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<document id="0AD31FC880E7D7069C2CC503ECC550A8" ID-CLB-Dataset="22061" ID-DOI="10.5852/ejt.2017.347" ID-GBIF-Dataset="6f1a06d8-dbae-462e-8415-0cb51016c64a" ID-ISSN="2118-9773" ID-Zenodo-Dep="3832630" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589831392081" checkinUser="carolina" docAuthor="Berning, Björn, Harmelin, Jean-Georges &amp; Bader, Beate" docDate="2017" docId="546F87A1FF98FFBC092090FD343FFD28" docLanguage="en" docName="ejt-2017-347.pdf.imf" docOrigin="European Journal of Taxonomy 347" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Calvetopora otapostasis Berning &amp; Harmelin &amp; Bader &amp; Cibio 2017, gen. et sp. nov." docType="treatment" docUuid="701FB911-9BE3-45A4-88A6-F6BB1E47CEFD" docUuidSource="ZooBank" docVersion="8" lastPageNumber="43" masterDocId="A856FFD9FFBFFF960B0F95243731FFCE" masterDocTitle="New Cheilostomata (Bryozoa) from NE Atlantic seamounts, islands, and the continental slope: evidence for deep-sea endemism" masterLastPageNumber="51" masterPageNumber="1" pageNumber="40" updateTime="1698840148996" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="3934F2A7EE207AF6BC6672C019085E5A">New Cheilostomata (Bryozoa) from NE Atlantic seamounts, islands, and the continental slope: evidence for deep-sea endemism</mods:title>
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<mods:namePart id="1ACB80677610C6B5E8D530C28BE789CA">Berning, Björn</mods:namePart>
<mods:nameIdentifier id="62AD2FE3153594718E74B8AD2F235E18" type="LSID">30D7D0DB-F379-4006-B727-E75A0720BD93</mods:nameIdentifier>
<mods:affiliation id="1277BB9C076038349FDE3A4E1A2517B3">Oberösterreichisches Landesmuseum, Geowissenschaftliche Sammlungen, 4060 Leonding, Austria. &amp; CIBIO, Centro de Investigação em Biodiversidade e Recursos Genéticos, InBIO Laboratório Associado, Pólo dos Açores, 9501 - 801 Ponta Delgada, Açores, Portugal. &amp; urn: lsid: zoobank. org: author: 30 D 7 D 0 DB-F 379 - 4006 - B 727 - E 75 A 0720 BD 93 &amp; Corresponding author: b. berning @ landesmuseum. at</mods:affiliation>
<mods:nameIdentifier id="E7976236C86938BE3475B7E462C3E0D0" type="email">b.berning@landesmuseum.at</mods:nameIdentifier>
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<mods:roleTerm id="8373C1942E960264B4BABC5FCF20F17B">Author</mods:roleTerm>
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<mods:namePart id="562DC9FC3B5F35CEA7EAC987B75C7223">Harmelin, Jean-Georges</mods:namePart>
<mods:nameIdentifier id="D8224AA38341F3F950719A4C14008EFB" type="LSID">D11AE07A-CFD9-41EE-B3F9-6E0472150300</mods:nameIdentifier>
<mods:affiliation id="4F8274D3636418FC08F42D4DD3815A26">Aix-Marseille University, Mediterranean Institute of Oceanography, OSU Pytheas, Station Marine d’Endoume, 13007 Marseille, France. &amp; Email: jean-georges. harmelin @ univ-amu. fr &amp; urn: lsid: zoobank. org: author: D 11 AE 07 A-CFD 9 - 41 EE-B 3 F 9 - 6 E 0472150300</mods:affiliation>
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<mods:roleTerm id="40DED6FB7B09FDE9A9C46DE3B1FCA53A">Author</mods:roleTerm>
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<mods:namePart id="13FD5575ECE9313EAB59E7A60E6ABB3A">Bader, Beate</mods:namePart>
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<mods:affiliation id="9DF4BF5E586D81DA2E33CBBDE85C65B8">Institut für Geowissenschaften, Christian-Albrechts-Universität, 24118 Kiel, Germany. &amp; Email: bbader @ online. no &amp; urn: lsid: zoobank. org: author: AA 3 BCFDC- 524 D- 4648 - 9268 - F 0 F 1 C 94 B 9 A 68</mods:affiliation>
<mods:nameIdentifier id="871A4133726C579900EF0156DF053CCC" type="email">bbader@online.no</mods:nameIdentifier>
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<mods:title id="95797CBA146C5C1F2CF7708C22D72578">European Journal of Taxonomy</mods:title>
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<mods:date id="5B711D56AECF0A94C0C1E48F36D0B09B">2017</mods:date>
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<mods:number id="9C636EB7EAF434AE3C1E22B56CE1709D">2017-08-31</mods:number>
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<treatment id="546F87A1FF98FFBC092090FD343FFD28" ID-DOI="http://doi.org/10.5281/zenodo.3850651" ID-GBIF-Taxon="164167292" ID-Zenodo-Dep="3850651" LSID="urn:lsid:zoobank.org:act:701FB911-9BE3-45A4-88A6-F6BB1E47CEFD" httpUri="http://treatment.plazi.org/id/546F87A1FF98FFBC092090FD343FFD28" lastPageId="42" lastPageNumber="43" pageId="39" pageNumber="40">
<subSubSection id="94DC653CFF98FFB1092090FD3334FA3A" box="[559,1029,1497,1524]" pageId="39" pageNumber="40" type="nomenclature">
<paragraph id="DC7936B7FF98FFB1092090FD3334FA3A" blockId="39.[364,1223,1497,1594]" box="[559,1029,1497,1524]" pageId="39" pageNumber="40">
<heading id="873181DBFF98FFB1092090FD3334FA3A" box="[559,1029,1497,1524]" centered="true" fontSize="11" level="2" pageId="39" pageNumber="40" reason="2">
<taxonomicName id="1BC64D34FF98FFB1092090FD347CFA3D" authority="Berning &amp; Harmelin &amp; Bader &amp; Cibio, 2017" authorityName="Berning &amp; Harmelin &amp; Bader &amp; Cibio" authorityYear="2017" box="[559,845,1497,1524]" class="Gymnolaemata" family="Atlantisinidae" genus="Calvetopora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="39" pageNumber="40" phylum="Bryozoa" rank="species" species="otapostasis" status="gen. et sp. nov.">
<emphasis id="EEB2EAA5FF98FFB1092090FD347CFA3D" bold="true" box="[559,845,1497,1524]" italics="true" pageId="39" pageNumber="40">Calvetopora otapostasis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F58157DEFF98FFB1085A90FE3334FA3A" box="[853,1029,1498,1524]" pageId="39" pageNumber="40" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="94DC653CFF98FFB10A6390D834A7F9F4" pageId="39" pageNumber="40" type="description">
<paragraph id="DC7936B7FF98FFB10A6390D833F6F9D9" blockId="39.[364,1223,1497,1594]" box="[364,1223,1532,1559]" pageId="39" pageNumber="40">
<uri id="A8573AB5FF98FFB10A6390D833F6F9D9" box="[364,1223,1532,1559]" pageId="39" pageNumber="40">
urn:lsid:zoobank.org:act:
<uuid id="A8600C62FF98FFB1098090D833F6F9D9" box="[655,1223,1532,1559]" pageId="39" pageNumber="40">701FB911-9BE3-45A4-88A6-F6BB1E47CEFD</uuid>
</uri>
</paragraph>
<paragraph id="DC7936B7FF98FFB10992930434A7F9F4" blockId="39.[364,1223,1497,1594]" box="[669,918,1568,1594]" pageId="39" pageNumber="40">
<figureCitation id="44FD2A32FF98FFB10992930435DFF9F4" box="[669,750,1568,1594]" captionStart="Fig" captionStartId="40.[189,232,1652,1678]" captionTargetBox="[189,1399,696,1611]" captionTargetId="figure@40.[189,1399,696,1611]" captionTargetPageId="40" captionText="Fig. 14. Calvetopora otapostasis gen. et sp. nov., Atlantis Smt. A. Overview of the holotype (MNHN- IB-2014-78). B. Close-up of an ooecium and the suboral avicularia (holotype MNHN-IB-2014-78). C. Slightly oblique view of an autozooid at the growth margin showing the communication pores in the lateral walls as well as marginal areolar pores (black arrow) and the roughly crescentically arranged pseudopores (white arrows) in the frontal shield (paratype MNHN-IB-2014-280). D. Close-up of orifice; note that the condyles are so short that usually only one can be seen (paratype OLL 2016/153). E. Ancestrula and early astogenetic autozooids (paratype MNHN-IB-2014-81). F. Interior frontal shield showing the lateral areolar pores and the central pseudopores (paratype OLL 2016/153). Scale bars: A = 500 µm; B, D, F = 100 µm; C, E = 200 µm." figureDoi="http://doi.org/10.5281/zenodo.3832658" httpUri="https://zenodo.org/record/3832658/files/figure.png" pageId="39" pageNumber="40">Fig. 14</figureCitation>
A–F,
<tableCitation id="9144030CFF98FFB1083E930434A7F9F4" box="[817,918,1568,1594]" captionStart="Table 14" captionStartId="41.[189,258,268,294]" captionTargetPageId="41" captionText="Table 14. Measurements of Calvetopora otapostasis gen. et sp. nov." pageId="39" pageNumber="40">Table 14</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="94DC653CFF98FFB10BB29346330AF8D8" pageId="39" pageNumber="40" type="diagnosis">
<paragraph id="DC7936B7FF98FFB10BB293463607F9B2" blockId="39.[189,310,1634,1660]" box="[189,310,1634,1660]" pageId="39" pageNumber="40">
<heading id="873181DBFF98FFB10BB293463607F9B2" bold="true" box="[189,310,1634,1660]" fontSize="11" level="3" pageId="39" pageNumber="40" reason="3">
<emphasis id="EEB2EAA5FF98FFB10BB293463607F9B2" bold="true" box="[189,310,1634,1660]" pageId="39" pageNumber="40">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="DC7936B7FF98FFB10BB293B5330AF8D8" blockId="39.[189,1399,1680,1814]" pageId="39" pageNumber="40">
<taxonomicName id="1BC64D34FF98FFB10BB293B536ECF964" authority="Berning &amp; Harmelin &amp; Bader &amp; Cibio, 2017" authorityName="Berning &amp; Harmelin &amp; Bader &amp; Cibio" authorityYear="2017" box="[189,477,1681,1707]" class="Gymnolaemata" family="Atlantisinidae" genus="Calvetopora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="39" pageNumber="40" phylum="Bryozoa" rank="species" species="otapostasis" status="gen. et sp. nov.">
<emphasis id="EEB2EAA5FF98FFB10BB293B536ECF964" box="[189,477,1681,1707]" italics="true" pageId="39" pageNumber="40">Calvetopora otapostasis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F58157DEFF98FFB10AE993B535ADF965" box="[486,668,1681,1707]" pageId="39" pageNumber="40" rank="species">gen. et sp. nov.</taxonomicNameLabel>
differs from the
<typeStatus id="037D8815FF98FFB1086693B534ADF965" box="[873,924,1681,1707]" pageId="39" pageNumber="40">type</typeStatus>
species
<taxonomicName id="1BC64D34FF98FFB10F0D93B53340F965" authorityName="Berning &amp; Harmelin &amp; Bader &amp; Cibio" authorityYear="2017" baseAuthorityName="Calvet" baseAuthorityYear="1906" box="[1026,1137,1681,1707]" class="Gymnolaemata" family="Atlantisinidae" genus="Calvetopora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="39" pageNumber="40" phylum="Bryozoa" rank="species" species="inflata" status="gen. et comb. nov.">
<emphasis id="EEB2EAA5FF98FFB10F0D93B53340F965" box="[1026,1137,1681,1707]" italics="true" pageId="39" pageNumber="40">C. inflata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F58157DEFF98FFB10F7593B53266F965" box="[1146,1367,1680,1707]" pageId="39" pageNumber="40" rank="species">gen. et comb. nov.</taxonomicNameLabel>
in having distinctly larger zooids, an orifice that is longer than wide and has only three to six oral spines, in the frontal shield with its granular surface structure and a higher number of central pseudopores, and in the ooecium, which is covered by a calcified ectooecium in its lower part.
</paragraph>
</subSubSection>
<subSubSection id="94DC653CFF98FFB10BB29219359BF867" pageId="39" pageNumber="40" type="etymology">
<paragraph id="DC7936B7FF98FFB10BB292193675F899" blockId="39.[189,324,1853,1879]" box="[189,324,1853,1879]" pageId="39" pageNumber="40">
<heading id="873181DBFF98FFB10BB292193675F899" bold="true" box="[189,324,1853,1879]" fontSize="11" level="3" pageId="39" pageNumber="40" reason="3">
<emphasis id="EEB2EAA5FF98FFB10BB292193675F899" bold="true" box="[189,324,1853,1879]" pageId="39" pageNumber="40">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="DC7936B7FF98FFB10BB2924F359BF867" blockId="39.[189,1399,1899,1962]" pageId="39" pageNumber="40">The name alludes to the two avicularia lateral to the orifice, which look like protruding ears (Latin: otapostasis); used as a noun in apposition.</paragraph>
</subSubSection>
<subSubSection id="94DC653CFF97FFBF0BB2942F3352FD3C" lastPageId="41" lastPageNumber="42" pageId="40" pageNumber="41" type="materials_examined">
<paragraph id="DC7936B7FF97FFBE0BB2942F369BFEEB" blockId="40.[189,426,267,293]" box="[189,426,267,293]" pageId="40" pageNumber="41">
<heading id="873181DBFF97FFBE0BB2942F369BFEEB" bold="true" box="[189,426,267,293]" fontSize="11" level="3" pageId="40" pageNumber="41" reason="3">
<emphasis id="EEB2EAA5FF97FFBE0BB2942F369BFEEB" bold="true" box="[189,426,267,293]" pageId="40" pageNumber="41">Material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="6CAE3CEAFF97FFBE0BE5941E33CCFEB9" ID-GBIF-Occurrence="2620523306" collectingDate="2014" collectingDateMax="2078" collectingDateMin="2014" collectionCode="SMT" pageId="40" pageNumber="41" specimenCode="MNHN-IB-2014-78" specimenCount="1" typeStatus="holotype">
<paragraph id="DC7936B7FF97FFBE0BE5941E366DFE9A" blockId="40.[189,1277,314,375]" box="[234,348,314,340]" pageId="40" pageNumber="41">
<emphasis id="EEB2EAA5FF97FFBE0BE5941E366DFE9A" bold="true" box="[234,348,314,340]" pageId="40" pageNumber="41">
<typeStatus id="037D8815FF97FFBE0BE5941E366DFE9A" box="[234,348,314,340]" pageId="40" pageNumber="41" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="DC7936B7FF97FFBE0BB2947933CCFEB9" blockId="40.[189,1277,314,375]" box="[189,1277,348,375]" pageId="40" pageNumber="41">
ATLANTIS SMT: 1 ovicellate colony on sandstone substratum, Stn 7 (
<specimenCode id="8C609ECCFF97FFBE08F2947933C1FEB9" box="[1021,1264,349,375]" pageId="40" pageNumber="41">MNHN-IB-2014-78</specimenCode>
).
</paragraph>
</materialsCitation>
<paragraph id="DC7936B7FF97FFBE0BE594BA3656FE76" blockId="40.[189,1399,414,617]" box="[234,359,414,440]" pageId="40" pageNumber="41">
<emphasis id="EEB2EAA5FF97FFBE0BE594BA3656FE76" bold="true" box="[234,359,414,440]" pageId="40" pageNumber="41">
<typeStatus id="037D8815FF97FFBE0BE594BA3656FE76" box="[234,359,414,440]" pageId="40" pageNumber="41" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="DC7936B7FF97FFBE0BB294E5334FFDA6" blockId="40.[189,1399,414,617]" pageId="40" pageNumber="41">
ATLANTIS
<collectionCode id="BAD7AE72FF97FFBE0A5F94E536BEFE15" box="[336,399,449,475]" pageId="40" pageNumber="41">SMT</collectionCode>
: 1 colony on stylasterid skeleton, Stn 7 (MNHN-IB-
<date id="A8781077FF97FFBE08FD94E53364FE15" box="[1010,1109,449,475]" pageId="40" pageNumber="41" value="2014" valueMax="2079">2014-79</date>
); 1 isolated colony, Stn 7 (MNHN-IB-
<date id="A8781077FF97FFBE0A5994C13689FE31" box="[342,440,485,511]" pageId="40" pageNumber="41" value="2014" valueMax="2080">2014-80</date>
); 1 colony on stylasterid skeleton showing the ancestrula and early astogenesis, Stn 7 (MNHN-IB-
<date id="A8781077FF97FFBE0AA9972C3536FDEC" box="[422,519,520,546]" pageId="40" pageNumber="41" value="2014" valueMax="2081">2014-81</date>
); 1 coated colony on stylasterid skeleton, Stn 7 (MNHN-IB-2014-280); 3 colonies on stylasterid skeleton, Stn 7 (
<collectionCode id="BAD7AE72FF97FFBE09AE970F35EAFD8B" box="[673,731,555,581]" pageId="40" pageNumber="41">OLL</collectionCode>
2016/127); 1 colony on stylasterid skeleton, Stn 7 (
<collectionCode id="BAD7AE72FF97FFBE0E32970F3246FD8B" box="[1341,1399,555,581]" pageId="40" pageNumber="41">OLL</collectionCode>
2016/153); 1 ovicellate colony on
<taxonomicName id="1BC64D34FF97FFBE095F976A35CBFDA6" authorityName="de Blainville" authorityYear="1830" box="[592,762,590,616]" class="Anthozoa" family="Dendrophylliidae" genus="Dendrophyllia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scleractinia" pageId="40" pageNumber="41" phylum="Cnidaria" rank="genus">
<emphasis id="EEB2EAA5FF97FFBE095F976A35CBFDA6" box="[592,762,590,616]" italics="true" pageId="40" pageNumber="41">Dendrophyllia</emphasis>
</taxonomicName>
sp., Stn 3 (MNHN-IB-
<date id="A8781077FF97FFBE0F1F976A3343FDA6" box="[1040,1138,590,616]" pageId="40" pageNumber="41" value="2014" valueMax="2082">2014-82</date>
).
</paragraph>
<caption id="88B9663FFF97FFBE0BB29350358FF867" ID-DOI="http://doi.org/10.5281/zenodo.3832658" ID-Zenodo-Dep="3832658" httpUri="https://zenodo.org/record/3832658/files/figure.png" pageId="40" pageNumber="41" startId="40.[189,232,1652,1678]" targetBox="[189,1399,696,1611]" targetPageId="40">
<paragraph id="DC7936B7FF97FFBE0BB29350358FF867" blockId="40.[189,1399,1652,1961]" pageId="40" pageNumber="41">
<emphasis id="EEB2EAA5FF97FFBE0BB29350362BF940" bold="true" box="[189,282,1652,1678]" pageId="40" pageNumber="41">Fig. 14.</emphasis>
<taxonomicName id="1BC64D34FF97FFBE0A2D93513570F940" authority="Berning &amp; Harmelin &amp; Bader &amp; Cibio, 2017" authorityName="Berning &amp; Harmelin &amp; Bader &amp; Cibio" authorityYear="2017" box="[290,577,1653,1679]" class="Gymnolaemata" family="Atlantisinidae" genus="Calvetopora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="40" pageNumber="41" phylum="Bryozoa" rank="species" species="otapostasis" status="gen. et sp. nov.">
<emphasis id="EEB2EAA5FF97FFBE0A2D93513570F940" box="[290,577,1653,1679]" italics="true" pageId="40" pageNumber="41">Calvetopora otapostasis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F58157DEFF97FFBE0946935135CFF941" box="[585,766,1653,1679]" pageId="40" pageNumber="41" rank="species">gen. et sp. nov.</taxonomicNameLabel>
, Atlantis Smt.
<emphasis id="EEB2EAA5FF97FFBE08A2935034F3F940" bold="true" box="[941,962,1652,1678]" pageId="40" pageNumber="41">A</emphasis>
. Overview of the holotype (MNHN- IB-2014-78).
<emphasis id="EEB2EAA5FF97FFBE0A6D93BC3647F97C" bold="true" box="[354,374,1688,1714]" pageId="40" pageNumber="41">B</emphasis>
. Close-up of an ooecium and the suboral avicularia (holotype MNHN-IB-2014-78).
<emphasis id="EEB2EAA5FF97FFBE0BB2939F37E3F91B" bold="true" box="[189,210,1723,1749]" pageId="40" pageNumber="41">C</emphasis>
. Slightly oblique view of an autozooid at the growth margin showing the communication pores in the lateral walls as well as marginal areolar pores (black arrow) and the roughly crescentically arranged pseudopores (white arrows) in the frontal shield (paratype MNHN-IB-2014-280).
<emphasis id="EEB2EAA5FF97FFBE0FB3922633E0F8D2" bold="true" box="[1212,1233,1794,1820]" pageId="40" pageNumber="41">D</emphasis>
. Close-up of orifice; note that the condyles are so short that usually only one can be seen (paratype OLL 2016/153).
<emphasis id="EEB2EAA5FF97FFBE0BB2926C37E0F8AC" bold="true" box="[189,209,1864,1890]" pageId="40" pageNumber="41">E</emphasis>
. Ancestrula and early astogenetic autozooids (paratype MNHN-IB-2014-81).
<emphasis id="EEB2EAA5FF97FFBE0F52926C335EF8AC" bold="true" box="[1117,1135,1864,1890]" pageId="40" pageNumber="41">F</emphasis>
. Interior frontal shield showing the lateral areolar pores and the central pseudopores (paratype OLL 2016/153). Scale bars: A = 500 µm; B, D, F = 100 µm; C, E = 200 µm.
</paragraph>
</caption>
<caption id="88B9663FFF96FFBF0BB2942834ECFEE8" box="[189,989,267,294]" pageId="41" pageNumber="42" targetBox="[215,1357,352,486]" targetIsTable="true" targetPageId="41">
<paragraph id="DC7936B7FF96FFBF0BB2942834ECFEE8" blockId="41.[189,989,267,294]" box="[189,989,267,294]" pageId="41" pageNumber="42">
<emphasis id="EEB2EAA5FF96FFBF0BB29428361FFEEB" bold="true" box="[189,302,267,294]" pageId="41" pageNumber="42">Table 14.</emphasis>
Measurements of
<taxonomicName id="1BC64D34FF96FFBF0908942F3414FEEB" authority="Berning &amp; Harmelin &amp; Bader &amp; Cibio, 2017" authorityName="Berning &amp; Harmelin &amp; Bader &amp; Cibio" authorityYear="2017" box="[519,805,267,293]" class="Gymnolaemata" family="Atlantisinidae" genus="Calvetopora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="41" pageNumber="42" phylum="Bryozoa" rank="species" species="otapostasis" status="gen. et sp. nov.">
<emphasis id="EEB2EAA5FF96FFBF0908942F3414FEEB" box="[519,805,267,293]" italics="true" pageId="41" pageNumber="42">Calvetopora otapostasis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F58157DEFF96FFBF0822942834ECFEE8" box="[813,989,268,294]" pageId="41" pageNumber="42" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="DC7936B7FF96FFBF0A6A9445320EFE2B" pageId="41" pageNumber="42">
<table id="AEC6C417FF9600690BD89444327CFE28" box="[215,1357,352,486]" gridcols="9" gridrows="4" pageId="41" pageNumber="42" rowsContinueFrom="39.[215,1357,349,482]" rowsContinueIn="43.[239,1340,322,455]">
<tr id="62F634F5FF9600690BD89444327CFEB5" box="[215,1357,352,379]" gridrow="0" pageId="41" pageNumber="42" rowspan-0="1">
<th id="21275D89FF9600690A54944436A7FEB5" box="[347,406,352,379]" gridcol="1" gridrow="0" pageId="41" pageNumber="42">
<emphasis id="EEB2EAA5FF96FFBF0A6A944536BDFEB5" bold="true" box="[357,396,353,379]" pageId="41" pageNumber="42">ZL</emphasis>
</th>
<th id="21275D89FF9600690AD094443510FEB5" box="[479,545,352,379]" gridcol="2" gridrow="0" pageId="41" pageNumber="42">
<emphasis id="EEB2EAA5FF96FFBF0AE794453528FEB5" bold="true" box="[488,537,353,379]" pageId="41" pageNumber="42">ZW</emphasis>
</th>
<th id="21275D89FF960069097D944435AFFEB5" box="[626,670,352,379]" gridcol="3" gridrow="0" pageId="41" pageNumber="42">
<emphasis id="EEB2EAA5FF96FFBF097B944435AFFEB4" bold="true" box="[628,670,352,378]" pageId="41" pageNumber="42">OL</emphasis>
</th>
<th id="21275D89FF96006909F99444341BFEB5" box="[758,810,352,379]" gridcol="4" gridrow="0" pageId="41" pageNumber="42">
<emphasis id="EEB2EAA5FF96FFBF09F99444341BFEB4" bold="true" box="[758,810,352,378]" pageId="41" pageNumber="42">OW</emphasis>
</th>
<th id="21275D89FF960069087294443487FEB5" box="[893,950,352,379]" gridcol="5" gridrow="0" pageId="41" pageNumber="42">
<emphasis id="EEB2EAA5FF96FFBF087294443487FEB4" bold="true" box="[893,950,352,378]" pageId="41" pageNumber="42">OvL</emphasis>
</th>
<th id="21275D89FF96006908F094443373FEB5" box="[1023,1090,352,379]" gridcol="6" gridrow="0" pageId="41" pageNumber="42">
<emphasis id="EEB2EAA5FF96FFBF08F094443373FEB4" bold="true" box="[1023,1090,352,378]" pageId="41" pageNumber="42">OvW</emphasis>
</th>
<th id="21275D89FF9600690F81944433F4FEB5" box="[1166,1221,352,379]" gridcol="7" gridrow="0" pageId="41" pageNumber="42">
<emphasis id="EEB2EAA5FF96FFBF0F81944433F4FEB4" bold="true" box="[1166,1221,352,378]" pageId="41" pageNumber="42">aAL</emphasis>
</th>
<th id="21275D89FF9600690E1D9444327CFEB5" box="[1298,1357,352,379]" gridcol="8" gridrow="0" pageId="41" pageNumber="42">
<emphasis id="EEB2EAA5FF96FFBF0E1D9444327CFEB4" bold="true" box="[1298,1357,352,378]" pageId="41" pageNumber="42">aAW</emphasis>
</th>
</tr>
<tr id="62F634F5FF9600690BD894A0327CFE51" box="[215,1357,388,415]" gridrow="1" pageId="41" pageNumber="42">
<th id="21275D89FF9600690BD894A0362BFE51" box="[215,282,388,415]" gridcol="0" gridrow="1" pageId="41" pageNumber="42">Mean</th>
<td id="21275D89FF9600690A5494A036A7FE51" box="[347,406,388,415]" gridcol="1" gridrow="1" pageId="41" pageNumber="42">1081</td>
<td id="21275D89FF9600690AD094A03510FE51" box="[479,545,388,415]" gridcol="2" gridrow="1" pageId="41" pageNumber="42">844</td>
<td id="21275D89FF960069097D94A035AFFE51" box="[626,670,388,415]" gridcol="3" gridrow="1" pageId="41" pageNumber="42">222</td>
<td id="21275D89FF96006909F994A0341BFE51" box="[758,810,388,415]" gridcol="4" gridrow="1" pageId="41" pageNumber="42">195</td>
<td id="21275D89FF960069087294A03487FE51" box="[893,950,388,415]" gridcol="5" gridrow="1" pageId="41" pageNumber="42">283</td>
<td id="21275D89FF96006908F094A03373FE51" box="[1023,1090,388,415]" gridcol="6" gridrow="1" pageId="41" pageNumber="42">291</td>
<td id="21275D89FF9600690F8194A033F4FE51" box="[1166,1221,388,415]" gridcol="7" gridrow="1" pageId="41" pageNumber="42">134</td>
<td id="21275D89FF9600690E1D94A0327CFE51" box="[1298,1357,388,415]" gridcol="8" gridrow="1" pageId="41" pageNumber="42">93</td>
</tr>
<tr id="62F634F5FF9600690BD89483327CFE0C" box="[215,1357,423,450]" gridrow="2" pageId="41" pageNumber="42">
<th id="21275D89FF9600690BD89483362BFE0C" box="[215,282,423,450]" gridcol="0" gridrow="2" pageId="41" pageNumber="42">SD</th>
<td id="21275D89FF9600690A54948336A7FE0C" box="[347,406,423,450]" gridcol="1" gridrow="2" pageId="41" pageNumber="42">± 97</td>
<td id="21275D89FF9600690AD094833510FE0C" box="[479,545,423,450]" gridcol="2" gridrow="2" pageId="41" pageNumber="42">± 115</td>
<td id="21275D89FF960069097D948335AFFE0C" box="[626,670,423,450]" gridcol="3" gridrow="2" pageId="41" pageNumber="42">± 8</td>
<td id="21275D89FF96006909F99483341BFE0C" box="[758,810,423,450]" gridcol="4" gridrow="2" pageId="41" pageNumber="42">± 7</td>
<td id="21275D89FF960069087294833487FE0C" box="[893,950,423,450]" gridcol="5" gridrow="2" pageId="41" pageNumber="42">± 17</td>
<td id="21275D89FF96006908F094833373FE0C" box="[1023,1090,423,450]" gridcol="6" gridrow="2" pageId="41" pageNumber="42">± 13</td>
<td id="21275D89FF9600690F81948333F4FE0C" box="[1166,1221,423,450]" gridcol="7" gridrow="2" pageId="41" pageNumber="42">± 22</td>
<td id="21275D89FF9600690E1D9483327CFE0C" box="[1298,1357,423,450]" gridcol="8" gridrow="2" pageId="41" pageNumber="42">± 13</td>
</tr>
<tr id="62F634F5FF9600690BD894EF327CFE28" box="[215,1357,459,486]" gridrow="3" pageId="41" pageNumber="42">
<th id="21275D89FF9600690BD894EF362BFE28" box="[215,282,459,486]" gridcol="0" gridrow="3" pageId="41" pageNumber="42">#</th>
<td id="21275D89FF9600690A5494EF36A7FE28" box="[347,406,459,486]" gridcol="1" gridrow="3" pageId="41" pageNumber="42">10</td>
<td id="21275D89FF9600690AD094EF3510FE28" box="[479,545,459,486]" gridcol="2" gridrow="3" pageId="41" pageNumber="42">10</td>
<td id="21275D89FF960069097D94EF35AFFE28" box="[626,670,459,486]" gridcol="3" gridrow="3" pageId="41" pageNumber="42">10</td>
<td id="21275D89FF96006909F994EF341BFE28" box="[758,810,459,486]" gridcol="4" gridrow="3" pageId="41" pageNumber="42">10</td>
<td id="21275D89FF960069087294EF3487FE28" box="[893,950,459,486]" gridcol="5" gridrow="3" pageId="41" pageNumber="42">6</td>
<td id="21275D89FF96006908F094EF3373FE28" box="[1023,1090,459,486]" gridcol="6" gridrow="3" pageId="41" pageNumber="42">6</td>
<td id="21275D89FF9600690F8194EF33F4FE28" box="[1166,1221,459,486]" gridcol="7" gridrow="3" pageId="41" pageNumber="42">26</td>
<td id="21275D89FF9600690E1D94EF327CFE28" box="[1298,1357,459,486]" gridcol="8" gridrow="3" pageId="41" pageNumber="42">26</td>
</tr>
</table>
</paragraph>
<paragraph id="DC7936B7FF96FFBF0BB2971936C8FD99" blockId="41.[189,505,573,599]" box="[189,505,573,599]" pageId="41" pageNumber="42">
<heading id="873181DBFF96FFBF0BB2971936C8FD99" bold="true" box="[189,505,573,599]" fontSize="11" level="3" pageId="41" pageNumber="42" reason="3">
<emphasis id="EEB2EAA5FF96FFBF0BB2971936C8FD99" bold="true" box="[189,505,573,599]" pageId="41" pageNumber="42">Other material examined</emphasis>
</heading>
</paragraph>
<paragraph id="DC7936B7FF96FFBF0BB297483352FD3C" blockId="41.[189,1399,620,754]" pageId="41" pageNumber="42">
ATLANTIS
<collectionCode id="BAD7AE72FF96FFBF0A5F974836BEFD48" box="[336,399,620,646]" pageId="41" pageNumber="42">SMT</collectionCode>
: 4 colonies on stylasterid skeleton, Stn 7 (unregistered
<collectionCode id="BAD7AE72FF96FFBF0F1B9749335FFD49" box="[1044,1134,621,647]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="41" pageNumber="42">MNHN</collectionCode>
material); 1 ovicellate colony, Stn 8 (unregistered
<collectionCode id="BAD7AE72FF96FFBF090F97B5356BFD65" box="[512,602,657,683]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="41" pageNumber="42">MNHN</collectionCode>
material); 1 colony on stylasterid skeleton, with
<taxonomicName id="1BC64D34FF96FFBF0F9997B43247FD64" authority="Berning &amp; Harmelin &amp; Bader &amp; Cibio, 2017" authorityName="Berning &amp; Harmelin &amp; Bader &amp; Cibio" authorityYear="2017" box="[1174,1398,656,682]" class="Gymnolaemata" family="Atlantisinidae" genus="Atlantisina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="41" pageNumber="42" phylum="Bryozoa" rank="species" species="atlantis" status="gen. et sp. nov.">
<emphasis id="EEB2EAA5FF96FFBF0F9997B43247FD64" box="[1174,1398,656,682]" italics="true" pageId="41" pageNumber="42">Atlantisina atlantis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F58157DEFF96FFBF0BB29790365AFD00" box="[189,363,692,718]" pageId="41" pageNumber="42" rank="species">gen. et sp. nov.</taxonomicNameLabel>
, Stn 7 (
<collectionCode id="BAD7AE72FF96FFBF0AB1979036C9FD00" box="[446,504,692,718]" pageId="41" pageNumber="42">OLL</collectionCode>
2016/128); 1 colony on bioclast, with
<taxonomicName id="1BC64D34FF96FFBF08BC9790338EFD00" authority="Berning &amp; Harmelin &amp; Bader &amp; Cibio, 2017" authorityName="Berning &amp; Harmelin &amp; Bader &amp; Cibio" authorityYear="2017" box="[947,1215,692,718]" class="Gymnolaemata" family="Atlantisinidae" genus="Bathycyclopora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="41" pageNumber="42" phylum="Bryozoa" rank="species" species="suroiti" status="gen. et sp. nov.">
<emphasis id="EEB2EAA5FF96FFBF08BC9790338EFD00" box="[947,1215,692,718]" italics="true" pageId="41" pageNumber="42">Bathycyclopora suroiti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F58157DEFF96FFBF0FCB97903243FD00" box="[1220,1394,692,718]" pageId="41" pageNumber="42" rank="species">gen. et sp. nov.</taxonomicNameLabel>
, Stn 7 (
<collectionCode id="BAD7AE72FF96FFBF0A0697FC3672FD3C" box="[265,323,728,754]" pageId="41" pageNumber="42">OLL</collectionCode>
2016/151); 1 colony on stylasterid skeleton, Stn 7 (
<collectionCode id="BAD7AE72FF96FFBF08AE97FC34EAFD3C" box="[929,987,728,754]" pageId="41" pageNumber="42">OLL</collectionCode>
2016/154).
</paragraph>
</subSubSection>
<subSubSection id="94DC653CFF96FFBF0BB2963D326DF825" pageId="41" pageNumber="42" type="description">
<paragraph id="DC7936B7FF96FFBF0BB2963D367FFCFD" blockId="41.[189,334,793,819]" box="[189,334,793,819]" pageId="41" pageNumber="42">
<heading id="873181DBFF96FFBF0BB2963D367FFCFD" bold="true" box="[189,334,793,819]" fontSize="11" level="3" pageId="41" pageNumber="42" reason="3">
<emphasis id="EEB2EAA5FF96FFBF0BB2963D367FFCFD" bold="true" box="[189,334,793,819]" pageId="41" pageNumber="42">Description</emphasis>
</heading>
</paragraph>
<paragraph id="DC7936B7FF96FFBF0BB2966D35C8FB23" blockId="41.[189,1399,841,1261]" pageId="41" pageNumber="42">
Colony encrusting, unilaminar, multiserial, forming relatively large patches (
<figureCitation id="44FD2A32FF96FFBF0F54966D33FDFCAD" box="[1115,1228,841,867]" captionStart="Fig" captionStartId="40.[189,232,1652,1678]" captionTargetBox="[189,1399,696,1611]" captionTargetId="figure@40.[189,1399,696,1611]" captionTargetPageId="40" captionText="Fig. 14. Calvetopora otapostasis gen. et sp. nov., Atlantis Smt. A. Overview of the holotype (MNHN- IB-2014-78). B. Close-up of an ooecium and the suboral avicularia (holotype MNHN-IB-2014-78). C. Slightly oblique view of an autozooid at the growth margin showing the communication pores in the lateral walls as well as marginal areolar pores (black arrow) and the roughly crescentically arranged pseudopores (white arrows) in the frontal shield (paratype MNHN-IB-2014-280). D. Close-up of orifice; note that the condyles are so short that usually only one can be seen (paratype OLL 2016/153). E. Ancestrula and early astogenetic autozooids (paratype MNHN-IB-2014-81). F. Interior frontal shield showing the lateral areolar pores and the central pseudopores (paratype OLL 2016/153). Scale bars: A = 500 µm; B, D, F = 100 µm; C, E = 200 µm." figureDoi="http://doi.org/10.5281/zenodo.3832658" httpUri="https://zenodo.org/record/3832658/files/figure.png" pageId="41" pageNumber="42">Fig. 14A</figureCitation>
), occasionally thickened owing to self-overgrowth, sometimes growing free of substratum, producing variably thick calcified extensions from basal wall that may or may not reach substratum; cuticle iridescent in dried colonies. Zooids large, oval to pentagonal or polygonal in outline, separated by shallow grooves (
<figureCitation id="44FD2A32FF96FFBF0BC996FC3607FC3C" box="[198,310,984,1010]" captionStart="Fig" captionStartId="40.[189,232,1652,1678]" captionTargetBox="[189,1399,696,1611]" captionTargetId="figure@40.[189,1399,696,1611]" captionTargetPageId="40" captionText="Fig. 14. Calvetopora otapostasis gen. et sp. nov., Atlantis Smt. A. Overview of the holotype (MNHN- IB-2014-78). B. Close-up of an ooecium and the suboral avicularia (holotype MNHN-IB-2014-78). C. Slightly oblique view of an autozooid at the growth margin showing the communication pores in the lateral walls as well as marginal areolar pores (black arrow) and the roughly crescentically arranged pseudopores (white arrows) in the frontal shield (paratype MNHN-IB-2014-280). D. Close-up of orifice; note that the condyles are so short that usually only one can be seen (paratype OLL 2016/153). E. Ancestrula and early astogenetic autozooids (paratype MNHN-IB-2014-81). F. Interior frontal shield showing the lateral areolar pores and the central pseudopores (paratype OLL 2016/153). Scale bars: A = 500 µm; B, D, F = 100 µm; C, E = 200 µm." figureDoi="http://doi.org/10.5281/zenodo.3832658" httpUri="https://zenodo.org/record/3832658/files/figure.png" pageId="41" pageNumber="42">Fig. 14C</figureCitation>
); lateral walls well developed, communication via two or more basal pore chambers per neighbouring zooid, gymnocystal surface reduced, extensive areas of cryptocystal calcification with a reticulate surface, each framing a large suborbicular pore window (
<figureCitation id="44FD2A32FF96FFBF08F991043352FBF4" box="[1014,1123,1056,1082]" captionStart="Fig" captionStartId="40.[189,232,1652,1678]" captionTargetBox="[189,1399,696,1611]" captionTargetId="figure@40.[189,1399,696,1611]" captionTargetPageId="40" captionText="Fig. 14. Calvetopora otapostasis gen. et sp. nov., Atlantis Smt. A. Overview of the holotype (MNHN- IB-2014-78). B. Close-up of an ooecium and the suboral avicularia (holotype MNHN-IB-2014-78). C. Slightly oblique view of an autozooid at the growth margin showing the communication pores in the lateral walls as well as marginal areolar pores (black arrow) and the roughly crescentically arranged pseudopores (white arrows) in the frontal shield (paratype MNHN-IB-2014-280). D. Close-up of orifice; note that the condyles are so short that usually only one can be seen (paratype OLL 2016/153). E. Ancestrula and early astogenetic autozooids (paratype MNHN-IB-2014-81). F. Interior frontal shield showing the lateral areolar pores and the central pseudopores (paratype OLL 2016/153). Scale bars: A = 500 µm; B, D, F = 100 µm; C, E = 200 µm." figureDoi="http://doi.org/10.5281/zenodo.3832658" httpUri="https://zenodo.org/record/3832658/files/figure.png" pageId="41" pageNumber="42">Fig. 14C</figureCitation>
). Frontal shield matted vitreous, usually very slightly convex, entirely made of thick cryptocystal-type calcification with five to eight marginal pores and between 20 to 40 pseudopores of similar size, 12–16 outermost ones generally arranged in a peripheral U-shaped row, innermost pores occasionally arranged in a similar pattern or more scattered around a central imperforated area, each pore encircled with a faint rim; surface structure rugose with superposed fine granules (
<figureCitation id="44FD2A32FF96FFBF098F91F735DDFB23" box="[640,748,1235,1261]" captionStart="Fig" captionStartId="40.[189,232,1652,1678]" captionTargetBox="[189,1399,696,1611]" captionTargetId="figure@40.[189,1399,696,1611]" captionTargetPageId="40" captionText="Fig. 14. Calvetopora otapostasis gen. et sp. nov., Atlantis Smt. A. Overview of the holotype (MNHN- IB-2014-78). B. Close-up of an ooecium and the suboral avicularia (holotype MNHN-IB-2014-78). C. Slightly oblique view of an autozooid at the growth margin showing the communication pores in the lateral walls as well as marginal areolar pores (black arrow) and the roughly crescentically arranged pseudopores (white arrows) in the frontal shield (paratype MNHN-IB-2014-280). D. Close-up of orifice; note that the condyles are so short that usually only one can be seen (paratype OLL 2016/153). E. Ancestrula and early astogenetic autozooids (paratype MNHN-IB-2014-81). F. Interior frontal shield showing the lateral areolar pores and the central pseudopores (paratype OLL 2016/153). Scale bars: A = 500 µm; B, D, F = 100 µm; C, E = 200 µm." figureDoi="http://doi.org/10.5281/zenodo.3832658" httpUri="https://zenodo.org/record/3832658/files/figure.png" pageId="41" pageNumber="42">Fig. 14C</figureCitation>
).
</paragraph>
<paragraph id="DC7936B7FF96FFBF0BB2903136A3FA70" blockId="41.[189,1399,1301,1470]" pageId="41" pageNumber="42">
Orifice D-shaped, longer than wide, proximal margin fairly straight or shallowly curved (
<figureCitation id="44FD2A32FF96FFBF0FD490313259FAFE" box="[1243,1384,1301,1328]" captionStart="Fig" captionStartId="40.[189,232,1652,1678]" captionTargetBox="[189,1399,696,1611]" captionTargetId="figure@40.[189,1399,696,1611]" captionTargetPageId="40" captionText="Fig. 14. Calvetopora otapostasis gen. et sp. nov., Atlantis Smt. A. Overview of the holotype (MNHN- IB-2014-78). B. Close-up of an ooecium and the suboral avicularia (holotype MNHN-IB-2014-78). C. Slightly oblique view of an autozooid at the growth margin showing the communication pores in the lateral walls as well as marginal areolar pores (black arrow) and the roughly crescentically arranged pseudopores (white arrows) in the frontal shield (paratype MNHN-IB-2014-280). D. Close-up of orifice; note that the condyles are so short that usually only one can be seen (paratype OLL 2016/153). E. Ancestrula and early astogenetic autozooids (paratype MNHN-IB-2014-81). F. Interior frontal shield showing the lateral areolar pores and the central pseudopores (paratype OLL 2016/153). Scale bars: A = 500 µm; B, D, F = 100 µm; C, E = 200 µm." figureDoi="http://doi.org/10.5281/zenodo.3832658" httpUri="https://zenodo.org/record/3832658/files/figure.png" pageId="41" pageNumber="42">Fig. 14D, F</figureCitation>
); condyles inconspicuous in frontal view, formed by a slightly rounded thickening of proximal ends of lateral orifice margin; adult non-maternal zooids usually with three oral spines on distal orifice margin (
<figureCitation id="44FD2A32FF96FFBF0BC990A43600FA55" box="[198,305,1408,1435]" captionStart="Fig" captionStartId="40.[189,232,1652,1678]" captionTargetBox="[189,1399,696,1611]" captionTargetId="figure@40.[189,1399,696,1611]" captionTargetPageId="40" captionText="Fig. 14. Calvetopora otapostasis gen. et sp. nov., Atlantis Smt. A. Overview of the holotype (MNHN- IB-2014-78). B. Close-up of an ooecium and the suboral avicularia (holotype MNHN-IB-2014-78). C. Slightly oblique view of an autozooid at the growth margin showing the communication pores in the lateral walls as well as marginal areolar pores (black arrow) and the roughly crescentically arranged pseudopores (white arrows) in the frontal shield (paratype MNHN-IB-2014-280). D. Close-up of orifice; note that the condyles are so short that usually only one can be seen (paratype OLL 2016/153). E. Ancestrula and early astogenetic autozooids (paratype MNHN-IB-2014-81). F. Interior frontal shield showing the lateral areolar pores and the central pseudopores (paratype OLL 2016/153). Scale bars: A = 500 µm; B, D, F = 100 µm; C, E = 200 µm." figureDoi="http://doi.org/10.5281/zenodo.3832658" httpUri="https://zenodo.org/record/3832658/files/figure.png" pageId="41" pageNumber="42">Fig. 14D</figureCitation>
), early astogenetic zooids may have up to eight spines (
<figureCitation id="44FD2A32FF96FFBF08BF90A43329FA55" box="[944,1048,1408,1435]" captionStart="Fig" captionStartId="40.[189,232,1652,1678]" captionTargetBox="[189,1399,696,1611]" captionTargetId="figure@40.[189,1399,696,1611]" captionTargetPageId="40" captionText="Fig. 14. Calvetopora otapostasis gen. et sp. nov., Atlantis Smt. A. Overview of the holotype (MNHN- IB-2014-78). B. Close-up of an ooecium and the suboral avicularia (holotype MNHN-IB-2014-78). C. Slightly oblique view of an autozooid at the growth margin showing the communication pores in the lateral walls as well as marginal areolar pores (black arrow) and the roughly crescentically arranged pseudopores (white arrows) in the frontal shield (paratype MNHN-IB-2014-280). D. Close-up of orifice; note that the condyles are so short that usually only one can be seen (paratype OLL 2016/153). E. Ancestrula and early astogenetic autozooids (paratype MNHN-IB-2014-81). F. Interior frontal shield showing the lateral areolar pores and the central pseudopores (paratype OLL 2016/153). Scale bars: A = 500 µm; B, D, F = 100 µm; C, E = 200 µm." figureDoi="http://doi.org/10.5281/zenodo.3832658" httpUri="https://zenodo.org/record/3832658/files/figure.png" pageId="41" pageNumber="42">Fig. 14E</figureCitation>
), only two spines in ovicellate zooids (
<figureCitation id="44FD2A32FF96FFBF0A15908036B4FA70" box="[282,389,1444,1470]" captionStart="Fig" captionStartId="40.[189,232,1652,1678]" captionTargetBox="[189,1399,696,1611]" captionTargetId="figure@40.[189,1399,696,1611]" captionTargetPageId="40" captionText="Fig. 14. Calvetopora otapostasis gen. et sp. nov., Atlantis Smt. A. Overview of the holotype (MNHN- IB-2014-78). B. Close-up of an ooecium and the suboral avicularia (holotype MNHN-IB-2014-78). C. Slightly oblique view of an autozooid at the growth margin showing the communication pores in the lateral walls as well as marginal areolar pores (black arrow) and the roughly crescentically arranged pseudopores (white arrows) in the frontal shield (paratype MNHN-IB-2014-280). D. Close-up of orifice; note that the condyles are so short that usually only one can be seen (paratype OLL 2016/153). E. Ancestrula and early astogenetic autozooids (paratype MNHN-IB-2014-81). F. Interior frontal shield showing the lateral areolar pores and the central pseudopores (paratype OLL 2016/153). Scale bars: A = 500 µm; B, D, F = 100 µm; C, E = 200 µm." figureDoi="http://doi.org/10.5281/zenodo.3832658" httpUri="https://zenodo.org/record/3832658/files/figure.png" pageId="41" pageNumber="42">Fig. 14B</figureCitation>
).
</paragraph>
<paragraph id="DC7936B7FF96FFBF0BB290C235B7F95E" blockId="41.[189,1398,1510,1680]" pageId="41" pageNumber="42">
Ooecium produced by zooid distal to maternal one, hyperstomial, resting on the distal zooid’s frontal shield, globular, about as wide as long, with a short peristome wedged in between a pair of spines, terminating at distal orifice margin (
<figureCitation id="44FD2A32FF96FFBF097E930A35EEF986" box="[625,735,1582,1608]" captionStart="Fig" captionStartId="40.[189,232,1652,1678]" captionTargetBox="[189,1399,696,1611]" captionTargetId="figure@40.[189,1399,696,1611]" captionTargetPageId="40" captionText="Fig. 14. Calvetopora otapostasis gen. et sp. nov., Atlantis Smt. A. Overview of the holotype (MNHN- IB-2014-78). B. Close-up of an ooecium and the suboral avicularia (holotype MNHN-IB-2014-78). C. Slightly oblique view of an autozooid at the growth margin showing the communication pores in the lateral walls as well as marginal areolar pores (black arrow) and the roughly crescentically arranged pseudopores (white arrows) in the frontal shield (paratype MNHN-IB-2014-280). D. Close-up of orifice; note that the condyles are so short that usually only one can be seen (paratype OLL 2016/153). E. Ancestrula and early astogenetic autozooids (paratype MNHN-IB-2014-81). F. Interior frontal shield showing the lateral areolar pores and the central pseudopores (paratype OLL 2016/153). Scale bars: A = 500 µm; B, D, F = 100 µm; C, E = 200 µm." figureDoi="http://doi.org/10.5281/zenodo.3832658" httpUri="https://zenodo.org/record/3832658/files/figure.png" pageId="41" pageNumber="42">Fig. 14B</figureCitation>
); ectooecium partially calcified, covering lower part of ooecium; exposed endooecium with a similar surface structure as that of frontal shield; ooecial aperture suborbicular, not closed by operculum.
</paragraph>
<paragraph id="DC7936B7FF96FFBF0BB2939C35EDF8AF" blockId="41.[189,1398,1720,1890]" pageId="41" pageNumber="42">
Adventitious avicularia paired, ovate, situated directly lateral to orifice on a slightly raised cystid (
<figureCitation id="44FD2A32FF96FFBF0BC993FF3626F938" box="[198,279,1755,1782]" captionStart="Fig" captionStartId="40.[189,232,1652,1678]" captionTargetBox="[189,1399,696,1611]" captionTargetId="figure@40.[189,1399,696,1611]" captionTargetPageId="40" captionText="Fig. 14. Calvetopora otapostasis gen. et sp. nov., Atlantis Smt. A. Overview of the holotype (MNHN- IB-2014-78). B. Close-up of an ooecium and the suboral avicularia (holotype MNHN-IB-2014-78). C. Slightly oblique view of an autozooid at the growth margin showing the communication pores in the lateral walls as well as marginal areolar pores (black arrow) and the roughly crescentically arranged pseudopores (white arrows) in the frontal shield (paratype MNHN-IB-2014-280). D. Close-up of orifice; note that the condyles are so short that usually only one can be seen (paratype OLL 2016/153). E. Ancestrula and early astogenetic autozooids (paratype MNHN-IB-2014-81). F. Interior frontal shield showing the lateral areolar pores and the central pseudopores (paratype OLL 2016/153). Scale bars: A = 500 µm; B, D, F = 100 µm; C, E = 200 µm." figureDoi="http://doi.org/10.5281/zenodo.3832658" httpUri="https://zenodo.org/record/3832658/files/figure.png" pageId="41" pageNumber="42">Fig. 14</figureCitation>
B–D); rostrum semi-elliptical but occasionally asymmetric and slightly curved inwards, directing distally or distolaterally, often at an acute angle to frontal plane; distal uncalcified area suborbicular, palate a narrow distal shelf, mandible hinged on a pair of long and thin condyles occasionally fusing at centre, proximal uncalcified area semicircular.
</paragraph>
<paragraph id="DC7936B7FF96FFBF0BB292AD326DF825" blockId="41.[189,1398,1929,2027]" pageId="41" pageNumber="42">
Ancestrula tatiform with 11 spines, opesia oval (ca
<quantity id="1B3E9B52FF96FFBF080992AD346EF86A" box="[774,863,1929,1956]" metricMagnitude="-1" metricUnit="m" metricValue="3.05" pageId="41" pageNumber="42" unit="mm" value="305.0">305 µm</quantity>
long,
<quantity id="1B3E9B52FF96FFBF08AB92AD34CCF86A" box="[932,1021,1929,1956]" metricMagnitude="-1" metricUnit="m" metricValue="2.35" pageId="41" pageNumber="42" unit="mm" value="235.0">235 µm</quantity>
wide), constricted in distal third, gymnocyst narrowing distolaterally, cryptocyst invisible (
<figureCitation id="44FD2A32FF96FFBF0860928934ECF809" box="[879,989,1965,1991]" captionStart="Fig" captionStartId="40.[189,232,1652,1678]" captionTargetBox="[189,1399,696,1611]" captionTargetId="figure@40.[189,1399,696,1611]" captionTargetPageId="40" captionText="Fig. 14. Calvetopora otapostasis gen. et sp. nov., Atlantis Smt. A. Overview of the holotype (MNHN- IB-2014-78). B. Close-up of an ooecium and the suboral avicularia (holotype MNHN-IB-2014-78). C. Slightly oblique view of an autozooid at the growth margin showing the communication pores in the lateral walls as well as marginal areolar pores (black arrow) and the roughly crescentically arranged pseudopores (white arrows) in the frontal shield (paratype MNHN-IB-2014-280). D. Close-up of orifice; note that the condyles are so short that usually only one can be seen (paratype OLL 2016/153). E. Ancestrula and early astogenetic autozooids (paratype MNHN-IB-2014-81). F. Interior frontal shield showing the lateral areolar pores and the central pseudopores (paratype OLL 2016/153). Scale bars: A = 500 µm; B, D, F = 100 µm; C, E = 200 µm." figureDoi="http://doi.org/10.5281/zenodo.3832658" httpUri="https://zenodo.org/record/3832658/files/figure.png" pageId="41" pageNumber="42">Fig. 14E</figureCitation>
); first zooid budded distolaterally, with eight long spines, frontal shield with ca 12 large pseudopores and one adventitious avicularium.
</paragraph>
</subSubSection>
<subSubSection id="94DC653CFF95FFBC0BB2942F34C3FE2F" pageId="42" pageNumber="43" type="discussion">
<paragraph id="DC7936B7FF95FFBC0BB2942F361EFEEB" blockId="42.[189,303,267,293]" box="[189,303,267,293]" pageId="42" pageNumber="43">
<heading id="873181DBFF95FFBC0BB2942F361EFEEB" bold="true" box="[189,303,267,293]" fontSize="11" level="3" pageId="42" pageNumber="43" reason="3">
<emphasis id="EEB2EAA5FF95FFBC0BB2942F361EFEEB" bold="true" box="[189,303,267,293]" pageId="42" pageNumber="43">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="DC7936B7FF95FFBC0BB2941E34C3FE2F" blockId="42.[189,1399,314,482]" pageId="42" pageNumber="43">
With its characteristic, large-sized and strongly calcified zooids,
<taxonomicName id="1BC64D34FF95FFBC08A7941E33F4FE9A" authority="Berning &amp; Harmelin &amp; Bader &amp; Cibio, 2017" authorityName="Berning &amp; Harmelin &amp; Bader &amp; Cibio" authorityYear="2017" box="[936,1221,314,340]" class="Gymnolaemata" family="Atlantisinidae" genus="Calvetopora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="42" pageNumber="43" phylum="Bryozoa" rank="species" species="otapostasis" status="gen. et sp. nov.">
<emphasis id="EEB2EAA5FF95FFBC08A7941E33F4FE9A" box="[936,1221,314,340]" italics="true" pageId="42" pageNumber="43">Calvetopora otapostasis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F58157DEFF95FFBC0FC4941E3246FE9A" box="[1227,1399,314,340]" pageId="42" pageNumber="43" rank="species">gen. et sp. nov.</taxonomicNameLabel>
forms conspicuous and extensive encrusting colonies. Self-overgrowth often resulted in plurilaminar colonies, and the species is able to bridge gaps in the substratum while forming basal struts of highly variable length and width. Some zooids also showed intramural buds within otherwise undamaged zooecia, indicated by the presence of a second orifice rim with spines.
</paragraph>
</subSubSection>
<subSubSection id="94DC653CFF95FFBC0BB2972D36E5FDB8" pageId="42" pageNumber="43" type="biology_ecology">
<paragraph id="DC7936B7FF95FFBC0BB2972D3611FDED" blockId="42.[189,288,521,547]" box="[189,288,521,547]" pageId="42" pageNumber="43">
<heading id="873181DBFF95FFBC0BB2972D3611FDED" bold="true" box="[189,288,521,547]" fontSize="11" level="3" pageId="42" pageNumber="43" reason="3">
<emphasis id="EEB2EAA5FF95FFBC0BB2972D3611FDED" bold="true" box="[189,288,521,547]" pageId="42" pageNumber="43">Ecology</emphasis>
</heading>
</paragraph>
<paragraph id="DC7936B7FF95FFBC0BB2971C36E5FDB8" blockId="42.[189,1398,568,630]" pageId="42" pageNumber="43">
<taxonomicName id="1BC64D34FF95FFBC0BB2971C36D1FD9C" authority="Berning &amp; Harmelin &amp; Bader &amp; Cibio, 2017" authorityName="Berning &amp; Harmelin &amp; Bader &amp; Cibio" authorityYear="2017" box="[189,480,568,594]" class="Gymnolaemata" family="Atlantisinidae" genus="Calvetopora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="42" pageNumber="43" phylum="Bryozoa" rank="species" species="otapostasis" status="gen. et sp. nov.">
<emphasis id="EEB2EAA5FF95FFBC0BB2971C36D1FD9C" box="[189,480,568,594]" italics="true" pageId="42" pageNumber="43">Calvetopora otapostasis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F58157DEFF95FFBC0AE4971D3598FD9D" box="[491,681,569,595]" pageId="42" pageNumber="43" rank="species">gen. et sp. nov.</taxonomicNameLabel>
was mostly found encrusting dead stylasterid skeletons at depths of
<quantity id="1B3E9B52FF95FFBC0A3E977836FEFDB8" box="[305,463,603,630]" metricMagnitude="2" metricUnit="m" metricValue="3.7" metricValueMax="4.6" metricValueMin="2.8" pageId="42" pageNumber="43" unit="m" value="370.0" valueMax="460.0" valueMin="280.0">
280 to
<specimenCount id="CAC0FD3EFF95FFBC0A8D977F36FEFDB8" box="[386,463,603,630]" pageId="42" pageNumber="43" type="generic">460 m</specimenCount>
</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="94DC653CFF95FFBC0BB297B9343FFD28" box="[189,782,669,743]" pageId="42" pageNumber="43" type="distribution">
<paragraph id="DC7936B7FF95FFBC0BB297B93667FD79" blockId="42.[189,342,669,695]" box="[189,342,669,695]" pageId="42" pageNumber="43">
<heading id="873181DBFF95FFBC0BB297B93667FD79" bold="true" box="[189,342,669,695]" fontSize="11" level="3" pageId="42" pageNumber="43" reason="3">
<emphasis id="EEB2EAA5FF95FFBC0BB297B93667FD79" bold="true" box="[189,342,669,695]" pageId="42" pageNumber="43">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="DC7936B7FF95FFBC0BB297E8343FFD28" blockId="42.[189,782,716,743]" box="[189,782,716,743]" pageId="42" pageNumber="43">The species is apparently endemic to Atlantis Smt.</paragraph>
</subSubSection>
</treatment>
</document>
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