212 lines
29 KiB
XML
212 lines
29 KiB
XML
<document id="88F38AF4E79C02A3AC0EEE9829B07971" ID="10.11646/zootaxa.4196.1.1" ID-DOI="10.11646/zootaxa.4196.1.1" ID-GBIF-Dataset="b7651a7b-df24-4347-9585-0633b88dba51" ID-ISSN="1175-5326" ID-Zenodo-Dep="167648" ID-ZooBank="026D88BD-86C0-4811-814A-CB5CACB3104F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1479712233041" checkinUser="plazi" docAuthor="Sokolov, Igor M." docDate="2016" docId="8D3787848901FFD621CA3D40FF46FB5A" docLanguage="en" docName="zootaxa.4196.1.1.pdf" docOrigin="Zootaxa 4196 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Zeanillus montivagus Sokolov, 2016, sp.n." docType="treatment" docVersion="8" lastPageNumber="30" masterDocId="710EFFFC891DFFCB215D3B54FFBCFFFA" masterDocTitle="A taxonomic review of the anilline genus Zeanillus Jeannel (Coleoptera: Carabidae: Bembidiini) of New Zealand, with descriptions of seven new species, re-classification of the species, and notes on their biogeography and evolution" masterLastPageNumber="37" masterPageNumber="1" pageNumber="29" updateTime="1698393955637" updateUser="plazi">
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<mods:titleInfo id="317766176EA2F663E7A59EFB349E34C9">
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<mods:title id="2EB4BD8BC3166BCE84A707A442145D37">A taxonomic review of the anilline genus Zeanillus Jeannel (Coleoptera: Carabidae: Bembidiini) of New Zealand, with descriptions of seven new species, re-classification of the species, and notes on their biogeography and evolution</mods:title>
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<mods:name id="E2FBF0466BB490F5B3108C0AD73AD3F3" type="personal">
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<mods:roleTerm id="470E85C2C8BAB7224D81EDE0B3368A69">Author</mods:roleTerm>
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<mods:namePart id="6602909B50BBD969C472DC6F3C1D8B67">Sokolov, Igor M.</mods:namePart>
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<mods:typeOfResource id="E2F40FAE7058A5B35F9673068A5E9167">text</mods:typeOfResource>
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<mods:titleInfo id="678E75396099A7D54A5BF8216F962686">
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<mods:title id="DAC07036E77382F69C9B130C1A40F3D6">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="185ED83AFCCD6EA36A98BF5D764D99AE">
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<mods:date id="08975FCBC2E36FC34CAB4A6208BD2B3A">2016</mods:date>
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<mods:detail id="369F8660D3A09AD00D6FA0A365660B79" type="volume">
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<mods:number id="02B6464E59EE957C00AC2028659E9B16">4196</mods:number>
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</mods:detail>
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<mods:detail id="9D6AC7DE4E5849781358EB8C3291BCF0" type="issue">
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<mods:number id="B41C7AE3C0833422C247846FDB7D5749">1</mods:number>
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<mods:start id="75DB33BEF8CB3BE846B5116CB317B211">1</mods:start>
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<mods:end id="5AEF78EB7DC13B9368FA8345EF36A57C">37</mods:end>
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<mods:classification id="1ED7B5FABD5058416C6DF739F5A773A3">journal article</mods:classification>
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<mods:identifier id="D9CFB2C4186B5A93218DA42424A41618" type="DOI">10.11646/zootaxa.4196.1.1</mods:identifier>
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<mods:identifier id="BA146A7565DC118EB4A198D421376EAC" type="GBIF-Dataset">b7651a7b-df24-4347-9585-0633b88dba51</mods:identifier>
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<mods:identifier id="D167D07EB1E472C4691E870B05F262AB" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="B09A20AEFC06F66A558AB371D3E472B5" type="Zenodo-Dep">167648</mods:identifier>
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<mods:identifier id="C8621A169DC8636B10BC734A730C016F" type="ZooBank">026D88BD-86C0-4811-814A-CB5CACB3104F</mods:identifier>
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<treatment id="8D3787848901FFD621CA3D40FF46FB5A" ID-DOI="http://doi.org/10.5281/zenodo.6089694" ID-GBIF-Taxon="124814323" ID-Zenodo-Dep="6089694" LSID="urn:lsid:plazi:treatment:8D3787848901FFD621CA3D40FF46FB5A" httpUri="http://treatment.plazi.org/id/8D3787848901FFD621CA3D40FF46FB5A" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29">
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<subSubSection id="4D8465198901FFD721CA3D40FDD1F9B5" pageId="28" pageNumber="29" type="nomenclature">
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<paragraph id="052136928901FFD721CA3D40FE59F9D4" blockId="28.[151,621,1556,1615]" box="[151,485,1556,1582]" pageId="28" pageNumber="29">
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<heading id="5E6981FE8901FFD721CA3D40FE59F9D4" bold="true" box="[151,485,1556,1582]" fontSize="11" level="1" pageId="28" pageNumber="29" reason="1">
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<emphasis id="37EAEA808901FFD721CA3D40FE59F9D4" bold="true" box="[151,485,1556,1582]" pageId="28" pageNumber="29">
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<emphasis id="37EAEA808901FFD721CA3D40FE1FF9D4" bold="true" box="[151,419,1556,1582]" italics="true" pageId="28" pageNumber="29">
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<taxonomicName id="C29E4D118901FFD721CA3D40FE21F9D4" ID-CoL="8MK2Q" box="[151,413,1556,1582]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="montivagus" status="sp. nov.">Zeanillus montivagus</taxonomicName>
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,
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</emphasis>
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<taxonomicNameLabel id="2CD957FB8901FFD720F63D40FE59F9D4" box="[427,485,1556,1582]" pageId="28" pageNumber="29" rank="species">sp.n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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||
</paragraph>
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<paragraph id="052136928901FFD721CA3D62FDD1F9B5" blockId="28.[151,621,1556,1615]" box="[151,621,1590,1615]" pageId="28" pageNumber="29">
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<figureCitation id="9DA52A178901FFD721CA3D62FF5FF9B4" box="[151,227,1590,1615]" captionStart="FIGURE 7" captionStartId="14.[151,250,1204,1226]" captionTargetBox="[151,1428,193,1181]" captionTargetId="figure@14.[151,1435,193,1182]" captionTargetPageId="14" captionText="FIGURE 7. Digital images of pterothorax and first abdominal ventrites, ventral aspect, of Zeanillus species. A— Z. phyllobius; B— Z. punctigerus; C— Z. nunni; D— Z. pallidus; E— Z. pellucidus; F— Z. brouni; G— Z. lescheni; H— Z. montivagus; I— Z. nanus. Legend: ipa—intercoxal process of abdominal ventrite 2; mes—metendosternite; mte—metanepisternum; mtvmetaventrite; v 1 – v 2 — abdominal ventrites 1 – 2. Scale bars = 0.2 mm." httpUri="https://zenodo.org/record/167655/files/figure.png" pageId="28" pageNumber="29">Figs. 7</figureCitation>
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H,
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<date id="712010528901FFD720583D63FE90F9B5" box="[261,300,1591,1615]" pageId="28" pageNumber="29" value="2011-01">11I</date>
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, 13M–P, 14G,
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<date id="712010528901FFD7208D3D62FE4BF9B5" box="[464,503,1590,1615]" pageId="28" pageNumber="29" value="2015-01">15I</date>
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, 16G, 18B
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</paragraph>
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</subSubSection>
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<subSubSection id="4D8465198901FFD721CA3D2AFB3EF8F9" pageId="28" pageNumber="29" type="materials_examined">
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<paragraph id="052136928901FFD721CA3D2AFE5BF925" blockId="28.[151,1436,1662,2011]" pageId="28" pageNumber="29">
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<materialsCitation id="B5F63CCF8901FFD721CA3D2AFB80F941" ID-GBIF-Occurrence="1341350594" collectingDate="1984-12-08" collectionCode="NZAC" collectorName="CO North Rough Ridge & B. I. P. Barratt & NZ Arthropod Collection" country="New Zealand" elevation="1000" location="North Rough Ridge" pageId="28" pageNumber="29" specimenCount="2" typeStatus="holotype">
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<emphasis id="37EAEA808901FFD721CA3D2AFEF5F96D" bold="true" box="[151,329,1662,1687]" pageId="28" pageNumber="29">
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<typeStatus id="DA2588308901FFD721CA3D2AFF6EF96D" box="[151,210,1662,1687]" pageId="28" pageNumber="29">Type</typeStatus>
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material.
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</emphasis>
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<typeStatus id="DA2588308901FFD720093D2BFE57F96D" box="[340,491,1663,1687]" pageId="28" pageNumber="29" type="holotype">HOLOTYPE</typeStatus>
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, male, in
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<collectionCode id="638FAE578901FFD7233E3D2BFD0EF96D" box="[611,690,1663,1687]" country="New Zealand" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34070" name="New Zealand Arthropod Collection" pageId="28" pageNumber="29">NZAC</collectionCode>
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, point-mounted, undissected, labeled: /
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<collectingCountry id="7D8976028901FFD725D93D2BFAA1F96D" box="[1156,1309,1662,1687]" name="New Zealand" pageId="28" pageNumber="29">New Zealand</collectingCountry>
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<collectorName id="A86B53448901FFD724743D2BFE8AF941" pageId="28" pageNumber="29">
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CO
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<location id="004160498901FFD724063D2AFE8AF941" LSID="urn:lsid:plazi:treatment:8D3787848901FFD621CA3D40FF46FB5A:004160498901FFD724063D2AFE8AF941" country="New Zealand" name="North Rough Ridge" pageId="28" pageNumber="29">North Rough Ridge</location>
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</collectorName>
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<collectingDate id="6164E9BA8901FFD720153DF7FE62F941" box="[328,478,1699,1723]" pageId="28" pageNumber="29" value="1984-12-08">
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<date id="712010528901FFD720153DF7FE62F941" box="[328,478,1699,1723]" pageId="28" pageNumber="29" value="1984-12-08">8 Dec 1984</date>
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</collectingDate>
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<elevation id="8EB3D1A18901FFD720AD3DF7FD83F941" box="[496,575,1699,1723]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="28" pageNumber="29" unit="m" value="1000.0">
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||
<quantity id="C2669B778901FFD720AD3DF7FD83F941" box="[496,575,1699,1723]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="28" pageNumber="29" unit="m" value="1000.0">1000m</quantity>
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</elevation>
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<collectorName id="A86B53448901FFD7230D3DF7FD67F940" box="[592,731,1699,1722]" pageId="28" pageNumber="29">B.I.P.Barratt</collectorName>
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/
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<collectorName id="A86B53448901FFD7225B3DF7FB80F941" box="[774,1084,1698,1723]" pageId="28" pageNumber="29">NZ Arthropod Collection</collectorName>
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</materialsCitation>
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,
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<materialsCitation id="B5F63CCF8901FFD7250C3DF7FE5BF925" ID-GBIF-Occurrence="1341350599" collectionCode="NZAC" country="New Zealand" location="AUCKLAND" pageId="28" pageNumber="29" specimenCount="1" stateProvince="AUCKLAND" typeStatus="holotype">
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<collectionCode id="638FAE578901FFD7250C3DF7FB1CF941" box="[1105,1184,1699,1723]" country="New Zealand" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34070" name="New Zealand Arthropod Collection" pageId="28" pageNumber="29">NZAC</collectionCode>
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, Private Bag 92170
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<collectingRegion id="C75AF8708901FFD721CA3D93FE88F925" box="[151,308,1735,1759]" country="New Zealand" name="Auckland" pageId="28" pageNumber="29">AUCKLAND</collectingRegion>
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<collectingCountry id="7D8976028901FFD720663D93FE6DF925" box="[315,465,1734,1759]" name="New Zealand" pageId="28" pageNumber="29">New Zealand</collectingCountry>
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/.
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||
</materialsCitation>
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||
</paragraph>
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||
<paragraph id="052136928901FFD721CA3DBFFB3EF8F9" blockId="28.[151,1436,1662,2011]" box="[151,1154,1770,1795]" pageId="28" pageNumber="29">PARATYPES (5 specimens, 3 dissected), 4 males and 1 female labeled same as holotype.</paragraph>
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||
</subSubSection>
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||
<subSubSection id="4D8465198901FFD7219A3C5AFC9FF895" pageId="28" pageNumber="29" type="description">
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||
<paragraph id="052136928901FFD7219A3C5AFC9FF895" blockId="28.[151,1436,1662,2011]" pageId="28" pageNumber="29">
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<emphasis id="37EAEA808901FFD7219A3C5AFE3BF8DD" bold="true" box="[199,391,1806,1831]" pageId="28" pageNumber="29">Specific epithet.</emphasis>
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The specific epithet is a Latinized adjective in the masculine form and is derived from the Latin adjective
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<taxonomicName id="C29E4D118901FFD720103C67FE72F8B0" box="[333,462,1843,1866]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="montivagus">
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<emphasis id="37EAEA808901FFD720103C67FE72F8B0" box="[333,462,1843,1866]" italics="true" pageId="28" pageNumber="29">montivagus</emphasis>
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</taxonomicName>
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meaning " wandering over the mountains ". The epithet refers to the mountainous landscape at the locality, where this species was collected.
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</paragraph>
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</subSubSection>
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<subSubSection id="4D8465198901FFD7219A3C2EFEFAF821" pageId="28" pageNumber="29" type="materials_examined">
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<paragraph id="052136928901FFD7219A3C2EFC5EF869" blockId="28.[151,1436,1662,2011]" box="[199,994,1914,1939]" pageId="28" pageNumber="29">
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<materialsCitation id="B5F63CCF8901FFD7219A3C2EFC62F869" ID-GBIF-Occurrence="1341350596" box="[199,990,1914,1939]" country="New Zealand" location="North Rough Ridge" municipality="South Island" pageId="28" pageNumber="29" specimenCount="1" stateProvince="Otago" typeStatus="holotype">
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<emphasis id="37EAEA808901FFD7219A3C2EFED9F869" bold="true" box="[199,357,1914,1939]" pageId="28" pageNumber="29">
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<typeStatus id="DA2588308901FFD7219A3C2EFEBEF869" box="[199,258,1914,1939]" pageId="28" pageNumber="29">Type</typeStatus>
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locality.
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</emphasis>
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<collectingCountry id="7D8976028901FFD720313C2FFDB9F869" box="[364,517,1914,1939]" name="New Zealand" pageId="28" pageNumber="29">New Zealand</collectingCountry>
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,
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<collectingMunicipality id="E545ACE88901FFD7234D3C2EFD23F869" box="[528,671,1914,1939]" pageId="28" pageNumber="29">South Island</collectingMunicipality>
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,
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<collectingRegion id="C75AF8708901FFD723F63C2FFD4EF869" box="[683,754,1915,1939]" country="New Zealand" name="Otago" pageId="28" pageNumber="29">Otago</collectingRegion>
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,
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<location id="004160498901FFD723A03C2EFC62F869" LSID="urn:lsid:plazi:treatment:8D3787848901FFD621CA3D40FF46FB5A:004160498901FFD723A03C2EFC62F869" box="[765,990,1914,1939]" country="New Zealand" municipality="South Island" name="North Rough Ridge" pageId="28" pageNumber="29" stateProvince="Otago">North Rough Ridge</location>
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="052136928901FFD7219A3CCAFEFAF821" blockId="28.[151,1436,1662,2011]" pageId="28" pageNumber="29">
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<emphasis id="37EAEA808901FFD7219A3CCAFEE1F84D" bold="true" box="[199,349,1950,1975]" pageId="28" pageNumber="29">Recognition.</emphasis>
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Adults of this species (
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<figureCitation id="9DA52A178901FFD7233D3CCAFD13F84D" box="[608,687,1950,1975]" captionStart="FIGURE 11" captionStartId="20.[151,250,1887,1909]" captionTargetBox="[198,1372,193,1855]" captionTargetId="figure@20.[198,1374,193,1857]" captionTargetPageId="20" captionText="FIGURE 11. Digital images of habitus of Zeanillus species, dorsal aspect. A— Z. phyllobius (NZ, Canterbury, Coopers Knob); B— Z. punctigerus (NZ, Canterbury, Sharplin Falls); C— Z. nunni (NZ, Otago, Highway 87 at Rocklands Road); D— Z. pallidus (NZ, Otago, Taieri Mouth); E— Z. pellucidus (NZ, Otago, Oamaru, Glen Warren Reserve); F— Z. brouni (NZ, Otago, Oamaru, Glen Warren Reserve); G— Z. carltoni (NZ, Otago, Mount Watkin); H— Z. lescheni (NZ, Otago, Waikaia Forest); I— Z. montivagus (NZ, Otago, North Rough Ridge): J— Z. nanus (NZ, Otago, Pisa Range). Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/167659/files/figure.png" pageId="28" pageNumber="29">
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Fig.
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<date id="712010528901FFD723C93CCBFD13F84D" box="[660,687,1951,1975]" pageId="28" pageNumber="29" value="2011-01">11</date>
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</figureCitation>
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I) are distinguished from other congeners by the characters listed in the species key.
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</paragraph>
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</subSubSection>
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<subSubSection id="4D8465198900FFD6219A3BC3FF46FB5A" pageId="29" pageNumber="30" type="description">
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||
<paragraph id="052136928900FFD6219A3BC3FB51FF4A" blockId="29.[151,1437,151,1184]" box="[199,1261,151,176]" pageId="29" pageNumber="30">
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<emphasis id="37EAEA808900FFD6219A3BC3FEE5FF4A" bold="true" box="[199,345,151,176]" pageId="29" pageNumber="30">Description.</emphasis>
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Size. Medium for genus (SBL range
|
||
<quantity id="C2669B778900FFD623A13BC3FC1DFF55" box="[764,929,151,176]" metricMagnitude="-3" metricUnit="m" metricValue="1.685" metricValueMax="1.73" metricValueMin="1.64" pageId="29" pageNumber="30" unit="mm" value="1.685" valueMax="1.73" valueMin="1.64">1.64–1.73 mm</quantity>
|
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, mean 1.69±
|
||
<quantity id="C2669B778900FFD625723BC3FB19FF55" box="[1071,1189,151,176]" metricMagnitude="-5" metricUnit="m" metricValue="4.3999999999999995" pageId="29" pageNumber="30" unit="mm" value="0.044">0.044 mm</quantity>
|
||
, n=6).
|
||
</paragraph>
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||
<paragraph id="052136928900FFD6219A3BE8FE17FEE6" blockId="29.[151,1437,151,1184]" pageId="29" pageNumber="30">Habitus. Body form slightly convex, ovoid, moderately elongate (WE/SBL 0.35±0.013), head of moderate width for genus compared to pronotum (WH/WPm 0.73±0.009), pronotum rather wide in comparison to elytra (WPm/WE 0.78±0.029).</paragraph>
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||
<paragraph id="052136928900FFD6219A3A73FCF7FEC5" blockId="29.[151,1437,151,1184]" box="[199,843,295,320]" pageId="29" pageNumber="30">Color. Body color rufotestaceous, appendages testaceous.</paragraph>
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<paragraph id="052136928900FFD6219A3A18FCEBFE56" blockId="29.[151,1437,151,1184]" pageId="29" pageNumber="30">Prothorax. Pronotum slightly long (LP/LE 0.38±0.010) and moderately elongate (WPm/LP 1.28±0.023), with lateral margins rectilinear constricted posteriorly (WPm/WPp 1.68±0.130). Width between anterior angles much greater than between posterior angles (WPa/WPp 1.31±0.074).</paragraph>
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<paragraph id="052136928900FFD6219A3AE3FECAFDE2" blockId="29.[151,1437,151,1184]" pageId="29" pageNumber="30">Elytra. Slightly depressed along suture, comparatively long (LE/SBL 0.55±0.012) and slightly wide (WE/LE 0.63±0.028). Lateral margins slightly divergent at basal third, subparallel at middle third and evenly rounded to apex in apical third.</paragraph>
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<paragraph id="052136928900FFD6219A3970FDA8FD36" blockId="29.[151,1437,151,1184]" pageId="29" pageNumber="30">
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Male genitalia. Median lobe (
|
||
<figureCitation id="9DA52A178900FFD6237D3970FDCFFDC6" box="[544,627,548,573]" captionStart="FIGURE 13" captionStartId="22.[151,250,1677,1699]" captionTargetBox="[162,1426,562,1641]" captionTargetId="figure@22.[151,1436,553,1656]" captionTargetPageId="22" captionText="FIGURE 13. Line drawings of aedeagus of Zeanillus species. Z. carltoni (NZ, Otago, Mount Watkin): A—median lobe, right lateral aspect, B—apical part of median lobe, dorsal aspect, C—left paramere, left lateral aspect, D—right paramere, right lateral aspect; Z. lescheni (NZ, Otago, Waikaia Forest): E—median lobe, right lateral aspect, F—apical part of median lobe, dorsal aspect, G—left paramere, left lateral aspect, H—right paramere, right lateral aspect; Z. nanus (NZ, Otago, Balclutha): I—median lobe, right lateral aspect, J—apical part of median lobe, dorsal aspect, K—left paramere, left lateral aspect, L—right paramere, right lateral aspect; Z. montivagus (NZ, Otago, North Rough Ridge): M—median lobe, right lateral aspect, Napical part of median lobe, dorsal aspect, O—left paramere, left lateral aspect, P—right paramere, right lateral aspect; Z. brouni (NZ, Otago, Oamaru, Glen Warren Reserve): Q—median lobe, right lateral aspect, R—apical part of median lobe, dorsal aspect, S—left paramere, left lateral aspect, T—right paramere, right lateral aspect. Legend: ds—dorsal sclerites; of—ostial field. Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/167661/files/figure.png" pageId="29" pageNumber="30">Fig. 13</figureCitation>
|
||
M) with broadly concave ventral margin and short semicircular apex. Apical orifice area with a pair of ostial fields. Sclerites of internal sac separate, uneven: dorsal is strongly sclerotized and bifurcated basally, ventral is weakly sclerotized, narrow and linear; sclerites shorter than the half of the distance between apical and basal orifices. Left paramere with apical constriction (
|
||
<figureCitation id="9DA52A178900FFD625D539DBFB60FD52" box="[1160,1244,655,680]" captionStart="FIGURE 13" captionStartId="22.[151,250,1677,1699]" captionTargetBox="[162,1426,562,1641]" captionTargetId="figure@22.[151,1436,553,1656]" captionTargetPageId="22" captionText="FIGURE 13. Line drawings of aedeagus of Zeanillus species. Z. carltoni (NZ, Otago, Mount Watkin): A—median lobe, right lateral aspect, B—apical part of median lobe, dorsal aspect, C—left paramere, left lateral aspect, D—right paramere, right lateral aspect; Z. lescheni (NZ, Otago, Waikaia Forest): E—median lobe, right lateral aspect, F—apical part of median lobe, dorsal aspect, G—left paramere, left lateral aspect, H—right paramere, right lateral aspect; Z. nanus (NZ, Otago, Balclutha): I—median lobe, right lateral aspect, J—apical part of median lobe, dorsal aspect, K—left paramere, left lateral aspect, L—right paramere, right lateral aspect; Z. montivagus (NZ, Otago, North Rough Ridge): M—median lobe, right lateral aspect, Napical part of median lobe, dorsal aspect, O—left paramere, left lateral aspect, P—right paramere, right lateral aspect; Z. brouni (NZ, Otago, Oamaru, Glen Warren Reserve): Q—median lobe, right lateral aspect, R—apical part of median lobe, dorsal aspect, S—left paramere, left lateral aspect, T—right paramere, right lateral aspect. Legend: ds—dorsal sclerites; of—ostial field. Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/167661/files/figure.png" pageId="29" pageNumber="30">Fig. 13</figureCitation>
|
||
O). Ring sclerite without visible handle (
|
||
<figureCitation id="9DA52A178900FFD620C239E0FE4CFD36" box="[415,496,692,717]" captionStart="FIGURE 14" captionStartId="23.[151,250,1033,1055]" captionTargetBox="[165,1404,198,1020]" captionTargetId="figure@23.[162,1414,193,1024]" captionTargetPageId="23" captionText="FIGURE 14. Line drawings of ring sclerite of Zeanillus species, male genitalia, dorsal aspect. A— Z. phyllobius (NZ, Canterbury, Ahuriri Scenic Reserve); B— Z. punctigerus (NZ, Canterbury, Orari Gorge); C— Z. nunni (NZ, Otago, Highway 87 at Rocklands Rd); D— Z. pallidus (NZ, Southland, Thomsons Bush); E— Z. pellucidus (NZ, Otago, Oamaru, Glen Warren Reserve); F— Z. nanus (NZ, Otago, Pisa Range); G— Z. montivagus (NZ, Otago, North Rough Ridge); H— Z. brouni (NZ, Otago, Oamaru, Glen Warren Reserve); I— Z. lescheni (NZ, Otago, Waikaia Forest); J— Z. carltoni (NZ, Otago, Mount Watkin). Legend: hd—handle of ring sclerite. Scale bar = 0.1 mm." httpUri="https://zenodo.org/record/167662/files/figure.png" pageId="29" pageNumber="30">Fig. 14</figureCitation>
|
||
G).
|
||
</paragraph>
|
||
<paragraph id="052136928900FFD6219A3983FBFAFCC2" blockId="29.[151,1437,151,1184]" pageId="29" pageNumber="30">
|
||
Female internal genitalia. Goncoxite 2 with short ensiform setae (
|
||
<figureCitation id="9DA52A178900FFD622EF3983FBBAFD0A" box="[946,1030,727,752]" captionStart="FIGURE 15" captionStartId="24.[151,250,1566,1588]" captionTargetBox="[159,1415,184,1535]" captionTargetId="figure@24.[150,1436,175,1545]" captionTargetPageId="24" captionText="FIGURE 15. Line drawings of right ovipositor sclerite, lateral aspect, of Zeanillus species. A— Z. phyllobius (NZ, Canterbury, Ahuriri Scenic Reserve); B— Z. punctigerus (NZ, Canterbury, Sharplin Falls); C— Z. nunni (NZ, Otago, Woodside Glen); D— Z. pallidus (NZ, Otago, Taieri Mouth); E— Z. pellucidus (NZ, Otago, Oamaru, Glen Warren Reserve); F— Z. brouni (NZ, Otago, Oamaru, Glen Warren Reserve); G— Z. carltoni (NZ, Otago, Mount Watkin); H— Z. lescheni (NZ, Otago, Waikaia Forest); I— Z. montivagus (NZ, Otago, North Rough Ridge); J— Z. nanus (NZ, Otago, Balclutha). Legend: es—ensiform seta; gc 1 — gonocoxite 1; gc 2 — gonocoxite 2; lt—laterotergite; ns—nematiform seta. Scale bar = 0.05 mm." httpUri="https://zenodo.org/record/167663/files/figure.png" pageId="29" pageNumber="30">
|
||
Fig.
|
||
<date id="712010528900FFD622B53983FBBAFD0A" box="[1000,1030,727,752]" pageId="29" pageNumber="30" value="2015-01">15</date>
|
||
</figureCitation>
|
||
I). Spermatheca sclerotized, slightly elongate, ovoid, with two compartments (
|
||
<figureCitation id="9DA52A178900FFD6232A39A8FD77FCEF" box="[631,715,764,789]" captionStart="FIGURE 16" captionStartId="25.[151,250,1600,1622]" captionTargetBox="[171,1418,211,1571]" captionTargetId="figure@25.[151,1435,193,1579]" captionTargetPageId="25" captionText="FIGURE 16. Line drawings of spermatheca, spermathecal duct, and spermathecal gland of Zeanillus species. A— Z. phyllobius (NZ, Canterbury, Ahuriri Scenic Reserve); B— Z. punctigerus (NZ, Canterbury, Sharplin Falls); C— Z. nunni (NZ, Otago, Woodside Glen); D— Z. pellucidus (NZ, Otago, Oamaru, Glen Warren Reserve); E— Z. pallidus (NZ, Otago, Taieri Mouth); F— Z. brouni (NZ, Otago, Oamaru, Glen Warren Reserve); G— Z. montivagus (NZ, Otago, North Rough Ridge); H— Z. nanus (NZ, Otago, Balclutha). Legend: sd—spermathecal duct; sg—spermathecal gland; sp—spermatheca. Scale bar = 0.05 mm." httpUri="https://zenodo.org/record/167664/files/figure.png" pageId="29" pageNumber="30">Fig. 16</figureCitation>
|
||
G). Spermathecal duct short and without coils. Attachments of spermathecal duct and gland to spermatheca at a moderate distance from each other.
|
||
</paragraph>
|
||
<paragraph id="052136928900FFD6219A3810FDA9FC7A" blockId="29.[151,1437,151,1184]" pageId="29" pageNumber="30">
|
||
<emphasis id="37EAEA808900FFD6219A3810FDB8FCA7" bold="true" box="[199,516,836,861]" pageId="29" pageNumber="30">Geographical distribution.</emphasis>
|
||
This species is known from one locality only in the North Rough Ridge at
|
||
<quantity id="C2669B778900FFD624133810FA21FCA6" box="[1358,1437,836,860]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="29" pageNumber="30" unit="m" value="1000.0">1000m</quantity>
|
||
altitude (
|
||
<figureCitation id="9DA52A178900FFD621A73833FEF7FC7A" box="[250,331,871,896]" captionStart="FIGURE 18" captionStartId="30.[151,250,1918,1940]" captionTargetBox="[198,1387,199,1885]" captionTargetId="figure@30.[189,1398,193,1897]" captionTargetPageId="30" captionText="FIGURE 18. Map of southern half of South Island, New Zealand, showing positions of locality records for the species of Zeanillus. A: white circles, Z. phyllobius; green circles, Z. punctigerus; red circles, Z. nunni; blue diamond, Z. pellucidus; green diamond, Z. pallidus. B: white triangles, Z. brouni; green triangles, Z. nanus; red triangles, Z. lescheni; yellow triangle, Z. carltoni; black triangle, Z. montivagus. Elevation scale bars are given in meters." httpUri="https://zenodo.org/record/167666/files/figure.png" pageId="29" pageNumber="30">Fig. 18</figureCitation>
|
||
B, black triangle).
|
||
</paragraph>
|
||
<paragraph id="052136928900FFD6219A38D8FCABFC5F" blockId="29.[151,1437,151,1184]" box="[199,791,908,933]" pageId="29" pageNumber="30">
|
||
<emphasis id="37EAEA808900FFD6219A38D8FEF3FC5F" bold="true" box="[199,335,908,933]" pageId="29" pageNumber="30">Way of life.</emphasis>
|
||
No precise data on habitat are available.
|
||
</paragraph>
|
||
<paragraph id="052136928900FFD6219A38FBFF46FB5A" blockId="29.[151,1437,151,1184]" pageId="29" pageNumber="30">
|
||
<emphasis id="37EAEA808900FFD6219A38FBFECCFC32" bold="true" box="[199,368,943,968]" pageId="29" pageNumber="30">Relationships.</emphasis>
|
||
In the structure of copulatory sclerites of median lobe, males of this species are distinct from those of the other members of subgenus. In the shape of round sclerite, males of this species are similar to those of the interior
|
||
<taxonomicName id="C29E4D118900FFD6204338ADFE25FBEA" box="[286,409,1016,1040]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="lescheni">
|
||
<emphasis id="37EAEA808900FFD6204338ADFE25FBEA" box="[286,409,1016,1040]" italics="true" pageId="29" pageNumber="30">Z. lescheni</emphasis>
|
||
</taxonomicName>
|
||
and coastal
|
||
<taxonomicName id="C29E4D118900FFD6237638ADFD28FBEA" box="[555,660,1016,1040]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="brouni">
|
||
<emphasis id="37EAEA808900FFD6237638ADFD28FBEA" box="[555,660,1016,1040]" italics="true" pageId="29" pageNumber="30">Z. brouni</emphasis>
|
||
</taxonomicName>
|
||
. Members of
|
||
<taxonomicName id="C29E4D118900FFD6226538ADFC65FBF5" box="[824,985,1016,1040]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="montivagus">
|
||
<emphasis id="37EAEA808900FFD6226538ADFC65FBF5" box="[824,985,1016,1040]" italics="true" pageId="29" pageNumber="30">Z. montivagus</emphasis>
|
||
</taxonomicName>
|
||
are similar to those of
|
||
<taxonomicName id="C29E4D118900FFD625B538ADFAEDFBEA" box="[1256,1361,1016,1040]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="brouni">
|
||
<emphasis id="37EAEA808900FFD625B538ADFAEDFBEA" box="[1256,1361,1016,1040]" italics="true" pageId="29" pageNumber="30">Z. brouni</emphasis>
|
||
</taxonomicName>
|
||
in the presence of ostial fields at apical orifice in the male median lobe and in the shape and proportions of the spermatheca of females (i.e., rounded on one side and tapered on the other). Based on these morphological evidences and species ranges, I suggest that
|
||
<taxonomicName id="C29E4D118900FFD623F73F31FCF1FB86" box="[682,845,1125,1148]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="montivagus">
|
||
<emphasis id="37EAEA808900FFD623F73F31FCF1FB86" box="[682,845,1125,1148]" italics="true" pageId="29" pageNumber="30">Z. montivagus</emphasis>
|
||
</taxonomicName>
|
||
shares a common ancestry with
|
||
<taxonomicName id="C29E4D118900FFD625843F31FAF9FB86" box="[1241,1349,1124,1148]" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="brouni">
|
||
<emphasis id="37EAEA808900FFD625843F31FAF9FB86" box="[1241,1349,1124,1148]" italics="true" pageId="29" pageNumber="30">Z. brouni</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="C29E4D118900FFD624D83F31FF4FFB5A" class="Insecta" family="Carabidae" genus="Zeanillus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="lescheni">
|
||
<emphasis id="37EAEA808900FFD624D83F31FF4FFB5A" italics="true" pageId="29" pageNumber="30">Z. lescheni</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |