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<document id="13CA1F4188EC8184291DD143AB67C2DC" ID-DOI="10.11646/zootaxa.5446.4.7" ID-ISSN="1175-5326" ID-Zenodo-Dep="11102260" ID-ZooBank="905DDE73-8CC9-4E88-BBAB-68B56F14A1F9" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1714636254393" checkinUser="plazi" docAuthor="Perkovsky, Evgeny E. &amp; Vasilenko, Dmitry V." docDate="2024" docId="7A6B87DB782BAB755EC3FBCBB6D3F839" docLanguage="en" docName="zootaxa.5446.4.7.pdf" docOrigin="Zootaxa 5446 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5446.4.7" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Adicophasma hafniensis Perkovsky &amp; Storozhenko 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="558" masterDocId="8652FFA3782FAB705E54FFDEB27CFF95" masterDocTitle="New and little-known Mantophasmatidae (Insecta: Mantophasmatodea) from European amber" masterLastPageNumber="563" masterPageNumber="553" pageNumber="557" updateTime="1714637384925" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="4F2CA2DC0D8FB9E0DDD7C7A11BF22619">New and little-known Mantophasmatidae (Insecta: Mantophasmatodea) from European amber</mods:title>
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<mods:name id="B5E5DCDE46365702DC296FE63E7A84D3" type="personal">
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<mods:namePart id="454C08C0F1F2B01DD16BD45BFD37ADD5">Perkovsky, Evgeny E.</mods:namePart>
<mods:affiliation id="859CF233A1023D60A584F3922CE0E4BB">Natural History Museum of Denmark, Universitetsparken 15, Copenhagen 2100, Denmark.</mods:affiliation>
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<mods:namePart id="8BBA0A8B4BB4C71D28DF502D2565EB38">Vasilenko, Dmitry V.</mods:namePart>
<mods:affiliation id="CEAF24BA3AA55ED7DF3A4BB86F68D343">Cherepovets State University, Lunacharsky Prospect 5, Cherepovets, Vologda Region, 162600, Russia. &amp; Borissiak Paleontological Institute, Russian Academy of Sciences, Moscow, 117647 Russia. &amp; Federal Scientific Center of the East Asia Terrestrial Biodiversity, Far Eastern Branch of the Russian Academy of Sciences, 690022 Vladivostok, Russia.</mods:affiliation>
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<mods:title id="9A77A912FD589C27720575AE781EEA7F">Zootaxa</mods:title>
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<mods:date id="6181FD4E96995B97DC0ED2013F4DA673">2024</mods:date>
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<mods:identifier id="4D3B314F8B74F65160D4D9B898CF5B55" type="DOI">10.11646/zootaxa.5446.4.7</mods:identifier>
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<treatment id="7A6B87DB782BAB755EC3FBCBB6D3F839" LSID="urn:lsid:plazi:treatment:7A6B87DB782BAB755EC3FBCBB6D3F839" httpUri="http://treatment.plazi.org/id/7A6B87DB782BAB755EC3FBCBB6D3F839" lastPageId="5" lastPageNumber="558" pageId="4" pageNumber="557">
<subSubSection id="BAD86546782BAB745EC3FBCBB101FBBA" box="[151,893,1045,1072]" pageId="4" pageNumber="557" type="nomenclature">
<paragraph id="F27D36CD782BAB745EC3FBCBB101FBBA" blockId="4.[151,893,1045,1107]" box="[151,893,1045,1072]" pageId="4" pageNumber="557">
<heading id="A93581A1782BAB745EC3FBCBB101FBBA" bold="true" box="[151,893,1045,1072]" fontSize="11" level="1" pageId="4" pageNumber="557" reason="1">
<emphasis id="C0B6EADF782BAB745EC3FBCBB101FBBA" bold="true" box="[151,893,1045,1072]" pageId="4" pageNumber="557">
<taxonomicName id="35C24D4E782BAB745EC3FBCBB16EFBBA" authority="Perkovsky &amp; Storozhenko" authorityName="Perkovsky &amp; Storozhenko" authorityYear="2024" box="[151,786,1045,1072]" class="Insecta" family="Mantophasmatidae" genus="Adicophasma" kingdom="Animalia" order="Mantophasmatodea" pageId="4" pageNumber="557" phylum="Arthropoda" rank="species" species="hafniensis" status="sp. nov.">
<emphasis id="C0B6EADF782BAB745EC3FBCBB3BDFBA5" bold="true" box="[151,449,1045,1072]" italics="true" pageId="4" pageNumber="557">Adicophasma hafniensis</emphasis>
Perkovsky &amp; Storozhenko
</taxonomicName>
,
<taxonomicNameLabel id="DB8557A4782BAB745D4BFBCBB101FBBA" box="[799,893,1045,1071]" pageId="4" pageNumber="557" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="BAD86546782BAB745EC3FBE7B37BFBC6" box="[151,263,1081,1107]" pageId="4" pageNumber="557" type="description">
<paragraph id="F27D36CD782BAB745EC3FBE7B37BFBC6" blockId="4.[151,893,1045,1107]" box="[151,263,1081,1107]" pageId="4" pageNumber="557">
<figureCitation id="6AF92A48782BAB745EC3FBE7B37BFBC6" box="[151,263,1081,1107]" captionStart-0="FIGURES 9–12" captionStart-1="FIGURES 13–16" captionStart-2="FIGURES 17–23" captionStartId-0="5.[152,270,1220,1244]" captionStartId-1="6.[152,270,1706,1730]" captionStartId-2="7.[152,270,1831,1855]" captionTargetBox-0="[170,1409,184,1190]" captionTargetBox-1="[247,1338,187,1676]" captionTargetBox-2="[158,1429,188,1802]" captionTargetId-0="figure-19@5.[170,1418,179,1194]" captionTargetId-1="figure-26@6.[177,1410,179,1680]" captionTargetId-2="figure-19@7.[151,1436,181,1807]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionText-0="FIGURES 9–12. Adicophasma hafniensis sp. nov. 9. Thoracic and abdominal tergites, dorsal view; 10. Fifth tarsal segment and arolium, dorsal view; 11. Anterior half of body, lateral view; 12. Head and profemur, frontal view." captionText-1="FIGURES 13–16. Adicophasma hafniensis sp. nov. 13. Body, fronto-ventral view; 14. Apex of abdomen, ventral view; 15. Mesofemur from inner side, lateral view; 16. Fore leg from inner side, lateral view." captionText-2="FIGURES 17–23. Adicophasma hafniensis sp. nov. 17. Head, basal segments of antennae, thoracic segments and coxae, lateral view; 18. Head, frontal view; 19. Fore leg, lateral view; 20. Metafemur, lateral view; 21. Thoracic and abdominal tergites; 22. Apex of abdomen, ventral view; 23. Fifth tarsal segment and arolium, dorsal view." figureDoi-0="http://doi.org/10.5281/zenodo.11102266" figureDoi-1="http://doi.org/10.5281/zenodo.11102268" figureDoi-2="http://doi.org/10.5281/zenodo.11102270" httpUri-0="https://zenodo.org/record/11102266/files/figure.png" httpUri-1="https://zenodo.org/record/11102268/files/figure.png" httpUri-2="https://zenodo.org/record/11102270/files/figure.png" pageId="4" pageNumber="557">Figs 9–23</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="BAD86546782BAB745EC3FB5FB285FB76" pageId="4" pageNumber="557" type="etymology">
<paragraph id="F27D36CD782BAB745EC3FB5FB285FB76" blockId="4.[151,1437,1153,1827]" pageId="4" pageNumber="557">
<emphasis id="C0B6EADF782BAB745EC3FB5FB362FB0E" bold="true" box="[151,286,1153,1179]" pageId="4" pageNumber="557">Etymology.</emphasis>
The species name is after Universitas Hafniensis (Københavns Universitet), the oldest university in
<collectingCountry id="8AD5765D782BAB745EECFB7BB35FFB2A" box="[184,291,1189,1215]" name="Denmark" pageId="4" pageNumber="557">Denmark</collectingCountry>
, where the
<typeStatus id="2D79886F782BAB745FFAFB78B3A2FB55" box="[430,478,1190,1216]" pageId="4" pageNumber="557">type</typeStatus>
was found at NHMD (the former Zoological Museum of Copenhagen University, ZMUC).
</paragraph>
</subSubSection>
<subSubSection id="BAD86546782BAB745E93FB33B3ABFAE6" pageId="4" pageNumber="557" type="materials_examined">
<paragraph id="F27D36CD782BAB745E93FB33B3ABFAE6" blockId="4.[151,1437,1153,1827]" pageId="4" pageNumber="557">
<emphasis id="C0B6EADF782BAB745E93FB33B308FA92" bold="true" box="[199,372,1261,1287]" pageId="4" pageNumber="557">Type material</emphasis>
.
<materialsCitation id="42AA3C90782BAB745FDCFB33B3ABFAE6" collectingDate="1969-02-01" collectionCode="V" pageId="4" pageNumber="557" specimenCount="1" typeStatus="holotype">
<typeStatus id="2D79886F782BAB745FDCFB33B38CFA92" box="[392,496,1261,1287]" pageId="4" pageNumber="557" type="holotype">Holotype</typeStatus>
ZMUC-901825, with old label ‘Phasmoidea 1825 C.
<collectionCode id="94D3AE08782BAB745A38FB33B6FEFA92" box="[1132,1154,1261,1287]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="4" pageNumber="557" type="Museum">V</collectionCode>
. Henningsen
<date id="867C100D782BAB745B71FB33B7EFFA92" box="[1317,1427,1261,1287]" pageId="4" pageNumber="557" value="1969-02-01">
<collectingDate id="9638E9E5782BAB745B71FB33B7EFFA92" box="[1317,1427,1261,1287]" pageId="4" pageNumber="557" value="1969-02-01">1-2-1969</collectingDate>
</date>
’, Danish amber, late Eocene. Nymph, probably mid instar. The sex unknown, because of in the nymphs of extant
<taxonomicName id="35C24D4E782BAB745EC3FAEBB30EFADA" authorityName="Klass, Zompro, Kristensen &amp; Adis" authorityYear="2002" box="[151,370,1333,1359]" class="Insecta" kingdom="Animalia" order="Mantophasmatodea" pageId="4" pageNumber="557" phylum="Arthropoda" rank="order">Mantophasmatodea</taxonomicName>
the apical abdominal sternites and cerci are not strongly modified such as in adult specimens. Syninclusions: stellate hairs.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="BAD86546782BAB745E93FAA3B620F9DE" pageId="4" pageNumber="557" type="diagnosis">
<paragraph id="F27D36CD782BAB745E93FAA3B620F9DE" blockId="4.[151,1437,1153,1827]" pageId="4" pageNumber="557">
<emphasis id="C0B6EADF782BAB745E93FAA3B346FA02" bold="true" box="[199,314,1405,1431]" pageId="4" pageNumber="557">Diagnosis</emphasis>
. New species is most similar to
<taxonomicName id="35C24D4E782BAB745CF7FAA3B18FFA02" authorityName="Arillo &amp; Engel" authorityYear="2006" box="[675,1011,1405,1431]" class="Insecta" family="Mantophasmatidae" genus="Adicophasma" kingdom="Animalia" order="Mantophasmatodea" pageId="4" pageNumber="557" phylum="Arthropoda" rank="species" species="grylloblattoides">
<emphasis id="C0B6EADF782BAB745CF7FAA3B18FFA02" box="[675,1011,1405,1431]" italics="true" pageId="4" pageNumber="557">Adicophasma grylloblattoides</emphasis>
</taxonomicName>
but differs from the latter by enlarged pedicel and chaetotaxy of abdomen (in
<taxonomicName id="35C24D4E782BAB745C35FA7CB148FA2E" authorityName="Arillo &amp; Engel" authorityYear="2006" box="[609,820,1441,1468]" class="Insecta" family="Mantophasmatidae" genus="Adicophasma" kingdom="Animalia" order="Mantophasmatodea" pageId="4" pageNumber="557" phylum="Arthropoda" rank="species" species="grylloblattoides">
<emphasis id="C0B6EADF782BAB745C35FA7CB148FA2E" box="[609,820,1441,1468]" italics="true" pageId="4" pageNumber="557">A. grylloblattoides</emphasis>
</taxonomicName>
, pedicel as long as wide and sternites with scattered, simple setae). In
<taxonomicName id="35C24D4E782BAB745F08FA18B39BFA4A" authorityName="Engel &amp; Grimaldi" authorityYear="2004" box="[348,487,1478,1504]" class="Insecta" family="Mantophasmatidae" genus="Adicophasma" kingdom="Animalia" order="Mantophasmatodea" pageId="4" pageNumber="557" phylum="Arthropoda" rank="species" species="spinosum">
<emphasis id="C0B6EADF782BAB745F08FA18B39BFA4A" box="[348,487,1478,1504]" italics="true" pageId="4" pageNumber="557">A. spinosum</emphasis>
</taxonomicName>
pedicel elongated, at least 3 times longer than wide, scapus shorter than elongated pedicel, anterior side of procoxa with 3 long bristles, and sternites with numerous sparse setae of moderated length. The features separated
<taxonomicName id="35C24D4E782BAB745FC8F9D0B06EF9B2" authorityName="Zompro" authorityYear="2008" box="[412,530,1549,1576]" class="Insecta" family="Mantophasmatidae" genus="Adicophasma" kingdom="Animalia" order="Mantophasmatodea" pageId="4" pageNumber="557" phylum="Arthropoda" rank="species" species="groehni">
<emphasis id="C0B6EADF782BAB745FC8F9D0B06EF9B2" box="[412,530,1549,1576]" italics="true" pageId="4" pageNumber="557">A. groehni</emphasis>
</taxonomicName>
from new species are follow: the posterior margin of pro-, meso- and metatorax with 6 long bristles and the posterior margin of sternites with a row of up to 8 bristles.
</paragraph>
</subSubSection>
<subSubSection id="BAD86546782BAB755E93F98BB183F81C" lastPageId="5" lastPageNumber="558" pageId="4" pageNumber="557" type="description">
<paragraph id="F27D36CD782BAB745E93F98BB602F8B6" blockId="4.[151,1437,1153,1827]" pageId="4" pageNumber="557">
<emphasis id="C0B6EADF782BAB745E93F98BB32DF9FA" bold="true" box="[199,337,1621,1647]" pageId="4" pageNumber="557">Description</emphasis>
. Head hypognathous; the compound eyes large and protrude slightly above the head; vertical diameter of the eye significantly exceeds the height of the cheeks. Frontoclipeal suture broadly rounded; clipeus triangular; labrum narrow triangular with acutely rounded apex. Antennae 16-segmented; scapus cylindrical, 1.5 times as long as wide, about equal in size to pedicel; pedicel 1.5 times as long as wide; third antennomere (first flagellomere) elongated, 3.5 times as long as wide and 1.4 times as long as scapus. Maxillary palpus 5-segmented, second and fifth palpomeres longest, other palpomeres short. Labial palpus 3-segmented.
</paragraph>
<caption id="A6BD6645782AAB755ECCFB1AB604FA95" ID-DOI="http://doi.org/10.5281/zenodo.11102266" ID-Zenodo-Dep="11102266" httpUri="https://zenodo.org/record/11102266/files/figure.png" pageId="5" pageNumber="558" startId="5.[152,270,1220,1244]" targetBox="[170,1409,184,1190]" targetPageId="5" targetType="figure">
<paragraph id="F27D36CD782AAB755ECCFB1AB604FA95" blockId="5.[151,1437,1220,1280]" pageId="5" pageNumber="558">
<emphasis id="C0B6EADF782AAB755ECCFB1AB33AFB49" bold="true" box="[152,326,1220,1244]" pageId="5" pageNumber="558">FIGURES 9–12</emphasis>
.
<taxonomicName id="35C24D4E782AAB755F05FB1AB03BFB49" authority="Perkovsky &amp; Vasilenko, 2024" authorityName="Perkovsky &amp; Vasilenko" authorityYear="2024" box="[337,583,1220,1244]" class="Insecta" family="Mantophasmatidae" genus="Adicophasma" kingdom="Animalia" order="Mantophasmatodea" pageId="5" pageNumber="558" phylum="Arthropoda" rank="species" species="hafniensis" status="sp. nov.">
<emphasis id="C0B6EADF782AAB755F05FB1AB03BFB49" box="[337,583,1220,1244]" italics="true" pageId="5" pageNumber="558">Adicophasma hafniensis</emphasis>
</taxonomicName>
<emphasis id="C0B6EADF782AAB755C18FB1AB0E7FB49" bold="true" box="[588,667,1220,1244]" pageId="5" pageNumber="558">
<taxonomicNameLabel id="DB8557A4782AAB755C18FB1AB0E7FB49" box="[588,667,1220,1244]" pageId="5" pageNumber="558" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
9. Thoracic and abdominal tergites, dorsal view; 10. Fifth tarsal segment and arolium, dorsal view; 11. Anterior half of body, lateral view; 12. Head and profemur, frontal view.
</paragraph>
</caption>
<paragraph id="F27D36CD782AAB755E93FAF0B332FA4D" blockId="5.[151,1437,1326,1965]" pageId="5" pageNumber="558">Thorax. Pronotum distinctly shorter than head, almost as long as meso- and metanotum, with almost straight posterior margin armed by four bristles. Dorsal side of pro-, meso- and metatorax with numerous, mostly irregularly situated, spine-like bristles. Thoracic tergites distinctly marginated laterally and posteriorly; posterior margin of mesothorax straight and with a row of six bristles. Metatorax narrower than mesothorax, with a few irregularly situated bristles.</paragraph>
<paragraph id="F27D36CD782AAB755E93FA3DB6B6F924" blockId="5.[151,1437,1326,1965]" pageId="5" pageNumber="558">
Legs. Pro- and metafemora slightly broadened and longer than more slender mesofemora. Ventral side of pro- and mesofemora with two rows of spine-like bristles. Protibia with numerous hair-like setae, inner side apically bearing 3-4 setae with cuticular thickenings at base. Mesotibia not visible. Coxae elongate, anterior side of procoxa with a long bristle. Tarsi 5-segmented, basitarsus as long as second tarsomere; third segments slightly longer than second; fourth tarsomere as long as third; third and fourth tarsomere with wide distal process in the middle; fifth segment elongate; arolium large, with typical for the
<taxonomicName id="35C24D4E782AAB755CBEF948B1B9F925" authorityName="Klass, Zompro, Kristensen &amp; Adis" authorityYear="2002" box="[746,965,1686,1712]" class="Insecta" kingdom="Animalia" order="Mantophasmatodea" pageId="5" pageNumber="558" phylum="Arthropoda" rank="order">Mantophasmatodea</taxonomicName>
preapical row of setae.
</paragraph>
<paragraph id="F27D36CD782AAB755E93F964B7F6F8D4" blockId="5.[151,1437,1326,1965]" pageId="5" pageNumber="558">Abdomen. Abdominal tergites transverse, with a few irregularly situated very long setae; anal plate trangular. Abdomem below with visible four apical sternites and subgenital plate. Hind margin of 8-th and 9-th sternites with two strong bristles; apex of subgenital plate with deep excision; paraprocts considerably extending beyond apex of subgenital plate; cerci simple. Lateral sides of sternites and cerci with very long, thin and densely arranged setae.</paragraph>
<paragraph id="F27D36CD782AAB755E93F894B183F81C" blockId="5.[151,1437,1326,1965]" pageId="5" pageNumber="558">Lengths (mm): Total: 3.2; head: 0,56; antennae: 3.35; pronotum: 0,43; mesonotum: 0,41; metanotum: 0,4; profemora: 0,75; protibiae: 0.82; protarsi: 0,65; mesofemora: 0,76; cerci 0,15.</paragraph>
</subSubSection>
<subSubSection id="BAD86546782AAB755E93F84CB6D3F839" box="[199,1199,1938,1965]" pageId="5" pageNumber="558" type="discussion">
<paragraph id="F27D36CD782AAB755E93F84CB6D3F839" blockId="5.[151,1437,1326,1965]" box="[199,1199,1938,1965]" pageId="5" pageNumber="558">
<emphasis id="C0B6EADF782AAB755E93F84CB348F839" bold="true" box="[199,308,1938,1964]" pageId="5" pageNumber="558">Remarks</emphasis>
. This is first record of the genus
<taxonomicName id="35C24D4E782AAB755CF2F84CB142F839" authorityName="Engel &amp; Grimaldi" authorityYear="2004" box="[678,830,1938,1964]" class="Insecta" family="Mantophasmatidae" genus="Adicophasma" kingdom="Animalia" order="Mantophasmatodea" pageId="5" pageNumber="558" phylum="Arthropoda" rank="genus">
<emphasis id="C0B6EADF782AAB755CF2F84CB142F839" box="[678,830,1938,1964]" italics="true" pageId="5" pageNumber="558">Adicophasma</emphasis>
</taxonomicName>
from late Eocene Danish amber.
</paragraph>
</subSubSection>
</treatment>
</document>
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