186 lines
22 KiB
XML
186 lines
22 KiB
XML
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<mods:title id="63FA33BEFA83D73EFF2A4918BF69C32C">Arboreal eurymeline leafhoppers (Hemiptera: Cicadellidae: Eurymelinae) of the Indian subcontinent with description of new genera and eight new species</mods:title>
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<mods:namePart id="8682663F0EE63A78BB45EF5699ECAE0A">VIRAKTAMATH, C. A.</mods:namePart>
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<mods:namePart id="4BE1C9FBF0847575FC750400158070FD">YESHWANTH, H. M.</mods:namePart>
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<mods:title id="DB518ADB9228C7C048B53992296939EF">Zootaxa</mods:title>
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<mods:date id="D795F71F4CAAF3CC6FF92D0901D8D1AD">2024</mods:date>
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<mods:url id="A426E697283410E480B41B015DA3BCBC">http://dx.doi.org/10.11646/zootaxa.5462.1.1</mods:url>
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<treatment id="4D73DA02FFA3FFE0FF17F9BBEDCDFF1F" ID-DOI="http://doi.org/10.5281/zenodo.11627015" ID-GBIF-Taxon="232743677" ID-Zenodo-Dep="11627015" LSID="urn:lsid:plazi:treatment:4D73DA02FFA3FFE0FF17F9BBEDCDFF1F" httpUri="http://treatment.plazi.org/id/4D73DA02FFA3FFE0FF17F9BBEDCDFF1F" lastPageId="57" lastPageNumber="58" pageId="54" pageNumber="55">
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<subSubSection id="8DC0389FFFA3FFEFFF17F9BBEA4CF9AD" box="[151,531,1585,1612]" pageId="54" pageNumber="55" type="nomenclature">
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<paragraph id="C5656B14FFA3FFEFFF17F9BBEA4CF9AD" blockId="54.[151,531,1585,1612]" box="[151,531,1585,1612]" pageId="54" pageNumber="55">
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<emphasis id="F7AEB706FFA3FFEFFF17F9BBEA4CF9AD" bold="true" box="[151,531,1585,1612]" pageId="54" pageNumber="55">
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Genus
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<taxonomicName id="02DA1097FFA3FFEFFF6EF9B8EA4CF9AD" ID-CoL="8HL45" authority="Viraktamath" authorityName="Viraktamath" box="[238,531,1585,1612]" class="Insecta" family="Cicadellidae" genus="Jogocerus" kingdom="Animalia" order="Hemiptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="genus">
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<emphasis id="F7AEB706FFA3FFEFFF6EF9B8E935F9AA" bold="true" box="[238,362,1586,1612]" italics="true" pageId="54" pageNumber="55">Jogocerus</emphasis>
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Viraktamath
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</taxonomicName>
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="8DC0389FFFA3FFEFFF17F9FDEC99F968" box="[151,1222,1654,1678]" pageId="54" pageNumber="55" type="reference_group">
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<paragraph id="C5656B14FFA3FFEFFF17F9FDEC99F968" blockId="54.[151,1222,1654,1678]" box="[151,1222,1654,1678]" pageId="54" pageNumber="55">
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<treatmentCitationGroup id="E5CA4C3AFFA3FFEFFF17F9FDEC99F968" box="[151,1222,1654,1678]" pageId="54" pageNumber="55">
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<treatmentCitation id="447B4D05FFA3FFEFFF17F9FDE9AFF968" author="Viraktamath, C. A." box="[151,496,1654,1678]" page="17" pageId="54" pageNumber="55" year="1979">
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<taxonomicName id="02DA1097FFA3FFEFFF17F9FDE9AFF968" ID-CoL="8HL45" authority="Viraktamath 1979 a: 17" authorityName="Viraktamath" authorityPageNumber="17" authorityYear="1979" box="[151,496,1654,1678]" class="Insecta" family="Cicadellidae" genus="Jogocerus" kingdom="Animalia" order="Hemiptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="genus">
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||
<emphasis id="F7AEB706FFA3FFEFFF17F9FDE8A0F968" box="[151,255,1655,1678]" italics="true" pageId="54" pageNumber="55">Jogocerus</emphasis>
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<bibRefCitation id="A14B16E5FFA3FFEFFE85F9FCE9AFF968" author="Viraktamath, C. A." box="[261,496,1654,1678]" pageId="54" pageNumber="55" pagination="17 - 26" refId="ref60730" refString="Viraktamath, C. A. (1979 a) Jogocerus gen. nov. and new species of idiocerine leafhoppers from southern India (Homoptera: Cicadellidae). Entomon, 4, 17 - 26." type="journal article" year="1979">Viraktamath 1979a: 17</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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<typeStatus id="1A61D5B6FFA3FFEFFE79F9FCEA74F968" box="[505,555,1654,1678]" pageId="54" pageNumber="55">Type</typeStatus>
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species:
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<taxonomicName id="02DA1097FFA3FFEFFD09F9FDEB94F968" ID-CoL="3QPYV" authority="Viraktamath" authorityName="Viraktamath" box="[649,971,1654,1678]" class="Insecta" family="Cicadellidae" genus="Jogocerus" kingdom="Animalia" order="Hemiptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="freytagi">
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<emphasis id="F7AEB706FFA3FFEFFD09F9FDEB1AF968" box="[649,837,1654,1678]" italics="true" pageId="54" pageNumber="55">Jogocerus freytagi</emphasis>
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Viraktamath
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</taxonomicName>
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, by original designation.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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||
<subSubSection id="8DC0389FFFA3FFE1FF17F937ED64FEFB" lastPageId="56" lastPageNumber="57" pageId="54" pageNumber="55" type="diagnosis">
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||
<paragraph id="C5656B14FFA3FFE1FF17F937ED64FEFB" blockId="54.[151,1437,1724,2039]" lastBlockId="56.[151,1437,151,285]" lastPageId="56" lastPageNumber="57" pageId="54" pageNumber="55">
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<emphasis id="F7AEB706FFA3FFEFFF17F937E956F930" box="[151,265,1725,1750]" italics="true" pageId="54" pageNumber="55">Diagnosis</emphasis>
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. Among the genera with a rugose pronotum from the
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<taxonomicName id="02DA1097FFA3FFEFFCF7F936EBE0F930" box="[887,959,1724,1750]" class="Aves" family="Phasianidae" genus="Indian" kingdom="Animalia" order="Galliformes" pageId="54" pageNumber="124" phylum="Chordata" rank="genus">Indian</taxonomicName>
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subcontinent (namely
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<emphasis id="F7AEB706FFA3FFEFFB44F937ED34F930" box="[1220,1387,1725,1750]" italics="true" pageId="54" pageNumber="55">Tasnimoceerus</emphasis>
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and
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<emphasis id="F7AEB706FFA3FFEFFF17F96AE968F91C" box="[151,311,1760,1786]" italics="true" pageId="54" pageNumber="55">
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<taxonomicName id="02DA1097FFA3FFEFFF17F96AE970F91C" box="[151,303,1760,1786]" class="Insecta" family="Cicadellidae" genus="Nilgiriscopus" kingdom="Animalia" order="Hemiptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="genus">Nilgiriscopus</taxonomicName>
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)
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</emphasis>
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, it differs in the shape of the clypellus, forewing venation and male genitalia. From
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<taxonomicName id="02DA1097FFA3FFEFFB6BF96BEDD9F91C" box="[1259,1414,1761,1786]" class="Insecta" family="Cicadellidae" genus="Tasnimocerus" kingdom="Animalia" order="Hemiptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="genus">
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<emphasis id="F7AEB706FFA3FFEFFB6BF96BEDD9F91C" box="[1259,1414,1761,1786]" italics="true" pageId="54" pageNumber="55">Tasnimocerus</emphasis>
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</taxonomicName>
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it differs in having the clypellus narrow at base (broad at base in
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<taxonomicName id="02DA1097FFA3FFEFFCB9F88FEB8BF8F8" box="[825,980,1797,1822]" class="Insecta" family="Cicadellidae" genus="Tasnimocerus" kingdom="Animalia" order="Hemiptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="genus">
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||
<emphasis id="F7AEB706FFA3FFEFFCB9F88FEB8BF8F8" box="[825,980,1797,1822]" italics="true" pageId="54" pageNumber="55">Tasnimocerus</emphasis>
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</taxonomicName>
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), outer (third) subapical cell less than half as long as second subapical cells and forewing veins uniformaly colored (outer subapical cell more than half as long as third and forewing veins alternately marked with dark brown and white in
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<taxonomicName id="02DA1097FFA3FFEFFC39F8C7EC0BF880" box="[953,1108,1869,1894]" class="Insecta" family="Cicadellidae" genus="Tasnimocerus" kingdom="Animalia" order="Hemiptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="genus">
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||
<emphasis id="F7AEB706FFA3FFEFFC39F8C7EC0BF880" box="[953,1108,1869,1894]" italics="true" pageId="54" pageNumber="55">Tasnimocerus</emphasis>
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</taxonomicName>
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), dorsal apodeme of aedeagus well-developed (absent in
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<taxonomicName id="02DA1097FFA3FFEFFE42F8FBEA02F86C" box="[450,605,1905,1930]" class="Insecta" family="Cicadellidae" genus="Tasnimocerus" kingdom="Animalia" order="Hemiptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="genus">
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<emphasis id="F7AEB706FFA3FFEFFE42F8FBEA02F86C" box="[450,605,1905,1930]" italics="true" pageId="54" pageNumber="55">Tasnimocerus</emphasis>
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</taxonomicName>
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) and the style with ventral margin crenulate (smooth in
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<taxonomicName id="02DA1097FFA3FFEFFB59F8FBED2BF86C" box="[1241,1396,1905,1930]" class="Insecta" family="Cicadellidae" genus="Tasnimocerus" kingdom="Animalia" order="Hemiptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="genus">
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<emphasis id="F7AEB706FFA3FFEFFB59F8FBED2BF86C" box="[1241,1396,1905,1930]" italics="true" pageId="54" pageNumber="55">Tasnimocerus</emphasis>
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</taxonomicName>
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). It differs from
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<taxonomicName id="02DA1097FFA3FFEFFEA3F81EE9E5F848" box="[291,442,1940,1966]" class="Insecta" family="Cicadellidae" genus="Nilgiriscopus" kingdom="Animalia" order="Hemiptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="genus">
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<emphasis id="F7AEB706FFA3FFEFFEA3F81EE9E5F848" box="[291,442,1940,1966]" italics="true" pageId="54" pageNumber="55">Nilgiriscopus</emphasis>
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</taxonomicName>
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in having a narrow based clypellus (parallel sided in
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<taxonomicName id="02DA1097FFA3FFEFFB8BF81EECFDF848" box="[1035,1186,1940,1966]" class="Insecta" family="Cicadellidae" genus="Nilgiriscopus" kingdom="Animalia" order="Hemiptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="genus">
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<emphasis id="F7AEB706FFA3FFEFFB8BF81EECFDF848" box="[1035,1186,1940,1966]" italics="true" pageId="54" pageNumber="55">Nilgiriscopus</emphasis>
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</taxonomicName>
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), forewing with tthree subapical cells(two in
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<taxonomicName id="02DA1097FFA3FFEFFE08F832EA40F834" box="[392,543,1976,2002]" class="Insecta" family="Cicadellidae" genus="Nilgiriscopus" kingdom="Animalia" order="Hemiptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="genus">
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<emphasis id="F7AEB706FFA3FFEFFE08F832EA40F834" box="[392,543,1976,2002]" italics="true" pageId="54" pageNumber="55">Nilgiriscopus</emphasis>
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||
</taxonomicName>
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||
)and male anal collar process and pygofer ventral processes well-developed(absent in
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<taxonomicName id="02DA1097FFA3FFEFFF32F856E916F810" box="[178,329,2012,2038]" class="Insecta" family="Cicadellidae" genus="Nilgiriscopus" kingdom="Animalia" order="Hemiptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F7AEB706FFA3FFEFFF32F856E916F810" box="[178,329,2012,2038]" italics="true" pageId="54" pageNumber="55">Nilgiriscopus</emphasis>
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</taxonomicName>
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). Meta basitrsomere distal transverse row with 2 platellae flanked by one sta on either side (
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<figureCitation id="5DE17791FFA3FFEFFAADF856EDCFF810" box="[1325,1424,2012,2038]" captionStart="FIGURES 29" captionStartId="56.[151,264,1934,1958]" captionTargetBox="[163,1431,314,1896]" captionTargetId="figure-88@56.[151,1436,309,1911]" captionTargetPageId="56" captionText="FIGURES 29A–I. Jogocerus freytagi Viraktamath, holotype male: A–B, habitus, dorsal and lateral views; C, face; D, label data; E, metabasitarsomere, ventral view; F, male genital capsule, lateral view; G, styles, connective and aedeagus, ventral view; H, connective and aedeagus, ventral view; I, connective and aedeagus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.11610511" httpUri="https://zenodo.org/record/11610511/files/figure.png" pageId="54" pageNumber="55">Fig. 29E</figureCitation>
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). Female sternite VII twice as wide at base as long medially, narrowed posteriorly with posterior margin slightly concave medially (
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<figureCitation id="5DE17791FFADFFE1FEE9FF31E98FFF33" box="[361,464,187,213]" captionStart="FIGURES 61" captionStartId="108.[151,264,1695,1719]" captionTargetBox="[156,1430,186,1666]" captionTargetId="figure-22@108.[151,1435,181,1671]" captionTargetPageId="108" captionText="FIGURES 61A–R. Idiocerini of the Indian subcontinent, female sternite VII: A, Idiocerus cedarae; B, Metidioceus deodarae; C, M. gnezdilovai; D, Idiocerus sharmai; E, Idioscopus bimaculatus; F, I. clypealis. G, I. confuscous; H, I. irenae; I, I. nagpurensis; J, I. nitidulus; K, I. pretiosus; L, I. shillongensis; M, I. virescens; N, Jogocerus freytagi; O, Keralus webbi; P, Matangia capriliana; Q, M. inidca; R, M. thapai." figureDoi="http://doi.org/10.5281/zenodo.11610624" httpUri="https://zenodo.org/record/11610624/files/figure.png" pageId="56" pageNumber="57">Fig. 61N</figureCitation>
|
||
). Valvula I slightly curved, sculptured area occupying distal half length, strigae delicate, oblique and occupying dorsal 0.5 width of valvula (
|
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<figureCitation id="5DE17791FFADFFE1FD7AFF55EB0BFF1F" box="[762,852,223,249]" captionStart="FIGURES 68" captionStartId="115.[151,264,1771,1795]" captionTargetBox="[156,1430,276,1743]" captionTargetId="figure-17@115.[151,1435,271,1748]" captionTargetPageId="115" captionText="FIGURES 68A–P. Idiocerini of the Indian subcontinent, female valvula I, magnified view and valvula II, magnified view:A–D, Jogocerus freytagi; E–H, Keralus webbi; I–L, Matangia capriliana; M–P, M. indica." figureDoi="http://doi.org/10.5281/zenodo.11610646" httpUri="https://zenodo.org/record/11610646/files/figure.png" pageId="56" pageNumber="57">Figs 68</figureCitation>
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AB). Valvula II slightly curved, with toothed area occupying distal half length, teeth prominent, squre tipped, spaced less than their own width (
|
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<figureCitation id="5DE17791FFADFFE1FB2FFE89ED5AFEFB" box="[1199,1285,259,285]" captionStart="FIGURES 68" captionStartId="115.[151,264,1771,1795]" captionTargetBox="[156,1430,276,1743]" captionTargetId="figure-17@115.[151,1435,271,1748]" captionTargetPageId="115" captionText="FIGURES 68A–P. Idiocerini of the Indian subcontinent, female valvula I, magnified view and valvula II, magnified view:A–D, Jogocerus freytagi; E–H, Keralus webbi; I–L, Matangia capriliana; M–P, M. indica." figureDoi="http://doi.org/10.5281/zenodo.11610646" httpUri="https://zenodo.org/record/11610646/files/figure.png" pageId="56" pageNumber="57">Figs 68</figureCitation>
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CD).
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</paragraph>
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</subSubSection>
|
||
<caption id="91A53B9CFFA2FFEEFF17F81DEBCBF81E" ID-DOI="http://doi.org/10.5281/zenodo.11610508" ID-Zenodo-Dep="11610508" httpUri="https://zenodo.org/record/11610508/files/figure.png" pageId="55" pageNumber="56" startId="55.[151,264,1943,1967]" targetBox="[158,1429,186,1914]" targetPageId="55" targetType="figure">
|
||
<paragraph id="C5656B14FFA2FFEEFF17F81DEBCBF81E" blockId="55.[151,1436,1943,2040]" pageId="55" pageNumber="56">
|
||
<emphasis id="F7AEB706FFA2FFEEFF17F81DE907F849" bold="true" box="[151,344,1943,1967]" pageId="55" pageNumber="56">FIGURES 28A–I.</emphasis>
|
||
<taxonomicName id="02DA1097FFA2FFEEFEDFF812EA96F856" authority="(Viraktamath)" authorityName="VIRAKTAMATH & YESHWANTH" authorityYear="2024" baseAuthorityName="VIRAKTAMATH & YESHWANTH" baseAuthorityYear="2024" box="[351,713,1944,1968]" class="Insecta" family="Cicadellidae" genus="Jogina" kingdom="Animalia" order="Hemiptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="robustipennis" status="comb. nov.">
|
||
<emphasis id="F7AEB706FFA2FFEEFEDFF812EA6BF856" box="[351,564,1944,1968]" italics="true" pageId="55" pageNumber="56">Jogina robustipennis</emphasis>
|
||
(Viraktamath)
|
||
</taxonomicName>
|
||
<emphasis id="F7AEB706FFA2FFEEFD50F812EB1AF856" bold="true" box="[720,837,1944,1968]" pageId="55" pageNumber="56">
|
||
<taxonomicNameLabel id="EC9D0A7DFFA2FFEEFD50F812EB1AF856" box="[720,837,1944,1968]" pageId="55" pageNumber="56" rank="species">comb. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype male: A–B, habitus, dorsal and lateral views; C, face; D, label data; E, metabasitarsomere, ventral view; F, male genital capsule, lateral view; G, style, connective and aedeagus, dorsal view; H, style, lateral view; I, connective and aedeagus, lateral view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="91A53B9CFFADFFE1FF17F804EBBBF809" ID-DOI="http://doi.org/10.5281/zenodo.11610511" ID-Zenodo-Dep="11610511" httpUri="https://zenodo.org/record/11610511/files/figure.png" pageId="56" pageNumber="57" startId="56.[151,264,1934,1958]" targetBox="[163,1431,314,1896]" targetPageId="56" targetType="figure">
|
||
<paragraph id="C5656B14FFADFFE1FF17F804EBBBF809" blockId="56.[151,1436,1934,2031]" pageId="56" pageNumber="57">
|
||
<emphasis id="F7AEB706FFADFFE1FF17F804E906F840" bold="true" box="[151,345,1934,1958]" pageId="56" pageNumber="57">FIGURES 29A–I.</emphasis>
|
||
<taxonomicName id="02DA1097FFADFFE1FEE1F805EAF7F841" authority="Viraktamath" authorityName="Viraktamath" box="[353,680,1935,1959]" class="Insecta" family="Cicadellidae" genus="Jogocerus" kingdom="Animalia" order="Hemiptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="freytagi">
|
||
<emphasis id="F7AEB706FFADFFE1FEE1F805EA7FF841" box="[353,544,1935,1959]" italics="true" pageId="56" pageNumber="57">Jogocerus freytagi</emphasis>
|
||
Viraktamath
|
||
</taxonomicName>
|
||
, holotype male: A–B, habitus, dorsal and lateral views; C, face; D, label data; E, metabasitarsomere, ventral view; F, male genital capsule, lateral view; G, styles, connective and aedeagus, ventral view; H, connective and aedeagus, ventral view; I, connective and aedeagus, lateral view.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="8DC0389FFFACFFE0FF47FF1DEDCDFF1F" pageId="57" pageNumber="58" type="discussion">
|
||
<paragraph id="C5656B14FFACFFE0FF47FF1DEDCDFF1F" blockId="57.[151,1436,151,249]" pageId="57" pageNumber="58">
|
||
<emphasis id="F7AEB706FFACFFE0FF47FF1DE96DFF57" bold="true" box="[199,306,151,177]" pageId="57" pageNumber="58">Remarks</emphasis>
|
||
. Viraktamath (1979) has adequately described the genus. The genus at present contains only the
|
||
<typeStatus id="1A61D5B6FFACFFE0FAEDFF1DEDC3FF57" box="[1389,1436,151,177]" pageId="57" pageNumber="58">type</typeStatus>
|
||
species from the
|
||
<taxonomicName id="02DA1097FFACFFE0FECFFF31E9CAFF33" box="[335,405,187,213]" class="Aves" family="Phasianidae" genus="Indian" kingdom="Animalia" order="Galliformes" pageId="57" pageNumber="124" phylum="Chordata" rank="genus">Indian</taxonomicName>
|
||
subcontinent. However,
|
||
<bibRefCitation id="A14B16E5FFACFFE0FD23FF31EB2DFF33" author="Zhang, B. & Li, Z. Z." box="[675,882,187,213]" pageId="57" pageNumber="58" pagination="66 - 68" refId="ref62066" refString="Zhang, B. & Li, Z. Z. (2008) First report of the leafhopper genus Jogocerus Viraktamath (Hemiptera: Cicadellidae) from China, with description of one new species. Zootaxa, 1936 (1), 66 - 68. https: // doi. org / 10.11646 / zootaxa. 1936.1.4" type="journal article" year="2008">Zhang & Li (2008)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="A14B16E5FFACFFE0FC25FF31ECD6FF33" author="Xue, Q. Q. & Zhang, Y. L." box="[933,1161,187,213]" pageId="57" pageNumber="58" pagination="174 - 180" refId="ref61952" refString="Xue, Q. Q. & Zhang, Y. L. (2017) Two new species of Jogocerus Viraktamath (Hemiptera: Cicadellidae: Iiocerinae) from China. Zootaxa, 4247 (2), 174 - 180. https: // doi. org / 10.11646 / zootaxa. 4021.4.4" type="journal article" year="2017">Xue & Zhang (2017)</bibRefCitation>
|
||
described new species of the genus from
|
||
<collectingCountry id="BDCD2B84FFACFFE0FEC3FF55E9D7FF1F" box="[323,392,223,249]" name="China" pageId="57" pageNumber="58">China</collectingCountry>
|
||
. The genus now contains four species one from the
|
||
<taxonomicName id="02DA1097FFACFFE0FC40FF55EC59FF1F" box="[960,1030,223,249]" class="Aves" family="Phasianidae" genus="Indian" kingdom="Animalia" order="Galliformes" pageId="57" pageNumber="124" phylum="Chordata" rank="genus">Indian</taxonomicName>
|
||
subcontinent and three from
|
||
<collectingCountry id="BDCD2B84FFACFFE0FAC8FF55EDD2FF1F" box="[1352,1421,223,249]" name="China" pageId="57" pageNumber="58">China</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |