244 lines
35 KiB
XML
244 lines
35 KiB
XML
<document id="7F8880517F753532D9A641AD65464204" ID-CLB-Dataset="27622" ID-DOI="10.11646/zootaxa.4546.1.1" ID-GBIF-Dataset="aa93de4b-c9ed-4b3e-a1c2-3f5d90faba57" ID-ISSN="1175-5326" ID-Zenodo-Dep="2619020" ID-ZooBank="D2C9F49A-9B76-40AE-9A60-CAE9B99BA547" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553964486275" checkinUser="plazi" docAuthor="Huber, Bernhard A. & Carvalho, Leonardo S." docDate="2019" docId="E21587DBFFC9FF80FF11F8F54A82FF58" docLanguage="en" docName="zootaxa.4546.1.1.pdf" docOrigin="Zootaxa 4546 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Panjange togutil Huber & Carvalho 2019, sp. n." docType="treatment" docVersion="16" lastPageNumber="84" masterDocId="1E2CFFA3FF98FFD4FF86FFCD4B03FFA0" masterDocTitle="Filling the gaps: descriptions of unnamed species included in the latest molecular phylogeny of Pholcidae (Araneae)" masterLastPageNumber="96" masterPageNumber="1" pageNumber="82" updateTime="1698716013903" updateUser="ExternalLinkService">
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<mods:title id="C80BDD70F0C432CB5E9A4E136BB192EE">Filling the gaps: descriptions of unnamed species included in the latest molecular phylogeny of Pholcidae (Araneae)</mods:title>
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<mods:namePart id="B03D536692C6C2BDEEC1465A7A9CB9B6">Huber, Bernhard A.</mods:namePart>
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<mods:namePart id="27E0370E979F7D64BE1D473AF945B88E">Carvalho, Leonardo S.</mods:namePart>
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<mods:title id="1CBF92BB3A74FF9F62A0D51E9E3F52DF">Zootaxa</mods:title>
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<mods:date id="DFC42A5DAD9759CA1D1C0310C5B48DEB">2019</mods:date>
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<mods:number id="0897F668E7DA7B129D6701AD3950DC00">4546</mods:number>
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<treatment id="E21587DBFFC9FF80FF11F8F54A82FF58" ID-DOI="http://doi.org/10.5281/zenodo.5449799" ID-GBIF-Taxon="154531201" ID-Zenodo-Dep="5449799" LSID="urn:lsid:plazi:treatment:E21587DBFFC9FF80FF11F8F54A82FF58" httpUri="http://treatment.plazi.org/id/E21587DBFFC9FF80FF11F8F54A82FF58" lastPageId="84" lastPageNumber="84" pageId="81" pageNumber="82">
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<subSubSection id="22A66546FFC9FF85FF11F8F54901F8F3" box="[151,514,1848,1875]" pageId="81" pageNumber="82" type="nomenclature">
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<paragraph id="6A0336CDFFC9FF85FF11F8F54901F8F3" blockId="81.[151,528,1848,1908]" box="[151,514,1848,1875]" pageId="81" pageNumber="82">
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<heading id="314B81A1FFC9FF85FF11F8F54901F8F3" bold="true" box="[151,514,1848,1875]" fontSize="11" level="1" pageId="81" pageNumber="82" reason="1">
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<emphasis id="58C8EADFFFC9FF85FF11F8F54901F8F3" bold="true" box="[151,514,1848,1875]" pageId="81" pageNumber="82">
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<taxonomicName id="ADBC4D4EFFC9FF85FF11F8F54ABAF8F3" ID-CoL="4CFSQ" authority="Huber" authorityName="Huber & Carvalho" authorityYear="2019" box="[151,441,1848,1875]" class="Arachnida" family="Pholcidae" genus="Panjange" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="togutil" status="sp. nov.">
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<emphasis id="58C8EADFFFC9FF85FF11F8F54A63F8F2" bold="true" box="[151,352,1848,1874]" italics="true" pageId="81" pageNumber="82">Panjange togutil</emphasis>
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Huber
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</taxonomicName>
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<taxonomicNameLabel id="43FB57A4FFC9FF85FE46F8F44901F8F3" box="[448,514,1849,1875]" pageId="81" pageNumber="82" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="22A66546FFC9FF86FF11F896486BFB06" lastPageId="82" lastPageNumber="83" pageId="81" pageNumber="82" type="description">
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<paragraph id="6A0336CDFFC9FF85FF11F8964913F8D4" blockId="81.[151,528,1848,1908]" box="[151,528,1883,1908]" pageId="81" pageNumber="82">
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<figureCitation id="F2872A48FFC9FF85FF11F8964A31F8D4" box="[151,306,1883,1908]" captionStart="FIGURES 300–307" captionStartId="80.[151,264,1515,1537]" captionTargetBox="[151,1436,193,1494]" captionTargetId="figure@80.[151,1436,193,1494]" captionTargetPageId="80" captionText="FIGURES 300–307. Panjange spp. live specimens. 300–303. P. thomi sp. n., adult and penultimate instar males, and female with egg sac, from Gunung Meja. 304–307. P. togutil sp. n., adult and penultimate instar males, females with and without eggsac, from near Sofifi." httpUri="https://zenodo.org/record/2619136/files/figure.png" pageId="81" pageNumber="82">Figs 304–307</figureCitation>
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,
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<figureCitation id="F2872A48FFC9FF85FEB8F8964AA0F8D4" box="[318,419,1883,1908]" captionStart-0="FIGURES 312–313" captionStart-1="FIGURES 314–315" captionStartId-0="83.[151,264,1654,1676]" captionStartId-1="84.[151,264,1804,1826]" captionTargetBox-0="[158,1431,410,1618]" captionTargetBox-1="[152,1433,928,1782]" captionTargetId-0="figure@83.[151,1436,409,1633]" captionTargetId-1="figure@84.[151,1436,911,1782]" captionTargetPageId-0="83" captionTargetPageId-1="84" captionText-0="FIGURES 312–313. Panjange togutil sp. n. (ZFMK, Ar 20664), left male palp, prolateral and retrolateral views. ba, bulbal apophysis; em, embolus; gb, genital bulb; pr, procursus; tp, tarsal process. Scale line: 0.5." captionText-1="FIGURES 314–315. Panjange togutil sp. n. (ZFMK, Ar 20664–65), male prosoma and chelicerae, oblique frontal view, and cleared female genitalia, dorsal view. Scale lines: 0.3." httpUri-0="https://zenodo.org/record/2619142/files/figure.png" httpUri-1="https://zenodo.org/record/2619144/files/figure.png" pageId="81" pageNumber="82">312–315</figureCitation>
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,
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<figureCitation id="F2872A48FFC9FF85FE28F8914913F8D4" box="[430,528,1884,1908]" captionStart="FIGURES 316–321" captionStartId="85.[151,264,1397,1419]" captionTargetBox="[151,1436,193,1376]" captionTargetId="figure@85.[151,1436,193,1376]" captionTargetPageId="85" captionText="FIGURES 316–321. Panjange spp., epigyna in ventral views, and cleared female genitalia in ventral and dorsal views. 316– 318. P. thomi sp. n. (ZFMK, Ar 20662). 319–321. P. togutil sp. n. (ZFMK, Ar 20665). Scale lines: 0.3." httpUri="https://zenodo.org/record/2619146/files/figure.png" pageId="81" pageNumber="82">319–321</figureCitation>
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</paragraph>
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<paragraph id="6A0336CDFFC9FF86FF11F869486BFB06" blockId="81.[151,969,1955,1977]" lastBlockId="82.[151,1437,1129,2018]" lastPageId="82" lastPageNumber="83" pageId="81" pageNumber="82">
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<taxonomicName id="ADBC4D4EFFC9FF85FF11F8694A20F819" authority="Ind" authorityName="Deeleman-Reinhold & Deeleman" authorityYear="1983" box="[151,291,1955,1977]" class="Arachnida" family="Pholcidae" genus="Panjange" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="81" pageNumber="82" phylum="Arthropoda" rank="genus">
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<emphasis id="58C8EADFFFC9FF85FF11F8694BFBF819" box="[151,248,1956,1977]" italics="true" pageId="81" pageNumber="82">Panjange</emphasis>
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Ind
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</taxonomicName>
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109:
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<bibRefCitation id="0E2D4B3CFFC9FF85FED5F86E4904F819" author="Eberle, J. & Dimitrov, D. & Valdez-Mondragon, A. & Huber, B. A." box="[339,519,1955,1977]" pageId="81" pageNumber="82" pagination="141" refId="ref37668" refString="Eberle, J., Dimitrov, D., Valdez-Mondragon, A. & Huber, B. A. (2018) Microhabitat change drives diversification in pholcid spiders. BMC Evolutionary Biology, 18, 141. https: // doi. org / 10.1186 / s 12862 - 018 - 1244 - 8" type="journal article" year="2018">
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Eberle
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<emphasis id="58C8EADFFFC9FF85FE1AF8694ACCF819" box="[412,463,1955,1977]" italics="true" pageId="81" pageNumber="82">et al.</emphasis>
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2018
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</bibRefCitation>
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(molecular data); Huber
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<emphasis id="58C8EADFFFC9FF85FC81F8694839F819" box="[775,826,1955,1977]" italics="true" pageId="81" pageNumber="82">et al.</emphasis>
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2018: fig. 10.
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<emphasis id="58C8EADFFFCAFF86FF11FBA44A3DFB22" bold="true" box="[151,318,1129,1154]" pageId="82" pageNumber="83">Type material</emphasis>
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.
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<collectingCountry id="12AB765DFFCAFF86FECAFBA44AEAFB21" box="[332,489,1129,1153]" name="Indonesia" pageId="82" pageNumber="83">INDONESIA</collectingCountry>
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:
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<specimenCount id="7CBAFD44FFCAFF86FE73FBA74909FB21" box="[501,522,1130,1153]" pageId="82" pageNumber="83" type="male">♂</specimenCount>
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<typeStatus id="B507886FFFCAFF86FD97FBA44975FB22" box="[529,630,1129,1154]" pageId="82" pageNumber="83" type="holotype">holotype</typeStatus>
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, ZFMK (Ar 20663), Halmahera, degraded forest along river near Sofifi (
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<geoCoordinate id="0F88500AFFCAFF86FF19FB404A02FB06" box="[159,257,1165,1190]" direction="north" orientation="latitude" pageId="82" pageNumber="83" precision="55" value="0.715">0.715°N</geoCoordinate>
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,
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<geoCoordinate id="0F88500AFFCAFF86FE8AFB404A8BFB06" box="[268,392,1165,1190]" direction="east" orientation="longitude" pageId="82" pageNumber="83" precision="55" value="127.586">127.586°E</geoCoordinate>
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),
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<quantity id="AD449B28FFCAFF86FE1DFB434919FB05" box="[411,538,1166,1190]" metricMagnitude="2" metricUnit="m" metricValue="1.55" metricValueMax="1.8" metricValueMin="1.3" pageId="82" pageNumber="83" unit="m" value="155.0" valueMax="180.0" valueMin="130.0">130–180 m</quantity>
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a.s.l.,
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<date id="1E02100DFFCAFF86FDE5FB4049D9FB06" box="[611,730,1165,1190]" pageId="82" pageNumber="83" value="2009-11-11">11.xi.2009</date>
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(S. Sutono).
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</paragraph>
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</subSubSection>
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<caption id="3EC36645FFCAFF86FF11FBC049B4FBE2" httpUri="https://zenodo.org/record/2619140/files/figure.png" pageId="82" pageNumber="83" startId="82.[151,264,1037,1059]" targetBox="[173,1431,215,1012]" targetPageId="82">
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<paragraph id="6A0336CDFFCAFF86FF11FBC049B4FBE2" blockId="82.[151,1435,1037,1090]" pageId="82" pageNumber="83">
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<emphasis id="58C8EADFFFCAFF86FF11FBC04A6FFB83" bold="true" box="[151,364,1037,1059]" pageId="82" pageNumber="83">FIGURES 310–311.</emphasis>
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<taxonomicName id="ADBC4D4EFFCAFF86FEF3FBC3491BFB83" authority="Huber & Carvalho, 2019" authorityName="Huber & Carvalho" authorityYear="2019" box="[373,536,1037,1059]" class="Arachnida" family="Pholcidae" genus="Panjange" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="thomi" status="sp. nov.">
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<emphasis id="58C8EADFFFCAFF86FEF3FBC3491BFB83" box="[373,536,1037,1059]" italics="true" pageId="82" pageNumber="83">Panjange thomi</emphasis>
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</taxonomicName>
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<emphasis id="58C8EADFFFCAFF86FDA6FBC04958FB83" bold="true" box="[544,603,1037,1059]" pageId="82" pageNumber="83">
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<taxonomicNameLabel id="43FB57A4FFCAFF86FDA6FBC04958FB83" box="[544,603,1037,1059]" pageId="82" pageNumber="83" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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(ZFMK, Ar 20661–62), male prosoma and chelicerae, oblique frontal view, and cleared female genitalia, dorsal view. Scale lines: 0.3.
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</paragraph>
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</caption>
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<subSubSection id="22A66546FFCAFF86FF41FB7C4959FB4E" pageId="82" pageNumber="83" type="materials_examined">
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<paragraph id="6A0336CDFFCAFF86FF41FB7C4959FB4E" blockId="82.[151,1437,1129,2018]" pageId="82" pageNumber="83">
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<emphasis id="58C8EADFFFCAFF86FF41FB7C4AF4FB6A" bold="true" box="[199,503,1201,1226]" pageId="82" pageNumber="83">Other material examined</emphasis>
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.
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<materialsCitation id="DAD43C90FFCAFF86FD81FB7C4955FB4E" ID-GBIF-Occurrence="2147633829" country="Indonesia" pageId="82" pageNumber="83" specimenCount="8" specimenCount-female="5" specimenCount-male="3">
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<collectingCountry id="12AB765DFFCAFF86FD81FB7C49A7FB69" box="[519,676,1201,1225]" name="Indonesia" pageId="82" pageNumber="83">INDONESIA</collectingCountry>
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||
:
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<specimenCount id="7CBAFD44FFCAFF86FD36FB7C49D0FB69" box="[688,723,1201,1225]" pageId="82" pageNumber="83" type="male">3♂</specimenCount>
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<specimenCount id="7CBAFD44FFCAFF86FD5AFB7C49FCFB6A" box="[732,767,1201,1226]" pageId="82" pageNumber="83" type="female">5♀</specimenCount>
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7 juvs, ZFMK (Ar 20664–65), and 7 juvs in pure ethanol, ZFMK (Ind203), same data as holotype
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="22A66546FFCAFF86FF41FB344A33FA96" pageId="82" pageNumber="83" type="etymology">
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<paragraph id="6A0336CDFFCAFF86FF41FB344A33FA96" blockId="82.[151,1437,1129,2018]" pageId="82" pageNumber="83">
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<emphasis id="58C8EADFFFCAFF86FF41FB344A45FAB2" bold="true" box="[199,326,1273,1298]" pageId="82" pageNumber="83">Etymology</emphasis>
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. The species is named for the Togutil people in the interiors of north and central Halmahera; noun in apposition.
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</paragraph>
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</subSubSection>
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<subSubSection id="22A66546FFCAFF86FF41FA8C4966FA49" pageId="82" pageNumber="83" type="diagnosis">
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<paragraph id="6A0336CDFFCAFF86FF41FA8C4966FA49" blockId="82.[151,1437,1129,2018]" pageId="82" pageNumber="83">
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<emphasis id="58C8EADFFFCAFF86FF41FA8C4A39FAFA" bold="true" box="[199,314,1345,1370]" pageId="82" pageNumber="83">Diagnosis</emphasis>
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. Males are easily distinguished from other known representatives of the
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<taxonomicName id="ADBC4D4EFFCAFF86FBEBFA8C4FCFFAFA" authorityName="Deeleman-Reinhold & Deeleman" authorityYear="1983" box="[1133,1228,1345,1370]" class="Arachnida" family="Pholcidae" genus="Panjange" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="cavicola">
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<emphasis id="58C8EADFFFCAFF86FBEBFA8C4FCFFAFA" box="[1133,1228,1345,1370]" italics="true" pageId="82" pageNumber="83">cavicola</emphasis>
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</taxonomicName>
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group by strongly widening distal part of procursus (
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<figureCitation id="F2872A48FFCAFF86FD9DFAA849B9FADE" box="[539,698,1381,1406]" captionStart="FIGURES 312–313" captionStartId="83.[151,264,1654,1676]" captionTargetBox="[158,1431,410,1618]" captionTargetId="figure@83.[151,1436,409,1633]" captionTargetPageId="83" captionText="FIGURES 312–313. Panjange togutil sp. n. (ZFMK, Ar 20664), left male palp, prolateral and retrolateral views. ba, bulbal apophysis; em, embolus; gb, genital bulb; pr, procursus; tp, tarsal process. Scale line: 0.5." httpUri="https://zenodo.org/record/2619142/files/figure.png" pageId="82" pageNumber="83">Figs 312–313</figureCitation>
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; without dorsal process as in
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<taxonomicName id="ADBC4D4EFFCAFF86FB8EFAAB4F80FADE" authorityName="Deeleman-Reinhold & Deeleman" authorityYear="1983" box="[1032,1155,1381,1406]" class="Arachnida" family="Pholcidae" genus="Panjange" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="cavicola">
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<emphasis id="58C8EADFFFCAFF86FB8EFAAB4F80FADE" box="[1032,1155,1381,1406]" italics="true" pageId="82" pageNumber="83">P. cavicola</emphasis>
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</taxonomicName>
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) and by color pattern on carapace (
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<figureCitation id="F2872A48FFCAFF86FE8FFA444A68FA01" box="[265,363,1417,1442]" captionStart="FIGURES 300–307" captionStartId="80.[151,264,1515,1537]" captionTargetBox="[151,1436,193,1494]" captionTargetId="figure@80.[151,1436,193,1494]" captionTargetPageId="80" captionText="FIGURES 300–307. Panjange spp. live specimens. 300–303. P. thomi sp. n., adult and penultimate instar males, and female with egg sac, from Gunung Meja. 304–307. P. togutil sp. n., adult and penultimate instar males, females with and without eggsac, from near Sofifi." httpUri="https://zenodo.org/record/2619136/files/figure.png" pageId="82" pageNumber="83">Fig. 304</figureCitation>
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; anterior half pale whitish, posterior half dark brown); females differ (except from
|
||
<taxonomicName id="ADBC4D4EFFCAFF86FA91FA474E90FA02" authorityName="Deeleman-Reinhold & Deeleman" authorityYear="1983" box="[1303,1427,1417,1442]" class="Arachnida" family="Pholcidae" genus="Panjange" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="cavicola">
|
||
<emphasis id="58C8EADFFFCAFF86FA91FA474E90FA02" box="[1303,1427,1417,1442]" italics="true" pageId="82" pageNumber="83">P. cavicola</emphasis>
|
||
</taxonomicName>
|
||
) by short epigynal scape (
|
||
<figureCitation id="F2872A48FFCAFF86FE31FA60491CFA66" box="[439,543,1453,1478]" captionStart="FIGURES 314–315" captionStartId="84.[151,264,1804,1826]" captionTargetBox="[152,1433,928,1782]" captionTargetId="figure@84.[151,1436,911,1782]" captionTargetPageId="84" captionText="FIGURES 314–315. Panjange togutil sp. n. (ZFMK, Ar 20664–65), male prosoma and chelicerae, oblique frontal view, and cleared female genitalia, dorsal view. Scale lines: 0.3." httpUri="https://zenodo.org/record/2619144/files/figure.png" pageId="82" pageNumber="83">Figs 315</figureCitation>
|
||
,
|
||
<figureCitation id="F2872A48FFCAFF86FDAAFA634990FA66" box="[556,659,1454,1478]" captionStart="FIGURES 316–321" captionStartId="85.[151,264,1397,1419]" captionTargetBox="[151,1436,193,1376]" captionTargetId="figure@85.[151,1436,193,1376]" captionTargetPageId="85" captionText="FIGURES 316–321. Panjange spp., epigyna in ventral views, and cleared female genitalia in ventral and dorsal views. 316– 318. P. thomi sp. n. (ZFMK, Ar 20662). 319–321. P. togutil sp. n. (ZFMK, Ar 20665). Scale lines: 0.3." httpUri="https://zenodo.org/record/2619146/files/figure.png" pageId="82" pageNumber="83">319–320</figureCitation>
|
||
); both males and females also distinguished by posteriorly widened abdomen (in dorsal view;
|
||
<figureCitation id="F2872A48FFCAFF86FE3DFA1C495AFA49" box="[443,601,1489,1514]" captionStart="FIGURES 300–307" captionStartId="80.[151,264,1515,1537]" captionTargetBox="[151,1436,193,1494]" captionTargetId="figure@80.[151,1436,193,1494]" captionTargetPageId="80" captionText="FIGURES 300–307. Panjange spp. live specimens. 300–303. P. thomi sp. n., adult and penultimate instar males, and female with egg sac, from Gunung Meja. 304–307. P. togutil sp. n., adult and penultimate instar males, females with and without eggsac, from near Sofifi." httpUri="https://zenodo.org/record/2619136/files/figure.png" pageId="82" pageNumber="83">Figs 304–307</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="22A66546FFCAFF87FF41FA3848B2F804" lastPageId="83" lastPageNumber="84" pageId="82" pageNumber="83" type="description">
|
||
<paragraph id="6A0336CDFFCAFF86FF41FA3849E3F9F6" blockId="82.[151,1437,1129,2018]" pageId="82" pageNumber="83">
|
||
<emphasis id="58C8EADFFFCAFF86FF41FA384AA3F9AE" bold="true" box="[199,416,1525,1550]" pageId="82" pageNumber="83">Description. Male</emphasis>
|
||
(
|
||
<typeStatus id="B507886FFFCAFF86FE34FA384919F9AE" box="[434,538,1525,1550]" pageId="82" pageNumber="83" type="holotype">holotype</typeStatus>
|
||
). MEASUREMENTS. Total length 3.9, carapace width 1.0. Distance PME-PME 390 µm; diameter PME 90 µm; distance PME-ALE 30 µm; AME absent. Leg 1: 36.9 (8.5 + 0.4 + 8.7 + 17.4 + 1.9), tibia 2: 5.5, tibia 3: 3.0, tibia 4: 4.5; tibia 1 L/d: 109.
|
||
</paragraph>
|
||
<paragraph id="6A0336CDFFCAFF86FF41F9AC4892F962" blockId="82.[151,1437,1129,2018]" pageId="82" pageNumber="83">COLOR (in ethanol). Carapace anterior half pale whitish, posterior half dark brown; ocular area and clypeus also brown; sternum whitish; legs pale ochre-yellow, brown rings in patella area and at tibia-metatarsus joint. Abdomen pale ochre-yellow, with dark marks dorsally and laterally.</paragraph>
|
||
<paragraph id="6A0336CDFFCAFF86FF41F9034829F88E" blockId="82.[151,1437,1129,2018]" pageId="82" pageNumber="83">
|
||
BODY. Habitus as in
|
||
<figureCitation id="F2872A48FFCAFF86FE34F9004910F946" box="[434,531,1741,1766]" captionStart="FIGURES 300–307" captionStartId="80.[151,264,1515,1537]" captionTargetBox="[151,1436,193,1494]" captionTargetId="figure@80.[151,1436,193,1494]" captionTargetPageId="80" captionText="FIGURES 300–307. Panjange spp. live specimens. 300–303. P. thomi sp. n., adult and penultimate instar males, and female with egg sac, from Gunung Meja. 304–307. P. togutil sp. n., adult and penultimate instar males, females with and without eggsac, from near Sofifi." httpUri="https://zenodo.org/record/2619136/files/figure.png" pageId="82" pageNumber="83">Fig. 304</figureCitation>
|
||
. Eye triads on stalks (
|
||
<figureCitation id="F2872A48FFCAFF86FC8DF900486DF946" box="[779,878,1741,1766]" captionStart="FIGURES 314–315" captionStartId="84.[151,264,1804,1826]" captionTargetBox="[152,1433,928,1782]" captionTargetId="figure@84.[151,1436,911,1782]" captionTargetPageId="84" captionText="FIGURES 314–315. Panjange togutil sp. n. (ZFMK, Ar 20664–65), male prosoma and chelicerae, oblique frontal view, and cleared female genitalia, dorsal view. Scale lines: 0.3." httpUri="https://zenodo.org/record/2619144/files/figure.png" pageId="82" pageNumber="83">Fig. 314</figureCitation>
|
||
), each stalk with process ending in two tips each. Thoracic furrow absent. Clypeus unmodified. Sternum wider than long (0.62/0.50), unmodified. Abdomen cylindrical, widened posteriorly, pointed dorso-posteriorly.
|
||
</paragraph>
|
||
<paragraph id="6A0336CDFFCAFF86FF41F8F44FBAF8F2" blockId="82.[151,1437,1129,2018]" box="[199,1209,1849,1874]" pageId="82" pageNumber="83">
|
||
CHELICERAE. As in
|
||
<figureCitation id="F2872A48FFCAFF86FE21F8F44904F8F1" box="[423,519,1849,1874]" captionStart="FIGURES 314–315" captionStartId="84.[151,264,1804,1826]" captionTargetBox="[152,1433,928,1782]" captionTargetId="figure@84.[151,1436,911,1782]" captionTargetPageId="84" captionText="FIGURES 314–315. Panjange togutil sp. n. (ZFMK, Ar 20664–65), male prosoma and chelicerae, oblique frontal view, and cleared female genitalia, dorsal view. Scale lines: 0.3." httpUri="https://zenodo.org/record/2619144/files/figure.png" pageId="82" pageNumber="83">Fig. 314</figureCitation>
|
||
, with pair of simple weakly sclerotized processes proximally.
|
||
</paragraph>
|
||
<paragraph id="6A0336CDFFCAFF87FF41F8934AD6FF58" blockId="82.[151,1437,1129,2018]" lastBlockId="83.[151,1436,151,357]" lastPageId="83" lastPageNumber="84" pageId="82" pageNumber="83">
|
||
PALPS. As in
|
||
<figureCitation id="F2872A48FFCAFF86FEE7F8904AFDF8D6" box="[353,510,1885,1910]" captionStart="FIGURES 312–313" captionStartId="83.[151,264,1654,1676]" captionTargetBox="[158,1431,410,1618]" captionTargetId="figure@83.[151,1436,409,1633]" captionTargetPageId="83" captionText="FIGURES 312–313. Panjange togutil sp. n. (ZFMK, Ar 20664), left male palp, prolateral and retrolateral views. ba, bulbal apophysis; em, embolus; gb, genital bulb; pr, procursus; tp, tarsal process. Scale line: 0.5." httpUri="https://zenodo.org/record/2619142/files/figure.png" pageId="82" pageNumber="83">Figs 312–313</figureCitation>
|
||
, symmetric; coxa with small but distinct retrolateral-ventral apophysis, trochanter with retrolateral-dorsal process, femur with rounded dorsal process proximally; femur-patella joints strongly shifted towards prolateral side; tarsus with long whitish dorsal process with tarsal organ near tip; procursus complex, distal part apparently hinged, proximal part with two ventral processes, without ventral parallel ridges; brush of hairs dorsally between proximal and distal parts; distal part wide and flat; genital bulb oval, with two processes lying parallel to each other: semitransparent embolus and slightly sclerotized long bulbal apophysis (‘appendix’; ba in
|
||
<figureCitation id="F2872A48FFCBFF87FEE0FF124ACAFF58" box="[358,457,223,248]" captionStart="FIGURES 312–313" captionStartId="83.[151,264,1654,1676]" captionTargetBox="[158,1431,410,1618]" captionTargetId="figure@83.[151,1436,409,1633]" captionTargetPageId="83" captionText="FIGURES 312–313. Panjange togutil sp. n. (ZFMK, Ar 20664), left male palp, prolateral and retrolateral views. ba, bulbal apophysis; em, embolus; gb, genital bulb; pr, procursus; tp, tarsal process. Scale line: 0.5." httpUri="https://zenodo.org/record/2619142/files/figure.png" pageId="83" pageNumber="84">Fig. 312</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="6A0336CDFFCBFF87FF41FEC84A36FEC5" blockId="83.[151,1436,151,357]" pageId="83" pageNumber="84">LEGS. Without spines and curved hairs, few vertical hairs; retrolateral trichobothrium of tibia 1 at 3%; tibia 1 without prolateral trichobothrium (present on other tibiae); tarsus 1 with ~30 pseudosegments, only distally a few fairly distinct.</paragraph>
|
||
<caption id="3EC36645FFCBFF87FF11F9BB4F22F90A" httpUri="https://zenodo.org/record/2619142/files/figure.png" pageId="83" pageNumber="84" startId="83.[151,264,1654,1676]" targetBox="[158,1431,410,1618]" targetPageId="83">
|
||
<paragraph id="6A0336CDFFCBFF87FF11F9BB4F22F90A" blockId="83.[151,1435,1654,1706]" pageId="83" pageNumber="84">
|
||
<emphasis id="58C8EADFFFCBFF87FF11F9BB4A6DF92C" bold="true" box="[151,366,1654,1676]" pageId="83" pageNumber="84">FIGURES 312–313.</emphasis>
|
||
<taxonomicName id="ADBC4D4EFFCBFF87FEF0F9BA4921F92C" authority="Huber & Carvalho, 2019" authorityName="Huber & Carvalho" authorityYear="2019" box="[374,546,1654,1676]" class="Arachnida" family="Pholcidae" genus="Panjange" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="togutil" status="sp. nov.">
|
||
<emphasis id="58C8EADFFFCBFF87FEF0F9BA4921F92C" box="[374,546,1654,1676]" italics="true" pageId="83" pageNumber="84">Panjange togutil</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="58C8EADFFFCBFF87FDACF9BB4967F92C" bold="true" box="[554,612,1654,1676]" pageId="83" pageNumber="84">
|
||
<taxonomicNameLabel id="43FB57A4FFCBFF87FDACF9BB4967F92C" box="[554,612,1654,1676]" pageId="83" pageNumber="84" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(ZFMK, Ar 20664), left male palp, prolateral and retrolateral views. ba, bulbal apophysis; em, embolus; gb, genital bulb; pr, procursus; tp, tarsal process. Scale line: 0.5.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="6A0336CDFFCBFF87FF41F91A4837F950" blockId="83.[151,1436,1751,2028]" box="[199,820,1751,1776]" pageId="83" pageNumber="84">
|
||
<emphasis id="58C8EADFFFCBFF87FF41F91A4A07F950" bold="true" box="[199,260,1751,1776]" pageId="83" pageNumber="84">Male</emphasis>
|
||
(variation). Tibia
|
||
<quantity id="AD449B28FFCBFF87FE56F9154AF8F950" box="[464,507,1751,1776]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="83" pageNumber="84" unit="in" value="1.0">1 in</quantity>
|
||
3 other males: 9.0, 9.5, 9.9.
|
||
</paragraph>
|
||
<paragraph id="6A0336CDFFCBFF87FF41F93648B2F804" blockId="83.[151,1436,1751,2028]" pageId="83" pageNumber="84">
|
||
<emphasis id="58C8EADFFFCBFF87FF41F9364A1DF8B4" bold="true" box="[199,286,1787,1812]" pageId="83" pageNumber="84">Female</emphasis>
|
||
. In general similar to male (
|
||
<figureCitation id="F2872A48FFCBFF87FDF6F9364813F8B4" box="[624,784,1787,1812]" captionStart="FIGURES 300–307" captionStartId="80.[151,264,1515,1537]" captionTargetBox="[151,1436,193,1494]" captionTargetId="figure@80.[151,1436,193,1494]" captionTargetPageId="80" captionText="FIGURES 300–307. Panjange spp. live specimens. 300–303. P. thomi sp. n., adult and penultimate instar males, and female with egg sac, from Gunung Meja. 304–307. P. togutil sp. n., adult and penultimate instar males, females with and without eggsac, from near Sofifi." httpUri="https://zenodo.org/record/2619136/files/figure.png" pageId="83" pageNumber="84">Figs 306–307</figureCitation>
|
||
) but eye triads on low humps, closer together (distance PME-PME 210 µm) and clypeus not darkened (only dark band between eye triads). Carapace pattern in most females like in males (but with light median line dividing dark mark), in one female different (
|
||
<figureCitation id="F2872A48FFCBFF87FB3CF88E4E1EF8FC" box="[1210,1309,1859,1884]" captionStart="FIGURES 300–307" captionStartId="80.[151,264,1515,1537]" captionTargetBox="[151,1436,193,1494]" captionTargetId="figure@80.[151,1436,193,1494]" captionTargetPageId="80" captionText="FIGURES 300–307. Panjange spp. live specimens. 300–303. P. thomi sp. n., adult and penultimate instar males, and female with egg sac, from Gunung Meja. 304–307. P. togutil sp. n., adult and penultimate instar males, females with and without eggsac, from near Sofifi." httpUri="https://zenodo.org/record/2619136/files/figure.png" pageId="83" pageNumber="84">Fig. 307</figureCitation>
|
||
). Tibia
|
||
<quantity id="AD449B28FFCBFF87FAF7F8894E9FF8FC" box="[1393,1436,1859,1884]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="83" pageNumber="84" unit="in" value="1.0">1 in</quantity>
|
||
<specimenCount id="7CBAFD44FFCBFF87FF11F8AA4A04F820" box="[151,263,1895,1920]" pageId="83" pageNumber="84" type="female">5 females</specimenCount>
|
||
: 6.7–8.0 (mean 7.5). Epigynum weakly sclerotized (
|
||
<figureCitation id="F2872A48FFCBFF87FCD5F8AA48B5F820" box="[851,950,1895,1920]" captionStart="FIGURES 316–321" captionStartId="85.[151,264,1397,1419]" captionTargetBox="[151,1436,193,1376]" captionTargetId="figure@85.[151,1436,193,1376]" captionTargetPageId="85" captionText="FIGURES 316–321. Panjange spp., epigyna in ventral views, and cleared female genitalia in ventral and dorsal views. 316– 318. P. thomi sp. n. (ZFMK, Ar 20662). 319–321. P. togutil sp. n. (ZFMK, Ar 20665). Scale lines: 0.3." httpUri="https://zenodo.org/record/2619146/files/figure.png" pageId="83" pageNumber="84">Fig. 319</figureCitation>
|
||
), with short folded scape. Internal genitalia with small median pocket and pair of large pore plates (
|
||
<figureCitation id="F2872A48FFCBFF87FC80F846486FF804" box="[774,876,1931,1956]" captionStart="FIGURES 314–315" captionStartId="84.[151,264,1804,1826]" captionTargetBox="[152,1433,928,1782]" captionTargetId="figure@84.[151,1436,911,1782]" captionTargetPageId="84" captionText="FIGURES 314–315. Panjange togutil sp. n. (ZFMK, Ar 20664–65), male prosoma and chelicerae, oblique frontal view, and cleared female genitalia, dorsal view. Scale lines: 0.3." httpUri="https://zenodo.org/record/2619144/files/figure.png" pageId="83" pageNumber="84">Figs 315</figureCitation>
|
||
,
|
||
<figureCitation id="F2872A48FFCBFF87FCF1F84148A6F804" box="[887,933,1932,1956]" captionStart="FIGURES 316–321" captionStartId="85.[151,264,1397,1419]" captionTargetBox="[151,1436,193,1376]" captionTargetId="figure@85.[151,1436,193,1376]" captionTargetPageId="85" captionText="FIGURES 316–321. Panjange spp., epigyna in ventral views, and cleared female genitalia in ventral and dorsal views. 316– 318. P. thomi sp. n. (ZFMK, Ar 20662). 319–321. P. togutil sp. n. (ZFMK, Ar 20665). Scale lines: 0.3." httpUri="https://zenodo.org/record/2619146/files/figure.png" pageId="83" pageNumber="84">321</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="22A66546FFCBFF80FF41F8624A82FF58" lastPageId="84" lastPageNumber="85" pageId="83" pageNumber="84" type="distribution">
|
||
<paragraph id="6A0336CDFFCBFF87FF41F8624842F868" blockId="83.[151,1436,1751,2028]" box="[199,833,1967,1992]" pageId="83" pageNumber="84">
|
||
<emphasis id="58C8EADFFFCBFF87FF41F8624A5AF868" bold="true" box="[199,345,1967,1992]" pageId="83" pageNumber="84">Distribution</emphasis>
|
||
. Known from
|
||
<typeStatus id="B507886FFFCBFF87FE78F87D492DF868" box="[510,558,1968,1992]" pageId="83" pageNumber="84">type</typeStatus>
|
||
locality only (
|
||
<figureCitation id="F2872A48FFCBFF87FD57F8624836F868" box="[721,821,1967,1992]" captionStart="FIGURE 350" captionStartId="93.[151,250,926,948]" captionTargetBox="[151,1436,193,904]" captionTargetId="figure@93.[151,1436,193,904]" captionTargetPageId="93" captionText="FIGURE 350. Known distribution of Panjange in eastern Indonesia (not shown: record of P. alba from Ambon; see Huber 2011, fig. 435)." httpUri="https://zenodo.org/record/2619170/files/figure.png" pageId="83" pageNumber="84">Fig. 350</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="6A0336CDFFCBFF80FF41F81E4A82FF58" blockId="83.[151,1436,1751,2028]" lastBlockId="84.[151,1436,151,248]" lastPageId="84" lastPageNumber="85" pageId="83" pageNumber="84">
|
||
<emphasis id="58C8EADFFFCBFF87FF41F81E4A7CF84C" bold="true" box="[199,383,2003,2028]" pageId="83" pageNumber="84">Natural history</emphasis>
|
||
. The spiders were found in a disturbed forest (
|
||
<figureCitation id="F2872A48FFCBFF87FC27F81E4F07F84C" box="[929,1028,2003,2028]" captionStart="FIGURES 336–341" captionStartId="88.[151,264,1639,1661]" captionTargetBox="[151,1436,193,1618]" captionTargetId="figure@88.[151,1436,193,1618]" captionTargetPageId="88" captionText="FIGURES 336–341. Selection of collecting sites and habitats. 336. Forest above Mokwam (West Papua), type locality of Arnapa arfak sp. n. and Aetana mokwam sp. n. 337. Tinoor Fall near Tomohon (North Sulawesi), type locality of Arnapa tinoor sp. n. 338. Entrance to Caverna Porco do Mato II (Rio Grande do Norte), type locality of Pinocchio barauna sp. n. 339. Desert between Sur and Kamil (Oman), type locality of Magana velox sp. n. 340. Bromeliad at Reserva Particular do Patrimônio Nacional Serra Bonita (Bahia), type locality of Psilochorus bromelicolus sp. n. (note pair of spiders among leaves). 341. Disturbed forest near Sofifi (Halmahera), type locality of Giloloa sofifi sp. n. and Panjange togutil sp. n." httpUri="https://zenodo.org/record/2619152/files/figure.png" pageId="83" pageNumber="84">Fig. 341</figureCitation>
|
||
), on the undersides of green leaves about
|
||
<quantity id="AD449B28FFCCFF80FF5BFF554A27FF0F" box="[221,292,152,176]" metricMagnitude="0" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="84" pageNumber="85" unit="m" value="1.5" valueMax="2.0" valueMin="1.0">1–2 m</quantity>
|
||
above the ground. The indistinct domed webs were connected to the undersides of the leaves at their apex, with a few silk lines extending beyond the leaves. Most specimens were found in a dryer patch of forest higher up on the hill.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |