treatments-xml/data/1A/31/E1/1A31E1036F5EE46AFD8FFA3A9233FACB.xml

<document id="9BFC0CFD0214CD1339BDF2E1CCF3A0A5" ID-CLB-Dataset="54259" ID-DOI="10.5852/ejt.2023.870.2127" ID-GBIF-Dataset="ff5f84b3-f279-4892-9622-7c28f6440af3" ID-ISSN="2118-9773" ID-Zenodo-Dep="8006320" ID-ZooBank="60245C9C-34EE-4F68-A346-F7A684E18EB6" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1685973613176" checkinUser="felipe" docAuthor="Říčan, Oldřich &amp; Říčanová, Štěpánka" docDate="2023" docId="1A31E1036F5EE46AFD8FFA3A9233FACB" docLanguage="en" docName="ejt-870_167-201.pdf" docOrigin="European Journal of Taxonomy 870" docSource="http://dx.doi.org/10.5852/ejt.2023.870.2127" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Bujurquina omagua Říčan &amp; Říčanová 2023, sp. nov." docType="treatment" docUuid="8E478708-58F1-4E97-960C-9FCD93107C9A" docUuidSource="ZooBank" docVersion="7" lastPageNumber="194" masterDocId="E608997B6F57E471FFE2FFEF9744FFE1" masterDocTitle="A new highly apomorphic species of Bujurquina (Teleostei: Cichlidae) from a reverse flowing river in the Peruvian Amazon, with a key to the species in the genus" masterLastPageNumber="201" masterPageNumber="167" pageNumber="176" updateTime="1699528374517" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="20E2EBC7895887E9BDF36622C1CEAA2C">A new highly apomorphic species of Bujurquina (Teleostei: Cichlidae) from a reverse flowing river in the Peruvian Amazon, with a key to the species in the genus</mods:title>
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<mods:namePart id="EE90ED6604ED65C20FC4A402EDB27075">Říčan, Oldřich</mods:namePart>
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<mods:affiliation id="467F3C301AC870168E4B947287682090">University of South Bohemia, Faculty of Science, Department of Zoology, Branišovská 31, 370 05, České Budějovice, Czech Republic.</mods:affiliation>
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<mods:namePart id="585CBF5F9B64C0FF03D4A62DE725B8DC">Říčanová, Štěpánka</mods:namePart>
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<mods:affiliation id="C2E673F611D36C49F89E887438FE0719">University of South Bohemia, Faculty of Science, Department of Zoology, Branišovská 31, 370 05, České Budějovice, Czech Republic.</mods:affiliation>
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<treatment id="1A31E1036F5EE46AFD8FFA3A9233FACB" ID-DOI="http://doi.org/10.5281/zenodo.8006645" ID-GBIF-Taxon="209668944" ID-Zenodo-Dep="8006645" LSID="urn:lsid:zoobank.org:act:8E478708-58F1-4E97-960C-9FCD93107C9A" httpUri="http://treatment.plazi.org/id/1A31E1036F5EE46AFD8FFA3A9233FACB" lastPageId="27" lastPageNumber="194" pageId="9" pageNumber="176">
<subSubSection id="DA82039E6F5EE478FD8FFA3A9482FA11" box="[621,966,1493,1520]" pageId="9" pageNumber="176" type="nomenclature">
<paragraph id="922750156F5EE478FD8FFA3A9482FA11" blockId="9.[375,1214,1493,1590]" box="[621,966,1493,1520]" pageId="9" pageNumber="176">
<heading id="C96FE7796F5EE478FD8FFA3A9482FA11" box="[621,966,1493,1520]" centered="true" fontSize="11" level="2" pageId="9" pageNumber="176" reason="2">
<taxonomicName id="55982B966F5EE478FD8FFA3A9421FA0E" authority="Říčan &amp; Říčanová, 2023" authorityName="Říčan &amp; Říčanová" authorityYear="2023" box="[621,869,1493,1519]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="176" phylum="Chordata" rank="species" species="omagua" status="sp. nov.">
<emphasis id="A0EC8C076F5EE478FD8FFA3A9421FA0E" bold="true" box="[621,869,1493,1519]" italics="true" pageId="9" pageNumber="176">Bujurquina omagua</emphasis>
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<taxonomicNameLabel id="BBDF317C6F5EE478FC8EFA399482FA11" box="[876,966,1494,1520]" pageId="9" pageNumber="176" rank="species">sp. nov.</taxonomicNameLabel>
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<figureCitation id="0AA34C906F5EE478FD4FF9F49461F9D4" box="[685,805,1563,1589]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStartId-0="12.[189,232,1963,1989]" captionStartId-1="14.[189,232,1892,1918]" captionStartId-2="15.[189,232,1963,1989]" captionStartId-3="16.[189,232,1850,1876]" captionStartId-4="17.[189,232,1963,1989]" captionTargetBox-0="[192,1396,557,1920]" captionTargetBox-1="[369,1219,265,1851]" captionTargetBox-2="[189,1398,756,1920]" captionTargetBox-3="[189,1398,265,1799]" captionTargetBox-4="[236,1353,1164,1886]" captionTargetId-0="figure-83@12.[192,1396,557,1921]" captionTargetId-1="figure-16@14.[369,1219,265,1852]" captionTargetId-2="figure-138@15.[189,1398,756,1920]" captionTargetId-3="figure-15@16.[189,1398,265,1800]" captionTargetId-4="figure-263@17.[189,1398,1138,1913]" captionTargetPageId-0="12" captionTargetPageId-1="14" captionTargetPageId-2="15" captionTargetPageId-3="16" captionTargetPageId-4="17" captionText-0="Fig. 6. Bujurquina omaguasp. nov., holotype (MUSM 70225), 97.4 mm, ID tag 1189, Quebrada Sabalillo, cabeceras upper loc. P18-17 (3°27´08.4˝ S, 72°29´11.8˝ W). A. Left side. B. Right side reversed." captionText-1="Fig. 7. Bujurquina omagua sp. nov., paratypes. A. MUSM 70223, P18-16_1186, 92.6 mm. B. MUSM 70222, P18-16_1185, 79.9 mm. C. MUSM 70221, P18-15_1182, 78.7 mm. D. MUSM 70226, P18- 17_1190, 98.9 mm. E. MUSM 70229, P18-21_1200. F. MUSM 70227, P18-17_1191, 69.7 mm. G. MUSM 70228, P18-17_1192, 77.1 mm." captionText-2="Fig. 8. Bujurquina omagua sp. nov., color photos of preserved specimens. A. Holotype (MUSM 70225, ID tag 1189), 97.4 mm. B. Paratype (MUSM 70226, P18-17_1190), 98.9 mm." captionText-3="Fig. 9. Bujurquina omaguasp. nov., black and white photos of preserved specimens.A. Holotype (MUSM 70225, ID tag 1189), 97.4 mm. B. Paratype (MUSM 70226, P18-17_1190), 98.9 mm. C. Paratype (MUSM 70221, P18-15_1182), 78.7 mm." captionText-4="Fig. 10. Bujurquina omagua sp. nov., paratype (MUSM 70222, P18-16_1185, 79.9 mm), lower pharyngeal tooth plate in occlusal view. Scale bar = 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.8006334" figureDoi-1="http://doi.org/10.5281/zenodo.8006336" figureDoi-2="http://doi.org/10.5281/zenodo.8006338" figureDoi-3="http://doi.org/10.5281/zenodo.8006340" figureDoi-4="http://doi.org/10.5281/zenodo.8006342" httpUri-0="https://zenodo.org/record/8006334/files/figure.png" httpUri-1="https://zenodo.org/record/8006336/files/figure.png" httpUri-2="https://zenodo.org/record/8006338/files/figure.png" httpUri-3="https://zenodo.org/record/8006340/files/figure.png" httpUri-4="https://zenodo.org/record/8006342/files/figure.png" pageId="9" pageNumber="176">Figs 6–10</figureCitation>
; 
<tableCitation id="DF1A65AE6F5EE478FCD2F9F494C3F9D7" box="[816,903,1563,1590]" captionStart="Table 1" captionStartId="13.[189,259,265,291]" captionTargetPageId="13" captionText="Table 1. Proportional measurements in percent of standard length of nine specimens of Bujurquina omagua sp. nov. larger than 60 mm SL. Abbreviation: SD = standard deviation." httpUri="http://table.plazi.org/id/C6E7009D6F5AE47CFF5FFEE69325FEA7" pageId="9" pageNumber="176" tableUuid="C6E7009D6F5AE47CFF5FFEE69325FEA7">Table 1</tableCitation>
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<subSubSection id="DA82039E6F5EE478FF5FF9B195D1F97B" pageId="9" pageNumber="176" type="reference_group">
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<taxonomicName id="55982B966F5EE478FF5FF9B1962EF996" box="[189,362,1629,1655]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="176" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="A0EC8C076F5EE478FF5FF9B19606F996" box="[189,322,1630,1655]" italics="true" pageId="9" pageNumber="176">Bujurquina</emphasis>
sp.
</taxonomicName>
<collectingCountry id="EA8F10856F5EE478FE90F9B296E9F996" box="[370,429,1629,1655]" name="Algeria" pageId="9" pageNumber="176">Oran</collectingCountry>
<bibRefCitation id="F6092DE46F5EE478FE56F9B2953BF996" author="Rican O. &amp; Ricanova S. &amp; Rodriguez Haro L. R. &amp; Rodriguez Haro C. E." box="[436,639,1629,1657]" pageId="9" pageNumber="176" refId="ref15837" refString="Rican O., Ricanova S., Rodriguez Haro L. R. &amp; Rodriguez Haro C. E. 2022. Unrecognized species diversity and endemism in the cichlid genus Bujurquina (Teleostei: Cichlidae) together with a molecular phylogeny document large-scale transformation of the western Amazonian river network and reveal complex paleogeography of the Ecuadorian Amazon. Hydrobiologia. https: // doi. org / 10.1007 / s 10750 - 022 - 05019 - z" type="book" year="2022">
Říčan 
<emphasis id="A0EC8C076F5EE478FE1DF9B19577F996" box="[511,563,1629,1655]" italics="true" pageId="9" pageNumber="176">et al</emphasis>
. 2022
</bibRefCitation>
.
</paragraph>
<paragraph id="922750156F5EE478FF5FF96E95D1F97B" blockId="9.[189,661,1629,1693]" box="[189,661,1664,1693]" pageId="9" pageNumber="176">
<taxonomicName id="55982B966F5EE478FF5FF96E962EF97A" box="[189,362,1665,1691]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="176" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="A0EC8C076F5EE478FF5FF96E9606F97B" box="[189,322,1665,1690]" italics="true" pageId="9" pageNumber="176">Bujurquina</emphasis>
sp.
</taxonomicName>
<collectingCountry id="EA8F10856F5EE478FE90F96E96ECF97A" box="[370,424,1665,1691]" name="Peru" pageId="9" pageNumber="176">Peru</collectingCountry>
1 
<bibRefCitation id="F6092DE46F5EE478FE27F96E95D4F97B" author="Rican O. &amp; Ricanova S. &amp; Rodriguez Haro L. R. &amp; Rodriguez Haro C. E." box="[453,656,1664,1693]" pageId="9" pageNumber="176" refId="ref15837" refString="Rican O., Ricanova S., Rodriguez Haro L. R. &amp; Rodriguez Haro C. E. 2022. Unrecognized species diversity and endemism in the cichlid genus Bujurquina (Teleostei: Cichlidae) together with a molecular phylogeny document large-scale transformation of the western Amazonian river network and reveal complex paleogeography of the Ecuadorian Amazon. Hydrobiologia. https: // doi. org / 10.1007 / s 10750 - 022 - 05019 - z" type="book" year="2022">
Říčan 
<emphasis id="A0EC8C076F5EE478FDF2F96E9500F97B" box="[528,580,1664,1690]" italics="true" pageId="9" pageNumber="176">et al</emphasis>
. 2022
</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DA82039E6F5EE47BFF5FF92E933EFBC4" lastPageId="10" lastPageNumber="177" pageId="9" pageNumber="176" type="diagnosis">
<paragraph id="922750156F5EE478FF5FF92E9672F93A" blockId="9.[189,310,1729,1755]" box="[189,310,1729,1755]" pageId="9" pageNumber="176">
<heading id="C96FE7796F5EE478FF5FF92E9672F93A" bold="true" box="[189,310,1729,1755]" fontSize="11" level="3" pageId="9" pageNumber="176" reason="3">
<emphasis id="A0EC8C076F5EE478FF5FF92E9672F93A" bold="true" box="[189,310,1729,1755]" pageId="9" pageNumber="176">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="922750156F5EE47BFF5FF91E9435FD81" blockId="9.[188,1400,1776,2014]" lastBlockId="10.[189,1399,265,608]" lastPageId="10" lastPageNumber="177" pageId="9" pageNumber="176">
<taxonomicName id="55982B966F5EE478FF5FF91E96EFF8EB" authority="Říčan &amp; Říčanová, 2023" authorityName="Říčan &amp; Říčanová" authorityYear="2023" box="[189,427,1777,1802]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="176" phylum="Chordata" rank="species" species="omagua" status="sp. nov.">
<emphasis id="A0EC8C076F5EE478FF5FF91E96EFF8EB" box="[189,427,1777,1802]" italics="true" pageId="9" pageNumber="176">Bujurquina omagua</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBDF317C6F5EE478FE56F91E9556F8EA" box="[436,530,1777,1803]" pageId="9" pageNumber="176" rank="species">sp. nov.</taxonomicNameLabel>
is highly apomorphic compared to other described species in the genus because it is unique in several character states. These include: 1) shape and pattern of the suborbital stripe in adults, which is bend and with a blotch at the postero-ventral end (vs straight or absent, with or without a blotch), with an ontogenetic change with juveniles and subadults having a straight stripe running diagonally postero-ventrally and starting from below the eye in juveniles (i.e., not bend as in adults) and without a blotch, 2) orange cheek and lower head coloration with alternating opalescent lines and series of spots on snout and spotted on cheek and lower head, 3) spinous portion of dorsal fin with a single thin diagonal black line per membrane, or two lines per membrane in 
<typeStatus id="4D23EEB76F5DE47BFBDEFEE693E7FEC2" box="[1084,1187,265,291]" pageId="10" pageNumber="177" type="holotype">holotype</typeStatus>
and largest adults (vs unpatterned in all Southern group species, or with circular blotches, with several thick diagonal black lines per membrane, or two thick horizontal bands across the dorsal fin in all other Northern group species), 4) longest snout of all species without overlap (mean 15.5%, 14.4–16.4% of SL; followed by 
<taxonomicName id="55982B966F5DE47BFF07FE7896D0FE51" authorityName="Kullander" authorityYear="1987" box="[229,404,406,432]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="177" phylum="Chordata" rank="species" species="oenolaemus">
<emphasis id="A0EC8C076F5DE47BFF07FE7896D0FE51" box="[229,404,406,432]" italics="true" pageId="10" pageNumber="177">B. oenolaemus</emphasis>
</taxonomicName>
with a mean of 11.4%, 10.3–13.1 of SL), and 5) most delicate LPJ, expressed by being shortest of all species with widest horns (length/width ratio 0.42–0.45 vs 
<quantity id="5560FDF06F5DE47BFB94FE56924FFE32" box="[1142,1291,441,468]" metricMagnitude="-2" metricUnit="m" metricValue="1.4478" metricValueMax="1.6256" metricValueMin="1.27" pageId="10" pageNumber="177" unit="in" value="0.57" valueMax="0.64" valueMin="0.5">0.50–0.64 in</quantity>
all other species), one of the smallest dentigerous areas (length of dentigerous area/width of LPJ ratio 0.33–0.34 vs 0.34–0.44 except 
<taxonomicName id="55982B966F5DE47BFE4CFDEF951CFDF8" authorityName="Kullander" authorityYear="1986" box="[430,600,511,537]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="177" phylum="Chordata" rank="species" species="tambopatae">
<emphasis id="A0EC8C076F5DE47BFE4CFDEF951CFDF8" box="[430,600,511,537]" italics="true" pageId="10" pageNumber="177">B. tambopatae</emphasis>
</taxonomicName>
, 
<taxonomicName id="55982B966F5DE47BFD85FDEF95B9FDF8" authorityName="Kullander" authorityYear="1986" box="[615,765,512,537]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="177" phylum="Chordata" rank="species" species="moriorum">
<emphasis id="A0EC8C076F5DE47BFD85FDEF95B9FDF8" box="[615,765,512,537]" italics="true" pageId="10" pageNumber="177">B. moriorum</emphasis>
</taxonomicName>
and 
<taxonomicName id="55982B966F5DE47BFCD7FDEF948EFDF8" authorityName="Kullander" authorityYear="1986" box="[821,970,511,537]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="177" phylum="Chordata" rank="species" species="huallagae">
<emphasis id="A0EC8C076F5DE47BFCD7FDEF948EFDF8" box="[821,970,511,537]" italics="true" pageId="10" pageNumber="177">B. huallagae</emphasis>
</taxonomicName>
where also 0.33–0.34), and the least robust dorso-ventrally (bone at posterior end of dentigerous area less thick or equal to length of largest posterior teeth vs much thicker than teeth in other species).
</paragraph>
<paragraph id="922750156F5DE47BFF5FFD67933EFBC4" blockId="10.[189,1399,647,1061]" pageId="10" pageNumber="177">
<taxonomicName id="55982B966F5DE47BFF5FFD6796F4FD40" authority="Říčan &amp; Říčanová, 2023" authorityName="Říčan &amp; Říčanová" authorityYear="2023" box="[189,432,648,673]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="177" phylum="Chordata" rank="species" species="omagua" status="sp. nov.">
<emphasis id="A0EC8C076F5DE47BFF5FFD6796F4FD40" box="[189,432,648,673]" italics="true" pageId="10" pageNumber="177">Bujurquina omagua</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBDF317C6F5DE47BFE22FD679567FD43" box="[448,547,648,674]" pageId="10" pageNumber="177" rank="species">sp. nov.</taxonomicNameLabel>
is additionally diagnosed by the following unique combination of morphometric characters, the most pronounced of which characterize the robust head of the species: 1) second to third longest head (after 
<taxonomicName id="55982B966F5DE47BFD67FD219470FD09" authorityName="Kullander" authorityYear="1987" box="[645,820,718,744]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="177" phylum="Chordata" rank="species" species="oenolaemus">
<emphasis id="A0EC8C076F5DE47BFD67FD219470FD09" box="[645,820,718,744]" italics="true" pageId="10" pageNumber="177">B. oenolaemus</emphasis>
</taxonomicName>
and together with 
<taxonomicName id="55982B966F5DE47BFBFEFD2193D7FD09" authorityName="Kullander" authorityYear="1986" box="[1052,1171,718,744]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="177" phylum="Chordata" rank="species" species="labiosa">
<emphasis id="A0EC8C076F5DE47BFBFEFD2193D7FD09" box="[1052,1171,718,744]" italics="true" pageId="10" pageNumber="177">B. labiosa</emphasis>
</taxonomicName>
; both with a mean 37.0%, 36.3–38.0% of SL in 
<taxonomicName id="55982B966F5DE47BFDF0FD1D95D6FCEA" authorityName="Říčan &amp; Říčanová" authorityYear="2023" box="[530,658,754,779]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="177" phylum="Chordata" rank="species" species="omagua">
<emphasis id="A0EC8C076F5DE47BFDF0FD1D95D6FCEA" box="[530,658,754,779]" italics="true" pageId="10" pageNumber="177">B. omagua</emphasis>
</taxonomicName>
vs mean 39.1%, 38.6–40.4% of SL), 2) third longest preorbital distance (after 
<taxonomicName id="55982B966F5DE47BFE85FCFA969BFCCF" authorityName="Kullander" authorityYear="1986" box="[359,479,788,814]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="177" phylum="Chordata" rank="species" species="robusta">
<emphasis id="A0EC8C076F5DE47BFE85FCFA969BFCCF" box="[359,479,788,814]" italics="true" pageId="10" pageNumber="177">B. robusta</emphasis>
</taxonomicName>
and 
<taxonomicName id="55982B966F5DE47BFDF3FCFA95F8FCCF" authorityName="Kullander" authorityYear="1987" box="[529,700,788,814]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="177" phylum="Chordata" rank="species" species="oenolaemus">
<emphasis id="A0EC8C076F5DE47BFDF3FCFA95F8FCCF" box="[529,700,788,814]" italics="true" pageId="10" pageNumber="177">B. oenolaemus</emphasis>
</taxonomicName>
; mean 9.3%, 8.5–10.0% of SL), 3) large interocular distance (largest mean value of all species, 12.5% of SL vs 9.7–12.4% however with large overlap of ranges with the other species), 4) rather wide head (mean 19.9%, range 19.1–20.7% of SL), 5) rather long preorbital distance (mean 9.3%, range 8.5–10.0% of SL, third longest based on mean value), 6) robust head which has rather small eyes (mean 11.3%, range 10.3–12.0% of SL), 7) very deep body (mean 45.1%, range 43.2–46.8% of SL, based on mean the second after 
<taxonomicName id="55982B966F5DE47BFCECFC2A943CFC3F" baseAuthorityName="Heckel" baseAuthorityYear="1840" box="[782,888,965,990]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="177" phylum="Chordata" rank="species" species="vitatta">
<emphasis id="A0EC8C076F5DE47BFCECFC2A943CFC3F" box="[782,888,965,990]" italics="true" pageId="10" pageNumber="177">B. vitatta</emphasis>
</taxonomicName>
), but 8) with rather long and shallow caudal peduncle (CPL mean 13.8%, range 12.6–15.6% of SL; CPD mean 16.6%, range 15.5–17.1% of SL), and 9) rather short last spine in the dorsal fin (mean 15.6%, range 12.3–17.1% of SL).
</paragraph>
</subSubSection>
<subSubSection id="DA82039E6F5DE47BFF5FFBA39375FB3A" pageId="10" pageNumber="177" type="etymology">
<paragraph id="922750156F5DE47BFF5FFBA39600FB87" blockId="10.[189,324,1100,1126]" box="[189,324,1100,1126]" pageId="10" pageNumber="177">
<heading id="C96FE7796F5DE47BFF5FFBA39600FB87" bold="true" box="[189,324,1100,1126]" fontSize="11" level="3" pageId="10" pageNumber="177" reason="3">
<emphasis id="A0EC8C076F5DE47BFF5FFBA39600FB87" bold="true" box="[189,324,1100,1126]" pageId="10" pageNumber="177">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="922750156F5DE47BFF5FFB959375FB3A" blockId="10.[189,1399,1146,1243]" pageId="10" pageNumber="177">
The specific epithet ‘ 
<taxonomicName id="55982B966F5DE47BFE59FB94955EFB75" authorityName="Říčan &amp; Říčanová" authorityYear="2023" box="[443,538,1147,1172]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="177" phylum="Chordata" rank="species" species="omagua">
<emphasis id="A0EC8C076F5DE47BFE59FB94955EFB75" box="[443,538,1147,1172]" italics="true" pageId="10" pageNumber="177">omagua</emphasis>
</taxonomicName>
’ is a noun in apposition given after the Omagua people, which were at the time of first contact with Europeans in the 16 
<superScript id="65EDFD5D6F5DE47BFCE3FB72944AFB4A" attach="left" box="[769,782,1181,1195]" fontSize="6" pageId="10" pageNumber="177">th</superScript>
century the dominant people along the banks of the Amazon River upstream from the mouth of the Negro River well into 
<collectingCountry id="EA8F10856F5DE47BFC16FB2E936AFB3A" box="[1012,1070,1217,1243]" name="Peru" pageId="10" pageNumber="177">Peru</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="DA82039E6F5DE47AFF5FFAED943BFDDC" lastPageId="11" lastPageNumber="178" pageId="10" pageNumber="177" type="materials_examined">
<paragraph id="922750156F5DE47BFF5FFAED9628FAFD" blockId="10.[189,364,1282,1308]" box="[189,364,1282,1308]" pageId="10" pageNumber="177">
<heading id="C96FE7796F5DE47BFF5FFAED9628FAFD" bold="true" box="[189,364,1282,1308]" fontSize="11" level="3" pageId="10" pageNumber="177" reason="3">
<emphasis id="A0EC8C076F5DE47BFF5FFAED9628FAFD" bold="true" box="[189,364,1282,1308]" pageId="10" pageNumber="177">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="922750156F5DE47BFF08FADE9332FA39" blockId="10.[189,1399,1329,1496]" pageId="10" pageNumber="177">
<materialsCitation id="22F05A486F5DE47BFF08FADE9332FA39" ID-GBIF-Occurrence="4126387305" collectingDate="2018-06-28" collectionCode="MUSM" collectorName="Rican" country="Peru" county="Amazonas river" elevation="97" latitude="-3.4523335" location="Maynas Province" longLatPrecision="1" longitude="-72.48661" municipality="Department" pageId="10" pageNumber="177" specimenCode="70225" specimenCount="1" specimenCount-male="1" stateProvince="Department Loreto" typeStatus="holotype">
<emphasis id="A0EC8C076F5DE47BFF08FADE9618FAAA" bold="true" box="[234,348,1329,1355]" pageId="10" pageNumber="177">
<typeStatus id="4D23EEB76F5DE47BFF08FADE9618FAAA" box="[234,348,1329,1355]" pageId="10" pageNumber="177" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="EA8F10856F5DE47BFF5FFABB964CFA8F" box="[189,264,1364,1390]" name="Peru" pageId="10" pageNumber="177">PERU</collectingCountry>
• adult 
<specimenCount id="849E9B9C6F5DE47BFE8AFABB96C4FA8F" box="[360,384,1364,1390]" pageId="10" pageNumber="177" type="male">♂</specimenCount>
, 
<quantity id="5560FDF06F5DE47BFE6CFABB96BFFA8F" box="[398,507,1364,1390]" metricMagnitude="-2" metricUnit="m" metricValue="9.74" pageId="10" pageNumber="177" unit="mm" value="97.4">97.4 mm</quantity>
; 
<collectingCounty id="7B4628996F5DE47BFDE5FABB9587FA8F" box="[519,707,1364,1390]" pageId="10" pageNumber="177">Amazonas river</collectingCounty>
basin, 
<collectingRegion id="505C9EF76F5DE47BFCF9FABB94BBFA8F" box="[795,1023,1364,1390]" pageId="10" pageNumber="177">Department Loreto</collectingRegion>
, 
<location id="974706CE6F5DE47BFBEFFABB939AFA8F" LSID="urn:lsid:plazi:treatment:1A31E1036F5EE46AFD8FFA3A9233FACB:974706CE6F5DE47BFBEFFABB939AFA8F" box="[1037,1246,1364,1390]" country="Peru" county="Amazonas river" latitude="-3.4523335" longLatPrecision="1" longitude="-72.48661" municipality="Department" name="Maynas Province" pageId="10" pageNumber="177" stateProvince="Department Loreto">Maynas Province</location>
, 
<location id="974706CE6F5DE47BFB09FABB9678FA70" LSID="urn:lsid:plazi:treatment:1A31E1036F5EE46AFD8FFA3A9233FACB:974706CE6F5DE47BFB09FABB9678FA70" country="Peru" county="Amazonas river" latitude="-3.4523335" longLatPrecision="1" longitude="-72.48661" municipality="Department" name="District Las Amazonas" pageId="10" pageNumber="177" stateProvince="Department Loreto">District Las Amazonas</location>
, close to Oran village, 
<location id="974706CE6F5DE47BFDB6FA98947EFA70" LSID="urn:lsid:plazi:treatment:1A31E1036F5EE46AFD8FFA3A9233FACB:974706CE6F5DE47BFDB6FA98947EFA70" box="[596,826,1399,1425]" country="Peru" county="Amazonas river" latitude="-3.4523335" longLatPrecision="1" longitude="-72.48661" municipality="Department" name="Quebrada Sabalillo" pageId="10" pageNumber="177" stateProvince="Department Loreto">Quebrada Sabalillo</location>
, cabeceras upper locality P18-17; 
<geoCoordinate id="F7AC36D26F5DE47BFB35FA989235FA70" box="[1239,1393,1399,1427]" degrees="3" direction="south" minutes="27" orientation="latitude" pageId="10" pageNumber="177" precision="1" seconds="08.4" value="-3.4523335">
3° 
<emphasis id="A0EC8C076F5DE47BFB0CFA989212FA72" bold="true" box="[1262,1366,1399,1427]" pageId="10" pageNumber="177">27´08.4˝</emphasis>
S
</geoCoordinate>
, 
<geoCoordinate id="F7AC36D26F5DE47BFF5DFA75962AFA54" box="[191,366,1434,1462]" degrees="72" direction="west" minutes="29" orientation="longitude" pageId="10" pageNumber="177" precision="1" seconds="11.8" value="-72.48661">
72° 
<emphasis id="A0EC8C076F5DE47BFF04FA75960DFA57" bold="true" box="[230,329,1434,1462]" pageId="10" pageNumber="177">29´11.8˝</emphasis>
W
</geoCoordinate>
; 
<quantity id="5560FDF06F5DE47BFE9BFA7596F1FA54" box="[377,437,1434,1461]" metricMagnitude="1" metricUnit="m" metricValue="9.7" pageId="10" pageNumber="177" unit="m" value="97.0">
<elevation id="19B5B7266F5DE47BFE9BFA7596F1FA54" box="[377,437,1434,1461]" metricMagnitude="1" metricUnit="m" metricValue="9.7" pageId="10" pageNumber="177" unit="m" value="97.0">97 m</elevation>
</quantity>
a.s.l. based on GPS, 115 m a.s.l. based on Google Earth; 
<date id="E62676D56F5DE47BFBB1FA7593A8FA55" box="[1107,1260,1434,1461]" pageId="10" pageNumber="177" value="2018-06-28">
<collectingDate id="F6628F3D6F5DE47BFBB1FA7593A8FA55" box="[1107,1260,1434,1461]" pageId="10" pageNumber="177" value="2018-06-28">28 Jun. 2018</collectingDate>
</date>
; 
<collectorName id="3F6D35C36F5DE47BFB1AFA749278FA56" box="[1272,1340,1435,1463]" pageId="10" pageNumber="177">Říčan</collectorName>
leg.; clear-water stream with blackish tinge; ID tag 1189; 
<collectionCode id="F489C8D06F5DE47BFCC4FA5194C4FA39" box="[806,896,1470,1496]" pageId="10" pageNumber="177">MUSM</collectionCode>
<specimenCode id="C23EF86E6F5DE47BFC65FA529494FA39" box="[903,976,1469,1496]" pageId="10" pageNumber="177">70225</specimenCode>
(
<figureCitation id="0AA34C906F5DE47BFC02FA529369FA36" box="[992,1069,1469,1495]" captionStart="Fig" captionStartId="12.[189,232,1963,1989]" captionTargetBox="[192,1396,557,1920]" captionTargetId="figure-83@12.[192,1396,557,1921]" captionTargetPageId="12" captionText="Fig. 6. Bujurquina omaguasp. nov., holotype (MUSM 70225), 97.4 mm, ID tag 1189, Quebrada Sabalillo, cabeceras upper loc. P18-17 (3°27´08.4˝ S, 72°29´11.8˝ W). A. Left side. B. Right side reversed." figureDoi="http://doi.org/10.5281/zenodo.8006334" httpUri="https://zenodo.org/record/8006334/files/figure.png" pageId="10" pageNumber="177">Figs 6</figureCitation>
, 
<figureCitation id="0AA34C906F5DE47BFBDBFA519323FA39" box="[1081,1127,1470,1496]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="15.[189,232,1963,1989]" captionStartId-1="16.[189,232,1850,1876]" captionTargetBox-0="[189,1398,756,1920]" captionTargetBox-1="[189,1398,265,1799]" captionTargetId-0="figure-138@15.[189,1398,756,1920]" captionTargetId-1="figure-15@16.[189,1398,265,1800]" captionTargetPageId-0="15" captionTargetPageId-1="16" captionText-0="Fig. 8. Bujurquina omagua sp. nov., color photos of preserved specimens. A. Holotype (MUSM 70225, ID tag 1189), 97.4 mm. B. Paratype (MUSM 70226, P18-17_1190), 98.9 mm." captionText-1="Fig. 9. Bujurquina omaguasp. nov., black and white photos of preserved specimens.A. Holotype (MUSM 70225, ID tag 1189), 97.4 mm. B. Paratype (MUSM 70226, P18-17_1190), 98.9 mm. C. Paratype (MUSM 70221, P18-15_1182), 78.7 mm." figureDoi-0="http://doi.org/10.5281/zenodo.8006338" figureDoi-1="http://doi.org/10.5281/zenodo.8006340" httpUri-0="https://zenodo.org/record/8006338/files/figure.png" httpUri-1="https://zenodo.org/record/8006340/files/figure.png" pageId="10" pageNumber="177">8–9</figureCitation>
).
</materialsCitation>
</paragraph>
<materialsCitation id="22F05A486F5DE47BFF08FA1096BCF981" ID-GBIF-Occurrence="4126387311" country="Peru" location="Maynas Province" pageId="10" pageNumber="177" specimenCount="9" stateProvince="Department Loreto" typeStatus="paratype">
<paragraph id="922750156F5DE47BFF08FA109623F9F8" blockId="10.[189,1398,1535,1632]" box="[234,359,1535,1561]" pageId="10" pageNumber="177">
<emphasis id="A0EC8C076F5DE47BFF08FA109623F9F8" bold="true" box="[234,359,1535,1561]" pageId="10" pageNumber="177">
<typeStatus id="4D23EEB76F5DE47BFF08FA109623F9F8" box="[234,359,1535,1561]" pageId="10" pageNumber="177" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="922750156F5DE47BFF5FF9CD96BCF981" blockId="10.[189,1398,1535,1632]" pageId="10" pageNumber="177">
All from 
<collectingCountry id="EA8F10856F5DE47BFECDF9CC962CF9DC" box="[303,360,1571,1597]" name="Peru" pageId="10" pageNumber="177">Peru</collectingCountry>
, 
<locationDeviation id="839549376F5DE47BFE97F9CD9539F9DD" box="[373,637,1570,1596]" pageId="10" pageNumber="177">Amazonas river basin</locationDeviation>
, 
<collectingRegion id="505C9EF76F5DE47BFD69F9CC9434F9DC" box="[651,880,1571,1597]" pageId="10" pageNumber="177">Department Loreto</collectingRegion>
, 
<location id="974706CE6F5DE47BFC9DF9CC9314F9DD" LSID="urn:lsid:plazi:treatment:1A31E1036F5EE46AFD8FFA3A9233FACB:974706CE6F5DE47BFC9DF9CC9314F9DD" box="[895,1104,1570,1597]" country="Peru" name="Maynas Province" pageId="10" pageNumber="177" stateProvince="Department Loreto">Maynas Province</location>
, 
<location id="974706CE6F5DE47BFBBDF9CD9236F9DD" LSID="urn:lsid:plazi:treatment:1A31E1036F5EE46AFD8FFA3A9233FACB:974706CE6F5DE47BFBBDF9CD9236F9DD" box="[1119,1394,1570,1597]" country="Peru" name="District Las Amazonas" pageId="10" pageNumber="177" stateProvince="Department Loreto">District Las Amazonas</location>
, close to Oran village, 
<specimenCount id="849E9B9C6F5DE47BFE5DF9AA96BCF981" box="[447,504,1605,1632]" pageId="10" pageNumber="177" type="generic">9 ex.</specimenCount>
</paragraph>
</materialsCitation>
<paragraph id="922750156F5DE47AFF5FF968943BFDDC" blockId="10.[189,1399,1671,2014]" lastBlockId="11.[189,1399,265,573]" lastPageId="11" lastPageNumber="178" pageId="10" pageNumber="177">
<materialsCitation id="22F05A486F5DE47BFF5FF9689628F925" ID-GBIF-Occurrence="4126387301" collectionCode="MUSM" country="Peru" county="Amazonas river" latitude="-3.4523335" location="PERU" municipality="Department" pageId="10" pageNumber="177" specimenCode="MUSM 70224" specimenCount="1" specimenCount-adult="1" stateProvince="Loreto">
<collectingCountry id="EA8F10856F5DE47BFF5FF968964CF940" box="[189,264,1671,1697]" name="Peru" pageId="10" pageNumber="177">PERU</collectingCountry>
• 
<specimenCount id="849E9B9C6F5DE47BFEC4F9689639F940" box="[294,381,1671,1697]" pageId="10" pageNumber="177" type="adult">1 adult</specimenCount>
, 
<quantity id="5560FDF06F5DE47BFE69F96896B2F940" box="[395,502,1671,1697]" metricMagnitude="-2" metricUnit="m" metricValue="7.859999999999999" pageId="10" pageNumber="177" unit="mm" value="78.6">78.6 mm</quantity>
; same collection data as for holotype; ID tag 1188; GenBank: 
<accessionNumber id="8DCBCDF66F5DE47BFB13F9689236F940" box="[1265,1394,1671,1697]" pageId="10" pageNumber="177">OP436199</accessionNumber>
; 
<specimenCode id="C23EF86E6F5DE47BFF5FF9459628F925" box="[189,364,1706,1732]" collectionCode="MUSM" pageId="10" pageNumber="177">MUSM 70224</specimenCode>
</materialsCitation>
• 
<materialsCitation id="22F05A486F5DE47BFE73F945953DF906" ID-GBIF-Occurrence="4126387306" collectionCode="MUSM" country="Peru" county="Amazonas river" latitude="-3.4523335" location="GenBank" municipality="Department" pageId="10" pageNumber="177" specimenCode="MUSM 70226" specimenCount="1" specimenCount-adult="1" stateProvince="Loreto">
<specimenCount id="849E9B9C6F5DE47BFE73F94596AEF925" box="[401,490,1706,1732]" pageId="10" pageNumber="177" type="adult">1 adult</specimenCount>
, 
<quantity id="5560FDF06F5DE47BFE19F945952DF924" box="[507,617,1706,1733]" metricMagnitude="-2" metricUnit="m" metricValue="9.89" pageId="10" pageNumber="177" unit="mm" value="98.9">98.9 mm</quantity>
; same collection data as for holotype; ID tag 1190; GenBank: 
<accessionNumber id="8DCBCDF66F5DE47BFF5FF922967AF906" box="[189,318,1741,1767]" pageId="10" pageNumber="177">OP436200</accessionNumber>
; 
<specimenCode id="C23EF86E6F5DE47BFEA9F92296B2F906" box="[331,502,1741,1767]" collectionCode="MUSM" pageId="10" pageNumber="177">MUSM 70226</specimenCode>
(
<figureCitation id="0AA34C906F5DE47BFDE5F9229534F906" box="[519,624,1741,1768]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="14.[189,232,1892,1918]" captionStartId-1="15.[189,232,1963,1989]" captionStartId-2="16.[189,232,1850,1876]" captionTargetBox-0="[369,1219,265,1851]" captionTargetBox-1="[189,1398,756,1920]" captionTargetBox-2="[189,1398,265,1799]" captionTargetId-0="figure-16@14.[369,1219,265,1852]" captionTargetId-1="figure-138@15.[189,1398,756,1920]" captionTargetId-2="figure-15@16.[189,1398,265,1800]" captionTargetPageId-0="14" captionTargetPageId-1="15" captionTargetPageId-2="16" captionText-0="Fig. 7. Bujurquina omagua sp. nov., paratypes. A. MUSM 70223, P18-16_1186, 92.6 mm. B. MUSM 70222, P18-16_1185, 79.9 mm. C. MUSM 70221, P18-15_1182, 78.7 mm. D. MUSM 70226, P18- 17_1190, 98.9 mm. E. MUSM 70229, P18-21_1200. F. MUSM 70227, P18-17_1191, 69.7 mm. G. MUSM 70228, P18-17_1192, 77.1 mm." captionText-1="Fig. 8. Bujurquina omagua sp. nov., color photos of preserved specimens. A. Holotype (MUSM 70225, ID tag 1189), 97.4 mm. B. Paratype (MUSM 70226, P18-17_1190), 98.9 mm." captionText-2="Fig. 9. Bujurquina omaguasp. nov., black and white photos of preserved specimens.A. Holotype (MUSM 70225, ID tag 1189), 97.4 mm. B. Paratype (MUSM 70226, P18-17_1190), 98.9 mm. C. Paratype (MUSM 70221, P18-15_1182), 78.7 mm." figureDoi-0="http://doi.org/10.5281/zenodo.8006336" figureDoi-1="http://doi.org/10.5281/zenodo.8006338" figureDoi-2="http://doi.org/10.5281/zenodo.8006340" httpUri-0="https://zenodo.org/record/8006336/files/figure.png" httpUri-1="https://zenodo.org/record/8006338/files/figure.png" httpUri-2="https://zenodo.org/record/8006340/files/figure.png" pageId="10" pageNumber="177">Figs 7–9</figureCitation>
)
</materialsCitation>
• 
<materialsCitation id="22F05A486F5DE47BFD76F922946AF8EA" ID-GBIF-Occurrence="4126387303" collectionCode="MUSM" country="Peru" county="Amazonas river" latitude="-3.4523335" location="GenBank" municipality="Department" pageId="10" pageNumber="177" specimenCode="MUSM 70227" specimenCount="1" specimenCount-adult="1" stateProvince="Loreto">
<specimenCount id="849E9B9C6F5DE47BFD76F92295ADF906" box="[660,745,1741,1767]" pageId="10" pageNumber="177" type="adult">1 adult</specimenCount>
, 
<quantity id="5560FDF06F5DE47BFD17F922941BF909" box="[757,863,1741,1768]" metricMagnitude="-2" metricUnit="m" metricValue="6.97" pageId="10" pageNumber="177" unit="mm" value="69.7">69.7 mm</quantity>
; same collection data as for holotype; ID tag 1191; GenBank: 
<accessionNumber id="8DCBCDF66F5DE47BFE69F91F9548F8EB" box="[395,524,1776,1802]" pageId="10" pageNumber="177">OP436201</accessionNumber>
; 
<specimenCode id="C23EF86E6F5DE47BFDF9F91E958CF8EA" box="[539,712,1777,1803]" collectionCode="MUSM" pageId="10" pageNumber="177">MUSM 70227</specimenCode>
(
<figureCitation id="0AA34C906F5DE47BFD3EF91F9461F8EA" box="[732,805,1776,1803]" captionStart="Fig" captionStartId="14.[189,232,1892,1918]" captionTargetBox="[369,1219,265,1851]" captionTargetId="figure-16@14.[369,1219,265,1852]" captionTargetPageId="14" captionText="Fig. 7. Bujurquina omagua sp. nov., paratypes. A. MUSM 70223, P18-16_1186, 92.6 mm. B. MUSM 70222, P18-16_1185, 79.9 mm. C. MUSM 70221, P18-15_1182, 78.7 mm. D. MUSM 70226, P18- 17_1190, 98.9 mm. E. MUSM 70229, P18-21_1200. F. MUSM 70227, P18-17_1191, 69.7 mm. G. MUSM 70228, P18-17_1192, 77.1 mm." figureDoi="http://doi.org/10.5281/zenodo.8006336" httpUri="https://zenodo.org/record/8006336/files/figure.png" pageId="10" pageNumber="177">Fig. 7</figureCitation>
)
</materialsCitation>
• 
<materialsCitation id="22F05A486F5DE47BFCAFF91E94BFF8CF" ID-GBIF-Occurrence="4126387304" collectionCode="MUSM" country="Peru" county="Amazonas river" latitude="-3.4523335" location="GenBank" municipality="Department" pageId="10" pageNumber="177" specimenCode="MUSM 70228" specimenCount="1" specimenCount-adult="1" stateProvince="Loreto">
<specimenCount id="849E9B9C6F5DE47BFCAFF91E94E0F8EB" box="[845,932,1776,1803]" pageId="10" pageNumber="177" type="adult">1 adult</specimenCount>
, 
<quantity id="5560FDF06F5DE47BFC51F91E935BF8EA" box="[947,1055,1777,1803]" metricMagnitude="-2" metricUnit="m" metricValue="7.709999999999999" pageId="10" pageNumber="177" unit="mm" value="77.1">77.1 mm</quantity>
; same collection data as for holotype; ID tag 1192; GenBank: 
<accessionNumber id="8DCBCDF66F5DE47BFDB8F8FC959FF8CC" box="[602,731,1811,1837]" pageId="10" pageNumber="177">OP436202</accessionNumber>
; 
<specimenCode id="C23EF86E6F5DE47BFD08F8FB94D2F8CF" box="[746,918,1812,1838]" collectionCode="MUSM" pageId="10" pageNumber="177">MUSM 70228</specimenCode>
(
<figureCitation id="0AA34C906F5DE47BFC4BF8FB94B6F8CF" box="[937,1010,1812,1838]" captionStart="Fig" captionStartId="14.[189,232,1892,1918]" captionTargetBox="[369,1219,265,1851]" captionTargetId="figure-16@14.[369,1219,265,1852]" captionTargetPageId="14" captionText="Fig. 7. Bujurquina omagua sp. nov., paratypes. A. MUSM 70223, P18-16_1186, 92.6 mm. B. MUSM 70222, P18-16_1185, 79.9 mm. C. MUSM 70221, P18-15_1182, 78.7 mm. D. MUSM 70226, P18- 17_1190, 98.9 mm. E. MUSM 70229, P18-21_1200. F. MUSM 70227, P18-17_1191, 69.7 mm. G. MUSM 70228, P18-17_1192, 77.1 mm." figureDoi="http://doi.org/10.5281/zenodo.8006336" httpUri="https://zenodo.org/record/8006336/files/figure.png" pageId="10" pageNumber="177">Fig. 7</figureCitation>
)
</materialsCitation>
• 
<materialsCitation id="22F05A486F5DE47BFBF8F8FB9496F85A" ID-GBIF-Occurrence="4126387307" collectingDate="2018-06-27" collectionCode="MUSM" collectorName="Rican" country="Peru" county="Amazonas river" elevation="90" latitude="-3.4626667" location="Maynas Province" longLatPrecision="1" longitude="-72.48958" municipality="Department" pageId="10" pageNumber="177" specimenCode="MUSM 70221" specimenCount="1" specimenCount-adult="1" stateProvince="Loreto">
<specimenCount id="849E9B9C6F5DE47BFBF8F8FB9335F8CF" box="[1050,1137,1812,1838]" pageId="10" pageNumber="177" type="adult">1 adult</specimenCount>
, 
<quantity id="5560FDF06F5DE47BFB9DF8FB93AEF8CF" box="[1151,1258,1812,1838]" metricMagnitude="-2" metricUnit="m" metricValue="7.87" pageId="10" pageNumber="177" unit="mm" value="78.7">78.7 mm</quantity>
; 
<collectingCounty id="7B4628996F5DE47BFB19F8FB97B0F8B0" pageId="10" pageNumber="177">Amazonas river</collectingCounty>
basin, 
<collectingRegion id="505C9EF76F5DE47BFEB4F8D8957BF8B0" box="[342,575,1847,1873]" country="Peru" name="Loreto" pageId="10" pageNumber="177">Department Loreto</collectingRegion>
, 
<location id="974706CE6F5DE47BFDB3F8D89463F8B0" LSID="urn:lsid:plazi:treatment:1A31E1036F5EE46AFD8FFA3A9233FACB:974706CE6F5DE47BFDB3F8D89463F8B0" box="[593,807,1847,1873]" country="Peru" county="Amazonas river" latitude="-3.4626667" longLatPrecision="1" longitude="-72.48958" municipality="Department" name="Maynas Province" pageId="10" pageNumber="177" stateProvince="Loreto">Maynas Province</location>
, District Las Amazonas, close to 
<location id="974706CE6F5DE47BFB35F8D89236F8B0" LSID="urn:lsid:plazi:treatment:1A31E1036F5EE46AFD8FFA3A9233FACB:974706CE6F5DE47BFB35F8D89236F8B0" box="[1239,1394,1847,1873]" country="Peru" county="Amazonas river" latitude="-3.4626667" longLatPrecision="1" longitude="-72.48958" municipality="Department" name="Oran village" pageId="10" pageNumber="177" stateProvince="Loreto">Oran village</location>
, Qebrada Sabalillo on trail from mouth, P18-15; 
<geoCoordinate id="F7AC36D26F5DE47BFCEEF8B594ECF895" box="[780,936,1882,1910]" degrees="3" direction="south" minutes="27" orientation="latitude" pageId="10" pageNumber="177" precision="1" seconds="45.6" value="-3.4626667">
3° 
<emphasis id="A0EC8C076F5DE47BFCC6F8B594CFF897" bold="true" box="[804,907,1882,1910]" pageId="10" pageNumber="177">27´45.6˝</emphasis>
S
</geoCoordinate>
, 
<geoCoordinate id="F7AC36D26F5DE47BFC5BF8B5932BF894" box="[953,1135,1882,1910]" degrees="72" direction="west" minutes="29" orientation="longitude" pageId="10" pageNumber="177" precision="1" seconds="22.5" value="-72.48958">
72° 
<emphasis id="A0EC8C076F5DE47BFC03F8B59302F897" bold="true" box="[993,1094,1882,1910]" pageId="10" pageNumber="177">29´22.5˝</emphasis>
W
</geoCoordinate>
; 
<elevation id="19B5B7266F5DE47BFB9DF8B593FAF894" box="[1151,1214,1882,1909]" metricMagnitude="1" metricUnit="m" metricValue="9.0" pageId="10" pageNumber="177" unit="m" value="90.0">
<quantity id="5560FDF06F5DE47BFB9DF8B593FAF894" box="[1151,1214,1882,1909]" metricMagnitude="1" metricUnit="m" metricValue="9.0" pageId="10" pageNumber="177" unit="m" value="90.0">90 m</quantity>
</elevation>
a.s.l. based on GPS, 110 m a.s.l. based on Google Earth; 
<date id="E62676D56F5DE47BFD5EF892941EF876" box="[700,858,1917,1944]" pageId="10" pageNumber="177" value="2018-06-27">
<collectingDate id="F6628F3D6F5DE47BFD5EF892941EF876" box="[700,858,1917,1944]" pageId="10" pageNumber="177" value="2018-06-27">27 Jun. 2018</collectingDate>
</date>
; 
<collectorName id="3F6D35C36F5DE47BFC8AF89194E8F87B" box="[872,940,1918,1946]" pageId="10" pageNumber="177">Říčan</collectorName>
leg.; clear-water stream with blackish tinge; ID tag 1182; GenBank: OP436196; 
<specimenCode id="C23EF86E6F5DE47BFD4BF84E9417F85A" box="[681,851,1953,1979]" collectionCode="MUSM" pageId="10" pageNumber="177">MUSM 70221</specimenCode>
(
<figureCitation id="0AA34C906F5DE47BFC80F84F94EAF85A" box="[866,942,1952,1979]" captionStart="Fig" captionStartId="14.[189,232,1892,1918]" captionTargetBox="[369,1219,265,1851]" captionTargetId="figure-16@14.[369,1219,265,1852]" captionTargetPageId="14" captionText="Fig. 7. Bujurquina omagua sp. nov., paratypes. A. MUSM 70223, P18-16_1186, 92.6 mm. B. MUSM 70222, P18-16_1185, 79.9 mm. C. MUSM 70221, P18-15_1182, 78.7 mm. D. MUSM 70226, P18- 17_1190, 98.9 mm. E. MUSM 70229, P18-21_1200. F. MUSM 70227, P18-17_1191, 69.7 mm. G. MUSM 70228, P18-17_1192, 77.1 mm." figureDoi="http://doi.org/10.5281/zenodo.8006336" httpUri="https://zenodo.org/record/8006336/files/figure.png" pageId="10" pageNumber="177">Figs 7</figureCitation>
, 
<figureCitation id="0AA34C906F5DE47BFC5BF84E948CF85A" box="[953,968,1953,1979]" captionStart="Fig" captionStartId="16.[189,232,1850,1876]" captionTargetBox="[189,1398,265,1799]" captionTargetId="figure-15@16.[189,1398,265,1800]" captionTargetPageId="16" captionText="Fig. 9. Bujurquina omaguasp. nov., black and white photos of preserved specimens.A. Holotype (MUSM 70225, ID tag 1189), 97.4 mm. B. Paratype (MUSM 70226, P18-17_1190), 98.9 mm. C. Paratype (MUSM 70221, P18-15_1182), 78.7 mm." figureDoi="http://doi.org/10.5281/zenodo.8006340" httpUri="https://zenodo.org/record/8006340/files/figure.png" pageId="10" pageNumber="177">9</figureCitation>
)
</materialsCitation>
• 
<materialsCitation id="22F05A486F5DE47AFC08F84E94FDFE88" ID-GBIF-Occurrence="4126387308" collectingDate="2018-06-28" collectionCode="MUSM" collectorName="Rican" country="Peru" county="Amazonas river" elevation="75" lastPageId="11" lastPageNumber="178" latitude="-3.4548612" location="Maynas Province" longLatPrecision="1" longitude="-72.48966" municipality="Department" pageId="10" pageNumber="177" specimenCode="MUSM 70222" specimenCount="1" specimenCount-adult="1" stateProvince="Department Loreto">
<specimenCount id="849E9B9C6F5DE47BFC08F84E9379F85B" box="[1002,1085,1952,1979]" pageId="10" pageNumber="177" type="adult">1 adult</specimenCount>
, 
<quantity id="5560FDF06F5DE47BFBAAF84E93F4F85A" box="[1096,1200,1953,1979]" metricMagnitude="-2" metricUnit="m" metricValue="7.99" pageId="10" pageNumber="177" unit="mm" value="79.9">79.9 mm</quantity>
; 
<collectingCounty id="7B4628996F5DE47BFB5FF84E9232F85B" box="[1213,1398,1952,1979]" pageId="10" pageNumber="177">Amazonas river</collectingCounty>
basin, 
<collectingRegion id="505C9EF76F5DE47BFEE9F82B96ABF83F" box="[267,495,1988,2014]" pageId="10" pageNumber="177">Department Loreto</collectingRegion>
, 
<location id="974706CE6F5DE47BFE1FF82B958AF83F" LSID="urn:lsid:plazi:treatment:1A31E1036F5EE46AFD8FFA3A9233FACB:974706CE6F5DE47BFE1FF82B958AF83F" box="[509,718,1988,2014]" country="Peru" county="Amazonas river" latitude="-3.4548612" longLatPrecision="1" longitude="-72.48966" municipality="Department" name="Maynas Province" pageId="10" pageNumber="177" stateProvince="Department Loreto">Maynas Province</location>
, District Las Amazonas, close to 
<location id="974706CE6F5DE47BFB82F82B93B3F83F" LSID="urn:lsid:plazi:treatment:1A31E1036F5EE46AFD8FFA3A9233FACB:974706CE6F5DE47BFB82F82B93B3F83F" box="[1120,1271,1988,2014]" country="Peru" county="Amazonas river" latitude="-3.4548612" longLatPrecision="1" longitude="-72.48966" municipality="Department" name="Oran village" pageId="10" pageNumber="177" stateProvince="Department Loreto">Oran village</location>
, cabeceras of 
<location id="974706CE6F5CE47AFF02FEE6968CFEC2" LSID="urn:lsid:plazi:treatment:1A31E1036F5EE46AFD8FFA3A9233FACB:974706CE6F5CE47AFF02FEE6968CFEC2" box="[224,456,265,291]" country="Peru" county="Amazonas river" latitude="-3.4548612" longLatPrecision="1" longitude="-72.48966" municipality="Department" name="Quebrada Sabalillo" pageId="11" pageNumber="178" stateProvince="Department Loreto">Quebrada Sabalillo</location>
, lower loc., P18-16; 
<geoCoordinate id="F7AC36D26F5CE47AFD2BFEE69420FEC2" box="[713,868,265,293]" degrees="3" direction="south" minutes="27" orientation="latitude" pageId="11" pageNumber="178" precision="1" seconds="17.5" value="-3.4548612">
3° 
<emphasis id="A0EC8C076F5CE47AFD03FEE6940CFEC4" bold="true" box="[737,840,265,293]" pageId="11" pageNumber="178">27´17.5˝</emphasis>
S
</geoCoordinate>
, 
<geoCoordinate id="F7AC36D26F5CE47AFC97FEE6936EFEC2" box="[885,1066,265,293]" degrees="72" direction="west" minutes="29" orientation="longitude" pageId="11" pageNumber="178" precision="1" seconds="22.8" value="-72.48966">
72° 
<emphasis id="A0EC8C076F5CE47AFC7FFEE69346FEC4" bold="true" box="[925,1026,265,293]" pageId="11" pageNumber="178">29´22.8˝</emphasis>
W
</geoCoordinate>
; 
<elevation id="19B5B7266F5CE47AFBDBFEE6933CFEC2" box="[1081,1144,265,292]" metricMagnitude="1" metricUnit="m" metricValue="7.5" pageId="11" pageNumber="178" unit="m" value="75.0">
<quantity id="5560FDF06F5CE47AFBDBFEE6933CFEC2" box="[1081,1144,265,292]" metricMagnitude="1" metricUnit="m" metricValue="7.5" pageId="11" pageNumber="178" unit="m" value="75.0">75 m</quantity>
</elevation>
a.s.l. based on GPS, 117 m a.s.l. based on Google Earth; 
<date id="E62676D56F5CE47AFD91FEC39454FEA7" box="[627,784,300,327]" pageId="11" pageNumber="178" value="2018-06-28">
<collectingDate id="F6628F3D6F5CE47AFD91FEC39454FEA7" box="[627,784,300,327]" pageId="11" pageNumber="178" value="2018-06-28">28 Jun. 2018</collectingDate>
</date>
; 
<collectorName id="3F6D35C36F5CE47AFCFCFEC29426FEA8" box="[798,866,301,329]" pageId="11" pageNumber="178">Říčan</collectorName>
leg.; clear-water stream with blackish tinge; ID tag 1185; GenBank: OP436197; 
<specimenCode id="C23EF86E6F5CE47AFD91FEA09464FE88" box="[627,800,335,361]" collectionCode="MUSM" pageId="11" pageNumber="178">MUSM 70222</specimenCode>
(
<figureCitation id="0AA34C906F5CE47AFCD1FEA094C6FE8B" box="[819,898,335,362]" captionStart="Fig" captionStartId="14.[189,232,1892,1918]" captionTargetBox="[369,1219,265,1851]" captionTargetId="figure-16@14.[369,1219,265,1852]" captionTargetPageId="14" captionText="Fig. 7. Bujurquina omagua sp. nov., paratypes. A. MUSM 70223, P18-16_1186, 92.6 mm. B. MUSM 70222, P18-16_1185, 79.9 mm. C. MUSM 70221, P18-15_1182, 78.7 mm. D. MUSM 70226, P18- 17_1190, 98.9 mm. E. MUSM 70229, P18-21_1200. F. MUSM 70227, P18-17_1191, 69.7 mm. G. MUSM 70228, P18-17_1192, 77.1 mm." figureDoi="http://doi.org/10.5281/zenodo.8006336" httpUri="https://zenodo.org/record/8006336/files/figure.png" pageId="11" pageNumber="178">Figs 7</figureCitation>
, 
<figureCitation id="0AA34C906F5CE47AFC70FEA094F4FE88" box="[914,944,335,361]" captionStart="Fig" captionStartId="17.[189,232,1963,1989]" captionTargetBox="[236,1353,1164,1886]" captionTargetId="figure-263@17.[189,1398,1138,1913]" captionTargetPageId="17" captionText="Fig. 10. Bujurquina omagua sp. nov., paratype (MUSM 70222, P18-16_1185, 79.9 mm), lower pharyngeal tooth plate in occlusal view. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8006342" httpUri="https://zenodo.org/record/8006342/files/figure.png" pageId="11" pageNumber="178">10</figureCitation>
)
</materialsCitation>
• 
<materialsCitation id="22F05A486F5CE47AFC3AFEA0933AFE6C" ID-GBIF-Occurrence="4126387310" collectingDate="2018-06-28" collectionCode="MUSM" collectorName="Rican" country="Peru" county="Amazonas river" elevation="75" latitude="-3.4548612" location="GenBank" longLatPrecision="1" longitude="-72.48966" municipality="Department" pageId="11" pageNumber="178" specimenCode="MUSM 70223" specimenCount="1" specimenCount-adult="1" stateProvince="Loreto">
<specimenCount id="849E9B9C6F5CE47AFC3AFEA0936BFE88" box="[984,1071,335,361]" pageId="11" pageNumber="178" type="adult">1 adult</specimenCount>
, 
<quantity id="5560FDF06F5CE47AFBDCFEA093EEFE8B" box="[1086,1194,335,362]" metricMagnitude="-2" metricUnit="m" metricValue="9.26" pageId="11" pageNumber="178" unit="mm" value="92.6">92.6 mm</quantity>
; same collection data as for preceding; ID tag 1186; GenBank: 
<accessionNumber id="8DCBCDF66F5CE47AFD01FE9D9421FE6D" box="[739,869,370,396]" pageId="11" pageNumber="178">OP436198</accessionNumber>
; 
<specimenCode id="C23EF86E6F5CE47AFC90FE9C9359FE6C" box="[882,1053,371,397]" collectionCode="MUSM" pageId="11" pageNumber="178">MUSM 70223</specimenCode>
(
<figureCitation id="0AA34C906F5CE47AFBCCFE9D9331FE6C" box="[1070,1141,370,397]" captionStart="Fig" captionStartId="14.[189,232,1892,1918]" captionTargetBox="[369,1219,265,1851]" captionTargetId="figure-16@14.[369,1219,265,1852]" captionTargetPageId="14" captionText="Fig. 7. Bujurquina omagua sp. nov., paratypes. A. MUSM 70223, P18-16_1186, 92.6 mm. B. MUSM 70222, P18-16_1185, 79.9 mm. C. MUSM 70221, P18-15_1182, 78.7 mm. D. MUSM 70226, P18- 17_1190, 98.9 mm. E. MUSM 70229, P18-21_1200. F. MUSM 70227, P18-17_1191, 69.7 mm. G. MUSM 70228, P18-17_1192, 77.1 mm." figureDoi="http://doi.org/10.5281/zenodo.8006336" httpUri="https://zenodo.org/record/8006336/files/figure.png" pageId="11" pageNumber="178">Fig. 7</figureCitation>
)
</materialsCitation>
• 
<materialsCitation id="22F05A486F5CE47AFB78FE9C9366FDF8" ID-GBIF-Occurrence="4126387309" collectingDate="2018-07-03" collectionCode="MUSM" collectorName="Rican" country="Peru" county="Amazonas river" elevation="82" latitude="-3.455889" location="Maynas Province" longLatPrecision="1" longitude="-72.512" municipality="Department" pageId="11" pageNumber="178" specimenCode="MUSM 70229" specimenCount="1" specimenCount-adult="1" stateProvince="Loreto">
<specimenCount id="849E9B9C6F5CE47AFB78FE9C93A8FE6D" box="[1178,1260,370,397]" pageId="11" pageNumber="178" type="adult">1 adult</specimenCount>
; 
<collectingCounty id="7B4628996F5CE47AFB19FE9C97B0FE51" pageId="11" pageNumber="178">Amazonas river</collectingCounty>
basin, 
<collectingRegion id="505C9EF76F5CE47AFEA8FE799569FE51" box="[330,557,406,432]" country="Peru" name="Loreto" pageId="11" pageNumber="178">Department Loreto</collectingRegion>
, 
<location id="974706CE6F5CE47AFDD8FE79944DFE51" LSID="urn:lsid:plazi:treatment:1A31E1036F5EE46AFD8FFA3A9233FACB:974706CE6F5CE47AFDD8FE79944DFE51" box="[570,777,406,432]" country="Peru" county="Amazonas river" latitude="-3.455889" longLatPrecision="1" longitude="-72.512" municipality="Department" name="Maynas Province" pageId="11" pageNumber="178" stateProvince="Loreto">Maynas Province</location>
, District Las Amazonas, close to 
<location id="974706CE6F5CE47AFB77FE79926FFE51" LSID="urn:lsid:plazi:treatment:1A31E1036F5EE46AFD8FFA3A9233FACB:974706CE6F5CE47AFB77FE79926FFE51" box="[1173,1323,406,432]" country="Peru" county="Amazonas river" latitude="-3.455889" longLatPrecision="1" longitude="-72.512" municipality="Department" name="Oran village" pageId="11" pageNumber="178" stateProvince="Loreto">Oran village</location>
, small unnamed quebrada just before entering 
<location id="974706CE6F5CE47AFD74FE569459FE32" LSID="urn:lsid:plazi:treatment:1A31E1036F5EE46AFD8FFA3A9233FACB:974706CE6F5CE47AFD74FE569459FE32" box="[662,797,441,467]" country="Peru" county="Amazonas river" latitude="-3.455889" longLatPrecision="1" longitude="-72.512" municipality="Department" name="Oran creek" pageId="11" pageNumber="178" stateProvince="Loreto">Oran creek</location>
, P18-21; 
<geoCoordinate id="F7AC36D26F5CE47AFC72FE56936DFE32" box="[912,1065,441,469]" degrees="3" direction="south" minutes="27" orientation="latitude" pageId="11" pageNumber="178" precision="1" seconds="21.2" value="-3.455889">
3° 
<emphasis id="A0EC8C076F5CE47AFC45FE56934BFE34" bold="true" box="[935,1039,441,469]" pageId="11" pageNumber="178">27´21.2˝</emphasis>
S
</geoCoordinate>
, 
<geoCoordinate id="F7AC36D26F5CE47AFBDAFE5693A8FE32" box="[1080,1260,441,469]" degrees="72" direction="west" minutes="30" orientation="longitude" pageId="11" pageNumber="178" precision="1" seconds="43.2" value="-72.512">
72° 
<emphasis id="A0EC8C076F5CE47AFB82FE569382FE34" bold="true" box="[1120,1222,441,469]" pageId="11" pageNumber="178">30´43.2˝</emphasis>
W
</geoCoordinate>
; 
<elevation id="19B5B7266F5CE47AFB18FE569273FE32" box="[1274,1335,441,467]" metricMagnitude="1" metricUnit="m" metricValue="8.2" pageId="11" pageNumber="178" unit="m" value="82.0">
<quantity id="5560FDF06F5CE47AFB18FE569273FE32" box="[1274,1335,441,467]" metricMagnitude="1" metricUnit="m" metricValue="8.2" pageId="11" pageNumber="178" unit="m" value="82.0">82 m</quantity>
</elevation>
a.s.l. based on GPS, 104 m a.s.l. based on Google Earth; 
<date id="E62676D56F5CE47AFCD5FE339486FE17" box="[823,962,476,502]" pageId="11" pageNumber="178" value="2018-07-03">
<collectingDate id="F6628F3D6F5CE47AFCD5FE339486FE17" box="[823,962,476,502]" pageId="11" pageNumber="178" value="2018-07-03">3 Jul. 2018</collectingDate>
</date>
; 
<collectorName id="3F6D35C36F5CE47AFC33FE329351FE18" box="[977,1045,477,505]" pageId="11" pageNumber="178">Říčan</collectorName>
leg.; clear-water stream with blackish tinge; ID tag 1200; GenBank: OP436204; 
<specimenCode id="C23EF86E6F5CE47AFCFAFE109486FDF8" box="[792,962,511,537]" collectionCode="MUSM" pageId="11" pageNumber="178">MUSM 70229</specimenCode>
(
<figureCitation id="0AA34C906F5CE47AFC30FE10935CFDF8" box="[978,1048,511,537]" captionStart="Fig" captionStartId="14.[189,232,1892,1918]" captionTargetBox="[369,1219,265,1851]" captionTargetId="figure-16@14.[369,1219,265,1852]" captionTargetPageId="14" captionText="Fig. 7. Bujurquina omagua sp. nov., paratypes. A. MUSM 70223, P18-16_1186, 92.6 mm. B. MUSM 70222, P18-16_1185, 79.9 mm. C. MUSM 70221, P18-15_1182, 78.7 mm. D. MUSM 70226, P18- 17_1190, 98.9 mm. E. MUSM 70229, P18-21_1200. F. MUSM 70227, P18-17_1191, 69.7 mm. G. MUSM 70228, P18-17_1192, 77.1 mm." figureDoi="http://doi.org/10.5281/zenodo.8006336" httpUri="https://zenodo.org/record/8006336/files/figure.png" pageId="11" pageNumber="178">Fig. 7</figureCitation>
)
</materialsCitation>
• 
<materialsCitation id="22F05A486F5CE47AFBD9FE10943FFDDC" ID-GBIF-Occurrence="4126387302" collectingDate="2018-07-03" collectionCode="MUSM" collectorName="Rican" country="Peru" county="Amazonas river" elevation="82" latitude="-3.455889" location="GenBank" longLatPrecision="1" longitude="-72.512" municipality="Department" pageId="11" pageNumber="178" specimenCode="MUSM 70230" specimenCount="1" specimenCount-juvenile="1" stateProvince="Loreto">
<specimenCount id="849E9B9C6F5CE47AFBD9FE1093F7FDF8" box="[1083,1203,511,537]" pageId="11" pageNumber="178" type="juvenile">1 juvenile</specimenCount>
; same collection data as for preceding; GenBank: 
<accessionNumber id="8DCBCDF66F5CE47AFDA2FDCD9585FDDC" box="[576,705,546,573]" pageId="11" pageNumber="178">OP436205</accessionNumber>
; 
<specimenCode id="C23EF86E6F5CE47AFD2FFDCC943FFDDC" box="[717,891,547,573]" collectionCode="MUSM" pageId="11" pageNumber="178">MUSM 70230</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DA82039E6F5CE460FF5FFD8B9687FDE0" lastPageId="17" lastPageNumber="184" pageId="11" pageNumber="178" type="description">
<paragraph id="922750156F5CE47AFF5FFD8B960AFD9F" blockId="11.[189,334,612,638]" box="[189,334,612,638]" pageId="11" pageNumber="178">
<heading id="C96FE7796F5CE47AFF5FFD8B960AFD9F" bold="true" box="[189,334,612,638]" fontSize="11" level="3" pageId="11" pageNumber="178" reason="3">
<emphasis id="A0EC8C076F5CE47AFF5FFD8B960AFD9F" bold="true" box="[189,334,612,638]" pageId="11" pageNumber="178">Description</emphasis>
</heading>
</paragraph>
<paragraph id="922750156F5CE47AFF5FFD7D95C6FD31" blockId="11.[189,1398,658,720]" pageId="11" pageNumber="178">
Compound, based on 
<specimenCount id="849E9B9C6F5CE47AFE30FD7D95D1FD4D" box="[466,661,658,684]" pageId="11" pageNumber="178" type="generic">eight specimens</specimenCount>
from Sabalillo creek, 
<quantity id="5560FDF06F5CE47AFC55FD7D9328FD4C" box="[951,1132,658,685]" metricMagnitude="-2" metricUnit="m" metricValue="8.43" metricValueMax="9.89" metricValueMin="6.97" pageId="11" pageNumber="178" unit="mm" value="84.3" valueMax="98.9" valueMin="69.7">69.7–98.9 mm</quantity>
. Refer to 
<figureCitation id="0AA34C906F5CE47AFB11FD7D9237FD4C" box="[1267,1395,658,685]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStartId-0="12.[189,232,1963,1989]" captionStartId-1="14.[189,232,1892,1918]" captionStartId-2="15.[189,232,1963,1989]" captionStartId-3="16.[189,232,1850,1876]" captionStartId-4="17.[189,232,1963,1989]" captionTargetBox-0="[192,1396,557,1920]" captionTargetBox-1="[369,1219,265,1851]" captionTargetBox-2="[189,1398,756,1920]" captionTargetBox-3="[189,1398,265,1799]" captionTargetBox-4="[236,1353,1164,1886]" captionTargetId-0="figure-83@12.[192,1396,557,1921]" captionTargetId-1="figure-16@14.[369,1219,265,1852]" captionTargetId-2="figure-138@15.[189,1398,756,1920]" captionTargetId-3="figure-15@16.[189,1398,265,1800]" captionTargetId-4="figure-263@17.[189,1398,1138,1913]" captionTargetPageId-0="12" captionTargetPageId-1="14" captionTargetPageId-2="15" captionTargetPageId-3="16" captionTargetPageId-4="17" captionText-0="Fig. 6. Bujurquina omaguasp. nov., holotype (MUSM 70225), 97.4 mm, ID tag 1189, Quebrada Sabalillo, cabeceras upper loc. P18-17 (3°27´08.4˝ S, 72°29´11.8˝ W). A. Left side. B. Right side reversed." captionText-1="Fig. 7. Bujurquina omagua sp. nov., paratypes. A. MUSM 70223, P18-16_1186, 92.6 mm. B. MUSM 70222, P18-16_1185, 79.9 mm. C. MUSM 70221, P18-15_1182, 78.7 mm. D. MUSM 70226, P18- 17_1190, 98.9 mm. E. MUSM 70229, P18-21_1200. F. MUSM 70227, P18-17_1191, 69.7 mm. G. MUSM 70228, P18-17_1192, 77.1 mm." captionText-2="Fig. 8. Bujurquina omagua sp. nov., color photos of preserved specimens. A. Holotype (MUSM 70225, ID tag 1189), 97.4 mm. B. Paratype (MUSM 70226, P18-17_1190), 98.9 mm." captionText-3="Fig. 9. Bujurquina omaguasp. nov., black and white photos of preserved specimens.A. Holotype (MUSM 70225, ID tag 1189), 97.4 mm. B. Paratype (MUSM 70226, P18-17_1190), 98.9 mm. C. Paratype (MUSM 70221, P18-15_1182), 78.7 mm." captionText-4="Fig. 10. Bujurquina omagua sp. nov., paratype (MUSM 70222, P18-16_1185, 79.9 mm), lower pharyngeal tooth plate in occlusal view. Scale bar = 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.8006334" figureDoi-1="http://doi.org/10.5281/zenodo.8006336" figureDoi-2="http://doi.org/10.5281/zenodo.8006338" figureDoi-3="http://doi.org/10.5281/zenodo.8006340" figureDoi-4="http://doi.org/10.5281/zenodo.8006342" httpUri-0="https://zenodo.org/record/8006334/files/figure.png" httpUri-1="https://zenodo.org/record/8006336/files/figure.png" httpUri-2="https://zenodo.org/record/8006338/files/figure.png" httpUri-3="https://zenodo.org/record/8006340/files/figure.png" httpUri-4="https://zenodo.org/record/8006342/files/figure.png" pageId="11" pageNumber="178">Figs 6–10</figureCitation>
. Measurements summarized in 
<tableCitation id="DF1A65AE6F5CE47AFDC6FD5995C6FD31" box="[548,642,694,720]" captionStart="Table 1" captionStartId="13.[189,259,265,291]" captionTargetPageId="13" captionText="Table 1. Proportional measurements in percent of standard length of nine specimens of Bujurquina omagua sp. nov. larger than 60 mm SL. Abbreviation: SD = standard deviation." httpUri="http://table.plazi.org/id/C6E7009D6F5AE47CFF5FFEE69325FEA7" pageId="11" pageNumber="178" tableUuid="C6E7009D6F5AE47CFF5FFEE69325FEA7">Table 1.</tableCitation>
</paragraph>
<paragraph id="922750156F5CE47AFF5FFD1893D8FBAF" blockId="11.[188,1399,759,1103]" pageId="11" pageNumber="178">
<smallCapsWord id="94C1C6C96F5CE47AFF5FFD189647FCF1" baselines="778,779" box="[189,259,759,785]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Shape" pageId="11" pageNumber="178">SHAPE</smallCapsWord>
. Moderately elongate, rather robust, deep bodied (body depth mean 45.1%, range 43.2–46.8% of SL). Head especially robust, long (head length mean 37.0%, 36.3–38.0% of SL), wide (head width mean 19.9%, range 19.1–20.7% of SL), with large interocular distance (mean 12.5%, range 11.6–14.2% of SL). Frontal outline straight, nape and snout curved, anterior half of dorsal-fin base contour curved. Ventral outline curved. Snout long (mean 15.5%, 14.4–16.4% of SL). Preorbital distance long (mean 9.3%, range 8.5–10.0% of SL), preorbital bone much longer than high. Robust head with rather small eyes (mean 11.3%, range 10.3–12.0% of SL). Lips rather narrow, not turgid. Corner of mouth not reaching vertical from anterior margin of orbit, especially in largest specimens. Rather long and shallow caudal peduncle (CPL mean 13.8%, range 12.6–15.6% of SL; CPD mean 16.6%, range 15.5–17.1% of SL), and rather short last spine in dorsal fin (mean 15.6%, range 12.3–17.1% of SL).
</paragraph>
<paragraph id="922750156F5CE47AFF5FFB999363FB18" blockId="11.[189,1398,1141,1274]" pageId="11" pageNumber="178">
<smallCapsWord id="94C1C6C96F5CE47AFF5FFB99964BFB6F" baselines="1161,1161" box="[189,271,1142,1168]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Scales" pageId="11" pageNumber="178">SCALES</smallCapsWord>
. Squ. long. (E1) scales 24 (5), 25 (3); 2 scales between L1 and dorsal fin anteriorly, 2 scales between L1 and L2. Upper lateral line (L1) with 15 (1), 16 (7) scales. Lower lateral line (L2) with 9 (4), 10 (3), 11 (1) scales; canal bearing scales on caudal fin: 0 in dorsal, 
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median and 0 in ventral sequence. Cheek scales in 3 (8) series. Anterior portion of caudal fin scaly.
</paragraph>
<paragraph id="922750156F5CE47AFF5FFACE93D6FA29" blockId="11.[189,1399,1312,1480]" pageId="11" pageNumber="178">
<smallCapsWord id="94C1C6C96F5CE47AFF5FFACE97AAFAD8" baselines="1332,1332" box="[189,238,1313,1339]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Fins" pageId="11" pageNumber="178">FINS</smallCapsWord>
. Dorsal fin XII.11 (1), XIII.10 (1), XIII.11 (1), XIV.9 (1), XIV.10 (4). Soft anal fin pointed, 3 
<superScript id="65EDFD5D6F5CE47AFAD9FACF920DFACF" attach="left" box="[1339,1353,1312,1326]" fontSize="6" pageId="11" pageNumber="178">rd</superScript>
ray longest, reaching to middle of caudal fin or slightly beyond. Anal fin III.8 (8). Pectoral fin with rounded tip, reaching to above genital papilla to anal-fin origin (to 2 
<superScript id="65EDFD5D6F5CE47AFC8EFA899439FA95" attach="left" box="[876,893,1382,1396]" fontSize="6" pageId="11" pageNumber="178">nd</superScript>
spine in 
<specimenCount id="849E9B9C6F5CE47AFC07FA8893CEFA60" box="[997,1162,1383,1409]" pageId="11" pageNumber="178" type="generic">one specimen</specimenCount>
); Pectoral fin 13 (8). Pelvic fin pointed, first ray slightly produced, reaching to origin of anal fin or base of 1 
<superScript id="65EDFD5D6F5CE47AFB47FA6593F4FA79" attach="left" box="[1189,1200,1418,1432]" fontSize="6" pageId="11" pageNumber="178">st</superScript>
spine of anal fin. Caudal fin rounded, without well-developed streamers or with only short streamers.
</paragraph>
<paragraph id="922750156F5CE47AFF5FFA0097B3F9CD" blockId="11.[189,1399,1518,1580]" pageId="11" pageNumber="178">
<smallCapsWord id="94C1C6C96F5CE47AFF5FFA00960BF9E6" baselines="1538,1539,1538,1538" box="[189,335,1519,1545]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Gill-rakers" pageId="11" pageNumber="178">GILL-RAKERS</smallCapsWord>
. 1 epibranchial, 
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angle, and 5 (3), 6 (4), 7 (1) ceratobranchial rakers externally on first arch.
</paragraph>
<paragraph id="922750156F5CE47AFF5FF9BB9492F971" blockId="11.[189,1399,1619,1681]" pageId="11" pageNumber="178">
<smallCapsWord id="94C1C6C96F5CE47AFF5FF9BB97AEF98D" baselines="1639,1639" box="[189,234,1620,1646]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Jaw" pageId="11" pageNumber="178">JAW</smallCapsWord>
<smallCapsWord id="94C1C6C96F5CE47AFF0CF9B69674F98D" baselines="1639" box="[238,304,1625,1644]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="teeth" pageId="11" pageNumber="178">TEETH</smallCapsWord>
. An outer series of stronger, conical, sharply pointed, slightly recurved teeth, slightly increasing in size anteriorly, not worn, 
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outer hemiseries in upper jaw.
</paragraph>
<paragraph id="922750156F5CE47DFF5FF9579249FEA7" blockId="11.[189,1399,1720,1993]" lastBlockId="12.[189,1398,265,327]" lastPageId="12" lastPageNumber="179" pageId="11" pageNumber="178">
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<smallCapsWord id="94C1C6C96F5CE47AFE4AF9519695F930" baselines="1740" box="[424,465,1726,1745]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="jaw" pageId="11" pageNumber="178">JAW</smallCapsWord>
<smallCapsWord id="94C1C6C96F5CE47AFE3BF9519542F930" baselines="1740" box="[473,518,1726,1745]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="and" pageId="11" pageNumber="178">AND</smallCapsWord>
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. Lower pharyngeal jaw (LPJ) as examined in 
<specimenCount id="849E9B9C6F5CE47AFB96F9579266F933" box="[1140,1314,1720,1746]" pageId="11" pageNumber="178" type="generic">two specimens</specimenCount>
(
<quantity id="5560FDF06F5CE47AFAD1F9579663F914" metricMagnitude="-2" metricUnit="m" metricValue="8.934999999999999" metricValueMax="9.89" metricValueMin="7.9799999999999995" pageId="11" pageNumber="178" unit="mm" value="89.35" valueMax="98.9" valueMin="79.8">79.8– 98.9 mm</quantity>
) (
<figureCitation id="0AA34C906F5CE47AFEDFF93496D5F914" box="[317,401,1755,1781]" captionStart="Fig" captionStartId="17.[189,232,1963,1989]" captionTargetBox="[236,1353,1164,1886]" captionTargetId="figure-263@17.[189,1398,1138,1913]" captionTargetPageId="17" captionText="Fig. 10. Bujurquina omagua sp. nov., paratype (MUSM 70222, P18-16_1185, 79.9 mm), lower pharyngeal tooth plate in occlusal view. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8006342" httpUri="https://zenodo.org/record/8006342/files/figure.png" pageId="11" pageNumber="178">Fig. 10</figureCitation>
) very delicate, most delicate LPJ of all described species of 
<taxonomicName id="55982B966F5CE47AFBACF9339397F914" authorityName="Kullander" authorityYear="1986" box="[1102,1235,1756,1781]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="178" phylum="Chordata" rank="genus">
<emphasis id="A0EC8C076F5CE47AFBACF9339397F914" box="[1102,1235,1756,1781]" italics="true" pageId="11" pageNumber="178">Bujurquina</emphasis>
</taxonomicName>
, very shallow vertically dorso-ventrally (bone at posterior end of dentigerous area less thick than length of largest posterior teeth in the smaller specimen, almost so in the larger specimen vs thicker than the teeth in all other species). LPJ also shortest and widest of all species, length/width ratio 0.42–0.45 (vs 
<quantity id="5560FDF06F5CE47AFB01F8AA9233F8BE" box="[1251,1399,1861,1888]" metricMagnitude="-2" metricUnit="m" metricValue="1.4478" metricValueMax="1.6256" metricValueMin="1.27" pageId="11" pageNumber="178" unit="in" value="0.57" valueMax="0.64" valueMin="0.5">0.50–0.64 in</quantity>
all other species). LPJ tooth-plate (due to the longest arms within 
<taxonomicName id="55982B966F5CE47AFC2FF8869316F863" authorityName="Kullander" authorityYear="1986" box="[973,1106,1897,1922]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="178" phylum="Chordata" rank="genus">
<emphasis id="A0EC8C076F5CE47AFC2FF8869316F863" box="[973,1106,1897,1922]" italics="true" pageId="11" pageNumber="178">Bujurquina</emphasis>
</taxonomicName>
) also one of the shortest in genus, length of dentigerous area/width of LPJ ratio 0.33–0.34 (vs 0.34–0.44 except 
<taxonomicName id="55982B966F5CE47AFB21F863922BF844" authorityName="Kullander" authorityYear="1986" box="[1219,1391,1931,1957]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="178" phylum="Chordata" rank="species" species="tambopatae">
<emphasis id="A0EC8C076F5CE47AFB21F863922BF844" box="[1219,1391,1931,1957]" italics="true" pageId="11" pageNumber="178">B. tambopatae</emphasis>
</taxonomicName>
, 
<taxonomicName id="55982B966F5CE47AFF5FF840961EF829" authorityName="Kullander" authorityYear="1986" box="[189,346,1967,1992]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="178" phylum="Chordata" rank="species" species="moriorum">
<emphasis id="A0EC8C076F5CE47AFF5FF840961EF829" box="[189,346,1967,1992]" italics="true" pageId="11" pageNumber="178">B. moriorum</emphasis>
</taxonomicName>
and 
<taxonomicName id="55982B966F5CE47AFE40F840957BF829" authorityName="Kullander" authorityYear="1986" box="[418,575,1966,1992]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="178" phylum="Chordata" rank="species" species="huallagae">
<emphasis id="A0EC8C076F5CE47AFE40F840957BF829" box="[418,575,1966,1992]" italics="true" pageId="11" pageNumber="178">B. huallagae</emphasis>
</taxonomicName>
where also 0.33–0.34), Teeth slightly laterally compressed (most pronounced in posterior row), slightly bicuspid with main cusp posterior, central and posterior teeth largest, smaller towards outside, central teeth with flat-worn apex, others with pronounced tip.
</paragraph>
<paragraph id="922750156F5BE47DFF5FFE829681FE69" blockId="12.[189,453,365,392]" box="[189,453,365,392]" pageId="12" pageNumber="179">
<heading id="C96FE7796F5BE47DFF5FFE829681FE69" bold="true" box="[189,453,365,392]" fontSize="11" level="3" pageId="12" pageNumber="179" reason="3">
<emphasis id="A0EC8C076F5BE47DFF5FFE829681FE69" bold="true" box="[189,453,365,392]" pageId="12" pageNumber="179">Coloration in alcohol</emphasis>
</heading>
</paragraph>
<paragraph id="922750156F5BE47DFF5FFE739306FE1D" blockId="12.[189,1398,412,508]" pageId="12" pageNumber="179">Coloration limited to melanin patterns in preserved state, ground colour light yellowish-brownish, lighter towards whitish on chest and anterior abdomen, dusky greyish on anterior head, snout, and dorsum. Dark diffuse and wide interorbital band. Operculum silvery greyish.</paragraph>
<caption id="C6E7009D6F5BE47DFF5FF844926FF809" ID-DOI="http://doi.org/10.5281/zenodo.8006334" ID-Zenodo-Dep="8006334" httpUri="https://zenodo.org/record/8006334/files/figure.png" pageId="12" pageNumber="179" startId="12.[189,232,1963,1989]" targetBox="[192,1396,557,1920]" targetPageId="12">
<paragraph id="922750156F5BE47DFF5FF844926FF809" blockId="12.[189,1398,1963,2026]" pageId="12" pageNumber="179">
<emphasis id="A0EC8C076F5BE47DFF5FF844964CF824" bold="true" box="[189,264,1963,1989]" pageId="12" pageNumber="179">Fig. 6.</emphasis>
<taxonomicName id="55982B966F5BE47DFEEFF84396B1F824" authority="Říčan &amp; Říčanová, 2023" authorityName="Říčan &amp; Říčanová" authorityYear="2023" box="[269,501,1964,1989]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="179" phylum="Chordata" rank="species" species="omagua" status="sp. nov.">
<emphasis id="A0EC8C076F5BE47DFEEFF84396B1F824" box="[269,501,1964,1989]" italics="true" pageId="12" pageNumber="179">Bujurquina omagua</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBDF317C6F5BE47DFE14F8439515F827" box="[502,593,1964,1990]" pageId="12" pageNumber="179" rank="species">sp. nov.</taxonomicNameLabel>
, holotype (MUSM 70225), 97.4 mm, ID tag 1189, Quebrada Sabalillo, cabeceras upper loc. P18-17 (3° 
<emphasis id="A0EC8C076F5BE47DFDD2F82195DCF80B" bold="true" box="[560,664,1998,2026]" pageId="12" pageNumber="179">27´08.4˝</emphasis>
S, 72° 
<emphasis id="A0EC8C076F5BE47DFD04F821940EF80B" bold="true" box="[742,842,1998,2026]" pageId="12" pageNumber="179">29´11.8˝</emphasis>
W). 
<emphasis id="A0EC8C076F5BE47DFC67F82194DEF809" bold="true" box="[901,922,1998,2024]" pageId="12" pageNumber="179">A</emphasis>
. Left side. 
<emphasis id="A0EC8C076F5BE47DFBFDF8219377F809" bold="true" box="[1055,1075,1998,2024]" pageId="12" pageNumber="179">B</emphasis>
. Right side reversed.
</paragraph>
</caption>
<caption id="C6E7009D6F5AE47CFF5FFEE69325FEA7" ID-Table-UUID="C6E7009D6F5AE47CFF5FFEE69325FEA7" httpUri="http://table.plazi.org/id/C6E7009D6F5AE47CFF5FFEE69325FEA7" pageId="13" pageNumber="180" startId="13.[189,259,265,291]" targetBox="[254,1299,378,1085]" targetIsTable="true" targetPageId="13">
<paragraph id="922750156F5AE47CFF5FFEE69325FEA7" blockId="13.[189,1399,265,327]" pageId="13" pageNumber="180">
<emphasis id="A0EC8C076F5AE47CFF5FFEE69660FEC2" bold="true" box="[189,292,265,291]" pageId="13" pageNumber="180">Table 1.</emphasis>
Proportional measurements in percent of standard length of nine specimens of 
<taxonomicName id="55982B966F5AE47CFB10FEE59658FEA7" authority="Říčan &amp; Říčanová, 2023" authorityName="Říčan &amp; Říčanová" authorityYear="2023" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="180" phylum="Chordata" rank="species" species="omagua" status="sp. nov.">
<emphasis id="A0EC8C076F5AE47CFB10FEE59658FEA7" italics="true" pageId="13" pageNumber="180">Bujurquina omagua</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBDF317C6F5AE47CFEC1FEC29639FEA6" box="[291,381,301,327]" pageId="13" pageNumber="180" rank="species">sp. nov.</taxonomicNameLabel>
larger than 60 mm SL. Abbreviation: SD = standard deviation.
</paragraph>
</caption>
<paragraph id="922750156F5AE47CFC86FE959257FBDD" pageId="13" pageNumber="180">
<table id="E098A2B56F5A1B8EFF1CFE959257FBDC" box="[254,1299,378,1085]" gridcols="7" gridrows="16" pageId="13" pageNumber="180">
<tr id="2CA852576F5A1B8EFF1CFE959257FE75" box="[254,1299,378,404]" gridrow="0" pageId="13" pageNumber="180" rowspan-0="1" rowspan-1="1" rowspan-5="1" rowspan-6="1">
<th id="6F793B2B6F5A1B8EFCC2FE959309FE75" box="[800,1101,378,404]" colspan="3" colspanRight="2" gridcol="2" gridrow="0" pageId="13" pageNumber="180">
<emphasis id="A0EC8C076F5AE47CFC86FE959309FE75" bold="true" box="[868,1101,378,404]" pageId="13" pageNumber="180">
<taxonomicName id="55982B966F5AE47CFC86FE9594ADFE75" authority="Říčan &amp; Říčanová, 2023" authorityName="Říčan &amp; Říčanová" authorityYear="2023" box="[868,1001,378,404]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="180" phylum="Chordata" rank="species" species="omagua" status="sp. nov.">
<emphasis id="A0EC8C076F5AE47CFC86FE9594ADFE75" bold="true" box="[868,1001,378,404]" italics="true" pageId="13" pageNumber="180">B. omagua</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBDF317C6F5AE47CFC12FE959309FE75" box="[1008,1101,378,404]" pageId="13" pageNumber="180" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</th>
</tr>
<tr id="2CA852576F5A1B8EFF1CFE499257FE20" box="[254,1299,422,449]" gridrow="1" pageId="13" pageNumber="180" rowspan-0="1">
<td id="6F793B2B6F5A1B8EFD61FE4995B4FE20" box="[643,752,422,449]" gridcol="1" gridrow="1" pageId="13" pageNumber="180">Holotype</td>
<td id="6F793B2B6F5A1B8EFCC2FE499427FE20" box="[800,867,422,449]" gridcol="2" gridrow="1" pageId="13" pageNumber="180">Mean</td>
<td id="6F793B2B6F5A1B8EFC7FFE499309FE20" box="[925,1101,422,449]" colspan="2" colspanRight="1" gridcol="3" gridrow="1" pageId="13" pageNumber="180">Range</td>
<td id="6F793B2B6F5A1B8EFB62FE4993F7FE20" box="[1152,1203,422,449]" gridcol="5" gridrow="1" pageId="13" pageNumber="180">SD</td>
<td id="6F793B2B6F5A1B8EFAE6FE499257FE20" box="[1284,1299,422,449]" gridcol="6" gridrow="1" pageId="13" pageNumber="180">n</td>
</tr>
<tr id="2CA852576F5A1B8EFF1CFE3C9257FE0C" box="[254,1299,467,493]" gridrow="2" pageId="13" pageNumber="180" rowspan-0="1" rowspan-1="1" rowspan-2="1" rowspan-5="1" rowspan-6="1">
<td id="6F793B2B6F5A1B8EFC7FFE3C9494FE0C" box="[925,976,467,493]" gridcol="3" gridrow="2" pageId="13" pageNumber="180">min</td>
<td id="6F793B2B6F5A1B8EFBEDFE3C9309FE0C" box="[1039,1101,467,493]" gridcol="4" gridrow="2" pageId="13" pageNumber="180">max</td>
</tr>
<tr id="2CA852576F5A1B8EFF1CFE129257FDF9" box="[254,1299,509,536]" gridrow="3" pageId="13" pageNumber="180" rowspan-5="1">
<th id="6F793B2B6F5A1B8EFF1CFE12957AFDF9" box="[254,574,509,536]" gridcol="0" gridrow="3" pageId="13" pageNumber="180">Standard length (mm)</th>
<td id="6F793B2B6F5A1B8EFD61FE1295B4FDF9" box="[643,752,509,536]" gridcol="1" gridrow="3" pageId="13" pageNumber="180">97.4</td>
<td id="6F793B2B6F5A1B8EFCC2FE129427FDF9" box="[800,867,509,536]" gridcol="2" gridrow="3" pageId="13" pageNumber="180">84.1</td>
<td id="6F793B2B6F5A1B8EFC7FFE129494FDF9" box="[925,976,509,536]" gridcol="3" gridrow="3" pageId="13" pageNumber="180">69.7</td>
<td id="6F793B2B6F5A1B8EFBEDFE129309FDF9" box="[1039,1101,509,536]" gridcol="4" gridrow="3" pageId="13" pageNumber="180">98.9</td>
<td id="6F793B2B6F5A1B8EFAE6FE129257FDF9" box="[1284,1299,509,536]" gridcol="6" gridrow="3" pageId="13" pageNumber="180">8</td>
</tr>
<tr id="2CA852576F5A1B8EFF1CFDC49257FDA4" box="[254,1299,555,581]" gridrow="4" pageId="13" pageNumber="180">
<th id="6F793B2B6F5A1B8EFF1CFDC4957AFDA4" box="[254,574,555,581]" gridcol="0" gridrow="4" pageId="13" pageNumber="180">Head length</th>
<td id="6F793B2B6F5A1B8EFD61FDC495B4FDA4" box="[643,752,555,581]" gridcol="1" gridrow="4" pageId="13" pageNumber="180">36.3</td>
<td id="6F793B2B6F5A1B8EFCC2FDC49427FDA4" box="[800,867,555,581]" gridcol="2" gridrow="4" pageId="13" pageNumber="180">37.0</td>
<td id="6F793B2B6F5A1B8EFC7FFDC49494FDA4" box="[925,976,555,581]" gridcol="3" gridrow="4" pageId="13" pageNumber="180">36.3</td>
<td id="6F793B2B6F5A1B8EFBEDFDC49309FDA4" box="[1039,1101,555,581]" gridcol="4" gridrow="4" pageId="13" pageNumber="180">38.0</td>
<td id="6F793B2B6F5A1B8EFB62FDC493F7FDA4" box="[1152,1203,555,581]" gridcol="5" gridrow="4" pageId="13" pageNumber="180">0.69</td>
<td id="6F793B2B6F5A1B8EFAE6FDC49257FDA4" box="[1284,1299,555,581]" gridcol="6" gridrow="4" pageId="13" pageNumber="180">8</td>
</tr>
<tr id="2CA852576F5A1B8EFF1CFDB79257FD92" box="[254,1299,600,627]" gridrow="5" pageId="13" pageNumber="180">
<th id="6F793B2B6F5A1B8EFF1CFDB7957AFD92" box="[254,574,600,627]" gridcol="0" gridrow="5" pageId="13" pageNumber="180">Snout length</th>
<td id="6F793B2B6F5A1B8EFD61FDB795B4FD92" box="[643,752,600,627]" gridcol="1" gridrow="5" pageId="13" pageNumber="180">16.3</td>
<td id="6F793B2B6F5A1B8EFCC2FDB79427FD92" box="[800,867,600,627]" gridcol="2" gridrow="5" pageId="13" pageNumber="180">15.5</td>
<td id="6F793B2B6F5A1B8EFC7FFDB79494FD92" box="[925,976,600,627]" gridcol="3" gridrow="5" pageId="13" pageNumber="180">14.5</td>
<td id="6F793B2B6F5A1B8EFBEDFDB79309FD92" box="[1039,1101,600,627]" gridcol="4" gridrow="5" pageId="13" pageNumber="180">16.4</td>
<td id="6F793B2B6F5A1B8EFB62FDB793F7FD92" box="[1152,1203,600,627]" gridcol="5" gridrow="5" pageId="13" pageNumber="180">0.84</td>
<td id="6F793B2B6F5A1B8EFAE6FDB79257FD92" box="[1284,1299,600,627]" gridcol="6" gridrow="5" pageId="13" pageNumber="180">8</td>
</tr>
<tr id="2CA852576F5A1B8EFF1CFD699257FD40" box="[254,1299,646,673]" gridrow="6" pageId="13" pageNumber="180">
<th id="6F793B2B6F5A1B8EFF1CFD69957AFD40" box="[254,574,646,673]" gridcol="0" gridrow="6" pageId="13" pageNumber="180">Body depth</th>
<td id="6F793B2B6F5A1B8EFD61FD6995B4FD40" box="[643,752,646,673]" gridcol="1" gridrow="6" pageId="13" pageNumber="180">43.2</td>
<td id="6F793B2B6F5A1B8EFCC2FD699427FD40" box="[800,867,646,673]" gridcol="2" gridrow="6" pageId="13" pageNumber="180">45.1</td>
<td id="6F793B2B6F5A1B8EFC7FFD699494FD40" box="[925,976,646,673]" gridcol="3" gridrow="6" pageId="13" pageNumber="180">43.2</td>
<td id="6F793B2B6F5A1B8EFBEDFD699309FD40" box="[1039,1101,646,673]" gridcol="4" gridrow="6" pageId="13" pageNumber="180">46.8</td>
<td id="6F793B2B6F5A1B8EFB62FD6993F7FD40" box="[1152,1203,646,673]" gridcol="5" gridrow="6" pageId="13" pageNumber="180">1.00</td>
<td id="6F793B2B6F5A1B8EFAE6FD699257FD40" box="[1284,1299,646,673]" gridcol="6" gridrow="6" pageId="13" pageNumber="180">8</td>
</tr>
<tr id="2CA852576F5A1B8EFF1CFD5B9257FD2F" box="[254,1299,692,718]" gridrow="7" pageId="13" pageNumber="180">
<th id="6F793B2B6F5A1B8EFF1CFD5B957AFD2F" box="[254,574,692,718]" gridcol="0" gridrow="7" pageId="13" pageNumber="180">Orbital diameter</th>
<td id="6F793B2B6F5A1B8EFD61FD5B95B4FD2F" box="[643,752,692,718]" gridcol="1" gridrow="7" pageId="13" pageNumber="180">10.3</td>
<td id="6F793B2B6F5A1B8EFCC2FD5B9427FD2F" box="[800,867,692,718]" gridcol="2" gridrow="7" pageId="13" pageNumber="180">11.3</td>
<td id="6F793B2B6F5A1B8EFC7FFD5B9494FD2F" box="[925,976,692,718]" gridcol="3" gridrow="7" pageId="13" pageNumber="180">10.3</td>
<td id="6F793B2B6F5A1B8EFBEDFD5B9309FD2F" box="[1039,1101,692,718]" gridcol="4" gridrow="7" pageId="13" pageNumber="180">12.0</td>
<td id="6F793B2B6F5A1B8EFB62FD5B93F7FD2F" box="[1152,1203,692,718]" gridcol="5" gridrow="7" pageId="13" pageNumber="180">0.66</td>
<td id="6F793B2B6F5A1B8EFAE6FD5B9257FD2F" box="[1284,1299,692,718]" gridcol="6" gridrow="7" pageId="13" pageNumber="180">8</td>
</tr>
<tr id="2CA852576F5A1B8EFF1CFD0D9257FD1D" box="[254,1299,738,764]" gridrow="8" pageId="13" pageNumber="180">
<th id="6F793B2B6F5A1B8EFF1CFD0D957AFD1D" box="[254,574,738,764]" gridcol="0" gridrow="8" pageId="13" pageNumber="180">Head width</th>
<td id="6F793B2B6F5A1B8EFD61FD0D95B4FD1D" box="[643,752,738,764]" gridcol="1" gridrow="8" pageId="13" pageNumber="180">19.7</td>
<td id="6F793B2B6F5A1B8EFCC2FD0D9427FD1D" box="[800,867,738,764]" gridcol="2" gridrow="8" pageId="13" pageNumber="180">19.9</td>
<td id="6F793B2B6F5A1B8EFC7FFD0D9494FD1D" box="[925,976,738,764]" gridcol="3" gridrow="8" pageId="13" pageNumber="180">19.1</td>
<td id="6F793B2B6F5A1B8EFBEDFD0D9309FD1D" box="[1039,1101,738,764]" gridcol="4" gridrow="8" pageId="13" pageNumber="180">20.7</td>
<td id="6F793B2B6F5A1B8EFB62FD0D93F7FD1D" box="[1152,1203,738,764]" gridcol="5" gridrow="8" pageId="13" pageNumber="180">0.64</td>
<td id="6F793B2B6F5A1B8EFAE6FD0D9257FD1D" box="[1284,1299,738,764]" gridcol="6" gridrow="8" pageId="13" pageNumber="180">8</td>
</tr>
<tr id="2CA852576F5A1B8EFF1CFCFF9257FCCA" box="[254,1299,784,811]" gridrow="9" pageId="13" pageNumber="180">
<th id="6F793B2B6F5A1B8EFF1CFCFF957AFCCA" box="[254,574,784,811]" gridcol="0" gridrow="9" pageId="13" pageNumber="180">Interorbital width</th>
<td id="6F793B2B6F5A1B8EFD61FCFF95B4FCCA" box="[643,752,784,811]" gridcol="1" gridrow="9" pageId="13" pageNumber="180">12.6</td>
<td id="6F793B2B6F5A1B8EFCC2FCFF9427FCCA" box="[800,867,784,811]" gridcol="2" gridrow="9" pageId="13" pageNumber="180">12.5</td>
<td id="6F793B2B6F5A1B8EFC7FFCFF9494FCCA" box="[925,976,784,811]" gridcol="3" gridrow="9" pageId="13" pageNumber="180">11.6</td>
<td id="6F793B2B6F5A1B8EFBEDFCFF9309FCCA" box="[1039,1101,784,811]" gridcol="4" gridrow="9" pageId="13" pageNumber="180">14.2</td>
<td id="6F793B2B6F5A1B8EFB62FCFF93F7FCCA" box="[1152,1203,784,811]" gridcol="5" gridrow="9" pageId="13" pageNumber="180">0.90</td>
<td id="6F793B2B6F5A1B8EFAE6FCFF9257FCCA" box="[1284,1299,784,811]" gridcol="6" gridrow="9" pageId="13" pageNumber="180">8</td>
</tr>
<tr id="2CA852576F5A1B8EFF1CFCD29257FCB9" box="[254,1299,829,856]" gridrow="10" pageId="13" pageNumber="180">
<th id="6F793B2B6F5A1B8EFF1CFCD2957AFCB9" box="[254,574,829,856]" gridcol="0" gridrow="10" pageId="13" pageNumber="180">Preorbital depth</th>
<td id="6F793B2B6F5A1B8EFD61FCD295B4FCB9" box="[643,752,829,856]" gridcol="1" gridrow="10" pageId="13" pageNumber="180">9.6</td>
<td id="6F793B2B6F5A1B8EFCC2FCD29427FCB9" box="[800,867,829,856]" gridcol="2" gridrow="10" pageId="13" pageNumber="180">9.3</td>
<td id="6F793B2B6F5A1B8EFC7FFCD29494FCB9" box="[925,976,829,856]" gridcol="3" gridrow="10" pageId="13" pageNumber="180">8.5</td>
<td id="6F793B2B6F5A1B8EFBEDFCD29309FCB9" box="[1039,1101,829,856]" gridcol="4" gridrow="10" pageId="13" pageNumber="180">10.0</td>
<td id="6F793B2B6F5A1B8EFB62FCD293F7FCB9" box="[1152,1203,829,856]" gridcol="5" gridrow="10" pageId="13" pageNumber="180">0.54</td>
<td id="6F793B2B6F5A1B8EFAE6FCD29257FCB9" box="[1284,1299,829,856]" gridcol="6" gridrow="10" pageId="13" pageNumber="180">8</td>
</tr>
<tr id="2CA852576F5A1B8EFF1CFC849257FC67" box="[254,1299,875,902]" gridrow="11" pageId="13" pageNumber="180">
<th id="6F793B2B6F5A1B8EFF1CFC84957AFC67" box="[254,574,875,902]" gridcol="0" gridrow="11" pageId="13" pageNumber="180">Caudal-peduncle depth</th>
<td id="6F793B2B6F5A1B8EFD61FC8495B4FC67" box="[643,752,875,902]" gridcol="1" gridrow="11" pageId="13" pageNumber="180">15.5</td>
<td id="6F793B2B6F5A1B8EFCC2FC849427FC67" box="[800,867,875,902]" gridcol="2" gridrow="11" pageId="13" pageNumber="180">16.6</td>
<td id="6F793B2B6F5A1B8EFC7FFC849494FC67" box="[925,976,875,902]" gridcol="3" gridrow="11" pageId="13" pageNumber="180">15.5</td>
<td id="6F793B2B6F5A1B8EFBEDFC849309FC67" box="[1039,1101,875,902]" gridcol="4" gridrow="11" pageId="13" pageNumber="180">17.1</td>
<td id="6F793B2B6F5A1B8EFB62FC8493F7FC67" box="[1152,1203,875,902]" gridcol="5" gridrow="11" pageId="13" pageNumber="180">0.52</td>
<td id="6F793B2B6F5A1B8EFAE6FC849257FC67" box="[1284,1299,875,902]" gridcol="6" gridrow="11" pageId="13" pageNumber="180">8</td>
</tr>
<tr id="2CA852576F5A1B8EFF1CFC769257FC55" box="[254,1299,921,948]" gridrow="12" pageId="13" pageNumber="180">
<th id="6F793B2B6F5A1B8EFF1CFC76957AFC55" box="[254,574,921,948]" gridcol="0" gridrow="12" pageId="13" pageNumber="180">Caudal-peduncle length</th>
<td id="6F793B2B6F5A1B8EFD61FC7695B4FC55" box="[643,752,921,948]" gridcol="1" gridrow="12" pageId="13" pageNumber="180">12.7</td>
<td id="6F793B2B6F5A1B8EFCC2FC769427FC55" box="[800,867,921,948]" gridcol="2" gridrow="12" pageId="13" pageNumber="180">13.8</td>
<td id="6F793B2B6F5A1B8EFC7FFC769494FC55" box="[925,976,921,948]" gridcol="3" gridrow="12" pageId="13" pageNumber="180">12.6</td>
<td id="6F793B2B6F5A1B8EFBEDFC769309FC55" box="[1039,1101,921,948]" gridcol="4" gridrow="12" pageId="13" pageNumber="180">15.6</td>
<td id="6F793B2B6F5A1B8EFB62FC7693F7FC55" box="[1152,1203,921,948]" gridcol="5" gridrow="12" pageId="13" pageNumber="180">1.05</td>
<td id="6F793B2B6F5A1B8EFAE6FC769257FC55" box="[1284,1299,921,948]" gridcol="6" gridrow="12" pageId="13" pageNumber="180">8</td>
</tr>
<tr id="2CA852576F5A1B8EFF1CFC289257FC00" box="[254,1299,967,993]" gridrow="13" pageId="13" pageNumber="180">
<th id="6F793B2B6F5A1B8EFF1CFC28957AFC00" box="[254,574,967,993]" gridcol="0" gridrow="13" pageId="13" pageNumber="180">Pectoral-fin length</th>
<td id="6F793B2B6F5A1B8EFD61FC2895B4FC00" box="[643,752,967,993]" gridcol="1" gridrow="13" pageId="13" pageNumber="180">33.3</td>
<td id="6F793B2B6F5A1B8EFCC2FC289427FC00" box="[800,867,967,993]" gridcol="2" gridrow="13" pageId="13" pageNumber="180">33.4</td>
<td id="6F793B2B6F5A1B8EFC7FFC289494FC00" box="[925,976,967,993]" gridcol="3" gridrow="13" pageId="13" pageNumber="180">32.4</td>
<td id="6F793B2B6F5A1B8EFBEDFC289309FC00" box="[1039,1101,967,993]" gridcol="4" gridrow="13" pageId="13" pageNumber="180">34.8</td>
<td id="6F793B2B6F5A1B8EFB62FC2893F7FC00" box="[1152,1203,967,993]" gridcol="5" gridrow="13" pageId="13" pageNumber="180">0.81</td>
<td id="6F793B2B6F5A1B8EFAE6FC289257FC00" box="[1284,1299,967,993]" gridcol="6" gridrow="13" pageId="13" pageNumber="180">8</td>
</tr>
<tr id="2CA852576F5A1B8EFF1CFC1A9257FBF1" box="[254,1299,1013,1040]" gridrow="14" pageId="13" pageNumber="180">
<th id="6F793B2B6F5A1B8EFF1CFC1A957AFBF1" box="[254,574,1013,1040]" gridcol="0" gridrow="14" pageId="13" pageNumber="180">Pelvic-fin length</th>
<td id="6F793B2B6F5A1B8EFD61FC1A95B4FBF1" box="[643,752,1013,1040]" gridcol="1" gridrow="14" pageId="13" pageNumber="180">36.0</td>
<td id="6F793B2B6F5A1B8EFCC2FC1A9427FBF1" box="[800,867,1013,1040]" gridcol="2" gridrow="14" pageId="13" pageNumber="180">35.9</td>
<td id="6F793B2B6F5A1B8EFC7FFC1A9494FBF1" box="[925,976,1013,1040]" gridcol="3" gridrow="14" pageId="13" pageNumber="180">30.1</td>
<td id="6F793B2B6F5A1B8EFBEDFC1A9309FBF1" box="[1039,1101,1013,1040]" gridcol="4" gridrow="14" pageId="13" pageNumber="180">39.5</td>
<td id="6F793B2B6F5A1B8EFB62FC1A93F7FBF1" box="[1152,1203,1013,1040]" gridcol="5" gridrow="14" pageId="13" pageNumber="180">2.92</td>
<td id="6F793B2B6F5A1B8EFAE6FC1A9257FBF1" box="[1284,1299,1013,1040]" gridcol="6" gridrow="14" pageId="13" pageNumber="180">8</td>
</tr>
<tr id="2CA852576F5A1B8EFF1CFBCD9257FBDC" box="[254,1299,1058,1085]" gridrow="15" pageId="13" pageNumber="180">
<th id="6F793B2B6F5A1B8EFF1CFBCD957AFBDC" box="[254,574,1058,1085]" gridcol="0" gridrow="15" pageId="13" pageNumber="180">Last dorsal-fin spine length</th>
<td id="6F793B2B6F5A1B8EFD61FBCD95B4FBDC" box="[643,752,1058,1085]" gridcol="1" gridrow="15" pageId="13" pageNumber="180">15.7</td>
<td id="6F793B2B6F5A1B8EFCC2FBCD9427FBDC" box="[800,867,1058,1085]" gridcol="2" gridrow="15" pageId="13" pageNumber="180">15.6</td>
<td id="6F793B2B6F5A1B8EFC7FFBCD9494FBDC" box="[925,976,1058,1085]" gridcol="3" gridrow="15" pageId="13" pageNumber="180">12.3</td>
<td id="6F793B2B6F5A1B8EFBEDFBCD9309FBDC" box="[1039,1101,1058,1085]" gridcol="4" gridrow="15" pageId="13" pageNumber="180">17.1</td>
<td id="6F793B2B6F5A1B8EFB62FBCD93F7FBDC" box="[1152,1203,1058,1085]" gridcol="5" gridrow="15" pageId="13" pageNumber="180">1.59</td>
<td id="6F793B2B6F5A1B8EFAE6FBCD9257FBDC" box="[1284,1299,1058,1085]" gridcol="6" gridrow="15" pageId="13" pageNumber="180">8</td>
</tr>
</table>
</paragraph>
<paragraph id="922750156F5AE47CFF5FFB6E94CEFAAB" blockId="13.[189,1399,1152,1355]" pageId="13" pageNumber="180">Up to four clearly defined grey lines between eye and mouth corner and on anterioir cheek interspersed with spots, rest of cheek as well as opercular series richly spotted. Suborbital stripe rather marrow and of uniform width, diffuse and ghost-like, bend from behind eye forward to cheek and then postero-ventrally to a variously developed suborbital stripe blotch in adults (never ocellated and with diffuse borders). In one collected juvenile specimen and in subadults straight (i.e., not bend), running from below eye (vs from behind eye) diagonally postero-ventrally.</paragraph>
<paragraph id="922750156F5AE47CFF5FFA9D9341FA17" blockId="13.[189,1399,1394,1526]" pageId="13" pageNumber="180">
Midlateral band continuous from head to penultimate vertical bar on body, strongly pigmented and black, very wide anteriorly and gradually narrowing posteriorly. No interruption in vertical bar 5 (sensu 
<bibRefCitation id="F6092DE46F5AE47CFF5FFA5696CFFA32" author="Rican O. &amp; Musilova Z. &amp; Muska M. &amp; Novak J." box="[189,395,1464,1493]" pageId="13" pageNumber="180" pagination="1 - 46" refId="ref15407" refString="Rican O., Musilova Z., Muska M. &amp; Novak J. 2005. Development of coloration patterns in Neotropical cichlids (Perciformes: Cichlidae: Cichlasomatinae). Folia Zoologica 54: 1 - 46." type="journal article" year="2005">
Říčan 
<emphasis id="A0EC8C076F5AE47CFEEBFA569600FA33" box="[265,324,1464,1490]" italics="true" pageId="13" pageNumber="180">et al.</emphasis>
2005
</bibRefCitation>
), midlateral blotch (usually present and dominant in bar 
<quantity id="5560FDF06F5AE47CFBC7FA579316FA33" box="[1061,1106,1464,1491]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="13" pageNumber="180" unit="in" value="5.0">5 in</quantity>
cichlasomatine cichlids) is absent. Lateral band continued on head across nape close behind eye.
</paragraph>
<paragraph id="922750156F5AE47CFF5FF9F29677F906" blockId="13.[189,1399,1565,1767]" pageId="13" pageNumber="180">Body with five vertical bars plus one caudal peduncle bar and a caudal peduncle blotch (usually vertically elongated). The five body bars correspond to ontogenetic bars 2–5 with the anteriormost ontogenetic bars 6–7 fused into one wide bar below midlateral band but visible as two above it. Posterior body bars (2–4) narrow and tightly spaced, anterior bars (5, 6+ 7) wide and widely spaced. Bars darkest dorsally, reaching ventrally approximately to level of lower edge of caudal peduncle. Bars do not extend onto dorsal fin.</paragraph>
<paragraph id="922750156F5AE47CFF5FF8E19233F8C9" blockId="13.[189,1399,1806,1833]" box="[189,1399,1806,1833]" pageId="13" pageNumber="180">Body scale centres (below midlateral band) with dark blotches, more pronounced in zones of vertical bars.</paragraph>
<paragraph id="922750156F5AE47CFF5FF8A09236F832" blockId="13.[189,1399,1871,2004]" pageId="13" pageNumber="180">Dorsal fin light grey, lappets indistinct, with a single narrow grey diagonal stripe per membrane in spinous part, with blotches on ventral portion of soft part. Anal fin light grey, with spots on last about four membranes. Caudal fin light grey with minute slightly elongate dark dots on anterior half, distally membranes hyaline with grey edges. Pelvic fin light grey, edge and produced portion of first ray white.</paragraph>
<caption id="C6E7009D6F59E47FFF5FF88B95F3F808" ID-DOI="http://doi.org/10.5281/zenodo.8006336" ID-Zenodo-Dep="8006336" httpUri="https://zenodo.org/record/8006336/files/figure.png" pageId="14" pageNumber="181" startId="14.[189,232,1892,1918]" targetBox="[369,1219,265,1851]" targetPageId="14">
<paragraph id="922750156F59E47FFF5FF88B95F3F808" blockId="14.[189,1399,1892,2025]" pageId="14" pageNumber="181">
<emphasis id="A0EC8C076F59E47FFF5FF88B9648F89E" bold="true" box="[189,268,1892,1919]" pageId="14" pageNumber="181">Fig. 7.</emphasis>
<taxonomicName id="55982B966F59E47FFEF7F8899545F89E" authority="Říčan &amp; Říčanová, 2023" authorityName="Říčan &amp; Říčanová" authorityYear="2023" box="[277,513,1894,1919]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="181" phylum="Chordata" rank="species" species="omagua" status="sp. nov.">
<emphasis id="A0EC8C076F59E47FFEF7F8899545F89E" box="[277,513,1894,1919]" italics="true" pageId="14" pageNumber="181">Bujurquina omagua</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBDF317C6F59E47FFDE8F88A9520F89E" box="[522,612,1893,1919]" pageId="14" pageNumber="181" rank="species">sp. nov.</taxonomicNameLabel>
, paratypes. 
<emphasis id="A0EC8C076F59E47FFD17F88B944DF89F" bold="true" box="[757,777,1892,1918]" pageId="14" pageNumber="181">A</emphasis>
. MUSM 70223, P18-16_1186, 92.6 mm. 
<emphasis id="A0EC8C076F59E47FFB1BF88A9249F89E" bold="true" box="[1273,1293,1893,1919]" pageId="14" pageNumber="181">B</emphasis>
. MUSM 70222, P18-16_1185, 79.9 mm. 
<emphasis id="A0EC8C076F59E47FFDA7F867951EF843" bold="true" box="[581,602,1928,1954]" pageId="14" pageNumber="181">C</emphasis>
. MUSM 70221, P18-15_1182, 78.7 mm. 
<emphasis id="A0EC8C076F59E47FFBBAF8679329F843" bold="true" box="[1112,1133,1928,1954]" pageId="14" pageNumber="181">D</emphasis>
. MUSM 70226, P18- 17_1190, 98.9 mm. 
<emphasis id="A0EC8C076F59E47FFE5FF8449695F824" bold="true" box="[445,465,1963,1989]" pageId="14" pageNumber="181">E</emphasis>
. MUSM 70229, P18-21_1200. 
<emphasis id="A0EC8C076F59E47FFC82F8449436F824" bold="true" box="[864,882,1963,1989]" pageId="14" pageNumber="181">F</emphasis>
. MUSM 70227, P18-17_1191, 69.7 mm. 
<emphasis id="A0EC8C076F59E47FFF5FF8219790F809" bold="true" box="[189,212,1998,2024]" pageId="14" pageNumber="181">G</emphasis>
. MUSM 70228, P18-17_1192, 77.1 mm.
</paragraph>
</caption>
<paragraph id="922750156F58E47EFF5FFEE796D6FEC3" blockId="15.[189,402,264,290]" box="[189,402,264,290]" pageId="15" pageNumber="182">
<heading id="C96FE7796F58E47EFF5FFEE796D6FEC3" bold="true" box="[189,402,264,290]" fontSize="11" level="3" pageId="15" pageNumber="182" reason="3">
<emphasis id="A0EC8C076F58E47EFF5FFEE796D6FEC3" bold="true" box="[189,402,264,290]" pageId="15" pageNumber="182">Coloration in life</emphasis>
</heading>
</paragraph>
<paragraph id="922750156F58E47EFF5FFED8940CFDA6" blockId="15.[189,1401,311,583]" pageId="15" pageNumber="182">
Melanin patterns as in preserved. Lower head, cheek, and opercular series yellow-orange, with opalescent golden to azure blue lines and spots, arranged as up to four clearly defined lines between eye and mouth corner and on anterioir cheek interspersed with spots, rest of cheek as well as opercular series richly spotted. Fins orange, pelvic fin yellow, edge and produced portion of first ray white (opalescent). Spinous portion of dorsal fin with a single thin diagonal black line per membrane in juveniles and subadults, and with two lines per membrane in 
<typeStatus id="4D23EEB76F58E47EFDA0FE0895EDFDE0" box="[578,681,487,513]" pageId="15" pageNumber="182" type="holotype">holotype</typeStatus>
and large adults. Lower lip pale greyish to indistinctly light azure blue. Midlateral blotch present only as a paler reflective area within the dark midlateral band (hence absent from coloration of preserved specimens).
</paragraph>
<caption id="C6E7009D6F58E47EFF5FF8449314F808" ID-DOI="http://doi.org/10.5281/zenodo.8006338" ID-Zenodo-Dep="8006338" httpUri="https://zenodo.org/record/8006338/files/figure.png" pageId="15" pageNumber="182" startId="15.[189,232,1963,1989]" targetBox="[189,1398,756,1920]" targetPageId="15">
<paragraph id="922750156F58E47EFF5FF8449314F808" blockId="15.[189,1398,1963,2025]" pageId="15" pageNumber="182">
<emphasis id="A0EC8C076F58E47EFF5FF844964FF824" bold="true" box="[189,267,1963,1989]" pageId="15" pageNumber="182">Fig. 8.</emphasis>
<taxonomicName id="55982B966F58E47EFEF0F84396B8F824" authority="Říčan &amp; Říčanová, 2023" authorityName="Říčan &amp; Říčanová" authorityYear="2023" box="[274,508,1964,1989]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="182" phylum="Chordata" rank="species" species="omagua" status="sp. nov.">
<emphasis id="A0EC8C076F58E47EFEF0F84396B8F824" box="[274,508,1964,1989]" italics="true" pageId="15" pageNumber="182">Bujurquina omagua</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBDF317C6F58E47EFDE0F8439519F827" box="[514,605,1964,1990]" pageId="15" pageNumber="182" rank="species">sp. nov.</taxonomicNameLabel>
, color photos of preserved specimens. 
<emphasis id="A0EC8C076F58E47EFBC6F844937DF824" bold="true" box="[1060,1081,1963,1989]" pageId="15" pageNumber="182">A</emphasis>
. Holotype (MUSM 70225, ID tag 1189), 97.4 mm. 
<emphasis id="A0EC8C076F58E47EFE35F82196AFF809" bold="true" box="[471,491,1998,2024]" pageId="15" pageNumber="182">B</emphasis>
. Paratype (MUSM 70226, P18-17_1190), 98.9 mm.
</paragraph>
</caption>
<caption id="C6E7009D6F47E461FF5FF8D595E1F87A" ID-DOI="http://doi.org/10.5281/zenodo.8006340" ID-Zenodo-Dep="8006340" httpUri="https://zenodo.org/record/8006340/files/figure.png" pageId="16" pageNumber="183" startId="16.[189,232,1850,1876]" targetBox="[189,1398,265,1799]" targetPageId="16">
<paragraph id="922750156F47E461FF5FF8D595E1F87A" blockId="16.[189,1399,1850,1947]" pageId="16" pageNumber="183">
<emphasis id="A0EC8C076F47E461FF5FF8D5964CF8B5" bold="true" box="[189,264,1850,1876]" pageId="16" pageNumber="183">Fig. 9.</emphasis>
<taxonomicName id="55982B966F47E461FEEFF8D496B1F8B5" authority="Říčan &amp; Říčanová, 2023" authorityName="Říčan &amp; Říčanová" authorityYear="2023" box="[269,501,1851,1876]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="183" phylum="Chordata" rank="species" species="omagua" status="sp. nov.">
<emphasis id="A0EC8C076F47E461FEEFF8D496B1F8B5" box="[269,501,1851,1876]" italics="true" pageId="16" pageNumber="183">Bujurquina omagua</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBDF317C6F47E461FE14F8D49515F8B4" box="[502,593,1851,1877]" pageId="16" pageNumber="183" rank="species">sp. nov.</taxonomicNameLabel>
, black and white photos of preserved specimens. 
<emphasis id="A0EC8C076F47E461FB62F8D593D1F8B5" bold="true" box="[1152,1173,1850,1876]" pageId="16" pageNumber="183">A</emphasis>
. Holotype (MUSM 70225, ID tag 1189), 97.4 mm. 
<emphasis id="A0EC8C076F47E461FDA8F8B2951AF896" bold="true" box="[586,606,1885,1911]" pageId="16" pageNumber="183">B</emphasis>
. Paratype (MUSM 70226, P18-17_1190), 98.9 mm. 
<emphasis id="A0EC8C076F47E461FB0AF8B293B9F896" bold="true" box="[1256,1277,1885,1911]" pageId="16" pageNumber="183">C</emphasis>
. Paratype (MUSM 70221, P18-15_1182), 78.7 mm.
</paragraph>
</caption>
<paragraph id="922750156F46E460FF5FFEE79659FEC3" blockId="17.[189,285,264,290]" box="[189,285,264,290]" pageId="17" pageNumber="184">
<heading id="C96FE7796F46E460FF5FFEE79659FEC3" bold="true" box="[189,285,264,290]" fontSize="11" level="3" pageId="17" pageNumber="184" reason="3">
<emphasis id="A0EC8C076F46E460FF5FFEE79659FEC3" bold="true" box="[189,285,264,290]" pageId="17" pageNumber="184">Habitat</emphasis>
</heading>
</paragraph>
<paragraph id="922750156F46E460FF5FFED79687FDE0" blockId="17.[189,1399,311,514]" pageId="17" pageNumber="184">
<taxonomicName id="55982B966F46E460FF5FFED796E3FEB0" authority="Říčan &amp; Říčanová, 2023" authorityName="Říčan &amp; Říčanová" authorityYear="2023" box="[189,423,312,337]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="184" phylum="Chordata" rank="species" species="omagua" status="sp. nov.">
<emphasis id="A0EC8C076F46E460FF5FFED796E3FEB0" box="[189,423,312,337]" italics="true" pageId="17" pageNumber="184">Bujurquina omagua</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBDF317C6F46E460FE4EFED79541FEB3" box="[428,517,312,338]" pageId="17" pageNumber="184" rank="species">sp. nov.</taxonomicNameLabel>
was found only in small rainforest streams within the basins of the Sabalillo and 
<collectingCountry id="EA8F10856F46E460FF0DFEB5966EFE95" box="[239,298,346,372]" name="Algeria" pageId="17" pageNumber="184">Oran</collectingCountry>
creeks. The localities (
<figureCitation id="0AA34C906F46E460FDDFFEB595D5FE95" box="[573,657,346,372]" captionStart="Fig" captionStartId="18.[189,232,1857,1883]" captionTargetBox="[505,1087,267,1816]" captionTargetId="figure-15@18.[501,1087,265,1816]" captionTargetPageId="18" captionText="Fig. 11. Type localities of Bujurquina omagua sp. nov. A–B. P18-15, Qebrada Sabalillo on trail from mouth, (3°27´45.6˝ S, 72°29´22.5˝ W), 27th June 2018. C. P18-16, cabeceras of Quebrada Sabalillo, lower loc. (3°27´17.5˝ S, 72°29´22.8˝ W), 28th June 2018. D. P18-17, holotype locality, cabeceras of Quebrada Sabalillo, upper loc. (3°27´08.4˝ S, 72°29´11.8˝ W), 28th June 2018.E. P18-21, small unnamed quebrada just before entering Oran creek, (3°27´21.2˝ S, 72°30´43.2˝ W), 3rd July 2018." figureDoi="http://doi.org/10.5281/zenodo.8006344" httpUri="https://zenodo.org/record/8006344/files/figure.png" pageId="17" pageNumber="184">Fig. 11</figureCitation>
) were around 
<quantity id="5560FDF06F46E460FCDFFEB5942FFE94" box="[829,875,346,373]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="17" pageNumber="184" unit="m" value="2.0">2 m</quantity>
wide streams, with shallow water (less than 
<quantity id="5560FDF06F46E460FF5FFE9197A8FE79" box="[189,236,382,408]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="17" pageNumber="184" unit="m" value="1.0">1 m</quantity>
except in deeper pools) but deep mud and accumulated debris, blackish water, and only weak or moderate current. 
<taxonomicName id="55982B966F46E460FE6DFE4E953DFE5B" authorityName="Říčan &amp; Říčanová" authorityYear="2023" box="[399,633,417,442]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="184" phylum="Chordata" rank="species" species="omagua">
<emphasis id="A0EC8C076F46E460FE6DFE4E953DFE5B" box="[399,633,417,442]" italics="true" pageId="17" pageNumber="184">Bujurquina omagua</emphasis>
</taxonomicName>
has not been collected from the main streams of the 
<collectingCountry id="EA8F10856F46E460FB34FE4E9255FE5A" box="[1238,1297,417,443]" name="Algeria" pageId="17" pageNumber="184">Oran</collectingCountry>
creek or from the lower sections of the 
<collectingCountry id="EA8F10856F46E460FDC1FE2B951AFE3F" box="[547,606,452,478]" name="Algeria" pageId="17" pageNumber="184">Oran</collectingCountry>
and Sabalillo creeks (
<figureCitation id="0AA34C906F46E460FC80FE2B94FCFE3F" box="[866,952,452,478]" captionStart="Fig" captionStartId="19.[189,232,1892,1918]" captionTargetBox="[499,1089,1062,1848]" captionTargetId="figure-275@19.[499,1089,1062,1848]" captionTargetPageId="19" captionText="Fig. 12. Lower portions of Sabalillo and Oran streams with muddy water with a blackish tinge (June 2018). Bujurquina omagua sp. nov. was not encountered in these habitats, which are typical for B. syspilus (Cope, 1872) in this area (see Fig. 13). A. Quebrada Sabalillo close to mouth into Amazon River. B. Oran creek at mouth of P18-21." figureDoi="http://doi.org/10.5281/zenodo.8006346" httpUri="https://zenodo.org/record/8006346/files/figure.png" pageId="17" pageNumber="184">Fig. 12</figureCitation>
), which have muddy water and where 
<taxonomicName id="55982B966F46E460FF5FFE07967FFDE0" baseAuthorityName="Cope" baseAuthorityYear="1872" box="[189,315,487,513]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="184" phylum="Chordata" rank="species" species="syspilus">
<emphasis id="A0EC8C076F46E460FF5FFE07967FFDE0" box="[189,315,487,513]" italics="true" pageId="17" pageNumber="184">B. syspilus</emphasis>
</taxonomicName>
was found.
</paragraph>
</subSubSection>
<subSubSection id="DA82039E6F46E460FF5FFDC796A2FD3A" pageId="17" pageNumber="184" type="distribution">
<paragraph id="922750156F46E460FF5FFDC79612FDA3" blockId="17.[189,342,552,578]" box="[189,342,552,578]" pageId="17" pageNumber="184">
<heading id="C96FE7796F46E460FF5FFDC79612FDA3" bold="true" box="[189,342,552,578]" fontSize="11" level="3" pageId="17" pageNumber="184" reason="3">
<emphasis id="A0EC8C076F46E460FF5FFDC79612FDA3" bold="true" box="[189,342,552,578]" pageId="17" pageNumber="184">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="922750156F46E460FF5FFDB796A2FD3A" blockId="17.[189,1399,599,731]" pageId="17" pageNumber="184">
<taxonomicName id="55982B966F46E460FF5FFDB796EEFD90" authority="Říčan &amp; Říčanová, 2023" authorityName="Říčan &amp; Říčanová" authorityYear="2023" box="[189,426,600,625]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="184" phylum="Chordata" rank="species" species="omagua" status="sp. nov.">
<emphasis id="A0EC8C076F46E460FF5FFDB796EEFD90" box="[189,426,600,625]" italics="true" pageId="17" pageNumber="184">Bujurquina omagua</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBDF317C6F46E460FE51FDB7954BFD93" box="[435,527,600,626]" pageId="17" pageNumber="184" rank="species">sp. nov.</taxonomicNameLabel>
is only known from the Sabalillo and 
<collectingCountry id="EA8F10856F46E460FC02FDB8935FFD90" box="[992,1051,599,625]" name="Algeria" pageId="17" pageNumber="184">Oran</collectingCountry>
creeks which empty into the Amazon River on either side of the village of 
<collectingCountry id="EA8F10856F46E460FD09FD959462FD75" box="[747,806,634,660]" name="Algeria" pageId="17" pageNumber="184">Oran</collectingCountry>
located on the first high ground just downstream from the mouth of the Napo into the Amazon, District Las Amazonas, Maynas Province, Department 
<collectingRegion id="505C9EF76F46E460FF5FFD2E964AFD3A" box="[189,270,705,731]" country="Peru" name="Loreto" pageId="17" pageNumber="184">Loreto</collectingRegion>
, 
<collectingCountry id="EA8F10856F46E460FEF8FD2E9614FD3A" box="[282,336,705,731]" name="Peru" pageId="17" pageNumber="184">Perú</collectingCountry>
(
<figureCitation id="0AA34C906F46E460FEBDFD2E96EEFD3A" box="[351,426,705,731]" captionStart="Fig" captionStartId="4.[189,232,1857,1883]" captionTargetBox="[189,1398,907,1814]" captionTargetId="figure-237@4.[189,1398,907,1814]" captionTargetPageId="4" captionText="Fig. 1. Distribution and the two main groups of species of Bujurquina Kullander, 1986. Shown with red dotted lines are main geographic drainage basin divides (structural arches) of the western Amazon. Red arrows show the two main subducting ridges responsible for the origin of the formation of the structural arches of the western Amazon. All species were included in morphological analyses, species with an asterisk were also available for molecular analyses. Stars denote type localities for valid species." figureDoi="http://doi.org/10.5281/zenodo.8006322" httpUri="https://zenodo.org/record/8006322/files/figure.png" pageId="17" pageNumber="184">Figs 1</figureCitation>
, 
<figureCitation id="0AA34C906F46E460FE5AFD2E969CFD3A" box="[440,472,705,731]" captionStart="Fig" captionStartId="20.[189,232,1963,1989]" captionTargetBox="[191,1398,570,1920]" captionTargetId="figure-101@20.[189,1398,568,1923]" captionTargetPageId="20" captionText="Fig. 13. Distribution of Bujurquina omagua sp. nov. and parapatric species in the so far whole known area close to Oran village, District Las Amazonas, Maynas Province, Department Loreto, Perú." figureDoi="http://doi.org/10.5281/zenodo.8006348" httpUri="https://zenodo.org/record/8006348/files/figure.png" pageId="17" pageNumber="184">13</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="DA82039E6F46E468FF5FFCEE93DCF8A5" lastPageId="25" lastPageNumber="192" pageId="17" pageNumber="184" type="discussion">
<paragraph id="922750156F46E460FF5FFCEE94E5FCFD" blockId="17.[189,929,769,796]" box="[189,929,769,796]" pageId="17" pageNumber="184">
<heading id="C96FE7796F46E460FF5FFCEE94E5FCFD" bold="true" box="[189,929,769,796]" fontSize="11" level="3" pageId="17" pageNumber="184" reason="3">
<emphasis id="A0EC8C076F46E460FF5FFCEE94E5FCFD" bold="true" box="[189,929,769,796]" italics="true" pageId="17" pageNumber="184">
Morphometric differentiation of 
<taxonomicName id="55982B966F46E460FDAAFCED947BFCFD" authority="Říčan &amp; Říčanová, 2023" authorityName="Říčan &amp; Říčanová" authorityYear="2023" box="[584,831,770,796]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="184" phylum="Chordata" rank="species" species="omagua" status="sp. nov.">Bujurquina omagua</taxonomicName>
<taxonomicNameLabel id="BBDF317C6F46E460FCA4FCED94E5FCFD" box="[838,929,770,796]" pageId="17" pageNumber="184" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="922750156F46E460FF5FFCAB9646FBD0" blockId="17.[189,1399,835,1073]" pageId="17" pageNumber="184">
<taxonomicName id="55982B966F46E460FF5FFCAB96E8FCBC" authority="Říčan &amp; Říčanová, 2023" authorityName="Říčan &amp; Říčanová" authorityYear="2023" box="[189,428,836,861]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="184" phylum="Chordata" rank="species" species="omagua" status="sp. nov.">
<emphasis id="A0EC8C076F46E460FF5FFCAB96E8FCBC" box="[189,428,836,861]" italics="true" pageId="17" pageNumber="184">Bujurquina omagua</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBDF317C6F46E460FE5AFCAB9553FCBF" box="[440,535,836,862]" pageId="17" pageNumber="184" rank="species">sp. nov.</taxonomicNameLabel>
has a highly apomorphic head and body shape that sets it apart from the sympatric/parapatric species. When analyzed through PCA these species are arranged along the main tentic-lotic axis as corresponding to their lowland-upland habitats (
<figureCitation id="0AA34C906F46E460FBC0FC6593E8FC44" box="[1058,1196,906,933]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="21.[189,232,1822,1848]" captionStartId-1="22.[189,232,1787,1813]" captionTargetBox-0="[254,1340,289,1772]" captionTargetBox-1="[261,1333,287,1740]" captionTargetId-0="figure-11@21.[232,1355,265,1778]" captionTargetId-1="figure-15@22.[237,1350,265,1749]" captionTargetPageId-0="21" captionTargetPageId-1="22" captionText-0="Fig. 14. Morphometric variation and discrimination of Northern group species in Bujurquina Kullander, 1986 from the western Amazon river basin based on morphometric proportional values of SL in PCA based on specimens ≥ 60 mm SL. Colors represent individual species. Note main separation along PC1 which also corresponds to two main ecomorphs (lotic and lentic). Note that B. omagua sp. nov. is separated from these geographically closest species by its large head. Note that B. zamorensis (Regan, 1905) is the most lotic and B. syspilus (Cope, 1872) the most lentic species identified by the analysis." captionText-1="Fig. 15. Morphometric variation and discrimination of Northern group species in Bujurquina from the central westernAmazon river basin (i.e., excluding B. zamorensis(Regan, 1905)) based on morphometric proportional values of SL in PCA based on specimens ≥ 60 mm SL. Colors represent individual species. Note main separation along PC1 which also corresponds to two main ecomorphs (lotic and lentic). Note that B. omagua sp. nov. is separated from these geographically closest species by its large head and long snout. Note improved separation of B. huallagae Kullander, 1986, B. ortegai Kullander, 1986 and B. moriorum Kullander, 1986 after exclusion of B. zamorensis (cf. Fig. 14)." figureDoi-0="http://doi.org/10.5281/zenodo.8006350" figureDoi-1="http://doi.org/10.5281/zenodo.8006352" httpUri-0="https://zenodo.org/record/8006350/files/figure.png" httpUri-1="https://zenodo.org/record/8006352/files/figure.png" pageId="17" pageNumber="184">Figs 14–15</figureCitation>
). The lotic/lentic ecomorphology dichotomy is in the PCA analyses best correlated with caudal peduncle length (long vs short), caudal peduncle depth (narrow vs deep), body depth (narrow vs deep), the size of the eye (orbital diameter; small vs large), pectoral fin length (short vs long), and last dorsal fin spine length (short vs long).
</paragraph>
<caption id="C6E7009D6F46E460FF5FF8449429F808" ID-DOI="http://doi.org/10.5281/zenodo.8006342" ID-Zenodo-Dep="8006342" httpUri="https://zenodo.org/record/8006342/files/figure.png" pageId="17" pageNumber="184" startId="17.[189,232,1963,1989]" targetBox="[236,1353,1164,1886]" targetPageId="17">
<paragraph id="922750156F46E460FF5FF8449429F808" blockId="17.[189,1399,1963,2025]" pageId="17" pageNumber="184">
<emphasis id="A0EC8C076F46E460FF5FF8449666F824" bold="true" box="[189,290,1963,1989]" pageId="17" pageNumber="184">Fig. 10.</emphasis>
<taxonomicName id="55982B966F46E460FED3F8439567F824" authority="Říčan &amp; Říčanová, 2023" authorityName="Říčan &amp; Říčanová" authorityYear="2023" box="[305,547,1964,1989]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="184" phylum="Chordata" rank="species" species="omagua" status="sp. nov.">
<emphasis id="A0EC8C076F46E460FED3F8439567F824" box="[305,547,1964,1989]" italics="true" pageId="17" pageNumber="184">Bujurquina omagua</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBDF317C6F46E460FDD0F84395D0F827" box="[562,660,1964,1990]" pageId="17" pageNumber="184" rank="species">sp. nov.</taxonomicNameLabel>
, paratype (MUSM 70222, P18-16_1185, 79.9 mm), lower pharyngeal tooth plate in occlusal view. Scale bar = 1 mm.
</paragraph>
</caption>
<caption id="C6E7009D6F45E463FF5FF8AE9385F809" ID-DOI="http://doi.org/10.5281/zenodo.8006344" ID-Zenodo-Dep="8006344" httpUri="https://zenodo.org/record/8006344/files/figure.png" pageId="18" pageNumber="185" startId="18.[189,232,1857,1883]" targetBox="[505,1087,267,1816]" targetPageId="18">
<paragraph id="922750156F45E463FF5FF8AE9385F809" blockId="18.[189,1399,1857,2026]" pageId="18" pageNumber="185">
<emphasis id="A0EC8C076F45E463FF5FF8AE965EF8BA" bold="true" box="[189,282,1857,1883]" pageId="18" pageNumber="185">Fig. 11.</emphasis>
Type localities of 
<taxonomicName id="55982B966F45E463FE1EF8AD95ADF8BA" authority="Říčan &amp; Říčanová, 2023" authorityName="Říčan &amp; Říčanová" authorityYear="2023" box="[508,745,1858,1883]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="18" pageNumber="185" phylum="Chordata" rank="species" species="omagua" status="sp. nov.">
<emphasis id="A0EC8C076F45E463FE1EF8AD95ADF8BA" box="[508,745,1858,1883]" italics="true" pageId="18" pageNumber="185">Bujurquina omagua</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBDF317C6F45E463FD13F8AD9408F8BD" box="[753,844,1858,1884]" pageId="18" pageNumber="185" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="A0EC8C076F45E463FCB6F8AE94C8F8BA" bold="true" box="[852,908,1857,1883]" pageId="18" pageNumber="185">A–B</emphasis>
. P18-15, Qebrada Sabalillo on trail from mouth, (3° 
<emphasis id="A0EC8C076F45E463FED9F88A96E6F860" bold="true" box="[315,418,1893,1921]" pageId="18" pageNumber="185">27´45.6˝</emphasis>
S, 72° 
<emphasis id="A0EC8C076F45E463FE15F88A9518F860" bold="true" box="[503,604,1893,1921]" pageId="18" pageNumber="185">29´22.5˝</emphasis>
W), 27 
<superScript id="65EDFD5D6F45E463FD58F88B9583F893" attach="left" box="[698,711,1892,1906]" fontSize="6" pageId="18" pageNumber="185">th</superScript>
June 2018. 
<emphasis id="A0EC8C076F45E463FCBCF88B9437F89F" bold="true" box="[862,883,1892,1918]" pageId="18" pageNumber="185">C</emphasis>
. P18-16, cabeceras of Quebrada Sabalillo, lower loc. (3° 
<emphasis id="A0EC8C076F45E463FEBDF8679683F845" bold="true" box="[351,455,1928,1956]" pageId="18" pageNumber="185">27´17.5˝</emphasis>
S, 72° 
<emphasis id="A0EC8C076F45E463FDF8F867953BF845" bold="true" box="[538,639,1928,1956]" pageId="18" pageNumber="185">29´22.8˝</emphasis>
W), 28 
<superScript id="65EDFD5D6F45E463FD3EF86895ADF874" attach="left" box="[732,745,1927,1941]" fontSize="6" pageId="18" pageNumber="185">th</superScript>
June 2018. 
<emphasis id="A0EC8C076F45E463FC63F86794D0F843" bold="true" box="[897,916,1928,1954]" pageId="18" pageNumber="185">D</emphasis>
. P18-17, holotype locality, cabeceras of Quebrada Sabalillo, upper loc. (3° 
<emphasis id="A0EC8C076F45E463FDA1F84495EFF826" bold="true" box="[579,683,1963,1991]" pageId="18" pageNumber="185">27´08.4˝</emphasis>
S, 72° 
<emphasis id="A0EC8C076F45E463FD14F844941EF826" bold="true" box="[758,858,1963,1991]" pageId="18" pageNumber="185">29´11.8˝</emphasis>
W), 28 
<superScript id="65EDFD5D6F45E463FC4CF84594FFF859" attach="left" box="[942,955,1962,1976]" fontSize="6" pageId="18" pageNumber="185">th</superScript>
June 2018. 
<emphasis id="A0EC8C076F45E463FBA0F8449312F824" bold="true" box="[1090,1110,1963,1989]" pageId="18" pageNumber="185">E</emphasis>
. P18-21, small unnamed quebrada just before entering Oran creek, (3° 
<emphasis id="A0EC8C076F45E463FD2EF8219470F80B" bold="true" box="[716,820,1998,2026]" pageId="18" pageNumber="185">27´21.2˝</emphasis>
S, 72° 
<emphasis id="A0EC8C076F45E463FC60F82194ACF80B" bold="true" box="[898,1000,1998,2026]" pageId="18" pageNumber="185">30´43.2˝</emphasis>
W), 3 
<superScript id="65EDFD5D6F45E463FBD3F821937BF83D" attach="left" box="[1073,1087,1998,2012]" fontSize="6" pageId="18" pageNumber="185">rd</superScript>
July 2018.
</paragraph>
</caption>
<paragraph id="922750156F44E462FF5FFEE6961FFE8B" blockId="19.[189,1398,265,362]" pageId="19" pageNumber="186">
The remaining characters are not correlated with the main ecomorphological dichotomy (i.e., interocular distance, head width, head length, preorbital distance, snout length, and first ventral fin ray length) (
<figureCitation id="0AA34C906F44E462FF27FEA0960AFE8B" box="[197,334,335,362]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="21.[189,232,1822,1848]" captionStartId-1="22.[189,232,1787,1813]" captionTargetBox-0="[254,1340,289,1772]" captionTargetBox-1="[261,1333,287,1740]" captionTargetId-0="figure-11@21.[232,1355,265,1778]" captionTargetId-1="figure-15@22.[237,1350,265,1749]" captionTargetPageId-0="21" captionTargetPageId-1="22" captionText-0="Fig. 14. Morphometric variation and discrimination of Northern group species in Bujurquina Kullander, 1986 from the western Amazon river basin based on morphometric proportional values of SL in PCA based on specimens ≥ 60 mm SL. Colors represent individual species. Note main separation along PC1 which also corresponds to two main ecomorphs (lotic and lentic). Note that B. omagua sp. nov. is separated from these geographically closest species by its large head. Note that B. zamorensis (Regan, 1905) is the most lotic and B. syspilus (Cope, 1872) the most lentic species identified by the analysis." captionText-1="Fig. 15. Morphometric variation and discrimination of Northern group species in Bujurquina from the central westernAmazon river basin (i.e., excluding B. zamorensis(Regan, 1905)) based on morphometric proportional values of SL in PCA based on specimens ≥ 60 mm SL. Colors represent individual species. Note main separation along PC1 which also corresponds to two main ecomorphs (lotic and lentic). Note that B. omagua sp. nov. is separated from these geographically closest species by its large head and long snout. Note improved separation of B. huallagae Kullander, 1986, B. ortegai Kullander, 1986 and B. moriorum Kullander, 1986 after exclusion of B. zamorensis (cf. Fig. 14)." figureDoi-0="http://doi.org/10.5281/zenodo.8006350" figureDoi-1="http://doi.org/10.5281/zenodo.8006352" httpUri-0="https://zenodo.org/record/8006350/files/figure.png" httpUri-1="https://zenodo.org/record/8006352/files/figure.png" pageId="19" pageNumber="186">Figs 14–15</figureCitation>
).
</paragraph>
<paragraph id="922750156F44E462FF5FFE7E94DEFD9F" blockId="19.[189,1399,401,639]" pageId="19" pageNumber="186">
The lotic/lentic ecomorphology dichotomy is best visible in the analysis including all central western Amazonian species (
<figureCitation id="0AA34C906F44E462FE52FE5B96BFFE2F" box="[432,507,436,462]" captionStart="Fig" captionStartId="4.[189,232,1857,1883]" captionTargetBox="[189,1398,907,1814]" captionTargetId="figure-237@4.[189,1398,907,1814]" captionTargetPageId="4" captionText="Fig. 1. Distribution and the two main groups of species of Bujurquina Kullander, 1986. Shown with red dotted lines are main geographic drainage basin divides (structural arches) of the western Amazon. Red arrows show the two main subducting ridges responsible for the origin of the formation of the structural arches of the western Amazon. All species were included in morphological analyses, species with an asterisk were also available for molecular analyses. Stars denote type localities for valid species." figureDoi="http://doi.org/10.5281/zenodo.8006322" httpUri="https://zenodo.org/record/8006322/files/figure.png" pageId="19" pageNumber="186">Figs 1</figureCitation>
and 
<figureCitation id="0AA34C906F44E462FDD7FE5B9511FE2F" box="[565,597,436,462]" captionStart="Fig" captionStartId="21.[189,232,1822,1848]" captionTargetBox="[254,1340,289,1772]" captionTargetId="figure-11@21.[232,1355,265,1778]" captionTargetPageId="21" captionText="Fig. 14. Morphometric variation and discrimination of Northern group species in Bujurquina Kullander, 1986 from the western Amazon river basin based on morphometric proportional values of SL in PCA based on specimens ≥ 60 mm SL. Colors represent individual species. Note main separation along PC1 which also corresponds to two main ecomorphs (lotic and lentic). Note that B. omagua sp. nov. is separated from these geographically closest species by its large head. Note that B. zamorensis (Regan, 1905) is the most lotic and B. syspilus (Cope, 1872) the most lentic species identified by the analysis." figureDoi="http://doi.org/10.5281/zenodo.8006350" httpUri="https://zenodo.org/record/8006350/files/figure.png" pageId="19" pageNumber="186">14</figureCitation>
), while the diagnostic characters of 
<taxonomicName id="55982B966F44E462FC1CFE5A93ADFE2F" authority="Říčan &amp; Říčanová, 2023" authorityName="Říčan &amp; Říčanová" authorityYear="2023" box="[1022,1257,437,462]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="186" phylum="Chordata" rank="species" species="omagua" status="sp. nov.">
<emphasis id="A0EC8C076F44E462FC1CFE5A93ADFE2F" box="[1022,1257,437,462]" italics="true" pageId="19" pageNumber="186">Bujurquina omagua</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBDF317C6F44E462FB12FE5A920FFE2E" box="[1264,1355,437,463]" pageId="19" pageNumber="186" rank="species">sp. nov.</taxonomicNameLabel>
are best visible in the analysis excluding the westernmost and most highland species 
<taxonomicName id="55982B966F44E462FB80FE379240FE10" authorityName="" baseAuthorityName="Regan" baseAuthorityYear="1905" box="[1122,1284,472,497]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="186" phylum="Chordata" rank="species" species="zamorensis">
<emphasis id="A0EC8C076F44E462FB80FE379240FE10" box="[1122,1284,472,497]" italics="true" pageId="19" pageNumber="186">B. zamorensis</emphasis>
</taxonomicName>
(
<figureCitation id="0AA34C906F44E462FAF1FE38922CFE13" box="[1299,1384,471,498]" captionStart="Fig" captionStartId="22.[189,232,1787,1813]" captionTargetBox="[261,1333,287,1740]" captionTargetId="figure-15@22.[237,1350,265,1749]" captionTargetPageId="22" captionText="Fig. 15. Morphometric variation and discrimination of Northern group species in Bujurquina from the central westernAmazon river basin (i.e., excluding B. zamorensis(Regan, 1905)) based on morphometric proportional values of SL in PCA based on specimens ≥ 60 mm SL. Colors represent individual species. Note main separation along PC1 which also corresponds to two main ecomorphs (lotic and lentic). Note that B. omagua sp. nov. is separated from these geographically closest species by its large head and long snout. Note improved separation of B. huallagae Kullander, 1986, B. ortegai Kullander, 1986 and B. moriorum Kullander, 1986 after exclusion of B. zamorensis (cf. Fig. 14)." figureDoi="http://doi.org/10.5281/zenodo.8006352" httpUri="https://zenodo.org/record/8006352/files/figure.png" pageId="19" pageNumber="186">Fig. 15</figureCitation>
). 
<taxonomicName id="55982B966F44E462FF5FFE1496EDFDF5" authorityName="Říčan &amp; Říčanová" authorityYear="2023" box="[189,425,507,532]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="186" phylum="Chordata" rank="species" species="omagua">
<emphasis id="A0EC8C076F44E462FF5FFE1496EDFDF5" box="[189,425,507,532]" italics="true" pageId="19" pageNumber="186">Bujurquina omagua</emphasis>
</taxonomicName>
is in the PCAs distinguished from the sympatric/parapatric species predominantly by characters not correlated with the lotic-lentic, namely large head, wide head, long snout, and long preorbital dostance. All these diagnostic characters are in line with the diagnosis of the species based on means and ranges of the morphometric characters (see above).
</paragraph>
<paragraph id="922750156F44E462FF5FFD49930AFC8E" blockId="19.[189,1399,677,880]" pageId="19" pageNumber="186">
<taxonomicName id="55982B966F44E462FF5FFD4996EDFD5E" authority="Říčan &amp; Říčanová, 2023" authorityName="Říčan &amp; Říčanová" authorityYear="2023" box="[189,425,678,703]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="186" phylum="Chordata" rank="species" species="omagua" status="sp. nov.">
<emphasis id="A0EC8C076F44E462FF5FFD4996EDFD5E" box="[189,425,678,703]" italics="true" pageId="19" pageNumber="186">Bujurquina omagua</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBDF317C6F44E462FE53FD499549FD21" box="[433,525,678,704]" pageId="19" pageNumber="186" rank="species">sp. nov.</taxonomicNameLabel>
has an outlying LPJ shape that can be described as the most delicate LPJ found among the valid species of 
<taxonomicName id="55982B966F44E462FDBCFD2695A7FD03" authorityName="Kullander" authorityYear="1986" box="[606,739,713,738]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="186" phylum="Chordata" rank="genus">
<emphasis id="A0EC8C076F44E462FDBCFD2695A7FD03" box="[606,739,713,738]" italics="true" pageId="19" pageNumber="186">Bujurquina</emphasis>
</taxonomicName>
(
<figureCitation id="0AA34C906F44E462FD14FD269415FD03" box="[758,849,712,739]" captionStart="Fig" captionStartId="23.[189,232,1894,1920]" captionTargetBox="[205,1385,283,1835]" captionTargetId="figure-12@23.[189,1398,265,1845]" captionTargetPageId="23" captionText="Fig. 16. PCA based on morphometric proportional values of LPJ of all described species in Bujurquina Kullander, 1986 based on specimens ≥60 mm SL. Colors represent individual species. Note that B. omagua sp. nov. has the most outlying LPJ shape." figureDoi="http://doi.org/10.5281/zenodo.8006354" httpUri="https://zenodo.org/record/8006354/files/figure.png" pageId="19" pageNumber="186">Fig. 16</figureCitation>
). The LPJ of 
<taxonomicName id="55982B966F44E462FC1FFD2693A8FD03" authorityName="Říčan &amp; Říčanová" authorityYear="2023" box="[1021,1260,713,738]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="186" phylum="Chordata" rank="species" species="omagua">
<emphasis id="A0EC8C076F44E462FC1FFD2693A8FD03" box="[1021,1260,713,738]" italics="true" pageId="19" pageNumber="186">Bujurquina omagua</emphasis>
</taxonomicName>
(studied in 
<specimenCount id="849E9B9C6F44E462FF5FFD03962AFCE7" box="[189,366,748,774]" pageId="19" pageNumber="186" type="generic">two specimens</specimenCount>
, see above) as expressed by proportional measurements is the shortest and widest of all species (L/W ratio) and the LPJ tooth-plate is the shortest in the genus (Ld/W ratio). Additionaly to this the LPJ is very shallow vertically dorso-ventrally, but this character was scored as binary, in comparison to the length of teeth (see Results), and thus not included in the PCA analysis.
</paragraph>
<paragraph id="922750156F44E465FF5FFC78958EFEA7" blockId="19.[189,1399,919,981]" lastBlockId="20.[189,1398,265,327]" lastPageId="20" lastPageNumber="187" pageId="19" pageNumber="186">
Paradoxically, the most delicate LPJ of 
<taxonomicName id="55982B966F44E462FD74FC779452FC50" authority="Říčan &amp; Říčanová, 2023" authorityName="Říčan &amp; Říčanová" authorityYear="2023" box="[662,790,920,945]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="186" phylum="Chordata" rank="species" species="omagua" status="sp. nov.">
<emphasis id="A0EC8C076F44E462FD74FC779452FC50" box="[662,790,920,945]" italics="true" pageId="19" pageNumber="186">B. omagua</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBDF317C6F44E462FCFDFC78943FFC50" box="[799,891,919,945]" pageId="19" pageNumber="186" rank="species">sp. nov.</taxonomicNameLabel>
is associated with one of the largest heads among species of 
<taxonomicName id="55982B966F44E462FE7AFC549559FC35" authorityName="Kullander" authorityYear="1986" box="[408,541,955,980]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="186" phylum="Chordata" rank="genus">
<emphasis id="A0EC8C076F44E462FE7AFC549559FC35" box="[408,541,955,980]" italics="true" pageId="19" pageNumber="186">Bujurquina</emphasis>
</taxonomicName>
(together with 
<taxonomicName id="55982B966F44E462FD37FC54940AFC35" authorityName="Kullander" authorityYear="1986" box="[725,846,954,980]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="186" phylum="Chordata" rank="species" species="labiosa">
<emphasis id="A0EC8C076F44E462FD37FC54940AFC35" box="[725,846,954,980]" italics="true" pageId="19" pageNumber="186">B. labiosa</emphasis>
</taxonomicName>
) and with the longest snout among species of 
<taxonomicName id="55982B966F43E465FF5FFEE59606FEC2" authorityName="Kullander" authorityYear="1986" box="[189,322,266,291]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="20" pageNumber="187" phylum="Chordata" rank="genus">
<emphasis id="A0EC8C076F43E465FF5FFEE59606FEC2" box="[189,322,266,291]" italics="true" pageId="20" pageNumber="187">Bujurquina</emphasis>
</taxonomicName>
. A similar head and snout shape in 
<taxonomicName id="55982B966F43E465FD34FEE594C6FEC2" authorityName="Kullander" authorityYear="1987" box="[726,898,265,291]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="20" pageNumber="187" phylum="Chordata" rank="species" species="oenolaemus">
<emphasis id="A0EC8C076F43E465FD34FEE594C6FEC2" box="[726,898,265,291]" italics="true" pageId="20" pageNumber="187">B. oenolaemus</emphasis>
</taxonomicName>
is however associated with the most robust LPJ (
<figureCitation id="0AA34C906F43E465FF19FEC39614FEA7" box="[251,336,300,326]" captionStart="Fig" captionStartId="23.[189,232,1894,1920]" captionTargetBox="[205,1385,283,1835]" captionTargetId="figure-12@23.[189,1398,265,1845]" captionTargetPageId="23" captionText="Fig. 16. PCA based on morphometric proportional values of LPJ of all described species in Bujurquina Kullander, 1986 based on specimens ≥60 mm SL. Colors represent individual species. Note that B. omagua sp. nov. has the most outlying LPJ shape." figureDoi="http://doi.org/10.5281/zenodo.8006354" httpUri="https://zenodo.org/record/8006354/files/figure.png" pageId="20" pageNumber="187">Fig. 16</figureCitation>
) of this molluscivorous species.
</paragraph>
<caption id="C6E7009D6F44E462FF5FF88B95E5F809" ID-DOI="http://doi.org/10.5281/zenodo.8006346" ID-Zenodo-Dep="8006346" httpUri="https://zenodo.org/record/8006346/files/figure.png" pageId="19" pageNumber="186" startId="19.[189,232,1892,1918]" targetBox="[499,1089,1062,1848]" targetPageId="19">
<paragraph id="922750156F44E462FF5FF88B95E5F809" blockId="19.[189,1399,1892,2025]" pageId="19" pageNumber="186">
<emphasis id="A0EC8C076F44E462FF5FF88B9658F89E" bold="true" box="[189,284,1892,1919]" pageId="19" pageNumber="186">Fig. 12.</emphasis>
Lower portions of Sabalillo and Oran streams with muddy water with a blackish tinge (June 2018). 
<taxonomicName id="55982B966F44E462FEF5F866954CF843" authority="Říčan &amp; Říčanová, 2023" authorityName="Říčan &amp; Říčanová" authorityYear="2023" box="[279,520,1929,1954]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="186" phylum="Chordata" rank="species" species="omagua" status="sp. nov.">
<emphasis id="A0EC8C076F44E462FEF5F866954CF843" box="[279,520,1929,1954]" italics="true" pageId="19" pageNumber="186">Bujurquina omagua</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBDF317C6F44E462FDF7F8679533F843" box="[533,631,1928,1954]" pageId="19" pageNumber="186" rank="species">sp. nov.</taxonomicNameLabel>
was not encountered in these habitats, which are typical for 
<taxonomicName id="55982B966F44E462FF5FF843969BF824" authority="(Cope, 1872)" baseAuthorityName="Cope" baseAuthorityYear="1872" box="[189,479,1963,1989]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="186" phylum="Chordata" rank="species" species="syspilus">
<emphasis id="A0EC8C076F44E462FF5FF8439678F824" box="[189,316,1963,1989]" italics="true" pageId="19" pageNumber="186">B. syspilus</emphasis>
(Cope, 1872)
</taxonomicName>
in this area (see Fig. 13). 
<emphasis id="A0EC8C076F44E462FCFAF8449469F824" bold="true" box="[792,813,1963,1989]" pageId="19" pageNumber="186">A</emphasis>
. Quebrada Sabalillo close to mouth into Amazon River. 
<emphasis id="A0EC8C076F44E462FEE9F821965BF809" bold="true" box="[267,287,1998,2024]" pageId="19" pageNumber="186">B</emphasis>
. Oran creek at mouth of P18-21.
</paragraph>
</caption>
<paragraph id="922750156F43E465FF5FFE819594FE10" blockId="20.[189,1399,366,500]" pageId="20" pageNumber="187">
A similar combination of a long snout and large head with a very delicate LPJ as in 
<taxonomicName id="55982B966F43E465FB77FE819251FE66" authority="Říčan &amp; Říčanová, 2023" authorityName="Říčan &amp; Říčanová" authorityYear="2023" box="[1173,1301,366,391]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="20" pageNumber="187" phylum="Chordata" rank="species" species="omagua" status="sp. nov.">
<emphasis id="A0EC8C076F43E465FB77FE819251FE66" box="[1173,1301,366,391]" italics="true" pageId="20" pageNumber="187">B. omagua</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBDF317C6F43E465FAFEFE819233FE69" box="[1308,1399,366,392]" pageId="20" pageNumber="187" rank="species">sp. nov.</taxonomicNameLabel>
is however also known from the cichlid genus 
<taxonomicName id="55982B966F43E465FD12FE7E9388FE4A" authority="Rican &amp; Kullander, 2006" authorityName="Rican &amp; Kullander" authorityYear="2006" box="[752,1228,401,429]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="20" pageNumber="187" phylum="Chordata" rank="genus">
<emphasis id="A0EC8C076F43E465FD12FE7E94D1FE4A" box="[752,917,401,427]" italics="true" pageId="20" pageNumber="187">Australoheros</emphasis>
Říčan &amp; Kullander, 
<quantity id="5560FDF06F43E465FB73FE7E9388FE4A" box="[1169,1228,401,427]" metricMagnitude="1" metricUnit="m" metricValue="5.0952399999999995" pageId="20" pageNumber="187" unit="in" value="2006.0">2006</quantity>
</taxonomicName>
in the species 
<taxonomicName id="55982B966F43E465FF5FFE5B95CBFE2F" authority="Rican &amp; Kullander, 2008" authorityName="Rican &amp; Kullander" authorityYear="2008" box="[189,655,436,465]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="20" pageNumber="187" phylum="Chordata" rank="species" species="forquilha">
<emphasis id="A0EC8C076F43E465FF5FFE5B960AFE2F" box="[189,334,436,462]" italics="true" pageId="20" pageNumber="187">A. forquilha</emphasis>
<bibRefCitation id="F6092DE46F43E465FEB8FE5A95CBFE2F" author="Rican O. &amp; Kullander S. O." box="[346,655,436,465]" pageId="20" pageNumber="187" pagination="1 - 51" refId="ref15361" refString="Rican O. &amp; Kullander S. O. 2008. The Australoheros (Teleostei: Cichlidae) species of the Uruguay and Parana River drainages. Zootaxa 1724: 1 - 51. https: // doi. org / 10.11646 / zootaxa. 1724.1.1" type="journal article" year="2008">Říčan &amp; Kullander, 2008</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="55982B966F43E465FD32FE5B9233FE2F" authority="Rican, Pialeck, Almiron &amp; Casciotta, 2011" authorityName="Rican, Pialeck, Almiron &amp; Casciotta" authorityYear="2011" box="[720,1399,436,465]" class="Actinopterygii" family="Cichlidae" genus="Australoheros" kingdom="Animalia" order="Perciformes" pageId="20" pageNumber="187" phylum="Chordata" rank="species" species="ykeregua">
<emphasis id="A0EC8C076F43E465FD32FE5B941BFE2F" box="[720,863,436,462]" italics="true" pageId="20" pageNumber="187">A. ykeregua</emphasis>
Říčan, Piáleck, Almirón &amp; Casciotta, 2011
</taxonomicName>
(
<bibRefCitation id="F6092DE46F43E465FF27FE3796AFFE10" author="Rican O. &amp; Kullander S. O." box="[197,491,471,500]" pageId="20" pageNumber="187" pagination="1 - 51" refId="ref15361" refString="Rican O. &amp; Kullander S. O. 2008. The Australoheros (Teleostei: Cichlidae) species of the Uruguay and Parana River drainages. Zootaxa 1724: 1 - 51. https: // doi. org / 10.11646 / zootaxa. 1724.1.1" type="journal article" year="2008">Říčan &amp; Kullander 2008</bibRefCitation>
; 
<bibRefCitation id="F6092DE46F43E465FE15FE379587FE10" author="Rican O. &amp; Pialek L. &amp; Almiron A. &amp; Casciotta J." box="[503,707,471,500]" pageId="20" pageNumber="187" pagination="1 - 26" refId="ref15510" refString="Rican O., Pialek L., Almiron A. &amp; Casciotta J. 2011. Two new species of Australoheros (Teleostei: Cichlidae), with notes on diversity of the genus and biogeography of the Rio de la Plata basin. Zootaxa 2982: 1 - 26. https: // doi. org / 10.11646 / zootaxa. 2982.1.1" type="journal article" year="2011">
Říčan 
<emphasis id="A0EC8C076F43E465FDA1FE37953AFE10" box="[579,638,471,497]" italics="true" pageId="20" pageNumber="187">et al.</emphasis>
2011
</bibRefCitation>
).
</paragraph>
<caption id="C6E7009D6F43E465FF5FF844925DF809" ID-DOI="http://doi.org/10.5281/zenodo.8006348" ID-Zenodo-Dep="8006348" httpUri="https://zenodo.org/record/8006348/files/figure.png" pageId="20" pageNumber="187" startId="20.[189,232,1963,1989]" targetBox="[191,1398,570,1920]" targetPageId="20">
<paragraph id="922750156F43E465FF5FF844925DF809" blockId="20.[189,1399,1963,2025]" pageId="20" pageNumber="187">
<emphasis id="A0EC8C076F43E465FF5FF844965FF824" bold="true" box="[189,283,1963,1989]" pageId="20" pageNumber="187">Fig. 13.</emphasis>
Distribution of 
<taxonomicName id="55982B966F43E465FE39F8439583F824" authority="Říčan &amp; Říčanová, 2023" authorityName="Říčan &amp; Říčanová" authorityYear="2023" box="[475,711,1964,1989]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="20" pageNumber="187" phylum="Chordata" rank="species" species="omagua" status="sp. nov.">
<emphasis id="A0EC8C076F43E465FE39F8439583F824" box="[475,711,1964,1989]" italics="true" pageId="20" pageNumber="187">Bujurquina omagua</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBDF317C6F43E465FD2DF843946EF827" box="[719,810,1964,1990]" pageId="20" pageNumber="187" rank="species">sp. nov.</taxonomicNameLabel>
and parapatric species in the so far whole known area close to Oran village, District Las Amazonas, Maynas Province, Department Loreto, Perú.
</paragraph>
</caption>
<caption id="C6E7009D6F42E464FF5FF8F19224F809" ID-DOI="http://doi.org/10.5281/zenodo.8006350" ID-Zenodo-Dep="8006350" httpUri="https://zenodo.org/record/8006350/files/figure.png" pageId="21" pageNumber="188" startId="21.[189,232,1822,1848]" targetBox="[254,1340,289,1772]" targetPageId="21">
<paragraph id="922750156F42E464FF5FF8F19224F809" blockId="21.[188,1399,1822,2025]" pageId="21" pageNumber="188">
<emphasis id="A0EC8C076F42E464FF5FF8F1965DF8D9" bold="true" box="[189,281,1822,1848]" pageId="21" pageNumber="188">Fig. 14.</emphasis>
Morphometric variation and discrimination of Northern group species in 
<taxonomicName id="55982B966F42E464FB92F8F097B3F8BA" authority="Kullander, 1986" authorityName="Kullander" authorityYear="1986" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="21" pageNumber="188" phylum="Chordata" rank="genus">
<emphasis id="A0EC8C076F42E464FB92F8F093B1F8D9" box="[1136,1269,1823,1848]" italics="true" pageId="21" pageNumber="188">Bujurquina</emphasis>
<bibRefCitation id="F6092DE46F42E464FB19F8F197B3F8BA" author="Kullander S." pageId="21" pageNumber="188" refId="ref14485" refString="Kullander S. 1986. Cichlid Fishes of the Amazon River Drainage of Peru. Swedish Museum of Natural History, Sweden." type="book" year="1986">Kullander, 1986</bibRefCitation>
</taxonomicName>
from the western Amazon river basin based on morphometric proportional values of SL in PCA based on specimens ≥ 60 mm SL. Colors represent individual species. Note main separation along PC1 which also corresponds to two main ecomorphs (lotic and lentic). Note that 
<taxonomicName id="55982B966F42E464FB88F86693AAF843" authority="Říčan &amp; Říčanová, 2023" authorityName="Říčan &amp; Říčanová" authorityYear="2023" box="[1130,1262,1929,1954]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="21" pageNumber="188" phylum="Chordata" rank="species" species="omagua" status="sp. nov.">
<emphasis id="A0EC8C076F42E464FB88F86693AAF843" box="[1130,1262,1929,1954]" italics="true" pageId="21" pageNumber="188">B. omagua</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBDF317C6F42E464FB1BF867921CF843" box="[1273,1368,1928,1954]" pageId="21" pageNumber="188" rank="species">sp. nov.</taxonomicNameLabel>
is separated from these geographically closest species by its large head. Note that 
<taxonomicName id="55982B966F42E464FB8CF8439645F808" authority="(Regan, 1905)" authorityName="" baseAuthorityName="Regan" baseAuthorityYear="1905" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="21" pageNumber="188" phylum="Chordata" rank="species" species="zamorensis">
<emphasis id="A0EC8C076F42E464FB8CF8439256F824" box="[1134,1298,1964,1989]" italics="true" pageId="21" pageNumber="188">B. zamorensis</emphasis>
(Regan, 1905)
</taxonomicName>
is the most lotic and 
<taxonomicName id="55982B966F42E464FE1EF8209458F809" authority="(Cope, 1872)" baseAuthorityName="Cope" baseAuthorityYear="1872" box="[508,796,1998,2024]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="21" pageNumber="188" phylum="Chordata" rank="species" species="syspilus">
<emphasis id="A0EC8C076F42E464FE1EF820953EF809" box="[508,634,1998,2024]" italics="true" pageId="21" pageNumber="188">B. syspilus</emphasis>
(Cope, 1872)
</taxonomicName>
the most lentic species identified by the analysis.
</paragraph>
</caption>
<caption id="C6E7009D6F41E467FF5FF9149376F809" ID-DOI="http://doi.org/10.5281/zenodo.8006352" ID-Zenodo-Dep="8006352" httpUri="https://zenodo.org/record/8006352/files/figure.png" pageId="22" pageNumber="189" startId="22.[189,232,1787,1813]" targetBox="[261,1333,287,1740]" targetPageId="22">
<paragraph id="922750156F41E467FF5FF9149376F809" blockId="22.[189,1399,1787,2024]" pageId="22" pageNumber="189">
<emphasis id="A0EC8C076F41E467FF5FF914965FF8F4" bold="true" box="[189,283,1787,1813]" pageId="22" pageNumber="189">Fig. 15.</emphasis>
Morphometric variation and discrimination of Northern group species in 
<taxonomicName id="55982B966F41E467FB67F913924EF8F4" authorityName="Kullander" authorityYear="1986" box="[1157,1290,1788,1813]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="22" pageNumber="189" phylum="Chordata" rank="genus">
<emphasis id="A0EC8C076F41E467FB67F913924EF8F4" box="[1157,1290,1788,1813]" italics="true" pageId="22" pageNumber="189">Bujurquina</emphasis>
</taxonomicName>
from the central western Amazon river basin (i.e., excluding 
<taxonomicName id="55982B966F41E467FCE9F8F0931BF8D8" authority="(Regan, 1905))" authorityName="" baseAuthorityName="Regan" baseAuthorityYear="1905" box="[779,1119,1822,1849]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="22" pageNumber="189" phylum="Chordata" rank="species" species="zamorensis">
<emphasis id="A0EC8C076F41E467FCE9F8F094E9F8D9" box="[779,941,1823,1848]" italics="true" pageId="22" pageNumber="189">B. zamorensis</emphasis>
(Regan, 1905))
</taxonomicName>
based on morphometric proportional values of SL in PCA based on specimens ≥ 60 mm SL. Colors represent individual species. Note main separation along PC1 which also corresponds to two main ecomorphs (lotic and lentic). Note that 
<taxonomicName id="55982B966F41E467FF11F8669632F843" authority="Říčan &amp; Říčanová, 2023" authorityName="Říčan &amp; Říčanová" authorityYear="2023" box="[243,374,1929,1954]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="22" pageNumber="189" phylum="Chordata" rank="species" species="omagua" status="sp. nov.">
<emphasis id="A0EC8C076F41E467FF11F8669632F843" box="[243,374,1929,1954]" italics="true" pageId="22" pageNumber="189">B. omagua</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBDF317C6F41E467FE9DF8679699F843" box="[383,477,1928,1954]" pageId="22" pageNumber="189" rank="species">sp. nov.</taxonomicNameLabel>
is separated from these geographically closest species by its large head and long snout. Note improved separation of 
<taxonomicName id="55982B966F41E467FD42F84394B9F824" authority="Kullander, 1986" authorityName="Kullander" authorityYear="1986" box="[672,1021,1963,1989]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="22" pageNumber="189" phylum="Chordata" rank="species" species="huallagae">
<emphasis id="A0EC8C076F41E467FD42F8439471F824" box="[672,821,1963,1989]" italics="true" pageId="22" pageNumber="189">B. huallagae</emphasis>
<bibRefCitation id="F6092DE46F41E467FCDCF84494B9F824" author="Kullander S." box="[830,1021,1963,1989]" pageId="22" pageNumber="189" refId="ref14485" refString="Kullander S. 1986. Cichlid Fishes of the Amazon River Drainage of Peru. Swedish Museum of Natural History, Sweden." type="book" year="1986">Kullander, 1986</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="55982B966F41E467FBE8F8439201F824" authority="Kullander, 1986" authorityName="Kullander" authorityYear="1986" box="[1034,1349,1963,1989]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="22" pageNumber="189" phylum="Chordata" rank="species" species="ortegai">
<emphasis id="A0EC8C076F41E467FBE8F84393C4F824" box="[1034,1152,1964,1989]" italics="true" pageId="22" pageNumber="189">B. ortegai</emphasis>
<bibRefCitation id="F6092DE46F41E467FB65F8449201F824" author="Kullander S." box="[1159,1349,1963,1989]" pageId="22" pageNumber="189" refId="ref14485" refString="Kullander S. 1986. Cichlid Fishes of the Amazon River Drainage of Peru. Swedish Museum of Natural History, Sweden." type="book" year="1986">Kullander, 1986</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="55982B966F41E467FF5FF820955CF809" authority="Kullander, 1986" authorityName="Kullander" authorityYear="1986" box="[189,536,1998,2024]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="22" pageNumber="189" phylum="Chordata" rank="species" species="moriorum">
<emphasis id="A0EC8C076F41E467FF5FF8209617F809" box="[189,339,1999,2024]" italics="true" pageId="22" pageNumber="189">B. moriorum</emphasis>
<bibRefCitation id="F6092DE46F41E467FEB8F821955CF809" author="Kullander S." box="[346,536,1998,2024]" pageId="22" pageNumber="189" refId="ref14485" refString="Kullander S. 1986. Cichlid Fishes of the Amazon River Drainage of Peru. Swedish Museum of Natural History, Sweden." type="book" year="1986">Kullander, 1986</bibRefCitation>
</taxonomicName>
after exclusion of 
<taxonomicName id="55982B966F41E467FD16F82094D3F809" authorityName="" baseAuthorityName="Regan" baseAuthorityYear="1905" box="[756,919,1999,2024]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="22" pageNumber="189" phylum="Chordata" rank="species" species="zamorensis">
<emphasis id="A0EC8C076F41E467FD16F82094D3F809" box="[756,919,1999,2024]" italics="true" pageId="22" pageNumber="189">B. zamorensis</emphasis>
</taxonomicName>
(cf. Fig. 14).
</paragraph>
</caption>
<caption id="C6E7009D6F40E466FF5FF889946CF827" ID-DOI="http://doi.org/10.5281/zenodo.8006354" ID-Zenodo-Dep="8006354" httpUri="https://zenodo.org/record/8006354/files/figure.png" pageId="23" pageNumber="190" startId="23.[189,232,1894,1920]" targetBox="[205,1385,283,1835]" targetPageId="23">
<paragraph id="922750156F40E466FF5FF889946CF827" blockId="23.[189,1399,1894,1991]" pageId="23" pageNumber="190">
<emphasis id="A0EC8C076F40E466FF5FF889965EF861" bold="true" box="[189,282,1894,1920]" pageId="23" pageNumber="190">Fig. 16.</emphasis>
PCA based on morphometric proportional values of LPJ of all described species in 
<taxonomicName id="55982B966F40E466FB10F88896C5F842" authority="Kullander, 1986" authorityName="Kullander" authorityYear="1986" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="23" pageNumber="190" phylum="Chordata" rank="genus">
<emphasis id="A0EC8C076F40E466FB10F8889233F861" box="[1266,1399,1895,1920]" italics="true" pageId="23" pageNumber="190">Bujurquina</emphasis>
<bibRefCitation id="F6092DE46F40E466FF5FF86696C5F842" author="Kullander S." box="[189,385,1929,1955]" pageId="23" pageNumber="190" refId="ref14485" refString="Kullander S. 1986. Cichlid Fishes of the Amazon River Drainage of Peru. Swedish Museum of Natural History, Sweden." type="book" year="1986">Kullander, 1986</bibRefCitation>
</taxonomicName>
based on specimens ≥60 mm SL. Colors represent individual species. Note that 
<taxonomicName id="55982B966F40E466FF5FF8429679F827" authority="Říčan &amp; Říčanová, 2023" authorityName="Říčan &amp; Říčanová" authorityYear="2023" box="[189,317,1965,1990]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="23" pageNumber="190" phylum="Chordata" rank="species" species="omagua" status="sp. nov.">
<emphasis id="A0EC8C076F40E466FF5FF8429679F827" box="[189,317,1965,1990]" italics="true" pageId="23" pageNumber="190">B. omagua</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBDF317C6F40E466FEA1F84296DAF826" box="[323,414,1965,1991]" pageId="23" pageNumber="190" rank="species">sp. nov.</taxonomicNameLabel>
has the most outlying LPJ shape.
</paragraph>
</caption>
<paragraph id="922750156F4FE469FF5FFEE7924EFEC4" blockId="24.[189,1290,264,293]" box="[189,1290,264,293]" pageId="24" pageNumber="191">
<emphasis id="A0EC8C076F4FE469FF5FFEE7924EFEC4" bold="true" box="[189,1290,264,293]" italics="true" pageId="24" pageNumber="191">
Phylogeny of 
<taxonomicName id="55982B966F4FE469FE90FEE79549FEC4" authorityName="Kullander" authorityYear="1986" box="[370,525,264,293]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="24" pageNumber="191" phylum="Chordata" rank="genus">Bujurquina</taxonomicName>
and phylogenetic position of 
<taxonomicName id="55982B966F4FE469FC70FEE793DBFEC4" authority="Říčan &amp; Říčanová, 2023" authorityName="Říčan &amp; Říčanová" authorityYear="2023" box="[914,1183,264,293]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="24" pageNumber="191" phylum="Chordata" rank="species" species="omagua" status="sp. nov.">Bujurquina omagua</taxonomicName>
<taxonomicNameLabel id="BBDF317C6F4FE469FB45FEE6924EFEC4" box="[1191,1290,265,293]" pageId="24" pageNumber="191" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="922750156F4FE469FF5FFED296E0FDAC" blockId="24.[189,1399,316,589]" pageId="24" pageNumber="191">
Phylogenetic analyses of the 1055 characters 
<emphasis id="A0EC8C076F4FE469FD30FED29447FEB6" box="[722,771,317,343]" italics="true" pageId="24" pageNumber="191">cytb</emphasis>
data matrix performed with MP in PAUP* (including all ingroup sequences) and BI analyses in MrBayes and BEAST (performed with the corresponding haplotypes data matrix) provided robust and very similar results. BI runs in independent analyses with the best-fitting model of evolution (GTR+I+G) converged well (ESS&gt;200 for all parameters; burn-in 10%) and led to identical topologies under the same settings. The NJ and MP analyses were done with the complete dataset, the BI and BEAST analyses with the haplotype dataset. No phylogenetic differences were found between the MP, BI and BEAST analyses, and hence only the BEAST analysis is shown in 
<figureCitation id="0AA34C906F4FE469FEA8FDDC96E4FDAC" box="[330,416,563,589]" captionStart="Fig" captionStartId="24.[189,232,1892,1918]" captionTargetBox="[190,1398,1076,1839]" captionTargetId="figure-305@24.[189,1398,1072,1852]" captionTargetPageId="24" captionText="Fig. 17. Phylogenetic relationships and species delimitation within Bujurquina Kullander, 1986 based on the cytb marker from BEAST analysis. Numbers at nodes show age in Ma, grey bars at nodes show confidence intervals of age estimates (95% HPD). Bayesian posterior probabilities above 0.95 are shown as black points on nodes. Scale bar shows 5 My. Note the two main groups of Bujurquina." figureDoi="http://doi.org/10.5281/zenodo.8006356" httpUri="https://zenodo.org/record/8006356/files/figure.png" pageId="24" pageNumber="191">Fig. 17</figureCitation>
.
</paragraph>
<paragraph id="922750156F4FE469FF5FFD9B94F1FD19" blockId="24.[189,1399,628,761]" pageId="24" pageNumber="191">
The phylogenetic reconstructions divide 
<taxonomicName id="55982B966F4FE469FD49FD9A9474FD6F" authorityName="Kullander" authorityYear="1986" box="[683,816,629,654]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="24" pageNumber="191" phylum="Chordata" rank="genus">
<emphasis id="A0EC8C076F4FE469FD49FD9A9474FD6F" box="[683,816,629,654]" italics="true" pageId="24" pageNumber="191">Bujurquina</emphasis>
</taxonomicName>
into two main clades that can be diagnosed by patterned vs unpatterned dorsal fin, corresponding in geographical terms to Northern vs Southern group (
<figureCitation id="0AA34C906F4FE469FF27FD54964AFD34" box="[197,270,699,725]" captionStart="Fig" captionStartId="4.[189,232,1857,1883]" captionTargetBox="[189,1398,907,1814]" captionTargetId="figure-237@4.[189,1398,907,1814]" captionTargetPageId="4" captionText="Fig. 1. Distribution and the two main groups of species of Bujurquina Kullander, 1986. Shown with red dotted lines are main geographic drainage basin divides (structural arches) of the western Amazon. Red arrows show the two main subducting ridges responsible for the origin of the formation of the structural arches of the western Amazon. All species were included in morphological analyses, species with an asterisk were also available for molecular analyses. Stars denote type localities for valid species." figureDoi="http://doi.org/10.5281/zenodo.8006322" httpUri="https://zenodo.org/record/8006322/files/figure.png" pageId="24" pageNumber="191">Figs 1</figureCitation>
and 
<figureCitation id="0AA34C906F4FE469FEA1FD549627FD34" box="[323,355,699,725]" captionStart="Fig" captionStartId="24.[189,232,1892,1918]" captionTargetBox="[190,1398,1076,1839]" captionTargetId="figure-305@24.[189,1398,1072,1852]" captionTargetPageId="24" captionText="Fig. 17. Phylogenetic relationships and species delimitation within Bujurquina Kullander, 1986 based on the cytb marker from BEAST analysis. Numbers at nodes show age in Ma, grey bars at nodes show confidence intervals of age estimates (95% HPD). Bayesian posterior probabilities above 0.95 are shown as black points on nodes. Scale bar shows 5 My. Note the two main groups of Bujurquina." figureDoi="http://doi.org/10.5281/zenodo.8006356" httpUri="https://zenodo.org/record/8006356/files/figure.png" pageId="24" pageNumber="191">17</figureCitation>
). The main biogeographic dichotomy within the genus between the Southern and Northern groups is within the present upper 
<collectingRegion id="505C9EF76F4FE469FDB1FD3195EAFD19" box="[595,686,734,760]" country="Peru" name="Ucayali" pageId="24" pageNumber="191">Ucayali</collectingRegion>
basin in 
<collectingCountry id="EA8F10856F4FE469FCFBFD30940BFD18" box="[793,847,735,761]" name="Peru" pageId="24" pageNumber="191">Peru</collectingCountry>
(
<figureCitation id="0AA34C906F4FE469FCBCFD3194E7FD19" box="[862,931,734,760]" captionStart="Fig" captionStartId="4.[189,232,1857,1883]" captionTargetBox="[189,1398,907,1814]" captionTargetId="figure-237@4.[189,1398,907,1814]" captionTargetPageId="4" captionText="Fig. 1. Distribution and the two main groups of species of Bujurquina Kullander, 1986. Shown with red dotted lines are main geographic drainage basin divides (structural arches) of the western Amazon. Red arrows show the two main subducting ridges responsible for the origin of the formation of the structural arches of the western Amazon. All species were included in morphological analyses, species with an asterisk were also available for molecular analyses. Stars denote type localities for valid species." figureDoi="http://doi.org/10.5281/zenodo.8006322" httpUri="https://zenodo.org/record/8006322/files/figure.png" pageId="24" pageNumber="191">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="922750156F4FE468FF5FFCF0935AFEA7" blockId="24.[189,1398,799,999]" lastBlockId="25.[189,1398,265,326]" lastPageId="25" lastPageNumber="192" pageId="24" pageNumber="191">
The two main clades of 
<taxonomicName id="55982B966F4FE469FE37FCCF951EFCD8" authorityName="Kullander" authorityYear="1986" box="[469,602,800,825]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="24" pageNumber="191" phylum="Chordata" rank="genus">
<emphasis id="A0EC8C076F4FE469FE37FCCF951EFCD8" box="[469,602,800,825]" italics="true" pageId="24" pageNumber="191">Bujurquina</emphasis>
</taxonomicName>
are dated as having diverged at 16.6 Ma. Within the Southern group the two southern-most species from 
<collectingCountry id="EA8F10856F4FE469FD83FCAD959DFCBD" box="[609,729,834,860]" name="Argentina" pageId="24" pageNumber="191">Argentina</collectingCountry>
, 
<collectingCountry id="EA8F10856F4FE469FD06FCAD9416FCBD" box="[740,850,834,860]" name="Paraguay" pageId="24" pageNumber="191">Paraguay</collectingCountry>
, 
<collectingCountry id="EA8F10856F4FE469FCBFFCAD94E1FCBD" box="[861,933,834,860]" name="Brazil" pageId="24" pageNumber="191">Brazil</collectingCountry>
and 
<collectingCountry id="EA8F10856F4FE469FC39FCAD9375FCBD" box="[987,1073,834,860]" name="Bolivia" pageId="24" pageNumber="191">Bolivia</collectingCountry>
are sister species (
<taxonomicName id="55982B966F4FE469FAEEFCAD9232FCBA" baseAuthorityName="Heckel" baseAuthorityYear="1840" box="[1292,1398,834,859]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="24" pageNumber="191" phylum="Chordata" rank="species" species="vittata">
<emphasis id="A0EC8C076F4FE469FAEEFCAD9232FCBA" box="[1292,1398,834,859]" italics="true" pageId="24" pageNumber="191">B. vittata</emphasis>
</taxonomicName>
and 
<taxonomicName id="55982B966F4FE469FF0DFC8A96D8FC9F" authorityName="Kullander" authorityYear="1987" box="[239,412,868,894]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="24" pageNumber="191" phylum="Chordata" rank="species" species="oenolaemus">
<emphasis id="A0EC8C076F4FE469FF0DFC8A96D8FC9F" box="[239,412,868,894]" italics="true" pageId="24" pageNumber="191">B. oenolaemus</emphasis>
</taxonomicName>
), and they are a sister group to the clade composed of the Peruvian 
<taxonomicName id="55982B966F4FE469FB5FFC8A9200FC9F" authorityName="Kullander" authorityYear="1986" box="[1213,1348,868,894]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="24" pageNumber="191" phylum="Chordata" rank="species" species="eurhinus">
<emphasis id="A0EC8C076F4FE469FB5FFC8A9200FC9F" box="[1213,1348,868,894]" italics="true" pageId="24" pageNumber="191">B. eurhinus</emphasis>
</taxonomicName>
and 
<taxonomicName id="55982B966F4FE469FF5FFC679623FC40" authorityName="Kullander" authorityYear="1986" box="[189,359,903,929]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="24" pageNumber="191" phylum="Chordata" rank="species" species="tambopatae">
<emphasis id="A0EC8C076F4FE469FF5FFC679623FC40" box="[189,359,903,929]" italics="true" pageId="24" pageNumber="191">B. tambopatae</emphasis>
</taxonomicName>
as sister species, followed by the Bolivian 
<emphasis id="A0EC8C076F4FE469FCB7FC679423FC40" box="[853,871,904,929]" italics="true" pageId="24" pageNumber="191">B</emphasis>
. sp. 
<collectingCountry id="EA8F10856F4FE469FC79FC6894B5FC40" box="[923,1009,903,929]" name="Bolivia" pageId="24" pageNumber="191">Bolivia</collectingCountry>
and then the Peruvian 
<taxonomicName id="55982B966F4FE469FB1AFC679234FC40" authorityName="Kullander" authorityYear="1986" box="[1272,1392,903,929]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="24" pageNumber="191" phylum="Chordata" rank="species" species="robusta">
<emphasis id="A0EC8C076F4FE469FB1AFC679234FC40" box="[1272,1392,903,929]" italics="true" pageId="24" pageNumber="191">B. robusta</emphasis>
</taxonomicName>
. The phylogenetically basal-most species of the Southern group is the Peruvian 
<taxonomicName id="55982B966F4FE469FBB4FC459388FC25" authorityName="Kullander" authorityYear="1986" box="[1110,1228,938,964]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="24" pageNumber="191" phylum="Chordata" rank="species" species="labiosa">
<emphasis id="A0EC8C076F4FE469FBB4FC459388FC25" box="[1110,1228,938,964]" italics="true" pageId="24" pageNumber="191">B. labiosa</emphasis>
</taxonomicName>
. The Peruvian species are thus in basal positions within the Southern group, and they also have the longest internodes and the earliest dates of divergence based on molecular clock dating, with the time frame of divergence among the Peruvian species dated between 12.4 Ma and 3.0 Ma (
<figureCitation id="0AA34C906F4EE468FC5BFEC39354FEA7" box="[953,1040,300,326]" captionStart="Fig" captionStartId="24.[189,232,1892,1918]" captionTargetBox="[190,1398,1076,1839]" captionTargetId="figure-305@24.[189,1398,1072,1852]" captionTargetPageId="24" captionText="Fig. 17. Phylogenetic relationships and species delimitation within Bujurquina Kullander, 1986 based on the cytb marker from BEAST analysis. Numbers at nodes show age in Ma, grey bars at nodes show confidence intervals of age estimates (95% HPD). Bayesian posterior probabilities above 0.95 are shown as black points on nodes. Scale bar shows 5 My. Note the two main groups of Bujurquina." figureDoi="http://doi.org/10.5281/zenodo.8006356" httpUri="https://zenodo.org/record/8006356/files/figure.png" pageId="25" pageNumber="192">Fig. 17</figureCitation>
).
</paragraph>
<caption id="C6E7009D6F4FE469FF5FF88B939AF808" ID-DOI="http://doi.org/10.5281/zenodo.8006356" ID-Zenodo-Dep="8006356" httpUri="https://zenodo.org/record/8006356/files/figure.png" pageId="24" pageNumber="191" startId="24.[189,232,1892,1918]" targetBox="[190,1398,1076,1839]" targetPageId="24">
<paragraph id="922750156F4FE469FF5FF88B939AF808" blockId="24.[189,1399,1892,2025]" pageId="24" pageNumber="191">
<emphasis id="A0EC8C076F4FE469FF5FF88B9658F89E" bold="true" box="[189,284,1892,1919]" pageId="24" pageNumber="191">Fig. 17.</emphasis>
Phylogenetic relationships and species delimitation within 
<taxonomicName id="55982B966F4FE469FC3DF889926FF89E" authority="Kullander, 1986" authorityName="Kullander" authorityYear="1986" box="[991,1323,1893,1919]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="24" pageNumber="191" phylum="Chordata" rank="genus">
<emphasis id="A0EC8C076F4FE469FC3DF8899320F89E" box="[991,1124,1894,1919]" italics="true" pageId="24" pageNumber="191">Bujurquina</emphasis>
<bibRefCitation id="F6092DE46F4FE469FB8EF88A926FF89E" author="Kullander S." box="[1132,1323,1893,1919]" pageId="24" pageNumber="191" refId="ref14485" refString="Kullander S. 1986. Cichlid Fishes of the Amazon River Drainage of Peru. Swedish Museum of Natural History, Sweden." type="book" year="1986">Kullander, 1986</bibRefCitation>
</taxonomicName>
based on the cytb marker from BEAST analysis. Numbers at nodes show age in Ma, grey bars at nodes show confidence intervals of age estimates (95% HPD). Bayesian posterior probabilities above 0.95 are shown as black points on nodes. Scale bar shows 5 My. Note the two main groups of 
<taxonomicName id="55982B966F4FE469FBB0F8209393F809" authorityName="Kullander" authorityYear="1986" box="[1106,1239,1999,2024]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="24" pageNumber="191" phylum="Chordata" rank="genus">
<emphasis id="A0EC8C076F4FE469FBB0F8209393F809" box="[1106,1239,1999,2024]" italics="true" pageId="24" pageNumber="191">Bujurquina</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="922750156F4EE468FF5FFE83955CFC6F" blockId="25.[189,1399,364,911]" pageId="25" pageNumber="192">
The Northern group of 
<taxonomicName id="55982B966F4EE468FE26FE82950DFE67" authorityName="Kullander" authorityYear="1986" box="[452,585,365,390]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="25" pageNumber="192" phylum="Chordata" rank="genus">
<emphasis id="A0EC8C076F4EE468FE26FE82950DFE67" box="[452,585,365,390]" italics="true" pageId="25" pageNumber="192">Bujurquina</emphasis>
</taxonomicName>
is made up of two highly divergent clades (
<figureCitation id="0AA34C906F4EE468FBDBFE8393C8FE66" box="[1081,1164,364,391]" captionStart="Fig" captionStartId="24.[189,232,1892,1918]" captionTargetBox="[190,1398,1076,1839]" captionTargetId="figure-305@24.[189,1398,1072,1852]" captionTargetPageId="24" captionText="Fig. 17. Phylogenetic relationships and species delimitation within Bujurquina Kullander, 1986 based on the cytb marker from BEAST analysis. Numbers at nodes show age in Ma, grey bars at nodes show confidence intervals of age estimates (95% HPD). Bayesian posterior probabilities above 0.95 are shown as black points on nodes. Scale bar shows 5 My. Note the two main groups of Bujurquina." figureDoi="http://doi.org/10.5281/zenodo.8006356" httpUri="https://zenodo.org/record/8006356/files/figure.png" pageId="25" pageNumber="192">Fig. 17</figureCitation>
) unlike the situation in the Southern group. The first clade within the Northern group shows a younger basal divergence at 3.9 Ma and includes, as a basal species, the here described 
<taxonomicName id="55982B966F4EE468FC68FE5C9348FE2D" authority="Říčan &amp; Říčanová, 2023" authorityName="Říčan &amp; Říčanová" authorityYear="2023" box="[906,1036,435,460]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="25" pageNumber="192" phylum="Chordata" rank="species" species="omagua" status="sp. nov.">
<emphasis id="A0EC8C076F4EE468FC68FE5C9348FE2D" box="[906,1036,435,460]" italics="true" pageId="25" pageNumber="192">B. omagua</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBDF317C6F4EE468FBF5FE5D9330FE2D" box="[1047,1140,434,460]" pageId="25" pageNumber="192" rank="species">sp. nov.</taxonomicNameLabel>
, and then, among the Peruvian species, 
<taxonomicName id="55982B966F4EE468FE73FE3A9563FE0F" authorityName="Kullander" authorityYear="1986" box="[401,551,468,494]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="25" pageNumber="192" phylum="Chordata" rank="species" species="huallagae">
<emphasis id="A0EC8C076F4EE468FE73FE3A9563FE0F" box="[401,551,468,494]" italics="true" pageId="25" pageNumber="192">B. huallagae</emphasis>
</taxonomicName>
and 
<taxonomicName id="55982B966F4EE468FD80FE3A95A4FE0F" baseAuthorityName="Cope" baseAuthorityYear="1872" box="[610,736,468,494]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="25" pageNumber="192" phylum="Chordata" rank="species" species="syspilus">
<emphasis id="A0EC8C076F4EE468FD80FE3A95A4FE0F" box="[610,736,468,494]" italics="true" pageId="25" pageNumber="192">B. syspilus</emphasis>
</taxonomicName>
. Samples of the morphologically and biogeographically Southern group species 
<taxonomicName id="55982B966F4EE468FE34FE1795D5FDF0" authorityName="Kullander" authorityYear="1986" box="[470,657,503,529]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="25" pageNumber="192" phylum="Chordata" rank="species" species="megalospilus">
<emphasis id="A0EC8C076F4EE468FE34FE1795D5FDF0" box="[470,657,503,529]" italics="true" pageId="25" pageNumber="192">B. megalospilus</emphasis>
</taxonomicName>
are found within 
<taxonomicName id="55982B966F4EE468FC83FE17949AFDF0" baseAuthorityName="Cope" baseAuthorityYear="1872" box="[865,990,503,529]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="25" pageNumber="192" phylum="Chordata" rank="species" species="syspilus">
<emphasis id="A0EC8C076F4EE468FC83FE17949AFDF0" box="[865,990,503,529]" italics="true" pageId="25" pageNumber="192">B. syspilus</emphasis>
</taxonomicName>
together with some samples of the Southern group species 
<taxonomicName id="55982B966F4EE468FE37FDF49508FDD5" authorityName="Kullander" authorityYear="1986" box="[469,588,538,564]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="25" pageNumber="192" phylum="Chordata" rank="species" species="labiosa">
<emphasis id="A0EC8C076F4EE468FE37FDF49508FDD5" box="[469,588,538,564]" italics="true" pageId="25" pageNumber="192">B. labiosa</emphasis>
</taxonomicName>
and 
<taxonomicName id="55982B966F4EE468FD67FDF495BAFDD5" authorityName="Kullander" authorityYear="1986" box="[645,766,538,564]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="25" pageNumber="192" phylum="Chordata" rank="species" species="robusta">
<emphasis id="A0EC8C076F4EE468FD67FDF495BAFDD5" box="[645,766,538,564]" italics="true" pageId="25" pageNumber="192">B. robusta</emphasis>
</taxonomicName>
(Supp. file 2: 
<figureCitation id="0AA34C906F4EE468FC46FDF594B8FDD5" box="[932,1020,538,564]" captionStart="Fig" captionStartId="6.[189,232,1892,1918]" captionTargetBox="[189,1397,768,1849]" captionTargetId="figure-208@6.[189,1398,766,1851]" captionTargetPageId="6" captionText="Fig. 3. Detailed photographs of the spinous portion of the dorsal fin of species of Bujurquina Kullander, 1986 in the Southern group (see Figs 1–2). Note lack of patterning as opposed to various types of ornamentation in species of the Northern group. Note differences in the coloration of the dorsal fin lappets between some of the species, e.g., red-orange, vs white vs black." figureDoi="http://doi.org/10.5281/zenodo.8006328" httpUri="https://zenodo.org/record/8006328/files/figure.png" pageId="25" pageNumber="192">Fig. S3</figureCitation>
). These three species clearly are Southern group species whose mtDNA in these particular samples was swept by mtDNA from 
<taxonomicName id="55982B966F4EE468FB19FDD29233FDB7" baseAuthorityName="Cope" baseAuthorityYear="1872" box="[1275,1399,572,598]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="25" pageNumber="192" phylum="Chordata" rank="species" species="syspilus">
<emphasis id="A0EC8C076F4EE468FB19FDD29233FDB7" box="[1275,1399,572,598]" italics="true" pageId="25" pageNumber="192">B. syspilus</emphasis>
</taxonomicName>
following a single postulated hybridization event from 
<taxonomicName id="55982B966F4EE468FCB3FD8F9494FD98" baseAuthorityName="Cope" baseAuthorityYear="1872" box="[849,976,607,633]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="25" pageNumber="192" phylum="Chordata" rank="species" species="syspilus">
<emphasis id="A0EC8C076F4EE468FCB3FD8F9494FD98" box="[849,976,607,633]" italics="true" pageId="25" pageNumber="192">B. syspilus</emphasis>
</taxonomicName>
into the three introgressed species, dated in our phylogeny at 0.2 Ma (cf. 
<figureCitation id="0AA34C906F4EE468FD64FD6D959FFD7D" box="[646,731,642,668]" captionStart="Fig" captionStartId="24.[189,232,1892,1918]" captionTargetBox="[190,1398,1076,1839]" captionTargetId="figure-305@24.[189,1398,1072,1852]" captionTargetPageId="24" captionText="Fig. 17. Phylogenetic relationships and species delimitation within Bujurquina Kullander, 1986 based on the cytb marker from BEAST analysis. Numbers at nodes show age in Ma, grey bars at nodes show confidence intervals of age estimates (95% HPD). Bayesian posterior probabilities above 0.95 are shown as black points on nodes. Scale bar shows 5 My. Note the two main groups of Bujurquina." figureDoi="http://doi.org/10.5281/zenodo.8006356" httpUri="https://zenodo.org/record/8006356/files/figure.png" pageId="25" pageNumber="192">Fig. 17</figureCitation>
and Supp. file 2: 
<figureCitation id="0AA34C906F4EE468FC50FD6D934FFD7D" box="[946,1035,642,668]" captionStart="Fig" captionStartId="6.[189,232,1892,1918]" captionTargetBox="[189,1397,768,1849]" captionTargetId="figure-208@6.[189,1398,766,1851]" captionTargetPageId="6" captionText="Fig. 3. Detailed photographs of the spinous portion of the dorsal fin of species of Bujurquina Kullander, 1986 in the Southern group (see Figs 1–2). Note lack of patterning as opposed to various types of ornamentation in species of the Northern group. Note differences in the coloration of the dorsal fin lappets between some of the species, e.g., red-orange, vs white vs black." figureDoi="http://doi.org/10.5281/zenodo.8006328" httpUri="https://zenodo.org/record/8006328/files/figure.png" pageId="25" pageNumber="192">Fig. S3</figureCitation>
). Non-introgressed samples of 
<taxonomicName id="55982B966F4EE468FF5FFD4A9673FD5F" authorityName="Kullander" authorityYear="1986" box="[189,311,676,702]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="25" pageNumber="192" phylum="Chordata" rank="species" species="labiosa">
<emphasis id="A0EC8C076F4EE468FF5FFD4A9673FD5F" box="[189,311,676,702]" italics="true" pageId="25" pageNumber="192">B. labiosa</emphasis>
</taxonomicName>
and 
<taxonomicName id="55982B966F4EE468FE94FD4A96B7FD5F" authorityName="Kullander" authorityYear="1986" box="[374,499,676,702]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="25" pageNumber="192" phylum="Chordata" rank="species" species="robusta">
<emphasis id="A0EC8C076F4EE468FE94FD4A96B7FD5F" box="[374,499,676,702]" italics="true" pageId="25" pageNumber="192">B. robusta</emphasis>
</taxonomicName>
are found as separate Southern group species in all phylogenetic analyses (see 
<figureCitation id="0AA34C906F4EE468FF11FD28960CFD00" box="[243,328,711,737]" captionStart="Fig" captionStartId="24.[189,232,1892,1918]" captionTargetBox="[190,1398,1076,1839]" captionTargetId="figure-305@24.[189,1398,1072,1852]" captionTargetPageId="24" captionText="Fig. 17. Phylogenetic relationships and species delimitation within Bujurquina Kullander, 1986 based on the cytb marker from BEAST analysis. Numbers at nodes show age in Ma, grey bars at nodes show confidence intervals of age estimates (95% HPD). Bayesian posterior probabilities above 0.95 are shown as black points on nodes. Scale bar shows 5 My. Note the two main groups of Bujurquina." figureDoi="http://doi.org/10.5281/zenodo.8006356" httpUri="https://zenodo.org/record/8006356/files/figure.png" pageId="25" pageNumber="192">Fig. 17</figureCitation>
). Non-introgressed samples of 
<taxonomicName id="55982B966F4EE468FD57FD279434FD00" authorityName="Kullander" authorityYear="1986" box="[693,880,711,737]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="25" pageNumber="192" phylum="Chordata" rank="species" species="megalospilus">
<emphasis id="A0EC8C076F4EE468FD57FD279434FD00" box="[693,880,711,737]" italics="true" pageId="25" pageNumber="192">B. megalospilus</emphasis>
</taxonomicName>
have not been found in this study. The close phylogenetic position in mtDNA of the morphologically dissimilar yet clearly Northern group species 
<taxonomicName id="55982B966F4EE468FF5FFCE2967EFCC7" baseAuthorityName="Cope" baseAuthorityYear="1872" box="[189,314,780,806]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="25" pageNumber="192" phylum="Chordata" rank="species" species="syspilus">
<emphasis id="A0EC8C076F4EE468FF5FFCE2967EFCC7" box="[189,314,780,806]" italics="true" pageId="25" pageNumber="192">B. syspilus</emphasis>
</taxonomicName>
and 
<taxonomicName id="55982B966F4EE468FE90FCE29543FCC7" authorityName="Kullander" authorityYear="1986" box="[370,519,780,806]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="25" pageNumber="192" phylum="Chordata" rank="species" species="huallagae">
<emphasis id="A0EC8C076F4EE468FE90FCE29543FCC7" box="[370,519,780,806]" italics="true" pageId="25" pageNumber="192">B. huallagae</emphasis>
</taxonomicName>
may potentially also be a case of 
<taxonomicName id="55982B966F4EE468FC73FCE2934BFCC7" baseAuthorityName="Cope" baseAuthorityYear="1872" box="[913,1039,780,806]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="25" pageNumber="192" phylum="Chordata" rank="species" species="syspilus">
<emphasis id="A0EC8C076F4EE468FC73FCE2934BFCC7" box="[913,1039,780,806]" italics="true" pageId="25" pageNumber="192">B. syspilus</emphasis>
</taxonomicName>
introgression into the sampled specimen of 
<taxonomicName id="55982B966F4EE468FEBEFCDF96B2FCA8" authorityName="Kullander" authorityYear="1986" box="[348,502,815,841]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="25" pageNumber="192" phylum="Chordata" rank="species" species="huallagae">
<emphasis id="A0EC8C076F4EE468FEBEFCDF96B2FCA8" box="[348,502,815,841]" italics="true" pageId="25" pageNumber="192">B. huallagae</emphasis>
</taxonomicName>
. This specimen was sampled in the lowermost portion of its distribution closest to the natural distribution of 
<taxonomicName id="55982B966F4EE468FD96FCBC95B0FC8D" baseAuthorityName="Cope" baseAuthorityYear="1872" box="[628,756,850,876]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="25" pageNumber="192" phylum="Chordata" rank="species" species="syspilus">
<emphasis id="A0EC8C076F4EE468FD96FCBC95B0FC8D" box="[628,756,850,876]" italics="true" pageId="25" pageNumber="192">B. syspilus</emphasis>
</taxonomicName>
and not from the 
<typeStatus id="4D23EEB76F4EE468FC30FCBD9341FC8D" box="[978,1029,850,876]" pageId="25" pageNumber="192">type</typeStatus>
locality of 
<taxonomicName id="55982B966F4EE468FB71FCBC926FFC8D" authorityName="Kullander" authorityYear="1986" box="[1171,1323,850,876]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="25" pageNumber="192" phylum="Chordata" rank="species" species="huallagae">
<emphasis id="A0EC8C076F4EE468FB71FCBC926FFC8D" box="[1171,1323,850,876]" italics="true" pageId="25" pageNumber="192">B. huallagae</emphasis>
</taxonomicName>
much higher up the namesake river.
</paragraph>
<paragraph id="922750156F4EE468FF5FFC5996C4FBD6" blockId="25.[189,1399,949,1079]" pageId="25" pageNumber="192">In our present taxon sampling the first clade within the Northern group thus has a 10 my time gap from the basal node at 13.9 Ma to the presently sampled divergence at 3.9 Ma and this may suggest the existence of yet undiscovered older diverged species in this clade, or alternatively, it suggests extinction within the clade.</paragraph>
<paragraph id="922750156F4EE468FF5FFBB1957FFA51" blockId="25.[189,1399,1118,1456]" pageId="25" pageNumber="192">
The second clade within the Northern group shows a younger basal divergence at 7.9 Ma, hence also with a time gap and with similar potential implications, which is strenghtened by the isolated position of the basal-most species of this clade, 
<taxonomicName id="55982B966F4EE468FD6FFB4B9445FB5C" baseAuthorityName="Eigenmann" baseAuthorityYear="1923" box="[653,769,1188,1213]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="25" pageNumber="192" phylum="Chordata" rank="species" species="mariae">
<emphasis id="A0EC8C076F4EE468FD6FFB4B9445FB5C" box="[653,769,1188,1213]" italics="true" pageId="25" pageNumber="192">B. mariae</emphasis>
</taxonomicName>
(
<figureCitation id="0AA34C906F4EE468FCF0FB4B942DFB5F" box="[786,873,1188,1214]" captionStart="Fig" captionStartId="24.[189,232,1892,1918]" captionTargetBox="[190,1398,1076,1839]" captionTargetId="figure-305@24.[189,1398,1072,1852]" captionTargetPageId="24" captionText="Fig. 17. Phylogenetic relationships and species delimitation within Bujurquina Kullander, 1986 based on the cytb marker from BEAST analysis. Numbers at nodes show age in Ma, grey bars at nodes show confidence intervals of age estimates (95% HPD). Bayesian posterior probabilities above 0.95 are shown as black points on nodes. Scale bar shows 5 My. Note the two main groups of Bujurquina." figureDoi="http://doi.org/10.5281/zenodo.8006356" httpUri="https://zenodo.org/record/8006356/files/figure.png" pageId="25" pageNumber="192">Fig. 17</figureCitation>
). 
<taxonomicName id="55982B966F4EE468FC9DFB4B931BFB5C" baseAuthorityName="Eigenmann" baseAuthorityYear="1923" box="[895,1119,1188,1213]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="25" pageNumber="192" phylum="Chordata" rank="species" species="mariae">
<emphasis id="A0EC8C076F4EE468FC9DFB4B931BFB5C" box="[895,1119,1188,1213]" italics="true" pageId="25" pageNumber="192">Bujurquina mariae</emphasis>
</taxonomicName>
(from 
<collectingCountry id="EA8F10856F4EE468FB50FB4B926CFB5F" box="[1202,1320,1188,1214]" name="Colombia" pageId="25" pageNumber="192">Colombia</collectingCountry>
) is the only species so far sampled from the Orinoco basin to the north of the Amazon basin. The two remaining Peruvian species (
<taxonomicName id="55982B966F4EE468FE74FB059530FAE2" authorityName="Kullander" authorityYear="1986" box="[406,628,1257,1283]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="25" pageNumber="192" phylum="Chordata" rank="species" species="peregrinabunda">
<emphasis id="A0EC8C076F4EE468FE74FB059530FAE2" box="[406,628,1257,1283]" italics="true" pageId="25" pageNumber="192">B. peregrinabunda</emphasis>
</taxonomicName>
and 
<taxonomicName id="55982B966F4EE468FD53FB059403FAE2" authorityName="Kullander" authorityYear="1986" box="[689,839,1258,1283]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="25" pageNumber="192" phylum="Chordata" rank="species" species="moriorum">
<emphasis id="A0EC8C076F4EE468FD53FB059403FAE2" box="[689,839,1258,1283]" italics="true" pageId="25" pageNumber="192">B. moriorum</emphasis>
</taxonomicName>
) diverged at 4.6 Ma, but they are not found as closely related in the mtDNA phylogeny despite being more similar and lowland parapatric species. Instead, the lowland 
<taxonomicName id="55982B966F4EE468FE23FAC095E7FAA9" authorityName="Kullander" authorityYear="1986" box="[449,675,1326,1352]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="25" pageNumber="192" phylum="Chordata" rank="species" species="peregrinabunda">
<emphasis id="A0EC8C076F4EE468FE23FAC095E7FAA9" box="[449,675,1326,1352]" italics="true" pageId="25" pageNumber="192">B. peregrinabunda</emphasis>
</taxonomicName>
is found closely related to the allopatric Andean endemic 
<taxonomicName id="55982B966F4EE468FF5FFABD9627FA8A" authorityName="" baseAuthorityName="Regan" baseAuthorityYear="1905" box="[189,355,1362,1387]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="25" pageNumber="192" phylum="Chordata" rank="species" species="zamorensis">
<emphasis id="A0EC8C076F4EE468FF5FFABD9627FA8A" box="[189,355,1362,1387]" italics="true" pageId="25" pageNumber="192">B. zamorensis</emphasis>
</taxonomicName>
from 
<collectingCountry id="EA8F10856F4EE468FE4CFABE9554FA8A" box="[430,528,1361,1387]" name="Ecuador" pageId="25" pageNumber="192">Ecuador</collectingCountry>
plus one putative lowland Ecuadorian species, most likely also found in 
<collectingCountry id="EA8F10856F4EE468FF5FFA9B97B7FA6F" box="[189,243,1396,1422]" name="Peru" pageId="25" pageNumber="192">Peru</collectingCountry>
(
<emphasis id="A0EC8C076F4EE468FEE4FA9B965CFA6C" box="[262,280,1396,1421]" italics="true" pageId="25" pageNumber="192">B</emphasis>
. sp. 
<collectingCountry id="EA8F10856F4EE468FEB6FA9B96F2FA6F" box="[340,438,1396,1422]" name="Ecuador" pageId="25" pageNumber="192">Ecuador</collectingCountry>
13). 
<taxonomicName id="55982B966F4EE468FE15FA9B95BEFA6C" authorityName="Kullander" authorityYear="1986" box="[503,762,1396,1421]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="25" pageNumber="192" phylum="Chordata" rank="species" species="moriorum">
<emphasis id="A0EC8C076F4EE468FE15FA9B95BEFA6C" box="[503,762,1396,1421]" italics="true" pageId="25" pageNumber="192">Bujurquina moriorum</emphasis>
</taxonomicName>
is found in a clade with two other putative Peruvian species from the Marañón basin.
</paragraph>
<paragraph id="922750156F4EE468FF5FFA36961CF9FD" blockId="25.[189,1316,1497,1564]" pageId="25" pageNumber="192">
<emphasis id="A0EC8C076F4EE468FF5FFA36961CF9FD" bold="true" italics="true" pageId="25" pageNumber="192">
Phylogeny and coloration patterns: the two main groups and phylogenetic lineages of 
<taxonomicName id="55982B966F4EE468FF5FFA10961CF9FD" authorityName="Kullander" authorityYear="1986" box="[189,344,1535,1564]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="25" pageNumber="192" phylum="Chordata" rank="genus">Bujurquina</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="922750156F4EE468FF5FF9DB93DCF8A5" blockId="25.[189,1399,1588,1861]" pageId="25" pageNumber="192">
Our phylogenetic results have shown that 
<taxonomicName id="55982B966F4EE468FD4BF9DB946AF9AC" authorityName="Kullander" authorityYear="1986" box="[681,814,1588,1613]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="25" pageNumber="192" phylum="Chordata" rank="genus">
<emphasis id="A0EC8C076F4EE468FD4BF9DB946AF9AC" box="[681,814,1588,1613]" italics="true" pageId="25" pageNumber="192">Bujurquina</emphasis>
</taxonomicName>
is divided into two main clades. One clade, based on distribution patterns referred to us as the Northern group, is characterized by ornamented dorsal fins and a midlateral stripe running generally to the dorsal margin of the caudal peduncle (
<figureCitation id="0AA34C906F4EE468FB0DF995927FF975" box="[1263,1339,1658,1684]" captionStart="Fig" captionStartId="4.[189,232,1857,1883]" captionTargetBox="[189,1398,907,1814]" captionTargetId="figure-237@4.[189,1398,907,1814]" captionTargetPageId="4" captionText="Fig. 1. Distribution and the two main groups of species of Bujurquina Kullander, 1986. Shown with red dotted lines are main geographic drainage basin divides (structural arches) of the western Amazon. Red arrows show the two main subducting ridges responsible for the origin of the formation of the structural arches of the western Amazon. All species were included in morphological analyses, species with an asterisk were also available for molecular analyses. Stars denote type localities for valid species." figureDoi="http://doi.org/10.5281/zenodo.8006322" httpUri="https://zenodo.org/record/8006322/files/figure.png" pageId="25" pageNumber="192">Figs 1</figureCitation>
, 
<figureCitation id="0AA34C906F4EE468FAA8F9959232F974" box="[1354,1398,1658,1685]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="7.[189,232,1963,1989]" captionStartId-1="8.[189,232,1822,1848]" captionTargetBox-0="[190,1398,758,1923]" captionTargetBox-1="[189,1398,891,1781]" captionTargetId-0="figure-146@7.[189,1398,755,1923]" captionTargetId-1="figure-253@8.[189,1398,888,1781]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="Fig. 4. Representative live specimens of the valid species of Bujurquina Kullander, 1986 in the Northern group including the newly described species B. omagua sp. nov." captionText-1="Fig. 5. Detailed photographs of the spinous portion of the dorsal fin of species of Bujurquina Kullander, 1986 in the Northern group (see Figs 1 and 4). Note two main types of patterning of the dorsal fin, a blotched patterning vs lines in the spinous portion of the dorsal fin. Note additional differences in patterning of the spinous portion of the dorsal fin between the species, such as different numbers of lines per membrane, their thickness or angle. Also note differences in the coloration of the dorsal fin lappets between some of the species." figureDoi-0="http://doi.org/10.5281/zenodo.8006330" figureDoi-1="http://doi.org/10.5281/zenodo.8006332" httpUri-0="https://zenodo.org/record/8006330/files/figure.png" httpUri-1="https://zenodo.org/record/8006332/files/figure.png" pageId="25" pageNumber="192">4–5</figureCitation>
and 
<figureCitation id="0AA34C906F4EE468FF13F9729655F956" box="[241,273,1693,1719]" captionStart="Fig" captionStartId="24.[189,232,1892,1918]" captionTargetBox="[190,1398,1076,1839]" captionTargetId="figure-305@24.[189,1398,1072,1852]" captionTargetPageId="24" captionText="Fig. 17. Phylogenetic relationships and species delimitation within Bujurquina Kullander, 1986 based on the cytb marker from BEAST analysis. Numbers at nodes show age in Ma, grey bars at nodes show confidence intervals of age estimates (95% HPD). Bayesian posterior probabilities above 0.95 are shown as black points on nodes. Scale bar shows 5 My. Note the two main groups of Bujurquina." figureDoi="http://doi.org/10.5281/zenodo.8006356" httpUri="https://zenodo.org/record/8006356/files/figure.png" pageId="25" pageNumber="192">17</figureCitation>
). In contrast to this the Southern group is characterized by unpatterned (hyaline, i.e., without any markings) dorsal fins and a midlateral stripe that always runs towards the posterior insertion of the dorsal fin (
<figureCitation id="0AA34C906F4EE468FEDAF90B96DBF91F" box="[312,415,1764,1790]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="4.[189,232,1857,1883]" captionStartId-1="5.[189,232,1873,1899]" captionStartId-2="6.[189,232,1892,1918]" captionTargetBox-0="[189,1398,907,1814]" captionTargetBox-1="[189,1398,267,1822]" captionTargetBox-2="[189,1397,768,1849]" captionTargetId-0="figure-237@4.[189,1398,907,1814]" captionTargetId-1="figure-11@5.[189,1398,267,1823]" captionTargetId-2="figure-208@6.[189,1398,766,1851]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionText-0="Fig. 1. Distribution and the two main groups of species of Bujurquina Kullander, 1986. Shown with red dotted lines are main geographic drainage basin divides (structural arches) of the western Amazon. Red arrows show the two main subducting ridges responsible for the origin of the formation of the structural arches of the western Amazon. All species were included in morphological analyses, species with an asterisk were also available for molecular analyses. Stars denote type localities for valid species." captionText-1="Fig. 2. Representative live specimens of the valid species of Bujurquina Kullander, 1986 (plus B. sp. Bolivia) in the Southern group (no photo of a live specimen of B. cordemadi Kullander, 1986 is available and we are not aware that the species has ever been reliably identified and photographed alive)." captionText-2="Fig. 3. Detailed photographs of the spinous portion of the dorsal fin of species of Bujurquina Kullander, 1986 in the Southern group (see Figs 1–2). Note lack of patterning as opposed to various types of ornamentation in species of the Northern group. Note differences in the coloration of the dorsal fin lappets between some of the species, e.g., red-orange, vs white vs black." figureDoi-0="http://doi.org/10.5281/zenodo.8006322" figureDoi-1="http://doi.org/10.5281/zenodo.8006324" figureDoi-2="http://doi.org/10.5281/zenodo.8006328" httpUri-0="https://zenodo.org/record/8006322/files/figure.png" httpUri-1="https://zenodo.org/record/8006324/files/figure.png" httpUri-2="https://zenodo.org/record/8006328/files/figure.png" pageId="25" pageNumber="192">Figs 1–3</figureCitation>
and 
<figureCitation id="0AA34C906F4EE468FE34F90B96B2F91F" box="[470,502,1764,1790]" captionStart="Fig" captionStartId="24.[189,232,1892,1918]" captionTargetBox="[190,1398,1076,1839]" captionTargetId="figure-305@24.[189,1398,1072,1852]" captionTargetPageId="24" captionText="Fig. 17. Phylogenetic relationships and species delimitation within Bujurquina Kullander, 1986 based on the cytb marker from BEAST analysis. Numbers at nodes show age in Ma, grey bars at nodes show confidence intervals of age estimates (95% HPD). Bayesian posterior probabilities above 0.95 are shown as black points on nodes. Scale bar shows 5 My. Note the two main groups of Bujurquina." figureDoi="http://doi.org/10.5281/zenodo.8006356" httpUri="https://zenodo.org/record/8006356/files/figure.png" pageId="25" pageNumber="192">17</figureCitation>
). Lower lip coloration is another character that almost fully separates the two groups, since in majority of the Northern group species the lower lip is distinctly azure blue while in the Southern group species it is always the same color as the upper lip (i.e., never blue).
</paragraph>
</subSubSection>
<subSubSection id="DA82039E6F4EE46AFF5FF8829233FACB" lastPageId="27" lastPageNumber="194" pageId="25" pageNumber="192" type="key">
<paragraph id="922750156F4EE468FF5FF882952FF86B" blockId="25.[189,619,1900,1930]" box="[189,619,1900,1930]" pageId="25" pageNumber="192">
<emphasis id="A0EC8C076F4EE468FF5FF882952FF86B" bold="true" box="[189,619,1900,1930]" italics="true" pageId="25" pageNumber="192">
Key to the species of 
<taxonomicName id="55982B966F4EE468FE32F882952FF86B" ID-CoL="3DWL" authorityName="Kullander" authorityYear="1986" box="[464,619,1901,1930]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="25" pageNumber="192" phylum="Chordata" rank="genus">Bujurquina</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="922750156F4EE46BFF5FF84D9587FEA6" blockId="25.[189,1399,1953,2015]" lastBlockId="26.[189,1399,265,327]" lastPageId="26" lastPageNumber="193" pageId="25" pageNumber="192">
For the key we have chosen to use only readily visible external characters. All characters except where specifically noted are for large adult specimens (usually above 
<quantity id="5560FDF06F4EE468FC58F82B9354F83E" box="[954,1040,1988,2015]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="25" pageNumber="192" unit="mm" value="60.0">60 mm</quantity>
SL), especially the color and coloration pattern characters, where most are best visible in live specimens. In bold are the main distinguishing character states for each step.
</paragraph>
<key id="E60907246F4DE46BFF5FFE829233FA64" pageId="26" pageNumber="193">
<keyStep id="296C48B06F4DE46BFF5FFE829233FDB6" pageId="26" pageNumber="193">
<paragraph id="922750156F4DE46BFF5FFE829233FE0E" blockId="26.[189,1399,365,600]" pageId="26" pageNumber="193">
<keyLead id="2969F3206F4DE46BFF5FFE829233FE0E" pageId="26" pageNumber="193">
1. Spinous portion of the dorsal fin ornamented (i.e., with blotches, lines etc.); midlateral stripe on body oriented towards posterior insertion of dorsal fin except in one species (
<taxonomicName id="55982B966F4DE46BFBAAFE7E938CFE4B" ID-CoL="NPVS" baseAuthorityName="Cope" baseAuthorityYear="1872" box="[1096,1224,400,426]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="26" pageNumber="193" phylum="Chordata" rank="species" species="syspilus">
<emphasis id="A0EC8C076F4DE46BFBAAFE7E938CFE4B" box="[1096,1224,400,426]" italics="true" pageId="26" pageNumber="193">B. syspilus</emphasis>
</taxonomicName>
), i.e., oriented nearly horizontally; flanks with dark, squarish or wedge-shaped spots, in overlapping region between scale base and overlying scale margin except in two species (
<taxonomicName id="55982B966F4DE46BFC5AFE399368FE0E" ID-CoL="NPVH" baseAuthorityName="Eigenmann" baseAuthorityYear="1923" box="[952,1068,470,495]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="26" pageNumber="193" phylum="Chordata" rank="species" species="mariae">
<emphasis id="A0EC8C076F4DE46BFC5AFE399368FE0E" box="[952,1068,470,495]" italics="true" pageId="26" pageNumber="193">B. mariae</emphasis>
</taxonomicName>
and 
<taxonomicName id="55982B966F4DE46BFB87FE39939EFE0E" ID-CoL="NPVM" authorityName="Kullander" authorityYear="1986" box="[1125,1242,470,495]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="26" pageNumber="193" phylum="Chordata" rank="species" species="ortegai">
<emphasis id="A0EC8C076F4DE46BFB87FE39939EFE0E" box="[1125,1242,470,495]" italics="true" pageId="26" pageNumber="193">B. ortegai</emphasis>
</taxonomicName>
) ................ 2
</keyLead>
</paragraph>
<paragraph id="922750156F4DE46BFF5FFE179233FDB6" blockId="26.[189,1399,365,600]" pageId="26" pageNumber="193">
<keyLead id="2969F3206F4DE46BFF5FFE179233FDB6" pageId="26" pageNumber="193">
– Spinous portion of the dorsal fin 
<emphasis id="A0EC8C076F4DE46BFD8AFE179447FDF3" bold="true" box="[616,771,504,530]" pageId="26" pageNumber="193">unpatterned</emphasis>
(i.e., hyaline without any markings); midlateral stripe on body oriented 
<emphasis id="A0EC8C076F4DE46BFE20FDF49583FDD5" bold="true" box="[450,711,538,565]" pageId="26" pageNumber="193">towards soft portion</emphasis>
of dorsal fin, i.e., oriented more dorsally, flanks without spots .................................................................................................................................................. 9
</keyLead>
</paragraph>
</keyStep>
<keyStep id="296C48B06F4DE46BFF5FFD919233FD3C" pageId="26" pageNumber="193">
<paragraph id="922750156F4DE46BFF5FFD919233FD79" blockId="26.[189,1399,638,734]" box="[189,1399,638,664]" pageId="26" pageNumber="193">
<keyLead id="2969F3206F4DE46BFF5FFD919233FD79" box="[189,1399,638,664]" pageId="26" pageNumber="193">
2. Spinous portion of the dorsal fin with 
<emphasis id="A0EC8C076F4DE46BFD4AFD919410FD79" bold="true" box="[680,852,638,664]" pageId="26" pageNumber="193">diagonal lines</emphasis>
(i.e., dark lines on a hyaline background) ... 3
</keyLead>
</paragraph>
<paragraph id="922750156F4DE46BFF5FFD4E9233FD3C" blockId="26.[189,1399,638,734]" pageId="26" pageNumber="193">
<keyLead id="2969F3206F4DE46BFF5FFD4E9233FD3C" pageId="26" pageNumber="193">
– Spinous portion of the dorsal fin with 
<emphasis id="A0EC8C076F4DE46BFD01FD4F9488FD5B" bold="true" box="[739,972,672,698]" pageId="26" pageNumber="193">circular markings</emphasis>
(i.e., hyaline blotches on a dark background) ...................................................................................................................................... 6
</keyLead>
</paragraph>
</keyStep>
<keyStep id="296C48B06F4DE46BFF5FFCEB9233FC48" pageId="26" pageNumber="193">
<paragraph id="922750156F4DE46BFF5FFCEB9232FC67" blockId="26.[189,1399,772,937]" pageId="26" pageNumber="193">
<keyLead id="2969F3206F4DE46BFF5FFCEB9232FC67" pageId="26" pageNumber="193">
3. Diagonal lines on spinous portion of the dorsal fin 
<emphasis id="A0EC8C076F4DE46BFCD5FCEB9433FCFF" bold="true" box="[823,887,772,798]" pageId="26" pageNumber="193">thick</emphasis>
, sometimes barely recognisable as diagonal line but approaching blotches in shape, arranged as one longitudinal series with one line per membrane, dorsal-fin lappets orange; midlateral stripe on body oriented 
<emphasis id="A0EC8C076F4DE46BFBB3FCA59210FC82" bold="true" box="[1105,1364,841,868]" pageId="26" pageNumber="193">towards soft portion</emphasis>
of dorsal fin; suborbital stripe vertical, in adults barely visible ....................... 
<taxonomicName id="55982B966F4DE46BFBB6FC829232FC67" ID-CoL="NPVS" authority="(Cope, 1872)" baseAuthorityName="Cope" baseAuthorityYear="1872" box="[1108,1398,876,903]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="26" pageNumber="193" phylum="Chordata" rank="species" species="syspilus">
<emphasis id="A0EC8C076F4DE46BFBB6FC829391FC66" bold="true" box="[1108,1237,876,903]" italics="true" pageId="26" pageNumber="193">B. syspilus</emphasis>
(Cope, 1872)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="922750156F4DE46BFF5FFC609233FC48" blockId="26.[189,1399,772,937]" box="[189,1399,911,937]" pageId="26" pageNumber="193">
<keyLead id="2969F3206F4DE46BFF5FFC609233FC48" box="[189,1399,911,937]" pageId="26" pageNumber="193">
– Diagonal lines on spinous portion of the dorsal fin 
<emphasis id="A0EC8C076F4DE46BFCDCFC609435FC48" bold="true" box="[830,881,911,937]" pageId="26" pageNumber="193">thin</emphasis>
.................................................................. 4
</keyLead>
</paragraph>
</keyStep>
<keyStep id="296C48B06F4DE46BFF5FFC3F9233FBB2" pageId="26" pageNumber="193">
<paragraph id="922750156F4DE46BFF5FFC3F9232FBCE" blockId="26.[189,1399,976,1107]" pageId="26" pageNumber="193">
<keyLead id="2969F3206F4DE46BFF5FFC3F9232FBCE" pageId="26" pageNumber="193">
4. 
<emphasis id="A0EC8C076F4DE46BFF08FC3F9670FC0B" bold="true" box="[234,308,976,1002]" pageId="26" pageNumber="193">Three</emphasis>
diagonal lines vertically on each membrane of spinous portion of the dorsal fin, each line spanning three consecutive membranes, lines oriented distinctly diagonal, about 45%, dorsal-fin lappets orange; suborbital stripe vertical, in adults distinct................. 
<taxonomicName id="55982B966F4DE46BFBF4FBFA9232FBCE" ID-CoL="NPVH" authority="(Eigenmann, 1923)" baseAuthorityName="Eigenmann" baseAuthorityYear="1923" box="[1046,1398,1045,1071]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="26" pageNumber="193" phylum="Chordata" rank="species" species="mariae">
<emphasis id="A0EC8C076F4DE46BFBF4FBFA93C9FBCE" bold="true" box="[1046,1165,1045,1071]" italics="true" pageId="26" pageNumber="193">B. mariae</emphasis>
(Eigenmann, 1923)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="922750156F4DE46BFF5FFBD79233FBB2" blockId="26.[189,1399,976,1107]" box="[189,1399,1079,1107]" pageId="26" pageNumber="193">
<keyLead id="2969F3206F4DE46BFF5FFBD79233FBB2" box="[189,1399,1079,1107]" pageId="26" pageNumber="193">
– 
<emphasis id="A0EC8C076F4DE46BFF08FBD89630FBB3" bold="true" box="[234,372,1079,1106]" pageId="26" pageNumber="193">One to two</emphasis>
diagonal lines per membrane on spinous portion of the dorsal fin ............................... 5
</keyLead>
</paragraph>
</keyStep>
<keyStep id="296C48B06F4DE46BFF5FFB979233FA64" pageId="26" pageNumber="193">
<paragraph id="922750156F4DE46BFF5FFB979233FAFF" blockId="26.[189,1399,1144,1414]" pageId="26" pageNumber="193">
<keyLead id="2969F3206F4DE46BFF5FFB979233FAFF" pageId="26" pageNumber="193">
5. Lines per membrane oriented 
<emphasis id="A0EC8C076F4DE46BFDBAFB96942FFB73" bold="true" box="[600,875,1144,1171]" pageId="26" pageNumber="193">only slightly diagonal</emphasis>
, two lines only in largest adult specimens, one line in dorsal portion of membrane in all other sizes, dorsal-fin lappets dark, with orange tip; suborbital stripe bent from vertical to posteriad in juveniles, reduced to a preopercular spot in adults; large head, length equal or above 36.3% SL; longest snout of all species without overlap (mean 15.5%, 14.4–16.4% of SL) .................................................................................. 
<taxonomicName id="55982B966F4DE46BFB73FAEB9251FAFF" authority="Říčan &amp; Říčanová, 2023" authorityName="Říčan &amp; Říčanová" authorityYear="2023" box="[1169,1301,1284,1310]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="26" pageNumber="193" phylum="Chordata" rank="species" species="omagua" status="sp. nov.">
<emphasis id="A0EC8C076F4DE46BFB73FAEB9251FAFF" bold="true" box="[1169,1301,1284,1310]" italics="true" pageId="26" pageNumber="193">B. omagua</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBDF317C6F4DE46BFAFEFAEB9233FAFF" box="[1308,1399,1284,1310]" pageId="26" pageNumber="193" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="922750156F4DE46BFF5FFAC89233FA64" blockId="26.[189,1399,1144,1414]" pageId="26" pageNumber="193">
<keyLead id="2969F3206F4DE46BFF5FFAC89233FA64" pageId="26" pageNumber="193">
– Lines per membrane oriented 
<emphasis id="A0EC8C076F4DE46BFDBBFAC99400FAA1" bold="true" box="[601,836,1318,1344]" pageId="26" pageNumber="193">distinctly diagonal</emphasis>
, about 45%, dorsal-fin lappets nearly hyaline, only very slightly tinged orange; suborbital stripe slightly bent from vertical to posteriad, in adults well developed ......................................................................................... 
<taxonomicName id="55982B966F4DE46BFBD9FA839233FA64" ID-CoL="NPVM" authority="Kullander, 1986" authorityName="Kullander" authorityYear="1986" box="[1083,1399,1387,1414]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="26" pageNumber="193" phylum="Chordata" rank="species" species="ortegai">
<emphasis id="A0EC8C076F4DE46BFBD9FA8393F6FA64" bold="true" box="[1083,1202,1387,1414]" italics="true" pageId="26" pageNumber="193">B. ortegai</emphasis>
<bibRefCitation id="F6092DE46F4DE46BFB5BFA849233FA64" author="Kullander S." box="[1209,1399,1387,1413]" pageId="26" pageNumber="193" refId="ref14485" refString="Kullander S. 1986. Cichlid Fishes of the Amazon River Drainage of Peru. Swedish Museum of Natural History, Sweden." type="book" year="1986">Kullander, 1986</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
<paragraph id="922750156F4DE46BFF5FFA439233FA26" blockId="26.[189,1399,1452,1617]" box="[189,1399,1452,1479]" pageId="26" pageNumber="193">
<keyLead id="2969F3206F4DE46BFF5FFA439233FA26" box="[189,1399,1452,1479]" pageId="26" pageNumber="193">
6. Suborbital stripe 
<emphasis id="A0EC8C076F4DE46BFE56FA439556FA27" bold="true" box="[436,530,1452,1478]" pageId="26" pageNumber="193">vertical</emphasis>
.................................................................................................................. 7
</keyLead>
</paragraph>
<paragraph id="922750156F4DE46BFF5FFA209233FA08" blockId="26.[189,1399,1452,1617]" box="[189,1399,1487,1513]" pageId="26" pageNumber="193">
<keyLead id="2969F3206F4DE46BFF5FFA209233FA08" box="[189,1399,1487,1513]" pageId="26" pageNumber="193">
– Suborbital stripe 
<emphasis id="A0EC8C076F4DE46BFE56FA2096AFFA08" bold="true" box="[436,491,1487,1513]" pageId="26" pageNumber="193">bent</emphasis>
from vertical to posteriad .............................................................................. 8
</keyLead>
</paragraph>
<paragraph id="922750156F4DE46BFF5FFA1D9233F9B0" blockId="26.[189,1399,1452,1617]" pageId="26" pageNumber="193">
<keyLead id="2969F3206F4DE46BFF5FFA1D9233F9B0" pageId="26" pageNumber="193">
– Suborbital stripe 
<emphasis id="A0EC8C076F4DE46BFE5AFA1E955FF9EA" bold="true" box="[440,539,1521,1547]" pageId="26" pageNumber="193">inclined</emphasis>
posteriad, in adults reduced to its ventral portion in the form of a short stripe or a preopercular spot; dorsal-fin lappets dark to black, not orange ......................................... ............................................................................................................. 
<taxonomicName id="55982B966F4DE46BFBF7F9D89233F9B0" ID-CoL="NPVK" authority="Kullander, 1986" authorityName="Kullander" authorityYear="1986" box="[1045,1399,1591,1617]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="26" pageNumber="193" phylum="Chordata" rank="species" species="moriorum">
<emphasis id="A0EC8C076F4DE46BFBF7F9D893F5F9B0" bold="true" box="[1045,1201,1591,1617]" italics="true" pageId="26" pageNumber="193">B. moriorum</emphasis>
<bibRefCitation id="F6092DE46F4DE46BFB5BF9D89233F9B0" author="Kullander S." box="[1209,1399,1591,1617]" pageId="26" pageNumber="193" refId="ref14485" refString="Kullander S. 1986. Cichlid Fishes of the Amazon River Drainage of Peru. Swedish Museum of Natural History, Sweden." type="book" year="1986">Kullander, 1986</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="922750156F4DE46BFF5FF9979233F936" blockId="26.[189,1399,1656,1821]" pageId="26" pageNumber="193">
<keyLead id="2969F3206F4DE46BFF5FF9979233F936" pageId="26" pageNumber="193">
7. Suborbital stripe 
<emphasis id="A0EC8C076F4DE46BFE5BF9979553F973" bold="true" box="[441,535,1656,1682]" pageId="26" pageNumber="193">vertical</emphasis>
, 
<emphasis id="A0EC8C076F4DE46BFDCBF9979455F973" bold="true" box="[553,785,1656,1682]" pageId="26" pageNumber="193">shifted posteriorly</emphasis>
to a position behind the eye, in adults always as a stripe, sometimes reduced only to its ventral portion; dorsal-fin lappets orange-brown, same color as rest of markings on dorsal fin, not orange ................................. 
<taxonomicName id="55982B966F4DE46BFC2FF9529233F936" ID-CoL="NPVQ" authority="Kullander, 1986" authorityName="Kullander" authorityYear="1986" box="[973,1399,1725,1752]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="26" pageNumber="193" phylum="Chordata" rank="species" species="peregrinabunda">
<emphasis id="A0EC8C076F4DE46BFC2FF95293F6F939" bold="true" box="[973,1202,1725,1752]" italics="true" pageId="26" pageNumber="193">B. peregrinabunda</emphasis>
<bibRefCitation id="F6092DE46F4DE46BFB5BF9529233F936" author="Kullander S." box="[1209,1399,1725,1751]" pageId="26" pageNumber="193" refId="ref14485" refString="Kullander S. 1986. Cichlid Fishes of the Amazon River Drainage of Peru. Swedish Museum of Natural History, Sweden." type="book" year="1986">Kullander, 1986</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="922750156F4DE46BFF5FF90F9233F8FD" blockId="26.[189,1399,1656,1821]" pageId="26" pageNumber="193">
<keyLead id="2969F3206F4DE46BFF5FF90F9233F8FD" pageId="26" pageNumber="193">
– Suborbital stripe 
<emphasis id="A0EC8C076F4DE46BFE5AF90F9552F91B" bold="true" box="[440,534,1760,1786]" pageId="26" pageNumber="193">vertical</emphasis>
, in adults 
<emphasis id="A0EC8C076F4DE46BFD7BF90F95A4F91B" bold="true" box="[665,736,1760,1786]" pageId="26" pageNumber="193">whole</emphasis>
; dorsal-fin lappets orange-brown, same color as rest of markings on dorsal fin, not orange ..................................................... 
<taxonomicName id="55982B966F4DE46BFBEDF8EC9233F8FD" ID-CoL="NPVF" authority="Kullander, 1986" authorityName="Kullander" authorityYear="1986" box="[1039,1399,1794,1821]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="26" pageNumber="193" phylum="Chordata" rank="species" species="huallagae">
<emphasis id="A0EC8C076F4DE46BFBEDF8EC93EFF8FC" bold="true" box="[1039,1195,1794,1821]" italics="true" pageId="26" pageNumber="193">B. huallagae</emphasis>
<bibRefCitation id="F6092DE46F4DE46BFB5BF8ED9233F8FD" author="Kullander S." box="[1209,1399,1794,1820]" pageId="26" pageNumber="193" refId="ref14485" refString="Kullander S. 1986. Cichlid Fishes of the Amazon River Drainage of Peru. Swedish Museum of Natural History, Sweden." type="book" year="1986">Kullander, 1986</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="922750156F4DE46BFF5FF8AC9232F842" blockId="26.[189,1399,1859,2025]" pageId="26" pageNumber="193">
<keyLead id="2969F3206F4DE46BFF5FF8AC9232F842" pageId="26" pageNumber="193">
8. Suborbital stripe 
<emphasis id="A0EC8C076F4DE46BFE5AF8AC96ABF8BC" bold="true" box="[440,495,1859,1885]" pageId="26" pageNumber="193">bent</emphasis>
from vertical to posteriad, in adults 
<emphasis id="A0EC8C076F4DE46BFC47F8AC965AF861" bold="true" pageId="26" pageNumber="193">reduced to an ocellated preopercular spot</emphasis>
; dorsal-fin lappets orange-brown, same color as rest of markings on dorsal fin, not orange ..... .............................................................................................................. 
<taxonomicName id="55982B966F4DE46BFBFCF8669232F842" ID-CoL="NPVV" authority="(Regan, 1905)" authorityName="" baseAuthorityName="Regan" baseAuthorityYear="1905" box="[1054,1398,1928,1955]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="26" pageNumber="193" phylum="Chordata" rank="species" species="zamorensis">
<emphasis id="A0EC8C076F4DE46BFBFCF8669383F842" bold="true" box="[1054,1223,1929,1955]" italics="true" pageId="26" pageNumber="193">B. zamorensis</emphasis>
(Regan, 1905)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="922750156F4DE46BFF5FF8439233F809" blockId="26.[189,1399,1859,2025]" pageId="26" pageNumber="193">
<keyLead id="2969F3206F4DE46BFF5FF8439233F809" pageId="26" pageNumber="193">
– Suborbital stripe 
<emphasis id="A0EC8C076F4DE46BFE51F84496AEF824" bold="true" box="[435,490,1963,1989]" pageId="26" pageNumber="193">bent</emphasis>
from vertical to posteriad, in adults 
<emphasis id="A0EC8C076F4DE46BFC70F844949FF824" bold="true" box="[914,987,1963,1989]" pageId="26" pageNumber="193">whole</emphasis>
; dorsal-fin lappets 
<emphasis id="A0EC8C076F4DE46BFB5EF8449244F824" bold="true" box="[1212,1280,1963,1989]" pageId="26" pageNumber="193">white</emphasis>
with dark base ................................................... 
<taxonomicName id="55982B966F4DE46BFD44F8219233F809" ID-CoL="NPVP" authority="Arbour, Barriga Salazar &amp; Lopez-Fernandez, 2014" authorityName="Arbour, Barriga Salazar &amp; Lopez-Fernandez" authorityYear="2014" box="[678,1399,1997,2024]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="26" pageNumber="193" phylum="Chordata" rank="species" species="pardus">
<emphasis id="A0EC8C076F4DE46BFD44F821945FF809" bold="true" box="[678,795,1997,2024]" italics="true" pageId="26" pageNumber="193">B. pardus</emphasis>
<bibRefCitation id="F6092DE46F4DE46BFCC1F8219233F809" author="Arbour J. H. &amp; Barriga Salazar R. E. &amp; Lopez-Fernandez H." box="[803,1399,1998,2024]" pageId="26" pageNumber="193" pagination="79 - 86" refId="ref13136" refString="Arbour J. H., Barriga Salazar R. E. &amp; Lopez-Fernandez H. 2014. A new species of Bujurquina (Teleostei: Cichlidae) from the Rio Danta, Ecuador, with a key to the species in the genus. Copeia 2014: 79 - 86. https: // doi. org / 10.1643 / CI- 13 - 028" type="journal article" year="2014">Arbour, Barriga Salazar &amp; López-Fernández, 2014</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="922750156F4CE46AFF5FFEE69232FEC2" blockId="27.[189,1398,265,327]" box="[189,1398,265,291]" pageId="27" pageNumber="194">
<keyLead id="2969F3206F4CE46AFF5FFEE69232FEC2" box="[189,1398,265,291]" pageId="27" pageNumber="194">
9. Head 
<emphasis id="A0EC8C076F4CE46AFED2FEE69634FEC2" bold="true" box="[304,368,265,291]" pageId="27" pageNumber="194">large</emphasis>
, length equal or above 37% SL .................................................................................... 10
</keyLead>
</paragraph>
<paragraph id="922750156F4CE46AFF5FFEC29232FEA7" blockId="27.[189,1398,265,327]" box="[189,1398,300,327]" pageId="27" pageNumber="194">
<keyLead id="2969F3206F4CE46AFF5FFEC29232FEA7" box="[189,1398,300,327]" pageId="27" pageNumber="194">
– Head 
<emphasis id="A0EC8C076F4CE46AFED2FEC39637FEA7" bold="true" box="[304,371,300,326]" pageId="27" pageNumber="194">small</emphasis>
, mean length below 35.5% SL ......................................................................................11
</keyLead>
</paragraph>
<paragraph id="922750156F4CE46AFF5FFE819233FE2F" blockId="27.[189,1399,365,568]" pageId="27" pageNumber="194">
<keyLead id="2969F3206F4CE46AFF5FFE819233FE2F" pageId="27" pageNumber="194">
10. Mean length of head above 38.6% of SL, average 39.1%, lower pharyngeal teeth molariform, suborbital stripe in adults complete, dorsal-fin lappets white with dark base .................................... .......................................................................................................... 
<taxonomicName id="55982B966F4CE46AFC1CFE5B9233FE2F" ID-CoL="NPVL" authority="Kullander, 1987" authorityName="Kullander" authorityYear="1987" box="[1022,1399,435,462]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="27" pageNumber="194" phylum="Chordata" rank="species" species="oenolaemus">
<emphasis id="A0EC8C076F4CE46AFC1CFE5B93F6FE2F" bold="true" box="[1022,1202,435,462]" italics="true" pageId="27" pageNumber="194">B. oenolaemus</emphasis>
<bibRefCitation id="F6092DE46F4CE46AFB5BFE5B9233FE2F" author="Kullander S. O." box="[1209,1399,436,462]" pageId="27" pageNumber="194" pagination="195 - 205" refId="ref14508" refString="Kullander S. O. 1987. Cichlid fishes from the La Plata Basin. Part VI. Description of a new Bujurquina species from Bolivia. Cybium 1987: 195 - 205." type="journal article" year="1987">Kullander, 1987</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="922750156F4CE46AFF5FFE379233FDD6" blockId="27.[189,1399,365,568]" pageId="27" pageNumber="194">
<keyLead id="2969F3206F4CE46AFF5FFE379233FDD6" pageId="27" pageNumber="194">
– Mean length of head above 37%, lips hypertrophied, suborbital stripe in adults incomplete, limited to below eye or to middle of cheek (does not reach angle of preopercle), dorsal-fin lappets white with dark base .................................................................................................. 
<taxonomicName id="55982B966F4CE46AFBDBFDF19233FDD6" ID-CoL="NPVG" authority="Kullander, 1986" authorityName="Kullander" authorityYear="1986" box="[1081,1399,541,568]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="27" pageNumber="194" phylum="Chordata" rank="species" species="labiosa">
<emphasis id="A0EC8C076F4CE46AFBDBFDF193F5FDD9" bold="true" box="[1081,1201,541,568]" italics="true" pageId="27" pageNumber="194">B. labiosa</emphasis>
<bibRefCitation id="F6092DE46F4CE46AFB5BFDF19233FDD6" author="Kullander S." box="[1209,1399,541,568]" pageId="27" pageNumber="194" refId="ref14485" refString="Kullander S. 1986. Cichlid Fishes of the Amazon River Drainage of Peru. Swedish Museum of Natural History, Sweden." type="book" year="1986">Kullander, 1986</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="922750156F4CE46AFF5FFDB09232FD98" blockId="27.[189,1399,607,774]" box="[189,1398,607,634]" pageId="27" pageNumber="194">
<keyLead id="2969F3206F4CE46AFF5FFDB09232FD98" box="[189,1398,607,634]" pageId="27" pageNumber="194">
11. Suborbital stripe in adults 
<emphasis id="A0EC8C076F4CE46AFDC2FDB095D4FD98" bold="true" box="[544,656,607,633]" pageId="27" pageNumber="194">complete</emphasis>
............................................................................................... 12
</keyLead>
</paragraph>
<paragraph id="922750156F4CE46AFF5FFD6C9232FD7D" blockId="27.[189,1399,607,774]" box="[189,1398,642,669]" pageId="27" pageNumber="194">
<keyLead id="2969F3206F4CE46AFF5FFD6C9232FD7D" box="[189,1398,642,669]" pageId="27" pageNumber="194">
– Suborbital stripe in adults 
<emphasis id="A0EC8C076F4CE46AFDC2FD6D95EDFD7D" bold="true" box="[544,681,642,668]" pageId="27" pageNumber="194">incomplete</emphasis>
........................................................................................... 13
</keyLead>
</paragraph>
<paragraph id="922750156F4CE46AFF5FFD499232FD21" blockId="27.[189,1399,607,774]" box="[189,1398,677,704]" pageId="27" pageNumber="194">
<keyLead id="2969F3206F4CE46AFF5FFD499232FD21" box="[189,1398,677,704]" pageId="27" pageNumber="194">
– Suborbital stripe in adults 
<emphasis id="A0EC8C076F4CE46AFDC2FD4995C0FD21" bold="true" box="[544,644,678,704]" pageId="27" pageNumber="194">reduced</emphasis>
to preopercular spot ............................................................... 14
</keyLead>
</paragraph>
<paragraph id="922750156F4CE46AFF5FFD269233FCE7" blockId="27.[189,1399,607,774]" pageId="27" pageNumber="194">
<keyLead id="2969F3206F4CE46AFF5FFD269233FCE7" pageId="27" pageNumber="194">
– Suborbital stripe in adults 
<emphasis id="A0EC8C076F4CE46AFDC2FD269535FD02" bold="true" box="[544,625,713,739]" pageId="27" pageNumber="194">absent</emphasis>
, dorsal-fin lappets 
<emphasis id="A0EC8C076F4CE46AFCB0FD2794D2FD03" bold="true" box="[850,918,712,738]" pageId="27" pageNumber="194">white</emphasis>
with dark base ....................................... ............................................................................................................... 
<taxonomicName id="55982B966F4CE46AFBC0FD039233FCE7" ID-CoL="NPVD" authority="Kullander, 1986" authorityName="Kullander" authorityYear="1986" box="[1058,1399,747,774]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="27" pageNumber="194" phylum="Chordata" rank="species" species="eurhinus">
<emphasis id="A0EC8C076F4CE46AFBC0FD0393F5FCE7" bold="true" box="[1058,1201,747,774]" italics="true" pageId="27" pageNumber="194">B. eurhinus</emphasis>
<bibRefCitation id="F6092DE46F4CE46AFB5BFD039233FCE7" author="Kullander S." box="[1209,1399,748,774]" pageId="27" pageNumber="194" refId="ref14485" refString="Kullander S. 1986. Cichlid Fishes of the Amazon River Drainage of Peru. Swedish Museum of Natural History, Sweden." type="book" year="1986">Kullander, 1986</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="922750156F4CE46AFF5FFCC29232FC8A" blockId="27.[189,1399,813,981]" pageId="27" pageNumber="194">
<keyLead id="2969F3206F4CE46AFF5FFCC29232FC8A" pageId="27" pageNumber="194">
12. Suborbital stripe in adults 
<emphasis id="A0EC8C076F4CE46AFDC2FCC295D4FCA6" bold="true" box="[544,656,813,839]" pageId="27" pageNumber="194">complete</emphasis>
, 
<emphasis id="A0EC8C076F4CE46AFD7DFCC295B8FCA6" bold="true" box="[671,764,813,839]" pageId="27" pageNumber="194">distinct</emphasis>
, dorsal-fin lappets dark red to brown ......................... ...................................................................................................................... 
<taxonomicName id="55982B966F4CE46AFBB0FCBE9232FC8A" ID-CoL="694C5" authority="(Heckel, 1840)" baseAuthorityName="Heckel" baseAuthorityYear="1840" box="[1106,1398,848,875]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="27" pageNumber="194" phylum="Chordata" rank="species" species="vittata">
<emphasis id="A0EC8C076F4CE46AFBB0FCBE93FBFC8B" bold="true" box="[1106,1215,848,875]" italics="true" pageId="27" pageNumber="194">B. vittata</emphasis>
(Heckel, 1840)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="922750156F4CE46AFF5FFC9B9233FC50" blockId="27.[189,1399,813,981]" pageId="27" pageNumber="194">
<keyLead id="2969F3206F4CE46AFF5FFC9B9233FC50" pageId="27" pageNumber="194">
– Suborbital stripe in adults 
<emphasis id="A0EC8C076F4CE46AFDC2FC9B95D4FC6F" bold="true" box="[544,656,884,910]" pageId="27" pageNumber="194">complete</emphasis>
, 
<emphasis id="A0EC8C076F4CE46AFD7DFC9C9450FC6C" bold="true" box="[671,788,883,909]" pageId="27" pageNumber="194">indistinct</emphasis>
, dorsal-fin lappets distinctly 
<emphasis id="A0EC8C076F4CE46AFB88FC9B9384FC6F" bold="true" box="[1130,1216,884,910]" pageId="27" pageNumber="194">orange</emphasis>
...................... .................................................................................................................. 
<taxonomicName id="55982B966F4CE46AFBD6FC789233FC50" ID-CoL="NPVR" authority="Kullander, 1986" authorityName="Kullander" authorityYear="1986" box="[1076,1399,918,945]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="27" pageNumber="194" phylum="Chordata" rank="species" species="robusta">
<emphasis id="A0EC8C076F4CE46AFBD6FC7893F5FC50" bold="true" box="[1076,1201,918,945]" italics="true" pageId="27" pageNumber="194">B. robusta</emphasis>
<bibRefCitation id="F6092DE46F4CE46AFB5BFC789233FC50" author="Kullander S." box="[1209,1399,919,945]" pageId="27" pageNumber="194" refId="ref14485" refString="Kullander S. 1986. Cichlid Fishes of the Amazon River Drainage of Peru. Swedish Museum of Natural History, Sweden." type="book" year="1986">Kullander, 1986</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="922750156F4CE46AFF5FFC549233FC35" blockId="27.[189,1399,813,981]" box="[189,1399,954,981]" pageId="27" pageNumber="194">
<keyLead id="2969F3206F4CE46AFF5FFC549233FC35" box="[189,1399,954,981]" pageId="27" pageNumber="194">
– Suborbital stripe 
<emphasis id="A0EC8C076F4CE46AFE56FC559560FC35" bold="true" box="[436,548,954,980]" pageId="27" pageNumber="194">complete</emphasis>
, 
<emphasis id="A0EC8C076F4CE46AFDD1FC5595ECFC35" bold="true" box="[563,680,954,980]" pageId="27" pageNumber="194">indistinct</emphasis>
, dorsal-fin lappets 
<emphasis id="A0EC8C076F4CE46AFC68FC55948AFC35" bold="true" box="[906,974,954,980]" pageId="27" pageNumber="194">white</emphasis>
...... 
<taxonomicName id="55982B966F4CE46AFBEBFC559233FC35" ID-CoL="NPVC" authority="Kullander, 1986" authorityName="Kullander" authorityYear="1986" box="[1033,1399,954,981]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="27" pageNumber="194" phylum="Chordata" rank="species" species="cordemadi">
<emphasis id="A0EC8C076F4CE46AFBEBFC5593EEFC34" bold="true" box="[1033,1194,954,981]" italics="true" pageId="27" pageNumber="194">B. cordemadi</emphasis>
<bibRefCitation id="F6092DE46F4CE46AFB5BFC559233FC35" author="Kullander S." box="[1209,1399,954,980]" pageId="27" pageNumber="194" refId="ref14485" refString="Kullander S. 1986. Cichlid Fishes of the Amazon River Drainage of Peru. Swedish Museum of Natural History, Sweden." type="book" year="1986">Kullander, 1986</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="922750156F4CE46AFF5FFC139233FBD8" blockId="27.[189,1399,1019,1152]" pageId="27" pageNumber="194">
<keyLead id="2969F3206F4CE46AFF5FFC139233FBD8" pageId="27" pageNumber="194">
13. Suborbital stripe in adults 
<emphasis id="A0EC8C076F4CE46AFDC1FC1494F3FBF7" bold="true" box="[547,951,1019,1046]" pageId="27" pageNumber="194">incomplete, limited to below eye</emphasis>
, weakly developed, dorsal-fin lappets 
<emphasis id="A0EC8C076F4CE46AFF08FBF09650FBD8" bold="true" box="[234,276,1055,1081]" pageId="27" pageNumber="194">red</emphasis>
in anterior portion, 
<emphasis id="A0EC8C076F4CE46AFDE2FBF09500FBD8" bold="true" box="[512,580,1055,1081]" pageId="27" pageNumber="194">black</emphasis>
in posterior portion 
<emphasis id="A0EC8C076F4CE46AFCD9FBF094AFFBD9" box="[827,1003,1055,1080]" italics="true" pageId="27" pageNumber="194">........................</emphasis>
<taxonomicName id="55982B966F4CE46AFC13FBF09233FBD8" ID-CoL="NPVJ" authority="Kullander, 1986" authorityName="Kullander" authorityYear="1986" box="[1009,1399,1054,1081]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="27" pageNumber="194" phylum="Chordata" rank="species" species="megalospilus">
<emphasis id="A0EC8C076F4CE46AFC13FBF093F6FBD8" bold="true" box="[1009,1202,1054,1081]" italics="true" pageId="27" pageNumber="194">B. megalospilus</emphasis>
<bibRefCitation id="F6092DE46F4CE46AFB5BFBF09233FBD8" author="Kullander S." box="[1209,1399,1055,1081]" pageId="27" pageNumber="194" refId="ref14485" refString="Kullander S. 1986. Cichlid Fishes of the Amazon River Drainage of Peru. Swedish Museum of Natural History, Sweden." type="book" year="1986">Kullander, 1986</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="922750156F4CE46AFF5FFBAD9233FB9E" blockId="27.[189,1399,1019,1152]" pageId="27" pageNumber="194">
<keyLead id="2969F3206F4CE46AFF5FFBAD9233FB9E" pageId="27" pageNumber="194">
– Suborbital stripe in adults 
<emphasis id="A0EC8C076F4CE46AFDC1FBAD94F3FBBD" bold="true" box="[547,951,1090,1116]" pageId="27" pageNumber="194">incomplete, limited to below eye</emphasis>
, weakly developed, dorsal-fin lappets 
<emphasis id="A0EC8C076F4CE46AFF08FB8A966AFB9E" bold="true" box="[234,302,1125,1151]" pageId="27" pageNumber="194">white</emphasis>
................................................................................................. 
<taxonomicName id="55982B966F4CE46AFBE1FB8A9233FB9E" ID-CoL="NPVT" authority="Kullander, 1986" authorityName="Kullander" authorityYear="1986" box="[1027,1399,1125,1152]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="27" pageNumber="194" phylum="Chordata" rank="species" species="tambopatae">
<emphasis id="A0EC8C076F4CE46AFBE1FB8A93F6FB61" bold="true" box="[1027,1202,1125,1152]" italics="true" pageId="27" pageNumber="194">B. tambopatae</emphasis>
<bibRefCitation id="F6092DE46F4CE46AFB5BFB8A9233FB9E" author="Kullander S." box="[1209,1399,1125,1151]" pageId="27" pageNumber="194" refId="ref14485" refString="Kullander S. 1986. Cichlid Fishes of the Amazon River Drainage of Peru. Swedish Museum of Natural History, Sweden." type="book" year="1986">Kullander, 1986</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="922750156F4CE46AFF5FFB489233FB05" blockId="27.[189,1399,1190,1323]" pageId="27" pageNumber="194">
<keyLead id="2969F3206F4CE46AFF5FFB489233FB05" pageId="27" pageNumber="194">
14. Suborbital stripe in adults 
<emphasis id="A0EC8C076F4CE46AFDFDFB4895C7FB20" bold="true" box="[543,643,1191,1217]" pageId="27" pageNumber="194">reduced</emphasis>
to preopercular spot, 
<emphasis id="A0EC8C076F4CE46AFC60FB48937CFB20" bold="true" box="[898,1080,1191,1217]" pageId="27" pageNumber="194">well developed</emphasis>
, in most specimens clearly visible, dorsal-fin lappets with 
<emphasis id="A0EC8C076F4CE46AFDB7FB2595D5FB05" bold="true" box="[597,657,1226,1252]" pageId="27" pageNumber="194">dark</emphasis>
tips ................................................ 
<taxonomicName id="55982B966F4CE46AFBCCFB259233FB05" ID-CoL="694BT" authority="Kullander, 1986" authorityName="Kullander" authorityYear="1986" box="[1070,1399,1225,1252]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="27" pageNumber="194" phylum="Chordata" rank="species" species="hophrys">
<emphasis id="A0EC8C076F4CE46AFBCCFB2593F6FB05" bold="true" box="[1070,1202,1225,1252]" italics="true" pageId="27" pageNumber="194">B. hophrys</emphasis>
<bibRefCitation id="F6092DE46F4CE46AFB5BFB259233FB05" author="Kullander S." box="[1209,1399,1226,1252]" pageId="27" pageNumber="194" refId="ref14485" refString="Kullander S. 1986. Cichlid Fishes of the Amazon River Drainage of Peru. Swedish Museum of Natural History, Sweden." type="book" year="1986">Kullander, 1986</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="922750156F4CE46AFF5FFB029233FACB" blockId="27.[189,1399,1190,1323]" pageId="27" pageNumber="194">
<keyLead id="2969F3206F4CE46AFF5FFB029233FACB" pageId="27" pageNumber="194">
– Suborbital stripe in adults 
<emphasis id="A0EC8C076F4CE46AFDF9FB02953BFAE6" bold="true" box="[539,639,1261,1287]" pageId="27" pageNumber="194">reduced</emphasis>
to preopercular spot, 
<emphasis id="A0EC8C076F4CE46AFC9BFB0293D5FAE6" bold="true" box="[889,1169,1261,1287]" pageId="27" pageNumber="194">very weakly developed</emphasis>
, in most specimens almost invisible, dorsal-fin lappets with 
<emphasis id="A0EC8C076F4CE46AFD23FAFF941FFACB" bold="true" box="[705,859,1296,1322]" pageId="27" pageNumber="194">dark orange</emphasis>
tips ............. 
<taxonomicName id="55982B966F4CE46AFC1FFAFF9233FACB" ID-CoL="NPVB" authority="Kullander, 1986" authorityName="Kullander" authorityYear="1986" box="[1021,1399,1296,1322]" class="Actinopterygii" family="Cichlidae" genus="Bujurquina" kingdom="Animalia" order="Perciformes" pageId="27" pageNumber="194" phylum="Chordata" rank="species" species="apoparuana">
<emphasis id="A0EC8C076F4CE46AFC1FFAFF93F6FACB" bold="true" box="[1021,1202,1296,1322]" italics="true" pageId="27" pageNumber="194">B. apoparuana</emphasis>
<bibRefCitation id="F6092DE46F4CE46AFB5BFAFF9233FACB" author="Kullander S." box="[1209,1399,1296,1322]" pageId="27" pageNumber="194" refId="ref14485" refString="Kullander S. 1986. Cichlid Fishes of the Amazon River Drainage of Peru. Swedish Museum of Natural History, Sweden." type="book" year="1986">Kullander, 1986</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</subSubSection>
</treatment>
</document>