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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1136.72744" ID-Pensoft-Pub="1313-2970-1136-1" ID-Pensoft-UUID="5D0E6613B1D05E4293ABEE4BFD1CA18A" ID-ZooBank="11C6BBFB339A4672AE662CE2B1E6321E" ModsDocID="1313-2970-1136-1" checkinTime="1671287389350" checkinUser="pensoft" docAuthor="Miller, Kelly B." docDate="2022" docId="1F65DD4708BE58E3BBBEDCEFD8F011A8" docLanguage="en" docName="ZooKeys 1136: 1-56" docOrigin="ZooKeys 1136" docPubDate="2022-12-16" docSource="http://dx.doi.org/10.3897/zookeys.1136.72744" docTitle="Desmopachria lata Miller 2022, sp. nov." docType="treatment" docUuid="3D15426D-6C1A-461D-A567-B0A1F59E31D7" docUuidSource="ZooBank" docVersion="2" id="5D0E6613B1D05E4293ABEE4BFD1CA18A" lastPageNumber="1" masterDocId="5D0E6613B1D05E4293ABEE4BFD1CA18A" masterDocTitle="Nineteen new species of Desmopachria Babington, 1841 (Coleoptera, Adephaga, Dytiscidae, Hydroporinae, Hyphydrini) with notes on the taxonomy of the genus" masterLastPageNumber="56" masterPageNumber="1" pageNumber="1" updateTime="1671287691396" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Nineteen new species of Desmopachria Babington, 1841 (Coleoptera, Adephaga, Dytiscidae, Hydroporinae, Hyphydrini) with notes on the taxonomy of the genus</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Miller, Kelly B.</mods:namePart>
<mods:affiliation>Department of Biology and Museum of Southwestern Biology University of New Mexico, Albuquerque, NM 87131 - 0001, USA</mods:affiliation>
<mods:nameIdentifier type="email">kbmiller@unm.edu</mods:nameIdentifier>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-12-16</mods:number>
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<mods:detail type="volume">
<mods:number>1136</mods:number>
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<mods:start>1</mods:start>
<mods:end>56</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1136.72744</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1136.72744</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1136-1</mods:identifier>
<mods:identifier type="ZooBank">11C6BBFB339A4672AE662CE2B1E6321E</mods:identifier>
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<treatment LSID="urn:lsid:zoobank.org:act:3D15426D-6C1A-461D-A567-B0A1F59E31D7" httpUri="http://treatment.plazi.org/id/1F65DD4708BE58E3BBBEDCEFD8F011A8" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="http://zoobank.org/3D15426D-6C1A-461D-A567-B0A1F59E31D7" authority="Miller, 2022" authorityName="Miller" authorityYear="2022" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria lata" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="lata" status="sp. nov.">Desmopachria lata</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">
<figureCitation captionStart="Figures 2845" captionStartId="F3" captionText="Figures 28 - 45. Desmopachria species. (28 - 30) Desmopachria bifurcita 28 habitus 29 male median and lateral lobes, dorsal aspect 30 male lateral lobe, right lateral aspect (31 - 33) Desmopachria lata 31 habitus 32 male genitalia, ventral aspect 33 right lateral lobe, right lateral aspect. (34, 35) Desompachria bifasciata 34 habitus 35 male genitalia, ventral aspect (36 - 38) Desmopachria bolivari 36 male genitalia, dorsal aspect 37 right lateral lobe, right lateral aspect 38 median lobe, right lateral aspect 39 Desmopachria ovalis, male genitalia, dorsal aspect (40 - 42) Desmopachria varians 40 habitus 41 male genitalia, ventral aspect 42 right lateral lobe, right lateral aspect (43 - 45) Desmopachria pseudocavia 43 habitus 44 abdominal sternite VI, ventral aspect 45 male median lobe and right lateral lobe, ventral aspect. Scale bars: 1.0 mm for habitus drawings; 0.25 mm (44)." figureDoi="10.3897/zookeys.1136.72744.figures28-45" httpUri="https://binary.pensoft.net/fig/782455" pageId="0" pageNumber="1">Figures 31-33</figureCitation>
<figureCitation captionStart="Figure 78" captionStartId="F8" captionText="Figure 78. Desmopachria species distributions. Desmopachria bifurcita species group and Desmopachria convexa species group." figureDoi="10.3897/zookeys.1136.72744.figure78" httpUri="https://binary.pensoft.net/fig/782460" pageId="0" pageNumber="1">, 78</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type locality">
<paragraph pageId="0" pageNumber="1">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="1">
Brazil,
<normalizedToken originalValue="Pará">Para</normalizedToken>
State, Cachimbo.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
Males of this species have the median lobe short and simple (Fig.
<figureCitation captionStart="Figures 2845" captionStartId="F3" captionText="Figures 28 - 45. Desmopachria species. (28 - 30) Desmopachria bifurcita 28 habitus 29 male median and lateral lobes, dorsal aspect 30 male lateral lobe, right lateral aspect (31 - 33) Desmopachria lata 31 habitus 32 male genitalia, ventral aspect 33 right lateral lobe, right lateral aspect. (34, 35) Desompachria bifasciata 34 habitus 35 male genitalia, ventral aspect (36 - 38) Desmopachria bolivari 36 male genitalia, dorsal aspect 37 right lateral lobe, right lateral aspect 38 median lobe, right lateral aspect 39 Desmopachria ovalis, male genitalia, dorsal aspect (40 - 42) Desmopachria varians 40 habitus 41 male genitalia, ventral aspect 42 right lateral lobe, right lateral aspect (43 - 45) Desmopachria pseudocavia 43 habitus 44 abdominal sternite VI, ventral aspect 45 male median lobe and right lateral lobe, ventral aspect. Scale bars: 1.0 mm for habitus drawings; 0.25 mm (44)." figureDoi="10.3897/zookeys.1136.72744.figures28-45" httpUri="https://binary.pensoft.net/fig/782455" pageId="0" pageNumber="1">32</figureCitation>
). The lateral lobes are broad, long, and apically broadly expanded and truncate in dorsal aspect (Fig.
<figureCitation captionStart="Figures 2845" captionStartId="F3" captionText="Figures 28 - 45. Desmopachria species. (28 - 30) Desmopachria bifurcita 28 habitus 29 male median and lateral lobes, dorsal aspect 30 male lateral lobe, right lateral aspect (31 - 33) Desmopachria lata 31 habitus 32 male genitalia, ventral aspect 33 right lateral lobe, right lateral aspect. (34, 35) Desompachria bifasciata 34 habitus 35 male genitalia, ventral aspect (36 - 38) Desmopachria bolivari 36 male genitalia, dorsal aspect 37 right lateral lobe, right lateral aspect 38 median lobe, right lateral aspect 39 Desmopachria ovalis, male genitalia, dorsal aspect (40 - 42) Desmopachria varians 40 habitus 41 male genitalia, ventral aspect 42 right lateral lobe, right lateral aspect (43 - 45) Desmopachria pseudocavia 43 habitus 44 abdominal sternite VI, ventral aspect 45 male median lobe and right lateral lobe, ventral aspect. Scale bars: 1.0 mm for habitus drawings; 0.25 mm (44)." figureDoi="10.3897/zookeys.1136.72744.figures28-45" httpUri="https://binary.pensoft.net/fig/782455" pageId="0" pageNumber="1">32</figureCitation>
) and medially abruptly bent and apically sinuate with a narrowly rounded apex (Fig.
<figureCitation captionStart="Figures 2845" captionStartId="F3" captionText="Figures 28 - 45. Desmopachria species. (28 - 30) Desmopachria bifurcita 28 habitus 29 male median and lateral lobes, dorsal aspect 30 male lateral lobe, right lateral aspect (31 - 33) Desmopachria lata 31 habitus 32 male genitalia, ventral aspect 33 right lateral lobe, right lateral aspect. (34, 35) Desompachria bifasciata 34 habitus 35 male genitalia, ventral aspect (36 - 38) Desmopachria bolivari 36 male genitalia, dorsal aspect 37 right lateral lobe, right lateral aspect 38 median lobe, right lateral aspect 39 Desmopachria ovalis, male genitalia, dorsal aspect (40 - 42) Desmopachria varians 40 habitus 41 male genitalia, ventral aspect 42 right lateral lobe, right lateral aspect (43 - 45) Desmopachria pseudocavia 43 habitus 44 abdominal sternite VI, ventral aspect 45 male median lobe and right lateral lobe, ventral aspect. Scale bars: 1.0 mm for habitus drawings; 0.25 mm (44)." figureDoi="10.3897/zookeys.1136.72744.figures28-45" httpUri="https://binary.pensoft.net/fig/782455" pageId="0" pageNumber="1">33</figureCitation>
). The genitalia are similar to
<taxonomicName authorityName="K. B. Miller" authorityYear="1999" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria bolivari" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bolivari">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria bolivari</emphasis>
</taxonomicName>
Miller, but that species has the apices of the lateral lobes broadened and with a distinctive medially directed tooth or hook, among other more subtle genitalic shape differences (
<bibRefCitation DOI="https://doi.org/10.1163/187631200X00165" author="Miller, KB" journalOrPublisher="Entomologica Scandinavica" pageId="0" pageNumber="1" pagination="349 - 359" refId="B10" refString="Miller, KB, 1999. Description of eight new species of Desmopachria Babington (Coleoptera: Dytiscidae) from Bolivia. Entomologica Scandinavica 30: 349 - 359, DOI: https://doi.org/10.1163/187631200X00165" title="Description of eight new species of Desmopachria Babington (Coleoptera: Dytiscidae) from Bolivia." url="https://doi.org/10.1163/187631200X00165" volume="30" year="1999">Miller 1999</bibRefCitation>
: fig. 4).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Measurements.</emphasis>
TL = 2.3 mm, GW = 1.5-1.6 mm, PW = 1.2-1.3 mm, HW = 0.8 mm, EW = 0.4 mm, TL/GW = 1.4-1.5, HW/EW = 2.1-2.2. Body very broad, rounded, laterally broadly curved, lateral margins continuous between pronotum and elytron, body broadest across elytra near midlength of body (Fig.
<figureCitation captionStart="Figures 2845" captionStartId="F3" captionText="Figures 28 - 45. Desmopachria species. (28 - 30) Desmopachria bifurcita 28 habitus 29 male median and lateral lobes, dorsal aspect 30 male lateral lobe, right lateral aspect (31 - 33) Desmopachria lata 31 habitus 32 male genitalia, ventral aspect 33 right lateral lobe, right lateral aspect. (34, 35) Desompachria bifasciata 34 habitus 35 male genitalia, ventral aspect (36 - 38) Desmopachria bolivari 36 male genitalia, dorsal aspect 37 right lateral lobe, right lateral aspect 38 median lobe, right lateral aspect 39 Desmopachria ovalis, male genitalia, dorsal aspect (40 - 42) Desmopachria varians 40 habitus 41 male genitalia, ventral aspect 42 right lateral lobe, right lateral aspect (43 - 45) Desmopachria pseudocavia 43 habitus 44 abdominal sternite VI, ventral aspect 45 male median lobe and right lateral lobe, ventral aspect. Scale bars: 1.0 mm for habitus drawings; 0.25 mm (44)." figureDoi="10.3897/zookeys.1136.72744.figures28-45" httpUri="https://binary.pensoft.net/fig/782455" pageId="0" pageNumber="1">31</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Coloration</emphasis>
(Fig.
<figureCitation captionStart="Figures 2845" captionStartId="F3" captionText="Figures 28 - 45. Desmopachria species. (28 - 30) Desmopachria bifurcita 28 habitus 29 male median and lateral lobes, dorsal aspect 30 male lateral lobe, right lateral aspect (31 - 33) Desmopachria lata 31 habitus 32 male genitalia, ventral aspect 33 right lateral lobe, right lateral aspect. (34, 35) Desompachria bifasciata 34 habitus 35 male genitalia, ventral aspect (36 - 38) Desmopachria bolivari 36 male genitalia, dorsal aspect 37 right lateral lobe, right lateral aspect 38 median lobe, right lateral aspect 39 Desmopachria ovalis, male genitalia, dorsal aspect (40 - 42) Desmopachria varians 40 habitus 41 male genitalia, ventral aspect 42 right lateral lobe, right lateral aspect (43 - 45) Desmopachria pseudocavia 43 habitus 44 abdominal sternite VI, ventral aspect 45 male median lobe and right lateral lobe, ventral aspect. Scale bars: 1.0 mm for habitus drawings; 0.25 mm (44)." figureDoi="10.3897/zookeys.1136.72744.figures28-45" httpUri="https://binary.pensoft.net/fig/782455" pageId="0" pageNumber="1">31</figureCitation>
). Dorsal surface of head, pronotum and elytron dark red-brown, slightly but distinctly paler laterally on pronotum. Head appendages, pro- and mesothoracic legs, and prothoracic ventral surfaces red-orange, other ventral surfaces and metathoracic legs dark red-brown.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Sculpture and structure.</emphasis>
Head broad, anterior margin flattened, margined with distinctive narrow bead; surface of head shiny, finely and sparsely punctate; eyes large (Fig.
<figureCitation captionStart="Figures 2845" captionStartId="F3" captionText="Figures 28 - 45. Desmopachria species. (28 - 30) Desmopachria bifurcita 28 habitus 29 male median and lateral lobes, dorsal aspect 30 male lateral lobe, right lateral aspect (31 - 33) Desmopachria lata 31 habitus 32 male genitalia, ventral aspect 33 right lateral lobe, right lateral aspect. (34, 35) Desompachria bifasciata 34 habitus 35 male genitalia, ventral aspect (36 - 38) Desmopachria bolivari 36 male genitalia, dorsal aspect 37 right lateral lobe, right lateral aspect 38 median lobe, right lateral aspect 39 Desmopachria ovalis, male genitalia, dorsal aspect (40 - 42) Desmopachria varians 40 habitus 41 male genitalia, ventral aspect 42 right lateral lobe, right lateral aspect (43 - 45) Desmopachria pseudocavia 43 habitus 44 abdominal sternite VI, ventral aspect 45 male median lobe and right lateral lobe, ventral aspect. Scale bars: 1.0 mm for habitus drawings; 0.25 mm (44)." figureDoi="10.3897/zookeys.1136.72744.figures28-45" httpUri="https://binary.pensoft.net/fig/782455" pageId="0" pageNumber="1">31</figureCitation>
, HW/EW = 2.1-2.2); antennae short, scape and pedicel relatively large and rounded, flagellomere III long and slender, apically expanded, antennomeres IV-X short and broad, lobed at anterodorsal angle, antennomere XI elongate, apically pointed. Pronotum very short, lateral margins short, slightly curved with continuous narrow bead, slightly wider medially; surface shiny, impunctate medially, finely punctate around margins. Elytron very broad, laterally broadly curved; surface shiny, more coarsely and evenly punctate than pronotum, punctation distinctive and prominent, moderately fine. Prosternum extremely short, longitudinally compressed, medially slightly carinate; prosternal process slender anteriorly, with distinctive, small medial tubercle, apically short and moderately broad, medially evenly convex, apically broadly pointed. Metaventrite broad and evenly smoothly convex medially, surface shiny, impunctate; metaventrite wings extremely slender. Metacoxa with medial portion short, &lt;1/3 length of metaventrite medially, metacoxal lines indistinct, slightly divergent anteriorly; lateral portion of metacoxa extremely large, anteriorly strongly expanded; surface shiny, evenly, finely punctate. Metatrochanter large, subequal to length of ventral margin of metafemur; legs otherwise not noticeably modified. Abdomen with surfaces shiny and smooth, very finely and sparsely punctate.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Male genitalia.</emphasis>
Male median lobe in lateral aspect short, slightly curved, apically simple; in ventral aspect short, slender, parallel-sided to narrowly rounded apex (Fig.
<figureCitation captionStart="Figures 2845" captionStartId="F3" captionText="Figures 28 - 45. Desmopachria species. (28 - 30) Desmopachria bifurcita 28 habitus 29 male median and lateral lobes, dorsal aspect 30 male lateral lobe, right lateral aspect (31 - 33) Desmopachria lata 31 habitus 32 male genitalia, ventral aspect 33 right lateral lobe, right lateral aspect. (34, 35) Desompachria bifasciata 34 habitus 35 male genitalia, ventral aspect (36 - 38) Desmopachria bolivari 36 male genitalia, dorsal aspect 37 right lateral lobe, right lateral aspect 38 median lobe, right lateral aspect 39 Desmopachria ovalis, male genitalia, dorsal aspect (40 - 42) Desmopachria varians 40 habitus 41 male genitalia, ventral aspect 42 right lateral lobe, right lateral aspect (43 - 45) Desmopachria pseudocavia 43 habitus 44 abdominal sternite VI, ventral aspect 45 male median lobe and right lateral lobe, ventral aspect. Scale bars: 1.0 mm for habitus drawings; 0.25 mm (44)." figureDoi="10.3897/zookeys.1136.72744.figures28-45" httpUri="https://binary.pensoft.net/fig/782455" pageId="0" pageNumber="1">32</figureCitation>
). Lateral lobe in ventral aspect evenly broad throughout much of length, apex broadly expanded laterally, apex broad, somewhat obliquely truncate (Fig.
<figureCitation captionStart="Figures 2845" captionStartId="F3" captionText="Figures 28 - 45. Desmopachria species. (28 - 30) Desmopachria bifurcita 28 habitus 29 male median and lateral lobes, dorsal aspect 30 male lateral lobe, right lateral aspect (31 - 33) Desmopachria lata 31 habitus 32 male genitalia, ventral aspect 33 right lateral lobe, right lateral aspect. (34, 35) Desompachria bifasciata 34 habitus 35 male genitalia, ventral aspect (36 - 38) Desmopachria bolivari 36 male genitalia, dorsal aspect 37 right lateral lobe, right lateral aspect 38 median lobe, right lateral aspect 39 Desmopachria ovalis, male genitalia, dorsal aspect (40 - 42) Desmopachria varians 40 habitus 41 male genitalia, ventral aspect 42 right lateral lobe, right lateral aspect (43 - 45) Desmopachria pseudocavia 43 habitus 44 abdominal sternite VI, ventral aspect 45 male median lobe and right lateral lobe, ventral aspect. Scale bars: 1.0 mm for habitus drawings; 0.25 mm (44)." figureDoi="10.3897/zookeys.1136.72744.figures28-45" httpUri="https://binary.pensoft.net/fig/782455" pageId="0" pageNumber="1">32</figureCitation>
); in lateral aspect slender throughout length, abruptly curved medially, apex abruptly slightly expanded and apically narrowly rounded (Fig.
<figureCitation captionStart="Figures 2845" captionStartId="F3" captionText="Figures 28 - 45. Desmopachria species. (28 - 30) Desmopachria bifurcita 28 habitus 29 male median and lateral lobes, dorsal aspect 30 male lateral lobe, right lateral aspect (31 - 33) Desmopachria lata 31 habitus 32 male genitalia, ventral aspect 33 right lateral lobe, right lateral aspect. (34, 35) Desompachria bifasciata 34 habitus 35 male genitalia, ventral aspect (36 - 38) Desmopachria bolivari 36 male genitalia, dorsal aspect 37 right lateral lobe, right lateral aspect 38 median lobe, right lateral aspect 39 Desmopachria ovalis, male genitalia, dorsal aspect (40 - 42) Desmopachria varians 40 habitus 41 male genitalia, ventral aspect 42 right lateral lobe, right lateral aspect (43 - 45) Desmopachria pseudocavia 43 habitus 44 abdominal sternite VI, ventral aspect 45 male median lobe and right lateral lobe, ventral aspect. Scale bars: 1.0 mm for habitus drawings; 0.25 mm (44)." figureDoi="10.3897/zookeys.1136.72744.figures28-45" httpUri="https://binary.pensoft.net/fig/782455" pageId="0" pageNumber="1">33</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Sexual dimorphism.</emphasis>
No obvious sexual dimorphic features were discovered.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Variation.</emphasis>
There is some variation in the dorsal coloration with some specimens paler than others, but otherwise, little variation exists.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">
This species is named
<taxonomicName authorityName="Miller" authorityYear="2022" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria lata" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="lata">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria lata</emphasis>
</taxonomicName>
, Latin for broad, for the broad body in this species.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">
This species is known only from the type locality, Brazil,
<normalizedToken originalValue="Pará">Para</normalizedToken>
State, Cachimbo (Fig.
<figureCitation captionStart="Figure 78" captionStartId="F8" captionText="Figure 78. Desmopachria species distributions. Desmopachria bifurcita species group and Desmopachria convexa species group." figureDoi="10.3897/zookeys.1136.72744.figure78" httpUri="https://binary.pensoft.net/fig/782460" pageId="0" pageNumber="1">78</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type material">
<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph pageId="0" pageNumber="1">
Holotype in USNM, male labeled, &quot;Cachimbo, Para, Brasil, X.1959 M.Alvarenga/ USNMENT01190957 [barcode label]/ HOLOTYPE
<taxonomicName authorityName="Miller" authorityYear="2022" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria lata" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="lata">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria lata</emphasis>
</taxonomicName>
Miller, 2021 [red label with black line border].&quot; Paratypes, 21 in USNM and MSBA labeled same as holotype except with different barcode labels (Table
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Barcode label data for Desmopachria species." httpUri="http://table.plazi.org/id/B5FF9C5283064B11CB6B8D1966FB9A48" pageId="0" pageNumber="1" tableUuid="B5FF9C5283064B11CB6B8D1966FB9A48">1</tableCitation>
) and each with
<normalizedToken originalValue="“/…">&quot;/...</normalizedToken>
PARATYPE
<taxonomicName authorityName="Miller" authorityYear="2022" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria lata" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="lata">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria lata</emphasis>
</taxonomicName>
Miller, 2021 [blue label with black line border.&quot;
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="checklist of desmopachria bifurcita species group">
<paragraph pageId="0" pageNumber="1">
Checklist of
<taxonomicName authorityName="Miller" authorityYear="2022" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria bifurcita" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bifurcita">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria bifurcita</emphasis>
</taxonomicName>
species group
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Zimmermann" authorityYear="1921" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria bifasciata" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bifasciata">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria bifasciata</emphasis>
</taxonomicName>
Zimmermann, 1921 - Brazil (Figs
<figureCitation captionStart="Figures 2845" captionStartId="F3" captionText="Figures 28 - 45. Desmopachria species. (28 - 30) Desmopachria bifurcita 28 habitus 29 male median and lateral lobes, dorsal aspect 30 male lateral lobe, right lateral aspect (31 - 33) Desmopachria lata 31 habitus 32 male genitalia, ventral aspect 33 right lateral lobe, right lateral aspect. (34, 35) Desompachria bifasciata 34 habitus 35 male genitalia, ventral aspect (36 - 38) Desmopachria bolivari 36 male genitalia, dorsal aspect 37 right lateral lobe, right lateral aspect 38 median lobe, right lateral aspect 39 Desmopachria ovalis, male genitalia, dorsal aspect (40 - 42) Desmopachria varians 40 habitus 41 male genitalia, ventral aspect 42 right lateral lobe, right lateral aspect (43 - 45) Desmopachria pseudocavia 43 habitus 44 abdominal sternite VI, ventral aspect 45 male median lobe and right lateral lobe, ventral aspect. Scale bars: 1.0 mm for habitus drawings; 0.25 mm (44)." figureDoi="10.3897/zookeys.1136.72744.figures28-45" httpUri="https://binary.pensoft.net/fig/782455" pageId="0" pageNumber="1">34</figureCitation>
,
<figureCitation captionStart="Figures 2845" captionStartId="F3" captionText="Figures 28 - 45. Desmopachria species. (28 - 30) Desmopachria bifurcita 28 habitus 29 male median and lateral lobes, dorsal aspect 30 male lateral lobe, right lateral aspect (31 - 33) Desmopachria lata 31 habitus 32 male genitalia, ventral aspect 33 right lateral lobe, right lateral aspect. (34, 35) Desompachria bifasciata 34 habitus 35 male genitalia, ventral aspect (36 - 38) Desmopachria bolivari 36 male genitalia, dorsal aspect 37 right lateral lobe, right lateral aspect 38 median lobe, right lateral aspect 39 Desmopachria ovalis, male genitalia, dorsal aspect (40 - 42) Desmopachria varians 40 habitus 41 male genitalia, ventral aspect 42 right lateral lobe, right lateral aspect (43 - 45) Desmopachria pseudocavia 43 habitus 44 abdominal sternite VI, ventral aspect 45 male median lobe and right lateral lobe, ventral aspect. Scale bars: 1.0 mm for habitus drawings; 0.25 mm (44)." figureDoi="10.3897/zookeys.1136.72744.figures28-45" httpUri="https://binary.pensoft.net/fig/782455" pageId="0" pageNumber="1">35</figureCitation>
)
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Miller" authorityYear="2022" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria bifurcita" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bifurcita">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria bifurcita</emphasis>
</taxonomicName>
sp. nov. - Peru
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="K. B. Miller" authorityYear="1999" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria bolivari" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bolivari">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria bolivari</emphasis>
</taxonomicName>
Miller, 1999 - Bolivia (Figs
<figureCitation captionStart="Figures 2845" captionStartId="F3" captionText="Figures 28 - 45. Desmopachria species. (28 - 30) Desmopachria bifurcita 28 habitus 29 male median and lateral lobes, dorsal aspect 30 male lateral lobe, right lateral aspect (31 - 33) Desmopachria lata 31 habitus 32 male genitalia, ventral aspect 33 right lateral lobe, right lateral aspect. (34, 35) Desompachria bifasciata 34 habitus 35 male genitalia, ventral aspect (36 - 38) Desmopachria bolivari 36 male genitalia, dorsal aspect 37 right lateral lobe, right lateral aspect 38 median lobe, right lateral aspect 39 Desmopachria ovalis, male genitalia, dorsal aspect (40 - 42) Desmopachria varians 40 habitus 41 male genitalia, ventral aspect 42 right lateral lobe, right lateral aspect (43 - 45) Desmopachria pseudocavia 43 habitus 44 abdominal sternite VI, ventral aspect 45 male median lobe and right lateral lobe, ventral aspect. Scale bars: 1.0 mm for habitus drawings; 0.25 mm (44)." figureDoi="10.3897/zookeys.1136.72744.figures28-45" httpUri="https://binary.pensoft.net/fig/782455" pageId="0" pageNumber="1">36-38</figureCitation>
)
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Miller" authorityYear="2022" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria lata" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="lata">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria lata</emphasis>
</taxonomicName>
sp. nov. - Brazil
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Sharp" authorityYear="1882" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria ovalis" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="ovalis">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria ovalis</emphasis>
</taxonomicName>
Sharp, 1882 - Brazil (Fig.
<figureCitation captionStart="Figures 2845" captionStartId="F3" captionText="Figures 28 - 45. Desmopachria species. (28 - 30) Desmopachria bifurcita 28 habitus 29 male median and lateral lobes, dorsal aspect 30 male lateral lobe, right lateral aspect (31 - 33) Desmopachria lata 31 habitus 32 male genitalia, ventral aspect 33 right lateral lobe, right lateral aspect. (34, 35) Desompachria bifasciata 34 habitus 35 male genitalia, ventral aspect (36 - 38) Desmopachria bolivari 36 male genitalia, dorsal aspect 37 right lateral lobe, right lateral aspect 38 median lobe, right lateral aspect 39 Desmopachria ovalis, male genitalia, dorsal aspect (40 - 42) Desmopachria varians 40 habitus 41 male genitalia, ventral aspect 42 right lateral lobe, right lateral aspect (43 - 45) Desmopachria pseudocavia 43 habitus 44 abdominal sternite VI, ventral aspect 45 male median lobe and right lateral lobe, ventral aspect. Scale bars: 1.0 mm for habitus drawings; 0.25 mm (44)." figureDoi="10.3897/zookeys.1136.72744.figures28-45" httpUri="https://binary.pensoft.net/fig/782455" pageId="0" pageNumber="1">39</figureCitation>
)
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Wehncke" authorityYear="1877" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria varians" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="varians">
<emphasis italics="true" pageId="0" pageNumber="1">Desmopachria varians</emphasis>
</taxonomicName>
Wehncke, 1877 - Brazil (Figs
<figureCitation captionStart="Figures 2845" captionStartId="F3" captionText="Figures 28 - 45. Desmopachria species. (28 - 30) Desmopachria bifurcita 28 habitus 29 male median and lateral lobes, dorsal aspect 30 male lateral lobe, right lateral aspect (31 - 33) Desmopachria lata 31 habitus 32 male genitalia, ventral aspect 33 right lateral lobe, right lateral aspect. (34, 35) Desompachria bifasciata 34 habitus 35 male genitalia, ventral aspect (36 - 38) Desmopachria bolivari 36 male genitalia, dorsal aspect 37 right lateral lobe, right lateral aspect 38 median lobe, right lateral aspect 39 Desmopachria ovalis, male genitalia, dorsal aspect (40 - 42) Desmopachria varians 40 habitus 41 male genitalia, ventral aspect 42 right lateral lobe, right lateral aspect (43 - 45) Desmopachria pseudocavia 43 habitus 44 abdominal sternite VI, ventral aspect 45 male median lobe and right lateral lobe, ventral aspect. Scale bars: 1.0 mm for habitus drawings; 0.25 mm (44)." figureDoi="10.3897/zookeys.1136.72744.figures28-45" httpUri="https://binary.pensoft.net/fig/782455" pageId="0" pageNumber="1">40-42</figureCitation>
) (Braga and Ferreira-Jr. 2018)
</paragraph>
</subSubSection>
</treatment>
</document>