageId="0" pageNumber="79">C. oricula</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="79">sp. nov.</emphasis>
and the clade of 
<taxonomicName authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" family="Cicadellidae" kingdom="Animalia" lsidName="C. vilavelha" order="Hemiptera" pageId="0" pageNumber="79" rank="species" species="vilavelha">
<emphasis italics="true" pageId="0" pageNumber="79">C. vilavelha</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="79">sp. nov.</emphasis>
+ (
<taxonomicName authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" family="Cicadellidae" kingdom="Animalia" lsidName="C. nigromedia" order="Hemiptera" pageId="0" pageNumber="79" rank="species" species="nigromedia">
<emphasis italics="true" pageId="0" pageNumber="79">C. nigromedia</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="79">sp. nov.</emphasis>
+ 
<taxonomicName authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" family="Cicadellidae" kingdom="Animalia" lsidName="C. asymmetrica" order="Hemiptera" pageId="0" pageNumber="79" rank="species" species="asymmetrica">
<emphasis italics="true" pageId="0" pageNumber="79">C. asymmetrica</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="79">sp. nov.</emphasis>
). 
<taxonomicName authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" family="Cicadellidae" genus="Coarctana" kingdom="Animalia" lsidName="Coarctana striata" order="Hemiptera" pageId="0" pageNumber="79" rank="species" species="striata">
<emphasis italics="true" pageId="0" pageNumber="79">Coarctana striata</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="79">sp. nov.</emphasis>
is easily separated from its congeners by the aedeagus (Fig. 
<figureCitation captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Coarctana striata sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure26" httpUri="https://binary.pensoft.net/fig/798179" pageId="0" pageNumber="79">26L, M</figureCitation>
) with apodemal transversely striated on posterior margin and the shaft with pair of short apical processes, curved laterally and crossed over the posterior surface of shaft.
</paragraph>
</subSubSection>
</treatment>
</document>