treatments-xml/data/1F/20/F5/1F20F502FFB1FF94FF22FB248DA1FD64.xml

<document id="F08EEFC5EEFE8B1DFA35C4935D45C5EF" ID-CLB-Dataset="54879" ID-DOI="10.11646/zootaxa.5330.4.4" ID-GBIF-Dataset="c089657b-b4cb-469a-9e1f-94a669a51fb1" ID-ISSN="1175-5326" ID-Zenodo-Dep="8255293" ID-ZooBank="8EC3E31F-ED7B-48BD-901B-1EE8AE0BAF02" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="valdenar" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1692261687016" checkinUser="plazi" docAuthor="Puissant, Stephane &amp; Gurcel, Kevin" docDate="2023" docId="1F20F502FFB1FF94FF22FB248DA1FD64" docLanguage="en" docName="zootaxa.5330.4.4.pdf" docOrigin="Zootaxa 5330 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5330.4.4" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tibicina quadrisignata subsp. pilleti Puissant &amp; Gurcel 2023, ssp. n." docType="treatment" docVersion="4" lastPageNumber="550" masterDocId="E3198D7AFFB9FF9BFFB5FFA28F6AFFA9" masterDocTitle="The genus Tibicina Kolenati, 1857 in Morocco (Hemiptera: Cicadidae: Tibicininae): taxonomic assessment from integrative research" masterLastPageNumber="560" masterPageNumber="535" pageNumber="543" updateTime="1692299511565" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="B8D025EA04302A180614DE1EAFE80E51" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="4B92A8E3F983BE60400404E1C0822A09">
<mods:title id="E26DA0C54D10D298771D8A55E367F16E">The genus Tibicina Kolenati, 1857 in Morocco (Hemiptera: Cicadidae: Tibicininae): taxonomic assessment from integrative research</mods:title>
</mods:titleInfo>
<mods:name id="A197A3C66804E32DC8C549A1F5A64D92" type="personal">
<mods:role id="83E2242E87A3458B959860FBC58B805A">
<mods:roleTerm id="E30DADC48DD4A9593D425430A5B3A6F0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CF82778A598988759D1B49475478245B">Puissant, Stephane</mods:namePart>
<mods:nameIdentifier id="E5ECC8F95466FE913C8C6E8002E6CA56" type="ORCID">0000-0003-4179-3636</mods:nameIdentifier>
<mods:affiliation id="F459B40A586EEDAFA9807F40CF7F86AC">Muséum national d’Histoire naturelle, Département Systématique et Evolution, UMR CNRS 7205 OSEB, CP 50, 45 rue Buffon, F - 75005 Paris and Muséum d’Histoire naturelle de Dijon - Jardin de l’Arquebuse, 14 rue Jehan de Marville, F - 21000 Dijon, France. 10 chemin des Châtaigniers, F - 74150 Rumilly. kevin. gurcel @ orange. fr; https: // orcid. org / 0009 - 0006 - 7380 - 9072 Corresponding author. E-mail: spuissant @ ville-dijon. fr; https: // orcid. org / 0000 - 0003 - 4179 - 3636</mods:affiliation>
<mods:nameIdentifier id="8BCAAFC9EDA908EEC67CCB967A7413BD" type="email">spuissant@ville-dijon.fr</mods:nameIdentifier>
</mods:name>
<mods:name id="8E5BB13A25055B54B3181C71722BAAEE" type="personal">
<mods:role id="E56F7AC502E7248A86E84CB49824001E">
<mods:roleTerm id="CE8FFC7D811F71BB4E2375C047A3DD70">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="98D7C9F2AD4AAD11968B1C2038B18EF9">Gurcel, Kevin</mods:namePart>
<mods:nameIdentifier id="402EDC644DCD7A6D626ADF6CD354EE2C" type="ORCID">0009-0006-7380-9072</mods:nameIdentifier>
<mods:nameIdentifier id="0EA7127AC4E69C6486A0CE96680D450F" type="email">kevin.gurcel@orange.fr</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="4FB420EF9E9948A3501AA82D8C03718D">text</mods:typeOfResource>
<mods:relatedItem id="2C1EF1B9535C4733CC33FC56886EA976" type="host">
<mods:titleInfo id="F8E3C1B04ED7744B36CD5C75661E0000">
<mods:title id="9E6DA6B43718C0A4E02882DA9F3555DD">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="40BF8AD2C608988853C9A2DCEA71AD39">
<mods:date id="FC9112F34415ADFAA8EFE036E4A5F322">2023</mods:date>
<mods:detail id="607A5A05D8487EC1AC6F42A99628304D" type="pubDate">
<mods:number id="4495E3BB6F43933C02F2590FF371BB23">2023-08-17</mods:number>
</mods:detail>
<mods:detail id="C653AC49EFE25F2B79E7258481217587" type="volume">
<mods:number id="7952C4B4D178E1EE7CB6DA40351D58D8">5330</mods:number>
</mods:detail>
<mods:detail id="51737DA7ECE59FDD0AF56549879D9300" type="issue">
<mods:number id="6599BCE78EF852BA9BB1C72168333FAF">4</mods:number>
</mods:detail>
<mods:extent id="6DACBBC467DDBB61110CB703743D5901" unit="page">
<mods:start id="C9DB1730F09053303ECC256F5D49D330">535</mods:start>
<mods:end id="F43ECC5C449237D58D8BC401E752F721">560</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="794077013B4A907CF3A8D7DB47E2F7DA">
<mods:url id="F686E8A5C1E9D782D438421FF8866F32">http://dx.doi.org/10.11646/zootaxa.5330.4.4</mods:url>
</mods:location>
<mods:classification id="2B2EFA9C13ABE8295387D83E401BE655">journal article</mods:classification>
<mods:identifier id="D7A14E8D73CB6C0057E9DC22CFD9CEE5" type="CLB-Dataset">54879</mods:identifier>
<mods:identifier id="064D8BA4798D47E3BCF4A26BD23459D5" type="DOI">10.11646/zootaxa.5330.4.4</mods:identifier>
<mods:identifier id="5A0988E3D42CF4DB365F3B47B2EB693E" type="GBIF-Dataset">c089657b-b4cb-469a-9e1f-94a669a51fb1</mods:identifier>
<mods:identifier id="29F8BF15A98F453DFF1A40ACB8E472D4" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="86214F64581DF2C0B85FC2EA43A388C2" type="Zenodo-Dep">8255293</mods:identifier>
<mods:identifier id="BB24E5A2710C7682CBD2AAE01F89E6FE" type="ZooBank">8EC3E31F-ED7B-48BD-901B-1EE8AE0BAF02</mods:identifier>
</mods:mods>
<treatment id="1F20F502FFB1FF94FF22FB248DA1FD64" ID-DOI="http://doi.org/10.5281/zenodo.8255372" ID-Zenodo-Dep="8255372" LSID="urn:lsid:plazi:treatment:1F20F502FFB1FF94FF22FB248DA1FD64" httpUri="http://treatment.plazi.org/id/1F20F502FFB1FF94FF22FB248DA1FD64" lastPageId="15" lastPageNumber="550" pageId="8" pageNumber="543">
<subSubSection id="DF93179FFFB1FF93FF22FB248C18FB08" box="[151,882,1158,1185]" pageId="8" pageNumber="543" type="nomenclature">
<paragraph id="97364414FFB1FF93FF22FB248C18FB08" blockId="8.[151,882,1158,1185]" box="[151,882,1158,1185]" pageId="8" pageNumber="543">
<heading id="CC7EF378FFB1FF93FF22FB248C18FB08" bold="true" box="[151,882,1158,1185]" fontSize="11" level="1" pageId="8" pageNumber="543" reason="1">
<emphasis id="A5FD9806FFB1FF93FF22FB248C18FB08" bold="true" box="[151,882,1158,1185]" pageId="8" pageNumber="543">
<taxonomicName id="50893F97FFB1FF93FF22FB248C77FB09" authority="Puissant" authorityName="Puissant &amp; Gurcel" authorityYear="2023" box="[151,797,1158,1185]" class="Insecta" family="Cicadidae" genus="Tibicina" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="543" phylum="Arthropoda" rank="subSpecies" species="quadrisignata" status="ssp. n." subSpecies="pilleti">
<emphasis id="A5FD9806FFB1FF93FF22FB248EC0FB08" bold="true" box="[151,426,1158,1185]" italics="true" pageId="8" pageNumber="543">Tibicina quadrisignata</emphasis>
(
<bibRefCitation id="F31839E5FFB1FF93FE0FFB248D32FB09" author="Hagen, H. A." box="[442,600,1158,1184]" pageId="8" pageNumber="543" pagination="340 - 358" refId="ref13020" refString="Hagen, H. A. (1855) Die Sing-Cicaden Europa's. Jahresbericht des Entomologischen Vereins zu Stettin, 16, 340 - 358." type="journal article" year="1855">Hagen, 1855</bibRefCitation>
) 
<emphasis id="A5FD9806FFB1FF93FDD2FB248DC1FB08" bold="true" box="[615,683,1158,1185]" italics="true" pageId="8" pageNumber="543">pilleti</emphasis>
Puissant
</taxonomicName>
<taxonomicNameLabel id="BECE257DFFB1FF93FC91FB258C18FB08" box="[804,882,1159,1185]" pageId="8" pageNumber="543" rank="subSpecies">ssp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DF93179FFFB1FF94FF22FB698A17FEB4" lastPageId="15" lastPageNumber="550" pageId="8" pageNumber="543" type="materials_examined">
<paragraph id="97364414FFB1FF93FF22FB698E75FA96" blockId="8.[151,1437,1227,1343]" pageId="8" pageNumber="543">
<emphasis id="A5FD9806FFB1FF93FF22FB698E5FFB4A" bold="true" box="[151,309,1227,1251]" inLineHeading="true" pageId="8" pageNumber="543" reason="1">Type material.</emphasis>
<materialsCitation id="27E14E49FFB1FF93FE8BFB698E75FA96" collectingDate="2010-07-02" collectorName="A. Francois" country="Morocco" elevation="2135" latitude="33.66235" location="Pres" longLatPrecision="1" longitude="-4.13106" pageId="8" pageNumber="543" specimenCount="2" typeStatus="holotype">
<typeStatus id="4832FAB6FFB1FF93FE8BFB698EF6FB4A" box="[318,412,1227,1251]" pageId="8" pageNumber="543">Holotype</typeStatus>
male: 
<collectingCountry id="EF9E0484FFB1FF93FE51FB6E8D28FB4A" box="[484,578,1228,1251]" name="Morocco" pageId="8" pageNumber="543">Morocco</collectingCountry>
, “2135 m, 
<location id="925612CFFFB1FF93FD01FB698D8AFB4A" LSID="urn:lsid:plazi:treatment:1F20F502FFB1FF94FF22FB248DA1FD64:925612CFFFB1FF93FD01FB698D8AFB4A" box="[692,736,1227,1251]" country="Morocco" latitude="33.66235" longLatPrecision="1" longitude="-4.13106" name="Pres" pageId="8" pageNumber="543">Près</location>
d’un ruisseau, pente rocheuse” [label rectangular, pale yellow, with black margins, printed]; “ 
<date id="E33762D4FFB1FF93FE60FB488D27FAAB" box="[469,589,1258,1282]" pageId="8" pageNumber="543" value="2010-07-02">
<collectingDate id="F3739B3CFFB1FF93FE60FB488D27FAAB" box="[469,589,1258,1282]" pageId="8" pageNumber="543" value="2010-07-02">02/07/2010</collectingDate>
</date>
, Taffert, 
<geoCoordinate id="F2BD22D3FFB1FF93FD05FB488C65FAAB" box="[688,783,1258,1282]" degrees="33.66235" direction="north" orientation="latitude" pageId="8" pageNumber="543" precision="1" value="33.66235">33.66235</geoCoordinate>
/ 
<geoCoordinate id="F2BD22D3FFB1FF93FC9EFB488CE2FAAB" box="[811,904,1258,1282]" degrees="4.13106" direction="west" orientation="longitude" pageId="8" pageNumber="543" precision="1" value="-4.13106">-4.13106</geoCoordinate>
, 
<collectorName id="3A7C21C2FFB1FF93FC20FB488B64FAAB" box="[917,1038,1258,1282]" pageId="8" pageNumber="543">A. François</collectorName>
” [label rectangular, pale yellow, with black margins, printed] and “HOLOTYPE Ô, 
<taxonomicName id="50893F97FFB1FF93FD2DFAAA8CAAFA89" authority="Puissant &amp; Gurcel, 2023" authorityName="Puissant &amp; Gurcel" authorityYear="2023" box="[664,960,1288,1312]" class="Insecta" family="Cicadidae" genus="Tibicina" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="543" phylum="Arthropoda" rank="subSpecies" species="quadrisignata" status="ssp. n." subSpecies="pilleti">
<emphasis id="A5FD9806FFB1FF93FD2DFAAA8CAAFA89" box="[664,960,1288,1312]" italics="true" pageId="8" pageNumber="543">Tibicina quadrisignata pilleti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BECE257DFFB1FF93FC73FAAB8B6DFA88" box="[966,1031,1289,1313]" pageId="8" pageNumber="543" rank="subSpecies">ssp. n.</taxonomicNameLabel>
” [label rectangular, red, manuscript and printed].
</materialsCitation>
</paragraph>
<paragraph id="97364414FFB1FF93FF22FACC8C9EF9BE" blockId="8.[151,1437,1389,2028]" pageId="8" pageNumber="543">
<materialsCitation id="27E14E49FFB1FF93FF22FACC8D64FA02" country="Morocco" elevation="2135" location="Morocco" pageId="8" pageNumber="543" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<typeStatus id="4832FAB6FFB1FF93FF22FACC8E6EFA21" box="[151,260,1390,1416]" pageId="8" pageNumber="543">Paratypes</typeStatus>
(3 Ô, 
<specimenCount id="818F8F9DFFB1FF93FEE2FACF8EEBFA2E" box="[343,385,1389,1415]" pageId="8" pageNumber="543" type="female">1 ♀</specimenCount>
): 
<collectingCountry id="EF9E0484FFB1FF93FE2BFACC8D69FA21" box="[414,515,1390,1416]" name="Morocco" pageId="8" pageNumber="543">Morocco</collectingCountry>
3 Ô, “ 
<quantity id="5071E9F1FFB1FF93FDE2FACF8DDAFA21" box="[599,688,1389,1416]" metricMagnitude="3" metricUnit="m" metricValue="2.135" pageId="8" pageNumber="543" unit="m" value="2135.0">
<elevation id="1CA4A327FFB1FF93FDE2FACF8DDAFA21" box="[599,688,1389,1416]" metricMagnitude="3" metricUnit="m" metricValue="2.135" pageId="8" pageNumber="543" unit="m" value="2135.0">2135 m</elevation>
</quantity>
, Près d’un ruisseau, pente rocheuse” [Label rectangular, white, with black margins, manuscript]
</materialsCitation>
; “ 
<date id="E33762D4FFB1FF93FD92FA338DDAFA02" box="[551,688,1425,1451]" pageId="8" pageNumber="543" value="2022-06-27">27-VI-2022</date>
, Taffert, 
<geoCoordinate id="F2BD22D3FFB1FF93FCAEFA338CEEFA05" box="[795,900,1425,1452]" degrees="33.66235" direction="north" orientation="latitude" pageId="8" pageNumber="543" precision="1" value="33.66235">33.66235</geoCoordinate>
/ 
<geoCoordinate id="F2BD22D3FFB1FF93FC15FA338B62FA02" box="[928,1032,1425,1451]" degrees="4.13106" direction="west" orientation="longitude" pageId="8" pageNumber="543" precision="1" value="-4.13106">-4.13106</geoCoordinate>
, S. Puissant &amp; A. François” [Label rectangular, white, with black margins, manuscript] and “ 
<typeStatus id="4832FAB6FFB1FF93FCBAFA178CF7FA66" box="[783,925,1461,1487]" pageId="8" pageNumber="543">PARATYPE</typeStatus>
Ô 
<taxonomicName id="50893F97FFB1FF93FC0BFA178A69FA66" authority="Puissant &amp; Gurcel, 2023" authorityName="Puissant &amp; Gurcel" authorityYear="2023" box="[958,1283,1461,1487]" class="Insecta" family="Cicadidae" genus="Tibicina" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="543" phylum="Arthropoda" rank="subSpecies" species="quadrisignata" status="ssp. n." subSpecies="pilleti">
<emphasis id="A5FD9806FFB1FF93FC0BFA178A69FA66" box="[958,1283,1461,1487]" italics="true" pageId="8" pageNumber="543">Tibicina quadrisignata pilleti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BECE257DFFB1FF93FABCFA148A3BFA79" box="[1289,1361,1462,1488]" pageId="8" pageNumber="543" rank="subSpecies">ssp. n.</taxonomicNameLabel>
, [label rectangular, red, manuscript and printed]; 
<materialsCitation id="27E14E49FFB1FF93FDD9FA788C9AF9BE" country="Morocco" location="Morocco" pageId="8" pageNumber="543" specimenCount="3" specimenCount-female="2" typeStatus="paratype">
<collectingCountry id="EF9E0484FFB1FF93FDD9FA788DBBFA5D" box="[620,721,1498,1524]" name="Morocco" pageId="8" pageNumber="543">Morocco</collectingCountry>
<specimenCount id="818F8F9DFFB1FF93FD6DFA7B8C68FA5D" box="[728,770,1497,1524]" pageId="8" pageNumber="543" type="female">1 ♀</specimenCount>
, data as the 
<specimenCount id="818F8F9DFFB1FF93FC39FA7B8B5FFA5D" box="[908,1077,1497,1524]" pageId="8" pageNumber="543" type="generic" typeStatus="paratypes">three paratypes</specimenCount>
Ô and “ 
<typeStatus id="4832FAB6FFB1FF93FB25FA7B8A74FA5A" box="[1168,1310,1497,1523]" pageId="8" pageNumber="543">PARATYPE</typeStatus>
<specimenCount id="818F8F9DFFB1FF93FA90FA788A50FA5D" box="[1317,1338,1498,1524]" pageId="8" pageNumber="543" type="female">♀</specimenCount>
<taxonomicName id="50893F97FFB1FF93FAF5FA7B8E16F9BE" authority="Puissant &amp; Gurcel, 2023" authorityName="Puissant &amp; Gurcel" authorityYear="2023" class="Insecta" family="Cicadidae" genus="Tibicina" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="543" phylum="Arthropoda" rank="subSpecies" species="quadrisignata" status="ssp. n." subSpecies="pilleti">
<emphasis id="A5FD9806FFB1FF93FAF5FA7B8E16F9BE" italics="true" pageId="8" pageNumber="543">Tibicina quadrisignata pilleti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BECE257DFFB1FF93FE31FA5C8EA1F9BE" box="[388,459,1533,1560]" pageId="8" pageNumber="543" rank="subSpecies">ssp. n.</taxonomicNameLabel>
” [label rectangular, red, manuscript and printed]
</materialsCitation>
.
</paragraph>
<paragraph id="97364414FFB1FF92FF72F9838D63FEB4" blockId="8.[151,1437,1389,2028]" lastBlockId="9.[151,1437,150,2013]" lastPageId="9" lastPageNumber="544" pageId="8" pageNumber="543">
<typeStatus id="4832FAB6FFB1FF93FF72F9838EEFF992" box="[199,389,1569,1595]" isOtherTypeStatus="true" pageId="8" pageNumber="543">Other specimens</typeStatus>
used for morphological description but not include in the 
<typeStatus id="4832FAB6FFB1FF93FB95F9808B3AF995" box="[1056,1104,1570,1596]" pageId="8" pageNumber="543">type</typeStatus>
series (3 Ô, 
<specimenCount id="818F8F9DFFB1FF93FB51F9838A64F995" box="[1252,1294,1569,1596]" pageId="8" pageNumber="543" type="female">2 ♀</specimenCount>
, 2 exuviae): 
<materialsCitation id="27E14E49FFB1FF93FF22F9E48CF0F962" collectingDate="2015-06-28" collectorName="A. Francois &amp; B. Defaut" country="Morocco" county="d'Alep" elevation="2023" latitude="32.58735" location="Tizi n'Talghemt" longLatPrecision="1" longitude="-4.55191" municipality="Steppe" pageId="8" pageNumber="543" specimenCount="1">
<collectingCountry id="EF9E0484FFB1FF93FF22F9E48F96F9C9" box="[151,252,1606,1632]" name="Morocco" pageId="8" pageNumber="543">Morocco</collectingCountry>
1 Ô “ 
<location id="925612CFFFB1FF93FEF7F9E78E97F9F6" LSID="urn:lsid:plazi:treatment:1F20F502FFB1FF94FF22FB248DA1FD64:925612CFFFB1FF93FEF7F9E78E97F9F6" box="[322,509,1605,1631]" country="Morocco" county="d'Alep" latitude="32.58735" longLatPrecision="1" longitude="-4.55191" municipality="Steppe" name="Tizi n'Talghemt" pageId="8" pageNumber="543">Tizi n’Talghemt</location>
, 
<quantity id="5071E9F1FFB1FF93FDBFF9E78D0BF9C9" box="[522,609,1605,1632]" metricMagnitude="3" metricUnit="m" metricValue="2.023" pageId="8" pageNumber="543" unit="m" value="2023.0">
<elevation id="1CA4A327FFB1FF93FDBFF9E78D0BF9C9" box="[522,609,1605,1632]" metricMagnitude="3" metricUnit="m" metricValue="2.023" pageId="8" pageNumber="543" unit="m" value="2023.0">2023 m</elevation>
</quantity>
, 
<collectingMunicipality id="7752DE6EFFB1FF93FDDBF9E78DD0F9F6" box="[622,698,1605,1631]" pageId="8" pageNumber="543">Steppe</collectingMunicipality>
à armoises et plantation de pins 
<collectingCounty id="7E573C98FFB1FF93FB80F9E78BEFF9F6" box="[1077,1157,1605,1631]" pageId="8" pageNumber="543">d’Alep</collectingCounty>
” [label rectangular, pale yellow, with black margins, printed]; “ 
<date id="E33762D4FFB1FF93FDFAF9CB8DB9F92D" box="[591,723,1641,1668]" pageId="8" pageNumber="543" value="2015-06-28">
<collectingDate id="F3739B3CFFB1FF93FDFAF9CB8DB9F92D" box="[591,723,1641,1668]" pageId="8" pageNumber="543" value="2015-06-28">28/06/2015</collectingDate>
</date>
, Midelt, 
<geoCoordinate id="F2BD22D3FFB1FF93FC8EF9CB8CCEF92D" box="[827,932,1641,1668]" degrees="32.58735" direction="north" orientation="latitude" pageId="8" pageNumber="543" precision="1" value="32.58735">32.58735</geoCoordinate>
/ 
<geoCoordinate id="F2BD22D3FFB1FF93FC08F9CB8B4FF92D" box="[957,1061,1641,1668]" degrees="4.55191" direction="west" orientation="longitude" pageId="8" pageNumber="543" precision="1" value="-4.55191">-4.55191</geoCoordinate>
, 
<collectorName id="3A7C21C2FFB1FF93FB85F9C88BD8F92A" box="[1072,1202,1641,1668]" pageId="8" pageNumber="543">A. François</collectorName>
&amp; 
<collectorName id="3A7C21C2FFB1FF93FB6FF9C88A20F92A" box="[1242,1354,1641,1668]" pageId="8" pageNumber="543">B. Defaut</collectorName>
” [label rectangular, pale yellow, with black margins, printed] and “ 
<taxonomicName id="50893F97FFB1FF93FC83F92F8AF7F90E" authority=", Puissant &amp; Hertach" authorityName="Puissant &amp; Hertach" box="[822,1437,1677,1704]" class="Insecta" family="Cicadidae" genus="Tibicina" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="543" phylum="Arthropoda" rank="subSpecies" species="quadrisignata" subSpecies="pilleti">
<emphasis id="A5FD9806FFB1FF93FC83F92F8B5CF90E" box="[822,1078,1677,1703]" italics="true" pageId="8" pageNumber="543">Tibicina quadrisignata</emphasis>
ssp. 
<emphasis id="A5FD9806FFB1FF93FBC4F92F8BD8F90E" box="[1137,1202,1677,1703]" italics="true" pageId="8" pageNumber="543">pilleti</emphasis>
, Puissant &amp; Hertach
</taxonomicName>
dét. 2021” [label rectangular, white, with black margins, manuscript]
</materialsCitation>
; 
<materialsCitation id="27E14E49FFB1FF93FC10F9108E4AF89E" collectorName="Nadig" country="Morocco" elevation="1600" location="Tizi Tfri" pageId="8" pageNumber="543" specimenCount="1">
<collectingCountry id="EF9E0484FFB1FF93FC10F9108B60F965" box="[933,1034,1714,1740]" name="Morocco" pageId="8" pageNumber="543">Morocco</collectingCountry>
2 Ô “Maroc 68 MA 68:27, 
<location id="925612CFFFB1FF93FA8AF9138AF7F962" LSID="urn:lsid:plazi:treatment:1F20F502FFB1FF94FF22FB248DA1FD64:925612CFFFB1FF93FA8AF9138AF7F962" box="[1343,1437,1713,1739]" country="Morocco" name="Tizi Tfri" pageId="8" pageNumber="543">Tizi Tfri</location>
(S Targuist), 
<quantity id="5071E9F1FFB1FF93FE99F9778EE9F959" box="[300,387,1749,1776]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="8" pageNumber="543" unit="m" value="1600.0">
<elevation id="1CA4A327FFB1FF93FE99F9778EE9F959" box="[300,387,1749,1776]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="8" pageNumber="543" unit="m" value="1600.0">1600 m</elevation>
</quantity>
, 27.6.68, leg. 
<collectorName id="3A7C21C2FFB1FF93FD92F9778D04F946" box="[551,622,1749,1775]" pageId="8" pageNumber="543">Nadig</collectorName>
” [label rectangular, white, with black margins, manuscript and printed] and “ 
<taxonomicName id="50893F97FFB1FF93FF67F95B8C41F8BA" authority=", Puissant &amp; Hertach" authorityName="Puissant &amp; Hertach" box="[210,811,1785,1812]" class="Insecta" family="Cicadidae" genus="Tibicina" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="543" phylum="Arthropoda" rank="subSpecies" species="quadrisignata" subSpecies="pilleti">
<emphasis id="A5FD9806FFB1FF93FF67F95B8EBAF8BA" box="[210,464,1785,1811]" italics="true" pageId="8" pageNumber="543">Tibicina quadrisignata</emphasis>
ssp. 
<emphasis id="A5FD9806FFB1FF93FDB3F95B8D2DF8BA" box="[518,583,1785,1811]" italics="true" pageId="8" pageNumber="543">pilleti</emphasis>
, Puissant &amp; Hertach
</taxonomicName>
dét. 2021” [label rectangular, white, with black margins, manuscript]
</materialsCitation>
; 
<materialsCitation id="27E14E49FFB1FF93FE9AF8BC8B75F8D6" collectorName="Nadig" country="Morocco" elevation="600" location="Taounate-Ain" municipality="Aicha" pageId="8" pageNumber="543" specimenCount="1" specimenCount-female="1">
<collectingCountry id="EF9E0484FFB1FF93FE9AF8BC8EFEF891" box="[303,404,1822,1848]" name="Morocco" pageId="8" pageNumber="543">Morocco</collectingCountry>
<specimenCount id="818F8F9DFFB1FF93FE2BF8BF8EA2F891" box="[414,456,1821,1848]" pageId="8" pageNumber="543" type="female">1 ♀</specimenCount>
“Maroc 70 MA 70: 82, 
<location id="925612CFFFB1FF93FD50F8BD8CE8F893" LSID="urn:lsid:plazi:treatment:1F20F502FFB1FF94FF22FB248DA1FD64:925612CFFFB1FF93FD50F8BD8CE8F893" box="[741,898,1823,1850]" country="Morocco" municipality="Aicha" name="Taounate-Ain" pageId="8" pageNumber="543">Taounate-Âin</location>
, 
<collectingMunicipality id="7752DE6EFFB1FF93FC25F8BD8CB9F893" box="[912,979,1823,1850]" pageId="8" pageNumber="543">Âïcha</collectingMunicipality>
<quantity id="5071E9F1FFB1FF93FC68F8BF8B0BF891" box="[989,1121,1821,1848]" metricMagnitude="2" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="8" pageNumber="543" unit="m" value="600.0" valueMax="700.0" valueMin="500.0">
<elevation id="1CA4A327FFB1FF93FC68F8BF8B0BF891" box="[989,1121,1821,1848]" metricMagnitude="2" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="8" pageNumber="543" unit="m" value="600.0" valueMax="700.0" valueMin="500.0">500–700 m</elevation>
</quantity>
, 15.7.70 leg. 
<collectorName id="3A7C21C2FFB1FF93FAB4F8BF8A22F89E" box="[1281,1352,1821,1847]" pageId="8" pageNumber="543">Nadig</collectorName>
” [label rectangular, white, with black margins, manuscript and printed] and “ 
<taxonomicName id="50893F97FFB1FF93FC05F8E38E67F8D6" authority=", Puissant &amp; Hertach" authorityName="Puissant &amp; Hertach" class="Insecta" family="Cicadidae" genus="Tibicina" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="543" phylum="Arthropoda" rank="subSpecies" species="quadrisignata" subSpecies="pilleti">
<emphasis id="A5FD9806FFB1FF93FC05F8E38BDBF8F2" box="[944,1201,1857,1883]" italics="true" pageId="8" pageNumber="543">Tibicina quadrisignata</emphasis>
ssp. 
<emphasis id="A5FD9806FFB1FF93FB5BF8E38A45F8F2" box="[1262,1327,1857,1883]" italics="true" pageId="8" pageNumber="543">pilleti</emphasis>
, Puissant &amp; Hertach
</taxonomicName>
dét. 2021” [label rectangular, white, with black margins, manuscript]
</materialsCitation>
; 
<materialsCitation id="27E14E49FFB1FF93FB9EF8C48B16F842" collectingDate="2021-07-03" collectorName="A. Francois &amp; G. Lienart" country="Morocco" elevation="1655" latitude="32.9024" location="Morocco" longLatPrecision="5" longitude="-5.22426" pageId="8" pageNumber="543" specimenCount="1" specimenCount-female="1">
<collectingCountry id="EF9E0484FFB1FF93FB9EF8C48BFAF829" box="[1067,1168,1894,1920]" name="Morocco" pageId="8" pageNumber="543">Morocco</collectingCountry>
<specimenCount id="818F8F9DFFB1FF93FB22F8C78BABF829" box="[1175,1217,1893,1920]" pageId="8" pageNumber="543" type="female">1 ♀</specimenCount>
“ 
<quantity id="5071E9F1FFB1FF93FB61F8C78A46F829" box="[1236,1324,1893,1920]" metricMagnitude="3" metricUnit="m" metricValue="1.655" pageId="8" pageNumber="543" unit="m" value="1655.0">
<elevation id="1CA4A327FFB1FF93FB61F8C78A46F829" box="[1236,1324,1893,1920]" metricMagnitude="3" metricUnit="m" metricValue="1.655" pageId="8" pageNumber="543" unit="m" value="1655.0">1655 m</elevation>
</quantity>
, Cédraie” [label rectangular, pale yellow, with black margins, printed]; “ 
<date id="E33762D4FFB1FF93FCEBF82B8C88F80A" box="[862,994,1929,1955]" pageId="8" pageNumber="543" value="2021-07-03">
<collectingDate id="F3739B3CFFB1FF93FCEBF82B8C88F80A" box="[862,994,1929,1955]" pageId="8" pageNumber="543" value="2021-07-03">03/07/2021</collectingDate>
</date>
, Itzer, 
<geoCoordinate id="F2BD22D3FFB1FF93FB87F82B8BE7F80A" box="[1074,1165,1929,1955]" degrees="32.9024" direction="north" orientation="latitude" pageId="8" pageNumber="543" precision="5" value="32.9024">32.9024</geoCoordinate>
/ 
<geoCoordinate id="F2BD22D3FFB1FF93FB12F82B8A64F80D" box="[1191,1294,1929,1956]" degrees="5.22426" direction="west" orientation="longitude" pageId="8" pageNumber="543" precision="1" value="-5.22426">-5.22426</geoCoordinate>
, 
<collectorName id="3A7C21C2FFB1FF93FAAFF8288AF6F80A" box="[1306,1436,1929,1956]" pageId="8" pageNumber="543">A. François</collectorName>
&amp; 
<collectorName id="3A7C21C2FFB1FF93FF01F80F8E40F86E" box="[180,298,1965,1991]" pageId="8" pageNumber="543">G. Liénart</collectorName>
” [label rectangular, pale yellow, with black margins, printed] and “ 
<taxonomicName id="50893F97FFB1FF93FBAEF80F8E1BF842" authority=", Puissant &amp; Hertach" authorityName="Puissant &amp; Hertach" class="Insecta" family="Cicadidae" genus="Tibicina" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="543" phylum="Arthropoda" rank="subSpecies" species="quadrisignata" subSpecies="pilleti">
<emphasis id="A5FD9806FFB1FF93FBAEF80F8A71F86E" box="[1051,1307,1965,1991]" italics="true" pageId="8" pageNumber="543">Tibicina quadrisignata</emphasis>
ssp. 
<emphasis id="A5FD9806FFB1FF93FAE1F80F8AFFF86E" box="[1364,1429,1965,1991]" italics="true" pageId="8" pageNumber="543">pilleti</emphasis>
, Puissant &amp; Hertach
</taxonomicName>
dét. 2021” [label rectangular, white, with black margins, manuscript]
</materialsCitation>
; 
<materialsCitation id="27E14E49FFB1FF92FB32F8708D6EFEB4" collectingDate="2022-06-27" collectorName="S. Puissant &amp; A. Francois" country="Morocco" elevation="2135" lastPageId="9" lastPageNumber="544" latitude="33.66235" location="Label" longLatPrecision="1" longitude="-4.13106" pageId="8" pageNumber="543" specimenCount="1">
<collectingCountry id="EF9E0484FFB1FF93FB32F8708B86F845" box="[1159,1260,2002,2028]" name="Morocco" pageId="8" pageNumber="543">Morocco</collectingCountry>
2 exuviae from the 
<emphasis id="A5FD9806FFB0FF92FF70FF358E30FF19" box="[197,346,151,177]" italics="true" pageId="9" pageNumber="544">locus typicus</emphasis>
“ 
<quantity id="5071E9F1FFB0FF92FEDAFF348EA3FF18" box="[367,457,150,177]" metricMagnitude="3" metricUnit="m" metricValue="2.135" pageId="9" pageNumber="544" unit="m" value="2135.0">
<elevation id="1CA4A327FFB0FF92FEDAFF348EA3FF18" box="[367,457,150,177]" metricMagnitude="3" metricUnit="m" metricValue="2.135" pageId="9" pageNumber="544" unit="m" value="2135.0">2135 m</elevation>
</quantity>
, Près d’un ruisseau, pente rocheuse” [
<location id="925612CFFFB0FF92FC3FFF358CA7FF18" LSID="urn:lsid:plazi:treatment:1F20F502FFB1FF94FF22FB248DA1FD64:925612CFFFB0FF92FC3FFF358CA7FF18" box="[906,973,151,177]" country="Morocco" latitude="33.66235" longLatPrecision="1" longitude="-4.13106" name="Label" pageId="9" pageNumber="544">Label</location>
rectangular, white, with black margins, manuscript]; “ 
<date id="E33762D4FFB0FF92FE83FF198ED4FF7C" box="[310,446,187,213]" pageId="9" pageNumber="544" value="2022-06-27">
<collectingDate id="F3739B3CFFB0FF92FE83FF198ED4FF7C" box="[310,446,187,213]" pageId="9" pageNumber="544" value="2022-06-27">27-VI-2022</collectingDate>
</date>
, Taffert, 
<geoCoordinate id="F2BD22D3FFB0FF92FD97FF188DE1FF7C" box="[546,651,186,213]" degrees="33.66235" direction="north" orientation="latitude" pageId="9" pageNumber="544" precision="1" value="33.66235">33.66235</geoCoordinate>
/ 
<geoCoordinate id="F2BD22D3FFB0FF92FD15FF198C62FF7C" box="[672,776,187,213]" degrees="4.13106" direction="west" orientation="longitude" pageId="9" pageNumber="544" precision="1" value="-4.13106">-4.13106</geoCoordinate>
, 
<collectorName id="3A7C21C2FFB0FF92FCA7FF198CE7FF7C" box="[786,909,187,213]" pageId="9" pageNumber="544">S. Puissant</collectorName>
&amp; 
<collectorName id="3A7C21C2FFB0FF92FC1AFF198B5EFF7C" box="[943,1076,187,213]" pageId="9" pageNumber="544">A. François</collectorName>
” [Label rectangular, white, with black margins, manuscript] and “ 
<taxonomicName id="50893F97FFB0FF92FDBDFF7D8B6DFF50" authority=", S. Puissant" authorityName="S. Puissant" box="[520,1031,223,249]" class="Insecta" family="Cicadidae" genus="Tibicina" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="544" phylum="Arthropoda" rank="subSpecies" species="quadrisignata" subSpecies="pilleti">
<emphasis id="A5FD9806FFB0FF92FDBDFF7D8C6DFF50" box="[520,775,223,249]" italics="true" pageId="9" pageNumber="544">Tibicina quadrisignata</emphasis>
ssp. 
<emphasis id="A5FD9806FFB0FF92FC8BFF7D8C15FF50" box="[830,895,223,249]" italics="true" pageId="9" pageNumber="544">pilleti</emphasis>
, S. Puissant
</taxonomicName>
dét. 2022” [label rectangular, white, with black margins, manuscript]
</materialsCitation>
.
</paragraph>
<paragraph id="97364414FFB0FF92FF72FE858CF5FE04" blockId="9.[151,1437,150,2013]" pageId="9" pageNumber="544">
The 
<typeStatus id="4832FAB6FFB0FF92FF49FE858E0BFEE8" box="[252,353,295,321]" pageId="9" pageNumber="544">holotype</typeStatus>
, 
<specimenCount id="818F8F9DFFB0FF92FED8FE858EBFFEE8" box="[365,469,295,321]" pageId="9" pageNumber="544" type="male">one male</specimenCount>
collected by 
<collectorName id="3A7C21C2FFB0FF92FDC4FE858DBBFEE8" box="[625,721,295,321]" pageId="9" pageNumber="544">François</collectorName>
&amp; 
<collectorName id="3A7C21C2FFB0FF92FD4CFE858C2FFEE8" box="[761,837,295,321]" pageId="9" pageNumber="544">Defaut</collectorName>
and 
<specimenCount id="818F8F9DFFB0FF92FCCAFE858C97FEE8" box="[895,1021,295,321]" pageId="9" pageNumber="544" type="female">one female</specimenCount>
collected by 
<collectorName id="3A7C21C2FFB0FF92FB2DFE858B92FEE8" box="[1176,1272,295,321]" pageId="9" pageNumber="544">François</collectorName>
&amp; 
<collectorName id="3A7C21C2FFB0FF92FA95FE858A1BFEE8" box="[1312,1393,295,321]" pageId="9" pageNumber="544">Liénart</collectorName>
are deposited in the 
<collectionCode id="F198DCD1FFB0FF92FEFAFEE98ECFFECC" box="[335,421,331,357]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="9" pageNumber="544">MNHN</collectionCode>
(
<location id="925612CFFFB0FF92FE06FEE98E85FECC" LSID="urn:lsid:plazi:treatment:1F20F502FFB1FF94FF22FB248DA1FD64:925612CFFFB0FF92FE06FEE98E85FECC" box="[435,495,331,357]" country="France" name="Paris" pageId="9" pageNumber="544">Paris</location>
, 
<collectingCountry id="EF9E0484FFB0FF92FE4EFEE98D26FECC" box="[507,588,331,357]" name="France" pageId="9" pageNumber="544">France</collectingCountry>
). 
<collectorName id="3A7C21C2FFB0FF92FDEBFEE98DE0FECC" box="[606,650,331,357]" pageId="9" pageNumber="544">The</collectorName>
<specimenCount id="818F8F9DFFB0FF92FD25FEE98C5BFECC" box="[656,817,331,357]" pageId="9" pageNumber="544" type="generic" typeStatus="paratypes">four paratypes</specimenCount>
and the 2 exuviae collected by 
<collectorName id="3A7C21C2FFB0FF92FB27FEE98B85FECC" box="[1170,1263,331,357]" pageId="9" pageNumber="544">Puissant</collectorName>
&amp; 
<collectorName id="3A7C21C2FFB0FF92FAA6FEE98A19FECC" box="[1299,1395,331,357]" pageId="9" pageNumber="544">François</collectorName>
are deposited in the 
<collectorName id="3A7C21C2FFB0FF92FEF9FECD8D4CFE20" box="[332,550,367,393]" pageId="9" pageNumber="544">Museum d’Histoire</collectorName>
naturelle 
<collectorName id="3A7C21C2FFB0FF92FD26FECD8D99FE20" box="[659,755,367,393]" pageId="9" pageNumber="544">de Dijon</collectorName>
(
<collectionCode id="F198DCD1FFB0FF92FCB5FECD8C30FE20" box="[768,858,367,393]" pageId="9" pageNumber="544">MHND</collectionCode>
, 
<location id="925612CFFFB0FF92FCD1FECD8CCDFE20" LSID="urn:lsid:plazi:treatment:1F20F502FFB1FF94FF22FB248DA1FD64:925612CFFFB0FF92FCD1FECD8CCDFE20" box="[868,935,367,393]" country="France" name="Dijon" pageId="9" pageNumber="544">Dijon</location>
, 
<collectingCountry id="EF9E0484FFB0FF92FC04FECD8B68FE20" box="[945,1026,367,393]" name="France" pageId="9" pageNumber="544">France</collectingCountry>
). 
<materialsCitation id="27E14E49FFB0FF92FBA7FECD8CF1FE04" collectorName="Nadig" country="Switzerland" location="Geneva" pageId="9" pageNumber="544" specimenCount="1" stateProvince="Geneve" typeStatus="paratype">
The two other males and the female collected by 
<collectorName id="3A7C21C2FFB0FF92FE93FE318E00FE04" box="[294,362,403,429]" pageId="9" pageNumber="544">Nadig</collectorName>
are deposited in the MHN (
<collectingRegion id="554D8AF6FFB0FF92FD16FE318D97FE04" box="[675,765,403,429]" country="Switzerland" name="Geneve" pageId="9" pageNumber="544">Geneva</collectingRegion>
, 
<collectingCountry id="EF9E0484FFB0FF92FCBDFE318CF9FE04" box="[776,915,403,429]" name="Switzerland" pageId="9" pageNumber="544">Switzerland</collectingCountry>
)
</materialsCitation>
.
</paragraph>
<paragraph id="97364414FFB0FF92FF72FE148B5CFE5C" blockId="9.[151,1437,150,2013]" pageId="9" pageNumber="544">
<emphasis id="A5FD9806FFB0FF92FF72FE148ED5FE78" bold="true" box="[199,447,438,465]" pageId="9" pageNumber="544">Derivation of name.</emphasis>
Named in honor of our late colleague Jean-Marc Pillet 
<superScript id="60FCE95CFFB0FF92FBE2FE148B35FE6D" attach="left" box="[1111,1119,438,452]" fontSize="6" pageId="9" pageNumber="544">†</superScript>
(
<collectionCode id="F198DCD1FFB0FF92FBC6FE158BC2FE78" box="[1139,1192,439,465]" pageId="9" pageNumber="544">JMP</collectionCode>
<superScript id="60FCE95CFFB0FF92FB1CFE148BDBFE6D" attach="right" box="[1193,1201,438,452]" fontSize="6" pageId="9" pageNumber="544">†</superScript>
: 1952-2010), Swiss naturalist and herpetologist who collected cicadas from 
<collectingCountry id="EF9E0484FFB0FF92FCBCFE798C04FE5C" box="[777,878,475,501]" name="Morocco" pageId="9" pageNumber="544">Morocco</collectingCountry>
over many years.
</paragraph>
<paragraph id="97364414FFB0FF92FF72FE5C8C73FD64" blockId="9.[151,1437,150,2013]" pageId="9" pageNumber="544">
<emphasis id="A5FD9806FFB0FF92FF72FE5C8EC5FDB0" bold="true" box="[199,431,510,537]" pageId="9" pageNumber="544">Adults morphology</emphasis>
(
<figureCitation id="0FB25891FFB0FF92FE0AFE5D8D40FDB0" box="[447,554,510,537]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStartId-0="2.[151,250,1242,1266]" captionStartId-1="3.[151,250,1618,1642]" captionStartId-2="4.[151,250,1943,1967]" captionStartId-3="5.[151,250,1908,1932]" captionStartId-4="6.[151,250,1943,1967]" captionTargetBox-0="[151,1436,179,1216]" captionTargetBox-1="[151,1436,181,1594]" captionTargetBox-2="[177,1410,181,1919]" captionTargetBox-3="[151,1436,849,1885]" captionTargetBox-4="[213,1374,181,1919]" captionTargetId-0="figure-22@2.[151,1436,179,1216]" captionTargetId-1="figure-17@3.[151,1436,181,1594]" captionTargetId-2="figure-22@4.[177,1410,181,1919]" captionTargetId-3="figure-327@5.[151,1436,849,1885]" captionTargetId-4="figure-22@6.[213,1374,181,1919]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionTargetPageId-4="6" captionText-0="FIGURE 1. Tibicina quadrisignata pilleti Puissant ssp. n., Holotype male, Morocco. A, habitus, dorsal view. B, lateral view of pygofer, uncus curved with apex highly angular (↑). C, lateral view of dissected aedeagus, vesica of median lobe curved and inflated with apex short. D, distal end of aedeagus deeply curved and trilobed, the two lateral lobes small and edged, strongly elongated and rounded at their apex with numerous conjunctival claws (↑)." captionText-1="FIGURE 2. Tibicina maldesi Boulard, 1981, male, Morocco. A, habitus, dorsal view. B, lateral view of pygofer, uncus curved with apex slightly angular, vesica of median lobe curved and inflated with apex elongated. C, distal end of aedeagus deeply curved and trilobed, the two lateral lobes small, curved and rounded at their apex with numerous conjunctival claws (↑)." captionText-2="FIGURE 3. Right tymbal of T. maldesi. Series of six short ribs (SR) alternate with seven long ribs (LR). Tymbal plate (TP)." captionText-3="FIGURE 4. T. q. pilleti Puissant ssp. n., right lateral view of exuviae. A, habitus. B, forelimb robust with accessory tooth of femur (↑) long and flattened. C, head with bulging postclypeus (↑)." captionText-4="FIGURE 5. Male of T. q. pilleti Puissant ssp. n. in calling posture." figureDoi-0="http://doi.org/10.5281/zenodo.8255297" figureDoi-1="http://doi.org/10.5281/zenodo.8255299" figureDoi-2="http://doi.org/10.5281/zenodo.8255301" figureDoi-3="http://doi.org/10.5281/zenodo.8255305" figureDoi-4="http://doi.org/10.5281/zenodo.8255307" httpUri-0="https://zenodo.org/record/8255297/files/figure.png" httpUri-1="https://zenodo.org/record/8255299/files/figure.png" httpUri-2="https://zenodo.org/record/8255301/files/figure.png" httpUri-3="https://zenodo.org/record/8255305/files/figure.png" httpUri-4="https://zenodo.org/record/8255307/files/figure.png" pageId="9" pageNumber="544">Figs. 1–5</figureCitation>
). 
<emphasis id="A5FD9806FFB0FF92FDF5FE5D8D17FDB0" box="[576,637,511,537]" italics="true" pageId="9" pageNumber="544">Head</emphasis>
: slightly less broad than mesonotum; black except base of postclypeus brownish and lateral margins of postclypeus yellowish. Eyes brownish but greyish-bluish when specimen alive (
<figureCitation id="0FB25891FFB0FF92FF2AFDE58F8EFDC8" box="[159,228,582,609]" captionStart="FIGURE 5" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[213,1374,181,1919]" captionTargetId="figure-22@6.[213,1374,181,1919]" captionTargetPageId="6" captionText="FIGURE 5. Male of T. q. pilleti Puissant ssp. n. in calling posture." figureDoi="http://doi.org/10.5281/zenodo.8255307" httpUri="https://zenodo.org/record/8255307/files/figure.png" pageId="9" pageNumber="544">Fig. 5</figureCitation>
). Ocellus yellowish or ochraceous, the distance between lateral ocelli almost equal than the distance between each lateral ocelli and the nearest eye. Antennae blackish, supra-antennal plates yellowish. Anteclypeus black. Rostrum with the mentum brownish and the labium black. Apex of rostrum reaching base of meso-coxa. Gena and lorum blackish with dense and long light yellowish hairs.
</paragraph>
<paragraph id="97364414FFB0FF92FF72FD758A4AFC60" blockId="9.[151,1437,150,2013]" pageId="9" pageNumber="544">
<emphasis id="A5FD9806FFB0FF92FF72FD758E72FD58" box="[199,280,727,753]" italics="true" pageId="9" pageNumber="544">Thorax</emphasis>
: pronotum black except an interrupted yellowish median line, yellowish marks enclosing the posterior oblique fissures, a pair of yellowish marks between both pairs of oblique fissures, the yellowish posterior margin. Mesonotum black, scutum with a pair of anterior yellow fasciae in inverted triangular shape and a pair of posterior triangular-shaped, yellow fasciae. Cruciform elevation with posterior branches yellowish and anterior branches black, mesonotal scutellar cords yellowish; wing groove yellowish. Stridulatory file, 
<emphasis id="A5FD9806FFB0FF92FBF7FCC58BB8FC28" box="[1090,1234,871,897]" italics="true" pageId="9" pageNumber="544">pars stridens</emphasis>
, present on lateral angle of mesonotum. Metanotum yellowish with black anterior median margin. Metasternum blackish with a pair of lateral yellowish spots; opercula very distant from each other, black with posterior margin yellowish.
</paragraph>
<paragraph id="97364414FFB0FF92FF72FC718BE9FBF0" blockId="9.[151,1437,150,2013]" pageId="9" pageNumber="544">
<emphasis id="A5FD9806FFB0FF92FF72FC718E61FC45" box="[199,267,979,1004]" italics="true" pageId="9" pageNumber="544">Wings</emphasis>
: fore wing hyaline, with eight apical cells; anterior and posterior cubital veins black, median and cubital veins emerging separately from the apex of the basal cell; venation mostly blackish but slightly brownish at apical cells. Hind wing with six apical cells; venation blackish to brownish, cubitus anterior vein brownish, ambient vein slightly brownish; vannus slightly smoky and its margin with anal cell brown and orange.
</paragraph>
<paragraph id="97364414FFB0FF92FF72FBC18F82FA34" blockId="9.[151,1437,150,2013]" pageId="9" pageNumber="544">
<emphasis id="A5FD9806FFB0FF92FF72FBC18F96FBD5" box="[199,252,1123,1148]" italics="true" pageId="9" pageNumber="544">Legs</emphasis>
: yellowish-brownish and black. Fore legs: posterior side of coxa yellowish and anterior side black with long yellowish hairs; trochanter blackish with margin, fissures and posterior side yellowish to brownish; femora yellowish with black longitudinal bands, black edge with three spines blackish, the basal spine sharp and oblique, isolated from the two others, the second spine strong and sub-perpendicular, the apical spine very short and triangular; tibia blackish with yellowish spot on lateral sides; tarsus blackish to brownish. Mid legs: coxa yellowish except the latero-posterior side black; trochanter blackish, its base, ventral and lateral margin with spots yellowish; femora yellowish with four black longitudinal stripes of variable length, the thinner more or less interrupted; tibia black; tarsus blackish. Hind legs: similar to mid legs but lighter in colour and femora with only three black longitudinal stripes.
</paragraph>
<paragraph id="97364414FFB0FF92FF72FA058C73F8AC" blockId="9.[151,1437,150,2013]" pageId="9" pageNumber="544">
<emphasis id="A5FD9806FFB0FF92FF72FA058E5BFA68" box="[199,305,1447,1473]" italics="true" pageId="9" pageNumber="544">Abdomen</emphasis>
: tergites black, posterior margins yellowish; auditory capsules sub-circular, black, slightly convex. Tymbals bearing a basal plate which is as long as one third of the tymbal area and showing a series of eight short ribs alternating with nine long ribs, the sclerotized ribs running anterior to the tymbal plate. Sternites black with a posterior margin brownish; sternite VIII with a large blackish spot reaching the apical part, this large blackish spot bordered of yellow. Genitalia (
<figureCitation id="0FB25891FFB0FF92FE4EF9958DFEF9F8" box="[507,660,1591,1617]" captionStart="FIGURE 1" captionStartId="2.[151,250,1242,1266]" captionTargetBox="[151,1436,179,1216]" captionTargetId="figure-22@2.[151,1436,179,1216]" captionTargetPageId="2" captionText="FIGURE 1. Tibicina quadrisignata pilleti Puissant ssp. n., Holotype male, Morocco. A, habitus, dorsal view. B, lateral view of pygofer, uncus curved with apex highly angular (↑). C, lateral view of dissected aedeagus, vesica of median lobe curved and inflated with apex short. D, distal end of aedeagus deeply curved and trilobed, the two lateral lobes small and edged, strongly elongated and rounded at their apex with numerous conjunctival claws (↑)." figureDoi="http://doi.org/10.5281/zenodo.8255297" httpUri="https://zenodo.org/record/8255297/files/figure.png" pageId="9" pageNumber="544">Figs. 1B,C,D</figureCitation>
): pygopher curved without dorsal expansion but slightly V-shaped; uncus curved with an apex highly angular (
<figureCitation id="0FB25891FFB0FF92FD32F9F98D8AF9DC" box="[647,736,1627,1653]" captionStart="FIGURE 1" captionStartId="2.[151,250,1242,1266]" captionTargetBox="[151,1436,179,1216]" captionTargetId="figure-22@2.[151,1436,179,1216]" captionTargetPageId="2" captionText="FIGURE 1. Tibicina quadrisignata pilleti Puissant ssp. n., Holotype male, Morocco. A, habitus, dorsal view. B, lateral view of pygofer, uncus curved with apex highly angular (↑). C, lateral view of dissected aedeagus, vesica of median lobe curved and inflated with apex short. D, distal end of aedeagus deeply curved and trilobed, the two lateral lobes small and edged, strongly elongated and rounded at their apex with numerous conjunctival claws (↑)." figureDoi="http://doi.org/10.5281/zenodo.8255297" httpUri="https://zenodo.org/record/8255297/files/figure.png" pageId="9" pageNumber="544">Fig. 1B</figureCitation>
: ↑). Claspers absent. Ventrobasal pocket of aedeagus present. Basal plate of aedeagus undivided basally and attachment to the theca by sinuation completely chitinous with no mobility. Aedeagus elongated, deeply curved and trilobed at the distal end; the two lateral lobes small and edged, strongly elongated and rounded at their apex with numerous conjunctival claws (
<figureCitation id="0FB25891FFB0FF92FB97F9658B16F948" box="[1058,1148,1735,1761]" captionStart="FIGURE 1" captionStartId="2.[151,250,1242,1266]" captionTargetBox="[151,1436,179,1216]" captionTargetId="figure-22@2.[151,1436,179,1216]" captionTargetPageId="2" captionText="FIGURE 1. Tibicina quadrisignata pilleti Puissant ssp. n., Holotype male, Morocco. A, habitus, dorsal view. B, lateral view of pygofer, uncus curved with apex highly angular (↑). C, lateral view of dissected aedeagus, vesica of median lobe curved and inflated with apex short. D, distal end of aedeagus deeply curved and trilobed, the two lateral lobes small and edged, strongly elongated and rounded at their apex with numerous conjunctival claws (↑)." figureDoi="http://doi.org/10.5281/zenodo.8255297" httpUri="https://zenodo.org/record/8255297/files/figure.png" pageId="9" pageNumber="544">Fig. 1D</figureCitation>
: ↑); median lobe carrying the gonopore, vesica curved and inflated with apex short.
</paragraph>
<paragraph id="97364414FFB0FF91FF72F8AD8DA3FD64" blockId="9.[151,1437,150,2013]" lastBlockId="10.[151,1437,150,861]" lastPageId="10" lastPageNumber="545" pageId="9" pageNumber="544">
<emphasis id="A5FD9806FFB0FF92FF72F8AD8E01F880" bold="true" box="[199,363,1807,1833]" pageId="9" pageNumber="544">Measurement</emphasis>
. Seven males and 
<specimenCount id="818F8F9DFFB0FF92FD8BF8AD8DBEF880" box="[574,724,1807,1833]" pageId="9" pageNumber="544" type="female">three females</specimenCount>
are available. Length of body, from tip of head to end of wings in resting position: males 38.70 ± 
<quantity id="5071E9F1FFB0FF92FD9FF8918DE4F8E4" box="[554,654,1843,1869]" metricMagnitude="-3" metricUnit="m" metricValue="1.26" pageId="9" pageNumber="544" unit="mm" value="1.26">1.26 mm</quantity>
(37.00–40.50), females 40.00 ± 
<quantity id="5071E9F1FFB0FF92FBB8F8918B1BF8E4" box="[1037,1137,1843,1869]" metricMagnitude="-3" metricUnit="m" metricValue="3.16" pageId="9" pageNumber="544" unit="mm" value="3.16">3.16 mm</quantity>
(36.40–42.30). Length of body, from tip of head to end of abdomen: males 26.51 ± 
<quantity id="5071E9F1FFB0FF92FC98F8F48CFBF8D8" box="[813,913,1878,1905]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" pageId="9" pageNumber="544" unit="mm" value="1.35">1.35 mm</quantity>
(25.00–39.00), females 
<quantity id="5071E9F1FFB0FF92FB12F8F58A73F8D8" box="[1191,1305,1879,1905]" metricMagnitude="-2" metricUnit="m" metricValue="2.8129999999999997" pageId="9" pageNumber="544" unit="mm" value="28.13">28.13 mm</quantity>
± 
<quantity id="5071E9F1FFB0FF92FA8CF8F58AF7F8D8" box="[1337,1437,1879,1905]" metricMagnitude="-3" metricUnit="m" metricValue="2.04" pageId="9" pageNumber="544" unit="mm" value="2.04">2.04 mm</quantity>
(25.90–29.90). Length of head: male 2.67 ± 
<quantity id="5071E9F1FFB0FF92FD3FF8D98D84F83C" box="[650,750,1915,1941]" metricMagnitude="-4" metricUnit="m" metricValue="2.4" pageId="9" pageNumber="544" unit="mm" value="0.24">0.24 mm</quantity>
(2.40–2.90), females 2.44 ± 
<quantity id="5071E9F1FFB0FF92FB86F8D98BFDF83C" box="[1075,1175,1915,1941]" metricMagnitude="-4" metricUnit="m" metricValue="1.7" pageId="9" pageNumber="544" unit="mm" value="0.17">0.17 mm</quantity>
(2.30–2.63). Length of pronotum: males 4.06 ± 
<quantity id="5071E9F1FFB0FF92FE19F83D8D7AF810" box="[428,528,1951,1977]" metricMagnitude="-4" metricUnit="m" metricValue="2.8" pageId="9" pageNumber="544" unit="mm" value="0.28">0.28 mm</quantity>
(3.70–4.50), females 3.99 ± 
<quantity id="5071E9F1FFB0FF92FCE2F83D8CD1F810" box="[855,955,1951,1977]" metricMagnitude="-4" metricUnit="m" metricValue="6.2" pageId="9" pageNumber="544" unit="mm" value="0.62">0.62 mm</quantity>
(3.30–4.48). Length of mesonotum: males 6.95 ± 
<quantity id="5071E9F1FFB0FF92FF51F8618E22F874" box="[228,328,1987,2013]" metricMagnitude="-4" metricUnit="m" metricValue="3.4" pageId="9" pageNumber="544" unit="mm" value="0.34">0.34 mm</quantity>
(6.50–7.30), females 6.98 ± 
<quantity id="5071E9F1FFB0FF92FD3DF8618D86F874" box="[648,748,1987,2013]" metricMagnitude="-4" metricUnit="m" metricValue="9.399999999999999" pageId="9" pageNumber="544" unit="mm" value="0.94">0.94 mm</quantity>
(5.90–7.60). Length of head and thorax together: males 14.15 ± 
<quantity id="5071E9F1FFB3FF91FF1AFF358E79FF18" box="[175,275,151,177]" metricMagnitude="-4" metricUnit="m" metricValue="7.7" pageId="10" pageNumber="545" unit="mm" value="0.77">0.77 mm</quantity>
(13.20–15.40), females 13.79 ± 
<quantity id="5071E9F1FFB3FF91FD3DFF358D86FF18" box="[648,748,151,177]" metricMagnitude="-3" metricUnit="m" metricValue="1.64" pageId="10" pageNumber="545" unit="mm" value="1.64">1.64 mm</quantity>
(11.90–14.87). Length of abdomen: males 12.36 ± 
<quantity id="5071E9F1FFB3FF91FA8CFF358AF7FF18" box="[1337,1437,151,177]" metricMagnitude="-3" metricUnit="m" metricValue="1.18" pageId="10" pageNumber="545" unit="mm" value="1.18">1.18 mm</quantity>
(11.40–14.40), females 14.34 ± 
<quantity id="5071E9F1FFB3FF91FE4AFF198D09FF7C" box="[511,611,187,213]" metricMagnitude="-4" metricUnit="m" metricValue="8.4" pageId="10" pageNumber="545" unit="mm" value="0.84">0.84 mm</quantity>
(13.73–15.30). Ratio length of abdomen/head and thorax together: males 0.88 ± 
<quantity id="5071E9F1FFB3FF91FF58FF7D8E2DFF50" box="[237,327,223,249]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="545" unit="mm" value="0.1">0.1 mm</quantity>
(0.74–1.03), females 1.11± 
<quantity id="5071E9F1FFB3FF91FD26FF7D8D91FF50" box="[659,763,223,249]" metricMagnitude="-5" metricUnit="m" metricValue="8.0" pageId="10" pageNumber="545" unit="mm" value="0.08">0.08 mm</quantity>
(1.05–1.20). Width of head including eyes: males 8.67 ± 
<quantity id="5071E9F1FFB3FF91FF22FEA18F91FEB4" box="[151,251,259,285]" metricMagnitude="-4" metricUnit="m" metricValue="3.7" pageId="10" pageNumber="545" unit="mm" value="0.37">0.37 mm</quantity>
(8.00–9.12), females 8.87 ± 
<quantity id="5071E9F1FFB3FF91FDE6FEA18DDDFEB4" box="[595,695,259,285]" metricMagnitude="-4" metricUnit="m" metricValue="6.8" pageId="10" pageNumber="545" unit="mm" value="0.68">0.68 mm</quantity>
(8.10–9.40). Width of pronotum, supra-humeral plate included: males 10.63 ± 
<quantity id="5071E9F1FFB3FF91FE8EFE858EF5FEE8" box="[315,415,295,321]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" pageId="10" pageNumber="545" unit="mm" value="0.48">0.48 mm</quantity>
(10.00–11.40), females 10.90 ± 
<quantity id="5071E9F1FFB3FF91FCBFFE858C04FEE8" box="[778,878,295,321]" metricMagnitude="-3" metricUnit="m" metricValue="1.49" pageId="10" pageNumber="545" unit="mm" value="1.49">1.49 mm</quantity>
(9.20–12.00). Width of mesonotum: males 8.91 ± 
<quantity id="5071E9F1FFB3FF91FF22FEE98F91FECC" box="[151,251,331,357]" metricMagnitude="-4" metricUnit="m" metricValue="4.9" pageId="10" pageNumber="545" unit="mm" value="0.49">0.49 mm</quantity>
(8.00–9.40), females 9.29 ± 
<quantity id="5071E9F1FFB3FF91FD80FEE98DF3FECC" box="[565,665,331,357]" metricMagnitude="-4" metricUnit="m" metricValue="8.1" pageId="10" pageNumber="545" unit="mm" value="0.81">0.81 mm</quantity>
(8.40–9.98). Ratio width of head including eyes/width of mesonotum: males 0.98 ± 
<quantity id="5071E9F1FFB3FF91FE8FFECD8EF4FE20" box="[314,414,367,393]" metricMagnitude="-5" metricUnit="m" metricValue="2.0" pageId="10" pageNumber="545" unit="mm" value="0.02">0.02 mm</quantity>
(0.95–1.00), females 0.95 ± 
<quantity id="5071E9F1FFB3FF91FD40FECD8C33FE20" box="[757,857,367,393]" metricMagnitude="-5" metricUnit="m" metricValue="4.0" pageId="10" pageNumber="545" unit="mm" value="0.04">0.04 mm</quantity>
(0.91–0.99). Length of fore wing: males 31.17 ± 
<quantity id="5071E9F1FFB3FF91FF22FE318F91FE04" box="[151,251,403,429]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="10" pageNumber="545" unit="mm" value="1.4">1.40 mm</quantity>
(29.60–33.00), females 33.30 ± 
<quantity id="5071E9F1FFB3FF91FDCFFE318DB4FE04" box="[634,734,403,429]" metricMagnitude="-3" metricUnit="m" metricValue="2.62" pageId="10" pageNumber="545" unit="mm" value="2.62">2.62 mm</quantity>
(30.30–35.10). Width of fore wing: males 11.54 ± 
<quantity id="5071E9F1FFB3FF91FA8CFE318AF7FE04" box="[1337,1437,403,429]" metricMagnitude="-4" metricUnit="m" metricValue="6.6" pageId="10" pageNumber="545" unit="mm" value="0.66">0.66 mm</quantity>
(10.50–12.17), females 12.41 ± 
<quantity id="5071E9F1FFB3FF91FE4EFE158D35FE78" box="[507,607,439,465]" metricMagnitude="-4" metricUnit="m" metricValue="7.9" pageId="10" pageNumber="545" unit="mm" value="0.79">0.79 mm</quantity>
(11.80–13.30). Wingspan: males 68.93 ± 
<quantity id="5071E9F1FFB3FF91FB85FE158BFEFE78" box="[1072,1172,439,465]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="10" pageNumber="545" unit="mm" value="1.4">1.40 mm</quantity>
(67.00–71.00), females 75.10 ± 
<quantity id="5071E9F1FFB3FF91FF41FE798E32FE5C" box="[244,344,475,501]" metricMagnitude="-3" metricUnit="m" metricValue="7.64" pageId="10" pageNumber="545" unit="mm" value="7.64">7.64 mm</quantity>
(66.30–80.00). Ratio length of fore wing/width of fore wing: males 2.70 ± 
<quantity id="5071E9F1FFB3FF91FB1DFE798A66FE5C" box="[1192,1292,475,501]" metricMagnitude="-5" metricUnit="m" metricValue="9.0" pageId="10" pageNumber="545" unit="mm" value="0.09">0.09 mm</quantity>
(2.55–2.82), females 2.69 ± 
<quantity id="5071E9F1FFB3FF91FEF3FE5D8EC0FDB0" box="[326,426,511,537]" metricMagnitude="-4" metricUnit="m" metricValue="1.7999999999999998" pageId="10" pageNumber="545" unit="mm" value="0.18">0.18 mm</quantity>
(2.57–2.90). Ratio length of fore wing/width of pronotum, supra-humeral plate included: males 2.93 ± 
<quantity id="5071E9F1FFB3FF91FE9AFD818EF8FD94" box="[303,402,547,573]" metricMagnitude="-4" metricUnit="m" metricValue="1.1" pageId="10" pageNumber="545" unit="mm" value="0.11">0.11 mm</quantity>
(2.72–3.08), females 3.07 ± 
<quantity id="5071E9F1FFB3FF91FD63FD818C50FD94" box="[726,826,547,573]" metricMagnitude="-4" metricUnit="m" metricValue="2.1" pageId="10" pageNumber="545" unit="mm" value="0.21">0.21 mm</quantity>
(2.87–3.29). Distance between ocellus and compound eye of same side: males 1.27 ± 
<quantity id="5071E9F1FFB3FF91FDA6FDE58D1DFDC8" box="[531,631,583,609]" metricMagnitude="-5" metricUnit="m" metricValue="9.0" pageId="10" pageNumber="545" unit="mm" value="0.09">0.09 mm</quantity>
(1.10–1.35), females 1.41 ± 
<quantity id="5071E9F1FFB3FF91FC65FDE58B58FDC8" box="[976,1074,583,609]" metricMagnitude="-4" metricUnit="m" metricValue="1.1" pageId="10" pageNumber="545" unit="mm" value="0.11">0.11 mm</quantity>
(1.30–1.52). Distance between posterior lateral ocelli: males 1.35 ± 
<quantity id="5071E9F1FFB3FF91FDF6FDC98DCDFD2C" box="[579,679,619,645]" metricMagnitude="-5" metricUnit="m" metricValue="9.0" pageId="10" pageNumber="545" unit="mm" value="0.09">0.09 mm</quantity>
(1.20–1.44), females 1.46 ± 
<quantity id="5071E9F1FFB3FF91FC4CFDC98B37FD2C" box="[1017,1117,619,645]" metricMagnitude="-4" metricUnit="m" metricValue="2.1" pageId="10" pageNumber="545" unit="mm" value="0.21">0.21 mm</quantity>
(1.30–1.70). Ratio distance between ocellus and compound eye of same side/distance between posterior lateral ocelli: males 0.94 ± 
<quantity id="5071E9F1FFB3FF91FA8CFD2D8AF7FD00" box="[1337,1437,655,681]" metricMagnitude="-5" metricUnit="m" metricValue="6.0" pageId="10" pageNumber="545" unit="mm" value="0.06">0.06 mm</quantity>
(0.84–1.04), females 0.97 ± 
<quantity id="5071E9F1FFB3FF91FE61FD118D52FD64" box="[468,568,691,717]" metricMagnitude="-5" metricUnit="m" metricValue="7" pageId="10" pageNumber="545" unit="mm" value="0.07">0.07 mm</quantity>
(0.89–1.02).
</paragraph>
<paragraph id="97364414FFB3FF91FF72FD748B7AFCF4" blockId="10.[151,1437,150,861]" pageId="10" pageNumber="545">
<emphasis id="A5FD9806FFB3FF91FF72FD748E4DFD59" bold="true" box="[199,295,726,752]" pageId="10" pageNumber="545">Exuviae</emphasis>
(
<figureCitation id="0FB25891FFB3FF91FE89FD758EBFFD58" box="[316,469,727,753]" captionStart="FIGURE 4" captionStartId="5.[151,250,1908,1932]" captionTargetBox="[151,1436,849,1885]" captionTargetId="figure-327@5.[151,1436,849,1885]" captionTargetPageId="5" captionText="FIGURE 4. T. q. pilleti Puissant ssp. n., right lateral view of exuviae. A, habitus. B, forelimb robust with accessory tooth of femur (↑) long and flattened. C, head with bulging postclypeus (↑)." figureDoi="http://doi.org/10.5281/zenodo.8255305" httpUri="https://zenodo.org/record/8255305/files/figure.png" pageId="10" pageNumber="545">Figs. 4A,B,C</figureCitation>
). Very close of 
<emphasis id="A5FD9806FFB3FF91FD23FD7A8C00FD58" box="[662,874,727,753]" italics="true" pageId="10" pageNumber="545">
<collectionCode id="F198DCD1FFB3FF91FD23FD7A8DCDFD58" box="[662,679,728,753]" name="Tavera, Department of Geology and Geophysics" pageId="10" pageNumber="545">T</collectionCode>
. q. 
<taxonomicName id="50893F97FFB3FF91FD78FD758C00FD58" baseAuthorityName="Hagen" baseAuthorityYear="1855" box="[717,874,727,753]" class="Insecta" family="Cicadidae" genus="Tibicina" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="545" phylum="Arthropoda" rank="species" species="quadrisignata">quadrisignata</taxonomicName>
</emphasis>
exuviae, with sturdy shape. Head with bulging postclypeus (
<figureCitation id="0FB25891FFB3FF91FE9FFD598EE9FCBC" box="[298,387,763,789]" captionStart="FIGURE 4" captionStartId="5.[151,250,1908,1932]" captionTargetBox="[151,1436,849,1885]" captionTargetId="figure-327@5.[151,1436,849,1885]" captionTargetPageId="5" captionText="FIGURE 4. T. q. pilleti Puissant ssp. n., right lateral view of exuviae. A, habitus. B, forelimb robust with accessory tooth of femur (↑) long and flattened. C, head with bulging postclypeus (↑)." figureDoi="http://doi.org/10.5281/zenodo.8255305" httpUri="https://zenodo.org/record/8255305/files/figure.png" pageId="10" pageNumber="545">Fig. 4C</figureCitation>
: ↑), protruding as in some exuviae of 
<emphasis id="A5FD9806FFB3FF91FC93FD598C90FCBC" box="[806,1018,763,789]" italics="true" pageId="10" pageNumber="545">
<collectionCode id="F198DCD1FFB3FF91FC93FD598C5DFCBD" box="[806,823,763,788]" name="Tavera, Department of Geology and Geophysics" pageId="10" pageNumber="545">T</collectionCode>
. q. 
<taxonomicName id="50893F97FFB3FF91FCE8FD598C90FCBC" baseAuthorityName="Hagen" baseAuthorityYear="1855" box="[861,1018,763,789]" class="Insecta" family="Cicadidae" genus="Tibicina" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="545" phylum="Arthropoda" rank="species" species="quadrisignata">quadrisignata</taxonomicName>
</emphasis>
. Forelimb robust with femoral comb, intermediate tooth, posterior tooth well developed. Its accessory tooth of femur (
<figureCitation id="0FB25891FFB3FF91FBADFCBD8B1AFC90" box="[1048,1136,799,825]" captionStart="FIGURE 4" captionStartId="5.[151,250,1908,1932]" captionTargetBox="[151,1436,849,1885]" captionTargetId="figure-327@5.[151,1436,849,1885]" captionTargetPageId="5" captionText="FIGURE 4. T. q. pilleti Puissant ssp. n., right lateral view of exuviae. A, habitus. B, forelimb robust with accessory tooth of femur (↑) long and flattened. C, head with bulging postclypeus (↑)." figureDoi="http://doi.org/10.5281/zenodo.8255305" httpUri="https://zenodo.org/record/8255305/files/figure.png" pageId="10" pageNumber="545">Fig. 4B</figureCitation>
: ↑) long and flattened as in some exuviae of 
<emphasis id="A5FD9806FFB3FF91FEE3FCE18D40FCF4" box="[342,554,835,861]" italics="true" pageId="10" pageNumber="545">
<collectionCode id="F198DCD1FFB3FF91FEE3FCE18E0DFCF5" box="[342,359,835,860]" name="Tavera, Department of Geology and Geophysics" pageId="10" pageNumber="545">T</collectionCode>
. q. 
<taxonomicName id="50893F97FFB3FF91FE38FCE18D40FCF4" baseAuthorityName="Hagen" baseAuthorityYear="1855" box="[397,554,835,861]" class="Insecta" family="Cicadidae" genus="Tibicina" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="545" phylum="Arthropoda" rank="species" species="quadrisignata">quadrisignata</taxonomicName>
</emphasis>
. Apical tooth of tibiae long and acuminate.
</paragraph>
<caption id="C3F6149CFFB3FF91FF22F9F88BE0F93F" ID-DOI="http://doi.org/10.5281/zenodo.8255315" ID-Zenodo-Dep="8255315" httpUri="https://zenodo.org/record/8255315/files/figure.png" pageId="10" pageNumber="545" startId="10.[151,250,1626,1650]" targetBox="[160,1430,886,1603]" targetPageId="10" targetType="figure">
<paragraph id="97364414FFB3FF91FF22F9F88BE0F93F" blockId="10.[151,1436,1626,1687]" pageId="10" pageNumber="545">
<emphasis id="A5FD9806FFB3FF91FF22F9F88E79F9DB" bold="true" box="[151,275,1626,1650]" pageId="10" pageNumber="545">FIGURE 8.</emphasis>
Calling song recording of 
<emphasis id="A5FD9806FFB3FF91FD95F9F98DE7F9DB" box="[544,653,1626,1650]" italics="true" pageId="10" pageNumber="545">T. q. pilleti</emphasis>
<emphasis id="A5FD9806FFB3FF91FD26F9F88C5CF9DA" bold="true" box="[659,822,1626,1651]" pageId="10" pageNumber="545">
Puissant 
<taxonomicNameLabel id="BECE257DFFB3FF91FD41F9F98C5CF9DA" box="[756,822,1627,1651]" pageId="10" pageNumber="545" rank="subSpecies">ssp. n.</taxonomicNameLabel>
</emphasis>
with a digitization at 96 kHz sampling frequency. Frequency amplitude envelope peaking at 28 kHz, encapsulating the second upper harmonic at around 27 kHz.
</paragraph>
</caption>
<paragraph id="97364414FFB3FF96FF22F9668EC1F9AE" blockId="10.[151,1437,1732,2011]" lastBlockId="13.[151,1437,1445,2011]" lastPageId="13" lastPageNumber="548" pageId="10" pageNumber="545">
<emphasis id="A5FD9806FFB3FF91FF22F9668E30F977" bold="true" box="[151,346,1732,1758]" pageId="10" pageNumber="545">Sound behavior</emphasis>
(
<tableCitation id="DA0B71AFFFB3FF91FEDBF9678EAEF976" box="[366,452,1733,1759]" captionStart="TABLE 1" captionStartId="11.[151,239,619,645]" captionTargetPageId="11" captionText="TABLE 1. Mean ± standard deviation (minimum–maximum; sample size) for the temporal and frequency parameters of T. quadrisignata pilleti and T. maldesi calling signals, with: CD, call duration of a sequence (s); GPD, group of pulses duration (ms); GPP, group of pulses period (ms); NGP, number of group of pulses per second; NP, number of pulses per group. MF, frequencies with maximum amplitude; F1, peak of the first frequency band; F2, peak of the second frequency; F3, peak of the third frequency band; *carrier frequency band. See “Materials and methods” for more explanations." httpUri="http://table.plazi.org/id/C3F6149CFFB2FF90FF22FDC98C13FD58" pageId="10" pageNumber="545" tableUuid="C3F6149CFFB2FF90FF22FDC98C13FD58">Table 1</tableCitation>
; 
<figureCitation id="0FB25891FFB3FF91FE62F9678DB8F976" box="[471,722,1733,1759]" captionStart-0="FIGURE 8" captionStart-1="FIGURE 9" captionStartId-0="10.[151,250,1626,1650]" captionStartId-1="13.[151,250,150,174]" captionTargetBox-0="[160,1430,886,1603]" captionTargetId-0="figure-354@10.[151,1436,885,1603]" captionTargetId-1="figure-193@13.[151,1436,523,1243]" captionTargetPageId-0="10" captionTargetPageId-1="13" captionText-0="FIGURE 8. Calling song recording of T. q. pilleti Puissant ssp. n. with a digitization at 96 kHz sampling frequency. Frequency amplitude envelope peaking at 28 kHz, encapsulating the second upper harmonic at around 27 kHz." captionText-1="FIGURE 9. Calling song recording of T. q. pilleti Puissant ssp. n. with a digitization at 44.1 kHz sampling frequency.A, spectra (frequency vs. absolute amplitude). B, spectrogram and C, oscillogram (from top to bottom) of the beginning of the sequence. D, oscillogram of a part of C showing groups of pulses (slow amplitude modulation). E, detailed spectrogram and F, oscillogram of a part of D showing a train of groups of pulses in detail. F1, F2*, F3: three frequency peaks with F2* as carrier frequency; FFT: fast Fourier transform; GP: group of pulses (= syllable); MF: frequencies of maximum amplitude; N: number of males (N = 4) analyzed; n: number of measurements for sampling (n = 337); NAV: number of amplitude variations; light grey spectrum (at bottom): minimum amplitude spectrum plotted from the minimum amplitude value for each frequency; black spectrum (at middle): average linear amplitude spectrum plotted from the average amplitude value recorded for each frequency; grey spectrum (at top): maximum amplitude spectrum plotted from the maximum amplitude value recorded for each frequency." figureDoi-0="http://doi.org/10.5281/zenodo.8255315" figureDoi-1="http://doi.org/10.5281/zenodo.8257063" httpUri-0="https://zenodo.org/record/8255315/files/figure.png" httpUri-1="https://zenodo.org/record/8257063/files/figure.png" pageId="10" pageNumber="545">Figs. 8–9A,B,C,D,E,F</figureCitation>
– 
<figureCitation id="0FB25891FFB3FF91FD57F9678CC9F976" box="[738,931,1733,1759]" captionStart-0="FIGURE 10" captionStart-1="FIGURE 11" captionStartId-0="13.[151,250,1267,1291]" captionStartId-1="14.[151,250,1729,1753]" captionTargetBox-0="[160,1432,524,1243]" captionTargetBox-1="[164,1431,181,1705]" captionTargetId-0="figure-193@13.[151,1436,523,1243]" captionTargetId-1="figure-22@14.[151,1436,181,1705]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionText-0="FIGURE 10. Spectrogram and oscillogram (from top to bottom) of the rivalry song of T. q. pilleti Puissant ssp. n. One male producing an irregular train of echemes in the foreground (M2) is interrupted by another male producing a fast trains of rivalry song pulse groups in the background (M1)." captionText-1="FIGURE 11. Courtship song of T. q. pilleti Puissant ssp. n. A, spectrogram and B, oscillogram (from top to bottom) of a succession of phrases composed of two echemes. C, oscillogram of a part of B showing two phrases and their component echeme, a one long echeme (E1) and a one short terminal echeme (E2). D, detailed spectrogram and E, oscillogram of a part of C showing a phrase with, for each echeme, a low amplitude part duration (LPD) and a shorter high amplitude part duration (HPD)." figureDoi-0="http://doi.org/10.5281/zenodo.8255317" figureDoi-1="http://doi.org/10.5281/zenodo.8255321" httpUri-0="https://zenodo.org/record/8255317/files/figure.png" httpUri-1="https://zenodo.org/record/8255321/files/figure.png" pageId="10" pageNumber="545">10–11A,B,C,D,E</figureCitation>
). Three males were recorded on the 
<emphasis id="A5FD9806FFB3FF91FAD4F9678F8CF8AB" italics="true" pageId="10" pageNumber="545">locus typicus</emphasis>
by SP, then captured by SP with Alexandre François. Another male was recorded by SP without being caught. The males produce their calling song mainly from the low bushes to the tops of the trees. Densities of calling males can be very high; males usually call 
<quantity id="5071E9F1FFB3FF91FD66F8938C58F8E2" box="[723,818,1841,1867]" metricMagnitude="0" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="10" pageNumber="545" unit="m" value="2.0" valueMax="3.0" valueMin="1.0">1 to 3 m</quantity>
apart. When males emit their calling song, they raised their abdomen and lift up their wings from their body without decoupling them from the mesonotum (
<figureCitation id="0FB25891FFB3FF91FB4AF8F78A29F8C6" box="[1279,1347,1876,1903]" captionStart="FIGURE 5" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[213,1374,181,1919]" captionTargetId="figure-22@6.[213,1374,181,1919]" captionTargetPageId="6" captionText="FIGURE 5. Male of T. q. pilleti Puissant ssp. n. in calling posture." figureDoi="http://doi.org/10.5281/zenodo.8255307" httpUri="https://zenodo.org/record/8255307/files/figure.png" pageId="10" pageNumber="545">Fig. 5</figureCitation>
). Males produce relatively short, sustained calling songs of about one minute: CD = 67.4 ± 23.5 (46.3–98.0; 4), see 
<tableCitation id="DA0B71AFFFB3FF91FAEBF8DB8FCCF81E" captionStart="TABLE 1" captionStartId="11.[151,239,619,645]" captionTargetPageId="11" captionText="TABLE 1. Mean ± standard deviation (minimum–maximum; sample size) for the temporal and frequency parameters of T. quadrisignata pilleti and T. maldesi calling signals, with: CD, call duration of a sequence (s); GPD, group of pulses duration (ms); GPP, group of pulses period (ms); NGP, number of group of pulses per second; NP, number of pulses per group. MF, frequencies with maximum amplitude; F1, peak of the first frequency band; F2, peak of the second frequency; F3, peak of the third frequency band; *carrier frequency band. See “Materials and methods” for more explanations." httpUri="http://table.plazi.org/id/C3F6149CFFB2FF90FF22FDC98C13FD58" pageId="10" pageNumber="545" tableUuid="C3F6149CFFB2FF90FF22FDC98C13FD58">Table 1</tableCitation>
. Sometimes, a small number of amplitude modulated variations is emitted at the beginning of the call (
<figureCitation id="0FB25891FFB3FF91FA8AF83F8AF2F81E" box="[1343,1432,1949,1975]" captionStart="FIGURE 9" captionStartId="13.[151,250,150,174]" captionTargetId="figure-193@13.[151,1436,523,1243]" captionTargetPageId="13" captionText="FIGURE 9. Calling song recording of T. q. pilleti Puissant ssp. n. with a digitization at 44.1 kHz sampling frequency.A, spectra (frequency vs. absolute amplitude). B, spectrogram and C, oscillogram (from top to bottom) of the beginning of the sequence. D, oscillogram of a part of C showing groups of pulses (slow amplitude modulation). E, detailed spectrogram and F, oscillogram of a part of D showing a train of groups of pulses in detail. F1, F2*, F3: three frequency peaks with F2* as carrier frequency; FFT: fast Fourier transform; GP: group of pulses (= syllable); MF: frequencies of maximum amplitude; N: number of males (N = 4) analyzed; n: number of measurements for sampling (n = 337); NAV: number of amplitude variations; light grey spectrum (at bottom): minimum amplitude spectrum plotted from the minimum amplitude value for each frequency; black spectrum (at middle): average linear amplitude spectrum plotted from the average amplitude value recorded for each frequency; grey spectrum (at top): maximum amplitude spectrum plotted from the maximum amplitude value recorded for each frequency." figureDoi="http://doi.org/10.5281/zenodo.8257063" httpUri="https://zenodo.org/record/8257063/files/figure.png" pageId="10" pageNumber="545">Fig. 9C</figureCitation>
: 
<collectionCode id="F198DCD1FFB3FF91FF22F8638FBCF872" box="[151,214,1985,2011]" pageId="10" pageNumber="545">NAV</collectionCode>
). Group of pulses duration (
<collectionCode id="F198DCD1FFB3FF91FDBEF8638D2CF872" box="[523,582,1985,2011]" pageId="10" pageNumber="545">GPD</collectionCode>
) is notably constant: 13.02 ± 0.2 ms (12.2–13.6; 346). Group of pulse period (
<collectionCode id="F198DCD1FFB4FF96FF2AFA078FBCFA16" box="[159,214,1445,1471]" pageId="13" pageNumber="548">GPP</collectionCode>
) is 14.1 ± 0.2 ms (13.6–14.5; 86) and groups of pulses are produced a rate (
<collectionCode id="F198DCD1FFB4FF96FB89FA078B1DFA16" box="[1084,1143,1445,1471]" pageId="13" pageNumber="548">NGP</collectionCode>
) of 71.5 ± 
<geoCoordinate id="F2BD22D3FFB4FF96FB4CFA078A5FFA1A" box="[1273,1333,1445,1472]" degrees="0.9" direction="south" orientation="latitude" pageId="13" pageNumber="548" precision="5555" value="-0.9">
0.9 
<superScript id="60FCE95CFFB4FF96FA91FA048A5FFA1A" attach="left" box="[1316,1333,1445,1472]" fontSize="6" pageId="13" pageNumber="548">s-</superScript>
</geoCoordinate>
1 (70–73; 329). Pulses-groups (
<tableCitation id="DA0B71AFFFB4FF96FE36FA6B8EB0FA4A" box="[387,474,1481,1507]" captionStart="TABLE 1" captionStartId="11.[151,239,619,645]" captionTargetPageId="11" captionText="TABLE 1. Mean ± standard deviation (minimum–maximum; sample size) for the temporal and frequency parameters of T. quadrisignata pilleti and T. maldesi calling signals, with: CD, call duration of a sequence (s); GPD, group of pulses duration (ms); GPP, group of pulses period (ms); NGP, number of group of pulses per second; NP, number of pulses per group. MF, frequencies with maximum amplitude; F1, peak of the first frequency band; F2, peak of the second frequency; F3, peak of the third frequency band; *carrier frequency band. See “Materials and methods” for more explanations." httpUri="http://table.plazi.org/id/C3F6149CFFB2FF90FF22FDC98C13FD58" pageId="13" pageNumber="548" tableUuid="C3F6149CFFB2FF90FF22FDC98C13FD58">Table 1</tableCitation>
: NP per syllable) are made up of 8-11 pulses, typically 8 to 10 pulses per syllables, 
<emphasis id="A5FD9806FFB4FF96FACFFA688AF7FA4A" box="[1402,1437,1482,1507]" italics="true" pageId="13" pageNumber="548">i.e.</emphasis>
4 to 5 pulses per tymbal.
</paragraph>
<caption id="C3F6149CFFB2FF90FF22FDC98C13FD58" ID-Table-UUID="C3F6149CFFB2FF90FF22FDC98C13FD58" httpUri="http://table.plazi.org/id/C3F6149CFFB2FF90FF22FDC98C13FD58" pageId="11" pageNumber="546" startId="11.[151,239,619,645]" targetBox="[162,2017,768,1569]" targetIsTable="true" targetPageId="11" targetType="table">
<paragraph id="97364414FFB2FF90FF22FDC98C13FD58" blockId="11.[151,2037,619,753]" pageId="11" pageNumber="546">
<emphasis id="A5FD9806FFB2FF90FF22FDC98E61FD2C" bold="true" box="[151,267,619,645]" pageId="11" pageNumber="546">TABLE 1.</emphasis>
Mean ± standard deviation (minimum–maximum; sample size) for the temporal and frequency parameters of 
<taxonomicName id="50893F97FFB2FF90FA06FDC989C3FD2C" box="[1459,1705,619,645]" class="Insecta" family="Cicadidae" genus="Tibicina" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="546" phylum="Arthropoda" rank="subSpecies" species="quadrisignata" subSpecies="pilleti">
<emphasis id="A5FD9806FFB2FF90FA06FDC989C3FD2C" box="[1459,1705,619,645]" italics="true" pageId="11" pageNumber="546">T. quadrisignata pilleti</emphasis>
</taxonomicName>
and 
<taxonomicName id="50893F97FFB2FF90F96AFDC98826FD2C" authorityName="Boulard" authorityYear="1981" box="[1759,1868,619,645]" class="Insecta" family="Cicadidae" genus="Tibicina" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="546" phylum="Arthropoda" rank="species" species="maldesi">
<emphasis id="A5FD9806FFB2FF90F96AFDC98826FD2C" box="[1759,1868,619,645]" italics="true" pageId="11" pageNumber="546">T. maldesi</emphasis>
</taxonomicName>
calling signals, with: CD, call duration of a sequence (s); GPD, group of pulses duration (ms); GPP, group of pulses period (ms); NGP, number of group of pulses per second; NP, number of pulses per group. MF, frequencies with maximum amplitude; F1, peak of the first frequency band; F2, peak of the second frequency; F3, peak of the third frequency band; *carrier frequency band. See “Materials and methods” for more explanations.
</paragraph>
</caption>
<paragraph id="97364414FFB2FF90FEEFFCA2888BF9AA" pageId="11" pageNumber="546">
<table id="E589B6B4FFB20064FF17FCA2888BF988" box="[162,2017,768,1569]" gridcols="10" gridrows="13" pageId="11" pageNumber="546">
<tr id="29B94656FFB20064FF17FCA2888BFCB1" box="[162,2017,768,792]" gridrow="0" pageId="11" pageNumber="546" rowspan-0="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="6A682F2AFFB20064FEEFFCA28DB9FCB1" box="[346,723,768,792]" colspan="2" colspanRight="1" gridcol="1" gridrow="0" pageId="11" pageNumber="546">
<emphasis id="A5FD9806FFB2FF90FEEFFCA28D52FCB1" bold="true" box="[346,568,768,792]" pageId="11" pageNumber="546">Temporal parameters</emphasis>
</th>
<th id="6A682F2AFFB20064FA94FCA28962FCB1" box="[1313,1544,768,792]" gridcol="6" gridrow="0" pageId="11" pageNumber="546">
<emphasis id="A5FD9806FFB2FF90FA94FCA28962FCB1" bold="true" box="[1313,1544,768,792]" pageId="11" pageNumber="546">Frequency parameters</emphasis>
</th>
</tr>
<tr id="29B94656FFB20064FF17FC85888BFC96" box="[162,2017,807,831]" gridrow="1" pageId="11" pageNumber="546" rowspan-0="1">
<td id="6A682F2AFFB20064FEEFFC858E88FC96" box="[346,482,807,831]" gridcol="1" gridrow="1" pageId="11" pageNumber="546">
<emphasis id="A5FD9806FFB2FF90FE20FC858ED2FC96" bold="true" box="[405,440,807,831]" pageId="11" pageNumber="546">CD</emphasis>
</td>
<td id="6A682F2AFFB20064FD87FC858DB9FC96" box="[562,723,807,831]" gridcol="2" gridrow="1" pageId="11" pageNumber="546">
<emphasis id="A5FD9806FFB2FF90FDE0FC858DE2FC96" bold="true" box="[597,648,807,831]" pageId="11" pageNumber="546">GPD</emphasis>
</td>
<td id="6A682F2AFFB20064FCA6FC858CDEFC96" box="[787,948,807,831]" gridcol="3" gridrow="1" pageId="11" pageNumber="546">
<emphasis id="A5FD9806FFB2FF90FC82FC858C0DFC96" bold="true" box="[823,871,807,831]" pageId="11" pageNumber="546">GPP</emphasis>
</td>
<td id="6A682F2AFFB20064FC44FC858BECFC96" box="[1009,1158,807,831]" gridcol="4" gridrow="1" pageId="11" pageNumber="546">
<emphasis id="A5FD9806FFB2FF90FBBAFC858B28FC96" bold="true" box="[1039,1090,807,831]" pageId="11" pageNumber="546">NGP</emphasis>
</td>
<td id="6A682F2AFFB20064FB04FC858B87FC96" box="[1201,1261,807,831]" gridcol="5" gridrow="1" pageId="11" pageNumber="546">
<emphasis id="A5FD9806FFB2FF90FB00FC858BBFFC96" bold="true" box="[1205,1237,807,831]" pageId="11" pageNumber="546">NP</emphasis>
</td>
<td id="6A682F2AFFB20064FA94FC858962FC96" box="[1313,1544,807,831]" gridcol="6" gridrow="1" pageId="11" pageNumber="546">
<emphasis id="A5FD9806FFB2FF90FA37FC858ACDFC96" bold="true" box="[1410,1447,807,831]" pageId="11" pageNumber="546">MF</emphasis>
</td>
<td id="6A682F2AFFB20064F998FC8589F0FC96" box="[1581,1690,807,831]" gridcol="7" gridrow="1" pageId="11" pageNumber="546">
<emphasis id="A5FD9806FFB2FF90F9E3FC85891BFC96" bold="true" box="[1622,1649,807,831]" pageId="11" pageNumber="546">F1</emphasis>
</td>
<td id="6A682F2AFFB20064F972FC85882BFC96" box="[1735,1857,807,831]" gridcol="8" gridrow="1" pageId="11" pageNumber="546">
<emphasis id="A5FD9806FFB2FF90F942FC85887BFC96" bold="true" box="[1783,1809,807,831]" pageId="11" pageNumber="546">F2</emphasis>
</td>
<td id="6A682F2AFFB20064F8DDFC85888BFC96" box="[1896,2017,807,831]" gridcol="9" gridrow="1" pageId="11" pageNumber="546">
<emphasis id="A5FD9806FFB2FF90F822FC8588D8FC96" bold="true" box="[1943,1970,807,831]" pageId="11" pageNumber="546">F3</emphasis>
</td>
</tr>
<tr id="29B94656FFB20064FF17FCEC888BFCCC" box="[162,2017,846,869]" gridrow="2" pageId="11" pageNumber="546">
<th id="6A682F2AFFB20064FF17FCEC888BFCCC" box="[162,2017,846,869]" colspan="10" colspanRight="9" gridcol="0" gridrow="2" pageId="11" pageNumber="546">
<taxonomicName id="50893F97FFB2FF90FF17FCEC8EE8FCCC" box="[162,386,846,869]" class="Insecta" family="Cicadidae" genus="Tibicina" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="546" phylum="Arthropoda" rank="subSpecies" species="quadrisignata" subSpecies="pilleti">
<emphasis id="A5FD9806FFB2FF90FF17FCEC8EE8FCCC" bold="true" box="[162,386,846,869]" italics="true" pageId="11" pageNumber="546">T. quadrisignata pilleti</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="29B94656FFB20064FF17FCDA888BFC04" box="[162,2017,888,941]" gridrow="3" pageId="11" pageNumber="546">
<th id="6A682F2AFFB20064FF17FCDA8E6DFC04" box="[162,263,888,941]" gridcol="0" gridrow="3" pageId="11" pageNumber="546">
<emphasis id="A5FD9806FFB2FF90FF17FCDA8F82FC39" bold="true" box="[162,232,888,912]" pageId="11" pageNumber="546">Male 1</emphasis>
</th>
<td id="6A682F2AFFB20064FEEFFCDA8E88FC04" box="[346,482,888,941]" gridcol="1" gridrow="3" pageId="11" pageNumber="546">52.1</td>
<td id="6A682F2AFFB20064FD87FCDA8DB9FC04" box="[562,723,888,941]" gridcol="2" gridrow="3" pageId="11" pageNumber="546">12.8 ± 0.3 (12.2–13.4; 86)</td>
<td id="6A682F2AFFB20064FCA6FCDA8CDEFC04" box="[787,948,888,941]" gridcol="3" gridrow="3" pageId="11" pageNumber="546">14.1 ± 0.2 (13.6–14.5; 86)</td>
<td id="6A682F2AFFB20064FC44FCDA8BECFC04" box="[1009,1158,888,941]" gridcol="4" gridrow="3" pageId="11" pageNumber="546">70.8 ± 0.4 (70–72; 65)</td>
<td id="6A682F2AFFB20064FB04FCDA8B87FC04" box="[1201,1261,888,941]" gridcol="5" gridrow="3" pageId="11" pageNumber="546">8–9</td>
<td id="6A682F2AFFB20064FA94FCDA8962FC04" box="[1313,1544,888,941]" gridcol="6" gridrow="3" pageId="11" pageNumber="546">8908 ± 558 (7946–9798; 89)</td>
<td id="6A682F2AFFB20064F998FCDA89F0FC04" box="[1581,1690,888,941]" gridcol="7" gridrow="3" pageId="11" pageNumber="546">
<emphasis id="A5FD9806FFB2FF90F9F0FCDA89EBFC39" bold="true" box="[1605,1665,888,912]" pageId="11" pageNumber="546">*8419</emphasis>
</td>
<td id="6A682F2AFFB20064F972FCDA882BFC04" box="[1735,1857,888,941]" gridcol="8" gridrow="3" pageId="11" pageNumber="546">9195</td>
<td id="6A682F2AFFB20064F8DDFCDA888BFC04" box="[1896,2017,888,941]" gridcol="9" gridrow="3" pageId="11" pageNumber="546">9690</td>
</tr>
<tr id="29B94656FFB20064FF17FC1F888BFC58" box="[162,2017,957,1009]" gridrow="4" pageId="11" pageNumber="546">
<th id="6A682F2AFFB20064FF17FC1F8E6DFC58" box="[162,263,957,1009]" gridcol="0" gridrow="4" pageId="11" pageNumber="546">
<emphasis id="A5FD9806FFB2FF90FF17FC1F8F82FC7C" bold="true" box="[162,232,957,981]" pageId="11" pageNumber="546">Male 2</emphasis>
</th>
<td id="6A682F2AFFB20064FEEFFC1F8E88FC58" box="[346,482,957,1009]" gridcol="1" gridrow="4" pageId="11" pageNumber="546">46.3</td>
<td id="6A682F2AFFB20064FD87FC1F8DB9FC58" box="[562,723,957,1009]" gridcol="2" gridrow="4" pageId="11" pageNumber="546">13.2 ± 0.2 (13.0–13.6; 85)</td>
<td id="6A682F2AFFB20064FCA6FC1F8CDEFC58" box="[787,948,957,1009]" gridcol="3" gridrow="4" pageId="11" pageNumber="546">13.9 ± 0.2 (13.3–14.2; 91)</td>
<td id="6A682F2AFFB20064FC44FC1F8BECFC58" box="[1009,1158,957,1009]" gridcol="4" gridrow="4" pageId="11" pageNumber="546">72.4 ± 0.5 (71–73; 80)</td>
<td id="6A682F2AFFB20064FB04FC1F8B87FC58" box="[1201,1261,957,1009]" gridcol="5" gridrow="4" pageId="11" pageNumber="546">10–11</td>
<td id="6A682F2AFFB20064FA94FC1F8962FC58" box="[1313,1544,957,1009]" gridcol="6" gridrow="4" pageId="11" pageNumber="546">7867 ± 177 (7644–8871; 81)</td>
<td id="6A682F2AFFB20064F998FC1F89F0FC58" box="[1581,1690,957,1009]" gridcol="7" gridrow="4" pageId="11" pageNumber="546">
<emphasis id="A5FD9806FFB2FF90F9F0FC1F89EBFC7C" bold="true" box="[1605,1665,957,981]" pageId="11" pageNumber="546">*7817</emphasis>
</td>
<td id="6A682F2AFFB20064F972FC1F882BFC58" box="[1735,1857,957,1009]" gridcol="8" gridrow="4" pageId="11" pageNumber="546">8764</td>
<td id="6A682F2AFFB20064F8DDFC1F888BFC58" box="[1896,2017,957,1009]" gridcol="9" gridrow="4" pageId="11" pageNumber="546">9388</td>
</tr>
<tr id="29B94656FFB20064FF17FBA3888BFB9F" box="[162,2017,1025,1078]" gridrow="5" pageId="11" pageNumber="546">
<th id="6A682F2AFFB20064FF17FBA38E6DFB9F" box="[162,263,1025,1078]" gridcol="0" gridrow="5" pageId="11" pageNumber="546">
<emphasis id="A5FD9806FFB2FF90FF17FBA38F82FBB0" bold="true" box="[162,232,1025,1049]" pageId="11" pageNumber="546">Male 3</emphasis>
</th>
<td id="6A682F2AFFB20064FEEFFBA38E88FB9F" box="[346,482,1025,1078]" gridcol="1" gridrow="5" pageId="11" pageNumber="546">73.4</td>
<td id="6A682F2AFFB20064FD87FBA38DB9FB9F" box="[562,723,1025,1078]" gridcol="2" gridrow="5" pageId="11" pageNumber="546">13.0 ± 0.2 (12.6–13.3; 102)</td>
<td id="6A682F2AFFB20064FCA6FBA38CDEFB9F" box="[787,948,1025,1078]" gridcol="3" gridrow="5" pageId="11" pageNumber="546">14.0 ± 0.1 (13.7–14.5; 111)</td>
<td id="6A682F2AFFB20064FC44FBA38BECFB9F" box="[1009,1158,1025,1078]" gridcol="4" gridrow="5" pageId="11" pageNumber="546">71.8 ± 0.4 (71–72; 107)</td>
<td id="6A682F2AFFB20064FB04FBA38B87FB9F" box="[1201,1261,1025,1078]" gridcol="5" gridrow="5" pageId="11" pageNumber="546">8–11</td>
<td id="6A682F2AFFB20064FA94FBA38962FB9F" box="[1313,1544,1025,1078]" gridcol="6" gridrow="5" pageId="11" pageNumber="546">8929 ± 444 (7773–9647; 84)</td>
<td id="6A682F2AFFB20064F998FBA389F0FB9F" box="[1581,1690,1025,1078]" gridcol="7" gridrow="5" pageId="11" pageNumber="546">8204</td>
<td id="6A682F2AFFB20064F972FBA3882BFB9F" box="[1735,1857,1025,1078]" gridcol="8" gridrow="5" pageId="11" pageNumber="546">
<emphasis id="A5FD9806FFB2FF90F953FBA38848FBB0" bold="true" box="[1766,1826,1025,1049]" pageId="11" pageNumber="546">*9022</emphasis>
</td>
<td id="6A682F2AFFB20064F8DDFBA3888BFB9F" box="[1896,2017,1025,1078]" gridcol="9" gridrow="5" pageId="11" pageNumber="546">9561</td>
</tr>
<tr id="29B94656FFB20064FF17FBE4888BFBD2" box="[162,2017,1094,1147]" gridrow="6" pageId="11" pageNumber="546">
<th id="6A682F2AFFB20064FF17FBE48E6DFBD2" box="[162,263,1094,1147]" gridcol="0" gridrow="6" pageId="11" pageNumber="546">
<emphasis id="A5FD9806FFB2FF90FF17FBE48F82FBF7" bold="true" box="[162,232,1094,1118]" pageId="11" pageNumber="546">Male 4</emphasis>
</th>
<td id="6A682F2AFFB20064FEEFFBE48E88FBD2" box="[346,482,1094,1147]" gridcol="1" gridrow="6" pageId="11" pageNumber="546">98</td>
<td id="6A682F2AFFB20064FD87FBE48DB9FBD2" box="[562,723,1094,1147]" gridcol="2" gridrow="6" pageId="11" pageNumber="546">13.0 ± 0.2 (12.7–13.4; 73)</td>
<td id="6A682F2AFFB20064FCA6FBE48CDEFBD2" box="[787,948,1094,1147]" gridcol="3" gridrow="6" pageId="11" pageNumber="546">14.1 ± 0.3 (12,9–14.6; 77)</td>
<td id="6A682F2AFFB20064FC44FBE48BECFBD2" box="[1009,1158,1094,1147]" gridcol="4" gridrow="6" pageId="11" pageNumber="546">70.5 ± 0.5 (70–71; 77)</td>
<td id="6A682F2AFFB20064FB04FBE48B87FBD2" box="[1201,1261,1094,1147]" gridcol="5" gridrow="6" pageId="11" pageNumber="546">8–11</td>
<td id="6A682F2AFFB20064FA94FBE48962FBD2" box="[1313,1544,1094,1147]" gridcol="6" gridrow="6" pageId="11" pageNumber="546">8460 ± 358 (7429–9087; 83)</td>
<td id="6A682F2AFFB20064F998FBE489F0FBD2" box="[1581,1690,1094,1147]" gridcol="7" gridrow="6" pageId="11" pageNumber="546">8333</td>
<td id="6A682F2AFFB20064F972FBE4882BFBD2" box="[1735,1857,1094,1147]" gridcol="8" gridrow="6" pageId="11" pageNumber="546">
<emphasis id="A5FD9806FFB2FF90F953FBE48848FBF7" bold="true" box="[1766,1826,1094,1118]" pageId="11" pageNumber="546">*8807</emphasis>
</td>
<td id="6A682F2AFFB20064F8DDFBE4888BFBD2" box="[1896,2017,1094,1147]" gridcol="9" gridrow="6" pageId="11" pageNumber="546">9561</td>
</tr>
<tr id="29B94656FFB20064FF17FB29888BFB69" box="[162,2017,1163,1216]" gridrow="7" pageId="11" pageNumber="546">
<th id="6A682F2AFFB20064FF17FB298E6DFB69" box="[162,263,1163,1216]" gridcol="0" gridrow="7" pageId="11" pageNumber="546">
<emphasis id="A5FD9806FFB2FF90FF17FB298F83FB69" bold="true" box="[162,263,1163,1216]" pageId="11" pageNumber="546">Summary (N = 4)</emphasis>
</th>
<td id="6A682F2AFFB20064FEEFFB298E88FB69" box="[346,482,1163,1216]" gridcol="1" gridrow="7" pageId="11" pageNumber="546">67.4 ± 23.5</td>
<td id="6A682F2AFFB20064FD87FB298DB9FB69" box="[562,723,1163,1216]" gridcol="2" gridrow="7" pageId="11" pageNumber="546">13.02 ± 0.2 (12.2–13.6; 346)</td>
<td id="6A682F2AFFB20064FCA6FB298CDEFB69" box="[787,948,1163,1216]" gridcol="3" gridrow="7" pageId="11" pageNumber="546">14.0 ± 0.2 (12.9–14.6; 365)</td>
<td id="6A682F2AFFB20064FC44FB298BECFB69" box="[1009,1158,1163,1216]" gridcol="4" gridrow="7" pageId="11" pageNumber="546">71.5 ± 0.9 (70–73; 329)</td>
<td id="6A682F2AFFB20064FB04FB298B87FB69" box="[1201,1261,1163,1216]" gridcol="5" gridrow="7" pageId="11" pageNumber="546">8–11</td>
<td id="6A682F2AFFB20064FA94FB298962FB69" box="[1313,1544,1163,1216]" gridcol="6" gridrow="7" pageId="11" pageNumber="546">8553 ± 594 (7429–9798; 337)</td>
<td id="6A682F2AFFB20064F998FB2989F0FB69" box="[1581,1690,1163,1216]" gridcol="7" gridrow="7" pageId="11" pageNumber="546">8193 ± 266</td>
<td id="6A682F2AFFB20064F972FB29882BFB69" box="[1735,1857,1163,1216]" gridcol="8" gridrow="7" pageId="11" pageNumber="546">
<emphasis id="A5FD9806FFB2FF90F972FB29882BFB0A" bold="true" box="[1735,1857,1163,1187]" pageId="11" pageNumber="546">*8947 ± 200</emphasis>
</td>
<td id="6A682F2AFFB20064F8DDFB29888BFB69" box="[1896,2017,1163,1216]" gridcol="9" gridrow="7" pageId="11" pageNumber="546">9550 ± 124</td>
</tr>
<tr id="29B94656FFB20064FF17FB55888BFAA7" box="[162,2017,1271,1294]" gridrow="8" pageId="11" pageNumber="546">
<th id="6A682F2AFFB20064FF17FB55888BFAA7" box="[162,2017,1271,1294]" colspan="10" colspanRight="9" gridcol="0" gridrow="8" pageId="11" pageNumber="546">
<taxonomicName id="50893F97FFB2FF90FF17FB558E6DFAA7" authorityName="Boulard" authorityYear="1981" box="[162,263,1271,1294]" class="Insecta" family="Cicadidae" genus="Tibicina" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="546" phylum="Arthropoda" rank="species" species="maldesi">
<emphasis id="A5FD9806FFB2FF90FF17FB558E6DFAA7" bold="true" box="[162,263,1271,1294]" italics="true" pageId="11" pageNumber="546">T. maldesi</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="29B94656FFB20064FF17FABC888BFAFA" box="[162,2017,1310,1363]" gridrow="9" pageId="11" pageNumber="546">
<th id="6A682F2AFFB20064FF17FABC8E6DFAFA" box="[162,263,1310,1363]" gridcol="0" gridrow="9" pageId="11" pageNumber="546">
<emphasis id="A5FD9806FFB2FF90FF17FABC8F82FA9F" bold="true" box="[162,232,1310,1334]" pageId="11" pageNumber="546">Male 1</emphasis>
</th>
<td id="6A682F2AFFB20064FEEFFABC8E88FAFA" box="[346,482,1310,1363]" gridcol="1" gridrow="9" pageId="11" pageNumber="546">839.7</td>
<td id="6A682F2AFFB20064FD87FABC8DB9FAFA" box="[562,723,1310,1363]" gridcol="2" gridrow="9" pageId="11" pageNumber="546">8.3 ± 0.4 (7.6–9.0; 100)</td>
<td id="6A682F2AFFB20064FCA6FABC8CDEFAFA" box="[787,948,1310,1363]" gridcol="3" gridrow="9" pageId="11" pageNumber="546">10.9 ± 0.6 (9.6–12.3; 121)</td>
<td id="6A682F2AFFB20064FC44FABC8BECFAFA" box="[1009,1158,1310,1363]" gridcol="4" gridrow="9" pageId="11" pageNumber="546">102.9 ± 1.4 (100–106; 103)</td>
<td id="6A682F2AFFB20064FB04FABC8B87FAFA" box="[1201,1261,1310,1363]" gridcol="5" gridrow="9" pageId="11" pageNumber="546">8–10</td>
<td id="6A682F2AFFB20064FA94FABC8962FAFA" box="[1313,1544,1310,1363]" gridcol="6" gridrow="9" pageId="11" pageNumber="546">8593 ± 470 (7644–9367; 45)</td>
<td id="6A682F2AFFB20064F998FABC89F0FAFA" box="[1581,1690,1310,1363]" gridcol="7" gridrow="9" pageId="11" pageNumber="546">7881</td>
<td id="6A682F2AFFB20064F972FABC882BFAFA" box="[1735,1857,1310,1363]" gridcol="8" gridrow="9" pageId="11" pageNumber="546">
<emphasis id="A5FD9806FFB2FF90F953FABC8848FA9F" bold="true" box="[1766,1826,1310,1334]" pageId="11" pageNumber="546">*8699</emphasis>
</td>
<td id="6A682F2AFFB20064F8DDFABC888BFAFA" box="[1896,2017,1310,1363]" gridcol="9" gridrow="9" pageId="11" pageNumber="546">10357</td>
</tr>
<tr id="29B94656FFB20064FF17FAC0888BFA3E" box="[162,2017,1378,1431]" gridrow="10" pageId="11" pageNumber="546">
<th id="6A682F2AFFB20064FF17FAC08E6DFA3E" box="[162,263,1378,1431]" gridcol="0" gridrow="10" pageId="11" pageNumber="546">
<emphasis id="A5FD9806FFB2FF90FF17FAC08F82FAD3" bold="true" box="[162,232,1378,1402]" pageId="11" pageNumber="546">Male 2</emphasis>
</th>
<td id="6A682F2AFFB20064FEEFFAC08E88FA3E" box="[346,482,1378,1431]" gridcol="1" gridrow="10" pageId="11" pageNumber="546">874.5</td>
<td id="6A682F2AFFB20064FD87FAC08DB9FA3E" box="[562,723,1378,1431]" gridcol="2" gridrow="10" pageId="11" pageNumber="546">8.3 ± 0.4 (7.6–8.8; 103)</td>
<td id="6A682F2AFFB20064FCA6FAC08CDEFA3E" box="[787,948,1378,1431]" gridcol="3" gridrow="10" pageId="11" pageNumber="546">10.2 ± 0.6 (9.0–11.9; 131)</td>
<td id="6A682F2AFFB20064FC44FAC08BECFA3E" box="[1009,1158,1378,1431]" gridcol="4" gridrow="10" pageId="11" pageNumber="546">106.1 ± 0.5 (104–108; 106)</td>
<td id="6A682F2AFFB20064FB04FAC08B87FA3E" box="[1201,1261,1378,1431]" gridcol="5" gridrow="10" pageId="11" pageNumber="546">8–10</td>
<td id="6A682F2AFFB20064FA94FAC08962FA3E" box="[1313,1544,1378,1431]" gridcol="6" gridrow="10" pageId="11" pageNumber="546">9587 ± 469 (8506–10293; 50)</td>
<td id="6A682F2AFFB20064F998FAC089F0FA3E" box="[1581,1690,1378,1431]" gridcol="7" gridrow="10" pageId="11" pageNumber="546">8613</td>
<td id="6A682F2AFFB20064F972FAC0882BFA3E" box="[1735,1857,1378,1431]" gridcol="8" gridrow="10" pageId="11" pageNumber="546">
<emphasis id="A5FD9806FFB2FF90F953FAC08848FAD3" bold="true" box="[1766,1826,1378,1402]" pageId="11" pageNumber="546">*9087</emphasis>
</td>
<td id="6A682F2AFFB20064F8DDFAC0888BFA3E" box="[1896,2017,1378,1431]" gridcol="9" gridrow="10" pageId="11" pageNumber="546">10034</td>
</tr>
<tr id="29B94656FFB20064FF17FA05888BFA75" box="[162,2017,1447,1500]" gridrow="11" pageId="11" pageNumber="546">
<th id="6A682F2AFFB20064FF17FA058E6DFA75" box="[162,263,1447,1500]" gridcol="0" gridrow="11" pageId="11" pageNumber="546">
<emphasis id="A5FD9806FFB2FF90FF17FA058F82FA16" bold="true" box="[162,232,1447,1471]" pageId="11" pageNumber="546">Male 3</emphasis>
</th>
<td id="6A682F2AFFB20064FEEFFA058E88FA75" box="[346,482,1447,1500]" gridcol="1" gridrow="11" pageId="11" pageNumber="546">719.9</td>
<td id="6A682F2AFFB20064FD87FA058DB9FA75" box="[562,723,1447,1500]" gridcol="2" gridrow="11" pageId="11" pageNumber="546">8.3 ± 0.3 (7.6–8.8; 98)</td>
<td id="6A682F2AFFB20064FCA6FA058CDEFA75" box="[787,948,1447,1500]" gridcol="3" gridrow="11" pageId="11" pageNumber="546">10.1 ± 0.5 (9.2–11.9; 125)</td>
<td id="6A682F2AFFB20064FC44FA058BECFA75" box="[1009,1158,1447,1500]" gridcol="4" gridrow="11" pageId="11" pageNumber="546">104.8 ± 1.4 (102–108; 107)</td>
<td id="6A682F2AFFB20064FB04FA058B87FA75" box="[1201,1261,1447,1500]" gridcol="5" gridrow="11" pageId="11" pageNumber="546">8–10</td>
<td id="6A682F2AFFB20064FA94FA058962FA75" box="[1313,1544,1447,1500]" gridcol="6" gridrow="11" pageId="11" pageNumber="546">9172 ± 318 (8484–9668; 45)</td>
<td id="6A682F2AFFB20064F998FA0589F0FA75" box="[1581,1690,1447,1500]" gridcol="7" gridrow="11" pageId="11" pageNumber="546">8656</td>
<td id="6A682F2AFFB20064F972FA05882BFA75" box="[1735,1857,1447,1500]" gridcol="8" gridrow="11" pageId="11" pageNumber="546">
<emphasis id="A5FD9806FFB2FF90F953FA058848FA16" bold="true" box="[1766,1826,1447,1471]" pageId="11" pageNumber="546">*9388</emphasis>
</td>
<td id="6A682F2AFFB20064F8DDFA05888BFA75" box="[1896,2017,1447,1500]" gridcol="9" gridrow="11" pageId="11" pageNumber="546">10034</td>
</tr>
<tr id="29B94656FFB20064FF17FA4E888BF988" box="[162,2017,1516,1569]" gridrow="12" pageId="11" pageNumber="546">
<th id="6A682F2AFFB20064FF17FA4E8E6DF988" box="[162,263,1516,1569]" gridcol="0" gridrow="12" pageId="11" pageNumber="546">
<emphasis id="A5FD9806FFB2FF90FF17FA4E8F83F988" bold="true" box="[162,263,1516,1569]" pageId="11" pageNumber="546">Summary (N = 3)</emphasis>
</th>
<td id="6A682F2AFFB20064FEEFFA4E8E88F988" box="[346,482,1516,1569]" gridcol="1" gridrow="12" pageId="11" pageNumber="546">811.4 ± 81.1</td>
<td id="6A682F2AFFB20064FD87FA4E8DB9F988" box="[562,723,1516,1569]" gridcol="2" gridrow="12" pageId="11" pageNumber="546">8.3 ± 0.4 (7.6–9.0; 301)</td>
<td id="6A682F2AFFB20064FCA6FA4E8CDEF988" box="[787,948,1516,1569]" gridcol="3" gridrow="12" pageId="11" pageNumber="546">10.4 ± 0.7 (9.0–12.3; 377)</td>
<td id="6A682F2AFFB20064FC44FA4E8BECF988" box="[1009,1158,1516,1569]" gridcol="4" gridrow="12" pageId="11" pageNumber="546">104.6 ± 1.7 (100–108; 316)</td>
<td id="6A682F2AFFB20064FB04FA4E8B87F988" box="[1201,1261,1516,1569]" gridcol="5" gridrow="12" pageId="11" pageNumber="546">8–10</td>
<td id="6A682F2AFFB20064FA94FA4E8962F988" box="[1313,1544,1516,1569]" gridcol="6" gridrow="12" pageId="11" pageNumber="546">9134 ± 590 (7644–10293; 140)</td>
<td id="6A682F2AFFB20064F998FA4E89F0F988" box="[1581,1690,1516,1569]" gridcol="7" gridrow="12" pageId="11" pageNumber="546">8384 ± 436</td>
<td id="6A682F2AFFB20064F972FA4E882BF988" box="[1735,1857,1516,1569]" gridcol="8" gridrow="12" pageId="11" pageNumber="546">
<emphasis id="A5FD9806FFB2FF90F972FA4E882BF9AD" bold="true" box="[1735,1857,1516,1540]" pageId="11" pageNumber="546">*9058 ± 345</emphasis>
</td>
<td id="6A682F2AFFB20064F8DDFA4E888BF988" box="[1896,2017,1516,1569]" gridcol="9" gridrow="12" pageId="11" pageNumber="546">10142 ± 186</td>
</tr>
</table>
</paragraph>
<caption id="C3F6149CFFB4FF96FF22FF348A1AFE66" ID-DOI="http://doi.org/10.5281/zenodo.8257063" ID-Zenodo-Dep="8257063" httpUri="https://zenodo.org/record/8257063/files/figure.png" pageId="13" pageNumber="548" startId="13.[151,250,150,174]" targetBox="[160,1432,524,1243]" targetPageId="13" targetType="figure">
<paragraph id="97364414FFB4FF96FF22FF348A1AFE66" blockId="13.[151,1437,150,463]" pageId="13" pageNumber="548">
<emphasis id="A5FD9806FFB4FF96FF22FF348E79FF07" bold="true" box="[151,275,150,174]" pageId="13" pageNumber="548">FIGURE 9.</emphasis>
Calling song recording of 
<emphasis id="A5FD9806FFB4FF96FDA9FF358DE3FF06" box="[540,649,151,175]" italics="true" pageId="13" pageNumber="548">T. q. pilleti</emphasis>
<emphasis id="A5FD9806FFB4FF96FD3BFF348C5AFF06" bold="true" box="[654,816,150,175]" pageId="13" pageNumber="548">
Puissant 
<taxonomicNameLabel id="BECE257DFFB4FF96FD5BFF358C5AFF06" box="[750,816,151,175]" pageId="13" pageNumber="548" rank="subSpecies">ssp. n.</taxonomicNameLabel>
</emphasis>
with a digitization at 44.1 kHz sampling frequency.A, spectra (frequency 
<emphasis id="A5FD9806FFB4FF96FEBFFF198E75FF7B" box="[266,287,187,210]" italics="true" pageId="13" pageNumber="548">vs</emphasis>
. absolute amplitude). B, spectrogram and C, oscillogram (from top to bottom) of the beginning of the sequence. D, oscillogram of a part of C showing groups of pulses (slow amplitude modulation). E, detailed spectrogram and F, oscillogram of a part of D showing a train of groups of pulses in detail. F1, F2*, F3: three frequency peaks with F2* as carrier frequency; FFT: fast Fourier transform; GP: group of pulses (= syllable); MF: frequencies of maximum amplitude; N: number of males (N = 4) analyzed; n: number of measurements for sampling (n = 337); NAV: number of amplitude variations; light grey spectrum (at bottom): minimum amplitude spectrum plotted from the minimum amplitude value for each frequency; black spectrum (at middle): average linear amplitude spectrum plotted from the average amplitude value recorded for each frequency; grey spectrum (at top): maximum amplitude spectrum plotted from the maximum amplitude value recorded for each frequency.
</paragraph>
</caption>
<caption id="C3F6149CFFB4FF96FF22FB518D26FAFA" ID-DOI="http://doi.org/10.5281/zenodo.8255317" ID-Zenodo-Dep="8255317" httpUri="https://zenodo.org/record/8255317/files/figure.png" pageId="13" pageNumber="548" startId="13.[151,250,1267,1291]" targetBox="[160,1432,524,1243]" targetPageId="13" targetType="figure">
<paragraph id="97364414FFB4FF96FF22FB518D26FAFA" blockId="13.[151,1437,1267,1363]" pageId="13" pageNumber="548">
<emphasis id="A5FD9806FFB4FF96FF22FB518E4AFAA2" bold="true" box="[151,288,1267,1291]" pageId="13" pageNumber="548">FIGURE 10.</emphasis>
Spectrogram and oscillogram (from top to bottom) of the rivalry song of 
<emphasis id="A5FD9806FFB4FF96FBA2FB568BEEFAA2" box="[1047,1156,1267,1291]" italics="true" pageId="13" pageNumber="548">T. q. pilleti</emphasis>
<emphasis id="A5FD9806FFB4FF96FB3EFB518A58FAA2" bold="true" box="[1163,1330,1267,1291]" pageId="13" pageNumber="548">
Puissant 
<taxonomicNameLabel id="BECE257DFFB4FF96FB5BFB518A58FAA2" box="[1262,1330,1267,1291]" pageId="13" pageNumber="548" rank="subSpecies">ssp. n.</taxonomicNameLabel>
</emphasis>
One male producing an irregular train of echemes in the foreground (M2) is interrupted by another male producing a fast trains of rivalry song pulse groups in the background (M1).
</paragraph>
</caption>
<paragraph id="97364414FFB4FF96FF72F9B38D15F88E" blockId="13.[151,1437,1445,2011]" pageId="13" pageNumber="548">
Calling songs recording with digitization at 96 kHz sampling frequency (Hamming window width = 1024 points, frequency resolution = 94 Hz) shows a frequency amplitude envelope peaking at 28 kHz, encapsulating the second upper harmonic at around 27 kHz (
<figureCitation id="0FB25891FFB4FF96FDCDF9FB8DD6F9DA" box="[632,700,1625,1651]" captionStart="FIGURE 8" captionStartId="10.[151,250,1626,1650]" captionTargetBox="[160,1430,886,1603]" captionTargetId="figure-354@10.[151,1436,885,1603]" captionTargetPageId="10" captionText="FIGURE 8. Calling song recording of T. q. pilleti Puissant ssp. n. with a digitization at 96 kHz sampling frequency. Frequency amplitude envelope peaking at 28 kHz, encapsulating the second upper harmonic at around 27 kHz." figureDoi="http://doi.org/10.5281/zenodo.8255315" httpUri="https://zenodo.org/record/8255315/files/figure.png" pageId="13" pageNumber="548">Fig. 8</figureCitation>
). To be more accurate in the frequency domain, the calling songs recording were subsequently digitized at 44.1 kHz (Hamming window width = 1024 points, frequency resolution = 43 Hz). Power spectra (
<tableCitation id="DA0B71AFFFB4FF96FE00F9038D60F912" box="[437,522,1697,1723]" captionStart="TABLE 1" captionStartId="11.[151,239,619,645]" captionTargetPageId="11" captionText="TABLE 1. Mean ± standard deviation (minimum–maximum; sample size) for the temporal and frequency parameters of T. quadrisignata pilleti and T. maldesi calling signals, with: CD, call duration of a sequence (s); GPD, group of pulses duration (ms); GPP, group of pulses period (ms); NGP, number of group of pulses per second; NP, number of pulses per group. MF, frequencies with maximum amplitude; F1, peak of the first frequency band; F2, peak of the second frequency; F3, peak of the third frequency band; *carrier frequency band. See “Materials and methods” for more explanations." httpUri="http://table.plazi.org/id/C3F6149CFFB2FF90FF22FDC98C13FD58" pageId="13" pageNumber="548" tableUuid="C3F6149CFFB2FF90FF22FDC98C13FD58">Table 1</tableCitation>
; 
<figureCitation id="0FB25891FFB4FF96FDADF9038D05F912" box="[536,623,1697,1723]" captionStart="FIGURE 9" captionStartId="13.[151,250,150,174]" captionTargetId="figure-193@13.[151,1436,523,1243]" captionTargetPageId="13" captionText="FIGURE 9. Calling song recording of T. q. pilleti Puissant ssp. n. with a digitization at 44.1 kHz sampling frequency.A, spectra (frequency vs. absolute amplitude). B, spectrogram and C, oscillogram (from top to bottom) of the beginning of the sequence. D, oscillogram of a part of C showing groups of pulses (slow amplitude modulation). E, detailed spectrogram and F, oscillogram of a part of D showing a train of groups of pulses in detail. F1, F2*, F3: three frequency peaks with F2* as carrier frequency; FFT: fast Fourier transform; GP: group of pulses (= syllable); MF: frequencies of maximum amplitude; N: number of males (N = 4) analyzed; n: number of measurements for sampling (n = 337); NAV: number of amplitude variations; light grey spectrum (at bottom): minimum amplitude spectrum plotted from the minimum amplitude value for each frequency; black spectrum (at middle): average linear amplitude spectrum plotted from the average amplitude value recorded for each frequency; grey spectrum (at top): maximum amplitude spectrum plotted from the maximum amplitude value recorded for each frequency." figureDoi="http://doi.org/10.5281/zenodo.8257063" httpUri="https://zenodo.org/record/8257063/files/figure.png" pageId="13" pageNumber="548">Fig. 9A</figureCitation>
) are characterized by frequencies of maximum amplitude (
<collectionCode id="F198DCD1FFB4FF96FAB6F9038A44F912" box="[1283,1326,1697,1723]" pageId="13" pageNumber="548">MF</collectionCode>
) having a mean of 8553 ± 594 Hz (7429–9798; 337) with three frequency peaks: a peak of the carrier frequency band (F2*) at 8947 ± 200 Hz (8764–9195; 4) with two lateral frequency bands, F1 at about 8193 ± 266 Hz (7817–9419; 4) and F3 at about 9550 ± 124 Hz (9388–9690; 4).
</paragraph>
<paragraph id="97364414FFB4FF96FF72F8938F88F872" blockId="13.[151,1437,1445,2011]" pageId="13" pageNumber="548">
A rivalry sound (
<figureCitation id="0FB25891FFB4FF96FE30F8938EBDF8E2" box="[389,471,1841,1867]" captionStart="FIGURE 10" captionStartId="13.[151,250,1267,1291]" captionTargetBox="[160,1432,524,1243]" captionTargetId="figure-193@13.[151,1436,523,1243]" captionTargetPageId="13" captionText="FIGURE 10. Spectrogram and oscillogram (from top to bottom) of the rivalry song of T. q. pilleti Puissant ssp. n. One male producing an irregular train of echemes in the foreground (M2) is interrupted by another male producing a fast trains of rivalry song pulse groups in the background (M1)." figureDoi="http://doi.org/10.5281/zenodo.8255317" httpUri="https://zenodo.org/record/8255317/files/figure.png" pageId="13" pageNumber="548">Fig. 10</figureCitation>
) has been repeatedly noted when males are perched on the same bush or tree, close to each other. The rivalry signals were produced in continuous sequences of irregular duration, corresponding more or less to the duration of the encounter. Typically, rivalry signals consist of a relatively regular succession of short trains of pulse groups. The dominant frequency F2* is around 8750 Hz for two measurements taken manually in the signal.
</paragraph>
<caption id="C3F6149CFFB7FF95FF22F9638F88F8C0" ID-DOI="http://doi.org/10.5281/zenodo.8255321" ID-Zenodo-Dep="8255321" httpUri="https://zenodo.org/record/8255321/files/figure.png" pageId="14" pageNumber="549" startId="14.[151,250,1729,1753]" targetBox="[164,1431,181,1705]" targetPageId="14" targetType="figure">
<paragraph id="97364414FFB7FF95FF22F9638F88F8C0" blockId="14.[151,1437,1729,1897]" pageId="14" pageNumber="549">
<emphasis id="A5FD9806FFB7FF95FF22F9638E74F970" bold="true" box="[151,286,1729,1753]" pageId="14" pageNumber="549">FIGURE 11.</emphasis>
Courtship song of 
<emphasis id="A5FD9806FFB7FF95FE58F9608D30F970" box="[493,602,1729,1753]" italics="true" pageId="14" pageNumber="549">T. q. pilleti</emphasis>
<emphasis id="A5FD9806FFB7FF95FDD1F9638C64F970" bold="true" box="[612,782,1729,1753]" pageId="14" pageNumber="549">
Puissant 
<taxonomicNameLabel id="BECE257DFFB7FF95FD7CF9638C64F970" box="[713,782,1729,1753]" pageId="14" pageNumber="549" rank="subSpecies">ssp. n.</taxonomicNameLabel>
</emphasis>
A, spectrogram and B, oscillogram (from top to bottom) of a succession of phrases composed of two echemes. C, oscillogram of a part of B showing two phrases and their component echeme, a one long echeme (E1) and a one short terminal echeme (E2). D, detailed spectrogram and E, oscillogram of a part of C showing a phrase with, for each echeme, a low amplitude part duration (LPD) and a shorter high amplitude part duration (HPD).
</paragraph>
</caption>
<paragraph id="97364414FFB7FF94FF72F8358A17FEB4" blockId="14.[151,1436,1943,2042]" lastBlockId="15.[151,1437,151,717]" lastPageId="15" lastPageNumber="550" pageId="14" pageNumber="549">
A courtship behavior was also recorded in 
<specimenCount id="818F8F9DFFB7FF95FD07F83A8C73F818" box="[690,793,1943,1970]" pageId="14" pageNumber="549" type="male">one male</specimenCount>
(
<figureCitation id="0FB25891FFB7FF95FC9FF8358B69F818" box="[810,1027,1943,1969]" captionStart="FIGURE 11" captionStartId="14.[151,250,1729,1753]" captionTargetBox="[164,1431,181,1705]" captionTargetId="figure-22@14.[151,1436,181,1705]" captionTargetPageId="14" captionText="FIGURE 11. Courtship song of T. q. pilleti Puissant ssp. n. A, spectrogram and B, oscillogram (from top to bottom) of a succession of phrases composed of two echemes. C, oscillogram of a part of B showing two phrases and their component echeme, a one long echeme (E1) and a one short terminal echeme (E2). D, detailed spectrogram and E, oscillogram of a part of C showing a phrase with, for each echeme, a low amplitude part duration (LPD) and a shorter high amplitude part duration (HPD)." figureDoi="http://doi.org/10.5281/zenodo.8255321" httpUri="https://zenodo.org/record/8255321/files/figure.png" pageId="14" pageNumber="549">Figs. 11A,B,C,D,E</figureCitation>
). It occurred when a female arrived in the proximity of the calling male. Male courtship songs consisted of a succession of phrases composed of two echemes (
<figureCitation id="0FB25891FFB7FF95FEB2F87D8E07F850" box="[263,365,2015,2041]" captionStart="FIGURE 11" captionStartId="14.[151,250,1729,1753]" captionTargetBox="[164,1431,181,1705]" captionTargetId="figure-22@14.[151,1436,181,1705]" captionTargetPageId="14" captionText="FIGURE 11. Courtship song of T. q. pilleti Puissant ssp. n. A, spectrogram and B, oscillogram (from top to bottom) of a succession of phrases composed of two echemes. C, oscillogram of a part of B showing two phrases and their component echeme, a one long echeme (E1) and a one short terminal echeme (E2). D, detailed spectrogram and E, oscillogram of a part of C showing a phrase with, for each echeme, a low amplitude part duration (LPD) and a shorter high amplitude part duration (HPD)." figureDoi="http://doi.org/10.5281/zenodo.8255321" httpUri="https://zenodo.org/record/8255321/files/figure.png" pageId="14" pageNumber="549">Fig. 11C</figureCitation>
: E1 and E2): a first of about 3 s duration (E1; n = 6) and a second shorter of almost 1 s duration (E2; n = 6). Each echeme (
<figureCitation id="0FB25891FFB6FF94FE77FF358D49FF18" box="[450,547,151,177]" captionStart="FIGURE 11" captionStartId="14.[151,250,1729,1753]" captionTargetBox="[164,1431,181,1705]" captionTargetId="figure-22@14.[151,1436,181,1705]" captionTargetPageId="14" captionText="FIGURE 11. Courtship song of T. q. pilleti Puissant ssp. n. A, spectrogram and B, oscillogram (from top to bottom) of a succession of phrases composed of two echemes. C, oscillogram of a part of B showing two phrases and their component echeme, a one long echeme (E1) and a one short terminal echeme (E2). D, detailed spectrogram and E, oscillogram of a part of C showing a phrase with, for each echeme, a low amplitude part duration (LPD) and a shorter high amplitude part duration (HPD)." figureDoi="http://doi.org/10.5281/zenodo.8255321" httpUri="https://zenodo.org/record/8255321/files/figure.png" pageId="15" pageNumber="550">Fig. 11E</figureCitation>
) consists of a succession of groups of pulses arranged in two sub-groups: a low amplitude part duration (
<collectionCode id="F198DCD1FFB6FF94FE1FFF198E88FF7C" box="[426,482,187,213]" country="Argentina" lsid="urn:lsid:biocol.org:col:13082" name="Laboratorio de Botánica de la Dirección de Agricultura" pageId="15" pageNumber="550" type="Herbarium">LPD</collectionCode>
) and a shorter high amplitude part duration (
<collectionCode id="F198DCD1FFB6FF94FC7AFF198B67FF7C" box="[975,1037,187,213]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13184" name="Hampstead Scientific Society" pageId="15" pageNumber="550" type="Herbarium">HPD</collectionCode>
). Echemes E1 and E2 are regrouped together and compose a phrase of about 4.3 ± 
<geoCoordinate id="F2BD22D3FFB6FF94FD18FF7D8D88FF50" box="[685,738,223,249]" degrees="0.3" direction="south" orientation="latitude" pageId="15" pageNumber="550" precision="5555" value="-0.3">0.3 s</geoCoordinate>
(3.9–4.7; n = 6). During courtship behavior, fast wing-flicks preceding courtship song sequence are emitted. They produce audible clicks but were not recorded in this work.
</paragraph>
</subSubSection>
<subSubSection id="DF93179FFFB6FF94FF72FE848DA1FD64" pageId="15" pageNumber="550" type="distribution">
<paragraph id="97364414FFB6FF94FF72FE848DA1FD64" blockId="15.[151,1437,151,717]" pageId="15" pageNumber="550">
<emphasis id="A5FD9806FFB6FF94FF72FE848E85FEE9" bold="true" box="[199,495,294,321]" pageId="15" pageNumber="550">Distribution and habitat</emphasis>
(
<figureCitation id="0FB25891FFB6FF94FDB4FE858DE4FEE8" box="[513,654,295,321]" captionStart="FIGURE 14" captionStartId="18.[151,250,1938,1962]" captionTargetBox="[153,1435,416,1915]" captionTargetId="figure-153@18.[153,1435,416,1915]" captionTargetPageId="18" captionText="FIGURE 14. A, distribution map of T. q. quadrisignata (green and red dot) and T. q. pilleti Puissant ssp. n. (blue dot). B, distribution map of T. maldesi (black dot) and T. q. pilleti Puissant ssp. n. (blue dot). LC: locus typicus; RC: site of the recordings. Red dot represent stations that need to be confirmed (see explanation in text)." figureDoi="http://doi.org/10.5281/zenodo.8255327" httpUri="https://zenodo.org/record/8255327/files/figure.png" pageId="15" pageNumber="550">Figs. 14A,B</figureCitation>
: blue dot). 
<emphasis id="A5FD9806FFB6FF94FCA7FE858CE1FEE8" box="[786,907,295,321]" italics="true" pageId="15" pageNumber="550">T. q. pilleti</emphasis>
is currently known only from 
<collectingCountry id="EF9E0484FFB6FF94FB45FE858A32FEE8" box="[1264,1368,295,321]" name="Morocco" pageId="15" pageNumber="550">Morocco</collectingCountry>
, from five localities. Its geographical domain is located at the south of the nominal subspecies 
<emphasis id="A5FD9806FFB6FF94FB20FEE98A1AFECC" box="[1173,1392,331,357]" italics="true" pageId="15" pageNumber="550">T. q. quadrisagnata</emphasis>
, on the other side of the Strait of 
<collectingCountry id="EF9E0484FFB6FF94FE5DFECD8D25FE20" box="[488,591,367,393]" name="Gibraltar" pageId="15" pageNumber="550">Gibraltar</collectingCountry>
, Mediterranean Sea. Whereas 
<emphasis id="A5FD9806FFB6FF94FC1CFECD8B4BFE20" box="[937,1057,367,393]" italics="true" pageId="15" pageNumber="550">T. q. pilleti</emphasis>
has a restricted area distribution, 
<emphasis id="A5FD9806FFB6FF94FF22FE318E06FE04" box="[151,364,403,429]" italics="true" pageId="15" pageNumber="550">
T. q. 
<taxonomicName id="50893F97FFB6FF94FF7AFE318E06FE04" baseAuthorityName="Hagen" baseAuthorityYear="1855" box="[207,364,403,429]" class="Insecta" family="Cicadidae" genus="Tibicina" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="550" phylum="Arthropoda" rank="species" species="quadrisignata">quadrisignata</taxonomicName>
</emphasis>
is widely distributed throughout Western Europe and is known from 
<collectingCountry id="EF9E0484FFB6FF94FB23FE318B8FFE04" box="[1174,1253,403,429]" name="France" pageId="15" pageNumber="550">France</collectingCountry>
, 
<collectingCountry id="EF9E0484FFB6FF94FB41FE318A42FE04" box="[1268,1320,403,429]" name="Italy" pageId="15" pageNumber="550">Italy</collectingCountry>
, 
<collectingCountry id="EF9E0484FFB6FF94FA83FE318AF2FE04" box="[1334,1432,403,429]" name="Portugal" pageId="15" pageNumber="550">Portugal</collectingCountry>
, 
<collectingCountry id="EF9E0484FFB6FF94FF22FE158FBDFE78" box="[151,215,439,465]" name="Spain" pageId="15" pageNumber="550">Spain</collectingCountry>
and 
<collectingCountry id="EF9E0484FFB6FF94FEA4FE158EF0FE78" box="[273,410,439,465]" name="Switzerland" pageId="15" pageNumber="550">Switzerland</collectingCountry>
. In 
<collectingCountry id="EF9E0484FFB6FF94FE72FE158D45FE78" box="[455,559,439,465]" name="Morocco" pageId="15" pageNumber="550">Morocco</collectingCountry>
, 
<emphasis id="A5FD9806FFB6FF94FD89FE158DDEFE78" box="[572,692,439,465]" italics="true" pageId="15" pageNumber="550">T. q. pilleti</emphasis>
is present in the Mediterranean and sub-Mediterranean domains: SA3 stage, xeric semiarid class, temperate sub-class at 
<quantity id="5071E9F1FFB6FF94FCA3FE788C01FE5C" box="[790,875,474,501]" metricMagnitude="3" metricUnit="m" metricValue="2.025" pageId="15" pageNumber="550" unit="m" value="2025.0">2025 m</quantity>
in sagebrush steppe; SH2 stage, xeric subhumid class, warm sub-class, at 
<quantity id="5071E9F1FFB6FF94FE05FE5C8D44FDB0" box="[432,558,510,537]" metricMagnitude="2" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="15" pageNumber="550" unit="m" value="600.0" valueMax="700.0" valueMin="500.0">500–700 m</quantity>
above sea level; SX3 and SX4 stage, sub-axeric class, temperate and cool sub-class, at 
<quantity id="5071E9F1FFB6FF94FF43FD818E27FD94" box="[246,333,547,573]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="15" pageNumber="550" unit="m" value="1600.0">1600 m</quantity>
, 
<quantity id="5071E9F1FFB6FF94FEEDFD808EC7FD94" box="[344,429,546,573]" metricMagnitude="3" metricUnit="m" metricValue="1.655" pageId="15" pageNumber="550" unit="m" value="1655.0">1655 m</quantity>
and 
<quantity id="5071E9F1FFB6FF94FE56FD808D52FD94" box="[483,568,546,573]" metricMagnitude="3" metricUnit="m" metricValue="2.135" pageId="15" pageNumber="550" unit="m" value="2135.0">2135 m</quantity>
above sea level. The natural environment of the 
<emphasis id="A5FD9806FFB6FF94FBE3FD818B8DFD95" box="[1110,1255,547,573]" italics="true" pageId="15" pageNumber="550">locus typicus</emphasis>
is characterized by low but woody vegetation with a few isolated trees (
<figureCitation id="0FB25891FFB6FF94FC94FDE58C0FFDC8" box="[801,869,583,609]" captionStart="FIGURE 7" captionStartId="7.[151,250,2011,2035]" captionTargetBox="[151,1436,1129,1987]" captionTargetId="figure-67@7.[151,1436,1129,1987]" captionTargetPageId="7" captionText="FIGURE 7. Typical physical structure of vegetation where specimen of T. q. pilleti Puissant ssp. n. can be very numerous." figureDoi="http://doi.org/10.5281/zenodo.8255313" httpUri="https://zenodo.org/record/8255313/files/figure.png" pageId="15" pageNumber="550">Fig. 7</figureCitation>
). Physical structures of the vegetation inhabit by 
<emphasis id="A5FD9806FFB6FF94FF22FDC98E7AFD2C" box="[151,272,619,645]" italics="true" pageId="15" pageNumber="550">T. q. pilleti</emphasis>
in its area are scrub, sagebrush steppe and open wood of Aleppo pines and cedars with at least 40% of woody plants: habitat classes 4 to 8. For ecology, vegetation classes and phytoclimatic stages of 
<emphasis id="A5FD9806FFB6FF94FB74FD2D8AFFFD00" box="[1217,1429,655,681]" italics="true" pageId="15" pageNumber="550">
T. q. 
<taxonomicName id="50893F97FFB6FF94FB4DFD2D8AFFFD00" baseAuthorityName="Hagen" baseAuthorityYear="1855" box="[1272,1429,655,681]" class="Insecta" family="Cicadidae" genus="Tibicina" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="550" phylum="Arthropoda" rank="species" species="quadrisignata">quadrisignata</taxonomicName>
</emphasis>
, see 
<bibRefCitation id="F31839E5FFB6FF94FF77FD118E1BFD64" author="Puissant, S." box="[194,369,691,717]" pageId="15" pageNumber="550" refId="ref13943" refString="Puissant, S. (2006) Contribution a la connaissance des cigales de France: geonemie et ecologie des populations (Hemiptera, Cicadidae). ASCETE, Bedeilhac, 193 pp." type="book" year="2006">Puissant (2006)</bibRefCitation>
and 
<bibRefCitation id="F31839E5FFB6FF94FE12FD118DACFD64" author="Hertach, T. &amp; Gerber, S." box="[423,710,691,717]" pageId="15" pageNumber="550" pagination="7 - 25" refId="ref13105" refString="Hertach, T. &amp; Gerber, S. (2020) Schriller Larm im Fokus: Neues zu Verbreitung, Okologie und Schutz von Tibicina steveni und T. quadrisignata im Wallis (Cicadidae). Bulletin de la Murithienne, 137, 7 - 25." type="journal article" year="2020">Hertach &amp; Gerber (2020)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>