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<mods:title id="C88DACCBA58108AEA845611946BB6297">Taxonomic revision of the orb weaving spider genus Plato Coddington, 1986 (Araneae: Theridiosomatidae) with the description of three new species</mods:title>
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<mods:namePart id="77BC34E0499DDF31D6485A9790D642F0">Prete, Pedro H.</mods:namePart>
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<treatment id="715D87BA4D18287FFF17FF42FD0AFD53" ID-DOI="http://doi.org/10.5281/zenodo.12190441" ID-GBIF-Taxon="233317067" ID-Zenodo-Dep="12190441" LSID="urn:lsid:plazi:treatment:715D87BA4D18287FFF17FF42FD0AFD53" httpUri="http://treatment.plazi.org/id/715D87BA4D18287FFF17FF42FD0AFD53" lastPageNumber="7" pageId="6" pageNumber="7">
<subSubSection id="B1EE65274D18287FFF17FF42FD88F9C2" box="[151,569,151,178]" pageId="6" pageNumber="7" type="nomenclature">
<paragraph id="F94B36AC4D18287FFF17FF42FD88F9C2" blockId="6.[151,569,151,213]" box="[151,569,151,178]" pageId="6" pageNumber="7">
<heading id="A20381C04D18287FFF17FF42FD88F9C2" bold="true" box="[151,569,151,178]" fontSize="11" level="1" pageId="6" pageNumber="7" reason="1">
<taxonomicName id="3EF44D2F4D18287FFF17FF42FD88F9C2" ID-CoL="77NWW" authority="Coddington, 1986" authorityName="Coddington" authorityYear="1986" box="[151,569,151,178]" class="Arachnida" family="Theridiosomatidae" genus="Plato" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="troglodita">
<emphasis id="CB80EABE4D18287FFF17FF42FD88F9C2" bold="true" box="[151,569,151,178]" pageId="6" pageNumber="7">
<emphasis id="CB80EABE4D18287FFF17FF42FEE2F9C1" bold="true" box="[151,339,151,178]" italics="true" pageId="6" pageNumber="7">Plato troglodita</emphasis>
<bibRefCitation id="9D654B5D4D18287FFEDBFF42FD88F9C2" author="Coddington, J. A." box="[347,569,151,177]" pageId="6" pageNumber="7" pagination="1 - 96" refId="ref15738" refString="Coddington, J. A. (1986) The genera of the spider family Theridiosomatidae. S mithsonian Contributions to Zoology, 422, 1 - 96. https: // doi. org / 10.5479 / si. 00810282.422" type="journal article" year="1986">Coddington, 1986</bibRefCitation>
</emphasis>
</taxonomicName>
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</paragraph>
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<subSubSection id="B1EE65274D18287FFF17FF6EFEF7F9A6" box="[151,326,187,213]" pageId="6" pageNumber="7" type="description">
<paragraph id="F94B36AC4D18287FFF17FF6EFEF7F9A6" blockId="6.[151,569,151,213]" box="[151,326,187,213]" pageId="6" pageNumber="7">
<figureCitation id="61CF2A294D18287FFF17FF6EFEB0F9A6" box="[151,257,187,213]" captionStart="FIGURE 3" captionStartId="4.[151,250,1904,1928]" captionTargetBox="[164,1423,181,1881]" captionTargetId="figure-22@4.[164,1423,181,1881]" captionTargetPageId="4" captionText="FIGURE 3. Plato troglodita, from Morona-Santiago Province, Ecuador. A, male body, dorsal, illustration; B, C D, male palp, left, SEM; B, ventral view; C, mesal view, expanded; D, prolateral view (all adapted from Coddington (1986). Abbreviations, C, conductor. CA, conductor apophysis. Cy, cymbium. E, embolus. EA, embolic apophysis. MA, median apophysis. O, embolic opening. PC, paracymbium. ST, subtegulum. T, tegulum. Scales: A, 0.5; B, C, D, 0.1." figureDoi="http://doi.org/10.5281/zenodo.12189472" httpUri="https://zenodo.org/record/12189472/files/figure.png" pageId="6" pageNumber="7">Figures 3</figureCitation>
,
<figureCitation id="61CF2A294D18287FFE8EFF6EFEAFF9A6" box="[270,286,187,213]" captionStart="FIGURE 4" captionStartId="5.[151,250,1792,1816]" captionTargetBox="[151,1436,194,1769]" captionTargetId="figure-19@5.[151,1436,181,1769]" captionTargetPageId="5" captionText="FIGURE 4. Plato troglodita, from Morona-Santiago Province, Ecuador. A, female body, dorsal view; B–G, epigynum; B, ventral view; C, dorsal view; D, lateral view, clarified; E, lateral view, clarified; F, ventral view, SEM; G, anterior view, SEM. Abbreviations, At, atrium. CD, copulatory ducts. CP, central pit. DP, dorsal plate. FD, fertilization ducts. S, spermathecae. Su, epigynal support. TG, transversal groove. Scales: A, 2; B, 0.5; C, D, E 0.2; F, G, 0.1." figureDoi="http://doi.org/10.5281/zenodo.12189476" httpUri="https://zenodo.org/record/12189476/files/figure.png" pageId="6" pageNumber="7">4</figureCitation>
,
<figureCitation id="61CF2A294D18287FFEAAFF6EFEF7F9A6" box="[298,326,187,213]" captionStart="FIGURE 23" captionStartId="29.[151,250,1232,1256]" captionTargetBox="[151,1436,181,1208]" captionTargetId="figure-19@29.[151,1436,181,1208]" captionTargetPageId="29" captionText="FIGURE 23. Distribution of records of Plato (South America)." figureDoi="http://doi.org/10.5281/zenodo.12189530" httpUri="https://zenodo.org/record/12189530/files/figure.png" pageId="6" pageNumber="7">23</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="B1EE65274D18287FFF17FF2FFCB9F83C" pageId="6" pageNumber="7" type="reference_group">
<paragraph id="F94B36AC4D18287FFF17FF2FFCB9F83C" blockId="6.[151,1436,250,336]" pageId="6" pageNumber="7">
<treatmentCitationGroup id="D9E411824D18287FFF17FF2FFCB9F83C" pageId="6" pageNumber="7">
<treatmentCitation id="785510BD4D18287FFF17FF2FFDAAF861" author="Coddington, J. A." box="[151,539,250,274]" page="29" pageId="6" pageNumber="7" year="1986">
<taxonomicName id="3EF44D2F4D18287FFF17FF2FFDAAF861" ID-CoL="77NWW" authority="Coddington, 1986: 29" authorityName="Coddington" authorityPageNumber="29" authorityYear="1986" box="[151,539,250,274]" class="Arachnida" family="Theridiosomatidae" genus="Plato" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="troglodita">
<emphasis id="CB80EABE4D18287FFF17FF2FFE86F861" box="[151,311,250,274]" italics="true" pageId="6" pageNumber="7">Plato troglodita</emphasis>
<bibRefCitation id="9D654B5D4D18287FFEBDFF2FFDAAF861" author="Coddington, J. A." box="[317,539,250,274]" pageId="6" pageNumber="7" pagination="1 - 96" refId="ref15738" refString="Coddington, J. A. (1986) The genera of the spider family Theridiosomatidae. S mithsonian Contributions to Zoology, 422, 1 - 96. https: // doi. org / 10.5479 / si. 00810282.422" type="journal article" year="1986">Coddington, 1986: 29</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, figs. 10–12, 23–28 (male
<typeStatus id="264F880E4D18287FFCA5FF2FFCCFF861" box="[805,894,250,274]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
and female
<typeStatus id="264F880E4D18287FFC77FF2EFBFFF860" box="[1015,1102,251,275]" pageId="6" pageNumber="7" type="paratype">paratype</typeStatus>
from Cueva Los Tayos (
<geoCoordinate id="9CC0506B4D18287FFAC4FF2FFA29F861" box="[1348,1432,250,274]" degrees="03" direction="south" minutes="06" orientation="latitude" pageId="6" pageNumber="7" precision="925" value="-3.1">03º06’S</geoCoordinate>
,
<geoCoordinate id="9CC0506B4D18287FFF47FECCFE94F842" box="[199,293,281,305]" degrees="78" direction="west" minutes="12" orientation="longitude" pageId="6" pageNumber="7" precision="925" value="-78.2">78º12’W</geoCoordinate>
),
<collectingRegion id="3B30F84E4D18287FFEB8FECCFE5AF842" box="[312,491,281,305]" country="Ecuador" name="Morona-Santiago" pageId="6" pageNumber="7">Morona Santiago</collectingRegion>
,
<collectingCountry id="81E3763C4D18287FFE77FECCFDFCF842" box="[503,589,281,305]" name="Ecuador" pageId="6" pageNumber="7">Ecuador</collectingCountry>
, Ecuadorian British Los Tayos Expedition col., deposited in MCZ, not examined); Coddington, 1990: 19, fig. 104;
<treatmentCitation id="785510BD4D18287FFD8EFEEDFD73F823" author="Lopez, A." box="[526,706,311,336]" page="192" pageId="6" pageNumber="7" year="1996">
<bibRefCitation id="9D654B5D4D18287FFD8EFEEDFD73F823" author="Lopez, A." box="[526,706,311,336]" pageId="6" pageNumber="7" pagination="191 - 196" refId="ref16084" refString="Lopez, A. (1996) Plato juberthiei (Araneae: Theridiosomatidae), nouvelle araignee souterraine de la Guyana Francaise. Memoires de Biospeologie, 23, 191 - 196." type="journal article" year="1996">Lopez, 1996: 192</bibRefCitation>
</treatmentCitation>
, fig. 7.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="B1EE65274D18287FFF17FEA8FDDEF8CF" pageId="6" pageNumber="7" type="materials_examined">
<paragraph id="F94B36AC4D18287FFF17FEA8FDDEF8CF" blockId="6.[151,1437,381,1056]" pageId="6" pageNumber="7">
<emphasis id="CB80EABE4D18287FFF17FEA8FE65F8E4" bold="true" box="[151,468,381,407]" pageId="6" pageNumber="7">Other material examined.</emphasis>
<materialsCitation id="499C3CF14D18287FFE5FFEABFDDAF8CF" ID-GBIF-Occurrence="4897999316" collectingDate="1994-04-23" collectionCode="CAS" collectorName="V. Roth" country="Ecuador" latitude="-0.90861106" location="Caverna Oriente" longLatPrecision="1" longitude="-77.80972" municipality="Archidona" pageId="6" pageNumber="7" specimenCount="6" specimenCount-female="6" stateProvince="Napo">
<collectingCountry id="81E3763C4D18287FFE5FFEABFDDBF8EB" box="[479,618,382,408]" name="Ecuador" pageId="6" pageNumber="7">ECUADOR</collectingCountry>
.
<emphasis id="CB80EABE4D18287FFDF8FEAAFD04F8EB" box="[632,693,383,408]" italics="true" pageId="6" pageNumber="7">
<collectingRegion id="3B30F84E4D18287FFDF8FEAAFD04F8EB" box="[632,693,383,408]" country="Ecuador" name="Napo" pageId="6" pageNumber="7">Napo</collectingRegion>
</emphasis>
:
<collectingMunicipality id="192FACD64D18287FFD46FEABFC8EF8EB" box="[710,831,382,408]" pageId="6" pageNumber="7">Archidona</collectingMunicipality>
,
<location id="FC2B60774D18287FFCCFFEABFBBAF8EB" LSID="urn:lsid:plazi:treatment:715D87BA4D18287FFF17FF42FD0AFD53:FC2B60774D18287FFCCFFEABFBBAF8EB" box="[847,1035,382,408]" country="Ecuador" latitude="-0.90861106" longLatPrecision="1" longitude="-77.80972" municipality="Archidona" name="Caverna Oriente" pageId="6" pageNumber="7" stateProvince="Napo">Caverna Oriente</location>
(
<geoCoordinate id="9CC0506B4D18287FFB9EFEABFB19F8EA" box="[1054,1192,382,409]" degrees="0" direction="south" minutes="54" orientation="latitude" pageId="6" pageNumber="7" precision="1" seconds="31.0" value="-0.90861106">0°54'31.0&quot;S</geoCoordinate>
,
<geoCoordinate id="9CC0506B4D18287FFB38FEABFAEAF8EA" box="[1208,1371,382,409]" degrees="77" direction="west" minutes="48" orientation="longitude" pageId="6" pageNumber="7" precision="1" seconds="35.0" value="-77.80972">77°48'35.0&quot;W</geoCoordinate>
),
<specimenCount id="EFF2FD254D18287FFAF2FEABFA26F8EA" box="[1394,1431,382,409]" count="5" pageId="6" pageNumber="7" type="female">5♀</specimenCount>
,
<date id="8D4A106C4D18287FFF17FE77FEBEF8CF" box="[151,271,418,444]" pageId="6" pageNumber="7" value="1994-04-23">
<collectingDate id="9D0EE9844D18287FFF17FE77FEBEF8CF" box="[151,271,418,444]" pageId="6" pageNumber="7" value="1994-04-23">23.iv.1994</collectingDate>
</date>
,
<collectorName id="5401537A4D18287FFE9BFE77FEDEF8CF" box="[283,367,418,444]" pageId="6" pageNumber="7">V. Roth</collectorName>
col. (
<collectionCode id="9FE5AE694D18287FFE2FFE77FE5BF8CF" box="[431,490,418,444]" country="USA" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="6" pageNumber="7" type="Museum">CAS</collectionCode>
;
<specimenCount id="EFF2FD254D18287FFE75FE77FDA9F8CF" box="[501,536,418,444]" count="1" pageId="6" pageNumber="7" type="female">1♀</specimenCount>
MEV)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B1EE65274D18287FFF47FE10FDF2FB03" pageId="6" pageNumber="7" type="diagnosis">
<paragraph id="F94B36AC4D18287FFF47FE10FDF2FB03" blockId="6.[151,1437,381,1056]" pageId="6" pageNumber="7">
<emphasis id="CB80EABE4D18287FFF47FE10FEF3F8AC" bold="true" box="[199,322,453,479]" pageId="6" pageNumber="7">Diagnosis.</emphasis>
Male's palps resemble that of
<taxonomicName id="3EF44D2F4D18287FFD21FE13FC33F893" authority="Prete &amp; Brescovit, 2024" authorityName="Prete &amp; Brescovit" authorityYear="2024" box="[673,898,454,480]" class="Arachnida" family="Theridiosomatidae" genus="Plato" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="omnipraesens" status="sp. nov.">
<emphasis id="CB80EABE4D18287FFD21FE13FC33F893" box="[673,898,454,480]" italics="true" pageId="6" pageNumber="7">Plato omnipraesens</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D0B357C54D18287FFC0AFE13FC7BF893" box="[906,970,454,480]" pageId="6" pageNumber="7" rank="species">n. sp.</taxonomicNameLabel>
, especially the conductor apophysis (fig. 19B, C), but it differs by present a flattened conductor apophysis (fig. 3C), an elongated median apophysis and by the subrectrangular tip of embolus (fig. 3C). Females are recognized by the narrow, slightly curved transversal groove of the epigynum, with a support as thin as the transversal groove (fig. 4B, D), and the poorly sclerotized dorsal plate in the epigynum (fig. 4E).
</paragraph>
</subSubSection>
<subSubSection id="B1EE65274D18287FFF47FDACFB39FAC6" pageId="6" pageNumber="7" type="description">
<paragraph id="F94B36AC4D18287FFF47FDACFBBEFBE7" blockId="6.[151,1437,381,1056]" box="[199,1039,633,660]" pageId="6" pageNumber="7">
<emphasis id="CB80EABE4D18287FFF47FDACFEE8FBE0" bold="true" box="[199,345,633,659]" pageId="6" pageNumber="7">Description.</emphasis>
Male: see
<bibRefCitation id="9D654B5D4D18287FFE53FDAFFD1CFBE7" author="Coddington, J. A." box="[467,685,634,660]" pageId="6" pageNumber="7" pagination="1 - 96" refId="ref15738" refString="Coddington, J. A. (1986) The genera of the spider family Theridiosomatidae. S mithsonian Contributions to Zoology, 422, 1 - 96. https: // doi. org / 10.5479 / si. 00810282.422" type="journal article" year="1986">Coddington (1986)</bibRefCitation>
: 29, figs. 10–12, 23, 24, 27, 28.
</paragraph>
<paragraph id="F94B36AC4D18287FFF47FD4BFEF5FAE3" blockId="6.[151,1437,381,1056]" pageId="6" pageNumber="7">
Female (CAS). Carapace, legs, labium, endites and chelicerae brown. Sternum little darker than the rest of the prosoma. Abdomen gray, length: 1.86 Total length: 2.88. Carapace length: 1.32. Clypeus height: 0.24. Sternum:
<quantity id="3E0C9B494D18287FFF17FD33FF54FA73" box="[151,229,742,768]" metricMagnitude="-2" metricUnit="m" metricValue="1.8288" pageId="6" pageNumber="7" unit="in" value="0.72">0.72 in</quantity>
length and width. Eyes: AME 0.1; AME-AME 0.04; ALE-PLE contiguous (0.1 each); PME-PME separated by 0.09 (PME 0.1). Legs: I femur 1.55/ patella 0.57/ tibia 1.35/ metatarsus 0.82/ tarsus 0.68/ total 4.97; II 1.37/ 0.52/ 1.15/ 0.77/ 0.7/ 4.51; III 1/ 0.45/ 0.75/ 0.57/ 0.52/ 3.29; IV 1.25/ 0.37/ 0.8/ 0.55/ 0.51/ 3.48. Epigynum: dorsal plate posterior to spermathecae (fig. 4E). Copulatory ducts with a rounded curve in the anterior region (fig. 4C). Abdomen oval.
</paragraph>
<paragraph id="F94B36AC4D18287FFF47FC4CFB39FAC6" blockId="6.[151,1437,381,1056]" box="[199,1160,921,949]" pageId="6" pageNumber="7">
<emphasis id="CB80EABE4D18287FFF47FC4CFE8EFAC0" bold="true" box="[199,319,921,947]" pageId="6" pageNumber="7">Variation.</emphasis>
<specimenCount id="EFF2FD254D18287FFEC6FC4FFE06FAC7" box="[326,439,922,949]" count="5" pageId="6" pageNumber="7" type="female">5 females</specimenCount>
: total length: 2.48–2.88; carapace: 1.15–1.32; femur I: 1.4–1.75.
</paragraph>
</subSubSection>
<subSubSection id="B1EE65274D18287FFF47FC68FD0AFD53" pageId="6" pageNumber="7" type="distribution">
<paragraph id="F94B36AC4D18287FFF47FC68FB00FAAB" blockId="6.[151,1437,381,1056]" box="[199,1201,957,984]" pageId="6" pageNumber="7">
<emphasis id="CB80EABE4D18287FFF47FC68FED0FAA4" bold="true" box="[199,353,957,983]" pageId="6" pageNumber="7">Distribution.</emphasis>
Caves in the departments of
<collectingRegion id="3B30F84E4D18287FFD27FC6BFCD8FAAB" box="[679,873,958,984]" country="Ecuador" name="Morona-Santiago" pageId="6" pageNumber="7">Morona Santiago</collectingRegion>
and
<collectingRegion id="3B30F84E4D18287FFC1FFC6BFC6FFAAB" box="[927,990,958,984]" country="Ecuador" name="Napo" pageId="6" pageNumber="7">Napo</collectingRegion>
,
<collectingCountry id="81E3763C4D18287FFC6AFC6BFBF6FAAB" box="[1002,1095,958,984]" name="Ecuador" pageId="6" pageNumber="7">Ecuador</collectingCountry>
(fig. 23).
</paragraph>
<paragraph id="F94B36AC4D18287FFF47FC37FD0AFD53" blockId="6.[151,1437,381,1056]" pageId="6" pageNumber="7">
<emphasis id="CB80EABE4D18287FFF47FC37FEB7FA8F" bold="true" box="[199,262,994,1020]" pageId="6" pageNumber="7">Note.</emphasis>
Male of
<taxonomicName id="3EF44D2F4D18287FFEF3FC36FDB1FA8F" authorityName="Coddington" authorityYear="1986" box="[371,512,994,1020]" class="Arachnida" family="Theridiosomatidae" genus="Plato" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="troglodita">
<emphasis id="CB80EABE4D18287FFEF3FC36FDB1FA8F" box="[371,512,994,1020]" italics="true" pageId="6" pageNumber="7">P. troglodita</emphasis>
</taxonomicName>
was examined only by photos and illustrations presented by
<bibRefCitation id="9D654B5D4D18287FFB44FC37FA2DFA8F" author="Coddington, J. A." box="[1220,1436,994,1020]" pageId="6" pageNumber="7" pagination="1 - 96" refId="ref15738" refString="Coddington, J. A. (1986) The genera of the spider family Theridiosomatidae. S mithsonian Contributions to Zoology, 422, 1 - 96. https: // doi. org / 10.5479 / si. 00810282.422" type="journal article" year="1986">Coddington (1986)</bibRefCitation>
because there was no access to the
<typeStatus id="264F880E4D18287FFDA0FBD3FDE1FD53" box="[544,592,1030,1056]" pageId="6" pageNumber="7">type</typeStatus>
material.
</paragraph>
</subSubSection>
</treatment>
</document>
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