320 lines
40 KiB
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320 lines
40 KiB
XML
<document ID-DOI="10.5733/afin.056.0217" ID-ISSN="2305-2562" ID-Zenodo-Dep="7914310" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1683585688736" checkinUser="felipe" docAuthor="Zonstein, Sergei L. & Marusik, Yuri M." docDate="2015" docId="D80A4A02FFDDFFCCFE52FA31C4B2AB88" docLanguage="en" docName="AfrInvertebr.56.2.483-489.pdf" docOrigin="African Invertebrates 56 (2)" docSource="http://www.bioone.org/doi/10.5733/afin.056.0217" docStyle="DocumentStyle:B78C63EC8CEBBB75364A82B35BB1F83D.3:AfrInvertebr.2008-2015.journal_article.0cover" docStyleId="B78C63EC8CEBBB75364A82B35BB1F83D" docStyleName="AfrInvertebr.2008-2015.journal_article.0cover" docStyleVersion="3" docTitle="Andoharano ansieae Zonstein & Marusik 2015, sp. n." docType="treatment" docVersion="3" lastPageNumber="487" masterDocId="2433327AFFDCFFC8FFAEFE34C633AF49" masterDocTitle="The first record of Andoharano Lehtinen, 1967 (Araneae: Filistatidae) from mainland Africa" masterLastPageNumber="483" masterPageNumber="483" pageNumber="484" updateTime="1683635461506" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>The first record of Andoharano Lehtinen, 1967 (Araneae: Filistatidae) from mainland Africa</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Zonstein, Sergei L.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Marusik, Yuri M.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>African Invertebrates</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2015</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2015-08-12</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>56</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>2</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>483</mods:start>
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<mods:end>483</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://www.bioone.org/doi/10.5733/afin.056.0217</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.5733/afin.056.0217</mods:identifier>
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<mods:identifier type="ISSN">2305-2562</mods:identifier>
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<mods:identifier type="Zenodo-Dep">7914310</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:D80A4A02FFDDFFCCFE52FA31C4B2AB88" httpUri="http://treatment.plazi.org/id/D80A4A02FFDDFFCCFE52FA31C4B2AB88" lastPageId="4" lastPageNumber="487" pageId="1" pageNumber="484">
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<subSubSection box="[508,815,1029,1056]" pageId="1" pageNumber="484" type="nomenclature">
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<paragraph blockId="1.[181,1142,1029,1677]" box="[508,815,1029,1056]" pageId="1" pageNumber="484">
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<heading box="[508,815,1029,1056]" centered="true" fontSize="11" level="2" pageId="1" pageNumber="484" reason="2">
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<taxonomicName ID-CoL="DQJF" authority="Zonstein & Marusik, 2015" authorityName="Zonstein & Marusik" authorityYear="2015" box="[508,748,1029,1056]" class="Arachnida" family="Filistatidae" genus="Andoharano" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="484" phylum="Arthropoda" rank="species" species="ansieae" status="sp. n.">
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<emphasis bold="true" box="[508,748,1029,1056]" italics="true" pageId="1" pageNumber="484">Andoharano ansieae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[755,815,1030,1056]" pageId="1" pageNumber="484" rank="species">sp. n.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="4" lastPageNumber="487" pageId="1" pageNumber="484" type="description">
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<paragraph blockId="1.[181,1142,1029,1677]" box="[564,759,1070,1097]" pageId="1" pageNumber="484">
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<figureCitation box="[564,677,1070,1096]" captionStart-0="Figs 1–5" captionStart-1="Figs 6–13" captionStartId-0="2.[144,183,664,686]" captionStartId-1="3.[182,221,1540,1562]" captionTargetBox-0="[161,1091,204,645]" captionTargetBox-1="[193,1130,652,1515]" captionTargetId-0="figure-412@2.[157,1091,204,645]" captionTargetId-1="figure-262@3.[193,1130,652,1515]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="Figs 1–5. Habitus and somatic characters of Andoharano ansieae sp. n., (1) male habitus, lateral; (2) eye fielD, lateral; (3) female habitus, Dorsal; (4) female prosoma, Ventral; (5) calamistrum, retrolateral." captionText-1="Figs 6–13. Copulatory organs of Andoharano ansieae sp. n. (6–12) and A. grandidieri (13). (6, 7) male palp, prolateral; (8) male palp, retrolateral; (9) terminal part of male palp, prolateral; (10) terminal part of male palp and schematic course of spermophor, retrolateral; (11) endogyne, dorsal; (12) receptacle, dorsal; (13) terminal part of male palp, prolateral view and from above (after Lehtinen 1967). Abbreviations: Bc coil directed backward; Fl forelock." figureDoi-0="http://doi.org/10.5281/zenodo.7914312" figureDoi-1="http://doi.org/10.5281/zenodo.7914316" httpUri-0="https://zenodo.org/record/7914312/files/figure.png" httpUri-1="https://zenodo.org/record/7914316/files/figure.png" pageId="1" pageNumber="484">Figs 1–12</figureCitation>
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,
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<figureCitation box="[689,759,1070,1097]" captionStart="Figs 14, 15" captionStartId="4.[143,182,592,614]" captionTargetBox="[151,1095,205,574]" captionTargetId="figure-384@4.[151,1096,204,575]" captionTargetPageId="4" captionText="Figs 14, 15. Andoharano ansieae sp. n., female somatic characters.(14) junction of tibia and metatarsus IV showing calamistrum, retrolateral; (15) cribellum and spinnerets, ventral." figureDoi="http://doi.org/10.5281/zenodo.7914318" httpUri="https://zenodo.org/record/7914318/files/figure.png" pageId="1" pageNumber="484">14, 15</figureCitation>
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</paragraph>
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<paragraph blockId="1.[181,1142,1029,1677]" pageId="1" pageNumber="484">Etymology: The species is named in honour of Dr Ansie Dippenaar-Schoeman, for her immense contribution to the knowledge of African spiders.</paragraph>
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<paragraph blockId="1.[181,1142,1029,1677]" pageId="1" pageNumber="484">
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Diagnosis: The new species differs: from
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<taxonomicName authorityName="Fage" authorityYear="1945" box="[657,774,1184,1211]" class="Arachnida" family="Filistatidae" genus="Filistata" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="484" phylum="Arthropoda" rank="species" species="decaryi">
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<emphasis box="[657,774,1184,1211]" italics="true" pageId="1" pageNumber="484">A. decaryi</emphasis>
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</taxonomicName>
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by the absence of an hourglassshaped dark spot on the carapace (only eye tubercle is darkened); from
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<taxonomicName baseAuthorityName="Simon" baseAuthorityYear="1901" box="[958,1114,1216,1243]" class="Arachnida" family="Filistatidae" genus="Andoharano" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="484" phylum="Arthropoda" rank="species" species="grandidieri">
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<emphasis box="[958,1114,1216,1243]" italics="true" pageId="1" pageNumber="484">A. grandidieri</emphasis>
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</taxonomicName>
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in having the embolus sharply curved downward (vs. slightly curved sideward in the latter species), as well as by the male palpal tibia weakly dilated apically (vs. strongly dilated; cf.
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<figureCitation box="[216,397,1312,1339]" captionStart="Figs 6–13" captionStartId="3.[182,221,1540,1562]" captionTargetBox="[193,1130,652,1515]" captionTargetId="figure-262@3.[193,1130,652,1515]" captionTargetPageId="3" captionText="Figs 6–13. Copulatory organs of Andoharano ansieae sp. n. (6–12) and A. grandidieri (13). (6, 7) male palp, prolateral; (8) male palp, retrolateral; (9) terminal part of male palp, prolateral; (10) terminal part of male palp and schematic course of spermophor, retrolateral; (11) endogyne, dorsal; (12) receptacle, dorsal; (13) terminal part of male palp, prolateral view and from above (after Lehtinen 1967). Abbreviations: Bc coil directed backward; Fl forelock." figureDoi="http://doi.org/10.5281/zenodo.7914316" httpUri="https://zenodo.org/record/7914316/files/figure.png" pageId="1" pageNumber="484">Figs 6–8 and 13</figureCitation>
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); from
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<taxonomicName authorityName="Legendre" authorityYear="1971" box="[478,581,1312,1339]" class="Arachnida" family="Filistatidae" genus="Andoharano" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="484" phylum="Arthropoda" rank="species" species="milloti">
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<emphasis box="[478,581,1312,1339]" italics="true" pageId="1" pageNumber="484">A. milloti</emphasis>
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</taxonomicName>
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by the shorter male palp (2.5 and 2.0× shorter than the body, in the former and the latter species, respectively); and, from
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<taxonomicName authorityName="Legendre" authorityYear="1971" box="[954,1067,1344,1371]" class="Arachnida" family="Filistatidae" genus="Andoharano" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="484" phylum="Arthropoda" rank="species" species="monodi">
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<emphasis box="[954,1067,1344,1371]" italics="true" pageId="1" pageNumber="484">A. monodi</emphasis>
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</taxonomicName>
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by the diffuse darker colouration of the femora, which are adorned with contrasting brown rings in the latter species. In males of
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<taxonomicName authority="Zonstein & Marusik, 2015" authorityName="Zonstein & Marusik" authorityYear="2015" box="[607,724,1409,1435]" class="Arachnida" family="Filistatidae" genus="Andoharano" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="484" phylum="Arthropoda" rank="species" species="ansieae" status="sp. n.">
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<emphasis box="[607,724,1409,1435]" italics="true" pageId="1" pageNumber="484">A. ansieae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[730,790,1409,1435]" pageId="1" pageNumber="484" rank="species">sp. n.</taxonomicNameLabel>
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and
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<taxonomicName authorityName="Fage" authorityYear="1945" box="[844,959,1408,1435]" class="Arachnida" family="Filistatidae" genus="Filistata" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="484" phylum="Arthropoda" rank="species" species="decaryi">
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<emphasis box="[844,959,1408,1435]" italics="true" pageId="1" pageNumber="484">A. decaryi</emphasis>
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</taxonomicName>
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the leg I is 2.5× longer than the body, in
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<taxonomicName authorityName="Legendre" authorityYear="1971" box="[442,544,1440,1467]" class="Arachnida" family="Filistatidae" genus="Andoharano" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="484" phylum="Arthropoda" rank="species" species="milloti">
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<emphasis box="[442,544,1440,1467]" italics="true" pageId="1" pageNumber="484">A. milloti</emphasis>
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</taxonomicName>
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3.3×, and in
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<taxonomicName baseAuthorityName="Simon" baseAuthorityYear="1901" box="[683,839,1440,1467]" class="Arachnida" family="Filistatidae" genus="Andoharano" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="484" phylum="Arthropoda" rank="species" species="grandidieri">
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<emphasis box="[683,839,1440,1467]" italics="true" pageId="1" pageNumber="484">A. grandidieri</emphasis>
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</taxonomicName>
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6.2× (
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<bibRefCitation author="FAGE, L." box="[907,1027,1440,1467]" pageId="1" pageNumber="484" pagination="301 - 307" refId="ref3073" refString="FAGE, L. 1945. Arachnides cavernicoles nouveaux de Madagascar. Bulletin du Museum National d'Histoire Naturelle de Paris (2) 17: 301 - 307." type="journal article" year="1945">Fage 1945</bibRefCitation>
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;
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<bibRefCitation author="LEGENDRE, R." pageId="1" pageNumber="484" pagination="645 - 650" refId="ref3168" refString="LEGENDRE, R. 1971. Deux nouvelles araignees Filistatidae cavernicoles de Madagascar: Andoharano milloti n. sp. et Andoharano monodi n. sp. Bulletin du Museum National d'Histoire Naturelle de Paris 13: 645 - 650." type="journal article" year="1971">Legendre 1971</bibRefCitation>
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). None of the other species possess a thorn in the basal part of the embolus (cf.
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<figureCitation captionStart="Figs 6–13" captionStartId="3.[182,221,1540,1562]" captionTargetBox="[193,1130,652,1515]" captionTargetId="figure-262@3.[193,1130,652,1515]" captionTargetPageId="3" captionText="Figs 6–13. Copulatory organs of Andoharano ansieae sp. n. (6–12) and A. grandidieri (13). (6, 7) male palp, prolateral; (8) male palp, retrolateral; (9) terminal part of male palp, prolateral; (10) terminal part of male palp and schematic course of spermophor, retrolateral; (11) endogyne, dorsal; (12) receptacle, dorsal; (13) terminal part of male palp, prolateral view and from above (after Lehtinen 1967). Abbreviations: Bc coil directed backward; Fl forelock." figureDoi="http://doi.org/10.5281/zenodo.7914316" httpUri="https://zenodo.org/record/7914316/files/figure.png" pageId="1" pageNumber="484">Figs 9, 10</figureCitation>
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anD
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<bibRefCitation author="LEHTINEN, P. T." box="[291,457,1504,1530]" pageId="1" pageNumber="484" pagination="199 - 468" refId="ref3209" refString="LEHTINEN, P. T. 1967. Classification of the cribellate spiDers anD some allieD families, with notes on the evolution of the suborder Araneomorpha. Annales Zoologici Fennici 4: 199 - 468." type="journal article" year="1967">Lehtinen 1967</bibRefCitation>
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: fig. 24). ADDitionallY,
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<taxonomicName authority="Zonstein & Marusik, 2015" authorityName="Zonstein & Marusik" authorityYear="2015" box="[714,831,1505,1531]" class="Arachnida" family="Filistatidae" genus="Andoharano" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="484" phylum="Arthropoda" rank="species" species="ansieae" status="sp. n.">
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<emphasis box="[714,831,1505,1531]" italics="true" pageId="1" pageNumber="484">A. ansieae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[838,897,1505,1531]" pageId="1" pageNumber="484" rank="species">sp. n.</taxonomicNameLabel>
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may be distinguished from all previously described congeners, except
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<taxonomicName authorityName="Fage" authorityYear="1945" box="[733,849,1536,1563]" class="Arachnida" family="Filistatidae" genus="Filistata" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="484" phylum="Arthropoda" rank="species" species="decaryi">
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<emphasis box="[733,849,1536,1563]" italics="true" pageId="1" pageNumber="484">A. decaryi</emphasis>
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</taxonomicName>
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, by the weakly decorated (vs. uniformly coloured) dorsal opistosoma.
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</paragraph>
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<paragraph blockId="1.[181,1142,1029,1677]" box="[181,321,1610,1636]" pageId="1" pageNumber="484">Description:</paragraph>
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<paragraph blockId="1.[181,1142,1029,1677]" box="[181,370,1651,1677]" pageId="1" pageNumber="484">
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<emphasis box="[181,239,1651,1677]" italics="true" pageId="1" pageNumber="484">Male</emphasis>
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(
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<typeStatus box="[254,359,1651,1677]" pageId="1" pageNumber="484">holotype</typeStatus>
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).
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</paragraph>
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<caption ID-DOI="http://doi.org/10.5281/zenodo.7914312" ID-Zenodo-Dep="7914312" httpUri="https://zenodo.org/record/7914312/files/figure.png" pageId="2" pageNumber="485" startId="2.[144,183,664,686]" targetBox="[161,1091,204,645]" targetPageId="2" targetType="figure">
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<paragraph blockId="2.[144,1103,664,710]" pageId="2" pageNumber="485">
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Figs 1–5. Habitus and somatic characters of
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<taxonomicName authority="Zonstein & Marusik, 2015" authorityName="Zonstein & Marusik" authorityYear="2015" box="[552,741,664,686]" class="Arachnida" family="Filistatidae" genus="Andoharano" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="485" phylum="Arthropoda" rank="species" species="ansieae" status="sp. n.">
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<emphasis box="[552,741,664,686]" italics="true" pageId="2" pageNumber="485">Andoharano ansieae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[748,799,665,686]" pageId="2" pageNumber="485" rank="species">sp. n.</taxonomicNameLabel>
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, (1) male habitus, lateral; (2) eye fielD, lateral; (3) female habitus, Dorsal; (4) female prosoma, Ventral; (5) calamistrum, retrolateral.
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</paragraph>
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</caption>
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<paragraph blockId="2.[144,1104,751,1677]" pageId="2" pageNumber="485">
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Body length 3.15. Habitus as in
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<figureCitation box="[518,587,751,778]" captionStart="Figs 1–5" captionStartId="2.[144,183,664,686]" captionTargetBox="[161,1091,204,645]" captionTargetId="figure-412@2.[157,1091,204,645]" captionTargetPageId="2" captionText="Figs 1–5. Habitus and somatic characters of Andoharano ansieae sp. n., (1) male habitus, lateral; (2) eye fielD, lateral; (3) female habitus, Dorsal; (4) female prosoma, Ventral; (5) calamistrum, retrolateral." figureDoi="http://doi.org/10.5281/zenodo.7914312" httpUri="https://zenodo.org/record/7914312/files/figure.png" pageId="2" pageNumber="485">Fig. 1</figureCitation>
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. Colour in alcohol: prosoma and legs almost uniformly clay-yellow with weakly developed darker brownish spots and fasciae; eye tubercle with blackish spot acutely extended to clypeus; opistosoma yellowish-brown with very slightly darker and diffuse dorsal pattern consisting of few almost indistinct transverse fasciae. Carapace 1.27 long, 1.14 wide. Eye tubercle as shown in
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<figureCitation box="[1029,1100,877,904]" captionStart="Figs 1–5" captionStartId="2.[144,183,664,686]" captionTargetBox="[161,1091,204,645]" captionTargetId="figure-412@2.[157,1091,204,645]" captionTargetPageId="2" captionText="Figs 1–5. Habitus and somatic characters of Andoharano ansieae sp. n., (1) male habitus, lateral; (2) eye fielD, lateral; (3) female habitus, Dorsal; (4) female prosoma, Ventral; (5) calamistrum, retrolateral." figureDoi="http://doi.org/10.5281/zenodo.7914312" httpUri="https://zenodo.org/record/7914312/files/figure.png" pageId="2" pageNumber="485">Fig. 2</figureCitation>
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. Eye sizes and interdistances: AME 0.09, ALE 0.16, PLE 0.14, PME 0.10, AME–AME 0.02. Leg measurements as shown in Table 1. Leg spines absent. Paired claws with 5–8 fine Dense teeth. UnpaireD claw VerY small, curVeD. Palp as in
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<figureCitation box="[910,1027,972,998]" captionStart="Figs 6–13" captionStartId="3.[182,221,1540,1562]" captionTargetBox="[193,1130,652,1515]" captionTargetId="figure-262@3.[193,1130,652,1515]" captionTargetPageId="3" captionText="Figs 6–13. Copulatory organs of Andoharano ansieae sp. n. (6–12) and A. grandidieri (13). (6, 7) male palp, prolateral; (8) male palp, retrolateral; (9) terminal part of male palp, prolateral; (10) terminal part of male palp and schematic course of spermophor, retrolateral; (11) endogyne, dorsal; (12) receptacle, dorsal; (13) terminal part of male palp, prolateral view and from above (after Lehtinen 1967). Abbreviations: Bc coil directed backward; Fl forelock." figureDoi="http://doi.org/10.5281/zenodo.7914316" httpUri="https://zenodo.org/record/7914316/files/figure.png" pageId="2" pageNumber="485">Figs 6–10</figureCitation>
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; short, length equal to carapace length; all segments covered with long hairs; hairs on patella, tibia and cymbium longer than corresponding segments on legs; femur longer than patella+tibia, with long suberected ventral hairs, and decumbent dorsal hairs, longest hairs about half of the segment length; patella twice longer than wide, with long and dense dorsal hairs; tibia subconical, distal edge wider than proximal, and wider than femur width, dorsal hairs sparse in basal half and dense in distal part, some of hairs almost twice longer than tibia; cymbium short, its height shorter than diameter of bulb, apical part with dense brush of hairs forming kind of forelock (
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<emphasis box="[839,864,1224,1250]" italics="true" pageId="2" pageNumber="485">Fl</emphasis>
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) (
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<figureCitation box="[887,999,1224,1251]" captionStart="Figs 6–13" captionStartId="3.[182,221,1540,1562]" captionTargetBox="[193,1130,652,1515]" captionTargetId="figure-262@3.[193,1130,652,1515]" captionTargetPageId="3" captionText="Figs 6–13. Copulatory organs of Andoharano ansieae sp. n. (6–12) and A. grandidieri (13). (6, 7) male palp, prolateral; (8) male palp, retrolateral; (9) terminal part of male palp, prolateral; (10) terminal part of male palp and schematic course of spermophor, retrolateral; (11) endogyne, dorsal; (12) receptacle, dorsal; (13) terminal part of male palp, prolateral view and from above (after Lehtinen 1967). Abbreviations: Bc coil directed backward; Fl forelock." figureDoi="http://doi.org/10.5281/zenodo.7914316" httpUri="https://zenodo.org/record/7914316/files/figure.png" pageId="2" pageNumber="485">Figs 9, 10</figureCitation>
|
||
) hanging over the bulb, hairs 3× longer than cymbium; bulb with droplet-shaped tegular part and narrow embolic part; embolic part bent down, with small tooth like dorsal outgrowth in basal third; spermophor (sperm duct) with 1.5 coils, distal coil of unusual shape, directed backward (
|
||
<emphasis box="[366,396,1351,1376]" italics="true" pageId="2" pageNumber="485">Bc</emphasis>
|
||
) (
|
||
<figureCitation box="[420,535,1350,1377]" captionStart="Figs 6–13" captionStartId="3.[182,221,1540,1562]" captionTargetBox="[193,1130,652,1515]" captionTargetId="figure-262@3.[193,1130,652,1515]" captionTargetPageId="3" captionText="Figs 6–13. Copulatory organs of Andoharano ansieae sp. n. (6–12) and A. grandidieri (13). (6, 7) male palp, prolateral; (8) male palp, retrolateral; (9) terminal part of male palp, prolateral; (10) terminal part of male palp and schematic course of spermophor, retrolateral; (11) endogyne, dorsal; (12) receptacle, dorsal; (13) terminal part of male palp, prolateral view and from above (after Lehtinen 1967). Abbreviations: Bc coil directed backward; Fl forelock." figureDoi="http://doi.org/10.5281/zenodo.7914316" httpUri="https://zenodo.org/record/7914316/files/figure.png" pageId="2" pageNumber="485">Figs 9, 10</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="2.[144,1104,751,1677]" box="[144,357,1390,1416]" pageId="2" pageNumber="485">
|
||
<emphasis box="[144,228,1390,1416]" italics="true" pageId="2" pageNumber="485">Female</emphasis>
|
||
(
|
||
<typeStatus box="[242,345,1390,1416]" pageId="2" pageNumber="485">paratype</typeStatus>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="2.[144,1104,751,1677]" lastBlockId="4.[144,1104,680,1070]" lastPageId="4" lastPageNumber="487" pageId="2" pageNumber="485">
|
||
Body length 4.25. Habitus as in
|
||
<figureCitation box="[493,558,1429,1456]" captionStart="Figs 1–5" captionStartId="2.[144,183,664,686]" captionTargetBox="[161,1091,204,645]" captionTargetId="figure-412@2.[157,1091,204,645]" captionTargetPageId="2" captionText="Figs 1–5. Habitus and somatic characters of Andoharano ansieae sp. n., (1) male habitus, lateral; (2) eye fielD, lateral; (3) female habitus, Dorsal; (4) female prosoma, Ventral; (5) calamistrum, retrolateral." figureDoi="http://doi.org/10.5281/zenodo.7914312" httpUri="https://zenodo.org/record/7914312/files/figure.png" pageId="2" pageNumber="485">Fig. 3</figureCitation>
|
||
. Colour in alcohol and most somatic characters as in male; venter of prosoma uniformly yellowish (
|
||
<figureCitation box="[671,737,1461,1487]" captionStart="Figs 1–5" captionStartId="2.[144,183,664,686]" captionTargetBox="[161,1091,204,645]" captionTargetId="figure-412@2.[157,1091,204,645]" captionTargetPageId="2" captionText="Figs 1–5. Habitus and somatic characters of Andoharano ansieae sp. n., (1) male habitus, lateral; (2) eye fielD, lateral; (3) female habitus, Dorsal; (4) female prosoma, Ventral; (5) calamistrum, retrolateral." figureDoi="http://doi.org/10.5281/zenodo.7914312" httpUri="https://zenodo.org/record/7914312/files/figure.png" pageId="2" pageNumber="485">Fig. 4</figureCitation>
|
||
); dorsum of abdomen with poorly distinct pattern composed of transverse bands. Carapace 1.60 long, 1.37 wide. Eye sizes and interdistances: AME 0.09, ALE 0.16, PLE 0.13, PME 0.09, AME–AME 0.02. Leg measurements as shown in Table 1 (in parentheses). Calamistrum includes three sessile rows of narrow, weak, moderately long lamellose setae (
|
||
<figureCitation box="[779,853,1587,1614]" captionStart="Figs 1–5" captionStartId="2.[144,183,664,686]" captionTargetBox="[161,1091,204,645]" captionTargetId="figure-412@2.[157,1091,204,645]" captionTargetPageId="2" captionText="Figs 1–5. Habitus and somatic characters of Andoharano ansieae sp. n., (1) male habitus, lateral; (2) eye fielD, lateral; (3) female habitus, Dorsal; (4) female prosoma, Ventral; (5) calamistrum, retrolateral." figureDoi="http://doi.org/10.5281/zenodo.7914312" httpUri="https://zenodo.org/record/7914312/files/figure.png" pageId="2" pageNumber="485">Figs 5</figureCitation>
|
||
,
|
||
<figureCitation box="[865,895,1587,1613]" captionStart="Figs 14, 15" captionStartId="4.[143,182,592,614]" captionTargetBox="[151,1095,205,574]" captionTargetId="figure-384@4.[151,1096,204,575]" captionTargetPageId="4" captionText="Figs 14, 15. Andoharano ansieae sp. n., female somatic characters.(14) junction of tibia and metatarsus IV showing calamistrum, retrolateral; (15) cribellum and spinnerets, ventral." figureDoi="http://doi.org/10.5281/zenodo.7914318" httpUri="https://zenodo.org/record/7914318/files/figure.png" pageId="2" pageNumber="485">14</figureCitation>
|
||
), occupying about 1/5 of metatarsus IV length. Palpal claw large, with 7–8 long dense teeth. Endogyne as in
|
||
<figureCitation box="[204,333,1650,1676]" captionStart="Figs 6–13" captionStartId="3.[182,221,1540,1562]" captionTargetBox="[193,1130,652,1515]" captionTargetId="figure-262@3.[193,1130,652,1515]" captionTargetPageId="3" captionText="Figs 6–13. Copulatory organs of Andoharano ansieae sp. n. (6–12) and A. grandidieri (13). (6, 7) male palp, prolateral; (8) male palp, retrolateral; (9) terminal part of male palp, prolateral; (10) terminal part of male palp and schematic course of spermophor, retrolateral; (11) endogyne, dorsal; (12) receptacle, dorsal; (13) terminal part of male palp, prolateral view and from above (after Lehtinen 1967). Abbreviations: Bc coil directed backward; Fl forelock." figureDoi="http://doi.org/10.5281/zenodo.7914316" httpUri="https://zenodo.org/record/7914316/files/figure.png" pageId="2" pageNumber="485">Figs 11, 12</figureCitation>
|
||
; edge of epigastrum with long and dense hairs (
|
||
<figureCitation box="[876,958,1650,1676]" captionStart="Figs 6–13" captionStartId="3.[182,221,1540,1562]" captionTargetBox="[193,1130,652,1515]" captionTargetId="figure-262@3.[193,1130,652,1515]" captionTargetPageId="3" captionText="Figs 6–13. Copulatory organs of Andoharano ansieae sp. n. (6–12) and A. grandidieri (13). (6, 7) male palp, prolateral; (8) male palp, retrolateral; (9) terminal part of male palp, prolateral; (10) terminal part of male palp and schematic course of spermophor, retrolateral; (11) endogyne, dorsal; (12) receptacle, dorsal; (13) terminal part of male palp, prolateral view and from above (after Lehtinen 1967). Abbreviations: Bc coil directed backward; Fl forelock." figureDoi="http://doi.org/10.5281/zenodo.7914316" httpUri="https://zenodo.org/record/7914316/files/figure.png" pageId="2" pageNumber="485">Fig. 11</figureCitation>
|
||
); one pair of round receptacles, spaced
|
||
<emphasis box="[435,461,680,705]" italics="true" pageId="4" pageNumber="487">ca</emphasis>
|
||
. 3 diameters from each other, receptacles without distinct stems and porous glands, seemingly connected by transverse arch. Spinnerets as in
|
||
<figureCitation captionStart="Figs 14, 15" captionStartId="4.[143,182,592,614]" captionTargetBox="[151,1095,205,574]" captionTargetId="figure-384@4.[151,1096,204,575]" captionTargetPageId="4" captionText="Figs 14, 15. Andoharano ansieae sp. n., female somatic characters.(14) junction of tibia and metatarsus IV showing calamistrum, retrolateral; (15) cribellum and spinnerets, ventral." figureDoi="http://doi.org/10.5281/zenodo.7914318" httpUri="https://zenodo.org/record/7914318/files/figure.png" pageId="4" pageNumber="487">Fig. 15</figureCitation>
|
||
. Spinneret group separated from posterior tip of abdomen by about 1⁄3 of abdomen length. Cribellum large, bipartite, trapezoidal. ALS and PLS moderately short; PMS relatiVelY large, anteriorlY with short anD thick moDifieD setae.
|
||
</paragraph>
|
||
<paragraph blockId="3.[271,1053,204,257]" pageId="3" pageNumber="486">
|
||
<tableNote pageId="3" pageNumber="486" targetBox="[206,1131,284,546]" targetPageId="3">TABLE 1 Length of palp and leg segments: holotype male and paratype female (in parentheses).</tableNote>
|
||
</paragraph>
|
||
<paragraph pageId="3" pageNumber="486">
|
||
<table box="[206,1131,284,546]" gridcols="7" gridrows="6" pageId="3" pageNumber="486">
|
||
<tr box="[206,1131,284,306]" gridrow="0" pageId="3" pageNumber="486" rowspan-0="1">
|
||
<th box="[293,394,284,306]" gridcol="1" gridrow="0" pageId="3" pageNumber="486">Femur</th>
|
||
<th box="[439,539,284,306]" gridcol="2" gridrow="0" pageId="3" pageNumber="486">Patella</th>
|
||
<th box="[584,685,284,306]" gridcol="3" gridrow="0" pageId="3" pageNumber="486">Tibia</th>
|
||
<th box="[729,830,284,306]" gridcol="4" gridrow="0" pageId="3" pageNumber="486">Metatarsus</th>
|
||
<th box="[875,975,284,306]" gridcol="5" gridrow="0" pageId="3" pageNumber="486">Tarsus</th>
|
||
<th box="[1009,1131,284,306]" gridcol="6" gridrow="0" pageId="3" pageNumber="486">Total</th>
|
||
</tr>
|
||
<tr box="[206,1131,332,354]" gridrow="1" pageId="3" pageNumber="486">
|
||
<th box="[206,246,332,354]" gridcol="0" gridrow="1" pageId="3" pageNumber="486">Palp</th>
|
||
<td box="[293,394,332,354]" gridcol="1" gridrow="1" pageId="3" pageNumber="486">0.83 (1.23)</td>
|
||
<td box="[439,539,332,354]" gridcol="2" gridrow="1" pageId="3" pageNumber="486">0.32 (0.47)</td>
|
||
<td box="[584,685,332,354]" gridcol="3" gridrow="1" pageId="3" pageNumber="486">0.39 (0.80)</td>
|
||
<td box="[729,830,332,354]" gridcol="4" gridrow="1" pageId="3" pageNumber="486">-</td>
|
||
<td box="[875,975,332,354]" gridcol="5" gridrow="1" pageId="3" pageNumber="486">0.11 (0.91)</td>
|
||
<td box="[1009,1131,332,354]" gridcol="6" gridrow="1" pageId="3" pageNumber="486">1.65 (3.41)</td>
|
||
</tr>
|
||
<tr box="[206,1131,380,402]" gridrow="2" pageId="3" pageNumber="486">
|
||
<th box="[206,246,380,402]" gridcol="0" gridrow="2" pageId="3" pageNumber="486">I</th>
|
||
<td box="[293,394,380,402]" gridcol="1" gridrow="2" pageId="3" pageNumber="486">2.64 (2.42)</td>
|
||
<td box="[439,539,380,402]" gridcol="2" gridrow="2" pageId="3" pageNumber="486">0.55 (0.54)</td>
|
||
<td box="[584,685,380,402]" gridcol="3" gridrow="2" pageId="3" pageNumber="486">3.30 (3.43)</td>
|
||
<td box="[729,830,380,402]" gridcol="4" gridrow="2" pageId="3" pageNumber="486">2.88 (2.88)</td>
|
||
<td box="[875,975,380,402]" gridcol="5" gridrow="2" pageId="3" pageNumber="486">1.19 (1.22)</td>
|
||
<td box="[1009,1131,380,402]" gridcol="6" gridrow="2" pageId="3" pageNumber="486">10.56 (10.49)</td>
|
||
</tr>
|
||
<tr box="[206,1131,428,450]" gridrow="3" pageId="3" pageNumber="486">
|
||
<th box="[206,246,428,450]" gridcol="0" gridrow="3" pageId="3" pageNumber="486">II</th>
|
||
<td box="[293,394,428,450]" gridcol="1" gridrow="3" pageId="3" pageNumber="486">1.94 (1.78)</td>
|
||
<td box="[439,539,428,450]" gridcol="2" gridrow="3" pageId="3" pageNumber="486">0.52 (0.53)</td>
|
||
<td box="[584,685,428,450]" gridcol="3" gridrow="3" pageId="3" pageNumber="486">1.93 (1.59)</td>
|
||
<td box="[729,830,428,450]" gridcol="4" gridrow="3" pageId="3" pageNumber="486">1.62 (1.54)</td>
|
||
<td box="[875,975,428,450]" gridcol="5" gridrow="3" pageId="3" pageNumber="486">0.78 (0.90)</td>
|
||
<td box="[1009,1131,428,450]" gridcol="6" gridrow="3" pageId="3" pageNumber="486">6.79 (6.34)</td>
|
||
</tr>
|
||
<tr box="[206,1131,476,498]" gridrow="4" pageId="3" pageNumber="486">
|
||
<th box="[206,246,476,498]" gridcol="0" gridrow="4" pageId="3" pageNumber="486">III</th>
|
||
<td box="[293,394,476,498]" gridcol="1" gridrow="4" pageId="3" pageNumber="486">1.62 (1.52)</td>
|
||
<td box="[439,539,476,498]" gridcol="2" gridrow="4" pageId="3" pageNumber="486">0.45 (0.51)</td>
|
||
<td box="[584,685,476,498]" gridcol="3" gridrow="4" pageId="3" pageNumber="486">1.40 (1.32)</td>
|
||
<td box="[729,830,476,498]" gridcol="4" gridrow="4" pageId="3" pageNumber="486">1.14 (1.39)</td>
|
||
<td box="[875,975,476,498]" gridcol="5" gridrow="4" pageId="3" pageNumber="486">0.52 (0.78)</td>
|
||
<td box="[1009,1131,476,498]" gridcol="6" gridrow="4" pageId="3" pageNumber="486">5.13 (5.52)</td>
|
||
</tr>
|
||
<tr box="[206,1131,524,546]" gridrow="5" pageId="3" pageNumber="486">
|
||
<th box="[206,246,524,546]" gridcol="0" gridrow="5" pageId="3" pageNumber="486">IV</th>
|
||
<td box="[293,394,524,546]" gridcol="1" gridrow="5" pageId="3" pageNumber="486">1.88 (1.93)</td>
|
||
<td box="[439,539,524,546]" gridcol="2" gridrow="5" pageId="3" pageNumber="486">0.46 (0.59)</td>
|
||
<td box="[584,685,524,546]" gridcol="3" gridrow="5" pageId="3" pageNumber="486">1.85 (1.77)</td>
|
||
<td box="[729,830,524,546]" gridcol="4" gridrow="5" pageId="3" pageNumber="486">2.04 (1.71)</td>
|
||
<td box="[875,975,524,546]" gridcol="5" gridrow="5" pageId="3" pageNumber="486">0.60 (0.89)</td>
|
||
<td box="[1009,1131,524,546]" gridcol="6" gridrow="5" pageId="3" pageNumber="486">6.83 (6.98)</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7914316" ID-Zenodo-Dep="7914316" httpUri="https://zenodo.org/record/7914316/files/figure.png" pageId="3" pageNumber="486" startId="3.[182,221,1540,1562]" targetBox="[193,1130,652,1515]" targetPageId="3" targetType="figure">
|
||
<paragraph blockId="3.[182,1142,1540,1658]" pageId="3" pageNumber="486">
|
||
Figs 6–13. Copulatory organs of
|
||
<taxonomicName authority="Zonstein & Marusik, 2015" authorityName="Zonstein & Marusik" authorityYear="2015" box="[489,679,1540,1562]" class="Arachnida" family="Filistatidae" genus="Andoharano" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="486" phylum="Arthropoda" rank="species" species="ansieae" status="sp. n.">
|
||
<emphasis box="[489,679,1540,1562]" italics="true" pageId="3" pageNumber="486">Andoharano ansieae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[686,737,1541,1562]" pageId="3" pageNumber="486" rank="species">sp. n.</taxonomicNameLabel>
|
||
(6–12) and
|
||
<taxonomicName baseAuthorityName="Simon" baseAuthorityYear="1901" box="[851,981,1540,1562]" class="Arachnida" family="Filistatidae" genus="Andoharano" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="486" phylum="Arthropoda" rank="species" species="grandidieri">
|
||
<emphasis box="[851,865,1541,1562]" italics="true" pageId="3" pageNumber="486">A</emphasis>
|
||
.
|
||
<emphasis box="[878,981,1540,1561]" italics="true" pageId="3" pageNumber="486">grandidieri</emphasis>
|
||
</taxonomicName>
|
||
(13). (6, 7) male palp, prolateral; (8) male palp, retrolateral; (9) terminal part of male palp, prolateral; (10) terminal part of male palp and schematic course of spermophor, retrolateral; (11) endogyne, dorsal; (12) receptacle, dorsal; (13) terminal part of male palp, prolateral view and from above (after
|
||
<bibRefCitation author="LEHTINEN, P. T." pageId="3" pageNumber="486" pagination="199 - 468" refId="ref3209" refString="LEHTINEN, P. T. 1967. Classification of the cribellate spiDers anD some allieD families, with notes on the evolution of the suborder Araneomorpha. Annales Zoologici Fennici 4: 199 - 468." type="journal article" year="1967">Lehtinen 1967</bibRefCitation>
|
||
). Abbreviations:
|
||
<emphasis box="[449,473,1637,1658]" italics="true" pageId="3" pageNumber="486">Bc</emphasis>
|
||
coil directed backward;
|
||
<emphasis box="[697,717,1636,1657]" italics="true" pageId="3" pageNumber="486">Fl</emphasis>
|
||
forelock.
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7914318" ID-Zenodo-Dep="7914318" httpUri="https://zenodo.org/record/7914318/files/figure.png" pageId="4" pageNumber="487" startId="4.[143,182,592,614]" targetBox="[151,1095,205,574]" targetPageId="4" targetType="figure">
|
||
<paragraph blockId="4.[143,1104,592,638]" pageId="4" pageNumber="487">
|
||
Figs 14, 15.
|
||
<taxonomicName authority="Zonstein & Marusik, 2015" authorityName="Zonstein & Marusik" authorityYear="2015" box="[254,441,592,613]" class="Arachnida" family="Filistatidae" genus="Andoharano" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="487" phylum="Arthropoda" rank="species" species="ansieae" status="sp. n.">
|
||
<emphasis box="[254,441,592,613]" italics="true" pageId="4" pageNumber="487">Andoharano ansieae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[446,496,592,613]" pageId="4" pageNumber="487" rank="species">sp. n.</taxonomicNameLabel>
|
||
, female somatic characters. (14) junction of tibia and metatarsus IV showing calamistrum, retrolateral; (15) cribellum and spinnerets, ventral.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="4.[144,1104,680,1070]" pageId="4" pageNumber="487">
|
||
Note: The receptacles are very small and pale, almost invisible, and can only be seen after colouring with Chlorazol Black; female specimens can be easily mistaken for juveniles. Variation: Within the
|
||
<typeStatus box="[381,429,923,949]" pageId="4" pageNumber="487">type</typeStatus>
|
||
series the body length varies: in males 3.05–3.15; in females 4.10–4.25.
|
||
</paragraph>
|
||
<paragraph blockId="4.[144,1104,680,1070]" pageId="4" pageNumber="487">
|
||
<materialsCitation collectingDate="1979-05" collectionCode="MRAC" country="Namibia" latitude="-17.95" location="OkaVango" longLatPrecision="1277" longitude="19.716667" pageId="4" pageNumber="487" specimenCode="MRAC 152150" specimenCount="1" specimenCount-male="1" stateProvince="Kavango West" typeStatus="holotype">
|
||
<typeStatus box="[144,228,991,1013]" pageId="4" pageNumber="487">HolotYpe</typeStatus>
|
||
<specimenCount box="[231,251,991,1012]" pageId="4" pageNumber="487" type="male">♂</specimenCount>
|
||
:
|
||
<collectingCountry box="[259,361,991,1012]" name="Namibia" pageId="4" pageNumber="487">NAMIBIA</collectingCountry>
|
||
: RunDu-KaVango,
|
||
<collectingRegion box="[526,621,991,1013]" country="Namibia" name="Kavango West" pageId="4" pageNumber="487">OkaVango</collectingRegion>
|
||
,
|
||
<geoCoordinate box="[628,699,991,1013]" degrees="17" direction="south" minutes="57" orientation="latitude" pageId="4" pageNumber="487" precision="925" value="-17.95">17°57'S</geoCoordinate>
|
||
<geoCoordinate box="[703,777,991,1012]" degrees="19" direction="east" minutes="43" orientation="longitude" pageId="4" pageNumber="487" precision="925" value="19.716667">19°43'E</geoCoordinate>
|
||
,
|
||
<date box="[784,847,991,1012]" pageId="4" pageNumber="487" value="1979-05">
|
||
<collectingDate box="[784,847,991,1012]" pageId="4" pageNumber="487" value="1979-05">V.1979</collectingDate>
|
||
</date>
|
||
, leg. M.E. BaDDeleY (
|
||
<specimenCode collectionCode="MRAC" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="4" pageNumber="487" type="Museum">MRAC 152150</specimenCode>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph blockId="4.[144,1104,680,1070]" box="[144,877,1048,1070]" pageId="4" pageNumber="487">
|
||
<typeStatus box="[144,236,1049,1070]" pageId="4" pageNumber="487">ParatYpes</typeStatus>
|
||
:
|
||
<specimenCount box="[244,272,1048,1069]" pageId="4" pageNumber="487" type="male">1♂</specimenCount>
|
||
<specimenCount box="[278,306,1048,1069]" pageId="4" pageNumber="487" type="female">2♀</specimenCount>
|
||
collecteD together with the
|
||
<typeStatus box="[558,637,1048,1070]" pageId="4" pageNumber="487">holotYpe</typeStatus>
|
||
anD kept in the same Vial.
|
||
</paragraph>
|
||
<paragraph blockId="4.[144,1103,1086,1217]" box="[144,1059,1086,1112]" pageId="4" pageNumber="487">
|
||
Distribution: The species is currently known only from the
|
||
<typeStatus box="[808,856,1086,1112]" pageId="4" pageNumber="487">type</typeStatus>
|
||
locality (
|
||
<figureCitation box="[963,1046,1086,1112]" captionStart="Fig" captionStartId="5.[181,214,657,679]" captionTargetBox="[340,983,204,641]" captionTargetId="figure-354@5.[340,983,204,641]" captionTargetPageId="5" captionText="Fig.16.The known distributionof the genus Andoharano Lehtinen,1967 in mainlandAfrica and Madagascar." figureDoi="http://doi.org/10.5281/zenodo.7914320" httpUri="https://zenodo.org/record/7914320/files/figure.png" pageId="4" pageNumber="487">Fig. 16</figureCitation>
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).
|
||
</paragraph>
|
||
<paragraph blockId="4.[144,1103,1086,1217]" pageId="4" pageNumber="487">
|
||
Ecology: Unrecorded. Since all other congeners have been found inhabiting caves (see
|
||
<bibRefCitation author="LEGENDRE, R." box="[144,316,1159,1185]" pageId="4" pageNumber="487" pagination="645 - 650" refId="ref3168" refString="LEGENDRE, R. 1971. Deux nouvelles araignees Filistatidae cavernicoles de Madagascar: Andoharano milloti n. sp. et Andoharano monodi n. sp. Bulletin du Museum National d'Histoire Naturelle de Paris 13: 645 - 650." type="journal article" year="1971">Legendre 1971</bibRefCitation>
|
||
), we believe that the spiders might be found in small niches, cavities, and cracks in rock cliffs and rocky outcrops.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |