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<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.208.80382" ID-Pensoft-Pub="1314-2003-208-1" ID-Pensoft-UUID="F47A23EA345757E29C2800F6D51CEE05" ModsDocID="1314-2003-208-1" checkinTime="1663786009364" checkinUser="pensoft" docAuthor="Martinez-Dominguez, Lili, Nicolalde-Morejon, Fernando, Vergara-Silva, Francisco &amp; Stevenson, Dennis Wm." docDate="2022" docId="39A45C843C3E50ACBCF29799D3EB3B28" docLanguage="en" docName="PhytoKeys 208: 1-102" docOrigin="PhytoKeys 208" docPubDate="2022-09-21" docSource="http://dx.doi.org/10.3897/phytokeys.208.80382" docTitle="Ceratozamia fuscoviridis W. Bull., Retail List. 154: 4. 1879" docType="treatment" docVersion="1" id="F47A23EA345757E29C2800F6D51CEE05" lastPageNumber="1" masterDocId="F47A23EA345757E29C2800F6D51CEE05" masterDocTitle="Monograph of Ceratozamia (Zamiaceae, Cycadales): an endangered genus" masterLastPageNumber="102" masterPageNumber="1" pageNumber="1" updateTime="1663786009364" updateUser="pensoft">
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<mods:title>Monograph of Ceratozamia (Zamiaceae, Cycadales): an endangered genus</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Martinez-Dominguez, Lili</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-1158-1501</mods:nameIdentifier>
<mods:affiliation>Posgrado en Ciencias Biologicas, Instituto de Biologia, Universidad Nacional Autonoma de Mexico, 3 er. Circuito Exterior, Ciudad Universitaria, 04510, Coyoacan, CDMX, Mexico</mods:affiliation>
<mods:nameIdentifier type="email">lilimartinezd@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Nicolalde-Morejon, Fernando</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-1423-7474</mods:nameIdentifier>
<mods:affiliation>Laboratorio de Teoria Evolutiva e Historia de la Ciencia, Instituto de Biologia, Universidad Nacional Autonoma de Mexico, 3 er. Circuito Exterior, Ciudad Universitaria, 04510, Coyoacan, CDMX. Mexico</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Vergara-Silva, Francisco</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-5024-268X</mods:nameIdentifier>
<mods:affiliation>Posgrado en Ciencias Biologicas, Instituto de Biologia, Universidad Nacional Autonoma de Mexico, 3 er. Circuito Exterior, Ciudad Universitaria, 04510, Coyoacan, CDMX, Mexico</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Stevenson, Dennis Wm.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-2986-7076</mods:nameIdentifier>
<mods:affiliation>Laboratorio de Taxonomia Integrativa, Instituto de Investigaciones Biologicas, Universidad Veracruzana, Xalapa, 91190, Veracruz. Mexico</mods:affiliation>
<mods:nameIdentifier type="email">dws@nybg.org</mods:nameIdentifier>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>PhytoKeys</mods:title>
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<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-09-21</mods:number>
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<mods:detail type="volume">
<mods:number>208</mods:number>
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<mods:start>1</mods:start>
<mods:end>102</mods:end>
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<mods:url>http://dx.doi.org/10.3897/phytokeys.208.80382</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.208.80382</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2003-208-1</mods:identifier>
<mods:identifier type="Pensoft-UUID">F47A23EA345757E29C2800F6D51CEE05</mods:identifier>
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<treatment LSID="urn:lsid:plazi:treatment:39A45C843C3E50ACBCF29799D3EB3B28" httpUri="http://treatment.plazi.org/id/39A45C843C3E50ACBCF29799D3EB3B28" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
10.
<taxonomicName LSID="39A45C84-3C3E-50AC-BCF2-9799D3EB3B28" authority="W. Bull., Retail List. 154: 4. 1879" authorityName="W. Bull., Retail List. 154: 4." authorityYear="1879" class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ceratozamia fuscoviridis" order="Cycadales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="fuscoviridis">Ceratozamia fuscoviridis W.Bull., Retail List. 154: 4. 1879</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Comparison of qualitative characters of microsporophylls A Ceratozamia fuscoviridis B C. brevifrons C C. chamberlainii D C. kuesteriana." figureDoi="10.3897/phytokeys.208.80382.figure3" httpUri="https://binary.pensoft.net/fig/746062" pageId="0" pageNumber="1">Figs 3A</figureCitation>
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Color of leaves at emergence A Ceratozamia sancheziae B C. kuesteriana C C. fuscoviridis D C. miqueliana." figureDoi="10.3897/phytokeys.208.80382.figure9" httpUri="https://binary.pensoft.net/fig/746068" pageId="0" pageNumber="1">, 9C</figureCitation>
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Plants of Ceratozamia in habitat A C. alvarezii B C. aurantiaca C C. becerrae D C. brevifrons E C. chamberlainii F C. chimalapensis G C. decumbens H C. delucana I C. euryphyllidia J C. fuscoviridis K C. hondurensis L C. huastecorum." figureDoi="10.3897/phytokeys.208.80382.figure15" httpUri="https://binary.pensoft.net/fig/746074" pageId="0" pageNumber="1">, 15J</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="reference_group">
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Cycadopsida" family="Zamiaceae" form="fuscoviridis" genus="Ceratozamia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ceratozamia mexicana subsp. f." order="Cycadales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="form" species="mexicana" subSpecies="f.">Ceratozamia mexicana Brongn. f. fuscoviridis</taxonomicName>
(W.Bull.) J.Schust., Pflanzenr. (Engler) Heft 99, 4 fam 1: 132. 1932, as
<taxonomicName family="Zamiaceae" form="fuscoviridis" higherTaxonomySource="treatment-meta" kingdom="Plantae" lsidName="C. mexicana subsp. f. var. longifolia" order="Cycadales" pageId="0" pageNumber="1" rank="form" species="mexicana" subSpecies="f." variety="longifolia">C. mexicana var. longifolia f. fuscoviridis</taxonomicName>
. Type: Based on
<taxonomicName class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ceratozamia fuscoviridis" order="Cycadales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="fuscoviridis">Ceratozamia fuscoviridis</taxonomicName>
W.Bull.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">
<typeStatus>Type</typeStatus>
.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<materialsCitation collectingDate="1878-03-21" collectingDateMax="2015-03-31" collectingDateMin="1878-03-21" collectorName="D. Moore, L. Marti" country="Ireland" county="Cultivated" elevation="2" location="Molango-Xochicoatlan" municipality="Glasnevin" specimenCount="493" specimenCount-female="493" stateProvince="Hidalgo" typeStatus="neotype">
<collectingCounty>Cultivated</collectingCounty>
at
<collectingMunicipality>Glasnevin</collectingMunicipality>
,
<collectingCountry name="Ireland">Ireland</collectingCountry>
&quot;Hort. Bot. Glasnevin&quot;,
<collectingDate value="1878-03-21">21 Mar 1878</collectingDate>
(accessioned 1881),
<emphasis italics="true" pageId="0" pageNumber="1">
<collectorName>D. Moore</collectorName>
s.n.
</emphasis>
(
<typeStatus>neotype</typeStatus>
, designated by
<bibRefCitation DOI="https://doi.org/10.12705/661.9" author="Calonje, M" journalOrPublisher="Taxon" pageId="0" pageNumber="1" pagination="158 - 164" refId="B4" refString="Calonje, M, Sennikov, AN, 2017. In the process of saving plant names from oblivion: The revised nomenclature of Ceratozamia fuscoviridis (Zamiaceae). Taxon 66 (1): 158 - 164, DOI: https://doi.org/10.12705/661.9" title="In the process of saving plant names from oblivion: The revised nomenclature of Ceratozamia fuscoviridis (Zamiaceae)." url="https://doi.org/10.12705/661.9" volume="66" year="2017">Calonje and Sennikov 2017</bibRefCitation>
, pg. 161: K! [K000501714, K000501712, K000501713]).
<collectingCountry name="Mexico">Mexico</collectingCountry>
.
<collectingRegion country="Mexico" name="Hidalgo">Hidalgo</collectingRegion>
: Mun. Molango, km 3 carretera
<normalizedToken originalValue="Molango-Xochicoatlán">
<location LSID="urn:lsid:plazi:treatment:39A45C843C3E50ACBCF29799D3EB3B28:0ECA6091C50C0D3DFE01804648108315" country="Ireland" county="Cultivated" municipality="Glasnevin" name="Molango-Xochicoatlan" stateProvince="Hidalgo">Molango-Xochicoatlan</location>
</normalizedToken>
,
<quantity metricMagnitude="0" metricUnit="m" metricValue="1.86" unit="m" value="1.86">
<elevation metricMagnitude="0" metricUnit="m" metricValue="1.86" unit="m" value="1.86">1,860 m</elevation>
</quantity>
,
<collectingDate value="2015-03-31">31 Mar 2015</collectingDate>
,
<emphasis italics="true" pageId="0" pageNumber="1">
<collectorName>L. Marti</collectorName>
́nez-Domínguez et al.
<specimenCount type="female">493 ♀</specimenCount>
</emphasis>
(epitype, designated by
<bibRefCitation DOI="https://doi.org/10.3897/phytokeys.100.23152" author="Martinez-Dominguez, L" journalOrPublisher="PhytoKeys" pageId="0" pageNumber="1" pagination="91 - 124" refId="B25" refString="Martinez-Dominguez, L, Nicolalde-Morejon, F, Vergara-Silva, F, Stevenson, DW, 2018a. Taxonomic review of Ceratozamia (Zamiaceae) in the Sierra Madre Oriental, Mexico. PhytoKeys 100: 91 - 124, DOI: https://doi.org/10.3897/phytokeys.100.23152" title="Taxonomic review of Ceratozamia (Zamiaceae) in the Sierra Madre Oriental, Mexico." url="https://doi.org/10.3897/phytokeys.100.23152" volume="100" year="2018 a">
<normalizedToken originalValue="Martínez-Domínguez">Martinez-Dominguez</normalizedToken>
et al. 2018a
</bibRefCitation>
: 105: CIB! [acc. # 17465UV]; isoepitype: MEXU! [acc. # 1520508, 1520282, 1520519])
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Stem</emphasis>
20-90 cm long, 25-40 cm in diameter, epigeous, erect and decumbent.
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Cataphylls</emphasis>
2-4.5
<normalizedToken originalValue="×">x</normalizedToken>
2-3.5 cm wide at the base, persistent, narrowly triangular, reddish brown, densely brownish tomentose at emergence, tomentose at maturity, apex acuminate.
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Leaves</emphasis>
10-70, 92-215 cm long, ascending, light green and dark brown at emergence, with whitish gray trichomes, glabrous at maturity.
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
40-95 cm long, terete, linear, dark green in mature leaves; with 14-60 (66) thin prickles, 0.09-0.33 cm long.
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Rachis</emphasis>
65-150 cm long, terete, linear, green in mature leaves, with prickles.
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Leaflets</emphasis>
28-67 pairs, opposite to subopposite, insertion in one plane, lanceolate, abaxially curved longitudinally, basally falcate, papyraceous, flat, green, with adaxial and abaxial sides glabrous, distal end with entire margins, acuminate and symmetrical at the apex, attenuate at base, with conspicuous and green-light brown veins; median leaflets 16.6-42
<normalizedToken originalValue="×">x</normalizedToken>
1.3-2.1 cm, 0.6-2 cm between leaflets; articulations 0.6-1.3 cm wide, green and brown.
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Pollen strobili</emphasis>
22-30 cm long, 5-8 cm in diameter, solitary, cylindrical, erect, brownish yellow at emergence, greenish brown with reddish brown trichomes at maturity; peduncle 5-14.5 cm long, 1.6-2.3 cm in diameter, tomentose, reddish brown to brown; microsporophylls 1.6-2.4
<normalizedToken originalValue="×">x</normalizedToken>
1.2-1.6 cm, obconic with a non-recurved distal face and a deeply lobate fertile portion, infertile portion 0.45-0.56 cm long and orbicular with straight horns 0.16-0.29 cm long, 0.53-0.80 cm and a right angle between the horns.
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Ovulate strobili</emphasis>
24-35 cm long, 8.5-15 cm in diameter, solitary, cylindrical, erect, green with brown trichomes at emergence, brown-green with dark brown trichomes at maturity, acuminate apex; peduncle 4-15.5 cm long, 1.8-2.5 cm in diameter, erect or pendulous, tomentose, brown to reddish brown; megasporophylls 49-195, 7-15 orthostichies with 7-15 sporophylls per orthostichy, 2.3-3.5
<normalizedToken originalValue="×">x</normalizedToken>
3.0-4.0 cm, with a truncate distal face, horns straight and 0.70-0.99 cm long, 1.7-2.1 cm between horns and an obtuse angle between the horns.
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Seeds</emphasis>
2-2.6 cm long, 1.5-2 cm in diameter, ovate, sarcotesta whitish yellow to yellow when immature, light brown at maturity.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution and habitat.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ceratozamia fuscoviridis" order="Cycadales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="fuscoviridis">
<emphasis italics="true" pageId="0" pageNumber="1">Ceratozamia fuscoviridis</emphasis>
</taxonomicName>
is endemic to south to central Carso Huasteco in Mexico from Hidalgo to northwest of Veracruz States (Fig.
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Distribution of Ceratozamia species A C. euryphyllidia B C. fuscoviridis C C. hildae D C. hondurensis." figureDoi="10.3897/phytokeys.208.80382.figure17" httpUri="https://binary.pensoft.net/fig/746076" pageId="0" pageNumber="1">17B</figureCitation>
) in cloud forests between 1,300-1,900 m elevation.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">The epithet refers to the dark-brown color of the leaf at emergence.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="common names">
<paragraph pageId="0" pageNumber="1">Common names.</paragraph>
<paragraph pageId="0" pageNumber="1">
Mexico. Veracruz: teocintle, teocintli and tepecintli (
<bibRefCitation DOI="https://doi.org/10.1186/s13002-018-0282-z" author="Bonta, M" journalOrPublisher="Journal of Ethnobiology and Ethnomedicine" pageId="0" pageNumber="1" pagination=": 15" refId="B2" refString="Bonta, M, Pulido-Silva, MT, Diego-Vargas, T, Vite-Reyes, A, Vovides, AP, Cibrian-Jaramillo, A, 2019. Ethnobotany of Mexican and northern Central American cycads (Zamiaceae). Journal of Ethnobiology and Ethnomedicine 4 (1): 15, DOI: https://doi.org/10.1186/s13002-018-0282-z" title="Ethnobotany of Mexican and northern Central American cycads (Zamiaceae)." url="https://doi.org/10.1186/s13002-018-0282-z" volume="4" year="2019">Bonta et al. 2019</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="uses">
<paragraph pageId="0" pageNumber="1">Uses.</paragraph>
<paragraph pageId="0" pageNumber="1">This species has decorative uses associated with religious ceremonies and national holidays. The leaves are commonly used to make arches in the entrances of some Roman Catholic churches or decorate the walls during national holidays.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="conservation status">
<paragraph pageId="0" pageNumber="1">Conservation status.</paragraph>
<paragraph pageId="0" pageNumber="1">
(
<bibRefCitation author="Haynes, JL" journalOrPublisher="Phytotaxa" pageId="0" pageNumber="1" publicationUrl="https://www.iucnredlist.org" refId="B17" refString="2021. . https://www.iucnredlist.org" url="https://www.iucnredlist.org" year="2021">IUCN 2021</bibRefCitation>
).
<taxonomicName class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ceratozamia fuscoviridis" order="Cycadales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="fuscoviridis">
<emphasis italics="true" pageId="0" pageNumber="1">Ceratozamia fuscoviridis</emphasis>
</taxonomicName>
is listed as &quot;Critically Endangered&quot; under criteria B1ab(i,iii,iv,v). This species requires a reassessment of its conservation status because new populations have recently been recorded, including its current circumscription that includes populations from Veracruz State. Based on the number of populations, modelled potential geographical distribution from ecological niche and its occurrence in a Natural Protected Areas (Los
<normalizedToken originalValue="Mármoles">Marmoles</normalizedToken>
National Park),
<taxonomicName family="Zamiaceae" kingdom="Plantae" lsidName="C. fuscoviridis" order="Cycadales" pageId="0" pageNumber="1" rank="species" species="fuscoviridis">
<emphasis italics="true" pageId="0" pageNumber="1">C. fuscoviridis</emphasis>
</taxonomicName>
could be listed as &quot;Endangered (EN)&quot; (
<bibRefCitation DOI="https://doi.org/10.1017/S0030605320000204" author="Martinez-Dominguez, L" journalOrPublisher="Oryx" pageId="0" pageNumber="1" pagination="947 - 956" refId="B28" refString="Martinez-Dominguez, L, Nicolalde-Morejon, F, Stevenson, DW, Santiago-Jimenez, QJ, Rojas-Soto, O, Vergara-Silva, F, 2021. The need for multidisciplinary conservation: A case study of Ceratozamia (Zamiaceae, Cycadales) in eastern Mexico. Oryx 55 (6): 947 - 956, DOI: https://doi.org/10.1017/S0030605320000204" title="The need for multidisciplinary conservation: A case study of Ceratozamia (Zamiaceae, Cycadales) in eastern Mexico." url="https://doi.org/10.1017/S0030605320000204" volume="55" year="2021">
<normalizedToken originalValue="Martínez-Domínguez">Martinez-Dominguez</normalizedToken>
et al. 2021
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="discussion">
<paragraph pageId="0" pageNumber="1">Discussion.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ceratozamia fuscoviridis" order="Cycadales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="fuscoviridis">
<emphasis italics="true" pageId="0" pageNumber="1">Ceratozamia fuscoviridis</emphasis>
</taxonomicName>
is polymorphic within populations because individual plants may have either a light green or dark-brown leaf color at emergence. Individuals with dark-brown leaf at emergence also have a brownish abaxial side. This species is most geographically proximate to
<taxonomicName family="Zamiaceae" kingdom="Plantae" lsidName="C. chamberlainii" order="Cycadales" pageId="0" pageNumber="1" rank="species" species="chamberlainii">
<emphasis italics="true" pageId="0" pageNumber="1">C. chamberlainii</emphasis>
</taxonomicName>
, but it can be distinguished by lanceolate and papyraceous leaflets, a petiole armed with long and thin prickles; and ovulate strobili that are brownish green with dark trichomes at maturity.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<bibRefCitation author="Osborne, R" journalOrPublisher="Cornell University Press, Ithaca" pageId="0" pageNumber="1" refId="B43" refString="Osborne, R, Stevenson, DW, Vovides, AP, 2006. What is Ceratozamia fuscoviridis? Delpinoa 48: 5-10." title="What is Ceratozamia fuscoviridis? Delpinoa 48: 5 - 10." year="2006">Osborne et al. (2006)</bibRefCitation>
intended to validate
<taxonomicName class="Cycadopsida" family="Zamiaceae" genus="Ceratozamia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ceratozamia fuscoviridis" order="Cycadales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="fuscoviridis">
<emphasis italics="true" pageId="0" pageNumber="1">Ceratozamia fuscoviridis</emphasis>
</taxonomicName>
D.Moore, which was invalidly published in 1878. However, in 2017, Calonje &amp; Sennikov published the correction to this name using a brief description by William Bull in horticultural catalogues. This work is the valid publication for the specie and the name &quot;
<taxonomicName family="Zamiaceae" kingdom="Plantae" lsidName="C. fuscoviridis" order="Cycadales" pageId="0" pageNumber="1" rank="species" species="fuscoviridis">
<emphasis italics="true" pageId="0" pageNumber="1">C. fuscoviridis</emphasis>
</taxonomicName>
D.Moore&quot; is an isonym with no nomenclatural status.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Specimens examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
<materialsCitation collectingDate="1992-12-30" collectingDateMax="1995-03-18" collectingDateMin="1992-12-30" collectorName="I. Luna-Vega, J. L. Lopez-Garcia" country="Mexico" county="Mun. Eloxochitlan" location="Mun. Molango de Escamilla" municipality="Mun. Metztitlan" specimenCount="1" stateProvince="Hidalgo">
<collectingCountry name="Mexico">Mexico</collectingCountry>
.
<emphasis bold="true" pageId="0" pageNumber="1">
<collectingRegion country="Mexico" name="Hidalgo">Hidalgo</collectingRegion>
:
<collectingCounty>
Mun.
<normalizedToken originalValue="Eloxochitlán">Eloxochitlan</normalizedToken>
</collectingCounty>
</emphasis>
,
<collectingDate value="1995-03-18">18 Mar 1995</collectingDate>
,
<emphasis italics="true" pageId="0" pageNumber="1">
<collectorName>I. Luna-Vega</collectorName>
54716
</emphasis>
(FCME).
<emphasis bold="true" pageId="0" pageNumber="1">
<collectingMunicipality>
Mun.
<normalizedToken originalValue="Metztitlán">Metztitlan</normalizedToken>
</collectingMunicipality>
</emphasis>
,
<collectingDate value="1992-12-30">30 Dec 1992</collectingDate>
,
<emphasis italics="true" pageId="0" pageNumber="1">
<collectorName>
J.L.
<normalizedToken originalValue="López-García">Lopez-Garcia</normalizedToken>
</collectorName>
449
</emphasis>
(ENCB, IBUG, MEXU).
<emphasis bold="true" pageId="0" pageNumber="1">
<location LSID="urn:lsid:plazi:treatment:39A45C843C3E50ACBCF29799D3EB3B28:933D05076EF312D694462A911B38EDC1" country="Mexico" county="Mun. Eloxochitlan" municipality="Mun. Metztitlan" name="Mun. Molango de Escamilla" stateProvince="Hidalgo">Mun. Molango de Escamilla</location>
</emphasis>
</materialsCitation>
,
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.38" unit="m" value="1380.0">1,380 m</quantity>
,
<date value="1999-05-29">29 May 1999</date>
,
<emphasis italics="true" pageId="0" pageNumber="1">A.P. Vovides 1298</emphasis>
(XAL);
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.4" unit="m" value="1400.0">1,400 m</quantity>
,
<date value="1999-05-29">29 May 1999</date>
,
<emphasis italics="true" pageId="0" pageNumber="1">A.P. Vovides 1301</emphasis>
(XAL);
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.5" unit="m" value="1500.0">1,500 m</quantity>
,
<date value="2008-07-24">24 Jul 2008</date>
,
<emphasis italics="true" pageId="0" pageNumber="1">A. Vite-Reyes et al. 6</emphasis>
(XAL);
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.86" unit="m" value="1860.0">1,860 m</quantity>
,
<date value="2015-03-31">31 Mar 2015</date>
,
<emphasis italics="true" pageId="0" pageNumber="1">
F.
<normalizedToken originalValue="Nicolalde-Morejón">Nicolalde-Morejon</normalizedToken>
et al. 2209
</emphasis>
-
<emphasis italics="true" pageId="0" pageNumber="1">2211</emphasis>
(CIB);
<materialsCitation collectingDate="2015-03-31" collectorName="Martinez-Dominguez, T. W. Walters" country="Mexico" elevation="1860" location="Mun. Tenango de Doria" specimenCount="1" stateProvince="Hidalgo">
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.86" unit="m" value="1860.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.86" unit="m" value="1860.0">1,860 m</elevation>
</quantity>
,
<collectingDate value="2015-03-31">31 Mar 2015</collectingDate>
,
<emphasis italics="true" pageId="0" pageNumber="1">
L.
<normalizedToken originalValue="Martínez-Domínguez">
<collectorName>Martinez-Dominguez</collectorName>
</normalizedToken>
et al. 485
</emphasis>
-492, 494-
<emphasis italics="true" pageId="0" pageNumber="1">514</emphasis>
(CIB);
<emphasis italics="true" pageId="0" pageNumber="1">
<collectorName>T.W. Walters</collectorName>
2001-03-A
</emphasis>
(XAL).
<emphasis bold="true" pageId="0" pageNumber="1">
<location LSID="urn:lsid:plazi:treatment:39A45C843C3E50ACBCF29799D3EB3B28:B9D6F119D9697DDA3A011F72565F0D2C" country="Mexico" name="Mun. Tenango de Doria" stateProvince="Hidalgo">Mun. Tenango de Doria</location>
</emphasis>
</materialsCitation>
,
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.7" unit="m" value="1700.0">1,700 m</quantity>
,
<date value="1993-03-12">12 Mar 1993</date>
,
<emphasis italics="true" pageId="0" pageNumber="1">I. Luna-Vega 914</emphasis>
(FCME).
<emphasis bold="true" pageId="0" pageNumber="1">Mun. Tlanchinol</emphasis>
,
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.45" unit="m" value="1450.0">1,450 m</quantity>
,
<date value="1992-08-25">25 Aug 1992</date>
,
<emphasis italics="true" pageId="0" pageNumber="1">I. Luna-Vega s/n</emphasis>
(FCME);
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.42" unit="m" value="1420.0">1,420 m</quantity>
,
<date value="1992-10-07">7 Oct 1992</date>
,
<emphasis italics="true" pageId="0" pageNumber="1">I. Luna-Vega 625</emphasis>
(FCME, XAL);
<date value="1992-08-25">25 Aug 1992</date>
,
<emphasis italics="true" pageId="0" pageNumber="1">I. Luna-Vega 789</emphasis>
(XAL);
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.3119999999999998" unit="m" value="1312.0">1,312 m</quantity>
,
<date value="2015-03-31">31 Mar 2015</date>
,
<emphasis italics="true" pageId="0" pageNumber="1">
F.
<normalizedToken originalValue="Nicolalde-Morejón">Nicolalde-Morejon</normalizedToken>
et al. 2204-2208
</emphasis>
(CIB);
<materialsCitation collectingDate="2015-03-31" collectorName="Martinez-Dominguez, Jan, J. Rees" country="Mexico" elevation="1312" location="Mun. Zacualtipan de Angeles" specimenCount="1" stateProvince="Hidalgo">
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.3119999999999998" unit="m" value="1312.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.3119999999999998" unit="m" value="1312.0">1,312 m</elevation>
</quantity>
,
<collectingDate value="2015-03-31">31 Mar 2015</collectingDate>
,
<emphasis italics="true" pageId="0" pageNumber="1">
L.
<normalizedToken originalValue="Martínez-Domínguez">
<collectorName>Martinez-Dominguez</collectorName>
</normalizedToken>
et al. 455-484
</emphasis>
(CIB).
<emphasis bold="true" pageId="0" pageNumber="1">
<location LSID="urn:lsid:plazi:treatment:39A45C843C3E50ACBCF29799D3EB3B28:C68583E5C178A4341D09BAC2A1AD7606" country="Mexico" name="Mun. Zacualtipan de Angeles" stateProvince="Hidalgo">
Mun.
<normalizedToken originalValue="Zacualtipán">Zacualtipan</normalizedToken>
de
<normalizedToken originalValue="Ángeles">Angeles</normalizedToken>
</location>
</emphasis>
, 23
<collectorName>Jan</collectorName>
, 1983,
<emphasis italics="true" pageId="0" pageNumber="1">
<collectorName>J. Rees</collectorName>
389
</emphasis>
(CHAPA, FCME, MEXU)
</materialsCitation>
;
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.3599999999999999" unit="m" value="1360.0">1,360 m</quantity>
,
<date value="1974-12-04">4 Dec 1974</date>
,
<emphasis italics="true" pageId="0" pageNumber="1">J. Rees 1611</emphasis>
(CHIP, XAL);
<date value="1974-12-04">4 Dec 1974</date>
,
<emphasis italics="true" pageId="0" pageNumber="1">J. Rees 6339</emphasis>
(IEB).
<emphasis bold="true" pageId="0" pageNumber="1">
<collectingRegion country="Mexico" name="Veracruz de Ignacio de la Llave">Veracruz</collectingRegion>
: Mun. Huayacocotla
</emphasis>
,
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.913" unit="m" value="1913.0">1,913 m</quantity>
,
<date value="2005-02-23">23 Feb 2005</date>
,
<emphasis italics="true" pageId="0" pageNumber="1">
D. Saavedra
<normalizedToken originalValue="Millán">Millan</normalizedToken>
64
</emphasis>
(FCME);
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.85" unit="m" value="1850.0">1,850 m</quantity>
,
<date value="1980-03-13">13 Mar 1980</date>
,
<emphasis italics="true" pageId="0" pageNumber="1">J. Palma G. 63</emphasis>
(XAL);
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.844" unit="m" value="1844.0">1,844 m</quantity>
,
<date value="2015-04-01">1 Apr 2015</date>
,
<emphasis italics="true" pageId="0" pageNumber="1">
F.
<normalizedToken originalValue="Nicolalde-Morejón">Nicolalde-Morejon</normalizedToken>
et al. 2212
</emphasis>
-
<emphasis italics="true" pageId="0" pageNumber="1">2214</emphasis>
(CIB);
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.7" unit="m" value="1700.0">1,700 m</quantity>
,
<date value="1981-03-24">24 Mar 1981</date>
,
<emphasis italics="true" pageId="0" pageNumber="1">L. Ballesteros &amp; F. Ballesteros 460</emphasis>
(XAL);
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.55" unit="m" value="1550.0">1,550 m</quantity>
,
<date value="1981-04-23">23 Apr 1981</date>
,
<emphasis italics="true" pageId="0" pageNumber="1">
L.G.
<normalizedToken originalValue="Juárez">Juarez</normalizedToken>
G. 47
</emphasis>
(XAL);
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.844" unit="m" value="1844.0">1,844 m</quantity>
,
<date value="2015-04-01">1 Apr 2015</date>
,
<emphasis italics="true" pageId="0" pageNumber="1">
L.
<normalizedToken originalValue="Martínez-Domínguez">Martinez-Dominguez</normalizedToken>
et al. 515
</emphasis>
-
<emphasis italics="true" pageId="0" pageNumber="1">544</emphasis>
(CIB);
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.9" unit="m" value="1900.0">1,900 m</quantity>
,
<date value="1972-02-11">11 Feb 1972</date>
,
<emphasis italics="true" pageId="0" pageNumber="1">
R.
<normalizedToken originalValue="Hernández">Hernandez</normalizedToken>
M. 1507
</emphasis>
(MEXU, XAL);
<date value="1975-02-26">26 Feb 1975</date>
,
<emphasis italics="true" pageId="0" pageNumber="1">V. Sosa 59</emphasis>
(XAL).
</paragraph>
</subSubSection>
</treatment>
</document>