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<document ID-DOI="10.11646/palaeoentomology.5.5.8" ID-ISSN="2624-2834" ID-Zenodo-Dep="7333751" ID-ZooBank="B3D18635-38AD-496E-B681-E4CCC5CEE97B" approvalRequired="3" approvalRequired_for_taxonomicNames="2" approvalRequired_for_textStreams="1" checkinTime="1668767213831" checkinUser="plazi" docAuthor="WICHARD, WILFRIED &amp; MÜLLER, PATRICK" docDate="2022" docId="1F656719FF80FFC9FCDC2B03051A584A" docLanguage="en" docName="palaeoentomology.5.5.8.pdf" docOrigin="Palaeoentomology 5 (5)" docSource="http://dx.doi.org/10.11646/palaeoentomology.5.5.8" docStyle="DocumentStyle:4EE6FD780FF4521D541D0D0600705D0E.2:Palaeontomology.2018-.journal_article" docStyleId="4EE6FD780FF4521D541D0D0600705D0E" docStyleName="Palaeontomology.2018-.journal_article" docStyleVersion="2" docTitle="Palleptocerus grimaldii WICHARD &amp; MÜLLER 2022, sp. nov." docType="treatment" docVersion="2" lastPageNumber="470" masterDocId="E35C1F61FF81FFCBFFF62A71015A5F1F" masterDocTitle="Palleptoceridae fam. nov., an extinct leptoceroid family in mid-Cretaceous Burmese amber (Insecta, Trichoptera)" masterLastPageNumber="474" masterPageNumber="468" pageNumber="469" updateTime="1668767636559" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Palleptoceridae fam. nov., an extinct leptoceroid family in mid-Cretaceous Burmese amber (Insecta, Trichoptera)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>WICHARD, WILFRIED</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>MÜLLER, PATRICK</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Palaeoentomology</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-09-22</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>5</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>5</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>468</mods:start>
<mods:end>474</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.11646/palaeoentomology.5.5.8</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/palaeoentomology.5.5.8</mods:identifier>
<mods:identifier type="ISSN">2624-2834</mods:identifier>
<mods:identifier type="Zenodo-Dep">7333751</mods:identifier>
<mods:identifier type="ZooBank">B3D18635-38AD-496E-B681-E4CCC5CEE97B</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7333749" ID-Zenodo-Dep="7333749" LSID="urn:lsid:plazi:treatment:1F656719FF80FFC9FCDC2B03051A584A" httpUri="http://treatment.plazi.org/id/1F656719FF80FFC9FCDC2B03051A584A" lastPageId="2" lastPageNumber="470" pageId="1" pageNumber="469">
<subSubSection box="[810,1176,370,397]" pageId="1" pageNumber="469" type="nomenclature">
<paragraph blockId="1.[810,1176,370,432]" box="[810,1176,370,397]" pageId="1" pageNumber="469">
<heading bold="true" box="[810,1176,370,397]" fontSize="10" level="2" pageId="1" pageNumber="469" reason="3">
<emphasis bold="true" box="[810,1176,370,397]" pageId="1" pageNumber="469">
<taxonomicName authority="WICHARD &amp; MÜLLER, 2022" authorityName="WICHARD &amp; MÜLLER" authorityYear="2022" box="[810,1080,370,397]" class="Insecta" family="Leptoceridae" genus="Palleptocerus" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="469" phylum="Arthropoda" rank="species" species="grimaldii" status="sp. nov.">
<emphasis bold="true" box="[810,1080,370,397]" italics="true" pageId="1" pageNumber="469">Palleptocerus grimaldii</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1087,1176,370,396]" pageId="1" pageNumber="469" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[810,926,405,432]" pageId="1" pageNumber="469" type="description">
<paragraph blockId="1.[810,1176,370,432]" box="[810,926,405,432]" pageId="1" pageNumber="469">
(
<figureCitation box="[818,917,405,432]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="2.[136,235,1350,1374]" captionStartId-1="3.[136,235,1716,1740]" captionStartId-2="4.[136,235,1690,1714]" captionStartId-3="5.[136,235,1645,1669]" captionTargetBox-0="[248,1340,189,1326]" captionTargetBox-1="[248,1340,196,1682]" captionTargetBox-2="[271,1316,189,1667]" captionTargetBox-3="[282,1306,174,1622]" captionTargetId-0="figure-273@2.[248,1340,189,1326]" captionTargetId-1="figure-129@3.[248,1340,194,1692]" captionTargetId-2="figure-78@4.[271,1316,189,1667]" captionTargetId-3="figure-87@5.[282,1306,174,1622]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionText-0="FIGURE 1. Palleptocerus grimaldii sp. nov., mating. A, A mating pair, male holotype (right), female paratype 1 (left), ZFMK- TRI000831, in ventral view; scale bar = 2 mm. B, Male paratype, ZFMK-TRI000833, metatibia, each with a pair of apical and preapical spurs (arrows), circle = branching spurs; scale bar = 0.5 mm." captionText-1="FIGURE 2. Palleptocerus grimaldii sp. nov., characters. A, Maxillary palp of female paratype 1, ZFMK-TRI000831; scale bar = 0.1 mm. B, Wing venations, venations of male forewing, female forewing and hind wing; scale bar = 1 mm." captionText-2="FIGURE 3. Palleptocerus grimaldii sp. nov., genitalia. A, Male holotype, ZFMK-TRI000831, adult in dorsal view; scale bar = 1 mm. A Inset, Male genitalia with a pair of inferior appendages (inf ap); scale bar = 0.3 mm. B, Female paratype 1, ZFMK- TRI000831, adult in dorsal view; scale bar = 1 mm. B Inset, Female genitalia with a pair of lobes (dolo); scale bar = 0.2 mm. C, Mesoscutum sclerotized, running longitudinally, with dark marginal ridges on both sides; scale bar = 0.1 mm. C Inset, a row of setae, inside along of the marginal ridge of mesoscutum, scale bar = 0.05 mm" captionText-3="FIGURE 4. Palleptocerus grimaldii sp. nov., paratypes. A, Female paratype 2, ZFMK-TRI000832, adult in ventral view; scale bar = 1 mm. A Inset, Female genitalia with a pair of dorsal setose lobes (dolo); scale bar = 0.3 mm. B, Male paratype, ZFMK- TRI000833, adult in ventral view; scale bar = 1 mm. B Inset, Male genitalia with five-segmented maxillary palps (mx); scale bar = 0.3 mm." figureDoi-0="http://doi.org/10.5281/zenodo.7333755" figureDoi-1="http://doi.org/10.5281/zenodo.7333757" figureDoi-2="http://doi.org/10.5281/zenodo.7333759" figureDoi-3="http://doi.org/10.5281/zenodo.7333763" httpUri-0="https://zenodo.org/record/7333755/files/figure.png" httpUri-1="https://zenodo.org/record/7333757/files/figure.png" httpUri-2="https://zenodo.org/record/7333759/files/figure.png" httpUri-3="https://zenodo.org/record/7333763/files/figure.png" pageId="1" pageNumber="469">Figs 1–4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="469" type="materials_examined">
<paragraph blockId="1.[810,1452,476,2093]" pageId="1" pageNumber="469">
<emphasis bold="true" box="[810,994,476,502]" pageId="1" pageNumber="469">Type material.</emphasis>
Male
<typeStatus box="[1087,1185,476,502]" pageId="1" pageNumber="469">holotype</typeStatus>
and female
<typeStatus box="[1355,1451,477,503]" pageId="1" pageNumber="469">paratype</typeStatus>
1 are next to each other in small amber (
<figureCitation box="[1297,1394,511,538]" captionStart="FIGURE 1" captionStartId="2.[136,235,1350,1374]" captionTargetBox="[248,1340,189,1326]" captionTargetId="figure-273@2.[248,1340,189,1326]" captionTargetPageId="2" captionText="FIGURE 1. Palleptocerus grimaldii sp. nov., mating. A, A mating pair, male holotype (right), female paratype 1 (left), ZFMK- TRI000831, in ventral view; scale bar = 2 mm. B, Male paratype, ZFMK-TRI000833, metatibia, each with a pair of apical and preapical spurs (arrows), circle = branching spurs; scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7333755" httpUri="https://zenodo.org/record/7333755/files/figure.png" pageId="1" pageNumber="469">Figs 1A</figureCitation>
,
<figureCitation captionStart="FIGURE 3" captionStartId="4.[136,235,1690,1714]" captionTargetBox="[271,1316,189,1667]" captionTargetId="figure-78@4.[271,1316,189,1667]" captionTargetPageId="4" captionText="FIGURE 3. Palleptocerus grimaldii sp. nov., genitalia. A, Male holotype, ZFMK-TRI000831, adult in dorsal view; scale bar = 1 mm. A Inset, Male genitalia with a pair of inferior appendages (inf ap); scale bar = 0.3 mm. B, Female paratype 1, ZFMK- TRI000831, adult in dorsal view; scale bar = 1 mm. B Inset, Female genitalia with a pair of lobes (dolo); scale bar = 0.2 mm. C, Mesoscutum sclerotized, running longitudinally, with dark marginal ridges on both sides; scale bar = 0.1 mm. C Inset, a row of setae, inside along of the marginal ridge of mesoscutum, scale bar = 0.05 mm" figureDoi="http://doi.org/10.5281/zenodo.7333759" httpUri="https://zenodo.org/record/7333759/files/figure.png" pageId="1" pageNumber="469">3A, 3B</figureCitation>
): ZFMK-TRI000831. Female
<typeStatus box="[1214,1310,547,573]" pageId="1" pageNumber="469">paratype</typeStatus>
2: ZFMKTRI000832 (
<figureCitation box="[953,1043,582,608]" captionStart="FIGURE 4" captionStartId="5.[136,235,1645,1669]" captionTargetBox="[282,1306,174,1622]" captionTargetId="figure-87@5.[282,1306,174,1622]" captionTargetPageId="5" captionText="FIGURE 4. Palleptocerus grimaldii sp. nov., paratypes. A, Female paratype 2, ZFMK-TRI000832, adult in ventral view; scale bar = 1 mm. A Inset, Female genitalia with a pair of dorsal setose lobes (dolo); scale bar = 0.3 mm. B, Male paratype, ZFMK- TRI000833, adult in ventral view; scale bar = 1 mm. B Inset, Male genitalia with five-segmented maxillary palps (mx); scale bar = 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.7333763" httpUri="https://zenodo.org/record/7333763/files/figure.png" pageId="1" pageNumber="469">Fig. 4A</figureCitation>
). Male
<typeStatus box="[1125,1226,583,609]" pageId="1" pageNumber="469">paratype</typeStatus>
: ZFMK-TRI000833 (
<figureCitation box="[818,906,617,644]" captionStart="FIGURE 4" captionStartId="5.[136,235,1645,1669]" captionTargetBox="[282,1306,174,1622]" captionTargetId="figure-87@5.[282,1306,174,1622]" captionTargetPageId="5" captionText="FIGURE 4. Palleptocerus grimaldii sp. nov., paratypes. A, Female paratype 2, ZFMK-TRI000832, adult in ventral view; scale bar = 1 mm. A Inset, Female genitalia with a pair of dorsal setose lobes (dolo); scale bar = 0.3 mm. B, Male paratype, ZFMK- TRI000833, adult in ventral view; scale bar = 1 mm. B Inset, Male genitalia with five-segmented maxillary palps (mx); scale bar = 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.7333763" httpUri="https://zenodo.org/record/7333763/files/figure.png" pageId="1" pageNumber="469">Fig. 4B</figureCitation>
).
</paragraph>
<paragraph blockId="1.[810,1452,476,2093]" pageId="1" pageNumber="469">Bodies partially well preserved; forewings in dorsal view visible, hind wings partially covered by forewings. Long antennae incomplete in length. Head, thorax and abdomen can partially show signs of decomposition. Fore and hind wing venations largely visible, but crossveins are not or often barely visible, which is probably due to the preservation in amber.</paragraph>
<paragraph blockId="1.[810,1452,476,2093]" pageId="1" pageNumber="469">In dorsal and lateral views, the genitalia of males and females are usually obscured by the saddle-roofed and moderately transparent wings. In ventral view, the genitalia are more visible when not clouded by unfavourable preservation in amber. In male genitalia, only the anterior paired inferior appendages are often clearly visible ventrally for this reason.</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="469" type="etymology">
<paragraph blockId="1.[810,1452,476,2093]" pageId="1" pageNumber="469">
<emphasis bold="true" box="[863,998,1148,1174]" pageId="1" pageNumber="469">Etymology.</emphasis>
The new species is dedicated in honor of David Grimaldi on his 65
<superScript attach="left" box="[1096,1109,1183,1197]" fontSize="6" pageId="1" pageNumber="469">th</superScript>
birthday.
</paragraph>
</subSubSection>
<subSubSection box="[863,1182,1218,1245]" pageId="1" pageNumber="469" type="diagnosis">
<paragraph blockId="1.[810,1452,476,2093]" box="[863,1182,1218,1245]" pageId="1" pageNumber="469">
<emphasis bold="true" box="[863,985,1218,1244]" pageId="1" pageNumber="469">Diagnosis.</emphasis>
As for the genus.
</paragraph>
</subSubSection>
<subSubSection lastPageId="2" lastPageNumber="470" pageId="1" pageNumber="469" type="description">
<paragraph blockId="1.[810,1452,476,2093]" pageId="1" pageNumber="469">
<emphasis bold="true" box="[863,1009,1253,1279]" pageId="1" pageNumber="469">Description.</emphasis>
Head: Laterally protruded compound eyes; the outer eye distance is
<quantity box="[1148,1236,1289,1315]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="469" unit="mm" value="0.7">0.7 mm</quantity>
. Ocelli not present. Filiform antennae longer than forewings - embedded in amber and truncate, therefore incomplete in length; in
<typeStatus box="[844,940,1395,1421]" pageId="1" pageNumber="469">paratype</typeStatus>
male (
<figureCitation box="[1030,1122,1395,1421]" captionStart="FIGURE 4" captionStartId="5.[136,235,1645,1669]" captionTargetBox="[282,1306,174,1622]" captionTargetId="figure-87@5.[282,1306,174,1622]" captionTargetPageId="5" captionText="FIGURE 4. Palleptocerus grimaldii sp. nov., paratypes. A, Female paratype 2, ZFMK-TRI000832, adult in ventral view; scale bar = 1 mm. A Inset, Female genitalia with a pair of dorsal setose lobes (dolo); scale bar = 0.3 mm. B, Male paratype, ZFMK- TRI000833, adult in ventral view; scale bar = 1 mm. B Inset, Male genitalia with five-segmented maxillary palps (mx); scale bar = 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.7333763" httpUri="https://zenodo.org/record/7333763/files/figure.png" pageId="1" pageNumber="469">Fig. 4B</figureCitation>
) antennae twice as long as forewings, subsequently truncated. Scapus conically thickened; flagellomeres cylindrical, elongate. In both sexes, five-segmented maxillary palps present, palp segment lengths in a row: about 0.1, 0.2, 0.2, 0.12,
<quantity metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="469" unit="mm" value="0.3">0.3 mm</quantity>
(female:
<figureCitation box="[966,1058,1572,1598]" captionStart="FIGURE 2" captionStartId="3.[136,235,1716,1740]" captionTargetBox="[248,1340,196,1682]" captionTargetId="figure-129@3.[248,1340,194,1692]" captionTargetPageId="3" captionText="FIGURE 2. Palleptocerus grimaldii sp. nov., characters. A, Maxillary palp of female paratype 1, ZFMK-TRI000831; scale bar = 0.1 mm. B, Wing venations, venations of male forewing, female forewing and hind wing; scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7333757" httpUri="https://zenodo.org/record/7333757/files/figure.png" pageId="1" pageNumber="469">Fig. 2A</figureCitation>
; male:
<figureCitation box="[1145,1231,1572,1598]" captionStart="FIGURE 4" captionStartId="5.[136,235,1645,1669]" captionTargetBox="[282,1306,174,1622]" captionTargetId="figure-87@5.[282,1306,174,1622]" captionTargetPageId="5" captionText="FIGURE 4. Palleptocerus grimaldii sp. nov., paratypes. A, Female paratype 2, ZFMK-TRI000832, adult in ventral view; scale bar = 1 mm. A Inset, Female genitalia with a pair of dorsal setose lobes (dolo); scale bar = 0.3 mm. B, Male paratype, ZFMK- TRI000833, adult in ventral view; scale bar = 1 mm. B Inset, Male genitalia with five-segmented maxillary palps (mx); scale bar = 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.7333763" httpUri="https://zenodo.org/record/7333763/files/figure.png" pageId="1" pageNumber="469">Fig. 4B</figureCitation>
inset); 5
<superScript attach="left" box="[1335,1348,1571,1585]" fontSize="6" pageId="1" pageNumber="469">th</superScript>
segment flexible, not annulated. Labial palps three-segmented, terminal segment longest.
</paragraph>
<paragraph blockId="1.[810,1452,476,2093]" pageId="1" pageNumber="469">
Thorax: Mesoscutum (
<figureCitation box="[1120,1207,1678,1704]" captionStart="FIGURE 3" captionStartId="4.[136,235,1690,1714]" captionTargetBox="[271,1316,189,1667]" captionTargetId="figure-78@4.[271,1316,189,1667]" captionTargetPageId="4" captionText="FIGURE 3. Palleptocerus grimaldii sp. nov., genitalia. A, Male holotype, ZFMK-TRI000831, adult in dorsal view; scale bar = 1 mm. A Inset, Male genitalia with a pair of inferior appendages (inf ap); scale bar = 0.3 mm. B, Female paratype 1, ZFMK- TRI000831, adult in dorsal view; scale bar = 1 mm. B Inset, Female genitalia with a pair of lobes (dolo); scale bar = 0.2 mm. C, Mesoscutum sclerotized, running longitudinally, with dark marginal ridges on both sides; scale bar = 0.1 mm. C Inset, a row of setae, inside along of the marginal ridge of mesoscutum, scale bar = 0.05 mm" figureDoi="http://doi.org/10.5281/zenodo.7333759" httpUri="https://zenodo.org/record/7333759/files/figure.png" pageId="1" pageNumber="469">Fig. 3C</figureCitation>
) distinct, sclerotized, running longitudinally, with dark marginal ridges on both sides. Two bands with setiferous punctures along dorsal side of mesoscutum not completely visible, but as far as visible, some setae appear in row, visible along right band (
<figureCitation box="[818,903,1854,1880]" captionStart="FIGURE 3" captionStartId="4.[136,235,1690,1714]" captionTargetBox="[271,1316,189,1667]" captionTargetId="figure-78@4.[271,1316,189,1667]" captionTargetPageId="4" captionText="FIGURE 3. Palleptocerus grimaldii sp. nov., genitalia. A, Male holotype, ZFMK-TRI000831, adult in dorsal view; scale bar = 1 mm. A Inset, Male genitalia with a pair of inferior appendages (inf ap); scale bar = 0.3 mm. B, Female paratype 1, ZFMK- TRI000831, adult in dorsal view; scale bar = 1 mm. B Inset, Female genitalia with a pair of lobes (dolo); scale bar = 0.2 mm. C, Mesoscutum sclerotized, running longitudinally, with dark marginal ridges on both sides; scale bar = 0.1 mm. C Inset, a row of setae, inside along of the marginal ridge of mesoscutum, scale bar = 0.05 mm" figureDoi="http://doi.org/10.5281/zenodo.7333759" httpUri="https://zenodo.org/record/7333759/files/figure.png" pageId="1" pageNumber="469">Fig. 3C</figureCitation>
Inset).
</paragraph>
<paragraph blockId="1.[810,1452,476,2093]" pageId="1" pageNumber="469">
Tibial spurs: 2/4/4, metatibia, each with a pair of apical and preapical spurs (
<figureCitation box="[1115,1200,1925,1951]" captionStart="FIGURE 1" captionStartId="2.[136,235,1350,1374]" captionTargetBox="[248,1340,189,1326]" captionTargetId="figure-273@2.[248,1340,189,1326]" captionTargetPageId="2" captionText="FIGURE 1. Palleptocerus grimaldii sp. nov., mating. A, A mating pair, male holotype (right), female paratype 1 (left), ZFMK- TRI000831, in ventral view; scale bar = 2 mm. B, Male paratype, ZFMK-TRI000833, metatibia, each with a pair of apical and preapical spurs (arrows), circle = branching spurs; scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7333755" httpUri="https://zenodo.org/record/7333755/files/figure.png" pageId="1" pageNumber="469">Fig. 1B</figureCitation>
)
</paragraph>
<paragraph blockId="1.[810,1452,476,2093]" lastBlockId="2.[136,778,1515,2093]" lastPageId="2" lastPageNumber="470" pageId="1" pageNumber="469">
Wings: The forewings about
<quantity box="[1198,1286,1960,1987]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="1" pageNumber="469" unit="mm" value="4.5">4.5 mm</quantity>
long in males and 5.0 mm long in females. Sexual dimorphism is more pronounced in the venation (
<figureCitation box="[1200,1337,2031,2057]" captionStart="FIGURE 2" captionStartId="3.[136,235,1716,1740]" captionTargetBox="[248,1340,196,1682]" captionTargetId="figure-129@3.[248,1340,194,1692]" captionTargetPageId="3" captionText="FIGURE 2. Palleptocerus grimaldii sp. nov., characters. A, Maxillary palp of female paratype 1, ZFMK-TRI000831; scale bar = 0.1 mm. B, Wing venations, venations of male forewing, female forewing and hind wing; scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7333757" httpUri="https://zenodo.org/record/7333757/files/figure.png" pageId="1" pageNumber="469">Fig. 2B, 2C</figureCitation>
): forks III present in females, males without them. In both sexes Sc and R
<subScript attach="left" box="[203,211,1533,1547]" fontSize="6" pageId="2" pageNumber="470">1</subScript>
straight running parallel to the wing margin. R
<subScript attach="left" box="[746,754,1533,1547]" fontSize="6" pageId="2" pageNumber="470">2</subScript>
+ R
<subScript attach="left" box="[155,163,1570,1584]" fontSize="6" pageId="2" pageNumber="470">3</subScript>
forming fork I present and R
<subScript attach="left" box="[497,523,1570,1584]" fontSize="6" pageId="2" pageNumber="470">4+5</subScript>
simple, discoidal cells absent. Crossvein r-m incompletely indicated. In male M two-branched in M
<subScript attach="left" box="[396,421,1644,1658]" fontSize="6" pageId="2" pageNumber="470">1+2</subScript>
and M
<subScript attach="left" box="[510,536,1644,1658]" fontSize="6" pageId="2" pageNumber="470">3+4</subScript>
; in female M threebranched, M
<subScript attach="left" box="[278,286,1681,1695]" fontSize="6" pageId="2" pageNumber="470">1</subScript>
+ M
<subScript attach="left" box="[342,350,1681,1695]" fontSize="6" pageId="2" pageNumber="470">2</subScript>
forming fork III and M
<subScript attach="left" box="[618,644,1681,1695]" fontSize="6" pageId="2" pageNumber="470">3+4</subScript>
simple. Cu
<subScript attach="left" box="[770,778,1681,1695]" fontSize="6" pageId="2" pageNumber="470">1</subScript>
two-branched in Cu
<subScript attach="left" box="[357,372,1717,1731]" fontSize="6" pageId="2" pageNumber="470">1a</subScript>
and Cu
<subScript attach="left" box="[458,474,1717,1731]" fontSize="6" pageId="2" pageNumber="470">1b</subScript>
forming fork V. Crossvein m-cu closing the long thyridial cell. In male venation Cu
<subScript attach="left" box="[770,778,1754,1768]" fontSize="6" pageId="2" pageNumber="470">2</subScript>
and A
<subScript attach="left" box="[203,246,1791,1805]" fontSize="6" pageId="2" pageNumber="470">1+2+3</subScript>
running parallel and reaching the wing margin together at arculus. Crossvein between Cu
<subScript attach="left" box="[650,666,1827,1841]" fontSize="6" pageId="2" pageNumber="470">1b</subScript>
and Cu
<subScript attach="left" box="[770,778,1828,1842]" fontSize="6" pageId="2" pageNumber="470">2</subScript>
reaching arculus. In female Cu
<subScript attach="left" box="[476,484,1864,1878]" fontSize="6" pageId="2" pageNumber="470">2</subScript>
and A
<subScript attach="none" box="[555,598,1864,1878]" fontSize="6" pageId="2" pageNumber="470">1+2+3</subScript>
running parallel but reaching the wing margin at different distance. In both sexes simplified hind wing venation, forks I, II, III, IV absent, exclusively fork V present (
<figureCitation box="[546,638,1956,1982]" captionStart="FIGURE 2" captionStartId="3.[136,235,1716,1740]" captionTargetBox="[248,1340,196,1682]" captionTargetId="figure-129@3.[248,1340,194,1692]" captionTargetPageId="3" captionText="FIGURE 2. Palleptocerus grimaldii sp. nov., characters. A, Maxillary palp of female paratype 1, ZFMK-TRI000831; scale bar = 0.1 mm. B, Wing venations, venations of male forewing, female forewing and hind wing; scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7333757" httpUri="https://zenodo.org/record/7333757/files/figure.png" pageId="2" pageNumber="470">Fig. 2D</figureCitation>
), crossveins absent.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7333755" ID-Zenodo-Dep="7333755" httpUri="https://zenodo.org/record/7333755/files/figure.png" pageId="2" pageNumber="470" startId="2.[136,235,1350,1374]" targetBox="[248,1340,189,1326]" targetPageId="2">
<paragraph blockId="2.[136,1452,1350,1446]" pageId="2" pageNumber="470">
<emphasis bold="true" box="[136,261,1350,1374]" pageId="2" pageNumber="470">FIGURE 1.</emphasis>
<taxonomicName authority="WICHARD &amp; MÜLLER, 2022" authorityName="WICHARD &amp; MÜLLER" authorityYear="2022" box="[269,511,1350,1374]" class="Insecta" family="Leptoceridae" genus="Palleptocerus" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="470" phylum="Arthropoda" rank="species" species="grimaldii" status="sp. nov.">
<emphasis box="[269,511,1350,1374]" italics="true" pageId="2" pageNumber="470">Palleptocerus grimaldii</emphasis>
</taxonomicName>
<emphasis bold="true" box="[519,600,1350,1374]" pageId="2" pageNumber="470">
<taxonomicNameLabel box="[519,600,1350,1374]" pageId="2" pageNumber="470" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, mating.
<emphasis bold="true" box="[699,717,1350,1374]" pageId="2" pageNumber="470">A</emphasis>
, A mating pair, male holotype (right), female paratype 1 (left), ZFMKTRI000831, in ventral view; scale bar = 2 mm.
<emphasis bold="true" box="[633,650,1386,1410]" pageId="2" pageNumber="470">B</emphasis>
, Male paratype, ZFMK-TRI000833, metatibia, each with a pair of apical and preapical spurs (arrows), circle = branching spurs; scale bar = 0.5 mm.
</paragraph>
</caption>
<paragraph blockId="2.[136,778,1515,2093]" lastBlockId="2.[810,1451,1514,1878]" pageId="2" pageNumber="470">
Genitalia: Access and visibility of the male and female genital structures are difficult in amber. In the female, a pair of dorsal setose lobes visible (
<figureCitation box="[1301,1385,1514,1540]" captionStart="FIGURE 3" captionStartId="4.[136,235,1690,1714]" captionTargetBox="[271,1316,189,1667]" captionTargetId="figure-78@4.[271,1316,189,1667]" captionTargetPageId="4" captionText="FIGURE 3. Palleptocerus grimaldii sp. nov., genitalia. A, Male holotype, ZFMK-TRI000831, adult in dorsal view; scale bar = 1 mm. A Inset, Male genitalia with a pair of inferior appendages (inf ap); scale bar = 0.3 mm. B, Female paratype 1, ZFMK- TRI000831, adult in dorsal view; scale bar = 1 mm. B Inset, Female genitalia with a pair of lobes (dolo); scale bar = 0.2 mm. C, Mesoscutum sclerotized, running longitudinally, with dark marginal ridges on both sides; scale bar = 0.1 mm. C Inset, a row of setae, inside along of the marginal ridge of mesoscutum, scale bar = 0.05 mm" figureDoi="http://doi.org/10.5281/zenodo.7333759" httpUri="https://zenodo.org/record/7333759/files/figure.png" pageId="2" pageNumber="470">Fig. 3B</figureCitation>
inset, 4A inset); details of the gonopod plate not visible. In ventral view, the male genitalia equipped with a pair of inferior appendages (
<figureCitation box="[1049,1140,1626,1653]" captionStart="FIGURE 3" captionStartId="4.[136,235,1690,1714]" captionTargetBox="[271,1316,189,1667]" captionTargetId="figure-78@4.[271,1316,189,1667]" captionTargetPageId="4" captionText="FIGURE 3. Palleptocerus grimaldii sp. nov., genitalia. A, Male holotype, ZFMK-TRI000831, adult in dorsal view; scale bar = 1 mm. A Inset, Male genitalia with a pair of inferior appendages (inf ap); scale bar = 0.3 mm. B, Female paratype 1, ZFMK- TRI000831, adult in dorsal view; scale bar = 1 mm. B Inset, Female genitalia with a pair of lobes (dolo); scale bar = 0.2 mm. C, Mesoscutum sclerotized, running longitudinally, with dark marginal ridges on both sides; scale bar = 0.1 mm. C Inset, a row of setae, inside along of the marginal ridge of mesoscutum, scale bar = 0.05 mm" figureDoi="http://doi.org/10.5281/zenodo.7333759" httpUri="https://zenodo.org/record/7333759/files/figure.png" pageId="2" pageNumber="470">Fig. 3A</figureCitation>
), each consisting of a dark, rod-shaped coxopodite and an apically attached harpago that is shorter and lighter in color than the coxopodite and tapers and curves slightly toward the genital midline. As far as visible, a rod-shaped appendage nearly as long as the coxopodite tentatively interpreted as a meso-dorsal appendage of tergum 10.
</paragraph>
</subSubSection>
</treatment>
</document>
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