treatments-xml/data/1F/94/28/1F94286215D453C39B64BCE9C9965372.xml

<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.230.105878" ID-Pensoft-Pub="1314-2003-230-157" ID-Pensoft-UUID="2E95E2BCAF7955F8B2ABBD7F472DD9B0" ModsDocID="1314-2003-230-157" checkinTime="1691771898489" checkinUser="pensoft" docAuthor="Vigosa-Mercado, J. Luis, Delgado-Salinas, Alfonso, Alvarado Cardenas, Leonardo O. &amp; Eguiarte, Luis E." docDate="2023" docId="1F94286215D453C39B64BCE9C9965372" docLanguage="en" docName="PhytoKeys 230: 157-256" docOrigin="PhytoKeys 230" docPubDate="2023-08-11" docSource="http://dx.doi.org/10.3897/phytokeys.230.105878" docTitle="Agrostis turrialbae Mez, Repert. Spec. Nov. Regni Veg. 18 (1 – 3): 4. 1922." docType="treatment" docVersion="1" id="2E95E2BCAF7955F8B2ABBD7F472DD9B0" lastPageNumber="157" masterDocId="2E95E2BCAF7955F8B2ABBD7F472DD9B0" masterDocTitle="Revision of the genus Agrostis (Poaceae, Pooideae, Poeae) in Megamexico" masterLastPageNumber="256" masterPageNumber="157" pageNumber="157" updateTime="1691771898489" updateUser="pensoft">
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<mods:title id="C2C3E93AFAB7C99717B12D4396FED366">Revision of the genus Agrostis (Poaceae, Pooideae, Poeae) in Megamexico</mods:title>
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<mods:namePart id="6DCEBBBBDA66CD4A35937224359D4A8F">Vigosa-Mercado, J. Luis</mods:namePart>
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<mods:affiliation id="EA5E6745B4E75B4E30781BC31EC79B4B">Posgrado en Ciencias Biologicas, Universidad Nacional Autonoma de Mexico, Ciudad Universitaria, Av. Universidad 3000, Coyoacan, 04510, Cd. Mx., Mexico</mods:affiliation>
<mods:nameIdentifier id="E1C534D18BFF0D65D2D9217ABAE3873F" type="email">luis_vigosa@ciencias.unam.mx</mods:nameIdentifier>
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<mods:namePart id="117DBDB34EB1C6FCD18988BDE5618831">Delgado-Salinas, Alfonso</mods:namePart>
<mods:nameIdentifier id="CA6EACF34EC96AD6022832E15C843F4B" type="ORCID">https://orcid.org/0000-0002-9322-9968</mods:nameIdentifier>
<mods:affiliation id="25D187696238F36D784BC79A3BD84F1D">Posgrado en Ciencias Biologicas, Universidad Nacional Autonoma de Mexico, Ciudad Universitaria, Av. Universidad 3000, Coyoacan, 04510, Cd. Mx., Mexico</mods:affiliation>
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<mods:namePart id="9D430DB1D47505312C83168AF4EFDEBD">Alvarado Cardenas, Leonardo O.</mods:namePart>
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<mods:affiliation id="CC932156AF34B763B47796072F0195A7">Posgrado en Ciencias Biologicas, Universidad Nacional Autonoma de Mexico, Ciudad Universitaria, Av. Universidad 3000, Coyoacan, 04510, Cd. Mx., Mexico</mods:affiliation>
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<mods:namePart id="2C9D72CF8E01003E1BE4357A29D7AEA5">Eguiarte, Luis E.</mods:namePart>
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<mods:affiliation id="23209A1F189A2D741CA440B8BA1F07BF">Posgrado en Ciencias Biologicas, Universidad Nacional Autonoma de Mexico, Ciudad Universitaria, Av. Universidad 3000, Coyoacan, 04510, Cd. Mx., Mexico</mods:affiliation>
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<mods:date id="720D1D53E787528A14B94969DD8518B0">2023</mods:date>
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<subSubSection id="F99251E8732868EA015D817BA01DCB82" pageId="0" pageNumber="157" type="nomenclature">
<paragraph id="D3BBA65E09EEBF5B49FAA510428A1166" pageId="0" pageNumber="157">
19. 
<taxonomicName id="B38539B8652FF0CCEC3D7B31588BDD9D" ID-CoL="65TJN" LSID="1F942862-15D4-53C3-9B64-BCE9C9965372" authority="Mez, Repert. Spec. Nov. Regni Veg. 18 (1 - 3): 4. 1922." authorityName="Mez, Repert. Spec. Nov. Regni Veg. 18 (1 – 3): 4. 1922." class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis turrialbae" order="Poales" pageId="0" pageNumber="157" phylum="Tracheophyta" rank="species" species="turrialbae">Agrostis turrialbae Mez, Repert. Spec. Nov. Regni Veg. 18(1-3): 4. 1922.</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="62B22F1598652FA0D9B496ABF68287E2" pageId="0" pageNumber="157" type="description">
<paragraph id="7C0FF64AA2702EA69DEC15F292FD25B5" pageId="0" pageNumber="157">
<figureCitation id="DB446CD1FAA32B67E157376DF29FD727" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Florets of Agrostis species observed with SEM A A. bourgaei B A. calderoniae C A. capillaris D A. elliottiana E A. exarata F A. ghiesbreghtii G A. gigantea H A. hyemalis I A. laxissima J A. microphylla K A. pallens L A. perennans M A. scabra N A. stolonifera O A. subpatens P A. subrepens Q A. tolucensis R A. turrialbae S A. variabilis. Scale bars: 0.3 mm." figureDoi="10.3897/phytokeys.230.105878.figure4" httpUri="https://binary.pensoft.net/fig/891239" pageId="0" pageNumber="157">Figs 4R</figureCitation>
<figureCitation id="509FE740EA7506955EF7B467B60AA806" captionStart="Figure 30" captionStartId="F30" captionText="Figure 30. Calluses of Agrostis species observed with SEM A A. scabra B A. stolonifera C A. subpatens D A. subrepens E A. tolucensis F A. turrialbae G A. variabilis. Scale bars: 50 μm." figureDoi="10.3897/phytokeys.230.105878.figure30" httpUri="https://binary.pensoft.net/fig/891265" pageId="0" pageNumber="157">, 30F</figureCitation>
<figureCitation id="52C3C37E3C1EEA84343F8DC6F09C6D4B" captionStart="Figure 38" captionStartId="F38" captionText="Figure 38. Agrostis turrialbae A whole plant B ligular area C spikelet E floret, abaxial view F floret, adaxial view. Based on Guzman 5959 (MEXU). Scale bars: 3 cm (A); 1 mm (B); 0.5 mm (C); 0.2 mm (D, E)." figureDoi="10.3897/phytokeys.230.105878.figure38" httpUri="https://binary.pensoft.net/fig/891273" pageId="0" pageNumber="157">, 38</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="A541F54C9A2B3579ED0C2815D1037152" pageId="0" pageNumber="157" type="reference_group">
<paragraph id="E669EAD8E5F4449442B64910504A0CE4" pageId="0" pageNumber="157">
<taxonomicName id="EEE9B56C5248D2729CAC574B1E62E601" authorityName="Swallen" authorityYear="1950" class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis arcta" order="Poales" pageId="0" pageNumber="157" phylum="Tracheophyta" rank="species" species="arcta">= Agrostis arcta</taxonomicName>
Swallen, Contr. U.S. Natl. Herb. 29(9): 405. 1950. Type. Guatemala. Chimaltenango: moist roadside at Santa Elena, 17 Jul 1933, A.F. Skutch 422 (holotype: US (US00131720)).
</paragraph>
<paragraph id="E7DAA4143ACFEAEF71133EDB72EBCB51" pageId="0" pageNumber="157">
<taxonomicName id="69CF88FF60CE4CDF373B9B958A52C94F" authorityName="Swallen" authorityYear="1950" class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis vesca" order="Poales" pageId="0" pageNumber="157" phylum="Tracheophyta" rank="species" species="vesca">= Agrostis vesca</taxonomicName>
Swallen, Contr. U.S. Natl. Herb. 29(9): 405. 1950. Type. Guatemala. Chimaltenango: moist roadside at Santa Elena, 17 Jul 1933, A.F. Skutch 420 (holotype: US (US00131129)).
</paragraph>
</subSubSection>
<subSubSection id="E20CD48F16022BD7873680EA2DC62CB6" pageId="0" pageNumber="157" type="materials_examined">
<paragraph id="E9941131307E038EABB2DF43DC27EFD4" pageId="0" pageNumber="157">
<typeStatus id="FADCCBD798E854AE45F80BB6C79CB78F">Type</typeStatus>
.
</paragraph>
<paragraph id="55BAA3E5EE439453327EE3EF272C1FC6" pageId="0" pageNumber="157">
<materialsCitation id="809191D0EBB29BEC6D7459120E417B34" collectingDate="1884-01-27" collectorName="H. Pittier" country="US" elevation="2600" location="Turrialba" specimenCount="1" stateProvince="Cartago" typeStatus="holotype">
Costa Rica. 
<collectingRegion id="9F671808B8E8758B47901B3BE9BE3C68" country="Costa Rica" name="Cartago">Cartago</collectingRegion>
: plateau au field W du 
<location id="9D9370A94D39ADE66A9E20D01B9863B5" LSID="urn:lsid:plazi:treatment:1F94286215D453C39B64BCE9C9965372:9D9370A94D39ADE66A9E20D01B9863B5" country="US" name="Turrialba" stateProvince="Cartago">Turrialba</location>
, 
<quantity id="3431E51CEC33A8D8D254B5EC71FAD9DB" metricMagnitude="3" metricUnit="m" metricValue="2.6" unit="m" value="2600.0">
<elevation id="3E7AEF421EB9447F49758D6AC5274835" metricMagnitude="3" metricUnit="m" metricValue="2.6" unit="m" value="2600.0">2600 m</elevation>
</quantity>
alt., 
<collectingDate id="69CDC1EA8159AA51AFFDBAAFCA91E447" value="1884-01-27">27 Jan 1884</collectingDate>
, 
<collectorName id="45681FA497E777F1495D38DF6BA131F5">H. Pittier</collectorName>
855 (
<typeStatus id="1C9EB46EC9C65914DB6337FC1ADB012E">holotype</typeStatus>
: B; isotypes: US (
<collectingCountry id="679F467339DE0FA8128C3C5D551D020B" name="United States of America">US</collectingCountry>
00131127, US04023770 [image!]))
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="63DBA43E94B4722984F2E7E55B1529B9" pageId="0" pageNumber="157" type="description">
<paragraph id="B2BE297BE5CA0811E5062CA72D694BD3" pageId="0" pageNumber="157">Description.</paragraph>
<paragraph id="4376D82DB842EFD88A8A2F523FC8A44C" pageId="0" pageNumber="157">
<emphasis id="AACC6B60C06852D2F003D7DDEB9D02A2" bold="true" italics="true" pageId="0" pageNumber="157">Plants</emphasis>
perennial, caespitose. 
<emphasis id="253157E6AC79397E148F13EF154FB94A" bold="true" italics="true" pageId="0" pageNumber="157">Tillers</emphasis>
extravaginal, with cataphylls. 
<emphasis id="2761D950399C5E65B46096D213021CF2" bold="true" italics="true" pageId="0" pageNumber="157">Culms</emphasis>
13-32 cm long, erect, nodes 1-2, glabrous, internodes glabrous. 
<emphasis id="3EDB7245A2D5E5D3C4FF452545820A30" bold="true" italics="true" pageId="0" pageNumber="157">Leaves</emphasis>
mostly basal; sheaths 0.8-6.5(-10) cm long, usually longer than the internodes, glabrous or scaberulous; ligules 0.5-1.6 mm long, longer than wide, dorsally scaberulous, apices acute to rounded, erose or lacerate; blades 1-9 cm long, 0.2-0.5 mm wide, filiform, conduplicate to involute, rarely flat in the upper leaves, scaberulous on both surfaces. 
<emphasis id="69601C07312EBC19529EF338F4E85807" bold="true" italics="true" pageId="0" pageNumber="157">Panicles</emphasis>
3.8-10 cm long, 1.7-5(-8) cm wide, open, lax, ovate, exserted from the upper sheaths; branches ascending to spreading, rebranching about or slightly above mid-length, scaberulous, without spikelets near their base, inferior branches 0.7-2.5 cm long; pedicels 0.7-4 mm long, ascending to spreading, scaberulous. 
<emphasis id="B87A5F6DBBC09DED9F83C9254048A792" bold="true" italics="true" pageId="0" pageNumber="157">Spikelets</emphasis>
1.5-2.4 mm long, purplish; glumes subequal to unequal, lanceolate, apices acute, 1-veined, scaberulous on the keel, lower glume 1.5-2.4 mm long, upper glume 1.4-2.3 mm long; callus pubescent, with 2 bunches of trichomes; lemmas 1.3-1.8 mm long, elliptic, apices entire, acute, sometimes irregularly toothed, 5-veined, veins prominent, unawned, rarely awned near the apices, awn ca. 0.2 mm long, straight; paleas absent or up to 0.2 mm long, veinless, glabrous; anthers 3, ca. 0.7 mm long. 
<emphasis id="2D60E1A28A3E44332D0112E6A31EA949" bold="true" italics="true" pageId="0" pageNumber="157">Caryopsis</emphasis>
1.5-2.2 mm elliptic; endosperm solid. 2n= unknown.
</paragraph>
<caption id="52E3906CDD4601B84275E5870EABFEB8" doi="10.3897/phytokeys.230.105878.figure38" httpUri="https://binary.pensoft.net/fig/891273" pageId="0" pageNumber="157" start="Figure 38" startId="F38">
<paragraph id="8A26DDA71EE4542F3B088571E56B88F2" pageId="0" pageNumber="157">
<emphasis id="1546439BACEAF5128B4941D68B7F7135" bold="true" pageId="0" pageNumber="157">Figure 38.</emphasis>
<taxonomicName id="8C7676C3405C2BED74EF1F661A8757CE" class="Liliopsida" family="Gramineae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis turrialbae" order="Poales" pageId="0" pageNumber="157" phylum="Tracheophyta" rank="species" species="turrialbae">
<emphasis id="924F3ED724E8CB7C1127F6A6F9827E1D" italics="true" pageId="0" pageNumber="157">Agrostis turrialbae</emphasis>
</taxonomicName>
<emphasis id="170596F3F8FE940F0A38F606E0E1F400" bold="true" pageId="0" pageNumber="157">A</emphasis>
whole plant 
<emphasis id="AE8FE526155FA1C2AECA606B60537C1D" bold="true" pageId="0" pageNumber="157">B</emphasis>
ligular area 
<emphasis id="ED3CA45CC843CA32DE69C5FE44A99CF5" bold="true" pageId="0" pageNumber="157">C</emphasis>
spikelet 
<emphasis id="CD6DB7C9EA73A9667640F33294215797" bold="true" pageId="0" pageNumber="157">E</emphasis>
floret, abaxial view 
<emphasis id="8C493720848E44FC8D361C476BBD96B2" bold="true" pageId="0" pageNumber="157">F</emphasis>
floret, adaxial view. Based on 
<normalizedToken id="2F7EF81467913DDBAD926951AFA19A90" originalValue="Guzmán">Guzman</normalizedToken>
5959 (MEXU). Scale bars: 3 cm (
<emphasis id="414769E0E5E78CFC71169B28D4EA91D6" bold="true" pageId="0" pageNumber="157">A</emphasis>
); 1 mm (
<emphasis id="263E71365735DC15953D8913140EFCC6" bold="true" pageId="0" pageNumber="157">B</emphasis>
); 0.5 mm (
<emphasis id="FD5D0BE1C6F2B00E1BD6C276826BE398" bold="true" pageId="0" pageNumber="157">C</emphasis>
); 0.2 mm (
<emphasis id="938E3795E0A1853507CB7FB00D3FAAA0" bold="true" pageId="0" pageNumber="157">D, E</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="D73793BC3B6179F9F2F491B12315C76C" pageId="0" pageNumber="157" type="anatomy and micromorphology">
<paragraph id="5C67519CAF74220959377DA22F9B6E54" pageId="0" pageNumber="157">Anatomy and micromorphology.</paragraph>
<paragraph id="70C52E436DA4BAA326942711C5839E5F" pageId="0" pageNumber="157">
Leaf blades v-shaped to involute in transversal section; adaxial furrows deep, narrow; adaxial ribs rounded to triangular; keel absent; first order bundles circular in outline, sheath interrupted abaxially, abaxial and adaxial sclerenchyma in strands; second order bundles circular in outline, sheath interrupted abaxially, abaxial and adaxial sclerenchyma in strands; intercostal sclerenchyma present, abaxial; leaf margins with well-developed sclerenchyma caps, rounded; colorless cells absent (Fig. 
<figureCitation id="12E66AB726B7C3EF3B5574463D3EAB7C" captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Leaf blade anatomy in transversal section of Agrostis species, in general view, and details of lateral bundles. A, B A. tolucensis C, D A. turrialbae E, F A. variabilis. Scale bars: 0.1 mm." figureDoi="10.3897/phytokeys.230.105878.figure37" httpUri="https://binary.pensoft.net/fig/891272" pageId="0" pageNumber="157">37C, D</figureCitation>
). Lemmas with transversal thickenings irregular to oblong, wider than the unthickened portion of the wall; prickle hairs present, abundant (Fig. 
<figureCitation id="39DDF7536C4FDD8A06E25CBDDE96A556" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Lemma surface of Agrostis observed with SEM A A. scabra B A. stolonifera C A. subpatens D A. subrepens E A. tolucensis F A. turrialbae G A. variabilis. Scale bars: 15 μm." figureDoi="10.3897/phytokeys.230.105878.figure32" httpUri="https://binary.pensoft.net/fig/891267" pageId="0" pageNumber="157">32F</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="DB1C0F4C77F0A6A22EF01DF8A274BF41" pageId="0" pageNumber="157" type="distribution">
<paragraph id="060B51C5B0094C4475FDE70D39552C04" pageId="0" pageNumber="157">Distribution and habitat.</paragraph>
<paragraph id="3DE3FDEE963062C4B96627BB4A49B685" pageId="0" pageNumber="157">
<taxonomicName id="A38E65CF4713194A6B50586AFC552738" class="Liliopsida" family="Gramineae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis turrialbae" order="Poales" pageId="0" pageNumber="157" phylum="Tracheophyta" rank="species" species="turrialbae">
<emphasis id="F3B0581D44F9A156EC78D3D9CD72BE8D" italics="true" pageId="0" pageNumber="157">Agrostis turrialbae</emphasis>
</taxonomicName>
is distributed from central Mexico to Costa Rica. It has also been reported from Colombia and Venezuela (
<bibRefCitation id="EF3DC2844D13893A8720F6949E38CA12" author="Luteyn, JL" journalOrPublisher="Comision para el Conocimiento y Uso de la Biodiversidad, Mexico City" pageId="0" pageNumber="157" refId="B38" refString="Luteyn, JL, 1999. Paramos, a checklist of plant diversity, geographical distribution, and botanical literature. Memoirs of the New York Botanical Garden (Vol. 84). New Work Botanical Garden, New York, 1-278." title="Paramos, a checklist of plant diversity, geographical distribution, and botanical literature. Memoirs of the New York Botanical Garden (Vol. 84). New Work Botanical Garden, New York, 1 - 278." year="1999">Luteyn 1999</bibRefCitation>
; 
<bibRefCitation id="15128CCB33A6BDE147882CCBC7254FB6" author="Soreng, RJ" editor="Soreng, RJ" journalOrPublisher="Contributions from the United States National Herbarium" pageId="0" pageNumber="157" pagination="42 - 89" publicationUrl="https://archive.org/details/mobot39088011697067" refId="B64" refString="Soreng, RJ, Peterson, PM, 2003. Agrostis. In: Soreng, RJ, Peterson, PM, Davidse, G, Judziewicz, EJ, Zuloaga, FO, Filgueiras, TS, Morrone, O, Eds., Catalogue of New World Grasses (Poaceae): IV. Subfamily Pooideae. Contributions from the United States National Herbarium 48: 42 - 89, https://archive.org/details/mobot39088011697067" title="Agrostis." url="https://archive.org/details/mobot39088011697067" volume="48" volumeTitle="Catalogue of New World Grasses (Poaceae): IV. Subfamily Pooideae." year="2003">Soreng and Peterson 2003</bibRefCitation>
). In the study zone, this species has been collected in the Mexican states of Chiapas, 
<normalizedToken id="C43A78BB929B62CFE9A03F868BAA6EE9" originalValue="México">Mexico</normalizedToken>
, 
<normalizedToken id="574309B53E179357A50751C60BEBA733" originalValue="Querétaro">Queretaro</normalizedToken>
, Tlaxcala, and Veracruz, and the Guatemalan departments of Huehuetenango and San Marcos (Fig. 
<figureCitation id="C6EF76FD42F2C90C7D48438A367AD900" captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Map of known geographic distribution of Agrostis species, based on herbarium specimen data A A. turrialbae B A. variabilis." figureDoi="10.3897/phytokeys.230.105878.figure39" httpUri="https://binary.pensoft.net/fig/891274" pageId="0" pageNumber="157">39A</figureCitation>
). It has also been reported from Hidalgo (
<bibRefCitation id="D094BD3BF7AE8EE9BA3C112D6E91845C" DOI="https://doi.org/10.21829/abm126.2019.1379" author="Sanchez-Ken, JG" journalOrPublisher="PhytoKeys" pageId="0" pageNumber="157" refId="B61" refString="Sanchez-Ken, JG, 2019. Riqueza de especies, clasificacion y listado de las gramineas (Poaceae) de Mexico. Acta Botanica Mexicana 126: e1379. https://doi.org/10.21829/abm126.2019.1379" title="Riqueza de especies, clasificacion y listado de las gramineas (Poaceae) de Mexico. Acta Botanica Mexicana 126: e 1379." url="https://doi.org/10.21829/abm126.2019.1379" year="2019">
<normalizedToken id="3FC35E7CBB879AB226D9F7CCD09DBFCD" originalValue="Sánchez-Ken">Sanchez-Ken</normalizedToken>
2019
</bibRefCitation>
), but no specimens from this state were seen. 
<taxonomicName id="CA77535822BAD561FA0EEAD4897A64C4" class="Liliopsida" family="Gramineae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis turrialbae" order="Poales" pageId="0" pageNumber="157" phylum="Tracheophyta" rank="species" species="turrialbae">
<emphasis id="AB3F16F52A69F1AD79D0D6B88B50A569" italics="true" pageId="0" pageNumber="157">Agrostis turrialbae</emphasis>
</taxonomicName>
grows in open areas of temperate forests, with 
<taxonomicName id="2C684C4E5294FF7FD49191A3D625AEBF" class="Pinopsida" family="Pinaceae" genus="Pinus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Pinus" order="Pinales" pageId="0" pageNumber="157" phylum="Coniferophyta" rank="genus">
<emphasis id="83BA85F3A0876720A7C383776B42C023" italics="true" pageId="0" pageNumber="157">Pinus</emphasis>
</taxonomicName>
and 
<taxonomicName id="90187D28AD6696629A3875A0ADFF705B" class="Dicotyledoneae" family="Fagaceae" genus="Quercus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Quercus" order="Fagales" pageId="0" pageNumber="157" phylum="Angiospermae" rank="genus">
<emphasis id="913BCB2946684E5D4FDDB03FD566D9BF" italics="true" pageId="0" pageNumber="157">Quercus</emphasis>
</taxonomicName>
, and alpine grasslands, between 1600-4240 m a.s.l. (Fig. 
<figureCitation id="D79D283A011D14D7D52B2727A0F55CE5" captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Elevation histograms of Agrostis species A A. perennans B A. scabra C A. stolonifera D A. subpatens E A. subrepens F A. tolucensis G A. turrialbae H A. variabilis." figureDoi="10.3897/phytokeys.230.105878.figure27" httpUri="https://binary.pensoft.net/fig/891262" pageId="0" pageNumber="157">27G</figureCitation>
).
</paragraph>
<caption id="963B8578431216024B24FCCEC72FB03A" doi="10.3897/phytokeys.230.105878.figure39" httpUri="https://binary.pensoft.net/fig/891274" pageId="0" pageNumber="157" start="Figure 39" startId="F39">
<paragraph id="B58430C32A2C3CCB404CF0BC7DEAE05E" pageId="0" pageNumber="157">
<emphasis id="FEE174901831FCE7A761267B4F26EA12" bold="true" pageId="0" pageNumber="157">Figure 39.</emphasis>
Map of known geographic distribution of 
<taxonomicName id="9BBA8A168436FCA12974B89E501A0D76" class="Liliopsida" family="Gramineae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis" order="Poales" pageId="0" pageNumber="157" phylum="Tracheophyta" rank="genus">
<emphasis id="59652DD7EB03847EEAF401CDAE283632" italics="true" pageId="0" pageNumber="157">Agrostis</emphasis>
</taxonomicName>
species, based on herbarium specimen data 
<emphasis id="511C193A20BCF5F81F0AA894201EDF8F" bold="true" pageId="0" pageNumber="157">A</emphasis>
<taxonomicName id="F1AEAC98B8A1DFF18F9D0BDEB6A27C4A" family="Poaceae" kingdom="Plantae" lsidName="A. turrialbae" order="Poales" pageId="0" pageNumber="157" rank="species" species="turrialbae">
<emphasis id="8EEC704E2D744BAAB00AC3782379616F" italics="true" pageId="0" pageNumber="157">A. turrialbae</emphasis>
</taxonomicName>
<emphasis id="EAB0D0B14E4ABD647CF394109925540D" bold="true" pageId="0" pageNumber="157">B</emphasis>
<taxonomicName id="B4B289C9619C4BDAE08CD3B9799137C5" family="Poaceae" kingdom="Plantae" lsidName="A. variabilis" order="Poales" pageId="0" pageNumber="157" rank="species" species="variabilis">
<emphasis id="6D7D335110BC80415F0D4BAF573A7FB8" italics="true" pageId="0" pageNumber="157">A. variabilis</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="F36A78AFC56F69B1558ABB7919AA9D7B" pageId="0" pageNumber="157" type="phenology">
<paragraph id="D33264605AE931D0ECFAB8FE1B412DA5" pageId="0" pageNumber="157">Phenology.</paragraph>
<paragraph id="8B240C651D028CDB0A39330EDFDC1F8C" pageId="0" pageNumber="157">
Specimens with spikelets have been collected from June to February (Fig. 
<figureCitation id="BA2439D3A22063D401CD4BB17DC65242" captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Phenology histograms of Agrostis species A A. perennans B A. scabra C A. stolonifera D A. subpatens E A. subrepens F A. tolucensis G A. turrialbae H A. variabilis." figureDoi="10.3897/phytokeys.230.105878.figure28" httpUri="https://binary.pensoft.net/fig/891263" pageId="0" pageNumber="157">28G</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="B0B8E15786A9FF15B0CE66C67BD5526C" pageId="0" pageNumber="157" type="comments">
<paragraph id="FCD4D75934C1E9828A0D8B396355D186" pageId="0" pageNumber="157">Commentaries.</paragraph>
<paragraph id="55F1BAE530D33BAC7913829B3C527020" pageId="0" pageNumber="157">
Populations of this species from central Mexico have been confused with 
<taxonomicName id="AB88F7F8FFC5171EA203E836B2C81FC5" family="Poaceae" kingdom="Plantae" lsidName="A. subpatens" order="Poales" pageId="0" pageNumber="157" rank="species" species="subpatens">
<emphasis id="87D25FB861E42C7384EFF2A4AFD2B0CC" italics="true" pageId="0" pageNumber="157">A. subpatens</emphasis>
</taxonomicName>
by some authors (e.g., 
<bibRefCitation id="B1F005FC88EBA63397B72EBCF8FE03A3" author="Acosta, S" editor="Calderon de Rzedowski, G" journalOrPublisher=") Instituto de Ecologia, A. C., Comision Nacional para el Conocimiento y uso de la Biodiversidad, Patzcuaro, Michoacan" pageId="0" pageNumber="157" pagination="1014 - 1017" publicationUrl="https://www.biodiversidad.gob.mx/publicaciones/librosDig/pdf/Indice_Flora_del_Valle_de_Mx.pdf" refId="B1" refString="Acosta, S, 2001. Agrostis. In: Calderon de Rzedowski, G, Rzedowski, J, Eds., Flora fanerogamica del Valle de Mexico (2nd edn.) Instituto de Ecologia, A.C., Comision Nacional para el Conocimiento y uso de la Biodiversidad, Patzcuaro, Michoacan: 1014 - 1017, https://www.biodiversidad.gob.mx/publicaciones/librosDig/pdf/Indice_Flora_del_Valle_de_Mx.pdf" title="Agrostis." url="https://www.biodiversidad.gob.mx/publicaciones/librosDig/pdf/Indice_Flora_del_Valle_de_Mx.pdf" volumeTitle="Flora fanerogamica del Valle de Mexico (2 nd edn." year="2001">Acosta 2001</bibRefCitation>
; 
<bibRefCitation id="47ACDA18D8F7D27B063D916F18BA6163" DOI="https://doi.org/10.21829/fb.311.2020.219" author="Vigosa-Mercado, JL" editor="Rzedowski, J" journalOrPublisher="219) Instituto de Ecologia, A. C., Patzcuaro, Mexico" pageId="0" pageNumber="157" pagination="1 - 214" refId="B77" refString="Vigosa-Mercado, JL, Ruiz-Sanchez, E, 2020. Familia Gramineae, Subfamilia Pooideae. In: Rzedowski, J, Hernandez-Ledesma, P, Eds., Flora del Bajio y de Regiones Adyacentes (Vol. 219) Instituto de Ecologia, A.C., Patzcuaro, Mexico: 1 - 214, DOI: https://doi.org/10.21829/fb.311.2020.219" title="Familia Gramineae, Subfamilia Pooideae." url="https://doi.org/10.21829/fb.311.2020.219" volumeTitle="Flora del Bajio y de Regiones Adyacentes (Vol." year="2020">
Vigosa-Mercado and 
<normalizedToken id="9BBC69890E7BBAE4D0F34EF730782F73" originalValue="Ruiz-Sánchez">Ruiz-Sanchez</normalizedToken>
2020
</bibRefCitation>
) (see the note under the description of that species). 
<taxonomicName id="E494CEB96F6915F9B75A4E0E3DF48D6C" class="Liliopsida" family="Gramineae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis turrialbae" order="Poales" pageId="0" pageNumber="157" phylum="Tracheophyta" rank="species" species="turrialbae">
<emphasis id="A9807F08B3C72DD6DC4AFBDF2630CA6E" italics="true" pageId="0" pageNumber="157">Agrostis turrialbae</emphasis>
</taxonomicName>
is similar to 
<taxonomicName id="20E6C22DB5F725156A54B5B1900A1DBA" family="Poaceae" kingdom="Plantae" lsidName="A. perennans" order="Poales" pageId="0" pageNumber="157" rank="species" species="perennans">
<emphasis id="7429EE6310CEDE464ACEE8102E56BD10" italics="true" pageId="0" pageNumber="157">A. perennans</emphasis>
</taxonomicName>
sensu lato, and its identity has been put in doubt recently by 
<bibRefCitation id="A463A8921E6623D5871BC82989338CEB" DOI="https://doi.org/10.3897/phytokeys.151.50538" author="Sylvester, SP" journalOrPublisher="PhytoKeys" pageId="0" pageNumber="157" pagination="107 - 160" refId="B66" refString="Sylvester, SP, Cuta-Alarcon, LE, Bravo-Pedraza, WJ, Soreng, RJ, 2020a. Agrostis and Podagrostis from paramos of Boyaca, Colombia: Synoptic taxonomy including a key to Colombian species. PhytoKeys 151: 107 - 160, DOI: https://doi.org/10.3897/phytokeys.151.50538" title="Agrostis and Podagrostis from paramos of Boyaca, Colombia: Synoptic taxonomy including a key to Colombian species." url="https://doi.org/10.3897/phytokeys.151.50538" volume="151" year="2020 a">Sylvester et al. (2020a)</bibRefCitation>
. Both species share open panicles and unawned lemmas, as well as several lemma micromorphology characters. In the study zone, it has been found that the plants identified as 
<taxonomicName id="AE3C219A25416B32F74D1D12F71B41AD" family="Poaceae" kingdom="Plantae" lsidName="A. turrialbae" order="Poales" pageId="0" pageNumber="157" rank="species" species="turrialbae">
<emphasis id="58262BFD200AC3FE57E25CE793D443C6" italics="true" pageId="0" pageNumber="157">A. turrialbae</emphasis>
</taxonomicName>
are consistently different from 
<taxonomicName id="38757936242F555668D169634CF965B0" family="Poaceae" kingdom="Plantae" lsidName="A. perennans" order="Poales" pageId="0" pageNumber="157" rank="species" species="perennans">
<emphasis id="0C84F918A9A04E8FD6BBFBF426F2C0DF" italics="true" pageId="0" pageNumber="157">A. perennans</emphasis>
</taxonomicName>
, and are characterized by the small size of the plants, mostly basal leaves, leaf blades narrow and conduplicated to involute, with adaxial ribs rounded to triangular, with abaxial intercostal sclerenchyma, and purplish spikelets (vs. usually larger plants, usually basal and cauline leaves, but the basal ones often drying at anthesis, leaf blades wider and flat, with rounded adaxial ribs, without intercostal sclerenchyma, greenish to purplish spikelets in 
<taxonomicName id="CFF27C63C28D91095E23B4130C2A4444" family="Poaceae" kingdom="Plantae" lsidName="A. perennans" order="Poales" pageId="0" pageNumber="157" rank="species" species="perennans">
<emphasis id="D39BA556E4CD779D25840E411C417482" italics="true" pageId="0" pageNumber="157">A. perennans</emphasis>
</taxonomicName>
). This species is scarcely different from 
<taxonomicName id="7A741F52886780AD1F72D9F5A159A740" family="Poaceae" kingdom="Plantae" lsidName="A. idahoensis" order="Poales" pageId="0" pageNumber="157" rank="species" species="idahoensis">
<emphasis id="4EC6D5D05AA4C8924ECC7FD38166F416" italics="true" pageId="0" pageNumber="157">A. idahoensis</emphasis>
</taxonomicName>
, from the southern USA (see the note under the description of that species).
</paragraph>
</subSubSection>
<subSubSection id="5881842BA9219A129205FBBA71572BB4" pageId="0" pageNumber="157" type="conservation status">
<paragraph id="5D7A4341DABABDEB4BD4CAACBC2AB7C7" pageId="0" pageNumber="157">Conservation status.</paragraph>
<paragraph id="DEA6329FA04666AFE0CBAD70A4DB89AA" pageId="0" pageNumber="157">
<taxonomicName id="E6BD3AF1DE18059ED17C5B86F3E487E7" class="Liliopsida" family="Gramineae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis turrialbae" order="Poales" pageId="0" pageNumber="157" phylum="Tracheophyta" rank="species" species="turrialbae">
<emphasis id="7017316CBE6A1EA85510443B333471E0" italics="true" pageId="0" pageNumber="157">Agrostis turrialbae</emphasis>
</taxonomicName>
is a widespread species in the study zone. It is represented by 30 collections, with several populations occurring in six protected areas. The EOO is 183,426 km2 and the AOO is 84 km2. Following the IUCN criteria, the preliminary assessment category is Least Concern (LC).
</paragraph>
</subSubSection>
<subSubSection id="0721D109A6A77D739D05101462FEA4FE" pageId="0" pageNumber="157" type="materials_examined">
<paragraph id="223FE2DF8F1450338558B7B56A43286E" pageId="0" pageNumber="157">Representative specimens examined.</paragraph>
<paragraph id="4C4F20C2BB46CC9BED1AF068E5F0CB6E" pageId="0" pageNumber="157">
<materialsCitation id="8E566BEC47B5057611AFC9ACD03E81FD" collectingDate="1940-02-13" collectingDateMax="1942-08-02" collectingDateMin="1940-02-13" collectorName="J. A. Steyermark" country="Guatemala" county="Municipio Santa Eulalia" elevation="3125" latitude="15.048245" location="Chemalito" longLatPrecision="1" longitude="-91.86726" municipality="Santa Eulalia" specimenCount="1" stateProvince="Huehuetenango">
<emphasis id="02E150509BC5E5AFFF4C36FEF3C88F93" bold="true" pageId="0" pageNumber="157">
<collectingCountry id="74A80285045F43CE892A5E09037C5D4C" name="Guatemala">Guatemala</collectingCountry>
. 
<collectingRegion id="9206528878AD83F32654E91AC81DBB77" country="Guatemala" name="Huehuetenango">Huehuetenango</collectingRegion>
: 
<collectingCounty id="D77F927DEC3CF6800759C1C8BDC05415">
Municipio 
<collectingMunicipality id="67D250CDC34A266C3E8ADBE361765C01">Santa Eulalia</collectingMunicipality>
</collectingCounty>
</emphasis>
, top of cerro 
<location id="C2DA5467B7286DB763CFF1EF09237133" LSID="urn:lsid:plazi:treatment:1F94286215D453C39B64BCE9C9965372:C2DA5467B7286DB763CFF1EF09237133" country="Guatemala" county="Municipio Santa Eulalia" latitude="15.048245" longLatPrecision="1" longitude="-91.86726" municipality="Santa Eulalia" name="Chemalito" stateProvince="Huehuetenango">Chemalito</location>
, 
<location id="D2AFB53AAFAC63AA972A8ABAA799A5B8" LSID="urn:lsid:plazi:treatment:1F94286215D453C39B64BCE9C9965372:D2AFB53AAFAC63AA972A8ABAA799A5B8" country="Guatemala" county="Municipio Santa Eulalia" latitude="15.048245" longLatPrecision="1" longitude="-91.86726" municipality="Santa Eulalia" name="Sierra" stateProvince="Huehuetenango">Sierra</location>
de los 
<location id="F6A151840544EC6D19684EAEA8C03613" LSID="urn:lsid:plazi:treatment:1F94286215D453C39B64BCE9C9965372:F6A151840544EC6D19684EAEA8C03613" country="Guatemala" county="Municipio Santa Eulalia" latitude="15.048245" longLatPrecision="1" longitude="-91.86726" municipality="Santa Eulalia" name="Cuchumatanes" stateProvince="Huehuetenango">Cuchumatanes</location>
, 
<locationDeviation id="6C76340DBAEEB1EBCC71CE15E29FEE99" location="Santa Eulalia">
<quantity id="512F3B22298156AEE511DEDE8FEFE8CA" metricMagnitude="3" metricUnit="m" metricValue="5.632704" unit="mi" value="3.5">3.5 mi</quantity>
W of Santa Eulalia
</locationDeviation>
, [
<geoCoordinate id="9ADBD49EC1B630FB0F3DB1591CC8C4EE" degrees="15.7822223" direction="north" orientation="latitude" precision="1" value="15.782223">15.7822223°N</geoCoordinate>
, 
<geoCoordinate id="709A9ECB29263B627880DFBD6E401C7A" degrees="91.5111111" direction="west" orientation="longitude" precision="1" value="-91.51111">91.5111111°W</geoCoordinate>
], 
<quantity id="8BDB31E99B7362E96A96AB643CA7A6DD" metricMagnitude="3" metricUnit="m" metricValue="3.125" metricValueMax="3.15" metricValueMin="3.1" unit="m" value="3125.0" valueMax="3150.0" valueMin="3100.0">
<elevation id="4DE324BB3D5720115562D2E1C6AF7A79" metricMagnitude="3" metricUnit="m" metricValue="3.125" metricValueMax="3.15" metricValueMin="3.1" unit="m" value="3125.0" valueMax="3150.0" valueMin="3100.0">3100-3150 m</elevation>
</quantity>
alt., 
<collectingDate id="D47C4F8CF75E56C486CEE97DDDF6C6F1" value="1942-08-02">2 Aug 1942</collectingDate>
, 
<collectorName id="F8DB90381CCE54FB0691F2DBF9EA26B1">J.A. Steyermark</collectorName>
49905 (F, US). 
<emphasis id="32F7A1392E4E4DF0B38D992A08989BFE" bold="true" pageId="0" pageNumber="157">
<collectingRegion id="CE54213563AD899B5CFF44918F0B8425" country="Guatemala" name="San Marcos">San Marcos</collectingRegion>
: Municipio 
<collectingRegion id="35D2D87168DE21CAA14D6B2A40734281" country="Guatemala" name="San Marcos">San Marcos</collectingRegion>
</emphasis>
, between San 
<normalizedToken id="8D3851CCA540E7AC58724DF659339392" originalValue="Sebastián">Sebastian</normalizedToken>
and summit of 
<normalizedToken id="6C60D9B9E6F216FDE7227BB4DAA0E43A" originalValue="Volcán">Volcan</normalizedToken>
Tajumulco, [
<geoCoordinate id="2A7F8AC2970622F6F0AF87D9643B72B6" degrees="15.04824546" direction="north" orientation="latitude" precision="1" value="15.048245">15.04824546°N</geoCoordinate>
, 
<geoCoordinate id="821F2ACDF1C51B69119241E5FEBE6E04" degrees="91.86726456" direction="west" orientation="longitude" precision="1" value="-91.86726">91.86726456°W</geoCoordinate>
, 
<quantity id="9613289D4874867677E3BA7BD5321080" metricMagnitude="3" metricUnit="m" metricValue="3.353" unit="m" value="3353.0">
<elevation id="350E6B04AA4FD1DAA960B582EAAB852F" metricMagnitude="3" metricUnit="m" metricValue="3.353" unit="m" value="3353.0">3353 m</elevation>
</quantity>
alt.], 
<collectingDate id="B5FF153CCFF6DA23E4D49054BB6CF897" value="1940-02-13">13 Feb 1940</collectingDate>
, 
<collectorName id="DEF1ECF2669B69816F408D6D417CFFBD">J.A. Steyermark</collectorName>
35477 (F)
</materialsCitation>
. 
<materialsCitation id="9E6DA8CBE4603ADA0DE0687B2A6974E3" collectingDate="1981-11-24" collectorName="D. E. Breedlove, B. M. Bartholomew" country="Mexico" elevation="2750" latitude="15.419722" location="Municipio Motozintla" longLatPrecision="1" longitude="-92.33667" specimenCount="1" stateProvince="Chiapas">
<emphasis id="6CB6929CA081EFA974A5E60D875FC8C5" bold="true" pageId="0" pageNumber="157">
<collectingCountry id="A3185F049DC8D2E05CED1C42413FC124" name="Mexico">Mexico</collectingCountry>
. 
<collectingRegion id="93C00D842759BB69EB581F6515E1D0CD" country="Mexico" name="Chiapas">Chiapas</collectingRegion>
: 
<location id="26E9FD9D14DD017B8816511DB943874B" LSID="urn:lsid:plazi:treatment:1F94286215D453C39B64BCE9C9965372:26E9FD9D14DD017B8816511DB943874B" country="Mexico" latitude="15.419722" longLatPrecision="1" longitude="-92.33667" name="Municipio Motozintla" stateProvince="Chiapas">Municipio Motozintla</location>
</emphasis>
, near summit of cerro Mozotal., [
<geoCoordinate id="BAA40C5DF2531013F33E3B86E46CDF9F" degrees="15.419722" direction="north" orientation="latitude" precision="1" value="15.419722">15.419722°N</geoCoordinate>
, 
<geoCoordinate id="A0D5DD163372F7DED94010D15023B001" degrees="92.336667" direction="west" orientation="longitude" precision="1" value="-92.33667">92.336667°W</geoCoordinate>
], 
<quantity id="77E6CC5ACFB0542CA3A945391097D2D1" metricMagnitude="3" metricUnit="m" metricValue="2.75" unit="m" value="2750.0">
<elevation id="5CDAC799D120DEB9EFF15DD7527DAB5C" metricMagnitude="3" metricUnit="m" metricValue="2.75" unit="m" value="2750.0">2750 m</elevation>
</quantity>
alt., 
<collectingDate id="17F346CD06D570A52C1308896990D9A1" value="1981-11-24">24 Nov 1981</collectingDate>
, 
<collectorName id="1CAC8D3159D78B8D4BFA73C12425227E">D.E. Breedlove</collectorName>
and 
<collectorName id="DCD47F032C340EB72B08536841F09999">B.M. Bartholomew</collectorName>
55845 (MO)
</materialsCitation>
. 
<materialsCitation id="BA37A8F2EBB02003B35BECBDB96F4959" collectingDate="1981-11-24" collectingDateMax="2014-11-08" collectingDateMin="1981-11-24" collectorName="R. Hernandez, S. Villalobos, El Zamorano, A. Mora, J. Ramirez, R. Guzman, R. Hernandez-Cardenas, L. Arredondo-Amezcua, La Cuchilla, de Orizaba, See Suppl." country="Mexico" county="Municipio Amecameca" elevation="4050" latitude="19.067286" location="Portillo" longLatPrecision="1" longitude="-97.19152" municipality="Iztaccihuatl" specimenCount="1" stateProvince="Queretaro">
<emphasis id="4158471A1C8C3EB26CDDCB9ADC80A502" bold="true" pageId="0" pageNumber="157">
<normalizedToken id="D3F5566EC72EE4CD8B227F506530A9B4" originalValue="México">
<collectingCountry id="07F4F2469F8A8EA44E55A6EAB491C2BB" name="Mexico">Mexico</collectingCountry>
</normalizedToken>
: 
<collectingCounty id="6A86E6FC4B97A27A14C136402718AE0E">Municipio Amecameca</collectingCounty>
</emphasis>
: ladera SW del 
<normalizedToken id="C723D5CEAAC8E263074102FF13DFAC06" originalValue="volcán">volcan</normalizedToken>
<normalizedToken id="70AA5A9C2C2D0B8F2BBAF016D0E89ADE" originalValue="Iztaccíhuatl">
<collectingMunicipality id="0EB8EB355F16A7BE2C82D2A953FD0815">Iztaccihuatl</collectingMunicipality>
</normalizedToken>
, rumbo al primer 
<location id="3698665410B7450B7F62AA0256B31288" LSID="urn:lsid:plazi:treatment:1F94286215D453C39B64BCE9C9965372:3698665410B7450B7F62AA0256B31288" country="Mexico" county="Municipio Amecameca" latitude="19.067286" longLatPrecision="1" longitude="-97.19152" municipality="Iztaccihuatl" name="Portillo" stateProvince="Queretaro">Portillo</location>
, 
<geoCoordinate id="E7577830BFF99D15D152E7A2B5DE3389" degrees="19.13697222" direction="north" orientation="latitude" precision="1" value="19.136972">19.13697222°N</geoCoordinate>
, 
<geoCoordinate id="DB288F2D8A97FF70A75A5F7A4A41D1FB" degrees="98.64898611" direction="west" orientation="longitude" precision="1" value="-98.64899">98.64898611°W</geoCoordinate>
, 
<quantity id="CEB9EB80D801EEE914F8156BCC6A35EE" metricMagnitude="3" metricUnit="m" metricValue="4.05" unit="m" value="4050.0">
<elevation id="270C9350A634E4D318E08AC8CDBA652C" metricMagnitude="3" metricUnit="m" metricValue="4.05" unit="m" value="4050.0">4050 m</elevation>
</quantity>
alt., 
<collectingDate id="10BA233FAA6718D260AEAF589BA8C8C9" value="2014-11-08">8 Nov 2014</collectingDate>
, 
<collectorName id="33D5F2C1F132CCE05C7DA1DAAE0A11FD">
R. 
<normalizedToken id="24D625EFB95E065C5C5EDEA543B5BAAC" originalValue="Hernández">Hernandez</normalizedToken>
</collectorName>
and 
<collectorName id="D9447DB22FB5CA625596C3B3E32702F0">S. Villalobos</collectorName>
2057 (IEB, MEXU [**]). 
<emphasis id="D30C8D5E48E3B2899471D84A44FF9579" bold="true" pageId="0" pageNumber="157">
<normalizedToken id="60C2DD8898AE042FCE53AF673DE0B9E7" originalValue="Querétaro">
<collectingRegion id="FE5FA1C57316C6C1F0D5B32388440F72" country="Mexico" name="Queretaro de Arteaga">Queretaro</collectingRegion>
</normalizedToken>
: Municipio 
<normalizedToken id="7ADAFEE49CA7556754F6AAD8CC4B6FC9" originalValue="Colón">Colon</normalizedToken>
</emphasis>
, antena 
<collectorName id="45F7448AF547A0606B8DC85039899E1A">El Zamorano</collectorName>
, [
<geoCoordinate id="15C43158251A21048FB13C4EF740F97D" degrees="20.93305556" direction="north" orientation="latitude" precision="1" value="20.933056">20.93305556°N</geoCoordinate>
, 
<geoCoordinate id="6376CD7A2050C0FC4DE34F044FB5EC2B" degrees="100.1797222" direction="west" orientation="longitude" precision="1" value="-100.179726">100.1797222°W</geoCoordinate>
], 
<quantity id="D69101AF330FD7C886B6C728B7858DB6" metricMagnitude="3" metricUnit="m" metricValue="3.3549999999999995" unit="m" value="3355.0">
<elevation id="6312055747EB5BA239004F60A613A0B9" metricMagnitude="3" metricUnit="m" metricValue="3.3549999999999995" unit="m" value="3355.0">3355 m</elevation>
</quantity>
alt., 
<collectingDate id="E4B7FEB90C2F28ADC84674512DDF3C38" value="1981-11-24">24 Nov 1981</collectingDate>
, 
<collectorName id="0845039CF72EA95381181553D4E03A23">A. Mora</collectorName>
and 
<collectorName id="E389C7BC0B533BF5C492C29B53E66EB2">
J. 
<normalizedToken id="99A1771BB2CA02ED4A852CCBE8A26BF4" originalValue="Ramírez">Ramirez</normalizedToken>
</collectorName>
401-AMB (MEXU [*,**]). 
<emphasis id="2CB8D1C7F814A3BC4ECD5101E09D8760" bold="true" pageId="0" pageNumber="157">Municipio Landa</emphasis>
, Lobo, [
<geoCoordinate id="8DCC69B1B8F741366B6B0285C2F1157D" degrees="21.29275" direction="north" orientation="latitude" precision="1" value="21.29275">21.29275°N</geoCoordinate>
, 
<geoCoordinate id="1B28B56FD961B62C4725209EA8953AF8" degrees="99.11930833" direction="west" orientation="longitude" precision="1" value="-99.11931">99.11930833°W</geoCoordinate>
], 
<quantity id="F031D154E502CD1B2F42A23C81C59D48" metricMagnitude="3" metricUnit="m" metricValue="1.6" unit="m" value="1600.0">
<elevation id="81B39CF8C6FC798F5D944E064E3C4C44" metricMagnitude="3" metricUnit="m" metricValue="1.6" unit="m" value="1600.0">1600 m</elevation>
</quantity>
alt., 
<collectingDate id="4ADB41FAC816696EFA8D8C1D067A8BF5" value="1982-08-24">24 Aug 1982</collectingDate>
, 
<collectorName id="FAB1487373568CC0602E0AAA81A1759C">
R. 
<normalizedToken id="4F2667D9840E0B4A3E9AD7CF46BEECD6" originalValue="Guzmán">Guzman</normalizedToken>
</collectorName>
5959 (MEXU [*, **]). 
<emphasis id="5498145DC19691321BE68B71C82D254B" bold="true" pageId="0" pageNumber="157">
<collectingRegion id="3C67C40EBDE6F372FE4A8B3AEC7263B0" country="Mexico" name="Tlaxcala">Tlaxcala</collectingRegion>
: Municipio Huamantla
</emphasis>
, ladera N del 
<location id="590C8CD1C157183A526CD08D0B6FF524" LSID="urn:lsid:plazi:treatment:1F94286215D453C39B64BCE9C9965372:590C8CD1C157183A526CD08D0B6FF524" country="Mexico" county="Municipio Amecameca" latitude="19.067286" longLatPrecision="1" longitude="-97.19152" municipality="Iztaccihuatl" name="Volcan La Malinche" stateProvince="Queretaro">
<normalizedToken id="E8B41A069145333099B2882EE0CB08B9" originalValue="Volcán">Volcan</normalizedToken>
La Malinche
</location>
, 
<geoCoordinate id="5DE749FA02BBEAC46360D61FBC2C09F9" degrees="19.23486111" direction="north" orientation="latitude" precision="1" value="19.234861">19.23486111°N</geoCoordinate>
, 
<geoCoordinate id="5EA1E5F16418F3F75DE5B9E547653A98" degrees="98.03338889" direction="west" orientation="longitude" precision="1" value="-98.03339">98.03338889°W</geoCoordinate>
, 
<quantity id="69221215E6A05BCE6B3E453C425E2832" metricMagnitude="3" metricUnit="m" metricValue="4.1899999999999995" unit="m" value="4190.0">
<elevation id="4483BC9D00C5F67544C57539CDFF2798" metricMagnitude="3" metricUnit="m" metricValue="4.1899999999999995" unit="m" value="4190.0">4190 m</elevation>
</quantity>
alt., 
<collectingDate id="8C1830339DB931FF11E39862972EF256" value="2013-06-29">29 Jun 2013</collectingDate>
, 
<collectorName id="BF38A77F1271CB58443AD57C840FA9AC">
R. 
<normalizedToken id="1BA2DEF090DAB38A5226C269B42493D7" originalValue="Hernández-Cárdenas">Hernandez-Cardenas</normalizedToken>
</collectorName>
and 
<collectorName id="E5257326114910CD7BB842ADD414E408">L. Arredondo-Amezcua</collectorName>
1123 (IEB), 1929 (IEB). 
<emphasis id="76A57C02A88D8E588AA2F6139D1B5F3C" bold="true" pageId="0" pageNumber="157">
<collectingRegion id="D12E98D8C15A4B43C38AAD0C6C3C1922" country="Mexico" name="Veracruz de Ignacio de la Llave">Veracruz</collectingRegion>
: Municipio Calcahualco
</emphasis>
, 
<collectorName id="55F62936EE1CADDA88BC3BB239B3D778">La Cuchilla</collectorName>
, camino al 
<location id="3C477F47C7F855E60288886DEEAE2487" LSID="urn:lsid:plazi:treatment:1F94286215D453C39B64BCE9C9965372:3C477F47C7F855E60288886DEEAE2487" country="Mexico" county="Municipio Amecameca" latitude="19.067286" longLatPrecision="1" longitude="-97.19152" municipality="Iztaccihuatl" name="Pico de Orizaba" stateProvince="Queretaro">
Pico 
<collectorName id="BD7FA0B79AF46A76BB4A83859EDA6F13">de Orizaba</collectorName>
</location>
, por Coscomatepec, [
<geoCoordinate id="4665B0C040B9CA5B6A549ED088ECBDDF" degrees="19.06728503" direction="north" orientation="latitude" precision="1" value="19.067286">19.06728503°N</geoCoordinate>
, 
<geoCoordinate id="EF0A9AFECA28420CFCA144C4DD2B33DE" degrees="97.19152382" direction="west" orientation="longitude" precision="1" value="-97.19152">97.19152382°W</geoCoordinate>
], 
<quantity id="9912DA713C8837597BB671EA555177B3" metricMagnitude="3" metricUnit="m" metricValue="3.16" unit="m" value="3160.0">
<elevation id="863B4F1AF090AC026831A6D6A08AED8D" metricMagnitude="3" metricUnit="m" metricValue="3.16" unit="m" value="3160.0">3160 m</elevation>
</quantity>
alt., 
<collectingDate id="15565BC44DE6982F3337E47CE9753659" value="1982-07-22">22 Jul 1982</collectingDate>
, 
<collectorName id="D52BB0B716C9526652B46D09CA45A40B">
R. 
<normalizedToken id="3956290856887189246D256A984593E1" originalValue="Guzmán">Guzman</normalizedToken>
</collectorName>
5847 (MEXU). 
<collectorName id="032322ACE1C4BCA97EAC5E01FB4F34A1">See Suppl.</collectorName>
material 2 for the full list of examined specimens
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>