treatments-xml/data/46/4F/72/464F723BFFB0FF8D5CDCDDB9FE9AFEF0.xml

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<mods:title id="835E30198C76F15968CA4F5F915ED34A">Sinorhodeus microlepis, a new genus and species of bitterling from China (Teleostei: Cyprinidae: Acheilognathinae)</mods:title>
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<mods:namePart id="0FEA89CBA83E000CC83C227BBC9BEE4A">Liao, Te-Yu</mods:namePart>
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<subSubSection id="86FC90A6FFB0FF885CDCDDB9FD97F93B" box="[151,624,1724,1750]" pageId="1" pageNumber="70" type="nomenclature">
<paragraph id="CE59C32DFFB0FF885CDCDDB9FD97F93B" blockId="1.[151,624,1724,1750]" box="[151,624,1724,1750]" pageId="1" pageNumber="70">
<heading id="95117441FFB0FF885CDCDDB9FD97F93B" bold="true" box="[151,624,1724,1750]" fontSize="11" level="1" pageId="1" pageNumber="70" reason="1">
<emphasis id="FC921F3FFFB0FF885CDCDDB9FD97F93B" bold="true" box="[151,624,1724,1750]" pageId="1" pageNumber="70">
<taxonomicName id="09E6B8AEFFB0FF885CDCDDB9FED7F93B" authority="Li &amp; Liao &amp; Arai &amp; Zhao, 2017" authorityName="Li &amp; Liao &amp; Arai &amp; Zhao" authorityYear="2017" box="[151,304,1724,1750]" class="Actinopterygii" family="Cyprinidae" genus="Sinorhodeus" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="70" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="FC921F3FFFB0FF885CDCDDB9FED7F93B" bold="true" box="[151,304,1724,1750]" italics="true" pageId="1" pageNumber="70">Sinorhodeus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E7A1A244FFB0FF885D7CDDB9FE42F93B" box="[311,421,1724,1750]" pageId="1" pageNumber="70" rank="genus">gen. nov.</taxonomicNameLabel>
Li, Liao &amp; Arai
</emphasis>
</heading>
</paragraph>
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<subSubSection id="86FC90A6FFB0FF8B5CDCDC07F982FD70" lastPageId="2" lastPageNumber="71" pageId="1" pageNumber="70" type="diagnosis">
<paragraph id="CE59C32DFFB0FF8B5CDCDC07F8DEFE15" blockId="1.[151,1436,1794,2035]" lastBlockId="2.[159,1849,485,596]" lastPageId="2" lastPageNumber="71" pageId="1" pageNumber="70">
<emphasis id="FC921F3FFFB0FF885CDCDC07FEF6F8F6" bold="true" box="[151,273,1794,1819]" pageId="1" pageNumber="70">Diagnosis.</emphasis>
<taxonomicName id="09E6B8AEFFB0FF885D6BDC07FE4BF8F6" authority="Li &amp; Liao &amp; Arai &amp; Zhao, 2017" authorityName="Li &amp; Liao &amp; Arai &amp; Zhao" authorityYear="2017" box="[288,428,1794,1819]" class="Actinopterygii" family="Cyprinidae" genus="Sinorhodeus" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="70" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="FC921F3FFFB0FF885D6BDC07FE4BF8F6" box="[288,428,1794,1819]" italics="true" pageId="1" pageNumber="70">Sinorhodeus</emphasis>
</taxonomicName>
can be distinguished from all other genera of 
<taxonomicName id="09E6B8AEFFB0FF885FB6DC07FB24F8F6" authorityName="Bleeker" authorityYear="1863" box="[1021,1219,1794,1819]" class="Actinopterygii" family="Cyprinidae" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="70" phylum="Chordata" rank="subFamily" subFamily="Acheilognathinae">Acheilognathinae</taxonomicName>
by the following characters: pharyngeal teeth 0,0,4–4,0,0, longitudinal scales 41–46, transverse scales 16–18, white spots on dorsalfin rays absent, a black blotch on dorsal fin in juvenile absent, less developed wing-like yolk sac projections in larvae. Similar to 
<taxonomicName id="09E6B8AEFFB0FF885D15DC6BFE59F86A" authorityName="Agassiz" authorityYear="1832" box="[350,446,1902,1927]" class="Actinopterygii" family="Cyprinidae" genus="Rhodeus" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="70" phylum="Chordata" rank="genus">
<emphasis id="FC921F3FFFB0FF885D15DC6BFE59F86A" box="[350,446,1902,1927]" italics="true" pageId="1" pageNumber="70">Rhodeus</emphasis>
</taxonomicName>
in absence of barbels and incomplete lateral line, but distinguished from it by absence of white spots on dorsal-fin rays (vs. present), absence of a black blotch on dorsal fin in juvenile (vs. present), and less developed wing-like yolk sac projections in larvae (vs. well developed, 
<figureCitation id="56DDDFA8FFB0FF885F93DCB3FBD4F822" box="[984,1075,1974,1999]" captionStart="FIGURE1" captionStartId="4.[151,250,1719,1741]" captionTargetBox="[340,1247,311,1697]" captionTargetId="figure@4.[340,1247,311,1697]" captionTargetPageId="4" captionText="FIGURE1. Three types of wing-like yolk sac projections.Arrows show a wing-like yolk sac. A, welldeveloped inprolarva of Rhodeus albomarginatus. B, less developed in prolarva of Tanakia lanceolata. C, not developed in prolarva of Acheilognathus omeiensis." httpUri="https://zenodo.org/record/1064520/files/figure.png" pageId="1" pageNumber="70">Fig. 1A</figureCitation>
). Similar to 
<taxonomicName id="09E6B8AEFFB0FF88588CDCB3FAC7F822" authority="Li &amp; Liao &amp; Arai &amp; Zhao, 2017" authorityName="Li &amp; Liao &amp; Arai &amp; Zhao" authorityYear="2017" box="[1223,1312,1974,1999]" class="Actinopterygii" family="Cyprinidae" genus="Tanakia" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="70" phylum="Chordata" rank="genus" sensu="lato">
<emphasis id="FC921F3FFFB0FF88588CDCB3FAC7F822" box="[1223,1312,1974,1999]" italics="true" pageId="1" pageNumber="70">Tanakia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E7A1A244FFB0FF885961DCB2FA7BF822" box="[1322,1436,1974,1999]" pageId="1" pageNumber="70" sensu="lato">sensu lato</taxonomicNameLabel>
(
<taxonomicName id="09E6B8AEFFB0FF885CEBDCDFFF10F81E" authorityName="Li &amp; Liao &amp; Arai &amp; Zhao" authorityYear="2017" box="[160,247,2010,2035]" class="Actinopterygii" family="Cyprinidae" genus="Tanakia" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="70" phylum="Chordata" rank="genus">
<emphasis id="FC921F3FFFB0FF885CEBDCDFFF10F81E" box="[160,247,2010,2035]" italics="true" pageId="1" pageNumber="70">Tanakia</emphasis>
</taxonomicName>
, 
<taxonomicName id="09E6B8AEFFB0FF885D4CDCDFFE74F81E" box="[263,403,2010,2035]" class="Actinopterygii" family="Cyprinidae" genus="Paratanakia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="70" phylum="Chordata" rank="genus">
<emphasis id="FC921F3FFFB0FF885D4CDCDFFE74F81E" box="[263,403,2010,2035]" italics="true" pageId="1" pageNumber="70">Paratanakia</emphasis>
</taxonomicName>
, and 
<taxonomicName id="09E6B8AEFFB0FF885D9ADCDFFD9AF81E" box="[465,637,2010,2035]" class="Actinopterygii" family="Cyprinidae" genus="Pseudorhodeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="70" phylum="Chordata" rank="genus">
<emphasis id="FC921F3FFFB0FF885D9ADCDFFD9AF81E" box="[465,637,2010,2035]" italics="true" pageId="1" pageNumber="70">Pseudorhodeus</emphasis>
</taxonomicName>
) in absence of white spots on dorsal-fin rays, absence of a black blotch 
<emphasis id="FC921F3FFFB3FF8B583CDAE0FAF3FE14" box="[1143,1300,485,505]" italics="true" pageId="2" pageNumber="71">b b</emphasis>
! &quot;! &quot;# &quot;!
</paragraph>
<paragraph id="CE59C32DFFB3FF8B5CD4DAFBFB2CFDFC" blockId="2.[159,1849,485,596]" box="[159,1227,510,529]" pageId="2" pageNumber="71">%&amp; ’ &amp;! (&quot; &amp;) * &amp;</paragraph>
<paragraph id="CE59C32DFFB3FF8B5A1CD91EFA09FDB9" blockId="2.[159,1849,485,596]" pageId="2" pageNumber="71">
&quot; 
<emphasis id="FC921F3FFFB3FF8B5E17D945FD8FFDB9" box="[604,616,576,596]" italics="true" pageId="2" pageNumber="71">b</emphasis>
&quot; &quot;# $&quot; % (&quot; (&quot;&amp; (&quot;
</paragraph>
<paragraph id="CE59C32DFFB3FF8B5C33D98FF982FD70" blockId="2.[120,1790,650,1590]" box="[120,1637,650,670]" pageId="2" pageNumber="71">
<taxonomicName id="09E6B8AEFFB3FF8B5C33D98FFEB2FD73" authorityName="Li &amp; Liao &amp; Arai &amp; Zhao" authorityYear="2017" box="[120,341,650,670]" class="Actinopterygii" family="Cyprinidae" genus="Sinorhodeus" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="71" phylum="Chordata" rank="species" species="microlepis">
<emphasis id="FC921F3FFFB3FF8B5C33D98FFEB2FD73" box="[120,341,650,670]" italics="true" pageId="2" pageNumber="71">Sinorhodeus microlepis</emphasis>
</taxonomicName>
+, ­./01 +, ­.­ 2 +, ­.­ +, ­.­2/ +, ­./0­ +, ­.­2 +, ­.­23 4
</paragraph>
</subSubSection>
<subSubSection id="86FC90A6FFB3FF8A5C33D9A8FEC0FA86" lastPageId="3" lastPageNumber="72" pageId="2" pageNumber="71" type="description">
<paragraph id="CE59C32DFFB3FF8B5C33D9A8F982FD2C" blockId="2.[120,1790,650,1590]" box="[120,1637,685,705]" pageId="2" pageNumber="71">
<taxonomicName id="09E6B8AEFFB3FF8B5C33D9A8FEB2FD2C" authorityName="Li &amp; Liao &amp; Arai &amp; Zhao" authorityYear="2017" box="[120,341,685,705]" class="Actinopterygii" family="Cyprinidae" genus="Sinorhodeus" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="71" phylum="Chordata" rank="species" species="microlepis">
<emphasis id="FC921F3FFFB3FF8B5C33D9A8FEB2FD2C" box="[120,341,685,705]" italics="true" pageId="2" pageNumber="71">Sinorhodeus microlepis</emphasis>
</taxonomicName>
&amp; +, ­./0. +, ­.­ +, ­.­ 3 +, ­.­2­ +, ­./00 +, ­.­2&amp; +, ­.­21 4
</paragraph>
<paragraph id="CE59C32DFFB3FF8B5C33D9D5F982FD09" blockId="2.[120,1790,650,1590]" box="[120,1637,720,740]" pageId="2" pageNumber="71">
<taxonomicName id="09E6B8AEFFB3FF8B5C33D9D5FEB2FD09" authorityName="Li &amp; Liao &amp; Arai &amp; Zhao" authorityYear="2017" box="[120,341,720,740]" class="Actinopterygii" family="Cyprinidae" genus="Sinorhodeus" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="71" phylum="Chordata" rank="species" species="microlepis">
<emphasis id="FC921F3FFFB3FF8B5C33D9D5FEB2FD09" box="[120,341,720,740]" italics="true" pageId="2" pageNumber="71">Sinorhodeus microlepis</emphasis>
</taxonomicName>
+, ­./0/ +, ­.­ &amp; +, ­.­ 1 +, ­.­20 +, ­.­22 +, ­.­2 +, ­.­2. 4
</paragraph>
<paragraph id="CE59C32DFFB3FF8B5C33D9F1F91AF9D8" blockId="2.[120,1790,650,1590]" pageId="2" pageNumber="71">
<taxonomicName id="09E6B8AEFFB3FF8B5C33D9F1FE62FCEA" baseAuthorityName="Bleeker" baseAuthorityYear="1871" box="[120,389,756,776]" class="Actinopterygii" family="Cyprinidae" genus="Acheilognathus" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="71" phylum="Chordata" rank="species" species="macropterus">
<emphasis id="FC921F3FFFB3FF8B5C33D9F1FE62FCEA" box="[120,389,756,776]" italics="true" pageId="2" pageNumber="71">Acheilognathus macropterus</emphasis>
</taxonomicName>
5,3 2/&amp;­ 5,3 //­. 5,3&amp;0 ­0 5,3 3../ 5,33&amp; 5,3331­0 5,3.2 ­1 
<emphasis id="FC921F3FFFB3FF8B5AD1D9F0F92EFCE5" box="[1690,1737,756,776]" italics="true" pageId="2" pageNumber="71">et al.</emphasis>
&amp;2 3 
<taxonomicName id="09E6B8AEFFB3FF8B5C33D812FE9EFCC6" baseAuthorityName="Wu" baseAuthorityYear="1939" box="[120,377,791,811]" class="Actinopterygii" family="Cyprinidae" genus="Acheilognathus" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="71" phylum="Chordata" rank="species" species="meridianus">
<emphasis id="FC921F3FFFB3FF8B5C33D812FE9EFCC6" box="[120,377,791,811]" italics="true" pageId="2" pageNumber="71">Acheilognathus meridianus</emphasis>
</taxonomicName>
5,3 2/ &amp; 5,3 //02 5,3&amp;0 0 5,3 3./ 5,33&amp; / 5,33310 5,3.2 ­0 
<emphasis id="FC921F3FFFB3FF8B5AD1D81DF92EFCC6" box="[1690,1737,791,811]" italics="true" pageId="2" pageNumber="71">et al.</emphasis>
&amp;2 3 
<taxonomicName id="09E6B8AEFFB3FF8B5C33D83FFE8BFCA3" baseAuthorityName="Temminck &amp; Schlegel" baseAuthorityYear="1846" box="[120,364,826,846]" class="Actinopterygii" family="Cyprinidae" genus="Acheilognathus" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="71" phylum="Chordata" rank="species" species="rhombeus">
<emphasis id="FC921F3FFFB3FF8B5C33D83FFE8BFCA3" box="[120,364,826,846]" italics="true" pageId="2" pageNumber="71">Acheilognathus rhombeus</emphasis>
</taxonomicName>
5,3 2/ ­ 5,3 //0. 5,3&amp;0 0 0 5,3 3.// 5,33&amp; &amp; 5,333100 5,3.2 0 1 
<emphasis id="FC921F3FFFB3FF8B5AD1D839F92EFCA3" box="[1690,1737,826,846]" italics="true" pageId="2" pageNumber="71">et al.</emphasis>
&amp;2 3 
<taxonomicName id="09E6B8AEFFB3FF8B5C33D85BFE60FC9F" authority="tabira" box="[120,391,862,882]" class="Actinopterygii" family="Cyprinidae" genus="Acheilognathus" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="71" phylum="Chordata" rank="subSpecies" species="tabira" subSpecies="tabira">
<emphasis id="FC921F3FFFB3FF8B5C33D85BFE60FC9F" box="[120,391,862,882]" italics="true" pageId="2" pageNumber="71">Acheilognathus tabira tabira</emphasis>
</taxonomicName>
5,3 2/3 5,3 //00 5,3&amp;032&amp; 5,3 3.­2 5,33&amp; &amp;. 5,333.2&amp; 5,3.2 0­ 
<emphasis id="FC921F3FFFB3FF8B5AD1D85AF92EFC9F" box="[1690,1737,862,882]" italics="true" pageId="2" pageNumber="71">et al.</emphasis>
&amp;2 3 
<taxonomicName id="09E6B8AEFFB3FF8B5C33D884FEA6FC78" baseAuthorityName="Bleeker" baseAuthorityYear="1863" box="[120,321,897,917]" class="Actinopterygii" family="Cyprinidae" genus="Acheilognathus" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="71" phylum="Chordata" rank="species" species="typus">
<emphasis id="FC921F3FFFB3FF8B5C33D884FEA6FC78" box="[120,321,897,917]" italics="true" pageId="2" pageNumber="71">
Acheilognathus 
<typeStatus id="115D7D8FFFB3FF8B5D44D886FEA6FC78" box="[271,321,899,917]" pageId="2" pageNumber="71">typus</typeStatus>
</emphasis>
</taxonomicName>
5,3 2/3. 5,3 /­23 5,3&amp;032/ 5,3 3.­1 5,33&amp; 5,333.2/ 5,3.2&amp;2 
<emphasis id="FC921F3FFFB3FF8B5AD1D886F92EFC78" box="[1690,1737,897,917]" italics="true" pageId="2" pageNumber="71">et al.</emphasis>
&amp;2 3 
<taxonomicName id="09E6B8AEFFB3FF8B5CC9D8A0FEF3FC54" box="[130,276,933,953]" class="Actinopterygii" family="Cyprinidae" genus="Acheilognathus" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="71" phylum="Chordata" rank="genus">
<emphasis id="FC921F3FFFB3FF8B5CC9D8A0FEF3FC54" box="[130,276,933,953]" italics="true" pageId="2" pageNumber="71">Acheilognathus</emphasis>
</taxonomicName>
6 
<taxonomicName id="09E6B8AEFFB3FF8B5D68D8A3FE8EFC55" authorityName="Yang, Xiong, Tang &amp; Liu" authorityYear="2010" box="[291,361,934,952]" class="Actinopterygii" family="Cyprinidae" genus="Acheilognathus" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="71" phylum="Chordata" rank="species" species="striatus">
<emphasis id="FC921F3FFFB3FF8B5D68D8A3FE8EFC55" box="[291,361,934,952]" italics="true" pageId="2" pageNumber="71">striatus</emphasis>
</taxonomicName>
5,3 2/32 5,3 //0­ 5,3&amp;032 5,3 3./0 5,33&amp; &amp;1 5,333.2 5,3.2 0/ 
<emphasis id="FC921F3FFFB3FF8B5AD1D8A3F92EFC54" box="[1690,1737,933,953]" italics="true" pageId="2" pageNumber="71">et al.</emphasis>
&amp;2 3 
<taxonomicName id="09E6B8AEFFB3FF8B5C33D8CDFEF1FC31" authorityName="" baseAuthorityName="Miao" baseAuthorityYear="1934" box="[120,278,968,988]" class="Actinopterygii" family="Cyprinidae" genus="Paratanakia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="71" phylum="Chordata" rank="species" species="chii">
<emphasis id="FC921F3FFFB3FF8B5C33D8CDFEF1FC31" box="[120,278,968,988]" italics="true" pageId="2" pageNumber="71">Paratanakia chii</emphasis>
</taxonomicName>
5,3 2/00 5,3 /­1/ 5,3&amp;03.2 5,3 3/ ­ 5,33&amp; ­3 5,333..2 5,3.2&amp;1. 
<emphasis id="FC921F3FFFB3FF8B5AD1D8CFF92EFC31" box="[1690,1737,968,988]" italics="true" pageId="2" pageNumber="71">et al.</emphasis>
&amp;2 3 
<taxonomicName id="09E6B8AEFFB3FF8B5C33D8EEFEB8FC12" baseAuthorityName="Gunther" baseAuthorityYear="1868" box="[120,351,1003,1023]" class="Actinopterygii" family="Cyprinidae" genus="Paratanakia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="71" phylum="Chordata" rank="species" species="himantegus">
<emphasis id="FC921F3FFFB3FF8B5C33D8EEFEB8FC12" box="[120,351,1003,1023]" italics="true" pageId="2" pageNumber="71">Paratanakia himantegus</emphasis>
</taxonomicName>
5,3 2­23 5,3 /­.&amp; 5,3&amp;03.1 5,3 3/3 5,33&amp; ­0 5,333..1 5,3.2&amp;. 
<emphasis id="FC921F3FFFB3FF8B5AD1D8E8F92EFC12" box="[1690,1737,1003,1023]" italics="true" pageId="2" pageNumber="71">et al.</emphasis>
&amp;2 3 
<taxonomicName id="09E6B8AEFFB3FF8B5C33DF0AFEA8FBCF" authorityName="S. Tanaka" authorityYear="1909" box="[120,335,1039,1059]" class="Actinopterygii" family="Cyprinidae" genus="Pseudorhodeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="71" phylum="Chordata" rank="species" species="tanago">
<emphasis id="FC921F3FFFB3FF8B5C33DF0AFEA8FBCF" box="[120,335,1039,1059]" italics="true" pageId="2" pageNumber="71">Pseudorhodeus tanago</emphasis>
</taxonomicName>
5,3 2­ 5,3 /­/ 5,3&amp;03/3 5,3 3/1&amp; 5,33&amp; 0­ 5,333./3 5,3.2&amp;/2 
<emphasis id="FC921F3FFFB3FF8B5AD1DF15F92EFBCE" box="[1690,1737,1039,1059]" italics="true" pageId="2" pageNumber="71">et al.</emphasis>
&amp;2 3 
<taxonomicName id="09E6B8AEFFB3FF8B5C33DF37FE84FBAB" authorityName="Li &amp; Arai" authorityYear="2014" box="[120,355,1074,1094]" class="Actinopterygii" family="Cyprinidae" genus="Rhodeus" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="71" phylum="Chordata" rank="species" species="albomarginatus">
<emphasis id="FC921F3FFFB3FF8B5C33DF37FE84FBAB" box="[120,355,1074,1094]" italics="true" pageId="2" pageNumber="71">Rhodeus albomarginatus</emphasis>
</taxonomicName>
5,3 2/­2 5,3 /­ ­ 5,3&amp;033 5,3 3/ 0 5,33&amp;.1 5,333.3 5,3.2&amp; / 
<emphasis id="FC921F3FFFB3FF8B5AD1DF31F92EFBAB" box="[1690,1737,1074,1094]" italics="true" pageId="2" pageNumber="71">et al.</emphasis>
&amp;2 3 
<taxonomicName id="09E6B8AEFFB3FF8B5C33DF53FEF2FB84" baseAuthorityName="Bloch" baseAuthorityYear="1782" box="[120,277,1110,1130]" class="Actinopterygii" family="Cyprinidae" genus="Rhodeus" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="71" phylum="Chordata" rank="species" species="amarus">
<emphasis id="FC921F3FFFB3FF8B5C33DF53FEF2FB84" box="[120,277,1110,1130]" italics="true" pageId="2" pageNumber="71">Rhodeus amarus</emphasis>
</taxonomicName>
5,3 2/1 5,3/ ­20 5,3&amp;03 &amp; 5,3 3.0 2 5,33&amp;. 5,333. &amp; 5,3.2&amp;2­ 
<emphasis id="FC921F3FFFB3FF8B5AD1DF52F92EFB87" box="[1690,1737,1110,1130]" italics="true" pageId="2" pageNumber="71">et al.</emphasis>
&amp;2 3 
<taxonomicName id="09E6B8AEFFB3FF8B5C33DF7CFEC6FB61" baseAuthorityName="Jordan &amp; Thompson" baseAuthorityYear="1914" box="[120,289,1145,1165]" class="Actinopterygii" family="Cyprinidae" genus="Rhodeus" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="71" phylum="Chordata" rank="species" species="atremius">
<emphasis id="FC921F3FFFB3FF8B5C33DF7CFEC6FB61" box="[120,289,1145,1165]" italics="true" pageId="2" pageNumber="71">Rhodeus atremius</emphasis>
</taxonomicName>
5,3 2/10 5,3 /­ / 5,3&amp;03&amp;2 5,3 3.0­ 5,33&amp; 33 5,333.&amp;2 5,3.2&amp;. 
<emphasis id="FC921F3FFFB3FF8B5AD1DF7FF92EFB60" box="[1690,1737,1145,1165]" italics="true" pageId="2" pageNumber="71">et al.</emphasis>
&amp;2 3 
<taxonomicName id="09E6B8AEFFB3FF8B5C33DF98FEC0FB5C" authorityName="Bogutskaya &amp; Komlev" authorityYear="2001" box="[120,295,1181,1201]" class="Actinopterygii" family="Cyprinidae" genus="Rhodeus" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="71" phylum="Chordata" rank="species" species="colchicus">
<emphasis id="FC921F3FFFB3FF8B5C33DF98FEC0FB5C" box="[120,295,1181,1201]" italics="true" pageId="2" pageNumber="71">Rhodeus colchicus</emphasis>
</taxonomicName>
5,3 2/.&amp; 5,3 /­&amp;2 5,3&amp;03&amp; 5,3 3/2 5,33&amp; 3/ 5,333.&amp; 5,3.2&amp; 0 
<emphasis id="FC921F3FFFB3FF8B5AD1DF9BF92EFB5C" box="[1690,1737,1181,1201]" italics="true" pageId="2" pageNumber="71">et al.</emphasis>
&amp;2 3 
<taxonomicName id="09E6B8AEFFB3FF8B5C33DFC5FF18FB39" baseAuthorityName="Miao" baseAuthorityYear="1934" box="[120,255,1216,1236]" class="Actinopterygii" family="Cyprinidae" genus="Rhodeus" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="71" phylum="Chordata" rank="species" species="fangi">
<emphasis id="FC921F3FFFB3FF8B5C33DFC5FF18FB39" box="[120,255,1216,1236]" italics="true" pageId="2" pageNumber="71">Rhodeus fangi</emphasis>
</taxonomicName>
5,3 2/.3 5,3 /­&amp;&amp; 5,3&amp;03&amp;1 5,3 3/2 5,33&amp; 30 5,333.&amp;1 5,3.2&amp;&amp; 
<emphasis id="FC921F3FFFB3FF8B5AD1DFC4F92DFB39" box="[1690,1738,1216,1236]" italics="true" pageId="2" pageNumber="71">et al.</emphasis>
&amp;2 3 
<taxonomicName id="09E6B8AEFFB3FF8B5C33DFE6FEA2FB1A" authorityName="Karaman" authorityYear="1924" box="[120,325,1251,1271]" class="Actinopterygii" family="Cyprinidae" genus="Rhodeus" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="71" phylum="Chordata" rank="species" species="meridionalis">
<emphasis id="FC921F3FFFB3FF8B5C33DFE6FEA2FB1A" box="[120,325,1251,1271]" italics="true" pageId="2" pageNumber="71">Rhodeus meridionalis</emphasis>
</taxonomicName>
5,3 2/.1 5,3 /­&amp; 5,3&amp;03&amp;. 5,3 3/23 5,33&amp; 12 5,333.&amp;. 5,3.2&amp;&amp;&amp; 
<emphasis id="FC921F3FFFB3FF8B5AD1DFE1F92DFB1A" box="[1690,1738,1251,1271]" italics="true" pageId="2" pageNumber="71">et al.</emphasis>
&amp;2 3 
<taxonomicName id="09E6B8AEFFB3FF8B5C33DE02FE63FAF6" authorityName="Jordan &amp; Thompson" authorityYear="1914" box="[120,388,1287,1307]" class="Actinopterygii" family="Cyprinidae" genus="Rhodeus" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="71" phylum="Chordata" rank="subSpecies" species="ocellatus" subSpecies="kurumeus">
<emphasis id="FC921F3FFFB3FF8B5C33DE02FE63FAF6" box="[120,388,1287,1307]" italics="true" pageId="2" pageNumber="71">Rhodeus ocellatus kurumeus</emphasis>
</taxonomicName>
5,3 2/­&amp; 5,3 /­32 5,3&amp;033 5,3 3/&amp; 5,33&amp;./ 5,333.3 5,3.2&amp; 0 
<emphasis id="FC921F3FFFB3FF8B5AD1DE0DF92DFAF6" box="[1690,1738,1287,1307]" italics="true" pageId="2" pageNumber="71">et al.</emphasis>
&amp;2 3 
<taxonomicName id="09E6B8AEFFB3FF8B5C33DE2FFE9AFAD3" authority="ocellatus" box="[120,381,1322,1342]" class="Actinopterygii" family="Cyprinidae" genus="Rhodeus" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="71" phylum="Chordata" rank="subSpecies" species="ocellatus" subSpecies="ocellatus">
<emphasis id="FC921F3FFFB3FF8B5C33DE2FFE9AFAD3" box="[120,381,1322,1342]" italics="true" pageId="2" pageNumber="71">Rhodeus ocellatus ocellatus</emphasis>
</taxonomicName>
5,3 2//. 5,3 /­ 3 5,3&amp;03 / 5,3 3/ 1 5,33&amp;. 5,333. / 5,3.2&amp; 
<emphasis id="FC921F3FFFB3FF8B5AD1DE2EF92EFAD3" box="[1690,1737,1322,1342]" italics="true" pageId="2" pageNumber="71">et al.</emphasis>
&amp;2 3 
<taxonomicName id="09E6B8AEFFB3FF8B5C33DE48FEBAFA8C" authorityName="Arai, Jeon &amp; Ueda" authorityYear="2001" box="[120,349,1357,1377]" class="Actinopterygii" family="Cyprinidae" genus="Rhodeus" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="71" phylum="Chordata" rank="species" species="pseudosericeus">
<emphasis id="FC921F3FFFB3FF8B5C33DE48FEBAFA8C" box="[120,349,1357,1377]" italics="true" pageId="2" pageNumber="71">Rhodeus pseudosericeus</emphasis>
</taxonomicName>
5,3 2/­ 5,3 /­3 5,3&amp;0333 5,3 3/&amp;&amp; 5,33&amp;.­ 5,333.33 5,3.2&amp;32 
<emphasis id="FC921F3FFFB3FF8B5AD1DE4BF92EFA8C" box="[1690,1737,1357,1377]" italics="true" pageId="2" pageNumber="71">et al.</emphasis>
&amp;2 3 
<taxonomicName id="09E6B8AEFFB3FF8B5C33DE74FEFDFA69" baseAuthorityName="Pallas" baseAuthorityYear="1776" box="[120,282,1393,1413]" class="Actinopterygii" family="Cyprinidae" genus="Rhodeus" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="71" phylum="Chordata" rank="species" species="sericeus">
<emphasis id="FC921F3FFFB3FF8B5C33DE74FEFDFA69" box="[120,282,1393,1413]" italics="true" pageId="2" pageNumber="71">Rhodeus sericeus</emphasis>
</taxonomicName>
5,3 2/­1 5,3 /­3 5,3&amp;033. 5,3 3/&amp;3 5,33&amp; /2 5,333.3. 5,3.2&amp;3&amp; 
<emphasis id="FC921F3FFFB3FF8B5AD1DE77F92EFA68" box="[1690,1737,1393,1413]" italics="true" pageId="2" pageNumber="71">et al.</emphasis>
&amp;2 3 
<taxonomicName id="09E6B8AEFFB3FF8B5C33DE91FED6FA45" authorityName="Li &amp; Arai" authorityYear="2011" box="[120,305,1428,1448]" class="Actinopterygii" family="Cyprinidae" genus="Rhodeus" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="71" phylum="Chordata" rank="species" species="shitaiensis">
<emphasis id="FC921F3FFFB3FF8B5C33DE91FED6FA45" box="[120,305,1428,1448]" italics="true" pageId="2" pageNumber="71">Rhodeus shitaiensis</emphasis>
</taxonomicName>
5,3 2/­/ 5,3 /­31 5,3&amp;033­ 5,3 3/&amp;. 5,33&amp; /&amp; 5,333.3­ 5,3.2&amp;33 
<emphasis id="FC921F3FFFB3FF8B5AD1DE90F92EFA45" box="[1690,1737,1428,1448]" italics="true" pageId="2" pageNumber="71">et al.</emphasis>
&amp;2 3 
<taxonomicName id="09E6B8AEFFB3FF8B5C33DEB2FEFFFA27" authorityName="Gunther" authorityYear="1868" box="[120,280,1463,1483]" class="Actinopterygii" family="Cyprinidae" genus="Rhodeus" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="71" phylum="Chordata" rank="species" species="sinensis">
<emphasis id="FC921F3FFFB3FF8B5C33DEB2FEFFFA27" box="[120,280,1463,1483]" italics="true" pageId="2" pageNumber="71">Rhodeus sinensis</emphasis>
</taxonomicName>
5,3 2/­0 5,3 /­3/ 5,3&amp;0312 5,3 3/&amp;­ 5,33&amp; /3 5,333.12 5,3.2&amp;3. 
<emphasis id="FC921F3FFFB3FF8B5AD1DEBDF92EFA26" box="[1690,1737,1463,1483]" italics="true" pageId="2" pageNumber="71">et al.</emphasis>
&amp;2 3 
<taxonomicName id="09E6B8AEFFB3FF8B5C33DEDEFEC2FA03" baseAuthorityName="Mori" baseAuthorityYear="1935" box="[120,293,1499,1519]" class="Actinopterygii" family="Cyprinidae" genus="Rhodeus" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="71" phylum="Chordata" rank="species" species="suigensis">
<emphasis id="FC921F3FFFB3FF8B5C33DEDEFEC2FA03" box="[120,293,1499,1519]" italics="true" pageId="2" pageNumber="71">Rhodeus suigensis</emphasis>
</taxonomicName>
5,3 2/0­ 5,3 /­1. 5,3&amp;0310 5,3 3/ / 5,33&amp; ­ 5,333.10 5,3.2&amp;11 
<emphasis id="FC921F3FFFB3FF8B5AD1DED9F92EFA02" box="[1690,1737,1499,1519]" italics="true" pageId="2" pageNumber="71">et al.</emphasis>
&amp;2 3 
<taxonomicName id="09E6B8AEFFB3FF8B5C33DEFBFEC9F9FF" baseAuthorityName="Temminck &amp; Schlegel" baseAuthorityYear="1846" box="[120,302,1534,1554]" class="Actinopterygii" family="Cyprinidae" genus="Tanakia" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="71" phylum="Chordata" rank="species" species="lanceolata">
<emphasis id="FC921F3FFFB3FF8B5C33DEFBFEC9F9FF" box="[120,302,1534,1554]" italics="true" pageId="2" pageNumber="71">Tanakia lanceolata</emphasis>
</taxonomicName>
5,3 2­2/ 5,3 /­.1 5,3&amp;03.­ 5,3 3/3. 5,33&amp; 0&amp; 5,333..­ 5,3.2&amp;.3 
<emphasis id="FC921F3FFFB3FF8B5AD1DEFAF92EF9FF" box="[1690,1737,1534,1554]" italics="true" pageId="2" pageNumber="71">et al.</emphasis>
&amp;2 3 
<taxonomicName id="09E6B8AEFFB3FF8B5C33DD27FEF4F9DB" baseAuthorityName="Temminck &amp; Schlegel" baseAuthorityYear="1846" box="[120,275,1570,1590]" class="Actinopterygii" family="Cyprinidae" genus="Tanakia" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="71" phylum="Chordata" rank="species" species="limbata">
<emphasis id="FC921F3FFFB3FF8B5C33DD27FEF4F9DB" box="[120,275,1570,1590]" italics="true" pageId="2" pageNumber="71">Tanakia limbata</emphasis>
</taxonomicName>
5,3 2­20 5,3 /­./ 5,3&amp;03/2 5,3 3/3­ 5,33&amp; 0 3 5,333./2 5,3.2&amp;.. 
<emphasis id="FC921F3FFFB3FF8B5AD1DD26F92EF9DB" box="[1690,1737,1570,1590]" italics="true" pageId="2" pageNumber="71">et al.</emphasis>
&amp;2 3
</paragraph>
<paragraph id="CE59C32DFFB3FF8A5C33DD6FFE76FE65" blockId="2.[120,1789,1641,1696]" lastBlockId="3.[151,1437,151,392]" lastPageId="3" lastPageNumber="72" pageId="2" pageNumber="71">
<taxonomicName id="09E6B8AEFFB3FF8B5C33DD6FFEE7F990" baseAuthorityName="Hamilton" baseAuthorityYear="1822" box="[120,256,1641,1661]" class="Actinopterygii" family="Cyprinidae" genus="Danio" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="71" phylum="Chordata" rank="species" species="dangila">
<emphasis id="FC921F3FFFB3FF8B5C33DD6FFEE7F990" box="[120,256,1641,1661]" italics="true" pageId="2" pageNumber="71">Danio dangila</emphasis>
</taxonomicName>
<date id="BA58E5EDFFB3FF8B5E7EDD6CFD6AF991" box="[565,653,1641,1660]" pageId="2" pageNumber="71" value="1982-07-11">7 82 11</date>
&amp;1 (9&amp;0&amp;.0/ (9320..2 (9320..&amp; (9320/&amp;3 (9320/1. (9320/­­ 
<emphasis id="FC921F3FFFB3FF8B5AD1DD6FF92EF990" box="[1690,1737,1641,1661]" italics="true" pageId="2" pageNumber="71">et al.</emphasis>
&amp;2 3 
<taxonomicName id="09E6B8AEFFB3FF8B5C33DD88FEBEF94D" baseAuthorityName="Temminck &amp; Schlegel" baseAuthorityYear="1846" box="[120,345,1676,1696]" class="Actinopterygii" family="Cyprinidae" genus="Nipponocypris" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="71" phylum="Chordata" rank="species" species="sieboldii">
<emphasis id="FC921F3FFFB3FF8B5C33DD88FEBEF94D" box="[120,345,1676,1696]" italics="true" pageId="2" pageNumber="71">Nipponocypris sieboldii</emphasis>
</taxonomicName>
7 822­.1 (9&amp;0&amp;/,: 0/2.0,: 0/ &amp;2,:1 &amp;­,:1 &amp;,:1 3 
<emphasis id="FC921F3FFFB3FF8B5AD1DD88F92EF94D" box="[1690,1737,1676,1696]" italics="true" pageId="2" pageNumber="71">et al.</emphasis>
&amp;2 3 on dorsal fin in juvenile, and less developed wing-like yolk sac projections in larvae (
<figureCitation id="56DDDFA8FFB2FF8A5820DB92FB25FF5D" box="[1131,1218,151,176]" captionStart="FIGURE1" captionStartId="4.[151,250,1719,1741]" captionTargetBox="[340,1247,311,1697]" captionTargetId="figure@4.[340,1247,311,1697]" captionTargetPageId="4" captionText="FIGURE1. Three types of wing-like yolk sac projections.Arrows show a wing-like yolk sac. A, welldeveloped inprolarva of Rhodeus albomarginatus. B, less developed in prolarva of Tanakia lanceolata. C, not developed in prolarva of Acheilognathus omeiensis." httpUri="https://zenodo.org/record/1064520/files/figure.png" pageId="3" pageNumber="72">Fig. 1B</figureCitation>
), but distinguished from them by uninterrupted incomplete lateral line (vs. complete in 
<taxonomicName id="09E6B8AEFFB2FF8A5F94DBB9FBDFFF38" authorityName="Li &amp; Liao &amp; Arai &amp; Zhao" authorityYear="2017" box="[991,1080,188,213]" class="Actinopterygii" family="Cyprinidae" genus="Tanakia" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="72" phylum="Chordata" rank="genus">
<emphasis id="FC921F3FFFB2FF8A5F94DBB9FBDFFF38" box="[991,1080,188,213]" italics="true" pageId="3" pageNumber="72">Tanakia</emphasis>
</taxonomicName>
and 
<taxonomicName id="09E6B8AEFFB2FF8A5834DBB9FAECFF38" box="[1151,1291,188,213]" class="Actinopterygii" family="Cyprinidae" genus="Paratanakia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="72" phylum="Chordata" rank="genus">
<emphasis id="FC921F3FFFB2FF8A5834DBB9FAECFF38" box="[1151,1291,188,213]" italics="true" pageId="3" pageNumber="72">Paratanakia</emphasis>
</taxonomicName>
, interrupted incomplete in 
<taxonomicName id="09E6B8AEFFB2FF8A5D0DDBDAFE15FF15" box="[326,498,223,248]" class="Actinopterygii" family="Cyprinidae" genus="Pseudorhodeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="72" phylum="Chordata" rank="genus">
<emphasis id="FC921F3FFFB2FF8A5D0DDBDAFE15FF15" box="[326,498,223,248]" italics="true" pageId="3" pageNumber="72">Pseudorhodeus</emphasis>
</taxonomicName>
) and absence of barbels (vs. present). Distinguished from 
<taxonomicName id="09E6B8AEFFB2FF8A5888DBDAFA96FF15" box="[1219,1393,223,248]" class="Actinopterygii" family="Cyprinidae" genus="Acheilognathus" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="72" phylum="Chordata" rank="genus">
<emphasis id="FC921F3FFFB2FF8A5888DBDAFA96FF15" box="[1219,1393,223,248]" italics="true" pageId="3" pageNumber="72">Acheilognathus</emphasis>
</taxonomicName>
by incomplete lateral line (vs. complete, except 
<taxonomicName id="09E6B8AEFFB2FF8A5EDFDA01FD08FEF1" baseAuthorityName="Bleeker" baseAuthorityYear="1863" box="[660,751,260,284]" class="Actinopterygii" family="Cyprinidae" genus="Acheilognathus" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="72" phylum="Chordata" rank="species" species="typus">
<emphasis id="FC921F3FFFB2FF8A5EDFDA01FD08FEF1" box="[660,751,260,284]" italics="true" pageId="3" pageNumber="72">
A. 
<typeStatus id="115D7D8FFFB2FF8A5EFFDA00FD08FEF1" box="[692,751,261,284]" pageId="3" pageNumber="72">typus</typeStatus>
</emphasis>
</taxonomicName>
with incomplete lateral line), absence of white spots on finrays of dorsal fin (vs. present), and less developed wing-like yolk sac projections in larvae (vs. not developed, 
<figureCitation id="56DDDFA8FFB2FF8A593BDA22FF5DFE89" captionStart="FIGURE1" captionStartId="4.[151,250,1719,1741]" captionTargetBox="[340,1247,311,1697]" captionTargetId="figure@4.[340,1247,311,1697]" captionTargetPageId="4" captionText="FIGURE1. Three types of wing-like yolk sac projections.Arrows show a wing-like yolk sac. A, welldeveloped inprolarva of Rhodeus albomarginatus. B, less developed in prolarva of Tanakia lanceolata. C, not developed in prolarva of Acheilognathus omeiensis." httpUri="https://zenodo.org/record/1064520/files/figure.png" pageId="3" pageNumber="72">Fig. 1C</figureCitation>
). 
<taxonomicName id="09E6B8AEFFB2FF8A5C85DA49FEBDFE88" authority="Li &amp; Liao &amp; Arai &amp; Zhao, 2017" authorityName="Li &amp; Liao &amp; Arai &amp; Zhao" authorityYear="2017" box="[206,346,332,357]" class="Actinopterygii" family="Cyprinidae" genus="Sinorhodeus" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="70" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="FC921F3FFFB2FF8A5C85DA49FEBDFE88" box="[206,346,332,357]" italics="true" pageId="3" pageNumber="72">Sinorhodeus</emphasis>
</taxonomicName>
also can be further distinguished from all other genera by fewer pharyngeal teeth (0,0,4–4,0,0 vs. 0,0,5–5,0,0) (
<tableCitation id="8364F696FFB2FF8A5D66DA6AFE64FE65" box="[301,387,367,392]" captionStart="TABLE 3" captionStartId="3.[151,239,1451,1474]" captionText="TABLE 3. Diagnoses of six genera of the subfamily Acheilognathinae." pageId="3" pageNumber="72">Table 3</tableCitation>
).
</paragraph>
<caption id="9A9993A5FFB2FF8A5CDCDABDFE69FE1D" pageId="3" pageNumber="72">
<paragraph id="CE59C32DFFB2FF8A5CDCDABDFE69FE1D" blockId="3.[151,1435,440,496]" pageId="3" pageNumber="72">
<emphasis id="FC921F3FFFB2FF8A5CDCDABDFEEAFE22" bold="true" box="[151,269,440,463]" pageId="3" pageNumber="72">TABLE 2.</emphasis>
Models applied for each codon of seven genes. Numbers refer to codon positions. Refer to Table 1 for abbreviations of genes.
</paragraph>
</caption>
<paragraph id="CE59C32DFFB2FF8A5D0ED90EFA6EFDCC" blockId="3.[159,1417,523,1387]" box="[325,1417,523,545]" pageId="3" pageNumber="72">F81 F81+G F81+I GTR+G GTR+I GTR+I+G HKY+I JC+G JC+G+I HKY HKY+G</paragraph>
<paragraph id="CE59C32DFFB2FF8A5CD4D93FFEC0FDBD" blockId="3.[159,1417,523,1387]" box="[159,295,570,592]" pageId="3" pageNumber="72">
Cyt 
<emphasis id="FC921F3FFFB2FF8A5C81D93FFF30FDBD" box="[202,215,570,592]" italics="true" pageId="3" pageNumber="72">b</emphasis>
1
</paragraph>
<paragraph id="CE59C32DFFB2FF8A5D51D967FEC0FD95" blockId="3.[159,1417,523,1387]" box="[282,295,610,632]" pageId="3" pageNumber="72">2</paragraph>
<paragraph id="CE59C32DFFB2FF8A5D51D98FFEC0FD4D" blockId="3.[159,1417,523,1387]" box="[282,295,650,672]" pageId="3" pageNumber="72">3</paragraph>
<paragraph id="CE59C32DFFB2FF8A5CD4D9B4FEC0FD2A" blockId="3.[159,1417,523,1387]" box="[159,295,689,711]" pageId="3" pageNumber="72">EGR1 1</paragraph>
<paragraph id="CE59C32DFFB2FF8A5D51D9DCFEC0FD02" blockId="3.[159,1417,523,1387]" box="[282,295,729,751]" pageId="3" pageNumber="72">2</paragraph>
<paragraph id="CE59C32DFFB2FF8A5D51D804FEC0FCFA" blockId="3.[159,1417,523,1387]" box="[282,295,769,791]" pageId="3" pageNumber="72">3</paragraph>
<paragraph id="CE59C32DFFB2FF8A5CD4D82CFEC0FCD2" blockId="3.[159,1417,523,1387]" box="[159,295,809,831]" pageId="3" pageNumber="72">EGR2B 1</paragraph>
<paragraph id="CE59C32DFFB2FF8A5D51D855FEC0FC8B" blockId="3.[159,1417,523,1387]" box="[282,295,848,870]" pageId="3" pageNumber="72">2</paragraph>
<paragraph id="CE59C32DFFB2FF8A5D51D87DFEC0FC63" blockId="3.[159,1417,523,1387]" box="[282,295,888,910]" pageId="3" pageNumber="72">3</paragraph>
<paragraph id="CE59C32DFFB2FF8A5CD4D8A5FEC0FC5B" blockId="3.[159,1417,523,1387]" box="[159,295,928,950]" pageId="3" pageNumber="72">EGR3 1</paragraph>
<paragraph id="CE59C32DFFB2FF8A5D51D8C2FEC0FC30" blockId="3.[159,1417,523,1387]" box="[282,295,967,989]" pageId="3" pageNumber="72">2</paragraph>
<paragraph id="CE59C32DFFB2FF8A5D51D8EAFEC0FBE8" blockId="3.[159,1417,523,1387]" box="[282,295,1007,1029]" pageId="3" pageNumber="72">3</paragraph>
<paragraph id="CE59C32DFFB2FF8A5CD4DF12FEC0FBC0" blockId="3.[159,1417,523,1387]" box="[159,295,1047,1069]" pageId="3" pageNumber="72">IRBP2 1</paragraph>
<paragraph id="CE59C32DFFB2FF8A5D51DF3AFEC0FBB8" blockId="3.[159,1417,523,1387]" box="[282,295,1087,1109]" pageId="3" pageNumber="72">2</paragraph>
<paragraph id="CE59C32DFFB2FF8A5D51DF63FEC0FB91" blockId="3.[159,1417,523,1387]" box="[282,295,1126,1148]" pageId="3" pageNumber="72">3</paragraph>
<paragraph id="CE59C32DFFB2FF8A5CD4DF8BFEC0FB49" blockId="3.[159,1417,523,1387]" box="[159,295,1166,1188]" pageId="3" pageNumber="72">RAG1 1</paragraph>
<paragraph id="CE59C32DFFB2FF8A5D51DFB3FEC0FB21" blockId="3.[159,1417,523,1387]" box="[282,295,1206,1228]" pageId="3" pageNumber="72">2</paragraph>
<paragraph id="CE59C32DFFB2FF8A5D51DFDBFEC0FB19" blockId="3.[159,1417,523,1387]" box="[282,295,1246,1268]" pageId="3" pageNumber="72">3</paragraph>
<paragraph id="CE59C32DFFB2FF8A5CD4DE03FEC0FAF6" blockId="3.[159,1417,523,1387]" box="[159,295,1285,1307]" pageId="3" pageNumber="72">RH 1</paragraph>
<paragraph id="CE59C32DFFB2FF8A5D51DE28FEC0FAAE" blockId="3.[159,1417,523,1387]" box="[282,295,1325,1347]" pageId="3" pageNumber="72">2</paragraph>
<paragraph id="CE59C32DFFB2FF8A5D51DE50FEC0FA86" blockId="3.[159,1417,523,1387]" box="[282,295,1365,1387]" pageId="3" pageNumber="72">3</paragraph>
</subSubSection>
<caption id="9A9993A5FFB2FF8A5CDCDEAEFC68FA2F" box="[151,911,1451,1474]" pageId="3" pageNumber="72">
<paragraph id="CE59C32DFFB2FF8A5CDCDEAEFC68FA2F" blockId="3.[151,911,1451,1474]" box="[151,911,1451,1474]" pageId="3" pageNumber="72">
<emphasis id="FC921F3FFFB2FF8A5CDCDEAEFEEEFA2F" bold="true" box="[151,265,1451,1474]" pageId="3" pageNumber="72">TABLE 3.</emphasis>
Diagnoses of six genera of the subfamily 
<taxonomicName id="09E6B8AEFFB2FF8A5E87DEAEFC6BFA2F" authorityName="Bleeker" authorityYear="1863" box="[716,908,1451,1474]" class="Actinopterygii" family="Cyprinidae" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="72" phylum="Chordata" rank="subFamily" subFamily="Acheilognathinae">Acheilognathinae</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="CE59C32DFFB2FF8A5CD4DED8FA8DF80F" pageId="3" pageNumber="72">
<table id="BCE6318DFFB200765CD4DED8FA60F80F" box="[159,1415,1501,2018]" gridcols="6" gridrows="10" pageId="3" pageNumber="72">
<tr id="70D6C16FFFB200765CD4DED8FA60F9FF" box="[159,1415,1501,1554]" gridrow="0" pageId="3" pageNumber="72">
<th id="3307A813FFB200765CD4DED8FE2AF9FF" box="[159,461,1501,1554]" gridcol="0" gridrow="0" pageId="3" pageNumber="72">Character</th>
<th id="3307A813FFB200765DA8DED8FD9FF9FF" box="[483,632,1501,1554]" gridcol="1" gridrow="0" pageId="3" pageNumber="72">
<taxonomicName id="09E6B8AEFFB2FF8A5DA8DED8FD86FA1E" authority="Li &amp; Liao &amp; Arai &amp; Zhao, 2017" authorityName="Li &amp; Liao &amp; Arai &amp; Zhao" authorityYear="2017" box="[483,609,1501,1523]" class="Actinopterygii" family="Cyprinidae" genus="Sinorhodeus" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="70" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="FC921F3FFFB2FF8A5DA8DED8FD86FA1E" box="[483,609,1501,1523]" italics="true" pageId="3" pageNumber="72">Sinorhodeus</emphasis>
</taxonomicName>
</th>
<th id="3307A813FFB200765EDCDED8FCD6F9FF" box="[663,817,1501,1554]" gridcol="2" gridrow="0" pageId="3" pageNumber="72">
<taxonomicName id="09E6B8AEFFB2FF8A5EDCDED8FD09FA1E" authorityName="Agassiz" authorityYear="1832" box="[663,750,1501,1523]" class="Actinopterygii" family="Cyprinidae" genus="Rhodeus" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="72" phylum="Chordata" rank="genus">
<emphasis id="FC921F3FFFB2FF8A5EDCDED8FD09FA1E" box="[663,750,1501,1523]" italics="true" pageId="3" pageNumber="72">Rhodeus</emphasis>
</taxonomicName>
</th>
<th id="3307A813FFB200765F1ADED8FC02F9FF" box="[849,997,1501,1554]" gridcol="3" gridrow="0" pageId="3" pageNumber="72">
<emphasis id="FC921F3FFFB2FF8A5F1ADED8FC37F9FF" italics="true" pageId="3" pageNumber="72">
<taxonomicName id="09E6B8AEFFB2FF8A5F1ADED8FC45FA1E" authorityName="Li &amp; Liao &amp; Arai &amp; Zhao" authorityYear="2017" box="[849,930,1501,1523]" class="Actinopterygii" family="Cyprinidae" genus="Tanakia" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="72" phylum="Chordata" rank="genus">Tanakia</taxonomicName>
/ 
<taxonomicName id="09E6B8AEFFB2FF8A5F1ADEF9FC37F9FF" box="[849,976,1532,1554]" class="Actinopterygii" family="Cyprinidae" genus="Paratanakia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="72" phylum="Chordata" rank="genus">Paratanakia</taxonomicName>
</emphasis>
</th>
<th id="3307A813FFB20076585ADED8FB4AF9FF" box="[1041,1197,1501,1554]" gridcol="4" gridrow="0" pageId="3" pageNumber="72">
<taxonomicName id="09E6B8AEFFB2FF8A585ADED8FB4AFA1E" box="[1041,1197,1501,1523]" class="Actinopterygii" family="Cyprinidae" genus="Pseudorhodeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="72" phylum="Chordata" rank="genus">
<emphasis id="FC921F3FFFB2FF8A585ADED8FB4AFA1E" box="[1041,1197,1501,1523]" italics="true" pageId="3" pageNumber="72">Pseudorhodeus</emphasis>
</taxonomicName>
</th>
<th id="3307A813FFB200765890DED8FA60F9FF" box="[1243,1415,1501,1554]" gridcol="5" gridrow="0" pageId="3" pageNumber="72">
<taxonomicName id="09E6B8AEFFB2FF8A5890DED8FA9EFA1E" box="[1243,1401,1501,1523]" class="Actinopterygii" family="Cyprinidae" genus="Acheilognathus" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="72" phylum="Chordata" rank="genus">
<emphasis id="FC921F3FFFB2FF8A5890DED8FA9EFA1E" box="[1243,1401,1501,1523]" italics="true" pageId="3" pageNumber="72">Acheilognathus</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="70D6C16FFFB200765CD4DD20FA60F9D6" box="[159,1415,1573,1595]" gridrow="1" pageId="3" pageNumber="72" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<th id="3307A813FFB200765CD4DD20FE2AF9D6" box="[159,461,1573,1595]" gridcol="0" gridrow="1" pageId="3" pageNumber="72">
<emphasis id="FC921F3FFFB2FF8A5CD4DD20FE6FF9D6" bold="true" box="[159,392,1573,1595]" pageId="3" pageNumber="72">Morphology of adults</emphasis>
</th>
</tr>
<tr id="70D6C16FFFB200765CD4DD4AFA60F969" box="[159,1415,1615,1668]" gridrow="2" pageId="3" pageNumber="72">
<th id="3307A813FFB200765CD4DD4AFE2AF969" box="[159,461,1615,1668]" gridcol="0" gridrow="2" pageId="3" pageNumber="72">Lateral line</th>
<td id="3307A813FFB200765DA8DD4AFD9FF969" box="[483,632,1615,1668]" gridcol="1" gridrow="2" pageId="3" pageNumber="72">incomplete</td>
<td id="3307A813FFB200765EDCDD4AFCD6F969" box="[663,817,1615,1668]" gridcol="2" gridrow="2" pageId="3" pageNumber="72">incomplete</td>
<td id="3307A813FFB200765F1ADD4AFC02F969" box="[849,997,1615,1668]" gridcol="3" gridrow="2" pageId="3" pageNumber="72">complete</td>
<td id="3307A813FFB20076585ADD4AFB4AF969" box="[1041,1197,1615,1668]" gridcol="4" gridrow="2" pageId="3" pageNumber="72">interrupted incomplete</td>
<td id="3307A813FFB200765890DD4AFA60F969" box="[1243,1415,1615,1668]" gridcol="5" gridrow="2" pageId="3" pageNumber="72">
complete except 
<taxonomicName id="09E6B8AEFFB2FF8A5890DD6BFACBF969" baseAuthorityName="Bleeker" baseAuthorityYear="1863" box="[1243,1324,1646,1668]" class="Actinopterygii" family="Cyprinidae" genus="Acheilognathus" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="72" phylum="Chordata" rank="species" species="typus">
<emphasis id="FC921F3FFFB2FF8A5890DD6BFACBF969" box="[1243,1324,1646,1668]" italics="true" pageId="3" pageNumber="72">A. typus</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="70D6C16FFFB200765CD4DD92FA60F940" box="[159,1415,1687,1709]" gridrow="3" pageId="3" pageNumber="72">
<th id="3307A813FFB200765CD4DD92FE2AF940" box="[159,461,1687,1709]" gridcol="0" gridrow="3" pageId="3" pageNumber="72">Barbels</th>
<td id="3307A813FFB200765DA8DD92FD9FF940" box="[483,632,1687,1709]" gridcol="1" gridrow="3" pageId="3" pageNumber="72">absent</td>
<td id="3307A813FFB200765EDCDD92FCD6F940" box="[663,817,1687,1709]" gridcol="2" gridrow="3" pageId="3" pageNumber="72">absent</td>
<td id="3307A813FFB200765F1ADD92FC02F940" box="[849,997,1687,1709]" gridcol="3" gridrow="3" pageId="3" pageNumber="72">present</td>
<td id="3307A813FFB20076585ADD92FB4AF940" box="[1041,1197,1687,1709]" gridcol="4" gridrow="3" pageId="3" pageNumber="72">present</td>
<td id="3307A813FFB200765890DD92FA60F940" box="[1243,1415,1687,1709]" gridcol="5" gridrow="3" pageId="3" pageNumber="72">present or absent</td>
</tr>
<tr id="70D6C16FFFB200765CD4DDC5FA60F8FE" box="[159,1415,1728,1811]" gridrow="4" pageId="3" pageNumber="72">
<th id="3307A813FFB200765CD4DDC5FE2AF8FE" box="[159,461,1728,1811]" gridcol="0" gridrow="4" pageId="3" pageNumber="72">Whitish spots on fin-rays of dorsal fin form two transverse bands</th>
<td id="3307A813FFB200765DA8DDC5FD9FF8FE" box="[483,632,1728,1811]" gridcol="1" gridrow="4" pageId="3" pageNumber="72">absent</td>
<td id="3307A813FFB200765EDCDDC5FCD6F8FE" box="[663,817,1728,1811]" gridcol="2" gridrow="4" pageId="3" pageNumber="72">present</td>
<td id="3307A813FFB200765F1ADDC5FC02F8FE" box="[849,997,1728,1811]" gridcol="3" gridrow="4" pageId="3" pageNumber="72">absent</td>
<td id="3307A813FFB20076585ADDC5FB4AF8FE" box="[1041,1197,1728,1811]" gridcol="4" gridrow="4" pageId="3" pageNumber="72">absent</td>
<td id="3307A813FFB200765890DDC5FA60F8FE" box="[1243,1415,1728,1811]" gridcol="5" gridrow="4" pageId="3" pageNumber="72">present</td>
</tr>
<tr id="70D6C16FFFB200765CD4DC22FA60F8D0" box="[159,1415,1831,1853]" gridrow="5" pageId="3" pageNumber="72">
<th id="3307A813FFB200765CD4DC22FE2AF8D0" box="[159,461,1831,1853]" gridcol="0" gridrow="5" pageId="3" pageNumber="72">Pharyngeal teeth</th>
<td id="3307A813FFB200765DA8DC22FD9FF8D0" box="[483,632,1831,1853]" gridcol="1" gridrow="5" pageId="3" pageNumber="72">0,0,4–4,0,0</td>
<td id="3307A813FFB200765EDCDC22FCD6F8D0" box="[663,817,1831,1853]" gridcol="2" gridrow="5" pageId="3" pageNumber="72">0,0,5–5,0,0</td>
<td id="3307A813FFB200765F1ADC22FC02F8D0" box="[849,997,1831,1853]" gridcol="3" gridrow="5" pageId="3" pageNumber="72">0,0,5–5,0,0</td>
<td id="3307A813FFB20076585ADC22FB4AF8D0" box="[1041,1197,1831,1853]" gridcol="4" gridrow="5" pageId="3" pageNumber="72">0,0,5–5,0,0</td>
<td id="3307A813FFB200765890DC22FA60F8D0" box="[1243,1415,1831,1853]" gridcol="5" gridrow="5" pageId="3" pageNumber="72">0,0,5–5,0,0</td>
</tr>
<tr id="70D6C16FFFB200765CD4DC55FA60F88B" box="[159,1415,1872,1894]" gridrow="6" pageId="3" pageNumber="72" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<th id="3307A813FFB200765CD4DC55FE2AF88B" box="[159,461,1872,1894]" gridcol="0" gridrow="6" pageId="3" pageNumber="72">
<emphasis id="FC921F3FFFB2FF8A5CD4DC55FE41F88B" bold="true" box="[159,422,1872,1894]" pageId="3" pageNumber="72">Morphology of juveniles</emphasis>
</th>
</tr>
<tr id="70D6C16FFFB200765CD4DC7CFA60F862" box="[159,1415,1913,1935]" gridrow="7" pageId="3" pageNumber="72">
<th id="3307A813FFB200765CD4DC7CFE2AF862" box="[159,461,1913,1935]" gridcol="0" gridrow="7" pageId="3" pageNumber="72">A black blotch on dorsal fin</th>
<td id="3307A813FFB200765DA8DC7CFD9FF862" box="[483,632,1913,1935]" gridcol="1" gridrow="7" pageId="3" pageNumber="72">absent</td>
<td id="3307A813FFB200765EDCDC7CFCD6F862" box="[663,817,1913,1935]" gridcol="2" gridrow="7" pageId="3" pageNumber="72">present</td>
<td id="3307A813FFB200765F1ADC7CFC02F862" box="[849,997,1913,1935]" gridcol="3" gridrow="7" pageId="3" pageNumber="72">absent</td>
<td id="3307A813FFB20076585ADC7CFB4AF862" box="[1041,1197,1913,1935]" gridcol="4" gridrow="7" pageId="3" pageNumber="72">absent</td>
<td id="3307A813FFB200765890DC7CFA60F862" box="[1243,1415,1913,1935]" gridcol="5" gridrow="7" pageId="3" pageNumber="72">present or absent</td>
</tr>
<tr id="70D6C16FFFB200765CD4DCA6FA60F854" box="[159,1415,1955,1977]" gridrow="8" pageId="3" pageNumber="72" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<th id="3307A813FFB200765CD4DCA6FE2AF854" box="[159,461,1955,1977]" gridcol="0" gridrow="8" pageId="3" pageNumber="72">
<emphasis id="FC921F3FFFB2FF8A5CD4DCA6FE6DF854" bold="true" box="[159,394,1955,1977]" pageId="3" pageNumber="72">Morphology of larvae</emphasis>
</th>
</tr>
<tr id="70D6C16FFFB200765CD4DCC9FA60F80F" box="[159,1415,1996,2018]" gridrow="9" pageId="3" pageNumber="72">
<th id="3307A813FFB200765CD4DCC9FE2AF80F" box="[159,461,1996,2018]" gridcol="0" gridrow="9" pageId="3" pageNumber="72">Wing-like yolk sac projection</th>
<td id="3307A813FFB200765DA8DCC9FD9FF80F" box="[483,632,1996,2018]" gridcol="1" gridrow="9" pageId="3" pageNumber="72">less developed</td>
<td id="3307A813FFB200765EDCDCC9FCD6F80F" box="[663,817,1996,2018]" gridcol="2" gridrow="9" pageId="3" pageNumber="72">well developed</td>
<td id="3307A813FFB200765F1ADCC9FC02F80F" box="[849,997,1996,2018]" gridcol="3" gridrow="9" pageId="3" pageNumber="72">less developed</td>
<td id="3307A813FFB20076585ADCC9FB4AF80F" box="[1041,1197,1996,2018]" gridcol="4" gridrow="9" pageId="3" pageNumber="72">less developed</td>
<td id="3307A813FFB200765890DCC9FA60F80F" box="[1243,1415,1996,2018]" gridcol="5" gridrow="9" pageId="3" pageNumber="72">not developed</td>
</tr>
</table>
</paragraph>
<subSubSection id="86FC90A6FFB5FF8D5C8CDB92FD90FF42" box="[199,631,151,176]" pageId="4" pageNumber="73" type="materials_examined">
<paragraph id="CE59C32DFFB5FF8D5C8CDB92FD90FF42" blockId="4.[151,1436,151,285]" box="[199,631,151,176]" pageId="4" pageNumber="73">
<emphasis id="FC921F3FFFB5FF8D5C8CDB92FE85FF5D" bold="true" box="[199,354,151,176]" pageId="4" pageNumber="73">Type species.</emphasis>
<taxonomicName id="09E6B8AEFFB5FF8D5D22DB92FD97FF5D" authorityName="Li &amp; Liao &amp; Arai &amp; Zhao" authorityYear="2017" box="[361,624,151,176]" class="Actinopterygii" family="Cyprinidae" genus="Sinorhodeus" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="73" phylum="Chordata" rank="species" species="microlepis">
<emphasis id="FC921F3FFFB5FF8D5D22DB92FD97FF5D" box="[361,624,151,176]" italics="true" pageId="4" pageNumber="73">Sinorhodeus microlepis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="86FC90A6FFB5FF8D5C8CDBB9FE9AFEF0" pageId="4" pageNumber="73" type="etymology">
<paragraph id="CE59C32DFFB5FF8D5C8CDBB9FE9AFEF0" blockId="4.[151,1436,151,285]" pageId="4" pageNumber="73">
<emphasis id="FC921F3FFFB5FF8D5C8CDBB9FEA9FF38" bold="true" box="[199,334,188,213]" pageId="4" pageNumber="73">Etymology.</emphasis>
The generic name, 
<taxonomicName id="09E6B8AEFFB5FF8D5E61DBB9FD51FF38" authority="Li &amp; Liao &amp; Arai &amp; Zhao, 2017" authorityName="Li &amp; Liao &amp; Arai &amp; Zhao" authorityYear="2017" box="[554,694,188,213]" class="Actinopterygii" family="Cyprinidae" genus="Sinorhodeus" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="70" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="FC921F3FFFB5FF8D5E61DBB9FD51FF38" box="[554,694,188,213]" italics="true" pageId="4" pageNumber="73">Sinorhodeus</emphasis>
</taxonomicName>
, is derived from the Latin 
<emphasis id="FC921F3FFFB5FF8D5FABDBB8FBF5FF39" box="[992,1042,189,212]" italics="true" pageId="4" pageNumber="73">Sino</emphasis>
, meaning &quot;Chinese&quot;, and 
<taxonomicName id="09E6B8AEFFB5FF8D597FDBB9FA73FF38" authorityName="Agassiz" authorityYear="1832" box="[1332,1428,188,213]" class="Actinopterygii" family="Cyprinidae" genus="Rhodeus" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="73" phylum="Chordata" rank="genus">
<emphasis id="FC921F3FFFB5FF8D597FDBB9FA73FF38" box="[1332,1428,188,213]" italics="true" pageId="4" pageNumber="73">Rhodeus</emphasis>
</taxonomicName>
, a genus of bitterling fish, in reference to its distribution in 
<collectingCountry id="B6F183BDFFB5FF8D5F0ADBDAFC63FF15" box="[833,900,223,248]" name="China" pageId="4" pageNumber="73">China</collectingCountry>
and morphological similarity to 
<taxonomicName id="09E6B8AEFFB5FF8D58B4DBDAFAB8FF15" authorityName="Agassiz" authorityYear="1832" box="[1279,1375,223,248]" class="Actinopterygii" family="Cyprinidae" genus="Rhodeus" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="73" phylum="Chordata" rank="genus">
<emphasis id="FC921F3FFFB5FF8D58B4DBDAFAB8FF15" box="[1279,1375,223,248]" italics="true" pageId="4" pageNumber="73">Rhodeus</emphasis>
</taxonomicName>
. The gender is masculine.
</paragraph>
</subSubSection>
</treatment>
</document>