157 lines
22 KiB
XML
157 lines
22 KiB
XML
<document id="179D8ACA14D974776A556E497934C6DA" ID-DOI="10.11646/zootaxa.2457.1.1" ID-ISSN="11755334" ID-Zenodo-Dep="5313709" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629674756648" checkinUser="felipe" docAuthor="Oláh, János & Johanson, Kjell Arne" docDate="2010" docId="2B7E87E4FF94FFACF5A6FE2AFC37F9EB" docLanguage="en" docName="zt02457p128.pdf" docOrigin="Zootaxa 2457 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2457.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Anisocentropus (Anisomontropus) bungus Oláh & Johanson 2010, new species" docType="treatment" docVersion="3" lastPageNumber="60" masterDocId="D747FF9CFFADFF97F531FFFDFFD9FFE9" masterDocTitle="Description of 33 new species of Calamoceratidae, Molannidae, Odontoceridae and Philorheithridae (Trichoptera), with detailed presentation of their cephalic setal warts and grooves 2457" masterLastPageNumber="128" masterPageNumber="1" pageNumber="58" updateTime="1699094080944" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="1D6ADE86B8D0315F048C7D7D70707110">Description of 33 new species of Calamoceratidae, Molannidae, Odontoceridae and Philorheithridae (Trichoptera), with detailed presentation of their cephalic setal warts and grooves 2457</mods:title>
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<mods:namePart id="033C48C9C67E665DDF0F38A5B351D602">Oláh, János</mods:namePart>
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<mods:namePart id="DA8F41449DF513FE67378CA9EF1C2BAB">Johanson, Kjell Arne</mods:namePart>
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<mods:title id="830BEFB45BC041D21118369F23AB581C">Zootaxa</mods:title>
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<mods:part id="0F0C70F6A950907842D8D62CF58DF4C0">
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<mods:date id="DCCD9372AC1FC053DD2C5602172BFFBE">2010</mods:date>
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<mods:number id="FB9EFC2301FE17D9CDB13152522A38ED">2010-05-14</mods:number>
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<mods:number id="04974A58E6AD3B67761B5DEA922FC895">2457</mods:number>
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<mods:classification id="F13BF9F059B2C440A27F9449ECC9904B">journal article</mods:classification>
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<mods:identifier id="5BED8F635F711E77068B8D9E760B4BDF" type="DOI">10.11646/zootaxa.2457.1.1</mods:identifier>
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<subSubSection id="EBCD6579FF94FFAEF5A6FE2AFCEFFE18" box="[151,822,471,497]" pageId="57" pageNumber="58" type="nomenclature">
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<paragraph id="A36836F2FF94FFAEF5A6FE2AFCEFFE18" blockId="57.[151,822,471,532]" box="[151,822,471,497]" pageId="57" pageNumber="58">
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<heading id="F820819EFF94FFAEF5A6FE2AFCEFFE18" bold="true" box="[151,822,471,497]" fontSize="11" level="1" pageId="57" pageNumber="58" reason="1">
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<emphasis id="91A3EAE0FF94FFAEF5A6FE2AFCEFFE18" bold="true" box="[151,822,471,497]" pageId="57" pageNumber="58">
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<taxonomicName id="64D74D71FF94FFAEF5A6FE2AFD41FE18" authority="Oláh & Johanson, 2010" authorityName="Oláh & Johanson" authorityYear="2010" box="[151,664,471,497]" class="Insecta" family="Calamoceratidae" genus="Anisocentropus" kingdom="Animalia" order="Trichoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="bungus" status="sp. nov." subGenus="Anisomontropus">
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<emphasis id="91A3EAE0FF94FFAEF5A6FE2AFE8DFE18" bold="true" box="[151,340,471,497]" italics="true" pageId="57" pageNumber="58">Anisocentropus</emphasis>
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(
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<emphasis id="91A3EAE0FF94FFAEF454FE2AFDF4FE18" bold="true" box="[357,557,471,497]" italics="true" pageId="57" pageNumber="58">Anisomontropus</emphasis>
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)
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<emphasis id="91A3EAE0FF94FFAEF70FFE2AFD41FE18" bold="true" box="[574,664,471,497]" italics="true" pageId="57" pageNumber="58">bungus</emphasis>
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</taxonomicName>
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<emphasis id="91A3EAE0FF94FFAEF7A8FE2AFD46FE18" bold="true" box="[665,671,471,497]" italics="true" pageId="57" pageNumber="58">,</emphasis>
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<taxonomicNameLabel id="8A90579BFF94FFAEF797FE2AFCEFFE18" box="[678,822,471,497]" pageId="57" pageNumber="58" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="EBCD6579FF94FFACF5A6FE07FE2DFA8B" lastPageId="59" lastPageNumber="60" pageId="57" pageNumber="58" type="description">
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<paragraph id="A36836F2FF94FFAEF5A6FE07FEEFFDFD" blockId="57.[151,822,471,532]" box="[151,310,506,532]" pageId="57" pageNumber="58">
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<figureCitation id="3BEC2A77FF94FFAEF5A6FE07FEEFFDFD" box="[151,310,506,532]" captionStart="FIGURES 117–122" captionStartId="58.[151,269,1900,1924]" captionTargetBox="[165,1425,207,1849]" captionTargetId="figure-50@58.[159,1428,197,1873]" captionTargetPageId="58" captionText="FIGURES 117–122. Anisocentropus bungus, new species, holotype. 117 — head, frontal; 118 — genitalia, lateral; 119 — genitalia, dorsal; 120 — genitalia, ventral; 121 — phallus, lateral; 122 — phallus, ventral." figureDoi="http://doi.org/10.5281/zenodo.5313773" httpUri="https://zenodo.org/record/5313773/files/figure.png" pageId="57" pageNumber="58">Figs 117–122</figureCitation>
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</paragraph>
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<paragraph id="A36836F2FF94FFAEF5A6FDB7FB9DFCC5" blockId="57.[151,1437,586,2012]" pageId="57" pageNumber="58">
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This large, brown species is the only one in the subgenus without pattern on the forewing membrane. The genitalia resemble those of
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<taxonomicName id="64D74D71FF94FFAEF483FD8FFDCBFD62" authorityName="Malicky & Chantaramongkol" authorityYear="1994" box="[434,530,626,651]" class="Insecta" family="Calamoceratidae" genus="Anisocentropus" kingdom="Animalia" order="Trichoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="janus">
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<emphasis id="91A3EAE0FF94FFAEF483FD8FFDCBFD62" box="[434,530,626,651]" italics="true" pageId="57" pageNumber="58">A. janus</emphasis>
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</taxonomicName>
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from which it is separated by the more slender gonocoxites; and it has more regularly arched gonocoxites in lateral view, while the arch in
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<taxonomicName id="64D74D71FF94FFAEF65FFD67FC17FD5A" authorityName="Malicky & Chantaramongkol" authorityYear="1994" box="[878,974,666,691]" class="Insecta" family="Calamoceratidae" genus="Anisocentropus" kingdom="Animalia" order="Trichoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="janus">
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<emphasis id="91A3EAE0FF94FFAEF65FFD67FC17FD5A" box="[878,974,666,691]" italics="true" pageId="57" pageNumber="58">A. janus</emphasis>
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</taxonomicName>
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is dorsally flat; the preanal appendages are more robust and more strongly constricted at midlength in dorsal view; segment X has a broader dorsal interlobular gap with more rounded and slightly laterad directed lobes in dorsal view, while straight and tapering in
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<taxonomicName id="64D74D71FF94FFAEF5D3FCEFFE98FCC2" authorityName="Malicky & Chantaramongkol" authorityYear="1994" box="[226,321,786,811]" class="Insecta" family="Calamoceratidae" genus="Anisocentropus" kingdom="Animalia" order="Trichoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="janus">
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<emphasis id="91A3EAE0FF94FFAEF5D3FCEFFE98FCC2" box="[226,321,786,811]" italics="true" pageId="57" pageNumber="58">A. janus</emphasis>
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</taxonomicName>
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; in lateral view the dorsal ridge is more elevated than in
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<taxonomicName id="64D74D71FF94FFAEF6ECFCEFFBE4FCC2" authorityName="Malicky & Chantaramongkol" authorityYear="1994" box="[989,1085,786,811]" class="Insecta" family="Calamoceratidae" genus="Anisocentropus" kingdom="Animalia" order="Trichoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="janus">
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<emphasis id="91A3EAE0FF94FFAEF6ECFCEFFBE4FCC2" box="[989,1085,786,811]" italics="true" pageId="57" pageNumber="58">A. janus</emphasis>
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="A36836F2FF94FFACF5F6FCC7FEFAFD7B" blockId="57.[151,1437,586,2012]" lastBlockId="59.[151,1437,152,1538]" lastPageId="59" lastPageNumber="60" pageId="57" pageNumber="58">
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Male (in alcohol). Body large; body and forewings brown. Head rectangular in dorsal and facial view, almost as long as broad. Ocelli absent. Tentorium not visible. Facial groove pattern forming plate-like flange, or rim, connecting anterior tentorial pits. Theoretical line separating frons and clypeus forming oblique, dorsad continuation from anterior tentorial pits; almost horizontally merging with antennal grooves. Clypeogenal vertical grooves located ventrally of anterior tentorial pits, short, running slightly mesoventrad; plate-like flanges, and frontogenal and clypeogenal sutures forming complex of anterior tentorial arms (frontogenal septa). Small subantennal grooves horizontal, sinuous, located between frontogenal compact setal wart and palpifers, forming proximal articulation of palps on stipes. Subocular grooves invisible. Frontal groove pronoumced, short, present between antennae, joining vertexal medioantennal compact setose warts. Vertexal groove pattern reduced. Large, anterad directed, rounded elevation dominating on anterior half of vertex; reaching interantennal area; laterally delineated by antennal sockets; with corrugated or granulous surface near frontogenal compact setose wart. Epicranial groove (coronal groove) with vestigial stem. Antennal and ocular grooves accompanied by occipito-postgenal grooves, partly merging anteriorly; tangential to large occipital compact setal warts. Labrum vertically long, subquadrangular with rounded apex, freely hanging, sparsely setose. Mandible long, turning mesally, weakly pigmented, oriented laterad along labrum; lacinia broad, elongated, bearing few setae. Frontal setal warts absent; frontal interantennal warts absent due to anterad directed vertexal elevation. Pair of large, nearly triangular, frontogenal compact setal warts present on posterior pregenae, obliquely or almost horizontally from frontogenal grooves, representing only visible setal warts on face. Triangular elevation on vertex with fused vertexal medioantennal compact setose warts located anteriorly on head. Vertexal lateroantennal compact setal warts absent. Antennal sockets large; pair of small, rounded vertexal ocellar compact setose warts and equally large to larger pair vertexal medioocellar compact setose warts visible in middle of vertex. Pair of large, obliquely located, ovoid occipital compact setose warts dominating on posterior half of vertex. Pair of small postgenal compact setose warts visible between posterior section of ocular grooves and occipital compact setose warts. Maxillary palps filiform, covered by long setae; maxillary palp formula VI-IV-(II, V)-I-III. Scapes rounded, about half as long as head; pedicels 2/5ths as long as scapes. Two pairs pronotal warts present: 1 pair of large dorsal transverse ovoid, widely separated mesally with deep cleft; 1 pair small, rounded warts located deep laterally, visible in lateral view. Pair of mesoscutal warts arranged in longitudinal line running long along entire mesoscutum of bright yellow colour, almost indiscernible. Pair of mesoscutellar warts forming irregular, less pigmented area with 5 to 6 setal alveoli. Proepisternum with large, rounded setose wart located proximally above articulation of cervical sclerite; larger than setal wart on precoxale. Large, compact setal warts present mainly on membranous part of cervix, touching anterior arm of cervical sclerites. Lateral cervical sclerites composed of narrow anterior arm articulating anteriorly to back of head with occipital condyle above posterior tentorial pits, fused to posterior cervical sclerites. Posterior cervical sclerites forming narrow, elongated plates reaching prothoracic episternum, articulating with weakly sclerotized anteromedian band of prothoracic eusternum by thin, ventral intercervical sclerites. Legs with symmetrical claws; spur formula 2, 4, 2; each foreleg with posteroapical spur 2 times longer than anteroapical spur; midleg anteroapical and anterosubapical spurs 1/4th as long as other spurs; hind leg apical spurs equally long. Forewings: length 12.0 mm; membrane brown, without pattern; Sc hypertrophied with crossvein
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<emphasis id="91A3EAE0FF96FFACF4A6FE25FE1AFE18" box="[407,451,472,497]" italics="true" pageId="59" pageNumber="60">sc-r</emphasis>
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, forming deep furrow running along costal margin of wing; R1 confluent with, or recurrent into, R2 well before C, without visible continuation to C; base of discoidal cell located proximally of mid-length of wing; forks I, II, III, IV, V present; crossveins h,
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<emphasis id="91A3EAE0FF96FFACF645FDD5FB15FDA8" box="[884,1228,552,577]" italics="true" pageId="59" pageNumber="60">sc-r, s, r-m, m, m-cu, cu1, cu2</emphasis>
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and
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<emphasis id="91A3EAE0FF96FFACF035FDD5FAE1FDA8" box="[1284,1336,552,577]" italics="true" pageId="59" pageNumber="60">cu-a</emphasis>
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present; crossvein
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<emphasis id="91A3EAE0FF96FFACF43CFDADFEC1FD80" box="[269,280,592,617]" italics="true" pageId="59" pageNumber="60">r</emphasis>
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absent; postanal vein absent. Hind wings: R1 meeting R2; forks I, II, III, V present; fork I longer than fork II.
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</paragraph>
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<caption id="F7A8667AFF97FFADF5A6F891FBA0F84F" ID-DOI="http://doi.org/10.5281/zenodo.5313773" ID-Zenodo-Dep="5313773" httpUri="https://zenodo.org/record/5313773/files/figure.png" pageId="58" pageNumber="59" startId="58.[151,269,1900,1924]" targetBox="[165,1425,207,1849]" targetPageId="58">
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<paragraph id="A36836F2FF97FFADF5A6F891FBA0F84F" blockId="58.[151,1436,1900,1958]" pageId="58" pageNumber="59">
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<emphasis id="91A3EAE0FF97FFADF5A6F891FEAEF86D" bold="true" box="[151,375,1900,1924]" pageId="58" pageNumber="59">FIGURES 117–122.</emphasis>
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<taxonomicName id="64D74D71FF97FFADF44EF891FDA1F86A" authority="Oláh & Johanson, 2010" authorityName="Oláh & Johanson" authorityYear="2010" box="[383,632,1900,1923]" class="Insecta" family="Calamoceratidae" genus="Anisocentropus" kingdom="Animalia" order="Trichoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="bungus" status="sp. nov.">
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<emphasis id="91A3EAE0FF97FFADF44EF891FDA1F86A" box="[383,632,1900,1923]" italics="true" pageId="58" pageNumber="59">Anisocentropus bungus</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="8A90579BFF97FFADF7B7F891FCD3F86A" box="[646,778,1900,1923]" pageId="58" pageNumber="59" rank="species">new species</taxonomicNameLabel>
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, holotype. 117 — head, frontal; 118 — genitalia, lateral; 119 — genitalia, dorsal; 120 — genitalia, ventral; 121 — phallus, lateral; 122 — phallus, ventral.
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</paragraph>
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</caption>
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<paragraph id="A36836F2FF96FFACF5F6FD5DFE2DFA8B" blockId="59.[151,1437,152,1538]" pageId="59" pageNumber="60">Male genitalia. Abdominal segment IX fused annularly, without longitudinal groove separating dorsal, mesal and ventral parts; tergum shorter than venter; anterior margins of segment IX ventrally convex in lateral view, more developed ventrally; posterior margin forming large, regular circular apical lobe. Antecosta weakly developed, forming narrow, marginal rim, equally thin along margin, without conspicuous external groove of antecostal suture; tergum IX with narrow triangular mesal keel in dorsal view, forming continuation of sharp dorsal keel of segment X, characterised by lateral triangular projections; in dorsal view, tergite with rounded mesal lobe on anterior margin; similar excision present on anterior margins of venter IX; spine row on posterior lateral margins forming setal patches on almost entire apical lobe areas. Intersegmental depression between segment IX and segment X filled. Segment X about as long as gonocoxites, forming broad hood with blunt apex, sclerotized band running from apex to base of gonocoxites, near ventral margin. Apicoventral setose lobes forming setose surfaces before and at apex. Apicodorsal setose lobes with few tiny setae above middle of segment X. Dorsal interlobular gap broad, deeply triangular. Preanal appendages much longer than segment X; almost parallel-sided, slightly arching in lateral view; more robust and broader in dorsal view; constriction present at middle. Gonocoxites without harpago; slender, sinuous, broadening mesally in ventral view; forming regular arch in lateral view. Phallic apparatus straight in lateral and ventral view; with elongating, sclerotized apicoventral lobe, endotheca and phallicata membranous above apicoventral lobe. Phallotremal sclerites U-shaped in ventral view; apices curving mesad, tapering. Ejaculatory duct sinuous in lateral view; straight in ventral view.</paragraph>
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</subSubSection>
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<subSubSection id="EBCD6579FF96FFACF5F6FA92FF32FA5B" pageId="59" pageNumber="60" type="materials_examined">
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<paragraph id="A36836F2FF96FFACF5F6FA92FF32FA5B" blockId="59.[151,1437,152,1538]" pageId="59" pageNumber="60">
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<materialsCitation id="13BF3CAFFF96FFACF5F6FA92FF32FA5B" collectingDate="1991-01-10" collectionCode="OPC" collectorName="S. Andrikovics" country="Vietnam" location="Nghta Bihn Province" municipality="Bung" pageId="59" pageNumber="60" specimenCount="1" typeStatus="holotype">
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<emphasis id="91A3EAE0FF96FFACF5F6FA92FEA6FA60" bold="true" box="[199,383,1391,1417]" pageId="59" pageNumber="60">
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<typeStatus id="7C6C8850FF96FFACF5F6FA92FEE0FA60" box="[199,313,1391,1417]" pageId="59" pageNumber="60" type="holotype">Holotype</typeStatus>
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male
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</emphasis>
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:
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<emphasis id="91A3EAE0FF96FFACF4A3FA92FC91FA60" bold="true" box="[402,840,1391,1417]" pageId="59" pageNumber="60">
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<collectingCountry id="DBC07662FF96FFACF4A3FA92FDFDFA60" box="[402,548,1391,1417]" name="Vietnam" pageId="59" pageNumber="60">VIETNAM</collectingCountry>
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:
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<location id="A6086029FF96FFACF702FA92FC9AFA60" LSID="urn:lsid:plazi:treatment:2B7E87E4FF94FFACF5A6FE2AFC37F9EB:A6086029FF96FFACF702FA92FC9AFA60" box="[563,835,1391,1417]" country="Vietnam" municipality="Bung" name="Nghta Bihn Province" pageId="59" pageNumber="60">Nghta Bihn Province</location>
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:
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</emphasis>
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<collectingMunicipality id="430CAC88FF96FFACF662FA8DFC4BFA63" box="[851,914,1392,1418]" pageId="59" pageNumber="60">Bung</collectingMunicipality>
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stream,
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<date id="D7691032FF96FFACF6CDFA8DFBB4FA63" box="[1020,1133,1392,1418]" pageId="59" pageNumber="60" value="1991-01-10">
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<collectingDate id="C72DE9DAFF96FFACF6CDFA8DFBB4FA63" box="[1020,1133,1392,1418]" pageId="59" pageNumber="60" value="1991-01-10">10.i.1991</collectingDate>
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</date>
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, light [
|
||
<collectorName id="0E225324FF96FFACF1F7FA8DFAA5FA63" box="[1222,1404,1392,1418]" pageId="59" pageNumber="60">S. Andrikovics</collectorName>
|
||
] – (
|
||
<collectionCode id="C5C6AE37FF96FFACF5AEFA65FF07FA5B" box="[159,222,1432,1458]" pageId="59" pageNumber="60">OPC</collectionCode>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="EBCD6579FF96FFACF5F6FA42FE05FA33" box="[199,476,1471,1498]" pageId="59" pageNumber="60" type="distribution">
|
||
<paragraph id="A36836F2FF96FFACF5F6FA42FE05FA33" blockId="59.[151,1437,152,1538]" box="[199,476,1471,1498]" pageId="59" pageNumber="60">
|
||
<emphasis id="91A3EAE0FF96FFACF5F6FA42FEB0FA30" bold="true" box="[199,361,1471,1497]" pageId="59" pageNumber="60">Distribution:</emphasis>
|
||
<collectingCountry id="DBC07662FF96FFACF440FA3DFE01FA33" box="[369,472,1472,1498]" name="Vietnam" pageId="59" pageNumber="60">Vietnam</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="EBCD6579FF96FFACF5F6FA1AFC37F9EB" box="[199,1006,1511,1538]" pageId="59" pageNumber="60" type="etymology">
|
||
<paragraph id="A36836F2FF96FFACF5F6FA1AFC37F9EB" blockId="59.[151,1437,152,1538]" box="[199,1006,1511,1538]" pageId="59" pageNumber="60">
|
||
<emphasis id="91A3EAE0FF96FFACF5F6FA1AFE8EF9E8" bold="true" box="[199,343,1511,1537]" pageId="59" pageNumber="60">Etymology:</emphasis>
|
||
<emphasis id="91A3EAE0FF96FFACF46EFA15FE6EF9E8" box="[351,439,1512,1537]" italics="true" pageId="59" pageNumber="60">Bungus</emphasis>
|
||
, named after the
|
||
<typeStatus id="7C6C8850FF96FFACF7B2FA15FD6CF9EB" box="[643,693,1512,1538]" pageId="59" pageNumber="60">type</typeStatus>
|
||
locality, the Bung Stream.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |