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<document id="F44C3E07B1BA87D74447C53E86B64085" ID-CLB-Dataset="46105" ID-DOI="10.11646/zootaxa.3784.3.8" ID-GBIF-Dataset="3fe0e8f2-edf3-4ac9-afc9-f5266e2ededd" ID-ISSN="1175-5326" ID-Zenodo-Dep="228085" ID-ZooBank="413F693F-91C7-4014-9CCF-E13326BFDFF8" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460403565278" checkinUser="plazi" docAuthor="Munari, Lorenzo" docDate="2014" docId="03B387BD31598E3D4D93DCF48930FEE4" docLanguage="en" docName="zt03784p293.pdf" docOrigin="Zootaxa 3784 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pseudorhicnoessa longicerca Munari, 2014, sp. nov." docType="treatment" docVersion="9" lastPageNumber="291" masterDocId="FF8AFFC5315E8E374D04DE238A2EFFA4" masterDocTitle="New Indo-Pacific species of Dasyrhicnoessa Hendel, 1934 and Pseudorhicnoessa Malloch, 1914 (Diptera: Canacidae: Tethininae)" masterLastPageNumber="293" masterPageNumber="281" pageNumber="288" updateTime="1698299550856" updateUser="ExternalLinkService">
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<mods:title id="EDC8CB125E233E742E10A8208AC45E73">New Indo-Pacific species of Dasyrhicnoessa Hendel, 1934 and Pseudorhicnoessa Malloch, 1914 (Diptera: Canacidae: Tethininae)</mods:title>
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<mods:name id="D0B5DCF14D7D4BCCA1C25C48D8A2AF58" type="personal">
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<mods:namePart id="668AB98150B2EDFAB04ACD72D36A4BD7">Munari, Lorenzo</mods:namePart>
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<mods:title id="5806297ECB114DF5D32AE205B6AE10EB">Zootaxa</mods:title>
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<mods:date id="85825567A9A651A062692CDDD3427E4C">2014</mods:date>
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<mods:number id="852DBE35B9D2669546DEF2A53F78E71F">3784</mods:number>
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<mods:identifier id="15E9B5AA0D32ED67BB24CECF1D133C18" type="DOI">10.11646/zootaxa.3784.3.8</mods:identifier>
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<treatment id="03B387BD31598E3D4D93DCF48930FEE4" ID-DOI="http://doi.org/10.5281/zenodo.6144288" ID-GBIF-Taxon="119420614" ID-Zenodo-Dep="6144288" LSID="urn:lsid:plazi:treatment:03B387BD31598E3D4D93DCF48930FEE4" httpUri="http://treatment.plazi.org/id/03B387BD31598E3D4D93DCF48930FEE4" lastPageId="10" lastPageNumber="291" pageId="7" pageNumber="288">
<subSubSection id="C300652031598E304D93DCF48B24FCB7" pageId="7" pageNumber="288" type="nomenclature">
<paragraph id="8BA536AB31598E304D93DCF48872FD56" blockId="7.[151,604,727,787]" box="[151,604,727,754]" pageId="7" pageNumber="288">
<heading id="D0ED81C731598E304D93DCF48872FD56" bold="true" box="[151,604,727,754]" fontSize="11" level="1" pageId="7" pageNumber="288" reason="1">
<emphasis id="B96EEAB931598E304D93DCF48872FD56" bold="true" box="[151,604,727,754]" pageId="7" pageNumber="288">
<taxonomicName id="4C1A4D2831598E304D93DCF48BD9FD55" ID-CoL="8DJB3" box="[151,503,727,753]" class="Insecta" family="Canacidae" genus="Pseudorhicnoessa" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="288" phylum="Arthropoda" rank="species" species="longicerca" status="sp. nov.">
<emphasis id="B96EEAB931598E304D93DCF48BD9FD55" bold="true" box="[151,503,727,753]" italics="true" pageId="7" pageNumber="288">Pseudorhicnoessa longicerca</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25D57C231598E304CFADCFB8872FD56" box="[510,604,728,754]" pageId="7" pageNumber="288" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BA536AB31598E304D93DCD98B24FCB7" blockId="7.[151,604,727,787]" box="[151,266,762,787]" pageId="7" pageNumber="288">
(
<figureCitation id="13212A2E31598E304D9BDCD98B2DFCB7" box="[159,259,762,787]" captionStart="FIGURES 7 9" captionStartId="8.[151,264,1857,1880]" captionTargetBox="[163,1427,1191,1828]" captionTargetId="figure@8.[151,1436,1173,1836]" captionTargetPageId="8" captionText="FIGURES 7 9. ♂ terminalia of Pseudorhicnoessa longicerca sp. nov.. 7: lateral view; 8: posterior view; 9: left anterior surstylus, anterior view. Scale bar = 0.3 mm." httpUri="https://zenodo.org/record/228089/files/figure.png" pageId="7" pageNumber="288">Figs 79</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C300652031598E304D93DD6188CAFBAA" pageId="7" pageNumber="288" type="diagnosis">
<paragraph id="8BA536AB31598E304D93DD6188CAFBAA" blockId="7.[151,1437,834,2011]" pageId="7" pageNumber="288">
<emphasis id="B96EEAB931598E304D93DD618B3FFCFF" bold="true" box="[151,273,834,859]" pageId="7" pageNumber="288">Diagnosis.</emphasis>
This species is distinguished from congeners mainly by the following characters of the male terminalia: anterior surstylus distinctly curved backward, rounded apically, finger-like in lateral view, paddle-shaped in anterior and posterior views, bearing several long setae on ventral margin; posterior surstylus sickle-like (falciform) in lateral view, roughly subtriangular in posterior view; cercus conspicuously developed, slightly longer than anterior and posterior surstyli length, membranous part of both cerci apparently fused subapically, becoming widely separated and characteristically bifid at apex.
</paragraph>
</subSubSection>
<subSubSection id="C300652031598E3F4DC3DA398F70FBC5" lastPageId="8" lastPageNumber="289" pageId="7" pageNumber="288" type="description">
<paragraph id="8BA536AB31598E304DC3DA398E3CFB96" blockId="7.[151,1437,834,2011]" box="[199,1042,1050,1075]" pageId="7" pageNumber="288">
<emphasis id="B96EEAB931598E304DC3DA398B77FB97" bold="true" box="[199,345,1050,1075]" pageId="7" pageNumber="288">Description.</emphasis>
<emphasis id="B96EEAB931598E304C5BDA388BBAFB96" box="[351,404,1051,1074]" italics="true" pageId="7" pageNumber="288">Size.</emphasis>
Body length
<quantity id="4CE29B4E31598E304F2FDA3988FEFB96" box="[555,720,1050,1075]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" metricValueMax="3.52" metricValueMin="3.08" pageId="7" pageNumber="288" unit="mm" value="3.3" valueMax="3.52" valueMin="3.08">3.083.52 mm</quantity>
, wing length 2.00
<quantity id="4CE29B4E31598E304EA3DA388E20FB96" box="[935,1038,1051,1075]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="7" pageNumber="288" unit="mm" value="2.2">2.20 mm</quantity>
.
</paragraph>
<paragraph id="8BA536AB31598E304DC3DA1D8BF8FB3B" blockId="7.[151,1437,834,2011]" pageId="7" pageNumber="288">
<emphasis id="B96EEAB931598E304DC3DA1D8B0EFBF3" box="[199,288,1086,1111]" italics="true" pageId="7" pageNumber="288">Habitus</emphasis>
. Robust fly with dark grey (thorax) to brown (abdomen) body; legs mostly yellow, with apical parts of hind femur and tibia strongly infuscate; setal vestiture black; wing yellowish grey; male with large, globoid, broadly exposed epandrium.
</paragraph>
<paragraph id="8BA536AB31598E304DC3DA8988FEF9D7" blockId="7.[151,1437,834,2011]" pageId="7" pageNumber="288">
<emphasis id="B96EEAB931598E304DC3DA898B2AFB67" box="[199,260,1194,1219]" italics="true" pageId="7" pageNumber="288">Head</emphasis>
. Slightly higher than long; vertex and occiput cinereous, frons yellow, face apparently membranous, invested with pale whitish microtomentum, gena faintly yellow to whitish, evenly microtomentose; postocellar setae inclinate, somewhat spaced, about as long as dorsal postocular seta; medial vertical seta oriented inward, strong, about as long as lateral vertical seta, which is lateroclinate; postocular setae arranged in 2 distinct rows; ocellar triangle bearing some minute setulae between and behind ocelli, in addition to a pair of long, strong ocellar setae; 3 long fronto-orbital setae, posterior ones distinctly lateroclinate, anterior seta slightly reclinate; frontoorbitals intermixed with a few lateroclinate and inclinate short setulae; 3 pairs of proclinate and more or less slightly inclinate interfrontal setae, intermixed with some short additional setulae, anterior pair with setae weaker and shorter than posterior pairs; antenna yellow, with basal flagellomere widely dark brown; arista brown to black, microscopically pubescent; eye apparently glabrous, large, subcircular to slightly oblong, its vertical diameter about 4.6 times as long as genal height (
<typeStatus id="54A1880931598E304F6AD83188F8F98F" box="[622,726,1554,1579]" pageId="7" pageNumber="288" type="holotype">holotype</typeStatus>
); 6
<emphasis id="B96EEAB931598E304FFED8318926F98F" bold="true" box="[762,776,1554,1579]" pageId="7" pageNumber="288"></emphasis>
7 peristomal setae, including pseudovibrissa, increasingly longer towards vibrissal angle; clypeus distinctly exposed; mouthparts yellow (palpus) to brownish (proboscis), labellum slightly shorter than buccal cavity length.
</paragraph>
<paragraph id="8BA536AB31598E3F4DC3D85D8885FDC7" blockId="7.[151,1437,834,2011]" lastBlockId="8.[151,1436,151,1121]" lastPageId="8" lastPageNumber="289" pageId="7" pageNumber="288">
<emphasis id="B96EEAB931598E304DC3D85D8B36F933" box="[199,280,1662,1687]" italics="true" pageId="7" pageNumber="288">Thorax</emphasis>
. Blackish in ground colour, invested with grey microtomentum, excepting postpronotal lobe which is mostly yellow, whitish if viewed dorsally, and notopleuron faintly to distinctly yellowish; 1+3 dorsocentral setae, posterior one very long; acrostichal setulae arranged in 7
<emphasis id="B96EEAB931598E304E54D8E58970F97B" bold="true" box="[848,862,1734,1759]" pageId="7" pageNumber="288"></emphasis>
8 rows; acrostichal prescutellar setae noticeably developed; 3 postpronotal setae intermixed with several short setulae, each major seta with a different orientation; 1 presutural; 2 notopleurals; 2 supra-alars, ventral one distinctly shorter and weaker; 2 postalars; several rows of short setulae between dorsocentrals and the lateral side of scutum; scutellum with 2 pairs of long setae, several minute setulae on disc and on lateral margins; 1 proepisternal; 1 proepimeral; anepisternum densely setulose, with 2 long postero-marginal setae and 1 dorsally oriented, posterodorsal seta; katepisternum sparsely setulose, with 1 long, strong seta on posterodorsal margin; anepimeron and meron without setae and setulae.
<emphasis id="B96EEAB931598E3049ABD9BC8ECAF812" box="[1199,1252,1951,1974]" italics="true" pageId="7" pageNumber="288">Legs</emphasis>
. Mostly yellow, excepting apical parts of hind femur and tibia, and a patch on antero-apical area of forefemur, which are strongly dark brown; distal 2 tarsomeres distinctly black to brown infuscate; coxae with several long setae; forefemur slightly swollen, with some long setae dorsally and posterodorsally, a posteroventral row of long, fine setae on entire length of femur, an anteroventral row of fine, close, long setae on proximal two thirds, apical third bearing a ctenidial comb of short, close, pointed spinulae; foretibia and foretarsus evenly setulose; male mid femur with some anterior setae on distal half, a dense row of robust, long setae posteroventrally; mid tibia with 3
<emphasis id="B96EEAB931568E3F4884DF048FA0FEE4" bold="true" box="[1408,1422,295,320]" pageId="8" pageNumber="289"></emphasis>
4 posterodorsal setae, 1
<emphasis id="B96EEAB931568E3F4C8FDF6F8BB7FEC1" bold="true" box="[395,409,332,357]" pageId="8" pageNumber="289"></emphasis>
2 anterior to anterodorsal setae, 1 apicoventral, spur-like, long and strong seta; hind femur distinctly swollen in males, with 1
<emphasis id="B96EEAB931568E3F4F10DF4C880CFE2C" bold="true" box="[532,546,367,392]" pageId="8" pageNumber="289"></emphasis>
2 anterior setae approximately in the middle and 3
<emphasis id="B96EEAB931568E3F4956DF4C8E4EFE2C" bold="true" box="[1106,1120,367,392]" pageId="8" pageNumber="289"></emphasis>
4 dorsal setae on apical half, decreasing in length towards apex of femur; hind tibia bearing 2 dorsal to subdorsal, 2 anterodorsal, 2, sometimes 3, posterodorsal setae; 1 apicoventral spur-like seta.
<emphasis id="B96EEAB931568E3F4FE4DF9A8936FE74" box="[736,792,441,464]" italics="true" pageId="8" pageNumber="289">Wing</emphasis>
. Veins brownish yellow, membrane yellowish grey; alula with fringe formed by several, long, grey setae; subcostal break deeply notched; first costal sector bearing 2 long apical setae; costal vein reaching end of R4+5; R2+3 and R4+5 slightly divergent distally; R4+5 and M1 subparallel on distal half; crossvein
<emphasis id="B96EEAB931568E3F4C8CDC0B8B9EFD9B" box="[392,432,552,575]" italics="true" pageId="8" pageNumber="289">r-m</emphasis>
ending approximately near middle of cell
<emphasis id="B96EEAB931568E3F4E96DC05899AFD9B" box="[914,948,550,575]" italics="true" pageId="8" pageNumber="289">dm</emphasis>
; crossvein
<emphasis id="B96EEAB931568E3F4932DC058E52FD9B" box="[1078,1148,550,575]" italics="true" pageId="8" pageNumber="289">dm-cu</emphasis>
less than one third of the length of last section of CuA1; halter yellowish.
</paragraph>
<paragraph id="8BA536AB31568E3F4DC3DC528F70FBC5" blockId="8.[151,1436,151,1121]" pageId="8" pageNumber="289">
<emphasis id="B96EEAB931568E3F4DC3DC528B1FFD2E" box="[199,305,625,650]" italics="true" pageId="8" pageNumber="289">Abdomen</emphasis>
. Brown, invested with pale grey microtomentum, bearing several setulae dorsally and laterally, and long postero-marginal setae; tergite 6 bare dorsally, bearing a row of 5
<emphasis id="B96EEAB931568E3F4EA1DCB6899DFD0A" bold="true" box="[933,947,661,686]" pageId="8" pageNumber="289"></emphasis>
6 setae on postero-lateral margin; syntergite 1+2 and tergite 3 with obvious postero-marginal yellowish stripes; sternites yellow, pubescent.
<emphasis id="B96EEAB931568E3F49D9DC9A8FBBFD76" box="[1245,1429,697,722]" italics="true" pageId="8" pageNumber="289">
<collectingCountry id="F30D763B31568E3F49D9DC9A8F38FD76" box="[1245,1302,697,722]" name="Malta" pageId="8" pageNumber="289">Male</collectingCountry>
terminalia
</emphasis>
. Epandrium very large, globoid, densely setulose; anterior surstylus distinctly curved backward, rounded apically, finger-like in lateral view (fig. 7), paddle-shaped in anterior (fig. 9) and posterior views, bearing several long setae on ventral margin; posterior surstylus sharply sickle-like in lateral view (fig. 7), roughly subtriangular in posterior view (fig. 8); cercus conspicuously developed, slightly longer than length of anterior and posterior surstyli (figs 7 8), membranous part of both cerci apparently fused subapically, becoming widely separated and characteristically bifid at apex (fig. 8); phallus noticeably widened on proximal two thirds, ribbon-like on apical third; widened part strongly pubescent ventrally, minutely pubescent (papilla-like setulae) dorsally; ejaculatory apodeme and phallapodeme as usual for the subfamily.
<emphasis id="B96EEAB931568E3F4F6CDDFA8916FC56" box="[616,824,985,1010]" italics="true" pageId="8" pageNumber="289">Female terminalia</emphasis>
. Postabdomen and spermathecae with no considerable differences compared to
<taxonomicName id="4C1A4D2831568E3F4CC0DDDD886DFBB1" box="[452,579,1022,1045]" class="Insecta" family="Canacidae" genus="Pseudorhicnoessa" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="289" phylum="Arthropoda" rank="species" species="spinipes">
<emphasis id="B96EEAB931568E3F4CC0DDDD886DFBB1" box="[452,579,1022,1045]" italics="true" pageId="8" pageNumber="289">P. spinipes</emphasis>
</taxonomicName>
; apex of postabdomen telescopically retractile; spermathecae distinctly cylindrical, about as long as high in lateral view (a larger series of specimens is needed to verify the consistency of some very slight differences of the outlines of both the 7th tergite and sternite as well as of the spermathecae).
</paragraph>
</subSubSection>
<caption id="DF65662331568E3F4D93D9628872F8D2" httpUri="https://zenodo.org/record/228089/files/figure.png" pageId="8" pageNumber="289" targetBox="[163,1427,1191,1828]" targetPageId="8">
<paragraph id="8BA536AB31568E3F4D93D9628872F8D2" blockId="8.[151,1436,1857,1910]" pageId="8" pageNumber="289">
<emphasis id="B96EEAB931568E3F4D93D9628B10F8F3" bold="true" box="[151,318,1857,1880]" pageId="8" pageNumber="289">FIGURES 79.</emphasis>
♂ terminalia of
<taxonomicName id="4C1A4D2831568E3F4CF2D961890DF8FC" box="[502,803,1858,1880]" class="Insecta" family="Canacidae" genus="Pseudorhicnoessa" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="289" phylum="Arthropoda" rank="species" species="longicerca" status="sp. nov.">
<emphasis id="B96EEAB931568E3F4CF2D961890DF8FC" box="[502,803,1858,1880]" italics="true" pageId="8" pageNumber="289">Pseudorhicnoessa longicerca</emphasis>
</taxonomicName>
<emphasis id="B96EEAB931568E3F4E2AD96189ACF8F3" bold="true" box="[814,898,1858,1879]" pageId="8" pageNumber="289">
<taxonomicNameLabel id="A25D57C231568E3F4E2AD96189ACF8F3" box="[814,898,1858,1879]" pageId="8" pageNumber="289" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
. 7: lateral view; 8: posterior view; 9: left anterior surstylus, anterior view. Scale bar = 0.3 mm.
</paragraph>
</caption>
<caption id="DF65662331578E3E4D93DC4088D1FD13" httpUri="https://zenodo.org/record/228090/files/figure.png" pageId="9" pageNumber="290" targetBox="[162,1421,203,582]" targetPageId="9">
<paragraph id="8BA536AB31578E3E4D93DC4088D1FD13" blockId="9.[151,1436,611,695]" pageId="9" pageNumber="290">
<emphasis id="B96EEAB931578E3E4D93DC408B78FDDD" bold="true" box="[151,342,611,634]" pageId="9" pageNumber="290">FIGURES 1013.</emphasis>
♂ terminalia of
<taxonomicName id="4C1A4D2831578E3E4F0CDC4789AEFDDE" authority="Malloch." authorityName="Malloch." box="[520,896,612,634]" class="Insecta" family="Canacidae" genus="Pseudorhicnoessa" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="290" phylum="Arthropoda" rank="species" species="spinipes">
<emphasis id="B96EEAB931578E3E4F0CDC478932FDDE" box="[520,796,612,634]" italics="true" pageId="9" pageNumber="290">Pseudorhicnoessa spinipes</emphasis>
Malloch.
</taxonomicName>
10: anterior and posterior surstyli, lateral view; 11: posterior surstylus, lateral view from a slightly different inclination; 12: cerci and anterior and posterior surstyli, posterior view; 13: left anterior surstylus, anterior view. Scale bar = 0.3 mm.
</paragraph>
</caption>
<subSubSection id="C300652031578E3E4DC3DCFD8817FAB7" pageId="9" pageNumber="290" type="materials_examined">
<paragraph id="8BA536AB31578E3E4DC3DCFD8B2BFB27" blockId="9.[151,1436,734,2019]" pageId="9" pageNumber="290">
<emphasis id="B96EEAB931578E3E4DC3DCFD8B56FD53" bold="true" box="[199,376,734,759]" pageId="9" pageNumber="290">
<typeStatus id="54A1880931578E3E4DC3DCFD8B2CFD53" box="[199,258,734,759]" pageId="9" pageNumber="290">Type</typeStatus>
material.
</emphasis>
The
<typeStatus id="54A1880931578E3E4CBCDCFD8834FD53" box="[440,538,734,759]" pageId="9" pageNumber="290" type="holotype">holotype</typeStatus>
♂ of
<taxonomicName id="4C1A4D2831578E3E4F62DCFD899DFD53" box="[614,947,734,759]" class="Insecta" family="Canacidae" genus="Pseudorhicnoessa" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="290" phylum="Arthropoda" rank="species" species="longicerca" status="sp. nov.">
<emphasis id="B96EEAB931578E3E4F62DCFD899DFD53" box="[614,947,734,759]" italics="true" pageId="9" pageNumber="290">Pseudorhicnoessa longicerca</emphasis>
</taxonomicName>
<emphasis id="B96EEAB931578E3E4EB8DCFD8E34FD53" bold="true" box="[956,1050,734,759]" pageId="9" pageNumber="290">
<taxonomicNameLabel id="A25D57C231578E3E4EB8DCFD8E34FD53" box="[956,1050,734,759]" pageId="9" pageNumber="290" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is labelled “[printed white label] Papua-New-Guinea / Central Prov., / Gaba Gaba, [
<geoCoordinate id="EE2E506C31578E3E4FDBDD20897DFCBF" box="[735,851,771,795]" direction="south" orientation="latitude" pageId="9" pageNumber="290" precision="15" value="-9.803333">9°48'12&quot;S</geoCoordinate>
<geoCoordinate id="EE2E506C31578E3E4E58DD2089C3FCBF" box="[860,1005,771,795]" direction="east" orientation="longitude" pageId="9" pageNumber="290" precision="15" value="147.51334">147°30'48&quot;E</geoCoordinate>
] /
<date id="FFA4106B31578E3E490BDD218EA1FCBF" box="[1039,1167,770,795]" pageId="9" pageNumber="290" value="1982-12-12">12.xii.1982</date>
// [printed white label] J.W. Ismay / strand line / NHM 1995-E16 // [printed red label]
<typeStatus id="54A1880931578E3E4E5DDD0589E0FC9B" box="[857,974,806,831]" pageId="9" pageNumber="290" type="holotype">Holotypus</typeStatus>
/
<taxonomicName id="4C1A4D2831578E3E4EE0DD058E81FC9B" box="[996,1199,806,831]" class="Insecta" family="Canacidae" genus="Pseudorhicnoessa" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="290" phylum="Arthropoda" rank="genus">
<emphasis id="B96EEAB931578E3E4EE0DD058E81FC9B" box="[996,1199,806,831]" italics="true" pageId="9" pageNumber="290">Pseudorhicnoessa</emphasis>
</taxonomicName>
/
<taxonomicName id="4C1A4D2831578E3E49C1DD058F12FC9B" box="[1221,1340,806,831]" class="Insecta" family="Canacidae" genus="Pseudorhicnoessa" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="290" phylum="Arthropoda" rank="species" species="longicerca" status="sp. nov.">
<emphasis id="B96EEAB931578E3E49C1DD058F12FC9B" box="[1221,1340,806,831]" italics="true" pageId="9" pageNumber="290">longicerca</emphasis>
</taxonomicName>
<emphasis id="B96EEAB931578E3E4846DD058FB2FC9B" bold="true" box="[1346,1436,806,831]" pageId="9" pageNumber="290">
<taxonomicNameLabel id="A25D57C231578E3E4846DD058FB2FC9B" box="[1346,1436,806,831]" pageId="9" pageNumber="290" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
♂ / L. Munari des. 2013”. The
<typeStatus id="54A1880931578E3E4F10DD698858FCC7" box="[532,630,842,867]" pageId="9" pageNumber="290" type="holotype">holotype</typeStatus>
is double mounted (micropinned to a block of plastic matter), is in excellent condition (apical tarsomeres of left hind leg lost), with the abdomen removed, dissected, and stored in a small plastic tube filled with glycerol and pinned below the specimen.
<typeStatus id="54A1880931578E3E4EBADDB08E01FC0F" box="[958,1071,915,939]" pageId="9" pageNumber="290" type="paratype">Paratypes</typeStatus>
: same data as
<typeStatus id="54A1880931578E3E49D0DDB18F17FC0F" box="[1236,1337,914,939]" pageId="9" pageNumber="290" type="holotype">holotype</typeStatus>
, 3 ♂♂ 4 ♀♀;
<collectingCountry id="F30D763B31578E3E4DD4DD9B8B83FC6B" box="[208,429,950,975]" name="Papua New Guinea" pageId="9" pageNumber="290">Papua New Guinea</collectingCountry>
, Port Moresby, Ela Beach,
<date id="FFA4106B31578E3E4FE6DD958968FC6B" box="[738,838,950,975]" pageId="9" pageNumber="290" value="1963-10-06">6.x.1963</date>
, D.K. McAlpine, 4 ♂♂ (formerly misidentified as
<taxonomicName id="4C1A4D2831578E3E4883DD9B8ADDFC57" class="Insecta" family="Canacidae" genus="Pseudorhicnoessa" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="290" phylum="Arthropoda" rank="species" species="spinipes">
<emphasis id="B96EEAB931578E3E4883DD9B8ADDFC57" italics="true" pageId="9" pageNumber="290">P. spinipes</emphasis>
</taxonomicName>
). The
<typeStatus id="54A1880931578E3E4C47DDF98B8BFC57" box="[323,421,986,1011]" pageId="9" pageNumber="290" type="holotype">holotype</typeStatus>
and five
<typeStatus id="54A1880931578E3E4F1CDDF888ADFC57" box="[536,643,987,1011]" pageId="9" pageNumber="290" type="paratype">paratypes</typeStatus>
(2 ♂♂ 3 ♀♀), all from the same
<typeStatus id="54A1880931578E3E491DDDF88E67FC57" box="[1049,1097,987,1011]" pageId="9" pageNumber="290">type</typeStatus>
locality, are deposited in the Natural History Museum, London,
<collectingCountry id="F30D763B31578E3E4F2CDA23887AFBB3" box="[552,596,1024,1047]" name="United Kingdom" pageId="9" pageNumber="290">UK</collectingCountry>
; a pair of
<typeStatus id="54A1880931578E3E4FCEDA23891BFBBC" box="[714,821,1024,1048]" pageId="9" pageNumber="290" type="paratype">paratypes</typeStatus>
is deposited in the authors collection together with a third specimen (♂) formerly misidentified as
<taxonomicName id="4C1A4D2831578E3E4F91DA078923FB9F" box="[661,781,1060,1083]" class="Insecta" family="Canacidae" genus="Pseudorhicnoessa" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="290" phylum="Arthropoda" rank="species" species="spinipes">
<emphasis id="B96EEAB931578E3E4F91DA078923FB9F" box="[661,781,1060,1083]" italics="true" pageId="9" pageNumber="290">P. spinipes</emphasis>
</taxonomicName>
(see above) and already present in the authors collection; 3 additional males from Ela Beach (Port Moresby,
<collectingCountry id="F30D763B31578E3E4FE1DA648931FBFB" box="[741,799,1095,1119]" name="Papua New Guinea" pageId="9" pageNumber="290">PNG</collectingCountry>
) are also deposited in the Australian Museum, Sydney,
<collectingCountry id="F30D763B31578E3E4D93DA498B2FFB27" box="[151,257,1130,1155]" name="Australia" pageId="9" pageNumber="290">Australia</collectingCountry>
.
</paragraph>
<paragraph id="8BA536AB31578E3E4DC3DAB38817FAB7" blockId="9.[151,1436,734,2019]" pageId="9" pageNumber="290">
♀♀ of
<taxonomicName id="4C1A4D2831578E3E4C1DDAB38BBFFB03" box="[281,401,1168,1191]" class="Insecta" family="Canacidae" genus="Pseudorhicnoessa" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="290" phylum="Arthropoda" rank="species" species="spinipes">
<emphasis id="B96EEAB931578E3E4C1DDAB38BBFFB03" box="[281,401,1168,1191]" italics="true" pageId="9" pageNumber="290">P. spinipes</emphasis>
</taxonomicName>
are at present indistinguishable from those of
<taxonomicName id="4C1A4D2831578E3E4EA7DAB38E18FB03" box="[931,1078,1166,1191]" class="Insecta" family="Canacidae" genus="Pseudorhicnoessa" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="290" phylum="Arthropoda" rank="species" species="longicerca" status="sp. nov.">
<emphasis id="B96EEAB931578E3E4EA7DAB38E18FB03" box="[931,1078,1166,1191]" italics="true" pageId="9" pageNumber="290">P. longicerca</emphasis>
</taxonomicName>
<emphasis id="B96EEAB931578E3E493ADAAD8EB7FB03" bold="true" box="[1086,1177,1166,1191]" pageId="9" pageNumber="290">
<taxonomicNameLabel id="A25D57C231578E3E493ADAAD8EB7FB03" box="[1086,1177,1166,1191]" pageId="9" pageNumber="290" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
. Consequently, among the material identified as
<taxonomicName id="4C1A4D2831578E3E4CB6DA978807FB6F" box="[434,553,1204,1227]" class="Insecta" family="Canacidae" genus="Pseudorhicnoessa" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="290" phylum="Arthropoda" rank="species" species="spinipes">
<emphasis id="B96EEAB931578E3E4CB6DA978807FB6F" box="[434,553,1204,1227]" italics="true" pageId="9" pageNumber="290">P. spinipes</emphasis>
</taxonomicName>
, preserved in many scientific institutions, there might be occasional specimens of
<taxonomicName id="4C1A4D2831578E3E4DB0DAFB8B69FB4B" box="[180,327,1238,1263]" class="Insecta" family="Canacidae" genus="Pseudorhicnoessa" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="290" phylum="Arthropoda" rank="species" species="longicerca">
<emphasis id="B96EEAB931578E3E4DB0DAFB8B69FB4B" box="[180,327,1238,1263]" italics="true" pageId="9" pageNumber="290">P. longicerca</emphasis>
</taxonomicName>
intermixed with Mallochs species, these two species occurring in sympatry and syntopy, at least in the Papuan localities recorded above.
</paragraph>
</subSubSection>
<subSubSection id="C300652031578E3E4DC3DB3D8869FA93" box="[199,583,1310,1335]" pageId="9" pageNumber="290" type="distribution">
<paragraph id="8BA536AB31578E3E4DC3DB3D8869FA93" blockId="9.[151,1436,734,2019]" box="[199,583,1310,1335]" pageId="9" pageNumber="290">
<emphasis id="B96EEAB931578E3E4DC3DB3D8B4EFA93" bold="true" box="[199,352,1310,1335]" pageId="9" pageNumber="290">Distribution.</emphasis>
<collectingCountry id="F30D763B31578E3E4C63DB03886CFA93" box="[359,578,1310,1335]" name="Papua New Guinea" pageId="9" pageNumber="290">Papua New Guinea</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C300652031578E3E4DC3DB618930FADB" pageId="9" pageNumber="290" type="etymology">
<paragraph id="8BA536AB31578E3E4DC3DB618930FADB" blockId="9.[151,1436,734,2019]" pageId="9" pageNumber="290">
<emphasis id="B96EEAB931578E3E4DC3DB618B60FAFF" bold="true" box="[199,334,1346,1371]" pageId="9" pageNumber="290">Etymology.</emphasis>
The species epithet,
<taxonomicName id="4C1A4D2831578E3E4F39DB61889AFAFF" box="[573,692,1346,1371]" class="Insecta" family="Canacidae" genus="Pseudorhicnoessa" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="290" phylum="Arthropoda" rank="species" species="longicerca">
<emphasis id="B96EEAB931578E3E4F39DB61889AFAFF" box="[573,692,1346,1371]" italics="true" pageId="9" pageNumber="290">longicerca</emphasis>
</taxonomicName>
, is a compound name of Latin and ancient Greek derivations and refers to the remarkably long cerci of the male terminalia.
</paragraph>
</subSubSection>
<subSubSection id="C300652031578E3E4DC3DBA98FB4F847" pageId="9" pageNumber="290" type="discussion">
<paragraph id="8BA536AB31578E3E4DC3DBA98FB4F847" blockId="9.[151,1436,734,2019]" pageId="9" pageNumber="290">
<emphasis id="B96EEAB931578E3E4DC3DBA98B15FA07" bold="true" box="[199,315,1418,1443]" pageId="9" pageNumber="290">Remarks.</emphasis>
The genus
<taxonomicName id="4C1A4D2831578E3E4CC6DBA988A3FA07" box="[450,653,1418,1443]" class="Insecta" family="Canacidae" genus="Pseudorhicnoessa" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="290" phylum="Arthropoda" rank="genus">
<emphasis id="B96EEAB931578E3E4CC6DBA988A3FA07" box="[450,653,1418,1443]" italics="true" pageId="9" pageNumber="290">Pseudorhicnoessa</emphasis>
</taxonomicName>
includes two vicariant species so far, namely
<taxonomicName id="4C1A4D2831578E3E4999DBAF8FB9FA07" authority="Munari, 1981" authorityName="Munari" authorityYear="1981" box="[1181,1431,1418,1443]" class="Insecta" family="Canacidae" genus="Pseudorhicnoessa" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="290" phylum="Arthropoda" rank="species" species="rattii">
<emphasis id="B96EEAB931578E3E4999DBAF8EDCFA07" box="[1181,1266,1420,1443]" italics="true" pageId="9" pageNumber="290">P. rattii</emphasis>
<bibRefCitation id="EF8B4B5A31578E3E49FFDBA98FB9FA07" author="Munari" box="[1275,1431,1418,1443]" pageId="9" pageNumber="290" refString="Munari, L. (1981) Sul genere Pseudorhicnoessa Malloch, 1914 (Diptera Tethinidae). Societa Veneziana di Scienze Naturali - Lavori, 6, 91 - 96." type="journal article" year="1981">Munari, 1981</bibRefCitation>
</taxonomicName>
, distributed throughout the Indian Ocean and well characterized by structures of the male terminalia, and
<taxonomicName id="4C1A4D2831578E3E4821DB938B11FA4F" authority="Malloch, 1914" authorityName="Malloch" authorityYear="1914" class="Insecta" family="Canacidae" genus="Pseudorhicnoessa" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="290" phylum="Arthropoda" rank="species" species="spinipes">
<emphasis id="B96EEAB931578E3E4821DB938FB2FA63" box="[1317,1436,1456,1479]" italics="true" pageId="9" pageNumber="290">P. spinipes</emphasis>
<bibRefCitation id="EF8B4B5A31578E3E4D93DBF18B11FA4F" author="Malloch" box="[151,319,1490,1515]" pageId="9" pageNumber="290" refString="Malloch, J. R. (1914) Formosan Agromyzidae. Annales Musei Nationalis Hungarici, 12, 306 - 336." type="journal article" year="1914">Malloch, 1914</bibRefCitation>
</taxonomicName>
, with a wide distribution in the western Pacific Ocean.
<collectingCountry id="F30D763B31578E3E4EB9DBF089F2FA4F" box="[957,988,1491,1515]" name="American Samoa" pageId="9" pageNumber="290">As</collectingCountry>
mentioned previously, the new species occurs in sympatry, as well as in syntopy, with the common
<taxonomicName id="4C1A4D2831578E3E4E5CDBDB89FCF9AB" box="[856,978,1528,1551]" class="Insecta" family="Canacidae" genus="Pseudorhicnoessa" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="290" phylum="Arthropoda" rank="species" species="spinipes">
<emphasis id="B96EEAB931578E3E4E5CDBDB89FCF9AB" box="[856,978,1528,1551]" italics="true" pageId="9" pageNumber="290">P. spinipes</emphasis>
</taxonomicName>
in the eastern territories of
<collectingCountry id="F30D763B31578E3E481CDBDB8AC2F997" name="Papua New Guinea" pageId="9" pageNumber="290">Papua New Guinea</collectingCountry>
.
<taxonomicName id="4C1A4D2831578E3E4DFCD839886AF997" box="[248,580,1562,1587]" class="Insecta" family="Canacidae" genus="Pseudorhicnoessa" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="290" phylum="Arthropoda" rank="species" species="longicerca" status="sp. nov.">
<emphasis id="B96EEAB931578E3E4DFCD839886AF997" box="[248,580,1562,1587]" italics="true" pageId="9" pageNumber="290">Pseudorhicnoessa longicerca</emphasis>
</taxonomicName>
<emphasis id="B96EEAB931578E3E4F49D8398884F997" bold="true" box="[589,682,1562,1587]" pageId="9" pageNumber="290">
<taxonomicNameLabel id="A25D57C231578E3E4F49D8398884F997" box="[589,682,1562,1587]" pageId="9" pageNumber="290" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is strikingly similar and likewise very closely related to the very common
<taxonomicName id="4C1A4D2831578E3E4C05D8638B54F9F3" box="[257,378,1600,1623]" class="Insecta" family="Canacidae" genus="Pseudorhicnoessa" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="290" phylum="Arthropoda" rank="species" species="spinipes">
<emphasis id="B96EEAB931578E3E4C05D8638B54F9F3" box="[257,378,1600,1623]" italics="true" pageId="9" pageNumber="290">P. spinipes</emphasis>
</taxonomicName>
as to external features, especially chaetotaxy, but is distinguished from the latter (figs 7
<emphasis id="B96EEAB931578E3E486DD81D8F59F9F3" bold="true" box="[1385,1399,1598,1623]" pageId="9" pageNumber="290"></emphasis>
13) almost exclusively by structures of the external terminalia of male (figs 7
<emphasis id="B96EEAB931578E3E4EECD84189D8F9DF" bold="true" box="[1000,1014,1634,1659]" pageId="9" pageNumber="290"></emphasis>
9), in particular by having the cercal appendages strongly developed, longer than anterior and posterior surstyli length (figs 7
<emphasis id="B96EEAB931578E3E497BD8A58EA3F93B" bold="true" box="[1151,1165,1670,1695]" pageId="9" pageNumber="290"></emphasis>
8), with fused subapical part and distinctly bifid apex (fig. 8); in
<taxonomicName id="4C1A4D2831578E3E4F6CD88F88CFF967" box="[616,737,1708,1731]" class="Insecta" family="Canacidae" genus="Pseudorhicnoessa" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="290" phylum="Arthropoda" rank="species" species="spinipes">
<emphasis id="B96EEAB931578E3E4F6CD88F88CFF967" box="[616,737,1708,1731]" italics="true" pageId="9" pageNumber="290">P. spinipes</emphasis>
</taxonomicName>
these appendages are simple, short (fig. 12), as usual for the entire subfamily. Additional characters of the external male terminalia, such as the morphology of the anterior and posterior surstyli, are quite consistent, though to a lesser extent (figs 10
<emphasis id="B96EEAB931578E3E4EB9D8D189E5F8AF" bold="true" box="[957,971,1778,1803]" pageId="9" pageNumber="290"></emphasis>
13). Several male specimens of the allied
<taxonomicName id="4C1A4D2831578E3E4D93D93B8B20F88B" box="[151,270,1816,1839]" class="Insecta" family="Canacidae" genus="Pseudorhicnoessa" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="290" phylum="Arthropoda" rank="species" species="spinipes">
<emphasis id="B96EEAB931578E3E4D93D93B8B20F88B" box="[151,270,1816,1839]" italics="true" pageId="9" pageNumber="290">P. spinipes</emphasis>
</taxonomicName>
from northern Queensland (
<collectingCountry id="F30D763B31578E3E4F4DD935889DF88B" box="[585,691,1814,1839]" name="Australia" pageId="9" pageNumber="290">Australia</collectingCountry>
) (especially from localities facing the south-eastern coast of
<collectingCountry id="F30D763B31578E3E4853D93B8B08F8F7" name="Papua New Guinea" pageId="9" pageNumber="290">Papua New Guinea</collectingCountry>
) and from the Couburg Peninsula (Northern Territory,
<collectingCountry id="F30D763B31578E3E4E8AD91989D9F8F7" box="[910,1015,1850,1875]" name="Australia" pageId="9" pageNumber="290">Australia</collectingCountry>
) have also been examined in order to ascertain the possible occurrence of
<taxonomicName id="4C1A4D2831578E3E4F38D94388FCF8D3" box="[572,722,1886,1911]" class="Insecta" family="Canacidae" genus="Pseudorhicnoessa" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="290" phylum="Arthropoda" rank="species" species="longicerca" status="sp. nov.">
<emphasis id="B96EEAB931578E3E4F38D94388FCF8D3" box="[572,722,1886,1911]" italics="true" pageId="9" pageNumber="290">P. longicerca</emphasis>
</taxonomicName>
<emphasis id="B96EEAB931578E3E4FD8D97D8916F8D3" bold="true" box="[732,824,1886,1911]" pageId="9" pageNumber="290">
<taxonomicNameLabel id="A25D57C231578E3E4FD8D97D8916F8D3" box="[732,824,1886,1911]" pageId="9" pageNumber="290" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
among them.
<collectingCountry id="F30D763B31578E3E4EE5D97C8E2EF8D3" box="[993,1024,1887,1911]" name="American Samoa" pageId="9" pageNumber="290">As</collectingCountry>
a result, no specimens of the latter species were found from those northern regions of
<collectingCountry id="F30D763B31578E3E4FE5D9A18965F83F" box="[737,843,1922,1947]" name="Australia" pageId="9" pageNumber="290">Australia</collectingCountry>
. Thus, from the present state of knowledge, we can infer that the distribution of the new species might supposedly be strictly localized in the south-eastern coast of
<collectingCountry id="F30D763B31578E3E4D93D9E88B54F847" box="[151,378,1994,2019]" name="Papua New Guinea" pageId="9" pageNumber="290">Papua New Guinea</collectingCountry>
, or at most on the eastern seashores of that large island. While examining materials of
<emphasis id="B96EEAB931578E3E4883D9EF8FB4F847" box="[1415,1434,1996,2019]" italics="true" pageId="9" pageNumber="290">P.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C300652031548E3D4D93DEBA8930FEE4" pageId="10" pageNumber="291" type="reference_group">
<paragraph id="8BA536AB31548E3D4D93DEBA8930FEE4" blockId="10.[151,1436,151,320]" pageId="10" pageNumber="291">
<taxonomicName id="4C1A4D2831548E3D4D93DEBA8ADDFF14" ID-CoL="4P9QB" box="[151,243,153,176]" class="Insecta" family="Canacidae" genus="Pseudorhicnoessa" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="291" phylum="Arthropoda" rank="species" species="spinipes">
<emphasis id="B96EEAB931548E3D4D93DEBA8ADDFF14" box="[151,243,153,176]" italics="true" pageId="10" pageNumber="291">spinipes</emphasis>
</taxonomicName>
from the collection of the Australian Museum, Sydney, I was also able to study Mallochs
<typeStatus id="54A1880931548E3D481DDEB48F55FF14" box="[1305,1403,151,176]" pageId="10" pageNumber="291" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C1A4D2831548E3D4D93DE9F8BDDFF71" ID-CoL="8FDRD" box="[151,499,188,213]" class="Insecta" family="Canacidae" genus="Tethina" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="291" phylum="Arthropoda" rank="species" species="tibiseta" subGenus="Macrotethina">
<emphasis id="B96EEAB931548E3D4D93DE9F8AC4FF71" box="[151,234,188,213]" italics="true" pageId="10" pageNumber="291">Tethina</emphasis>
(
<emphasis id="B96EEAB931548E3D4DFEDE9F8BBFFF71" box="[250,401,188,213]" italics="true" pageId="10" pageNumber="291">Macrotethina</emphasis>
)
<emphasis id="B96EEAB931548E3D4CA5DE9F8BDDFF71" box="[417,499,188,213]" italics="true" pageId="10" pageNumber="291">tibiseta</emphasis>
</taxonomicName>
, which was formerly placed in synonymy under
<taxonomicName id="4C1A4D2831548E3D4910DE9E8F11FF70" ID-CoL="4P9QB" authority="Munari, 2002" authorityName="Munari" authorityYear="2002" box="[1044,1343,188,213]" class="Insecta" family="Canacidae" genus="Pseudorhicnoessa" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="291" phylum="Arthropoda" rank="species" species="spinipes">
<emphasis id="B96EEAB931548E3D4910DE9E8EA4FF70" box="[1044,1162,189,212]" italics="true" pageId="10" pageNumber="291">P. spinipes</emphasis>
(
<bibRefCitation id="EF8B4B5A31548E3D499DDE9F8F19FF70" author="Munari" box="[1177,1335,188,213]" pageId="10" pageNumber="291" refString="Munari, L. (2002) New species, records, and a new synonymy in the genera Dasyrhicnoessa Hendel, 1934 and Pseudorhicnoessa Malloch, 1914 (Diptera: Tethinidae). Studia dipterologica, [2001], 8 (2), 539 - 551." type="journal article" year="2002">Munari, 2002</bibRefCitation>
)
</taxonomicName>
. On that occasion, the synonymy was based on the examination of three
<typeStatus id="54A1880931548E3D4E6ADEC389F4FF5C" box="[878,986,224,248]" pageId="10" pageNumber="291" type="paratype">paratypes</typeStatus>
of
<taxonomicName id="4C1A4D2831548E3D4906DEC28E5CFF5C" ID-CoL="8FDRD" box="[1026,1138,223,248]" class="Insecta" family="Canacidae" genus="Tethina" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="291" phylum="Arthropoda" rank="species" species="tibiseta">
<emphasis id="B96EEAB931548E3D4906DEC28E5CFF5C" box="[1026,1138,223,248]" italics="true" pageId="10" pageNumber="291">T. tibiseta</emphasis>
</taxonomicName>
from the collection of the National Museum of Natural History, Smithsonian Institution, Washington D.C.. Thus, I can now corroborate that former synonymy, this time based on Mallochs
<typeStatus id="54A1880931548E3D4FB0DF048934FEE4" box="[692,794,295,320]" pageId="10" pageNumber="291" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
</treatment>
</document>