300 lines
37 KiB
XML
300 lines
37 KiB
XML
<document id="D737FA491424B410F8A39A6692821F59" ID-DOI="10.11646/zootaxa.3957.3.1" ID-GBIF-Dataset="31f8999c-e65b-4e76-93be-09f17c90bffe" ID-ISSN="1175-5326" ID-Zenodo-Dep="233401" ID-ZooBank="6CC62878-DBBD-4478-AAF2-9403B46C8C36" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461061343700" checkinUser="plazi" docAuthor="Zhao, Zong-Yi & Zhou, Hong-Zhang" docDate="2015" docId="A81387A57616204AF9AAFB5714FB4EF5" docLanguage="en" docName="zt03957p276.pdf" docOrigin="Zootaxa 3957 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Quedius" docType="treatment" docVersion="10" lastPageNumber="273" masterDocId="542AFFDD7602205CF93DFF8E16684D51" masterDocTitle="Phylogeny and taxonomic revision of the subgenus Velleius Leach (Coleoptera: Staphylinidae: Staphylininae)" masterLastPageNumber="276" masterPageNumber="251" pageNumber="271" updateTime="1698582440283" updateUser="plazi">
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<mods:titleInfo id="DD42DE27092F94259245A0C6A31DC3C5">
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<mods:title id="32B46C15761DEFBE47D0405C3E95EC98">Phylogeny and taxonomic revision of the subgenus Velleius Leach (Coleoptera: Staphylinidae: Staphylininae)</mods:title>
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<mods:namePart id="79F7C65FD5754609B4CE9D9070AC5130">Zhao, Zong-Yi</mods:namePart>
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<mods:namePart id="F10DC2346587642A96613559BF412D4A">Zhou, Hong-Zhang</mods:namePart>
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<mods:title id="4474C810C93BC01DC4D93A73E4EF9CC7">Zootaxa</mods:title>
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<mods:part id="E70F1B16022D3FE155831AB499ACA4DD">
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<mods:date id="32C618AB424AF0057FEA706EF0EE5023">2015</mods:date>
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<mods:number id="FEE34D157A8CFD68A58F8DFE3EE7AE9E">3957</mods:number>
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<treatment id="A81387A57616204AF9AAFB5714FB4EF5" ID-DOI="http://doi.org/10.5281/zenodo.5663293" ID-GBIF-Taxon="119614165" ID-Zenodo-Dep="5663293" LSID="urn:lsid:plazi:treatment:A81387A57616204AF9AAFB5714FB4EF5" httpUri="http://treatment.plazi.org/id/A81387A57616204AF9AAFB5714FB4EF5" lastPageId="22" lastPageNumber="273" pageId="20" pageNumber="271">
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<subSubSection id="68A0653876162048F9AAFB5714D649A2" box="[151,702,1241,1267]" pageId="20" pageNumber="271" type="nomenclature">
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<paragraph id="200536B376162048F9AAFB5714D649A2" blockId="20.[151,702,1241,1301]" box="[151,702,1241,1267]" pageId="20" pageNumber="271">
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<heading id="7B4D81DF76162048F9AAFB5714D649A2" bold="true" box="[151,702,1241,1267]" fontSize="11" level="1" pageId="20" pageNumber="271" reason="1">
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<emphasis id="12CEEAA176162048F9AAFB5714D649A2" bold="true" box="[151,702,1241,1267]" pageId="20" pageNumber="271">
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8.
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<taxonomicName id="E7BA4D3076162048F989FB57177149A2" ID-CoL="8VS2M" box="[180,281,1241,1267]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="271" phylum="Arthropoda" rank="genus">
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<emphasis id="12CEEAA176162048F989FB57177149A2" bold="true" box="[180,281,1241,1267]" italics="true" pageId="20" pageNumber="271">Quedius</emphasis>
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</taxonomicName>
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(
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<emphasis id="12CEEAA176162048F817FB5717ED49A2" bold="true" box="[298,389,1241,1267]" italics="true" pageId="20" pageNumber="271">Ve l le iu s</emphasis>
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)
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<taxonomicName id="E7BA4D3076162048F8A7FB5714D649A2" authority="Sharp 1874" authorityName="Sharp" authorityYear="1874" box="[410,702,1241,1267]" class="Insecta" family="Staphylinidae" genus="Velleius" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="271" phylum="Arthropoda" rank="species" species="pectinatus">
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<emphasis id="12CEEAA176162048F8A7FB57147E49A2" bold="true" box="[410,534,1241,1267]" italics="true" pageId="20" pageNumber="271">pectinatus</emphasis>
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(
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<bibRefCitation id="442B4B4276162048FB18FB5714DE49A2" author="Sharp" box="[549,694,1241,1267]" pageId="20" pageNumber="271" refString="Sharp, D. S. (1874) The Staphylinidae of Japan. Transactions of the Entomological Society of London, 1874, 1 - 103. http: // dx. doi. org / 10.1111 / j. 1365 - 2311.1874. tb 00159. x" type="journal article" year="1874">Sharp 1874</bibRefCitation>
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)
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="68A065387616204AF9AAFB7212004CD9" lastPageId="22" lastPageNumber="273" pageId="20" pageNumber="271" type="description">
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<paragraph id="200536B376162048F9AAFB7217684845" blockId="20.[151,702,1241,1301]" box="[151,256,1276,1301]" pageId="20" pageNumber="271">
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(
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<figureCitation id="B8812A3676162048F9A2FB72169B4845" box="[159,243,1276,1301]" captionStart="FIGURE 12. Q" captionStartId="21.[151,250,1626,1648]" captionTargetBox="[213,1366,706,1595]" captionTargetId="figure@21.[206,1381,697,1604]" captionTargetPageId="21" captionText="FIGURE 12. Q. (V.) pectinatus. A – C, aedeagus, (A) lateral view, (B) dorsal view, (C) underside of paramere, showing sensory peg setae; D – E, mandibles, dorsal view; F, male abdominal sternite IX; G, male abdominal tergite X; H, female abdominal tergite X. Scales = 0.5 mm." httpUri="https://zenodo.org/record/233413/files/figure.png" pageId="20" pageNumber="271">Fig. 12</figureCitation>
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).
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</paragraph>
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<paragraph id="200536B376162048F9AAFAB3137148C1" blockId="20.[151,1436,1341,1424]" pageId="20" pageNumber="271">
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<bibRefCitation id="442B4B4276162048F9AAFAB317794802" author="Sharp" box="[151,273,1341,1363]" pageId="20" pageNumber="271" refString="Sharp, D. S. (1874) The Staphylinidae of Japan. Transactions of the Entomological Society of London, 1874, 1 - 103. http: // dx. doi. org / 10.1111 / j. 1365 - 2311.1874. tb 00159. x" type="journal article" year="1874">Sharp 1874</bibRefCitation>
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: 24 (originally in
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<taxonomicName id="E7BA4D3076162048F8F2FAB314764802" box="[463,542,1341,1363]" class="Insecta" family="Staphylinidae" genus="Velleius" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="271" phylum="Arthropoda" rank="genus">
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<emphasis id="12CEEAA176162048F8F2FAB314764802" box="[463,542,1341,1363]" italics="true" pageId="20" pageNumber="271">Velleius</emphasis>
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</taxonomicName>
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;
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<typeStatus id="FF01881176162048FB10FAB014374802" box="[557,607,1342,1363]" pageId="20" pageNumber="271">Type</typeStatus>
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locality:
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<collectingCountry id="58AD762376162048FBF9FAB0156A4802" box="[708,770,1342,1363]" name="Japan" pageId="20" pageNumber="271">Japan</collectingCountry>
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: Hiogo; Nagasaki);
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<bibRefCitation id="442B4B4276162048FAEFFAB312084802" author="Gridelli" box="[978,1120,1341,1363]" pageId="20" pageNumber="271" refString="Gridelli, E. (1932) Studi sul genere Quedius Steph. (Coleopt. Staphyl.). 4 o contributo. Bollettino della Societa Entomologica Italiana, 64, 14 - 20." type="journal article" year="1932">Gridelli 1932</bibRefCitation>
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: 18;
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<bibRefCitation id="442B4B4276162048FDA8FAB313494802" author="Nakane" box="[1173,1313,1341,1363]" pageId="20" pageNumber="271" refString="Nakane, T. (1963) Staphylinidae. In: Nakane, T., Ohbayashi, K., Nomura, S. & Kurosawa, Y. (Eds.), Iconographia Insectorum Japonicorum, Colore naturali edita. Vol. 2. Hokuryukan, Tokyo, pp. 81 - 100." type="book" year="1963">Nakane 1963</bibRefCitation>
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: 94;
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<bibRefCitation id="442B4B4276162048FC6AFAB316954823" author="Naomi" pageId="20" pageNumber="271" refString="Naomi, S. I. (1986) Taxonomic study on the genus Velleius Mannerheim of Japan, with description of a new species (Coleoptera: Staphylinidae). Transactions of the Shikoku Entomological Society, 17 (4), 239 - 246." type="journal article" year="1986">Naomi 1986</bibRefCitation>
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: 244;
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<bibRefCitation id="442B4B4276162048F807FAD217A64823" author="Smetana" box="[314,462,1372,1394]" pageId="20" pageNumber="271" refString="Smetana, A. (1995) Revision of the tribes Quediini and Tanygnathinini. Part III. Taiwan. (Coleoptera: Staphylinidae). National Museum of Natural Science, Taichung, Taiwan. Special Publication, 6, 1 - 145." type="journal article" year="1995">Smetana 1995</bibRefCitation>
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: 110;
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<bibRefCitation id="442B4B4276162048FB37FAD214C14823" author="Watanabe" box="[522,681,1372,1394]" pageId="20" pageNumber="271" refString="Watanabe, Y. (1990) A revision of the Japanese species of the genus Velleius (Coleoptera, Staphylinidae). Elytra, 18 (1), 59 - 72." type="journal article" year="1990">Watanabe 1990</bibRefCitation>
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: 63;
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<bibRefCitation id="442B4B4276162048FBE7FAD215294823" author="Cho" box="[730,833,1372,1394]" pageId="20" pageNumber="271" refString="Cho, Y. B. (1996) Taxonomic study on the tribe Quediini (Coleoptera, Staphylinidae) from Korea. The Korean Journal of Systematic Zoology, 12 (2), 107 - 119." type="journal article" year="1996">Cho 1996</bibRefCitation>
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: 114;
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<bibRefCitation id="442B4B4276162048FA40FAD312634823" author="Herman" box="[893,1035,1372,1394]" pageId="20" pageNumber="271" refString="Herman, L. H. (2001) Catalog of the Staphylinidae (Insecta: Coleoptera) 1758 to the end of the second millennium. Parts I-VII. Bulletin of the American Museum of Natural History, 265, 1 - 4218. http: // dx. doi. org / 10.1206 / 0003 - 0090 (2001) 264 % 3 C 0003: NCITSI % 3 E 2.0. CO; 2" type="journal article" year="2001">Herman 2001</bibRefCitation>
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||
: 3307 (complete list of references).
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<emphasis id="12CEEAA176162048F9AAFAF4177948C1" bold="true" box="[151,273,1402,1424]" pageId="20" pageNumber="271">Synonymy:</emphasis>
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<taxonomicName id="E7BA4D3076162048F82AFAF4141248C1" authority="Fairmaire 1891" authorityName="Fairmaire" authorityYear="1891" box="[279,634,1402,1424]" class="Insecta" family="Staphylinidae" genus="Velleius" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="271" phylum="Arthropoda" rank="species" species="simillimus">
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<emphasis id="12CEEAA176162048F82AFAF417BC48C1" box="[279,468,1402,1424]" italics="true" pageId="20" pageNumber="271">Velleius simillimus</emphasis>
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<bibRefCitation id="442B4B4276162048F8E6FAF4141248C1" author="Fairmaire" box="[475,634,1402,1424]" pageId="20" pageNumber="271" refString="Fairmaire, L. (1891) Coleopteres de l'interieur de la Chine (suite: 7 e partie). Annales de la Societe Entomologique de Belgique, 35, clxxxviii - ccxix." type="journal volume" year="1891">Fairmaire 1891</bibRefCitation>
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</taxonomicName>
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: cxci;
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<bibRefCitation id="442B4B4276162048FB86FAF5152248C1" author="Herman" box="[699,842,1402,1424]" pageId="20" pageNumber="271" refString="Herman, L. H. (2001) Catalog of the Staphylinidae (Insecta: Coleoptera) 1758 to the end of the second millennium. Parts I-VII. Bulletin of the American Museum of Natural History, 265, 1 - 4218. http: // dx. doi. org / 10.1206 / 0003 - 0090 (2001) 264 % 3 C 0003: NCITSI % 3 E 2.0. CO; 2" type="journal article" year="2001">Herman 2001</bibRefCitation>
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||
: 3307 (complete list of references).
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<emphasis id="12CEEAA176162048FD88FAF4137148C1" bold="true" box="[1205,1305,1402,1424]" pageId="20" pageNumber="271">syn. nov..</emphasis>
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</paragraph>
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<paragraph id="200536B376162048F9AAFA38171748A2" blockId="20.[151,1436,1462,2023]" pageId="20" pageNumber="271">
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<emphasis id="12CEEAA176162048F9AAFA38172C489E" bold="true" box="[151,324,1462,1487]" pageId="20" pageNumber="271">Redescription.</emphasis>
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Body length about 20.0 mm, body width 5.0 mm (EW), HW/PW/EW/AW = 1.0:1.5: 1.6:1.4, HL/ PL/EL = 1.0:1.5:1.9.
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</paragraph>
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<paragraph id="200536B376162048F9FAFA73122A4B6B" blockId="20.[151,1436,1462,2023]" pageId="20" pageNumber="271">Head dark brown, pronotum and elytra reddish-brown, abdomen dark brown, antennae and all legs dark brown, maxillary and labial palpi with all segments reddish-brown and apices paler.</paragraph>
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<paragraph id="200536B376162048F9FAF9CB15204A40" blockId="20.[151,1436,1462,2023]" pageId="20" pageNumber="271">Head nearly round, wider than long, HW/HL = 1.0:0.8, eyes large and longer than tempora in dorsal view, HEL/HTL = 1.0:0.5; posterior frontal setiferous puncture positioned at the same level of posterior edge of eye and less than one diameter separated from posteromedial edge of eye, one basal setiferous puncture on each side and smaller than posterior frontal setiferous puncture, temporal setiferous puncture slightly closer to eye than to nuchal constriction; no oblique fovea behind insertion of antenna, dorsal surface covered with sparse fine punctures and very dense microsculpture consisting of isodiametric meshes.</paragraph>
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<paragraph id="200536B376162048F9FAF89212D94A09" blockId="20.[151,1436,1462,2023]" pageId="20" pageNumber="271">Antennae with apex of segment III wider than that of segment I, segment IV–XI each deeply bifurcated, segment III longer than segment II, last segment longer than 3 preceding segments combined.</paragraph>
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<paragraph id="200536B376162048F9FAF8ED12A54AF1" blockId="20.[151,1436,1462,2023]" pageId="20" pageNumber="271">
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Left mandible (
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<figureCitation id="B8812A3676162048F84AF8ED17A04A2D" box="[375,456,1891,1916]" captionStart="FIGURE 12. Q" captionStartId="21.[151,250,1626,1648]" captionTargetBox="[213,1366,706,1595]" captionTargetId="figure@21.[206,1381,697,1604]" captionTargetPageId="21" captionText="FIGURE 12. Q. (V.) pectinatus. A – C, aedeagus, (A) lateral view, (B) dorsal view, (C) underside of paramere, showing sensory peg setae; D – E, mandibles, dorsal view; F, male abdominal sternite IX; G, male abdominal tergite X; H, female abdominal tergite X. Scales = 0.5 mm." httpUri="https://zenodo.org/record/233413/files/figure.png" pageId="20" pageNumber="271">Fig. 12</figureCitation>
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D) with anterior tooth divided into two subteeth with posterior one larger, edge before anterior tooth smoothly curved, posterior tooth absent, right mandible (
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<figureCitation id="B8812A3676162048FA8EF809126B4ACE" box="[947,1027,1927,1952]" captionStart="FIGURE 12. Q" captionStartId="21.[151,250,1626,1648]" captionTargetBox="[213,1366,706,1595]" captionTargetId="figure@21.[206,1381,697,1604]" captionTargetPageId="21" captionText="FIGURE 12. Q. (V.) pectinatus. A – C, aedeagus, (A) lateral view, (B) dorsal view, (C) underside of paramere, showing sensory peg setae; D – E, mandibles, dorsal view; F, male abdominal sternite IX; G, male abdominal tergite X; H, female abdominal tergite X. Scales = 0.5 mm." httpUri="https://zenodo.org/record/233413/files/figure.png" pageId="20" pageNumber="271">Fig. 12</figureCitation>
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E) with two teeth.
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</paragraph>
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<paragraph id="200536B376162048F9FAF824131F4AB6" blockId="20.[151,1436,1462,2023]" pageId="20" pageNumber="271">Maxillary palpus with last segment lacking setae and nearly parallel-sided, surface covered with longitudinal narrow foveae on basal portion and many pores on apical portion, ratio of length of segments II–IV 1.0:0.9:1.0.</paragraph>
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<paragraph id="200536B376172049F9FAFF1917A94D85" blockId="21.[151,1436,151,645]" pageId="21" pageNumber="272">Labial palpus with last segment lacking setae and fusiform, surface covered with many pores, ratio of length of segments I–III 1.0:1.2:1.2.</paragraph>
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<paragraph id="200536B376172049F9FAFF5113194DA9" blockId="21.[151,1436,151,645]" box="[199,1393,223,248]" pageId="21" pageNumber="272">Neck surface covered with dense microsculpture consisting of meshes more transverse than those on head.</paragraph>
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<paragraph id="200536B376172049F9FAFE8B13594CD9" blockId="21.[151,1436,151,645]" pageId="21" pageNumber="272">Pronotum wider than long, PW/PL = 1.0:0.8; two setiferous punctures in each dorsal row with anterior puncture much smaller than posterior one, sometimes a small puncture accompanying the posterior puncture, one setiferous puncture in each sublateral row, large lateral setiferous puncture behind sublateral row but before posterior puncture in dorsal row; dorsal surface covered with dense and vague transverse microsculpture.</paragraph>
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<paragraph id="200536B376172049F9FAFE1A12C34C81" blockId="21.[151,1436,151,645]" pageId="21" pageNumber="272">Scutellum setose on apical half, surface covered with dense but shallow transverse microsculpture, anterior basal ridge with middle portion nearly straight, posterior basal ridge slightly arced backward.</paragraph>
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<paragraph id="200536B376172049F9FAFE5214D14F46" blockId="21.[151,1436,151,645]" pageId="21" pageNumber="272">Elytra. EW/EL = 1.0:0.9, EL/ESL = 1.0:0.5, evenly and more densely setose than scutellum, lateral face and posterior margin bearing many large black setae.</paragraph>
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<paragraph id="200536B376172049F9FAFDAA12F34F31" blockId="21.[151,1436,151,645]" pageId="21" pageNumber="272">Abdomen with each tergite evenly and densely setose, tergite III–VII bearing large middle lateral setae, tergite VII bearing whitish apical fringe, tergite VIII bearing many large black setae on apical half.</paragraph>
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<paragraph id="200536B376172049F9FAFDE214B84FD4" blockId="21.[151,1436,151,645]" box="[199,720,620,645]" pageId="21" pageNumber="272">Apices of meso- and metatarsomere V dilated.</paragraph>
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<caption id="74C5663B76172049F9AAF9D417C44BFC" httpUri="https://zenodo.org/record/233413/files/figure.png" pageId="21" pageNumber="272" targetBox="[213,1366,706,1595]" targetPageId="21">
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<paragraph id="200536B376172049F9AAF9D417C44BFC" blockId="21.[151,1436,1626,1709]" pageId="21" pageNumber="272">
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<emphasis id="12CEEAA176172049F9AAF9D417774B21" bold="true" box="[151,287,1626,1648]" pageId="21" pageNumber="272">FIGURE 12.</emphasis>
|
||
<taxonomicName id="E7BA4D3076172049F818F9D417BE4B21" box="[293,470,1626,1648]" class="Insecta" family="Staphylinidae" genus="Velleius" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="272" phylum="Arthropoda" rank="species" species="pectinatus" subGenus="Velleius">
|
||
<emphasis id="12CEEAA176172049F818F9D417564B3E" box="[293,318,1626,1647]" italics="true" pageId="21" pageNumber="272">Q.</emphasis>
|
||
(
|
||
<emphasis id="12CEEAA176172049F870F9D517364B21" box="[333,350,1627,1648]" italics="true" pageId="21" pageNumber="272">V.</emphasis>
|
||
)
|
||
<emphasis id="12CEEAA176172049F853F9D517BE4B21" box="[366,470,1627,1648]" italics="true" pageId="21" pageNumber="272">pectinatus</emphasis>
|
||
</taxonomicName>
|
||
. A–C, aedeagus, (A) lateral view, (B) dorsal view, (C) underside of paramere, showing sensory peg setae; D–E, mandibles, dorsal view; F, male abdominal sternite IX; G, male abdominal tergite X; H, female abdominal tergite X. Scales = 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="200536B37617204AF9FAF95A14674C4D" blockId="21.[151,1436,1748,2025]" lastBlockId="22.[151,1437,151,933]" lastPageId="22" lastPageNumber="273" pageId="21" pageNumber="272">
|
||
Male. Sternite VIII with a middle emargination on apical margin, a small hairless area before it; sternite IX (
|
||
<figureCitation id="B8812A3676172049F9A2F976169B4A40" box="[159,243,1784,1809]" captionStart="FIGURE 12. Q" captionStartId="21.[151,250,1626,1648]" captionTargetBox="[213,1366,706,1595]" captionTargetId="figure@21.[206,1381,697,1604]" captionTargetPageId="21" captionText="FIGURE 12. Q. (V.) pectinatus. A – C, aedeagus, (A) lateral view, (B) dorsal view, (C) underside of paramere, showing sensory peg setae; D – E, mandibles, dorsal view; F, male abdominal sternite IX; G, male abdominal tergite X; H, female abdominal tergite X. Scales = 0.5 mm." httpUri="https://zenodo.org/record/233413/files/figure.png" pageId="21" pageNumber="272">Fig. 12</figureCitation>
|
||
F) with basal portion very narrow and long, apical portion with a small middle emargination on apical margin, surface from widest portion to apex covered with dense setae gradually becoming longer backwards; tergite X (
|
||
<figureCitation id="B8812A3676172049F837F8CE17334A08" box="[266,347,1856,1881]" captionStart="FIGURE 12. Q" captionStartId="21.[151,250,1626,1648]" captionTargetBox="[213,1366,706,1595]" captionTargetId="figure@21.[206,1381,697,1604]" captionTargetPageId="21" captionText="FIGURE 12. Q. (V.) pectinatus. A – C, aedeagus, (A) lateral view, (B) dorsal view, (C) underside of paramere, showing sensory peg setae; D – E, mandibles, dorsal view; F, male abdominal sternite IX; G, male abdominal tergite X; H, female abdominal tergite X. Scales = 0.5 mm." httpUri="https://zenodo.org/record/233413/files/figure.png" pageId="21" pageNumber="272">Fig. 12</figureCitation>
|
||
G) with basal edge smoothly and broadly arcuated backwards, apical portion triangle-shaped but with apex truncated, surface of about apical 1/3 covered with many long setae; aedeagus in lateral view (
|
||
<figureCitation id="B8812A3676172049FC13F8EA13164A2C" box="[1326,1406,1892,1917]" captionStart="FIGURE 12. Q" captionStartId="21.[151,250,1626,1648]" captionTargetBox="[213,1366,706,1595]" captionTargetId="figure@21.[206,1381,697,1604]" captionTargetPageId="21" captionText="FIGURE 12. Q. (V.) pectinatus. A – C, aedeagus, (A) lateral view, (B) dorsal view, (C) underside of paramere, showing sensory peg setae; D – E, mandibles, dorsal view; F, male abdominal sternite IX; G, male abdominal tergite X; H, female abdominal tergite X. Scales = 0.5 mm." httpUri="https://zenodo.org/record/233413/files/figure.png" pageId="21" pageNumber="272">Fig. 12</figureCitation>
|
||
A) with apex of paramere protruding slightly beyond apex of median lobe, paramere gradually becoming wider from apex to base, not suddenly widened below apex, median lobe concaved below apex on side facing paramere, in dorsal view (
|
||
<figureCitation id="B8812A3676172049F817F85E17144AB8" box="[298,380,2000,2025]" captionStart="FIGURE 12. Q" captionStartId="21.[151,250,1626,1648]" captionTargetBox="[213,1366,706,1595]" captionTargetId="figure@21.[206,1381,697,1604]" captionTargetPageId="21" captionText="FIGURE 12. Q. (V.) pectinatus. A – C, aedeagus, (A) lateral view, (B) dorsal view, (C) underside of paramere, showing sensory peg setae; D – E, mandibles, dorsal view; F, male abdominal sternite IX; G, male abdominal tergite X; H, female abdominal tergite X. Scales = 0.5 mm." httpUri="https://zenodo.org/record/233413/files/figure.png" pageId="21" pageNumber="272">Fig. 12</figureCitation>
|
||
B) paramere gradually becoming wider from apex to base and narrower than median lobe for most length, apex of median lobe obtuse, apex of paramere (
|
||
<figureCitation id="B8812A367614204AFA79FF1915FD4DE1" box="[836,917,151,176]" captionStart="FIGURE 12. Q" captionStartId="21.[151,250,1626,1648]" captionTargetBox="[213,1366,706,1595]" captionTargetId="figure@21.[206,1381,697,1604]" captionTargetPageId="21" captionText="FIGURE 12. Q. (V.) pectinatus. A – C, aedeagus, (A) lateral view, (B) dorsal view, (C) underside of paramere, showing sensory peg setae; D – E, mandibles, dorsal view; F, male abdominal sternite IX; G, male abdominal tergite X; H, female abdominal tergite X. Scales = 0.5 mm." httpUri="https://zenodo.org/record/233413/files/figure.png" pageId="22" pageNumber="273">Fig. 12</figureCitation>
|
||
C) split in to two branches by a middle suture but the two branches still adjacent to each other, 2 long apical and 2 smaller lateral setae on each side, underside of paramere covered with many sensory peg setae forming two longitudinal groups extending from apex towards base beyond insertions of lateral setae.
|
||
</paragraph>
|
||
<paragraph id="200536B37614204AF9FAFEA912004CD9" blockId="22.[151,1437,151,933]" pageId="22" pageNumber="273">
|
||
Female. Tergite X (
|
||
<figureCitation id="B8812A367614204AF8A2FEA917874C11" box="[415,495,295,320]" captionStart="FIGURE 12. Q" captionStartId="21.[151,250,1626,1648]" captionTargetBox="[213,1366,706,1595]" captionTargetId="figure@21.[206,1381,697,1604]" captionTargetPageId="21" captionText="FIGURE 12. Q. (V.) pectinatus. A – C, aedeagus, (A) lateral view, (B) dorsal view, (C) underside of paramere, showing sensory peg setae; D – E, mandibles, dorsal view; F, male abdominal sternite IX; G, male abdominal tergite X; H, female abdominal tergite X. Scales = 0.5 mm." httpUri="https://zenodo.org/record/233413/files/figure.png" pageId="22" pageNumber="273">Fig. 12</figureCitation>
|
||
H) with basal side broadly and smoothly arcuated backwards, apical portion trilobed with two lateral lobes smaller than the middle lobe, apical portion of middle lobe triangle-shaped but with apex truncated, apical margin bearing a row of long setae, several smaller setae before them.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="68A065387614204AF9FAFE1A14584E0C" pageId="22" pageNumber="273" type="materials_examined">
|
||
<paragraph id="200536B37614204AF9FAFE1A15444F6D" blockId="22.[151,1437,151,933]" pageId="22" pageNumber="273">
|
||
<emphasis id="12CEEAA17614204AF9FAFE1A17954CFC" bold="true" box="[199,509,404,429]" pageId="22" pageNumber="273">
|
||
<typeStatus id="FF0188117614204AF9FAFE1A176A4CFC" box="[199,258,404,429]" pageId="22" pageNumber="273">Type</typeStatus>
|
||
materials.
|
||
<typeStatus id="FF0188117614204AF8B7FE1A17914CFC" box="[394,505,404,429]" pageId="22" pageNumber="273" type="holotype">Holotype</typeStatus>
|
||
,
|
||
</emphasis>
|
||
male,
|
||
<collectingCountry id="58AD76237614204AFB70FE1B14F84CFD" box="[589,656,405,428]" name="Japan" pageId="22" pageNumber="273">Japan</collectingCountry>
|
||
, collected by G. Lewis, deposited in NHM.
|
||
<emphasis id="12CEEAA17614204AFDBAFE1A12914CFC" bold="true" box="[1159,1273,404,429]" pageId="22" pageNumber="273">
|
||
<typeStatus id="FF0188117614204AFDBAFE1A129C4CFC" box="[1159,1268,404,429]" pageId="22" pageNumber="273" type="syntype">Syntypes</typeStatus>
|
||
,
|
||
</emphasis>
|
||
<specimenCount id="36BCFD3A7614204AFC3FFE1A13354CFC" box="[1282,1373,404,429]" pageId="22" pageNumber="273" type="male">2 males</specimenCount>
|
||
, with labels “Kiukiang,
|
||
<date id="540410737614204AF85DFE3617BD4C81" box="[352,469,440,464]" pageId="22" pageNumber="273" value="1887-05">May 1887</date>
|
||
, A. E. Pratt” / “
|
||
<typeStatus id="FF0188117614204AFBBDFE3614884C81" box="[640,736,440,464]" pageId="22" pageNumber="273" type="syntype">Syntype</typeStatus>
|
||
” / “Coll. et det. A. Fauvel,
|
||
<taxonomicName id="E7BA4D307614204AFD30FE3616BD4CA5" authority="Fairm., R. I. Sc. N. B." authorityName="Fairm., R. I. Sc. N. B." class="Insecta" family="Staphylinidae" genus="Velleius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="273" phylum="Arthropoda" rank="species" species="simillimus">
|
||
<emphasis id="12CEEAA17614204AFD30FE3612B64C81" box="[1037,1246,440,464]" italics="true" pageId="22" pageNumber="273">Velleius simillimus</emphasis>
|
||
, Fairm., R. I. Sc. N. B.
|
||
</taxonomicName>
|
||
17.479” / “Coll. et. det A. Fauvel,
|
||
<taxonomicName id="E7BA4D307614204AFB4FFE52123F4CA5" authority="Sharp., R. I. Sc. N. B." authorityName="Sharp., R. I. Sc. N. B." box="[626,1111,476,501]" class="Insecta" family="Staphylinidae" genus="Velleius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="273" phylum="Arthropoda" rank="species" species="pectinatus">
|
||
<emphasis id="12CEEAA17614204AFB4FFE52152E4CA5" box="[626,838,476,500]" italics="true" pageId="22" pageNumber="273">Velleius pectinatus</emphasis>
|
||
, Sharp., R. I. Sc. N. B.
|
||
</taxonomicName>
|
||
17.479”;
|
||
<specimenCount id="36BCFD3A7614204AFDF6FE5213744CA4" box="[1227,1308,476,501]" pageId="22" pageNumber="273" type="male">1 male</specimenCount>
|
||
, with label “Kiukiang,
|
||
<date id="540410737614204AF826FD8E17F94F49" box="[283,401,512,536]" pageId="22" pageNumber="273" value="1887-05">May 1887</date>
|
||
, A. E. Pratt” / “
|
||
<emphasis id="12CEEAA17614204AFB7BFD8E14D94F49" box="[582,689,512,536]" italics="true" pageId="22" pageNumber="273">simllimus</emphasis>
|
||
Fairm.
|
||
<typeStatus id="FF0188117614204AFA31FD8E15234F49" box="[780,843,512,536]" pageId="22" pageNumber="273">typus</typeStatus>
|
||
” / “
|
||
<collectingCountry id="58AD76237614204AFA45FD8E15A84F49" box="[888,960,512,536]" name="South Korea" pageId="22" pageNumber="273">Corée</collectingCountry>
|
||
, v. Heyden”/ “
|
||
<typeStatus id="FF0188117614204AFD58FD8E12AD4F49" box="[1125,1221,512,536]" pageId="22" pageNumber="273" type="syntype">Syntype</typeStatus>
|
||
” / “Coll. et. det A. Fauvel,
|
||
<taxonomicName id="E7BA4D307614204AF9CCFDAA14AC4F6D" authority="Sharp., R. I. Sc. N. B." authorityName="Sharp., R. I. Sc. N. B." box="[241,708,548,573]" class="Insecta" family="Staphylinidae" genus="Velleius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="273" phylum="Arthropoda" rank="species" species="pectinatus">
|
||
<emphasis id="12CEEAA17614204AF9CCFDAA17AA4F6D" box="[241,450,548,572]" italics="true" pageId="22" pageNumber="273">Velleius pectinatus</emphasis>
|
||
, Sharp., R. I. Sc. N. B.
|
||
</taxonomicName>
|
||
17.479”.
|
||
</paragraph>
|
||
<paragraph id="200536B37614204AF9FAFDC9130B4FA1" blockId="22.[151,1437,151,933]" pageId="22" pageNumber="273">
|
||
<emphasis id="12CEEAA17614204AF9FAFDC914234F31" bold="true" box="[199,587,583,608]" pageId="22" pageNumber="273">Additional specimens examined</emphasis>
|
||
.
|
||
<specimenCount id="36BCFD3A7614204AFB60FDC614D84F31" box="[605,688,583,608]" pageId="22" pageNumber="273" type="male">1 male</specimenCount>
|
||
,
|
||
<collectingCountry id="58AD76237614204AFBFCFDC9156E4F31" box="[705,774,583,608]" name="China" pageId="22" pageNumber="273">China</collectingCountry>
|
||
, Shanghai,
|
||
<date id="540410737614204AFAAFFDC6126E4F31" box="[914,1030,584,608]" pageId="22" pageNumber="273" value="1930-06-01">1.VI.1930</date>
|
||
, collected by A. Savio;
|
||
<specimenCount id="36BCFD3A7614204AFC16FDC913154F31" box="[1323,1405,583,608]" pageId="22" pageNumber="273" type="male">5 male</specimenCount>
|
||
,
|
||
<specimenCount id="36BCFD3A7614204AFCB3FDC6168E4FD4" pageId="22" pageNumber="273" type="female">1 female</specimenCount>
|
||
,
|
||
<collectingCountry id="58AD76237614204AF9C9FDE217524FD4" box="[244,314,620,645]" name="China" pageId="22" pageNumber="273">China</collectingCountry>
|
||
, Shanghai,
|
||
<date id="540410737614204AF8FCFDE214054FD4" box="[449,621,620,645]" pageId="22" pageNumber="273" value="1934-06-15" valueMax="1934-06-18" valueMin="1934-06-15">15–18.VI.1934</date>
|
||
, collected by O. Piel;
|
||
<specimenCount id="36BCFD3A7614204AFA49FDE215B34FD4" box="[884,987,620,645]" pageId="22" pageNumber="273" type="female">1 female</specimenCount>
|
||
,
|
||
<collectingCountry id="58AD76237614204AFAD7FDE212584FD4" box="[1002,1072,620,645]" name="China" pageId="22" pageNumber="273">China</collectingCountry>
|
||
, Haiyen,
|
||
<date id="540410737614204AFD9DFDE2134A4FD4" box="[1184,1314,620,645]" pageId="22" pageNumber="273" value="1935-06-17">17.VI.1935</date>
|
||
;
|
||
<specimenCount id="36BCFD3A7614204AFC0CFDE213F04FD4" box="[1329,1432,620,645]" pageId="22" pageNumber="273" type="female">1 female</specimenCount>
|
||
,
|
||
<collectingCountry id="58AD76237614204AF9AAFD0116B54FF9" box="[151,221,655,680]" name="China" pageId="22" pageNumber="273">China</collectingCountry>
|
||
, Emei Mountain,
|
||
<date id="540410737614204AF89AFD0114424FF9" box="[423,554,655,680]" pageId="22" pageNumber="273" value="1955-06-29">29.VI.1955</date>
|
||
, collected by Huangjin;
|
||
<specimenCount id="36BCFD3A7614204AFA02FD1E15E54FF9" box="[831,909,655,680]" pageId="22" pageNumber="273" type="male">1 male</specimenCount>
|
||
,
|
||
<specimenCount id="36BCFD3A7614204AFAA7FD1E15974FF9" box="[922,1023,655,680]" pageId="22" pageNumber="273" type="female">2 female</specimenCount>
|
||
,
|
||
<collectingCountry id="58AD76237614204AFD36FD0112394FF9" box="[1035,1105,655,680]" name="China" pageId="22" pageNumber="273">China</collectingCountry>
|
||
, Mokan Shan,
|
||
<date id="540410737614204AFDC6FD1E13FF4FF9" box="[1275,1431,656,680]" pageId="22" pageNumber="273" value="1937-06-09" valueMax="1937-06-11" valueMin="1937-06-09">9–11.VI.1937</date>
|
||
, collected by O. Piel;
|
||
<specimenCount id="36BCFD3A7614204AF8ABFD3A17964F9C" box="[406,510,692,717]" pageId="22" pageNumber="273" type="female">1 female</specimenCount>
|
||
,
|
||
<collectingCountry id="58AD76237614204AFB32FD3A143D4F9C" box="[527,597,692,717]" name="China" pageId="22" pageNumber="273">China</collectingCountry>
|
||
, Henan Province, Shangcheng, Suxianhe, Donghe village (
|
||
<geoCoordinate id="458E50747614204AFC37FD3A13FC4F9D" box="[1290,1428,692,716]" direction="north" orientation="latitude" pageId="22" pageNumber="273" precision="1" value="31.74098">31.74098°N</geoCoordinate>
|
||
/
|
||
<geoCoordinate id="458E50747614204AF9AAFD5917464FA1" box="[151,302,727,752]" direction="east" orientation="longitude" pageId="22" pageNumber="273" precision="1" value="115.55499">115.55499°E</geoCoordinate>
|
||
),
|
||
<quantity id="E7429B567614204AF87DFD5917EB4FA1" box="[320,387,727,752]" metricMagnitude="2" metricUnit="m" metricValue="5.04" pageId="22" pageNumber="273" unit="m" value="504.0">504m</quantity>
|
||
,
|
||
<date id="540410737614204AF8B3FD5914784FA1" box="[398,528,727,752]" pageId="22" pageNumber="273" value="2006-06-25">25.VI.2006</date>
|
||
, collected by Liu Ye & Liang Hongbin, all specimens deposited in IZ-CAS.
|
||
</paragraph>
|
||
<paragraph id="200536B37614204AF9FAFD7214584E0C" blockId="22.[151,1437,151,933]" pageId="22" pageNumber="273">
|
||
<emphasis id="12CEEAA17614204AF9FAFD72170A4E44" bold="true" box="[199,354,764,789]" pageId="22" pageNumber="273">Comparison.</emphasis>
|
||
<taxonomicName id="E7BA4D307614204AF854FD7217AE4E45" box="[361,454,764,788]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="273" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="12CEEAA17614204AF854FD7217AE4E45" box="[361,454,764,788]" italics="true" pageId="22" pageNumber="273">Quedius</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="12CEEAA17614204AF8EAFD7214444E45" box="[471,556,764,788]" italics="true" pageId="22" pageNumber="273">Vel leius</emphasis>
|
||
)
|
||
<taxonomicName id="E7BA4D307614204AFB02FD7314DA4E45" box="[575,690,765,788]" class="Insecta" family="Staphylinidae" genus="Velleius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="273" phylum="Arthropoda" rank="species" species="pectinatus">
|
||
<emphasis id="12CEEAA17614204AFB02FD7314DA4E45" box="[575,690,765,788]" italics="true" pageId="22" pageNumber="273">pectinatus</emphasis>
|
||
</taxonomicName>
|
||
is similar to other large-sized
|
||
<taxonomicName id="E7BA4D307614204AFD37FD7212364E45" box="[1034,1118,764,788]" class="Insecta" family="Staphylinidae" genus="Velleius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="273" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="12CEEAA17614204AFD37FD7212364E45" box="[1034,1118,764,788]" italics="true" pageId="22" pageNumber="273">Velleius</emphasis>
|
||
</taxonomicName>
|
||
species, but it can be easily recognized by deeply bifurcated antennal segments IV–X and the elongate last segment which is longer than preceding three segments combined.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="68A065387614204AF9FAFCE914FB4EF5" pageId="22" pageNumber="273" type="distribution">
|
||
<paragraph id="200536B37614204AF9FAFCE914FB4EF5" blockId="22.[151,1437,151,933]" pageId="22" pageNumber="273">
|
||
<emphasis id="12CEEAA17614204AF9FAFCE917084ED1" bold="true" box="[199,352,871,896]" pageId="22" pageNumber="273">Distribution.</emphasis>
|
||
<taxonomicName id="E7BA4D307614204AF856FCE614D24E2E" box="[363,698,872,896]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="273" phylum="Arthropoda" rank="species" species="pectinatus" subGenus="Velleius">
|
||
<emphasis id="12CEEAA17614204AF856FCE617A04ED1" box="[363,456,872,896]" italics="true" pageId="22" pageNumber="273">Quedius</emphasis>
|
||
(
|
||
<emphasis id="12CEEAA17614204AF8E1FCE614584ED1" box="[476,560,872,896]" italics="true" pageId="22" pageNumber="273">Velleius</emphasis>
|
||
)
|
||
<emphasis id="12CEEAA17614204AFB7AFCE614D24E2E" box="[583,698,872,895]" italics="true" pageId="22" pageNumber="273">pectinatus</emphasis>
|
||
</taxonomicName>
|
||
is distributed in
|
||
<collectingCountry id="58AD76237614204AFAB8FCE6159C4ED1" box="[901,1012,871,896]" name="China" pageId="22" pageNumber="273">East Asia</collectingCountry>
|
||
including
|
||
<collectingCountry id="58AD76237614204AFD4EFCE912DE4ED1" box="[1139,1206,871,896]" name="China" pageId="22" pageNumber="273">China</collectingCountry>
|
||
(Shanghai, Jiangxi, Zhejiang, Henan, Sichuan),
|
||
<collectingCountry id="58AD76237614204AF8ECFC03147D4EF5" box="[465,533,909,932]" name="South Korea" pageId="22" pageNumber="273">Korea</collectingCountry>
|
||
and
|
||
<collectingCountry id="58AD76237614204AFB71FC0314E74EF5" box="[588,655,909,932]" name="Japan" pageId="22" pageNumber="273">Japan</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |