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<document id="5D14EDCA4A5946722E74464117782355" ID-DOI="10.11646/zootaxa.3750.1.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="5271951" ID-ZooBank="3093B45D-78A2-4780-B965-448AB8A177DC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629951147495" checkinUser="felipe" docAuthor="Bilton, David T." docDate="2013" docId="163FA87CEF49FFE99BB4F9E3FF0373D5" docLanguage="en" docName="zt03750p036.pdf" docOrigin="Zootaxa 3750 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Sharphydrus kamiesbergensis Bilton 2013, sp. nov." docType="treatment" docVersion="4" lastPageNumber="34" masterDocId="EA06D004EF4CFFE19B23FF96FFF2750A" masterDocTitle="A taxonomic revision of South African Sharphydrus, with the description of two new species (Coleoptera: Dytiscidae: Bidessini)" masterLastPageNumber="36" masterPageNumber="26" pageNumber="31" updateTime="1699130920934" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="41215F3AD0F28A5EA48158BD42A21C54" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="7CA769D3ECCFAFC8D0D03089A3B0C50A">
<mods:title id="565326D657359229969F8C22214D9DB7">A taxonomic revision of South African Sharphydrus, with the description of two new species (Coleoptera: Dytiscidae: Bidessini)</mods:title>
</mods:titleInfo>
<mods:name id="1F9DADBC56137222D9AF60911EAB9290" type="personal">
<mods:role id="24BCE19556A1FD20B1E30341F96AE9D6">
<mods:roleTerm id="D83443498392592F3C2AF69BAE716E0C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5A1F6903DEE4840FDF527BC45573C29F">Bilton, David T.</mods:namePart>
</mods:name>
<mods:typeOfResource id="B268C14B90B364E246B4361DA014A560">text</mods:typeOfResource>
<mods:relatedItem id="F496D6822DA2C7FDE7E45F666EF0FB7F" type="host">
<mods:titleInfo id="0AD75DDCB0C01CB3A3CD45B1DBFDED0B">
<mods:title id="1573FAF9C5EF3530F6DB0E72CEF3FE19">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="22D7C7BFCB3878A7373204674DD20906">
<mods:date id="83CF5E24A0A2F0789303082540A0398F">2013</mods:date>
<mods:detail id="26BE57E7F75F5917DC7B15794CEEACA7" type="pubDate">
<mods:number id="33CCF68F5277E7BBB3EF31F7C2D52542">2013-12-17</mods:number>
</mods:detail>
<mods:detail id="9B65EC185437C6424C2E8362526A5A88" type="volume">
<mods:number id="678F0B29450F5EC2D826BEC365DCC01E">3750</mods:number>
</mods:detail>
<mods:detail id="67D7FBBBF9B970A2AC459B5E89BF4D77" type="issue">
<mods:number id="3EEEEF895FB29153414176DB4236B567">1</mods:number>
</mods:detail>
<mods:extent id="F036B7C600742AF9912DE24B08FAB35F" unit="page">
<mods:start id="CACD8BCCBCF94387BB0761F4F579900C">26</mods:start>
<mods:end id="F6DA46DADC735737743043ECBD84C343">36</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="F934E11A4A3F5964926D14055244A579">journal article</mods:classification>
<mods:identifier id="389B6E70887E7F11E45EDD935A7B772C" type="DOI">10.11646/zootaxa.3750.1.2</mods:identifier>
<mods:identifier id="EF19FBD56DE8CEC2A19C1DFC303EA0F5" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="F9543A3F3EAC9165F6E0052ED5A6F330" type="Zenodo-Dep">5271951</mods:identifier>
<mods:identifier id="A4D704621183A86D95DE63608938D87C" type="ZooBank">3093B45D-78A2-4780-B965-448AB8A177DC</mods:identifier>
</mods:mods>
<treatment id="163FA87CEF49FFE99BB4F9E3FF0373D5" ID-DOI="http://doi.org/10.5281/zenodo.5272165" ID-Zenodo-Dep="5272165" LSID="urn:lsid:plazi:treatment:163FA87CEF49FFE99BB4F9E3FF0373D5" httpUri="http://treatment.plazi.org/id/163FA87CEF49FFE99BB4F9E3FF0373D5" lastPageId="8" lastPageNumber="34" pageId="5" pageNumber="31">
<subSubSection id="D68C4AE1EF49FFE49BB4F9E3FD947385" box="[151,614,1653,1679]" pageId="5" pageNumber="31" type="nomenclature">
<paragraph id="9E29196AEF49FFE49BB4F9E3FD947385" blockId="5.[151,614,1653,1712]" box="[151,614,1653,1679]" pageId="5" pageNumber="31">
<heading id="C561AE06EF49FFE49BB4F9E3FD947385" bold="true" box="[151,614,1653,1679]" fontSize="11" level="1" pageId="5" pageNumber="31" reason="1">
<emphasis id="ACE2C578EF49FFE49BB4F9E3FD947385" bold="true" box="[151,614,1653,1679]" pageId="5" pageNumber="31">
<taxonomicName id="599662E9EF49FFE49BB4F9E3FDF37385" ID-CoL="8LZBS" authority="Bilton, 2013" authorityName="Bilton" authorityYear="2013" box="[151,513,1653,1679]" class="Insecta" family="Dytiscidae" genus="Sharphydrus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="31" phylum="Arthropoda" rank="species" species="kamiesbergensis" status="sp. nov.">
<emphasis id="ACE2C578EF49FFE49BB4F9E3FDF37385" bold="true" box="[151,513,1653,1679]" italics="true" pageId="5" pageNumber="31">Sharphydrus kamiesbergensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B7D17803EF49FFE4992BF9E3FD947385" box="[520,614,1653,1679]" pageId="5" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="D68C4AE1EF49FFE49BB4F901FEC273BA" box="[151,304,1687,1712]" pageId="5" pageNumber="31" type="description">
<paragraph id="9E29196AEF49FFE49BB4F901FEC273BA" blockId="5.[151,614,1653,1712]" box="[151,304,1687,1712]" pageId="5" pageNumber="31">
(
<figureCitation id="06AD05EFEF49FFE49BBCF901FF0773BA" box="[159,245,1687,1712]" captionStart="FIGURE 1" captionStartId="3.[151,249,1971,1993]" captionTargetBox="[226,1379,218,1931]" captionTargetId="figure-55@3.[201,1389,193,1949]" captionTargetPageId="3" captionText="FIGURE 1. Habitus of A) Sharphydrus brincki sp. nov. B) S. kamiesbergensis sp. nov. C) S. capensis. D) S. coriaceus. Dorsal and lateral views. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5271955" httpUri="https://zenodo.org/record/5271955/files/figure.png" pageId="5" pageNumber="31">Figs 1B</figureCitation>
–
<figureCitation id="06AD05EFEF49FFE49A25F901FED573BA" box="[262,295,1687,1712]" captionStart="FIGURE 3" captionStartId="7.[151,249,1826,1848]" captionTargetBox="[152,1435,193,1805]" captionTargetId="figure-73@7.[152,1435,193,1805]" captionTargetPageId="7" captionText="FIGURE 3. Type localities of: A) Sharphydrus brincki sp. nov. South Africa, Western Cape, Cederberg, Matjiesrivier Reserve, Matjies River ca. 3 km N of Cederberg Oasis; B) S. kamiesbergensis sp. nov. South Africa, Northern Cape, Kamiesberg, stream on Witwater–Langkloof road ca. 1 km S of junction. Photos D.T. Bilton." figureDoi="http://doi.org/10.5281/zenodo.5271961" httpUri="https://zenodo.org/record/5271961/files/figure.png" pageId="5" pageNumber="31">3B</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="D68C4AE1EF49FFE49BB4F949FF0372FF" pageId="5" pageNumber="31" type="materials_examined">
<paragraph id="9E29196AEF49FFE49BB4F949FCE27217" blockId="5.[151,1436,1759,2037]" pageId="5" pageNumber="31">
<emphasis id="ACE2C578EF49FFE49BB4F949FECB73FD" box="[151,313,1759,1784]" inLineHeading="true" pageId="5" pageNumber="31" reason="1">
<emphasis id="ACE2C578EF49FFE49BB4F949FEC073F2" bold="true" box="[151,306,1759,1784]" pageId="5" pageNumber="31">Type locality</emphasis>
.
</emphasis>
<materialsCitation id="2EFE1337EF49FFE49A67F949FCFE7217" country="South Africa" elevation="1097" latitude="-30.395267" location="Witwater-Langkloof" longLatPrecision="1" longitude="18.135666" municipality="Kamiesberg" pageId="5" pageNumber="31" specimenCount="1" stateProvince="Northern Cape">
<emphasis id="ACE2C578EF49FFE49A67F949FE1073F2" bold="true" box="[324,482,1759,1784]" pageId="5" pageNumber="31">
<collectingCountry id="E68159FAEF49FFE49A67F949FE1073F2" box="[324,482,1759,1784]" name="South Africa" pageId="5" pageNumber="31">South Africa</collectingCountry>
</emphasis>
,
<collectingRegion id="5C52D788EF49FFE49AD6F949FD5373F2" box="[501,673,1759,1784]" country="South Africa" name="Northern Cape" pageId="5" pageNumber="31">Northern Cape</collectingRegion>
,
<collectingMunicipality id="7E4D8310EF49FFE49992F949FCC973F2" box="[689,827,1759,1784]" pageId="5" pageNumber="31">Kamiesberg</collectingMunicipality>
, stream on
<location id="9B494FB1EF49FFE498EBF949FB5E73F2" LSID="urn:lsid:plazi:treatment:163FA87CEF49FFE99BB4F9E3FF0373D5:9B494FB1EF49FFE498EBF949FB5E73F2" box="[968,1196,1759,1784]" country="South Africa" latitude="-30.395267" longLatPrecision="1" longitude="18.135666" municipality="Kamiesberg" name="Witwater-Langkloof" pageId="5" pageNumber="31" stateProvince="Northern Cape">Witwater-Langkloof</location>
road ca.
<quantity id="596EB48FEF49FFE49E02F949FAAC73F2" box="[1313,1374,1759,1784]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="31" unit="km" value="1.0">1 km</quantity>
S of junction,
<geoCoordinate id="FBA27FADEF49FFE49A22F892FE677217" box="[257,405,1796,1821]" degrees="30" direction="south" minutes="23" orientation="latitude" pageId="5" pageNumber="31" precision="1" seconds="42.96" value="-30.395267">30 23 42.96S</geoCoordinate>
<geoCoordinate id="FBA27FADEF49FFE49ABFF892FDC77217" box="[412,565,1796,1821]" degrees="18" direction="east" minutes="08" orientation="longitude" pageId="5" pageNumber="31" precision="1" seconds="08.40" value="18.135666">18 08 08.40E</geoCoordinate>
,
<quantity id="596EB48FEF49FFE49963F892FD6A7211" box="[576,664,1796,1821]" metricMagnitude="3" metricUnit="m" metricValue="1.097" pageId="5" pageNumber="31" unit="m" value="1097.0">
<elevation id="15BBFE59EF49FFE49963F892FD6A7211" box="[576,664,1796,1821]" metricMagnitude="3" metricUnit="m" metricValue="1.097" pageId="5" pageNumber="31" unit="m" value="1097.0">1097 m</elevation>
</quantity>
. (
<figureCitation id="06AD05EFEF49FFE49988F892FCF17217" box="[683,771,1796,1821]" captionStart="FIGURE 3" captionStartId="7.[151,249,1826,1848]" captionTargetBox="[152,1435,193,1805]" captionTargetId="figure-73@7.[152,1435,193,1805]" captionTargetPageId="7" captionText="FIGURE 3. Type localities of: A) Sharphydrus brincki sp. nov. South Africa, Western Cape, Cederberg, Matjiesrivier Reserve, Matjies River ca. 3 km N of Cederberg Oasis; B) S. kamiesbergensis sp. nov. South Africa, Northern Cape, Kamiesberg, stream on Witwater–Langkloof road ca. 1 km S of junction. Photos D.T. Bilton." figureDoi="http://doi.org/10.5281/zenodo.5271961" httpUri="https://zenodo.org/record/5271961/files/figure.png" pageId="5" pageNumber="31">Fig. 3B</figureCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="9E29196AEF49FFE49BE4F8B1FB777282" blockId="5.[151,1436,1759,2037]" pageId="5" pageNumber="31">
<emphasis id="ACE2C578EF49FFE49BE4F8B1FE847235" box="[199,374,1831,1856]" inLineHeading="true" pageId="5" pageNumber="31" reason="1">
<emphasis id="ACE2C578EF49FFE49BE4F8B1FE9D724A" bold="true" box="[199,367,1831,1856]" pageId="5" pageNumber="31">Type material</emphasis>
.
</emphasis>
<materialsCitation id="2EFE1337EF49FFE49A5DF8B1FB777282" collectingDate="2010-09-19" collectionCode="ISAM" collectorName="D. T. Bilton" country="South Africa" location="South Africa" pageId="5" pageNumber="31" specimenCount="3" typeStatus="holotype">
<emphasis id="ACE2C578EF49FFE49A5DF8B1FE19724A" bold="true" box="[382,491,1831,1856]" pageId="5" pageNumber="31">
<typeStatus id="412DA7C8EF49FFE49A5DF8B1FE19724A" box="[382,491,1831,1856]" pageId="5" pageNumber="31" type="holotype">Holotype</typeStatus>
</emphasis>
(male): “
<date id="EA283FAAEF49FFE49974F8B1FD20724A" box="[599,722,1831,1856]" pageId="5" pageNumber="31" value="2010-09-19">
<collectingDate id="FA6CC642EF49FFE49974F8B1FD20724A" box="[599,722,1831,1856]" pageId="5" pageNumber="31" value="2010-09-19">19/ix/2010</collectingDate>
</date>
<collectingCountry id="E68159FAEF49FFE499F8F8B1FC99724A" box="[731,875,1831,1856]" name="South Africa" pageId="5" pageNumber="31">South Africa</collectingCountry>
NC, Kamiesberg, stream on Witwater-Langkloof road ca.
<quantity id="596EB48FEF49FFE49BDBF8DAFEC0726F" box="[248,306,1868,1893]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="31" unit="km" value="1.0">1 km</quantity>
S of junction,
<collectorName id="33637CBCEF49FFE49AF8F8DAFD90726F" box="[475,610,1868,1893]" pageId="5" pageNumber="31">D. T. Bilton</collectorName>
leg.” (genitalia extracted and mounted in DMHF on same card) with red printed
<typeStatus id="412DA7C8EF49FFE49BCEF8F9FEBD7282" box="[237,335,1903,1928]" pageId="5" pageNumber="31" type="holotype">holotype</typeStatus>
label “
<typeStatus id="412DA7C8EF49FFE49A84F8F9FDFD7282" box="[423,527,1903,1928]" pageId="5" pageNumber="31" type="holotype">Holotype</typeStatus>
<taxonomicName id="599662E9EF49FFE49935F8F9FC967282" authority="Bilton, 2013" authorityName="Bilton" authorityYear="2013" box="[534,868,1903,1928]" class="Insecta" family="Dytiscidae" genus="Sharphydrus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="31" phylum="Arthropoda" rank="species" species="kamiesbergensis" status="sp. nov.">
<emphasis id="ACE2C578EF49FFE49935F8F9FC967282" box="[534,868,1903,1928]" italics="true" pageId="5" pageNumber="31">Sharphydrus kamiesbergensis</emphasis>
</taxonomicName>
<emphasis id="ACE2C578EF49FFE49848F8F9FC377282" bold="true" box="[875,965,1903,1928]" pageId="5" pageNumber="31">
<taxonomicNameLabel id="B7D17803EF49FFE49848F8F9FC377282" box="[875,965,1903,1928]" pageId="5" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Bilton” (
<collectionCode id="F88781AFEF49FFE49F0EF8F9FB8B7282" box="[1069,1145,1903,1928]" pageId="5" pageNumber="31">ISAM</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="9E29196AEF49FFE49BE4F802FF0372FF" blockId="5.[151,1436,1759,2037]" pageId="5" pageNumber="31">
<materialsCitation id="2EFE1337EF49FFE49BE4F802FF1F72FF" collectingDate="2010-09-19" collectionCode="CDTB, NMW, UMO, ISAM, DMSA, SANC" collectorName="D. T. Bilton" country="South Africa" location="South Africa" pageId="5" pageNumber="31" specimenCount="26" specimenCount-female="12" specimenCount-male="14" typeStatus="paratype">
<emphasis id="ACE2C578EF49FFE49BE4F802FECD72A7" bold="true" box="[199,319,1940,1965]" pageId="5" pageNumber="31">
<typeStatus id="412DA7C8EF49FFE49BE4F802FECD72A7" box="[199,319,1940,1965]" pageId="5" pageNumber="31" type="paratype">Paratypes</typeStatus>
</emphasis>
(
<specimenCount id="8890D2E3EF49FFE49A73F802FE8772A7" box="[336,373,1940,1965]" pageId="5" pageNumber="31" type="male">7♂</specimenCount>
,
<specimenCount id="8890D2E3EF49FFE49AA0F802FE5572A7" box="[387,423,1940,1965]" pageId="5" pageNumber="31" type="female">6♀</specimenCount>
): Same data as holotype (
<specimenCount id="8890D2E3EF49FFE499F2F802FD0572A7" box="[721,759,1940,1965]" pageId="5" pageNumber="31" type="male">2♂</specimenCount>
,
<specimenCount id="8890D2E3EF49FFE49827F802FCD572A7" box="[772,807,1940,1965]" pageId="5" pageNumber="31" type="female">1♀</specimenCount>
<collectionCode id="F88781AFEF49FFE49813F802FC8C72A7" box="[816,894,1940,1965]" pageId="5" pageNumber="31">CDTB</collectionCode>
;
<specimenCount id="8890D2E3EF49FFE498A8F802FC5D72A7" box="[907,943,1940,1965]" pageId="5" pageNumber="31" type="male">1♂</specimenCount>
,
<specimenCount id="8890D2E3EF49FFE4989EF802FC1272A7" box="[957,992,1940,1965]" pageId="5" pageNumber="31" type="female">1♀</specimenCount>
<collectionCode id="F88781AFEF49FFE498CAF802FBC172A1" box="[1001,1075,1940,1963]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="5" pageNumber="31">NMW</collectionCode>
;
<specimenCount id="8890D2E3EF49FFE49F63F802FB9672A7" box="[1088,1124,1940,1965]" pageId="5" pageNumber="31" type="male">1♂</specimenCount>
,
<specimenCount id="8890D2E3EF49FFE49F50F802FB6772A7" box="[1139,1173,1940,1965]" pageId="5" pageNumber="31" type="female">1♀</specimenCount>
<collectionCode id="F88781AFEF49FFE49FBDF802FB1172A7" box="[1182,1251,1940,1965]" country="USA" httpUri="http://grbio.org/cool/asxa-80fx" name="University of Maine" pageId="5" pageNumber="31">UMO</collectionCode>
;
<specimenCount id="8890D2E3EF49FFE49FD3F802FAE672A7" box="[1264,1300,1940,1965]" pageId="5" pageNumber="31" type="male">1♂</specimenCount>
,
<specimenCount id="8890D2E3EF49FFE49E00F802FAB772A7" box="[1315,1349,1940,1965]" pageId="5" pageNumber="31" type="female">1♀</specimenCount>
<collectionCode id="F88781AFEF49FFE49E6DF802FA6A72A7" box="[1358,1432,1940,1965]" pageId="5" pageNumber="31">ISAM</collectionCode>
;
<specimenCount id="8890D2E3EF49FFE49BB4F82EFF4972DB" box="[151,187,1976,2001]" pageId="5" pageNumber="31" type="male">1♂</specimenCount>
,
<specimenCount id="8890D2E3EF49FFE49BE4F82EFF1872DB" box="[199,234,1976,2001]" pageId="5" pageNumber="31" type="female">1♀</specimenCount>
<collectionCode id="F88781AFEF49FFE49BD2F82EFEB772DB" box="[241,325,1976,2001]" country="South Africa" httpUri="http://grbio.org/cool/i1wq-4y1w" name="Durban Museum" pageId="5" pageNumber="31">DMSA</collectionCode>
;
<specimenCount id="8890D2E3EF49FFE49A73F82EFE8672DB" box="[336,372,1976,2001]" pageId="5" pageNumber="31" type="male">1♂</specimenCount>
,
<specimenCount id="8890D2E3EF49FFE49AA2F82EFE5672DB" box="[385,420,1976,2001]" pageId="5" pageNumber="31" type="female">1♀</specimenCount>
<collectionCode id="F88781AFEF49FFE49A89F82EFE0872DB" box="[426,506,1976,2001]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34248" lsid="urn:lsid:biocol.org:col:34248" name="Agricultural Research Council-Plant Protection Research Institute" pageId="5" pageNumber="31" type="Museum">SANC</collectionCode>
). With red printed
<typeStatus id="412DA7C8EF49FFE499E9F82FFCD872DB" box="[714,810,1977,2001]" pageId="5" pageNumber="31" type="paratype">paratype</typeStatus>
labels “
<typeStatus id="412DA7C8EF49FFE498A0F82EFC1A72DB" box="[899,1000,1976,2001]" pageId="5" pageNumber="31" type="paratype">Paratype</typeStatus>
<taxonomicName id="599662E9EF49FFE498CDF82EFACE72DB" authority="Bilton, 2013" authorityName="Bilton" authorityYear="2013" box="[1006,1340,1976,2001]" class="Insecta" family="Dytiscidae" genus="Sharphydrus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="31" phylum="Arthropoda" rank="species" species="kamiesbergensis" status="sp. nov.">
<emphasis id="ACE2C578EF49FFE498CDF82EFACE72DB" box="[1006,1340,1976,2001]" italics="true" pageId="5" pageNumber="31">Sharphydrus kamiesbergensis</emphasis>
</taxonomicName>
<emphasis id="ACE2C578EF49FFE49E60F82EFA6E72DB" bold="true" box="[1347,1436,1976,2001]" pageId="5" pageNumber="31">
<taxonomicNameLabel id="B7D17803EF49FFE49E60F82EFA6E72DB" box="[1347,1436,1976,2001]" pageId="5" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Bilton”
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="D68C4AE1EF4AFFE99BE4FF01FD827059" lastPageId="8" lastPageNumber="34" pageId="6" pageNumber="32" type="description">
<paragraph id="9E29196AEF4AFFE79BE4FF01FB7175DE" blockId="6.[151,1437,151,2035]" pageId="6" pageNumber="32">
<emphasis id="ACE2C578EF4AFFE79BE4FF01FEAA75A5" box="[199,344,151,176]" inLineHeading="true" pageId="6" pageNumber="32" reason="1">
<emphasis id="ACE2C578EF4AFFE79BE4FF01FEA375BA" bold="true" box="[199,337,151,176]" pageId="6" pageNumber="32">Description</emphasis>
.
</emphasis>
Size:
<typeStatus id="412DA7C8EF4AFFE79A80FF01FDE275BA" box="[419,528,151,176]" pageId="6" pageNumber="32" type="holotype">Holotype</typeStatus>
: body length (to elytral apices)
<quantity id="596EB48FEF4AFFE7985DFF01FC1575A5" box="[894,999,151,176]" metricMagnitude="-3" metricUnit="m" metricValue="2.85" pageId="6" pageNumber="32" unit="mm" value="2.85">2.85 mm</quantity>
; max. width (elytra)
<quantity id="596EB48FEF4AFFE79FF9FF01FAB175A5" box="[1242,1347,151,176]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" pageId="6" pageNumber="32" unit="mm" value="1.45">1.45 mm</quantity>
. Elytral length
<quantity id="596EB48FEF4AFFE79BC7FF2DFECF75D9" box="[228,317,187,212]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="6" pageNumber="32" unit="mm" value="2.1">2.1 mm</quantity>
. Same values for
<typeStatus id="412DA7C8EF4AFFE79920FF2AFD9C75DE" box="[515,622,188,212]" pageId="6" pageNumber="32" type="paratype">paratypes</typeStatus>
2.8–3.0, 1.45–1.5 and
<quantity id="596EB48FEF4AFFE7984CFF2DFC0275D9" box="[879,1008,187,212]" metricMagnitude="-3" metricUnit="m" metricValue="2.125" metricValueMax="2.25" metricValueMin="2.0" pageId="6" pageNumber="32" unit="mm" value="2.125" valueMax="2.25" valueMin="2.0">2–2.25 mm</quantity>
respectively.
</paragraph>
<paragraph id="9E29196AEF4AFFE79BE4FF49FBB774F8" blockId="6.[151,1437,151,2035]" pageId="6" pageNumber="32">
Colour (
<figureCitation id="06AD05EFEF4AFFE79A07FF49FE8875F2" box="[292,378,223,248]" captionStart="FIGURE 1" captionStartId="3.[151,249,1971,1993]" captionTargetBox="[226,1379,218,1931]" captionTargetId="figure-55@3.[201,1389,193,1949]" captionTargetPageId="3" captionText="FIGURE 1. Habitus of A) Sharphydrus brincki sp. nov. B) S. kamiesbergensis sp. nov. C) S. capensis. D) S. coriaceus. Dorsal and lateral views. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5271955" httpUri="https://zenodo.org/record/5271955/files/figure.png" pageId="6" pageNumber="32">Fig. 1B</figureCitation>
): Dorsal surface predominantly pale straw yellowish brown. Head and pronotum slightly more deeply pigmented than ground colour of elytra. Hind margin of pronotum with well-marked black band, this being thickest between lateral plicae, here approximately 1/6–1/5 pronotal length in extent. Black band narrower towards centre of pronotum, giving it a distinctly undulating appearance. Front margin of pronotum entirely pale. Elytra with diffuse dark markings, with rather ill-defined boundaries. Sublateral dark markings more well defined, present just behind middle, and just before apex. Palpi and legs uniformly pale yellowish brown. Antennae also pale yellowish brown, with segments 7–11 infuscated apically, this being particularly marked in segment 11. Venter pale yellowish brown, somewhat darkened around ventral junctions and coxal insertions.
</paragraph>
<paragraph id="9E29196AEF4AFFE79BE4FE6BFA61761A" blockId="6.[151,1437,151,2035]" pageId="6" pageNumber="32">Head: Broad, with large eyes. Clypeus somewhat thickened and evenly rounded, but lacking anterior border, and not separated from rest of head. Well-developed shallow, open fovea present on each side of head, anterior to eye, just behind clypeus. Cervical line well marked between hind portion of eyes. Head in front of cervical line with distinct, coarse punctures, each bearing a small peg-like seta at its centre. Punctures smaller and sparser on clypeus, here 2–3 puncture widths apart; becoming larger and denser towards cervical line, here approximately 3x diameter of those on anterior clypeus, and spaced 1/3–1/2 puncture width apart. Head microreticulate except on smooth central frons,. Microreticulation transverse on clypeus and behind cervical line; isodiametric elsewhere. Apical segments of maxiliary and labial palpi narrowly truncate, and somewhat flattened dorsoventrally at apices.</paragraph>
<paragraph id="9E29196AEF4AFFE79BE4FC8DFCB57100" blockId="6.[151,1437,151,2035]" pageId="6" pageNumber="32">Pronotum: Transverse rectangular. Sides rounded to front angles, straight behind; widest just before middle. Hind margins distinctly bisinuate around middle. Lateral plicae well developed, relatively deep, and open interiorly. Plicae deepest at posterior ends; sinuate and becoming shallow anteriorly, here extended to approximately 1/3 away from front margin of the pronotum. Pronotal surface rough, with a coriaceous appearance, due to coarse, dense, sometimes almost confluent punctation, each puncture bearing a small peg-like seta as on head. Punctures particularly dense towards centre of pronotal disc, here separated by ridges only. Pronotal surface smooth between punctures; without visible microreticulation.</paragraph>
<paragraph id="9E29196AEF4AFFE79BE4FB83FDC57064" blockId="6.[151,1437,151,2035]" pageId="6" pageNumber="32">
Elytra: Together forming an elongate oval, with point of maximum width around middle. Lateral margin rounded to front angles, somewhat subparallel in middle, and tapered towards apex in posterior half. Elytra with well-developed discal keels running posteriorly from base of pronotal lateral plicae to a point just over halfway along elytral length. Carinae robust; maximum height being towards middle of keel. Elytral suture also raised over discal region; area between keels and suture relatively flat (
<figureCitation id="06AD05EFEF4AFFE79876FB32FC4271B7" box="[853,944,1188,1213]" captionStart="FIGURE 1" captionStartId="3.[151,249,1971,1993]" captionTargetBox="[226,1379,218,1931]" captionTargetId="figure-55@3.[201,1389,193,1949]" captionTargetPageId="3" captionText="FIGURE 1. Habitus of A) Sharphydrus brincki sp. nov. B) S. kamiesbergensis sp. nov. C) S. capensis. D) S. coriaceus. Dorsal and lateral views. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5271955" httpUri="https://zenodo.org/record/5271955/files/figure.png" pageId="6" pageNumber="32">Fig. 1B</figureCitation>
). Elytral tracheation clearly visible in dry specimens. Each elytron with a row of long, fine, recumbent setae interior to keel, positioned approximately 1/3 way towards suture, and running for length of keel (setae apparently lost and invisible in some
<typeStatus id="412DA7C8EF4AFFE79FF7FB7DFAB6700E" box="[1236,1348,1259,1284]" pageId="6" pageNumber="32" type="paratype">paratypes</typeStatus>
). Entire elytral surface, including keels, with dense, coarse punctation, giving a coriaceous appearance. Punctures almost confluent in places (e.g. close to keels), and each puncture bearing a small peg-like seta. No visible microreticulation between punctures.
</paragraph>
<paragraph id="9E29196AEF4AFFE79BE4FAEDFE5472F8" blockId="6.[151,1437,151,2035]" pageId="6" pageNumber="32">Venter: Anterior margin of prementum with short, stout setae and scattered longer hair-like setae behind. Mentum will well developed blunt truncated medial tooth. Surface of mentum with fine transverse ridges, but otherwise lacking apparent microreticulation. Submentum with transverse furrows and scattered punctures, but lacking microreticulation. Gular sutures weak, but gular region distinct due to absence of microreticulation. Remainder of underside of head with distinct isodiametric reticulation, obsolete on genae and towards outer margins of mandibles. Pronotal hypomeron smooth and shining, with very few scattered punctures. Prosternum smooth and shining with scattered punctures, becoming dense and almost confluent at base of prosternal process, an area also furnished with long, hair-like setae. Prosternal process strongly raised, neck of process forming an angle of approximately 70° with prosternum. Process elongate, cordiform, flat, with a broad shallow groove occupying 3/4 of process width. Neck of process with long, hair-like setae; process itself with punctures bearing short, peg-like setae. Prosternal process reaching anteromedial process of the metaventrite. Metaventrite with weakly defined groove in rear half. Coxal lines well-marked, subparallel, diverging anteriorly. Hind coxal margin bisinuate; coxal lobes weak, not covering leg insertions. Ventral surface of meso- and metathorax and abdomen, elytral epipleurs with dense, coarse punctation, each puncture bearing a short peg-like seta. Punctures very dense and almost confluent on hind coxae and abdominal ventrites; elsewhere spaced at most 1–1.5 puncture widths apart. Punctures absent only from hind margins of hind coxae, where traces of isodiametric microreticulation are visible; spaces between punctures otherwise smooth. Fourth and fifth abdominal ventrites with long hair-like setal bunches at their centres.</paragraph>
<caption id="CAE949E2EF4BFFE69BB4F8B4FC86727F" ID-DOI="http://doi.org/10.5281/zenodo.5271961" ID-Zenodo-Dep="5271961" httpUri="https://zenodo.org/record/5271961/files/figure.png" pageId="7" pageNumber="33" startId="7.[151,249,1826,1848]" targetBox="[152,1435,193,1805]" targetPageId="7">
<paragraph id="9E29196AEF4BFFE69BB4F8B4FC86727F" blockId="7.[151,1436,1826,1909]" pageId="7" pageNumber="33">
<emphasis id="ACE2C578EF4BFFE69BB4F8B4FEE0723D" bold="true" box="[151,274,1826,1848]" pageId="7" pageNumber="33">FIGURE 3.</emphasis>
Type localities of: A)
<taxonomicName id="599662E9EF4BFFE69AD7F8B4FD317232" authority="Bilton, 2013" authorityName="Bilton" authorityYear="2013" box="[500,707,1826,1848]" class="Insecta" family="Dytiscidae" genus="Sharphydrus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="33" phylum="Arthropoda" rank="species" species="brincki" status="sp. nov.">
<emphasis id="ACE2C578EF4BFFE69AD7F8B4FD317232" box="[500,707,1826,1848]" italics="true" pageId="7" pageNumber="33">Sharphydrus brincki</emphasis>
</taxonomicName>
<emphasis id="ACE2C578EF4BFFE699EAF8B4FCEB723D" bold="true" box="[713,793,1826,1847]" pageId="7" pageNumber="33">
<taxonomicNameLabel id="B7D17803EF4BFFE699EAF8B4FCEB723D" box="[713,793,1826,1847]" pageId="7" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
South Africa, Western Cape, Cederberg, Matjiesrivier Reserve, Matjies River ca. 3 km N of Cederberg Oasis; B)
<taxonomicName id="599662E9EF4BFFE699ABF8D7FCB5725D" authority="Bilton, 2013" authorityName="Bilton" authorityYear="2013" box="[648,839,1857,1879]" class="Insecta" family="Dytiscidae" genus="Sharphydrus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="33" phylum="Arthropoda" rank="species" species="kamiesbergensis" status="sp. nov.">
<emphasis id="ACE2C578EF4BFFE699ABF8D7FCB5725D" box="[648,839,1857,1879]" italics="true" pageId="7" pageNumber="33">S. kamiesbergensis</emphasis>
</taxonomicName>
<emphasis id="ACE2C578EF4BFFE6986EF8D7FC6F725C" bold="true" box="[845,925,1857,1878]" pageId="7" pageNumber="33">
<taxonomicNameLabel id="B7D17803EF4BFFE6986EF8D7FC6F725C" box="[845,925,1857,1878]" pageId="7" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
South Africa, Northern Cape, Kamiesberg, stream on Witwater–Langkloof road ca. 1 km S of junction. Photos D.T. Bilton.
</paragraph>
</caption>
<caption id="CAE949E2EF44FFE99BB4FC0CFEED76E7" ID-DOI="http://doi.org/10.5281/zenodo.5271963" ID-Zenodo-Dep="5271963" httpUri="https://zenodo.org/record/5271963/files/figure.png" pageId="8" pageNumber="34" startId="8.[151,249,922,944]" targetBox="[158,1429,186,902]" targetPageId="8">
<paragraph id="9E29196AEF44FFE99BB4FC0CFEED76E7" blockId="8.[151,1435,922,1005]" pageId="8" pageNumber="34">
<emphasis id="ACE2C578EF44FFE99BB4FC0CFEE076A5" bold="true" box="[151,274,922,944]" pageId="8" pageNumber="34">FIGURE 4.</emphasis>
Distribution of
<taxonomicName id="599662E9EF44FFE99A97FC0CFD7776BA" box="[436,645,922,944]" class="Insecta" family="Dytiscidae" genus="Sharphydrus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="34" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="ACE2C578EF44FFE99A97FC0CFDC476BA" box="[436,566,922,944]" italics="true" pageId="8" pageNumber="34">Sharphydrus</emphasis>
species
</taxonomicName>
in South Africa: yellow symbols
<taxonomicName id="599662E9EF44FFE998F9FC0CFBC976BA" authorityName="Bilton" authorityYear="2013" box="[986,1083,922,944]" class="Insecta" family="Dytiscidae" genus="Sharphydrus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="34" phylum="Arthropoda" rank="species" species="brincki">
<emphasis id="ACE2C578EF44FFE998F9FC0CFBC976BA" box="[986,1083,922,944]" italics="true" pageId="8" pageNumber="34">S. brincki</emphasis>
</taxonomicName>
, green symbol
<taxonomicName id="599662E9EF44FFE99FF5FC0CFA6776BA" authorityName="Bilton" authorityYear="2013" box="[1238,1429,922,944]" class="Insecta" family="Dytiscidae" genus="Sharphydrus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="34" phylum="Arthropoda" rank="species" species="kamiesbergensis">
<emphasis id="ACE2C578EF44FFE99FF5FC0CFA6776BA" box="[1238,1429,922,944]" italics="true" pageId="8" pageNumber="34">S. kamiesbergensis</emphasis>
</taxonomicName>
, red symbols
<taxonomicName id="599662E9EF44FFE99A38FC2EFE7F76C4" baseAuthorityName="Thunberg" baseAuthorityYear="1781" box="[283,397,952,974]" class="Insecta" family="Gyrinidae" genus="Aulonogyrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="34" phylum="Arthropoda" rank="species" species="capensis">
<emphasis id="ACE2C578EF44FFE99A38FC2EFE7F76C4" box="[283,397,952,974]" italics="true" pageId="8" pageNumber="34">S. capensis</emphasis>
</taxonomicName>
, white symbols
<taxonomicName id="599662E9EF44FFE9991BFC2EFD4476C4" baseAuthorityName="Regimbart" baseAuthorityYear="1895" box="[568,694,952,974]" class="Insecta" family="Dytiscidae" genus="Sharphydrus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="34" phylum="Arthropoda" rank="species" species="coriaceus">
<emphasis id="ACE2C578EF44FFE9991BFC2EFD4476C4" box="[568,694,952,974]" italics="true" pageId="8" pageNumber="34">S. coriaceus</emphasis>
</taxonomicName>
. Closed circles indicate specimens examined, open circles records from the literature.
</paragraph>
</caption>
<paragraph id="9E29196AEF44FFE99BE4FB8CFE887194" blockId="8.[151,1436,1050,1759]" pageId="8" pageNumber="34">
Aedeagus: Median lobe characteristically shaped and pigmented (
<figureCitation id="06AD05EFEF44FFE99899FB8CFBE77139" box="[954,1045,1050,1075]" captionStart="FIGURE 2" captionStartId="4.[151,249,1368,1390]" captionTargetBox="[169,1418,177,1347]" captionTargetId="figure-301@4.[169,1418,161,1347]" captionTargetPageId="4" captionText="FIGURE 2. Male genitalia of Sharphydrus species: median lobe in lateral and ventral view, right paramere of: A) S. brincki sp. nov.; B) S. kamiesbergensis sp. nov.; C) S. capensis; D) S. coriaceus. Scale bar = 0.5 mm" figureDoi="http://doi.org/10.5281/zenodo.5271957" httpUri="https://zenodo.org/record/5271957/files/figure.png" pageId="8" pageNumber="34">Fig. 2B</figureCitation>
); trilobate at apex, with a curved, ventrally-directed process. Median lobe deeply pigmented, black, with exception of apical median process, this dark colouration being present in fresh and preserved specimens. Parameres (
<figureCitation id="06AD05EFEF44FFE998D5FBF4FBBE7171" box="[1014,1100,1122,1147]" captionStart="FIGURE 2" captionStartId="4.[151,249,1368,1390]" captionTargetBox="[169,1418,177,1347]" captionTargetId="figure-301@4.[169,1418,161,1347]" captionTargetPageId="4" captionText="FIGURE 2. Male genitalia of Sharphydrus species: median lobe in lateral and ventral view, right paramere of: A) S. brincki sp. nov.; B) S. kamiesbergensis sp. nov.; C) S. capensis; D) S. coriaceus. Scale bar = 0.5 mm" figureDoi="http://doi.org/10.5281/zenodo.5271957" httpUri="https://zenodo.org/record/5271957/files/figure.png" pageId="8" pageNumber="34">Fig. 2B</figureCitation>
) two-segmented, with bluntly curved, hooked tips.
</paragraph>
<paragraph id="9E29196AEF44FFE99BE4FB3CFAC07001" blockId="8.[151,1436,1050,1759]" pageId="8" pageNumber="34">Females: As males except punctures on pronotum somewhat sparser, with some isodiametric microreticulation visible between them. Similar, but much weaker and somewhat obsolete reticulation visible between elytral punctures. Fore and mid tarsi narrower and slightly shorter than in males, particularly last tarsal segment.</paragraph>
<paragraph id="9E29196AEF44FFE99BE4FA83FD827059" blockId="8.[151,1436,1050,1759]" pageId="8" pageNumber="34">
<emphasis id="ACE2C578EF44FFE99BE4FA83FEBE7027" box="[199,332,1301,1326]" inLineHeading="true" pageId="8" pageNumber="34" reason="1">
<emphasis id="ACE2C578EF44FFE99BE4FA83FEB77024" bold="true" box="[199,325,1301,1326]" pageId="8" pageNumber="34">Variability</emphasis>
.
</emphasis>
<typeStatus id="412DA7C8EF44FFE99A7BFA80FE377024" box="[344,453,1302,1326]" pageId="8" pageNumber="34" type="paratype">Paratypes</typeStatus>
vary slightly in size (see above) and the development of the elytral pattern, some specimens being darker than the
<typeStatus id="412DA7C8EF44FFE99925FAACFD9E7059" box="[518,620,1338,1363]" pageId="8" pageNumber="34" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="D68C4AE1EF44FFE99BE4FAC8FB8170E9" pageId="8" pageNumber="34" type="distribution">
<paragraph id="9E29196AEF44FFE99BE4FAC8FB8170E9" blockId="8.[151,1436,1050,1759]" pageId="8" pageNumber="34">
<emphasis id="ACE2C578EF44FFE99BE4FAC8FE92707C" box="[199,352,1374,1399]" inLineHeading="true" pageId="8" pageNumber="34" reason="1">
<emphasis id="ACE2C578EF44FFE99BE4FAC8FEAB707D" bold="true" box="[199,345,1374,1399]" pageId="8" pageNumber="34">Distribution</emphasis>
.
</emphasis>
To date only known from the
<typeStatus id="412DA7C8EF44FFE9998CFAC9FD2D707D" box="[687,735,1375,1399]" pageId="8" pageNumber="34">type</typeStatus>
locality (
<figureCitation id="06AD05EFEF44FFE9986BFAC8FC6D707D" box="[840,927,1374,1399]" captionStart="FIGURE 3" captionStartId="7.[151,249,1826,1848]" captionTargetBox="[152,1435,193,1805]" captionTargetId="figure-73@7.[152,1435,193,1805]" captionTargetPageId="7" captionText="FIGURE 3. Type localities of: A) Sharphydrus brincki sp. nov. South Africa, Western Cape, Cederberg, Matjiesrivier Reserve, Matjies River ca. 3 km N of Cederberg Oasis; B) S. kamiesbergensis sp. nov. South Africa, Northern Cape, Kamiesberg, stream on Witwater–Langkloof road ca. 1 km S of junction. Photos D.T. Bilton." figureDoi="http://doi.org/10.5281/zenodo.5271961" httpUri="https://zenodo.org/record/5271961/files/figure.png" pageId="8" pageNumber="34">Fig. 3B</figureCitation>
), a small stream flowing through Kamiesberg Granite Fynbos/Renosterveld (
<emphasis id="ACE2C578EF44FFE99ACCFA15FDDF7090" box="[495,557,1411,1434]" italics="true" pageId="8" pageNumber="34">sensu</emphasis>
Mucina &amp; Rutherford 2006). The high valleys and summits of the Kamiesberg range represent an isolated, northern outpost of the fynbos biome in relatively arid Namaqualand, and support other endemic aquatic beetles, including
<taxonomicName id="599662E9EF44FFE99902FA5CFCA670E9" authority="Sharp, 1882" authorityName="Sharp" authorityYear="1882" box="[545,852,1482,1507]" class="Insecta" family="Dytiscidae" genus="Andex" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="34" phylum="Arthropoda" rank="species" species="insignis">
<emphasis id="ACE2C578EF44FFE99902FA5CFD3770E8" box="[545,709,1482,1507]" italics="true" pageId="8" pageNumber="34">Andex insignis</emphasis>
<bibRefCitation id="FA07649BEF44FFE999EFFA5CFCA670E9" author="Sharp, D." box="[716,852,1482,1507]" pageId="8" pageNumber="34" pagination="179 - 1003" refId="ref7348" refString="Sharp, D. (1882) On aquatic carnivorous Coleoptera, or Dytiscidae. The Scientific Transactions of the Royal Dublin Society, (2), 2, 179 - 1003 + pls. 7 - 18." type="journal article" year="1882">Sharp, 1882</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="FA07649BEF44FFE99841FA5CFB9A70E9" author="Challet, G. &amp; Turner, C. R." box="[866,1128,1482,1507]" pageId="8" pageNumber="34" pagination="21 - 24" refId="ref6876" refString="Challet, G. &amp; Turner, C. R. (2006) Rediscovery of Coelhydrus brevicollis Sharp in South Africa with notes on Andex insignis Sharp (Coleptera: Dytiscidae). Latissimus, 21, 21 - 24." type="journal article" year="2006">Challet &amp; Turner 2006</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="D68C4AE1EF44FFE99BE4FA78FCA27321" pageId="8" pageNumber="34" type="etymology">
<paragraph id="9E29196AEF44FFE99BE4FA78FCA27321" blockId="8.[151,1436,1050,1759]" pageId="8" pageNumber="34">
<emphasis id="ACE2C578EF44FFE99BE4FA78FEBF730C" box="[199,333,1518,1543]" inLineHeading="true" pageId="8" pageNumber="34" reason="1">
<emphasis id="ACE2C578EF44FFE99BE4FA78FEB4730D" bold="true" box="[199,326,1518,1543]" pageId="8" pageNumber="34">Etymology</emphasis>
.
</emphasis>
Named after the Kamiesberg, an isolated granite massif in Namaqualand. The specific epithet is an adjective in the nominative singular standing in apposition.
</paragraph>
</subSubSection>
<subSubSection id="D68C4AE1EF44FFE99BE4F9A0FF0373D5" pageId="8" pageNumber="34" type="biology_ecology">
<paragraph id="9E29196AEF44FFE99BE4F9A0FF0373D5" blockId="8.[151,1436,1050,1759]" pageId="8" pageNumber="34">
<emphasis id="ACE2C578EF44FFE99BE4F9A0FED97344" box="[199,299,1590,1615]" inLineHeading="true" pageId="8" pageNumber="34" reason="1">
<emphasis id="ACE2C578EF44FFE99BE4F9A0FED67345" bold="true" box="[199,292,1590,1615]" pageId="8" pageNumber="34">Ecology</emphasis>
.
</emphasis>
Specimens were netted from relatively deep (ca.
<quantity id="596EB48FEF44FFE998ABF9A0FC277345" box="[904,981,1590,1615]" metricMagnitude="-1" metricUnit="m" metricValue="6.0" pageId="8" pageNumber="34" unit="cm" value="60.0">60 cm</quantity>
) water over gravel and silt in a pool immediately below a short riffle in a stream (
<figureCitation id="06AD05EFEF44FFE999B7F9CCFD1E7379" box="[660,748,1626,1651]" captionStart="FIGURE 3" captionStartId="7.[151,249,1826,1848]" captionTargetBox="[152,1435,193,1805]" captionTargetId="figure-73@7.[152,1435,193,1805]" captionTargetPageId="7" captionText="FIGURE 3. Type localities of: A) Sharphydrus brincki sp. nov. South Africa, Western Cape, Cederberg, Matjiesrivier Reserve, Matjies River ca. 3 km N of Cederberg Oasis; B) S. kamiesbergensis sp. nov. South Africa, Northern Cape, Kamiesberg, stream on Witwater–Langkloof road ca. 1 km S of junction. Photos D.T. Bilton." figureDoi="http://doi.org/10.5281/zenodo.5271961" httpUri="https://zenodo.org/record/5271961/files/figure.png" pageId="8" pageNumber="34">Fig. 3B</figureCitation>
). They occurred together with the water beetles
<taxonomicName id="599662E9EF44FFE99E2EF9CCFEF6739C" baseAuthorityName="Regimbart" baseAuthorityYear="1895" class="Insecta" family="Dytiscidae" genus="Sharphydrus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="34" phylum="Arthropoda" rank="species" species="coriaceus">
<emphasis id="ACE2C578EF44FFE99E2EF9CCFEF6739C" italics="true" pageId="8" pageNumber="34">Sharphydrus coriaceus</emphasis>
</taxonomicName>
,
<taxonomicName id="599662E9EF44FFE99A3DF9E8FC72739D" authority="(Gemminger &amp; Harold)" baseAuthorityName="Gemminger &amp; Harold" baseAuthorityYear="1868" box="[286,896,1662,1687]" class="Insecta" family="Dytiscidae" genus="Canthyporus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="34" phylum="Arthropoda" rank="species" species="hottentottus">
<emphasis id="ACE2C578EF44FFE99A3DF9E8FDB4739D" box="[286,582,1662,1687]" italics="true" pageId="8" pageNumber="34">Canthyporus hottentottus</emphasis>
(Gemminger &amp; Harold)
</taxonomicName>
,
<emphasis id="ACE2C578EF44FFE998B4F9E8FA92739C" box="[919,1376,1662,1687]" italics="true" pageId="8" pageNumber="34">
<taxonomicName id="599662E9EF44FFE998B4F9E8FBB8739C" authorityName="Sharp" authorityYear="1882" box="[919,1098,1662,1687]" class="Insecta" family="Dytiscidae" genus="Andex" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="34" phylum="Arthropoda" rank="species" species="insignis">Andex insignis</taxonomicName>
,
<taxonomicName id="599662E9EF44FFE99F41F9E8FA92739C" authorityName="Guignot" authorityYear="1936" box="[1122,1376,1662,1687]" class="Insecta" family="Dytiscidae" genus="Agabus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="34" phylum="Arthropoda" rank="species" species="ruwenzoricus">Agabus ruwenzoricus</taxonomicName>
</emphasis>
and
<taxonomicName id="599662E9EF44FFE99BB4F934FEA373B0" box="[151,337,1698,1723]" class="Insecta" family="Dytiscidae" genus="Nebrioporus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="34" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="ACE2C578EF44FFE99BB4F934FED773B1" box="[151,293,1698,1723]" italics="true" pageId="8" pageNumber="34">Nebrioporus</emphasis>
sp.
</taxonomicName>
One of the females collected retained an intact spermatophore tail, indicating recent mating activity.
</paragraph>
</subSubSection>
</treatment>
</document>
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