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<document id="6610F77331A29AE4BB1673FF66120292" ID-DOI="10.26879/1209" ID-ISSN="1094-8074" ID-Zenodo-Dep="11063294" ID-ZooBank="1726FDAE-2EE5-4145-A124-6D24287C0514" IM.bibliography_approvedBy="julia" IM.illustrations_approvedBy="julia" IM.materialsCitations_approvedBy="julia" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="julia" IM.taxonomicNames_approvedBy="julia" IM.treatments_approvedBy="julia" checkinTime="1714014612185" checkinUser="felipe" docAuthor="Zazhigin, Vladimir S. &amp; Voyta, Leonid L." docDate="2022" docId="71755174FFBB0329CE9AFCD8CC07FD7E" docLanguage="en" docName="PalaeontolElectron.25.3.a29.pdf" docOrigin="Palaeontologia Electronica (a 29) 25 (3)" docSource="http://dx.doi.org/10.26879/1209" docStyle="DocumentStyle:322FC57CC99045555090E8A84CFB029A.4:PalaeontolElectron.2018-.journal_article" docStyleId="322FC57CC99045555090E8A84CFB029A" docStyleName="PalaeontolElectron.2018-.journal_article" docStyleVersion="4" docTitle="Crusafontina undefined-1" docType="treatment" docVersion="3" lastPageNumber="17" masterDocId="8D4C290CFFAB0339CF1EFFE8C819FF81" masterDocTitle="New Neogene anourosoricin shrews from northern Asia" masterLastPageNumber="36" masterPageNumber="1" pageNumber="17" updateTime="1714663430529" updateUser="julia" zenodo-license-document="CC-BY-NC-SA-4.0">
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<mods:title id="1383B72F02C8B794D33854D803CD2817">New Neogene anourosoricin shrews from northern Asia</mods:title>
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<treatment id="71755174FFBB0329CE9AFCD8CC07FD7E" LSID="urn:lsid:plazi:treatment:71755174FFBB0329CE9AFCD8CC07FD7E" httpUri="http://treatment.plazi.org/id/71755174FFBB0329CE9AFCD8CC07FD7E" lastPageNumber="17" pageId="16" pageNumber="17">
<subSubSection id="B1C6B3E9FFBB0329CE9AFCD8CA4DFCCB" box="[388,596,816,842]" pageId="16" pageNumber="17" type="nomenclature">
<paragraph id="F963E062FFBB0329CE9AFCD8CA4DFCCB" blockId="16.[388,596,816,874]" box="[388,596,816,842]" pageId="16" pageNumber="17">
<heading id="A22B570EFFBB0329CE9AFCD8CA4DFCCB" box="[388,596,816,842]" centered="true" fontSize="10" level="2" pageId="16" pageNumber="17" reason="7">
<taxonomicName id="3EDC9BE1FFBB0329CE9AFCD8CA4DFCCB" box="[388,596,816,842]" class="Mammalia" family="Soricidae" genus="Crusafontina" kingdom="Animalia" order="Soricomorpha" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="undefined-1">
<emphasis id="CBA83C70FFBB0329CE9AFCD8CA02FCC8" box="[388,539,816,841]" italics="true" pageId="16" pageNumber="17">Crusafontina</emphasis>
sp.1
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="B1C6B3E9FFBB0329CEAFFCB9C94CFC7B" pageId="16" pageNumber="17" type="description">
<paragraph id="F963E062FFBB0329CEAFFCB9CA3FFCEB" blockId="16.[388,596,816,874]" box="[433,550,848,874]" pageId="16" pageNumber="17">
<figureCitation id="61E7FCE7FFBB0329CEAFFCB9CA3FFCEB" box="[433,550,848,874]" captionStart="FIGURE 6" captionStartId="11.[196,287,1080,1103]" captionTargetBox="[190,1441,166,1060]" captionTargetId="figure-235@11.[190,1441,166,1060]" captionTargetPageId="11" captionText="FIGURE 6. Teeth and bones of Paranourosorex seletiensis Storch and Zazhigin, 1996 (A, B), Crusafontina sp. 1, (C), Crusafontina sp. 2 (D) and Paranourosorex intermedius sp. nov. (E). A, GIN 951/1000 (holotype) from SLT/1A, left hemimandible fragment (cut image) with m1 and m2 in occlusal view (A2, magnified m2); B, GIN 951/1001 (paratype) from SLT/1A, left P4 in occlusal view; C, GIN 640/1004 from PVL/1A, left M1 in occlusal view; D, GIN 951/1003 SLT/ 1A, left P4 in occlusal view; E, GIN 948/1051 (holotype) from NST/1A, right dentary fragment with m1 and m2 (E1, occlusal view; E2, lateral view E3, medial view;). Scale bars equal 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11063306" httpUri="https://zenodo.org/record/11063306/files/figure.png" pageId="16" pageNumber="17">Figure 6C</figureCitation>
</paragraph>
<paragraph id="F963E062FFBB0329CFDEFC93C9D0FC35" blockId="16.[192,793,891,1888]" pageId="16" pageNumber="17">
1996
<taxonomicName id="3EDC9BE1FFBB0329CE3EFC93CA05FC15" box="[288,540,891,916]" class="Mammalia" family="Soricidae" genus="Paranourosorex" kingdom="Animalia" order="Soricomorpha" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="undefined-1">
<emphasis id="CBA83C70FFBB0329CE3EFC93C9C7FC15" box="[288,478,891,916]" italics="true" pageId="16" pageNumber="17">Paranourosorex</emphasis>
sp. 1
</taxonomicName>
Storch and Zazhigin, p. 261, fig. 4a.
</paragraph>
<paragraph id="F963E062FFBB0329CFDEFC29C94CFC7B" blockId="16.[192,793,891,1888]" pageId="16" pageNumber="17">
2004
<taxonomicName id="3EDC9BE1FFBB0329CE3EFC28CA7DFC5B" authorityName="van Dam" authorityYear="2004" box="[288,612,960,986]" class="Mammalia" family="Soricidae" genus="Crusafontina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="vandeweerdi">
<emphasis id="CBA83C70FFBB0329CE3EFC28C9AEFC58" box="[288,439,960,985]" italics="true" pageId="16" pageNumber="17">Crusafontina</emphasis>
?
<emphasis id="CBA83C70FFBB0329CED3FC29CA7DFC5B" box="[461,612,961,986]" italics="true" pageId="16" pageNumber="17">vandeweerdi</emphasis>
</taxonomicName>
? van Dam, p. 763.
</paragraph>
</subSubSection>
<subSubSection id="B1C6B3E9FFBB0329CFDEFBEECA74FBBE" pageId="16" pageNumber="17" type="materials_examined">
<paragraph id="F963E062FFBB0329CFDEFBEECA74FBBE" blockId="16.[192,793,891,1888]" pageId="16" pageNumber="17">
<emphasis id="CBA83C70FFBB0329CFDEFBEEC935FB9E" bold="true" box="[192,300,1030,1055]" pageId="16" pageNumber="17">Material.</emphasis>
<materialsCitation id="49B4EA3FFFBB0329CE22FBEECA70FBBE" pageId="16" pageNumber="17">Isolated left M1 (GIN 640/1004) from Pavlodar 1A locality (Appendix 2: 5)</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B1C6B3E9FFBB0329CFDEFBAECACEF9BE" pageId="16" pageNumber="17" type="description">
<paragraph id="F963E062FFBB0329CFDEFBAECAAFFA5E" blockId="16.[192,793,891,1888]" pageId="16" pageNumber="17">
<emphasis id="CBA83C70FFBB0329CFDEFBAEC943FBDE" bold="true" box="[192,346,1094,1119]" pageId="16" pageNumber="17">Description.</emphasis>
The large and moderately worn first upper molar displays clearly defined crown features. The buccal crests are expressed W-shaped line like a condition of
<taxonomicName id="3EDC9BE1FFBB0329CED9FB4ECA9DFB3E" authorityName="Rzebik-Kowalska" authorityYear="1975" box="[455,644,1190,1215]" class="Mammalia" family="Soricidae" genus="Paranourosorex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="16" pageNumber="17" phylum="Chordata" rank="genus">
<emphasis id="CBA83C70FFBB0329CED9FB4ECA9DFB3E" box="[455,644,1190,1215]" italics="true" pageId="16" pageNumber="17">Paranourosorex</emphasis>
</taxonomicName>
(
<figureCitation id="61E7FCE7FFBB0329CD8DFB4ECB01FB3E" box="[659,792,1190,1215]" captionStart="FIGURE 6" captionStartId="11.[196,287,1080,1103]" captionTargetBox="[190,1441,166,1060]" captionTargetId="figure-235@11.[190,1441,166,1060]" captionTargetPageId="11" captionText="FIGURE 6. Teeth and bones of Paranourosorex seletiensis Storch and Zazhigin, 1996 (A, B), Crusafontina sp. 1, (C), Crusafontina sp. 2 (D) and Paranourosorex intermedius sp. nov. (E). A, GIN 951/1000 (holotype) from SLT/1A, left hemimandible fragment (cut image) with m1 and m2 in occlusal view (A2, magnified m2); B, GIN 951/1001 (paratype) from SLT/1A, left P4 in occlusal view; C, GIN 640/1004 from PVL/1A, left M1 in occlusal view; D, GIN 951/1003 SLT/ 1A, left P4 in occlusal view; E, GIN 948/1051 (holotype) from NST/1A, right dentary fragment with m1 and m2 (E1, occlusal view; E2, lateral view E3, medial view;). Scale bars equal 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11063306" httpUri="https://zenodo.org/record/11063306/files/figure.png" pageId="16" pageNumber="17">Figures 6D</figureCitation>
cf. 7G). The metaconule is absent; the short metaloph is similar to the
<taxonomicName id="3EDC9BE1FFBB0329CD18FB0ECADDFB7E" authorityName="Rzebik-Kowalska" authorityYear="1975" box="[518,708,1254,1279]" class="Mammalia" family="Soricidae" genus="Paranourosorex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="16" pageNumber="17" phylum="Chordata" rank="genus">
<emphasis id="CBA83C70FFBB0329CD18FB0ECADDFB7E" box="[518,708,1254,1279]" italics="true" pageId="16" pageNumber="17">Paranourosorex</emphasis>
</taxonomicName>
condition and has a weak broadening of the base (poorly developed metaconule by
<bibRefCitation id="9D4D9D93FFBB0329CD7FFACEC8F6FADE" author="Van Dam, J. A." pageId="16" pageNumber="17" pagination="741 - 764" refId="ref17856" refString="Van Dam, J. A. 2004. Anourosoricini (Mammalia: Soricidae) from the Mediterranean region: A pre-Quaternary example of recurrent climate-controlled north - south range shifting. Journal of Paleontology, 78: 741 - 764. https: // www. jstor. org / stable / 4094903" type="journal article" year="2004">van Dam 2004: 752</bibRefCitation>
) and contacts with the metacone base. The hypocone is elongated in the base and shifts to the lingual margin of the flange. The posterior emargination is well-expressed; the hypoconal flange is protruded posteriorly and narrow in shape.
</paragraph>
<paragraph id="F963E062FFBB0329CFDEFA0ECACEF9BE" blockId="16.[192,793,891,1888]" pageId="16" pageNumber="17">
<emphasis id="CBA83C70FFBB0329CFDEFA0EC998FA7E" bold="true" box="[192,385,1510,1535]" pageId="16" pageNumber="17">Measurements.</emphasis>
BL(M1) =
<quantity id="3E244D87FFBB0329CD0AFA0ECA9EFA7E" box="[532,647,1510,1535]" metricMagnitude="-3" metricUnit="m" metricValue="2.54" pageId="16" pageNumber="17" unit="mm" value="2.54">2.54 mm</quantity>
; AW(M1) =
<quantity id="3E244D87FFBB0329CFDEF9EEC935F99E" box="[192,300,1542,1567]" metricMagnitude="-3" metricUnit="m" metricValue="2.68" pageId="16" pageNumber="17" unit="mm" value="2.68">2.68 mm</quantity>
; PW(M1) =
<quantity id="3E244D87FFBB0329CEA5F9EECA3EF99E" box="[443,551,1542,1567]" metricMagnitude="-3" metricUnit="m" metricValue="2.56" pageId="16" pageNumber="17" unit="mm" value="2.56">2.56 mm</quantity>
; LL(M1) =
<quantity id="3E244D87FFBB0329CDB6F9EECB0DF99E" box="[680,788,1542,1567]" metricMagnitude="-3" metricUnit="m" metricValue="2.42" pageId="16" pageNumber="17" unit="mm" value="2.42">2.42 mm</quantity>
; PE(M1) =
<quantity id="3E244D87FFBB0329CE27F9CEC9B9F9BE" box="[313,416,1574,1599]" metricMagnitude="-3" metricUnit="m" metricValue="2.07" pageId="16" pageNumber="17" unit="mm" value="2.07">2.07 mm</quantity>
(abbr. see in Appendix 3).
</paragraph>
</subSubSection>
<subSubSection id="B1C6B3E9FFBB0329CFDEF9AECC07FD7E" pageId="16" pageNumber="17" type="discussion">
<paragraph id="F963E062FFBB0329CFDEF9AECD13FDFE" blockId="16.[192,793,891,1888]" lastBlockId="16.[840,1441,166,767]" pageId="16" pageNumber="17">
<emphasis id="CBA83C70FFBB0329CFDEF9AEC921F9DE" bold="true" box="[192,312,1606,1631]" pageId="16" pageNumber="17">Remarks.</emphasis>
The M1 is slightly larger than the known M1 of
<taxonomicName id="3EDC9BE1FFBB0329CE10F98EC9D0F9FE" authorityName="van Dam" authorityYear="2004" box="[270,457,1638,1663]" class="Mammalia" family="Soricidae" genus="Crusafontina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="vandeweerdi">
<emphasis id="CBA83C70FFBB0329CE10F98EC9D0F9FE" box="[270,457,1638,1663]" italics="true" pageId="16" pageNumber="17">C. vandeweerdi</emphasis>
</taxonomicName>
(BL =
<quantity id="3E244D87FFBB0329CD00F98ECA93F9FE" box="[542,650,1638,1663]" metricMagnitude="-3" metricUnit="m" metricValue="2.48" pageId="16" pageNumber="17" unit="mm" value="2.48">2.48 mm</quantity>
, AW =
<quantity id="3E244D87FFBB0329CDFAF98EC8E9F91E" metricMagnitude="-3" metricUnit="m" metricValue="2.58" pageId="16" pageNumber="17" unit="mm" value="2.58">2.58 mm</quantity>
, see
<bibRefCitation id="9D4D9D93FFBB0329CE31F96ECA3AF91E" author="Van Dam, J. A." box="[303,547,1670,1695]" pageId="16" pageNumber="17" pagination="741 - 764" refId="ref17856" refString="Van Dam, J. A. 2004. Anourosoricini (Mammalia: Soricidae) from the Mediterranean region: A pre-Quaternary example of recurrent climate-controlled north - south range shifting. Journal of Paleontology, 78: 741 - 764. https: // www. jstor. org / stable / 4094903" type="journal article" year="2004">van Dam, 2004: 750</bibRefCitation>
) and distinctly larger than other species of
<taxonomicName id="3EDC9BE1FFBB0329CEDEF94ECA4EF93E" authorityName="Gibert" authorityYear="1974" box="[448,599,1702,1727]" class="Mammalia" family="Soricidae" genus="Crusafontina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="16" pageNumber="17" phylum="Chordata" rank="genus">
<emphasis id="CBA83C70FFBB0329CEDEF94ECA4EF93E" box="[448,599,1702,1727]" italics="true" pageId="16" pageNumber="17">Crusafontina</emphasis>
</taxonomicName>
. The crown features of M1 such as the buccal crests shape and the buccal outline are similar to
<taxonomicName id="3EDC9BE1FFBB0329CD44F90ECB01F97E" authorityName="Rzebik-Kowalska" authorityYear="1975" box="[602,792,1766,1791]" class="Mammalia" family="Soricidae" genus="Paranourosorex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="16" pageNumber="17" phylum="Chordata" rank="genus">
<emphasis id="CBA83C70FFBB0329CD44F90ECB01F97E" box="[602,792,1766,1791]" italics="true" pageId="16" pageNumber="17">Paranourosorex</emphasis>
</taxonomicName>
conditions. This fact did not lead
<bibRefCitation id="9D4D9D93FFBB0329CD58F8EEC922F8BE" author="Storch, G. &amp; Zazhigin, V. S." pageId="16" pageNumber="17" pagination="257 - 268" refId="ref17571" refString="Storch, G. and Zazhigin, V. S. 1996. Taxonomy and phylogeny of the Paranourosorex lineage, Neogene of Eurasia (Mammalia: Soricidae: Anourooricini). Palaontologische Zeitschrift, 70: 257 - 268. https: // doi. org / 10.1007 / BF 02988282" type="journal article" year="1996">Storch and Zazhigin (1996)</bibRefCitation>
to determine this molar as a
<taxonomicName id="3EDC9BE1FFBB0329CDBDF8CEC8F2F8DE" authorityName="Gibert" authorityYear="1974" class="Mammalia" family="Soricidae" genus="Crusafontina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="16" pageNumber="17" phylum="Chordata" rank="genus">
<emphasis id="CBA83C70FFBB0329CDBDF8CEC8F2F8DE" italics="true" pageId="16" pageNumber="17">Crusafontina</emphasis>
</taxonomicName>
. On the other hand, the authors could not associate this molar to
<taxonomicName id="3EDC9BE1FFBB0329CB76FF4ECD3CFF3E" authorityName="Rzebik-Kowalska" authorityYear="1975" box="[1128,1317,166,191]" class="Mammalia" family="Soricidae" genus="Paranourosorex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="16" pageNumber="17" phylum="Chordata" rank="genus">
<emphasis id="CBA83C70FFBB0329CB76FF4ECD3CFF3E" box="[1128,1317,166,191]" italics="true" pageId="16" pageNumber="17">Paranourosorex</emphasis>
</taxonomicName>
without a doubt because it has deep posterior emargination, contact between the metaloph and the base of the metacone and the narrow hypoconal flange. Later,
<bibRefCitation id="9D4D9D93FFBB0329CC56FECECC15FEBE" author="Van Dam, J. A." box="[840,1036,294,319]" pageId="16" pageNumber="17" pagination="741 - 764" refId="ref17856" refString="Van Dam, J. A. 2004. Anourosoricini (Mammalia: Soricidae) from the Mediterranean region: A pre-Quaternary example of recurrent climate-controlled north - south range shifting. Journal of Paleontology, 78: 741 - 764. https: // www. jstor. org / stable / 4094903" type="journal article" year="2004">van Dam (2004)</bibRefCitation>
supposed the conspecifity of the Pavlodars form and the new Spanish species
<taxonomicName id="3EDC9BE1FFBB0329CA9BFEAECBC6FEFE" authorityName="van Dam" authorityYear="2004" class="Mammalia" family="Soricidae" genus="Crusafontina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="vandeweerdi">
<emphasis id="CBA83C70FFBB0329CA9BFEAECBC6FEFE" italics="true" pageId="16" pageNumber="17">C. vandeweerdi</emphasis>
</taxonomicName>
(Teruel Basin, MN 12), pointed its intermediate feature conditions to both
<taxonomicName id="3EDC9BE1FFBB0329CA35FE6ECB6AFE3E" authorityName="Gibert" authorityYear="1974" class="Mammalia" family="Soricidae" genus="Crusafontina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="16" pageNumber="17" phylum="Chordata" rank="genus">
<emphasis id="CBA83C70FFBB0329CA35FE6ECB6AFE3E" italics="true" pageId="16" pageNumber="17">Crusafontina</emphasis>
</taxonomicName>
and
<taxonomicName id="3EDC9BE1FFBB0329CCACFE4ECC69FE3E" authorityName="Rzebik-Kowalska" authorityYear="1975" box="[946,1136,422,447]" class="Mammalia" family="Soricidae" genus="Paranourosorex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="16" pageNumber="17" phylum="Chordata" rank="genus">
<emphasis id="CBA83C70FFBB0329CCACFE4ECC69FE3E" box="[946,1136,422,447]" italics="true" pageId="16" pageNumber="17">Paranourosorex</emphasis>
</taxonomicName>
. The species has
<taxonomicName id="3EDC9BE1FFBB0329CA4EFE4ECB81FE5E" authorityName="Gibert" authorityYear="1974" class="Mammalia" family="Soricidae" genus="Crusafontina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="16" pageNumber="17" phylum="Chordata" rank="genus">
<emphasis id="CBA83C70FFBB0329CA4EFE4ECB81FE5E" italics="true" pageId="16" pageNumber="17">Crusafontina</emphasis>
</taxonomicName>
-like p4 with a central crest and distinct posterolingual basin; the ecto- and entocingulids of p4 are absent. We determined the Pavlodars molar as
<taxonomicName id="3EDC9BE1FFBB0329CCA6FDCECC8AFDBE" box="[952,1171,550,575]" class="Mammalia" family="Soricidae" genus="Crusafontina" kingdom="Animalia" order="Soricomorpha" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="undefined-1">
<emphasis id="CBA83C70FFBB0329CCA6FDCECC56FDBE" box="[952,1103,550,575]" italics="true" pageId="16" pageNumber="17">Crusafontina</emphasis>
sp. 1
</taxonomicName>
due to the absence of any tooth material for more detailed comparisons with
<typeStatus id="26675EC0FFBB0329CC61FD8ECBA8FDFE" box="[895,945,614,639]" pageId="16" pageNumber="17">type</typeStatus>
materials of
<taxonomicName id="3EDC9BE1FFBB0329CB54FD8ECD1AFDFE" authorityName="van Dam" authorityYear="2004" box="[1098,1283,614,639]" class="Mammalia" family="Soricidae" genus="Crusafontina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="vandeweerdi">
<emphasis id="CBA83C70FFBB0329CB54FD8ECD1AFDFE" box="[1098,1283,614,639]" italics="true" pageId="16" pageNumber="17">C. vandeweerdi</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="F963E062FFBB0329CC56FD6ECC07FD7E" blockId="16.[840,1441,166,767]" pageId="16" pageNumber="17">
<emphasis id="CBA83C70FFBB0329CC56FD6ECD06FD1E" bold="true" box="[840,1311,646,672]" pageId="16" pageNumber="17">Stratigraphic and geographic range.</emphasis>
<taxonomicName id="3EDC9BE1FFBB0329CA35FD6ECBACFD3E" class="Mammalia" family="Soricidae" genus="Crusafontina" kingdom="Animalia" order="Soricomorpha" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="undefined-1">
<emphasis id="CBA83C70FFBB0329CA35FD6ECB6AFD3E" italics="true" pageId="16" pageNumber="17">Crusafontina</emphasis>
sp. 1
</taxonomicName>
known only from the single Kazakh locality, Pavlodar 1A, late Miocene Pavlodar Formation (Turolian, MN 12).
</paragraph>
</subSubSection>
</treatment>
</document>