409 lines
40 KiB
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409 lines
40 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.796.22517" ID-GBIF-Dataset="cc0b7d13-b28b-438b-826d-9fc4d840f76e" ID-Pensoft-Pub="1313-2970-796-347" ID-ZBK="6F53022DE32F4E8987E46C65E9A3C7D7" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1313-2970-796-347" ModsDocOrigin="ZooKeys 796" ModsDocTitle="Review of the Neotropical genus Rhyncholepta with descriptions of three new species-group taxa (Hemiptera, Heteroptera, Pentatomidae)" checkinTime="1542332347104" checkinUser="pensoft" docAuthor="Kment, Petr, Eger, Joe E. & Rider 3, Jr. David A." docDate="2018" docId="75C424D22E9BB4A50250FA22EB748C3D" docLanguage="en" docName="ZooKeys 796: 347-395" docOrigin="ZooKeys 796" docSource="http://dx.doi.org/10.3897/zookeys.796.22517" docTitle="Rhyncholepta grandicallosa subsp. centroamericana Kment, Eger & 3, 2018, subsp. n." docType="treatment" docUuid="6B0F453F-819C-4360-BC13-9E87ED75901A" docUuidSource="ZooBank" docVersion="6" lastPageNumber="380" masterDocId="FFDFFFAB9F2AFF87FFF8FFE6FFA49E2D" masterDocTitle="Review of the Neotropical genus Rhyncholepta with descriptions of three new species-group taxa (Hemiptera, Heteroptera, Pentatomidae)" masterLastPageNumber="395" masterPageNumber="347" originalUpdateDomain="Boston" originalUpdateTime="1542332347104" originalUpdateUser="pensoft" pageNumber="365" updateTime="1643466164501" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Review of the Neotropical genus Rhyncholepta with descriptions of three new species-group taxa (Hemiptera, Heteroptera, Pentatomidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Kment, Petr</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Eger, Joe E.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Rider 3, Jr. David A.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2018</mods:date>
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<mods:detail type="volume">
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<mods:number>796</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>347</mods:start>
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<mods:end>395</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.796.22517</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.796.22517</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-796-347</mods:identifier>
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<mods:identifier type="ZBK">6F53022DE32F4E8987E46C65E9A3C7D7</mods:identifier>
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<mods:identifier type="ZooBank">6F53022DE32F4E8987E46C65E9A3C7D7</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="149667333" LSID="urn:lsid:zoobank.org:act:6B0F453F-819C-4360-BC13-9E87ED75901A" httpUri="http://treatment.plazi.org/id/75C424D22E9BB4A50250FA22EB748C3D" lastPageId="33" lastPageNumber="380" pageId="18" pageNumber="365">
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<subSubSection pageId="18" pageNumber="365" type="nomenclature">
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<paragraph pageId="18" pageNumber="365">
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<taxonomicName LSID="http://zoobank.org/6B0F453F-819C-4360-BC13-9E87ED75901A" class="Insecta" family="Pentatomidae" genus="Rhyncholepta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Rhyncholepta grandicallosa" order="Hemiptera" pageId="18" pageNumber="365" phylum="Arthropoda" rank="subSpecies" species="grandicallosa" subSpecies="centroamericana">Rhyncholepta grandicallosa centroamericana</taxonomicName>
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<taxonomicNameLabel pageId="18" pageNumber="365">subsp. n.</taxonomicNameLabel>
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Figs 5, 10, 13, 19-22, 26-28, 32-35, 44-47, 56-59, 67-68, 73-74, 81-83, 91, 95
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</paragraph>
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</subSubSection>
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<subSubSection pageId="19" pageNumber="366" type="reference_group">
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<paragraph pageId="19" pageNumber="366">
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<taxonomicName class="Insecta" family="Pentatomidae" genus="Rhyncholepta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Rhyncholepta grandicallosa" order="Hemiptera" pageId="19" pageNumber="366" phylum="Arthropoda" rank="species" species="grandicallosa">
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<pageBreakToken pageId="19" pageNumber="366" start="start">Rhyncholepta</pageBreakToken>
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grandicallosa
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</taxonomicName>
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:
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<bibRefCitation author="Becker, M" journalOrPublisher="Revista Brasileira de Biologia" pageId="41" pageNumber="388" pagination="389 - 399" title="Sobre o genero Rhyncholepta Bergroth, 1911, com a descricao de uma nova especie (Hemiptera, Pentatomidae, Pentatominae)." volume="31" year="1971">Becker and Grazia-Vieira (1971)</bibRefCitation>
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: 396 (partim, records from Panama);
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<bibRefCitation author="Froeschner, RC" journalOrPublisher="Memoirs of the American Entomological Institute" pageId="42" pageNumber="389" pagination="1 - 393" title="True bugs (Heteroptera) of Panama: A synoptic catalog as a contribution to the study of Panamanian biodiversity." volume="61" year="1999">Froeschner (1999)</bibRefCitation>
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: 185 (checklist);
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<bibRefCitation pageId="19" pageNumber="366">Arismendi (2002)</bibRefCitation>
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: 31 (distribution);
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<bibRefCitation author="Arismendi, N" journalOrPublisher="Insecta Mundi" pageId="41" pageNumber="388" pagination="218 - 236" title="Pentatomidae (Heteroptera) of Honduras: a checklist with description of a new ochlerine genus." volume="17" year="2003">Arismendi and Thomas (2003)</bibRefCitation>
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: 223, 230 (distribution, records);
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<bibRefCitation author="Cambra, RA" journalOrPublisher="Revista Nicaragueense de Entomologia" pageId="41" pageNumber="388" pagination="1 - 21" title="Los Pentatomidos (Hemiptera: Heteroptera) de Panama." volume="149" year="2018">Cambra et al. (2018)</bibRefCitation>
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: 13, 17: Figure 37 (list, photo).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="19" pageNumber="366" type="type locality">
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<paragraph pageId="19" pageNumber="366">Type locality.</paragraph>
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<paragraph pageId="19" pageNumber="366">
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Panama,
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<normalizedToken originalValue="Panamá">Panama</normalizedToken>
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Province, El Lano
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<normalizedToken originalValue="Cartí">Carti</normalizedToken>
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Road, km 8-11, 1100' [= 335 m a.s.l.], ca.
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<geoCoordinate direction="north" orientation="latitude" precision="925" value="9.283334">9°17'N</geoCoordinate>
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<geoCoordinate direction="west" orientation="longitude" precision="925" value="-78.96667">78°58'W</geoCoordinate>
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.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="23" lastPageNumber="370" pageId="19" pageNumber="366" type="type material">
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<paragraph pageId="19" pageNumber="366">Type material.</paragraph>
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<paragraph pageId="19" pageNumber="366">
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Holotype: ♂, "PAN. Panama Prv / El Lano Carti Rd k / 8-11 24
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<normalizedToken originalValue="May–">May-</normalizedToken>
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2 June / 1992 1100' JE Wappes [printed, white label] // DBT [printed, white label] // ♂ [printed, white label] // HOLOTYPUS / RHYNCHOLEPTA / grandicallosa subsp. / CENTROAMERICANA nov. / det. Kment, Eger, Rider 2017 [printed, red label]" (DBTC USNM). The holotype is pinned through scutellum, left distiflagellum is missing.
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</paragraph>
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<paragraph lastPageId="23" lastPageNumber="370" pageId="19" pageNumber="366">
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Paratypes: MEXICO: Chiapas: Palenque [
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="17.487501">17°29'15"N</geoCoordinate>
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<geoCoordinate direction="west" orientation="longitude" precision="15" value="-92.04639">92°02'47"W</geoCoordinate>
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], MV Light, 19.viii.1990, 1 ♂, P. J. Landolt lgt. (JEEC). - BELIZE: British Honduras, 5.iv.1937, 1 ♂, no collector, J. Grazia-Vieira 1973 det. (DARC); British Honduras, Rio Grande, viii.1935, 1 ♀, J. J. White lgt., B.M. 1935-597,
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<taxonomicName class="Insecta" family="Pentatomidae" genus="Rhyncholepta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Rhyncholepta grandicallosa" order="Hemiptera" pageId="19" pageNumber="366" phylum="Arthropoda" rank="species" species="grandicallosa">Rhyncholepta grandicallosa</taxonomicName>
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det. Ruckes 1961 (BMNH). - GUATEMALA: Izabal Province: 30 km SE of Morales, Finca Firmeza [
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<geoCoordinate direction="north" orientation="latitude" precision="55" value="15.379">15.379°N</geoCoordinate>
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<geoCoordinate direction="west" orientation="longitude" precision="55" value="-88.695">88.695°W</geoCoordinate>
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], at light, 7.-22.vii.2008, 4 ♂♂ 2 ♀♀, F. Skillman, C. and L.
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<normalizedToken originalValue="O’Brien">O'Brien</normalizedToken>
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lgt. (FSCA); Izabal, road up to Firmeza, 30 km SE Morales, 22.vii.2008. 1 ♂, C. W. and L. B.
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<normalizedToken originalValue="O’Brien">O'Brien</normalizedToken>
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and F. Skillman lgt. (JEEC); D. Izabel, Firmeza, 30 km SE Morales, at UV and metal halide light, 23.vii.2008, 1 ♀, C. W. and L. B.
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<normalizedToken originalValue="O’Brien">O'Brien</normalizedToken>
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and F. Skillman lgt. (JEEC). - COSTA RICA: Alajuela Province: 20 km S Upala [ca.
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="10.719722">10°43'11"N</geoCoordinate>
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<geoCoordinate direction="west" orientation="longitude" precision="15" value="-85.01833">85°01'06"W</geoCoordinate>
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], 11.-21.vi.1991, 1 ♀, F. D. Parker lgt. (DBTC); Ca. 15 km SW
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<normalizedToken originalValue="Volcán">Volcan</normalizedToken>
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Arenal, Arenal Vista Lodge [ca.
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="10.429723">10°25'47"N</geoCoordinate>
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<geoCoordinate direction="west" orientation="longitude" precision="15" value="-84.76111">84°45'40"W</geoCoordinate>
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], at mercury vapor and black light, 13.-15.viii.1995, 1 ♂ 1 ♀, J. E. Eger coll. [Costa Rica Collecting Permit No. 00113412] (1 ♂ JEEC, 1 ♀ INBIO);
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<normalizedToken originalValue="Caño">Cano</normalizedToken>
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Negro [ca.
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="10.893055">10°53'35"N</geoCoordinate>
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<geoCoordinate direction="west" orientation="longitude" precision="15" value="-84.79583">84°47'45"W</geoCoordinate>
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], R.N.V.S., 20 m a.s.l., 5.-28.ii.1995, 1 ♂, K. F. Flores lgt., L_N_319100_450200 #4424 (INBIO); Finca Monte Sele [ca.
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<geoCoordinate direction="north" orientation="latitude" precision="555" value="10.57">10.57°N</geoCoordinate>
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<geoCoordinate direction="west" orientation="longitude" precision="555" value="-85.25">85.25°W</geoCoordinate>
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]
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<pageBreakToken pageId="20" pageNumber="367" start="start">,</pageBreakToken>
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750 m a.s.l., ix.1994, 2 ♂♂, C. Moraga lgt., L N 326200_379900 #3204 (INBIO). Cartago Province: Monumento Nacional Guayabo [ca.
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="9.97111">9°58'16"N</geoCoordinate>
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<geoCoordinate direction="west" orientation="longitude" precision="15" value="-83.690834">83°41'27"W</geoCoordinate>
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], Turrialba, 1100 m a.s.l., ix.1994, 1 ♂, G. Fonseca lgt., L N 217200_570300 #3202 (INBIO); Paso Marcos env.,
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<geoCoordinate direction="north" orientation="latitude" precision="9" value="9.811833">9°48.71'N</geoCoordinate>
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<geoCoordinate direction="west" orientation="longitude" precision="9" value="-83.4975">83°29.85'W</geoCoordinate>
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, 2450' [= 747 m a.s.l.], 1 ♂, D. Thomas, D. Robacker and W. Warfield lgt. (DBTC). Guanacaste Province: Est. Pitilla, 9 km S Sta. Cecilia, P. N. Guanacaste, xi.1988, 1 ♂, C. Chaves and M. Espinosa lgt., L-N 330200, 380200 (INBIO); Est. Pitilla, 9 km S Sta. Cecilia, P. N. Guanacaste, i.-ii.1990, 1 ♂, GNP Biod. Survey, L-N 330200, 380200 (INBIO); Est. Pitilla, 9 km S Sta. Cecilia [ca.
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="10.98">10°58'48"N</geoCoordinate>
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<geoCoordinate direction="west" orientation="longitude" precision="15" value="-85.41334">85°24'48"W</geoCoordinate>
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], P. N. Guanacaste, x.1992, 1 ♂, C. Moraga lgt., L-N 330200, 380200 (INBIO); Hacienda El Oro, 450-500 m a.s.l., vii.1996, 1 ♂, A. Masis, M. M. Chavarria, C. Moraga, P.
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<normalizedToken originalValue="Ríos">Rios</normalizedToken>
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, de Luz [= light trap], L_N_332600_377400 #45263 (INBIO). Heredia Province: Est. El Ceibo, Braulio Carrillo N. P., 400-600 m a.s.l., v.1990, 1 ♀, C. Chaves lgt., L-N-527700, 256500 (INBIO); Est. Magsasay [ca.
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="10.405277">10°24'19"N</geoCoordinate>
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<geoCoordinate direction="west" orientation="longitude" precision="15" value="-84.055">84°03'18"W</geoCoordinate>
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], P.N. Braulio Carrillo, 200 m a.s.l., v.1991, 1 ♀, A.
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<normalizedToken originalValue="Fernández">Fernandez</normalizedToken>
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lgt., L-N-264600, 531000 (INBIO); Finca la Selva Verde, 12 km S. Puerto Viejo, 500 ft [= 152 m a.s.l.], 23.-26.ix.1986, 1 ♂ 1 ♀, J. E. Eger lgt. (JEEC); La Selva Biological Station, 3 km S Pto. Viejo,
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<geoCoordinate direction="north" orientation="latitude" precision="925" value="10.433333">10°26'N</geoCoordinate>
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<geoCoordinate direction="west" orientation="longitude" precision="925" value="-84.01667">84°01'W</geoCoordinate>
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,
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<pageBreakToken pageId="21" pageNumber="368" start="start">26</pageBreakToken>
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.vii.1992, 2 ♂♂, H. A. Hespenheide lgt., La Selva Project (DARC); Los Arbolitos [ca.
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="10.646389">10°38'47"N</geoCoordinate>
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<geoCoordinate direction="west" orientation="longitude" precision="15" value="-83.99333">83°59'36"W</geoCoordinate>
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], 30 m a.s.l., 20.-27.iii.1993, 2 ♂♂, F. Araya lgt., L N 536100_291400 #1952 (1 ♂ FSCA, 1 ♂ INBIO); near Puerto Viejo, La Selva Biological Station,
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<geoCoordinate direction="north" orientation="latitude" precision="925" value="10.416667">10°25'N</geoCoordinate>
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<geoCoordinate direction="west" orientation="longitude" precision="925" value="-84.0">84°00'W</geoCoordinate>
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, 179 ft [= 55 m a.s.l.], at light, 1.iii.2004, 1 ♂, C. R. Bartlett, J. Cryan and J. Urban lgt. (DARC).
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<normalizedToken originalValue="Limón">Limon</normalizedToken>
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Province: Amubri [ca.
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="9.526111">9°31'34"N</geoCoordinate>
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<geoCoordinate direction="west" orientation="longitude" precision="15" value="-82.95222">82°57'08"W</geoCoordinate>
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], 70 m a.s.l., 12.-31.x.1993, 1 ♂, G. Gallardo lgt., L S 385500_578000 #2407 (INBIO); Amubri, A. C. Amistad, 70 m a.s.l., 2.-20.ix.1993,
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<pageBreakToken pageId="22" pageNumber="369" start="start">1</pageBreakToken>
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♂, G. M. Gallardo lgt., L S 385500_578000 #2368 (INBIO); Cerro Tortuguero [ca.
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="10.583889">10°35'02"N</geoCoordinate>
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<geoCoordinate direction="west" orientation="longitude" precision="15" value="-83.52723">83°31'38"W</geoCoordinate>
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], P.N. Tortuguero, 100 m a.s.l., xi.1989, 1 ♂, J. Solano lgt., L-N 285000, 588000 (INBIO); Cerro Tortuguero, P.N. Tortuguero, 0-100 m a.s.l., v.1990, 1 ♀, J. Solano lgt., L-N 285000, 588000 (INBIO); Cerro Tortuguero, P.N. Tortuguero, 0-120 m a.s.l., vi.1991, 2 ♂♂, R. Delgado lgt., L-N 285000, 588000 (INBIO); Est. Cuatro Esquinas [ca.
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="10.456388">10°27'23"N</geoCoordinate>
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<geoCoordinate direction="west" orientation="longitude" precision="15" value="-83.67194">83°40'19"W</geoCoordinate>
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], P. N. Tortuguero, 0 m a.s.l., 27.iii.-29.iv.1992, 2 ♂♂, D. Garcia lgt., L-N 280000, 590500 (INBIO); Est. Hitoy Cerere, R. Cerere, Res. Biol. Hitoy [ca.
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="9.6472225">9°38'50"N</geoCoordinate>
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<geoCoordinate direction="west" orientation="longitude" precision="15" value="-83.07083">83°04'15"W</geoCoordinate>
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], 100 m a.s.l., 28.-12.[sic!].iv.1992, 1 ♂, E. Lopez lgt., L-N 184200, 643300 (INBIO);
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<normalizedToken originalValue="Río">Rio</normalizedToken>
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Sardinas, R.N.F.S. Barra del Colorado [ca.
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="10.768611">10°46'07"N</geoCoordinate>
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<geoCoordinate direction="west" orientation="longitude" precision="15" value="-83.585556">83°35'08"W</geoCoordinate>
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], 10 m a.s.l., 14.x.1992, 1 ♂, F. Araya lgt., L N 291500_564700 (INBIO);
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<normalizedToken originalValue="Río">Rio</normalizedToken>
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Sardinas, R.N.F.S. Barra del Colorado, 10 m a.s.l., 12.-30.ix.1993, 4 ♂♂ 4 ♀♀, F. Araya lgt., L N 291500_564700 #2355 (2 ♂♂ 2 ♀♀ INBIO, 1 ♂ BMNH, 1 ♂ 1 ♀ NHMW, 1 ♀ NMPC);
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<normalizedToken originalValue="Río">Rio</normalizedToken>
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Sardinas, R.N.F.S. Barra del Colorado, 10 m a.s.l., 11.-19.x.1993, 1 ♀, F. Araya lgt., L N 291500_564700 #2398 (INBIO);
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<normalizedToken originalValue="Río">Rio</normalizedToken>
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Sardinas, R.N.F.S. Barra del Colorado, 10 m a.s.l., 2.-12.i.1994, 1 ♂, F. Araya lgt., L N 291500_564700 #2552 (INBIO); R.V.S., Barra del Colorado, Camino a Linda Vista, 98 m a.s.l., 19.viii.2004, 1 ♂, B. Gamboa, W. Porras, D. Briceno, M. Moraga and Y.
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<normalizedToken originalValue="Cárdenas">Cardenas</normalizedToken>
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lgt., light trap, LN 284965 568835//77943 (INBIO); Sardinas, Barra del Colorado, 15 m a.s.l., 27.iii.-3.iv.1995, 2 ♂♂, F. Araya lgt., L N 291900 565900 #4414 (INBIO); Sector Cerro Cocori, Fca de E. Rojas [E. Rojas' farm;
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<geoCoordinate direction="north" orientation="latitude" precision="555" value="10.6">10.60°N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="555" value="-83.72">83.72°W</geoCoordinate>
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], xi.1990, 1 ♂, E. Rojas lgt., L-N-286000, 567500 (INBIO). Puntarenas Province: Esquipulas de Tasazu, 27 km NE of Quepos, 27.v.2003, 2 ♂♂ 1 ♀, J. C. Burne lgt. (DBTC); Est. Agujas [ca.
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="8.537998">8.537998°N</geoCoordinate>
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<geoCoordinate direction="west" orientation="longitude" precision="1" value="-83.47138">83.471381°W</geoCoordinate>
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], 300 m a.s.l., night collecting, 25.-28.x.1995, 1 ♂, A. Azofeifa lgt., L_S_276750_526550 #6373 (INBIO); Est. Quebrada Bonita, R. B. Carara [ca.
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||
<geoCoordinate direction="north" orientation="latitude" precision="15" value="9.78389">9°47'02"N</geoCoordinate>
|
||
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-84.56722">84°34'02"W</geoCoordinate>
|
||
], 80 m a.s.l., xi.1994, 1 ♂, J. C. Sobrio lgt., L N 194500_469850 #3290 (INBIO); Est. Sirena [ca.
|
||
<geoCoordinate direction="north" orientation="latitude" precision="15" value="8.543612">8°32'37"N</geoCoordinate>
|
||
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-83.52167">83°31'18"W</geoCoordinate>
|
||
], P. N. Corcovado, 0-100 m a.s.l., x.1989, 1 ♂, G. Fonseca lgt., L-S-270500, 508300 (INBIO FSCA); x.1989, 1 ♂, G. Fonseca lgt., L-S-270500, 508300 (FSCA); the same locality, x.1989, 1 ♂, C. Saborio lgt., L-S-270500, 508300 (INBIO FSCA); Est. Sirena, P. N. Corcovado, 1-100 m a.s.l., vi.1990, 1 ♂, G. Maass lgt., L-S-270500, 508300 (INBIO); Est. Sirena, P. N. Corcovado, 1-100 m a.s.l., vi.1990, 1 ♀, N. Obando lgt., L-S-270500, 508300 (INBIO); Est. Sirena, P. N. Corcovado, 1-100 m a.s.l., v.1994, 1 ♂, G. Fonseca lgt., L S 270500_508300 #2899 (INBIO); Sirena, Corcovado Nat. Pk., Osa Peninsula, 14.viii.1980, 1 ♂, H. Janzen and W. Hallwachs lgt., INBIO, CRI001 715796 (INBIO FSCA); Golfo Dulce, P.N. Corcovado, Est. Agujas [ca.
|
||
<geoCoordinate direction="north" orientation="latitude" precision="1" value="8.537998">8.537998°N</geoCoordinate>
|
||
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-83.47138">83.471381°W</geoCoordinate>
|
||
], 200-300 m a.s.l., 10.-12.x.2007, 1 ♂, J. A. Azofeifa lgt., Tp. Luz [= light trap], L_S_276750_526550 #92557 (INBIO); Lepanto,
|
||
<normalizedToken originalValue="Montaña">Montana</normalizedToken>
|
||
Grande, Estac. Karen Mogensen [ca.
|
||
<geoCoordinate direction="north" orientation="latitude" precision="15" value="9.870556">9°52'14"N</geoCoordinate>
|
||
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-85.058334">85°03'30"W</geoCoordinate>
|
||
], 320 m a.s.l., 25.-30.ix.2003, 1 ♂ 1 ♀, Y. Cardenas lgt., Tp. de Luz [= light trap], L_N_205600_420300 #75455 (INBIO); near Villa Neily [ca.
|
||
<geoCoordinate direction="north" orientation="latitude" precision="15" value="8.646389">8°38'47"N</geoCoordinate>
|
||
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-82.94556">82°56'44"W</geoCoordinate>
|
||
], 5.-11.viii.1963, 1 ♂, C. L. Hogue lgt. (DARC); Osa, Ciudad P[uer]to
|
||
<normalizedToken originalValue="Cortés">Cortes</normalizedToken>
|
||
[ca.
|
||
<geoCoordinate direction="north" orientation="latitude" precision="15" value="8.964444">8°57'52"N</geoCoordinate>
|
||
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-83.52583">83°31'33"W</geoCoordinate>
|
||
], Cuesta del Burro, 680 m a.s.l., 6.vi.2005, 1 ♂,
|
||
<pageBreakToken pageId="23" pageNumber="370" start="start">J</pageBreakToken>
|
||
. Montero, B. Gamboa, J.
|
||
<normalizedToken originalValue="Gutiérrez">Gutierrez</normalizedToken>
|
||
, M. Moraga, J. Azofeifa, Y.
|
||
<normalizedToken originalValue="Cárdenas">Cardenas</normalizedToken>
|
||
and J. Mata, Tp. Luz. [= light trap], L_S_330629_517352 #83476 (INBIO); Osa Peninsula, Sirena, Corcovado Nat. Pk., 13.viii.1980, 2 ♂♂, D. H. Janzen and W. Hallwachs lgt. (INBIO); R. Priv. Karen Mogensen, Alred.
|
||
<normalizedToken originalValue="Estación">Estacion</normalizedToken>
|
||
, 350 m a.s.l., 6.vii.2003, 1 ♀, M. A. Zumbado and W. Porras lgt., Tp. Luz. Mercurio [= mercury vapor light], L N 205600 420300 #74586 (INBIO); R.V.S.
|
||
<normalizedToken originalValue="Río">Rio</normalizedToken>
|
||
Piro, Golfito [ca.
|
||
<geoCoordinate direction="north" orientation="latitude" precision="15" value="8.619166">8°37'09"N</geoCoordinate>
|
||
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-83.14444">83°08'40"W</geoCoordinate>
|
||
], Finca Catalino, 200 m a.s.l., 16.ix.2004, 1 ♀, Y. Cardenas, D.
|
||
<normalizedToken originalValue="Briseño">Briseno</normalizedToken>
|
||
and B. Gamboa, Luz. [= light trap], L_S_264550_535590 #78215 (INBIO); Rancho Quemado [ca.
|
||
<geoCoordinate direction="north" orientation="latitude" precision="15" value="8.681945">8°40'55"N</geoCoordinate>
|
||
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-83.56056">83°33'38"W</geoCoordinate>
|
||
], Peninsula de Osa, x.1990, 6 ♂♂ 13 ♀♀, F. Quesada lgt., L-S-292500, 511000 (1 ♂ 5 ♀♀ FSCA, 2 ♂♂ 5 ♀♀ INBIO, 1 ♀ BMNH, 1 ♂ 1 ♀ HNHM, 2 ♂♂ 1 ♀ NMPC); Rancho Quemado, Pen.[insula de] Osa, x.-xi.1990, 1 ♀, B. Apu lgt., L-S-292500, 511000 (INBIO); Rancho Quemado, Pen.[insula de] Osa, i.1991, 3 ♂♂ 2 ♀♀, F. Quesada lgt., L-S-292500, 511000 (INBIO); Rancho Quemado, Pen.[insula de] Osa, iv.1991, 1 ♂, J. C. Saborio lgt., L-S-292500, 511000 (INBIO); Sirena, Corcovado Nat. Pk., Osa Penin.[sula], 5.-11.i.1981, 1 ♀, D. H. Janzen and W. Hallwachs lgt. (INBIO). San
|
||
<normalizedToken originalValue="José">Jose</normalizedToken>
|
||
Province: Est. Bijagual [ca.
|
||
<geoCoordinate direction="north" orientation="latitude" precision="15" value="9.728611">9°43'43"N</geoCoordinate>
|
||
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-84.56917">84°34'09"W</geoCoordinate>
|
||
], Res. Biol. Carara, 500 m a.s.l., ix.1990, 1 ♂, G. Varela lgt., L-N-192250, 474760 (INBIO). - PANAMA: Bocas del Toro Province: 12 km W Chiriqui Grande [ca.
|
||
<geoCoordinate direction="north" orientation="latitude" precision="15" value="8.992223">8°59'32"N</geoCoordinate>
|
||
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-82.25361">82°15'13"W</geoCoordinate>
|
||
], 10.-13.v.1999, 2 ♂♂, Morris and Wappes lgt. (JEEC).
|
||
<normalizedToken originalValue="Chiriquí">Chiriqui</normalizedToken>
|
||
Province: Finca La Suiza, 5.3 km N Los Planes,
|
||
<geoCoordinate direction="north" orientation="latitude" precision="925" value="8.65">8°39'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-82.2">82°12'W</geoCoordinate>
|
||
, 4500' [= 1372 m a.s.l.], 26.-30.v.1995, 1 ♂, B. Ratcliffe and M. Jameson lgt. (DBTC).
|
||
<normalizedToken originalValue="Coclé">Cocle</normalizedToken>
|
||
Province: Rio Indio Lodge, N El Valle,
|
||
<geoCoordinate direction="north" orientation="latitude" precision="1" value="8.6629715">8°39'46.7"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-80.118866">80°7'7.9"W</geoCoordinate>
|
||
, 575 m a.s.l., 23.-27.ii.2012, 1 ♀, J. B. Heppner lgt. (FSCA).
|
||
<normalizedToken originalValue="Colón">Colon</normalizedToken>
|
||
Province: Pipeline Road, km 0, 9°7'19"N, 79°42'53"W, beating of forest vegetation along strips for boat navigation signs, 31.viii.2010, 1 ♂ 2 ♀♀, L. Sekerka lgt. (MMBC).
|
||
<normalizedToken originalValue="Panamá">Panama</normalizedToken>
|
||
Province: Campana [= Cerro Campana], 2785 ft [= 849 m a.s.l.],
|
||
<geoCoordinate direction="north" orientation="latitude" precision="1" value="8.682">8°40.920'N</geoCoordinate>
|
||
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-79.92885">79°55.731'W</geoCoordinate>
|
||
, 28.v.-9.vi.2008, 1 ♀, D. C. Robacker lgt. (DBTC); 10-12 km N El Llano, 3.-8.vi.1986, 1 ♀, E. Giesbert lgt. (FSCA); El
|
||
<normalizedToken originalValue="Llano–Carti">Llano-Carti</normalizedToken>
|
||
Rd. [= road], km 10-13 [ca.
|
||
<geoCoordinate direction="north" orientation="latitude" precision="15" value="9.293612">9°17'37"N</geoCoordinate>
|
||
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-78.978615">78°58'43"W</geoCoordinate>
|
||
], 3.-7.vi.1984, 1 ♀, R. L. Penrose, F. T. Hovore and P. H. Sullivan lgt. (DBTC); K[m] 8-13 El
|
||
<normalizedToken originalValue="Llano–Carti">Llano-Carti</normalizedToken>
|
||
Rd. [= road], 10.-13.v.1996, 2 ♂♂ 1 ♀, Wappes, Huether and Morris lgt. (1 ♂ 1 ♀ DBTC, 1 ♂ JEEC); PN Chagres, Cerro Jeffe,
|
||
<geoCoordinate direction="north" orientation="latitude" precision="92" value="9.238334">9°14.3' N</geoCoordinate>
|
||
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-79.401665">79°24.1' W</geoCoordinate>
|
||
, 700-950 m a.s.l., lower montane forest; individual collecting, 18.-19.viii.2017, 2 ♂♂, M. Seidel and L. Sekerka lgt. (NMPC). San Blas Province: Nusagandi [
|
||
<geoCoordinate direction="north" orientation="latitude" precision="5" value="9.3489">9.3489°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="west" orientation="longitude" precision="55" value="-78.966">78.966°W</geoCoordinate>
|
||
], 18.-20.v.1993, UV light, 3 ♂♂, E. Riley lgt. (2 ♂♂ DARC, 1 ♂ DBTC); Nusagandi, at light, 250-350 m a.s.l., 26.vii.1995, 1 ♂ 3 ♀♀, C. W. and L. B.
|
||
<normalizedToken originalValue="O’Brien">O'Brien</normalizedToken>
|
||
lgt. (JEEC); San Blas, Punta Eseoses [=
|
||
<normalizedToken originalValue="Escocés">Escoces</normalizedToken>
|
||
;
|
||
<geoCoordinate direction="north" orientation="latitude" precision="555" value="8.85">8.85°N</geoCoordinate>
|
||
<geoCoordinate direction="west" orientation="longitude" precision="5" value="-77.6333">77.6333°W</geoCoordinate>
|
||
], Lt. Trap, ii.-iii.1979, 1 ♂ 1 ♀, Caroline Ash lgt., H. D. Engleman det. as
|
||
<taxonomicName class="Insecta" family="Pentatomidae" genus="Rhyncholepta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Rhyncholepta grandicallosa" order="Hemiptera" pageId="23" pageNumber="370" phylum="Arthropoda" rank="species" species="grandicallosa">Rhyncholepta grandicallosa</taxonomicName>
|
||
(JEEC). Veraguas Province: Santa
|
||
<normalizedToken originalValue="Fé">Fe</normalizedToken>
|
||
env., Cascada Alto de Piedra,
|
||
<geoCoordinate direction="north" orientation="latitude" precision="92" value="8.516666">8°31.0'N</geoCoordinate>
|
||
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-81.121666">81°07.3'W</geoCoordinate>
|
||
, 830 m a.s.l., lower montane forest, individual collecting, 12.ix.2017, 1 ♂, J.
|
||
<normalizedToken originalValue="Hájek">Hajek</normalizedToken>
|
||
lgt. (NMPC). - All the paratypes are bearing the following identification label: "PARATYPUS / RHYNCHOLEPTA / grandicallosa subsp. / CENTROAMERICANA nov. / det. Kment, Eger, Rider 2017 [printed, yellow label]".
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="29" lastPageNumber="376" pageId="24" pageNumber="371" type="diagnosis">
|
||
<paragraph pageId="24" pageNumber="371">
|
||
<pageBreakToken pageId="24" pageNumber="371" start="start">Diagnosis</pageBreakToken>
|
||
.
|
||
</paragraph>
|
||
<paragraph pageId="24" pageNumber="371">
|
||
Coloration, structure of head, thorax and pregenital abdomen, and vestiture as in other species of the genus (see redescription of
|
||
<taxonomicName class="Insecta" family="Pentatomidae" genus="Rhyncholepta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Rhyncholepta" order="Hemiptera" pageId="24" pageNumber="371" phylum="Arthropoda" rank="genus">Rhyncholepta</taxonomicName>
|
||
above) except the following characters.
|
||
</paragraph>
|
||
<paragraph lastPageId="25" lastPageNumber="372" pageId="24" pageNumber="371">
|
||
Apex of scutellum with anteapical black V-shaped stripe usually well developed, wide (Figure 20) to narrow (Figs 21-22), less frequently reduced to small black spot
|
||
<pageBreakToken pageId="25" pageNumber="372" start="start">on</pageBreakToken>
|
||
each lateral margin at anterior end of apical V-shaped callosity (Figure 19); all color forms can be syntopic (e.g. Costa Rica, Rio
|
||
<normalizedToken originalValue="Sárdinas">Sardinas</normalizedToken>
|
||
, INBIO). Apical callosity V-shaped, robust, branches of V forming ca. one third to one half of width, tip of scutellum with distinct triangular callosity (Figs 19-22).
|
||
</paragraph>
|
||
<paragraph lastPageId="28" lastPageNumber="375" pageId="25" pageNumber="372">
|
||
Male genitalia. Genital capsule in ventral view more or less constricted lateroapically (Figs 32: arrow, 34: arrow, 56: arrow, 58: arrow), posterolateral angles prominent, ca.
|
||
<pageBreakToken pageId="26" pageNumber="373" start="start">rectangular</pageBreakToken>
|
||
(Figs 32, 34, 56, 58); dorsal wall at base of posterolateral angles shallowly to deeply impressed (56: arrow, 58: arrow, 67-68). Ventral rim in ventral view bilobed apically, with shallow V-shaped notch medially (Figs 32-35); hypandrial projections not visible in ventral view (Figs 32-35). Hypandrium in posterior view with pair of large lobe-like anterior projections, apices appearing acute (Figs 44, 45: ap, 46, 47: ap) and short-pointed posterior projections directed posterolaterad (Figs 45: pp, 47: pp); lateral projections not visible in ventral view but site of their attachment apparent as obtuse angle laterally on anterior projections (Figure 45: lbp, 47: lbp). Anterior
|
||
<pageBreakToken pageId="27" pageNumber="374" start="start">hypandrial</pageBreakToken>
|
||
projections in most exposed (dorso-posterolateral) view smaller than in
|
||
<taxonomicName genus="Rh." lsidName="Rh. grandicallosa" pageId="27" pageNumber="374" rank="subSpecies" species="grandicallosa" subSpecies="grandicallosa">Rh. grandicallosa grandicallosa</taxonomicName>
|
||
, parabolic, with narrowly rounded apex (Figs 73: ap, 74: ap). In dorsal view, apices of anterior hypandrial projections directed anterodorsally, median outline straight (Figs 56, 57: ap, 58, 59: ap; in
|
||
<taxonomicName genus="Rh." lsidName="Rh. g." pageId="27" pageNumber="374" rank="subSpecies" species="g." subSpecies="grandicallosa">Rh. g. grandicallosa</taxonomicName>
|
||
convex); lateral hypandrial projections long (appearing longer than in
|
||
<taxonomicName genus="Rh." lsidName="Rh. g." pageId="27" pageNumber="374" rank="subSpecies" species="g." subSpecies="grandicallosa">Rh. g. grandicallosa</taxonomicName>
|
||
), golf-club "handles" nearly parallel, suddenly curved inwards (ca. in right angle) apically (Figs 57:
|
||
<pageBreakToken pageId="28" pageNumber="375" start="start">lp</pageBreakToken>
|
||
, 59: lp, 73: lp, 74: lp). Phallus (Figs 81-83) conjunctival sclerites (Figure 82: cjs) and aedeagus strongly S-shaped as in
|
||
<taxonomicName genus="Rh." lsidName="Rh. g." pageId="28" pageNumber="375" rank="subSpecies" species="g." subSpecies="grandicallosa">Rh. g. grandicallosa</taxonomicName>
|
||
(Figure 79;
|
||
<bibRefCitation author="Becker, M" journalOrPublisher="Revista Brasileira de Biologia" pageId="41" pageNumber="388" pagination="389 - 399" title="Sobre o genero Rhyncholepta Bergroth, 1911, com a descricao de uma nova especie (Hemiptera, Pentatomidae, Pentatominae)." volume="31" year="1971">Becker and Grazia-Vieira 1971</bibRefCitation>
|
||
: fig. 10).
|
||
</paragraph>
|
||
<paragraph pageId="29" pageNumber="376">
|
||
<pageBreakToken pageId="29" pageNumber="376" start="start">Female</pageBreakToken>
|
||
genitalia. Posterior edges of laterotergites VIII posteriorly as long as or slightly more prominent compared with laterotergites IX (Figure 91) (within variation of
|
||
<taxonomicName genus="Rh." lsidName="Rh. g." pageId="29" pageNumber="376" rank="subSpecies" species="g." subSpecies="grandicallosa">Rh. g. grandicallosa</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="29" pageNumber="376">Measurements. Table 1. Measurements of holotype (in mm): Body length 13.19, body length to segment VII 11.64, head length 2.56, head width 2.41, interocular width 1.11, length of antennomeres: I - 0.84, IIa - 1.43, IIb - 2.73, III - 3.44, IV - 2.73, pronotum length 2.71, pronotum width 7.88, scutellum length 4.65, scutellum width 3.83.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="31" lastPageNumber="378" pageId="29" pageNumber="376" type="variability">
|
||
<paragraph pageId="29" pageNumber="376">Variability.</paragraph>
|
||
<paragraph lastPageId="31" lastPageNumber="378" pageId="29" pageNumber="376">
|
||
Besides the usual variation in coloration and structure within
|
||
<taxonomicName class="Insecta" family="Pentatomidae" genus="Rhyncholepta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Rhyncholepta" order="Hemiptera" pageId="29" pageNumber="376" phylum="Arthropoda" rank="genus">Rhyncholepta</taxonomicName>
|
||
species, we observed some variability of
|
||
<taxonomicName genus="Rh." lsidName="Rh. g." pageId="29" pageNumber="376" rank="subSpecies" species="g." subSpecies="centroamericana">Rh. g. centroamericana</taxonomicName>
|
||
in the structure of the genital capsule and hypandrium, as illustrated in two males. One is from Panama, Pipeline Road (Figs 32-33, 44-45, 56-57, 67, 73; namely genital capsule slightly constricted posterolaterally [Figs 32, 56], anterior hypandrial projection longer, with apex
|
||
<pageBreakToken pageId="30" pageNumber="377" start="start">more</pageBreakToken>
|
||
prominent, and median margin slightly less concave medially [Figs 45: ap, 57: ap, 73: ap]). The second male is from Costa Rica, Rancho Quemado (Figs 34-35, 46-47, 58-59, 68, 74; namely genital capsule hardly constricted posterolaterally [Figs 34, 58], anterior hypandrial projection shorter, with apex less prominent, and median margin slightly more concave medially [Figs 47: ap, 59: ap, 74: ap]). The posterolateral constric
|
||
<pageBreakToken pageId="31" pageNumber="378" start="start">tion</pageBreakToken>
|
||
of the genital capsule in the specimen from Panama, Pipeline Road (Figs 32, 56: arrow) is somewhat intermediate between the typical
|
||
<taxonomicName genus="Rh." lsidName="Rh. g." pageId="31" pageNumber="378" rank="subSpecies" species="g." subSpecies="grandicallosa">Rh. g. grandicallosa</taxonomicName>
|
||
(see Figs 30, 54: arrow) and the population from Costa Rica, Rancho Quemado (Figs 34, 58: arrow).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="31" pageNumber="378" type="etymology">
|
||
<paragraph pageId="31" pageNumber="378">Etymology.</paragraph>
|
||
<paragraph pageId="31" pageNumber="378">The subspecies name is a Latin adjective centroamericanus (-a, -um) referring to its Central American distribution.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="32" pageNumber="379" type="bionomics">
|
||
<paragraph pageId="32" pageNumber="379">
|
||
<pageBreakToken pageId="32" pageNumber="379" start="start">Bionomics</pageBreakToken>
|
||
.
|
||
</paragraph>
|
||
<paragraph pageId="32" pageNumber="379">
|
||
Most specimens were collected by various types of light traps (UV light, mercury vapor light, metal halide light, black light) in or adjacent to dense forests. According to JE Eger, it has never been collected by hand catching, sweeping, or beating vegetation during the day or night. In Panama, three specimens were collected by "beating of forest vegetation along strips for boat navigation signs" (J Sekerka, pers. comm.) and one in "lower montane forest, [by] individual collecting" (J
|
||
<normalizedToken originalValue="Hájek">Hajek</normalizedToken>
|
||
, pers. comm.) but without detailed information. Collection dates indicate that
|
||
<taxonomicName genus="Rh." lsidName="Rh. g." pageId="32" pageNumber="379" rank="subSpecies" species="g." subSpecies="centroamericana">Rh. g. centroamericana</taxonomicName>
|
||
occurs year round, though most specimens were collected between May and October (Figure 95) (
|
||
<bibRefCitation author="Becker, M" journalOrPublisher="Revista Brasileira de Biologia" pageId="41" pageNumber="388" pagination="389 - 399" title="Sobre o genero Rhyncholepta Bergroth, 1911, com a descricao de uma nova especie (Hemiptera, Pentatomidae, Pentatominae)." volume="31" year="1971">Becker and Grazia-Vieira 1971</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Arismendi, N" journalOrPublisher="Insecta Mundi" pageId="41" pageNumber="388" pagination="218 - 236" title="Pentatomidae (Heteroptera) of Honduras: a checklist with description of a new ochlerine genus." volume="17" year="2003">Arismendi and Thomas 2003</bibRefCitation>
|
||
; present paper).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="33" pageNumber="380" type="distribution">
|
||
<paragraph pageId="33" pageNumber="380">
|
||
<pageBreakToken pageId="33" pageNumber="380" start="start">Distribution</pageBreakToken>
|
||
</paragraph>
|
||
<paragraph pageId="33" pageNumber="380">
|
||
(Figure 98). Mexico: Chiapas (new country record); Belize (new country record); Guatemala (new country record); Honduras (
|
||
<bibRefCitation pageId="33" pageNumber="380">Arismendi 2002</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Arismendi, N" journalOrPublisher="Insecta Mundi" pageId="41" pageNumber="388" pagination="218 - 236" title="Pentatomidae (Heteroptera) of Honduras: a checklist with description of a new ochlerine genus." volume="17" year="2003">Arismendi and Thomas 2003</bibRefCitation>
|
||
, both as
|
||
<taxonomicName genus="Rh." lsidName="Rh. grandicallosa" pageId="33" pageNumber="380" rank="species" species="grandicallosa">Rh. grandicallosa</taxonomicName>
|
||
); Costa Rica (
|
||
<bibRefCitation pageId="33" pageNumber="380">Arismendi 2002</bibRefCitation>
|
||
, as
|
||
<taxonomicName genus="Rh." lsidName="Rh. grandicallosa" pageId="33" pageNumber="380" rank="species" species="grandicallosa">Rh. grandicallosa</taxonomicName>
|
||
); Panama (
|
||
<bibRefCitation author="Becker, M" journalOrPublisher="Revista Brasileira de Biologia" pageId="41" pageNumber="388" pagination="389 - 399" title="Sobre o genero Rhyncholepta Bergroth, 1911, com a descricao de uma nova especie (Hemiptera, Pentatomidae, Pentatominae)." volume="31" year="1971">Becker and Grazia-Vieira 1971</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Froeschner, RC" journalOrPublisher="Memoirs of the American Entomological Institute" pageId="42" pageNumber="389" pagination="1 - 393" title="True bugs (Heteroptera) of Panama: A synoptic catalog as a contribution to the study of Panamanian biodiversity." volume="61" year="1999">Froeschner 1999</bibRefCitation>
|
||
,
|
||
<bibRefCitation pageId="33" pageNumber="380">Arismendi 2002</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Cambra, RA" journalOrPublisher="Revista Nicaragueense de Entomologia" pageId="41" pageNumber="388" pagination="1 - 21" title="Los Pentatomidos (Hemiptera: Heteroptera) de Panama." volume="149" year="2018">Cambra et al. 2018</bibRefCitation>
|
||
, all as
|
||
<taxonomicName genus="Rh." lsidName="Rh. grandicallosa" pageId="33" pageNumber="380" rank="species" species="grandicallosa">Rh. grandicallosa</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="33" pageNumber="380">
|
||
The subspecific identity of
|
||
<taxonomicName genus="Rh." lsidName="Rh. grandicallosa" pageId="33" pageNumber="380" rank="species" species="grandicallosa">Rh. grandicallosa</taxonomicName>
|
||
population from
|
||
<normalizedToken originalValue="Chocó">Choco</normalizedToken>
|
||
, Colombia requires revision.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |