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<document id="5572B051C0C903D5685CE65C83F90AAE" ID="10.11646/zootaxa.4170.3.3" ID-DOI="10.11646/zootaxa.4170.3.3" ID-GBIF-Dataset="e9fcf827-dcd7-4575-ae01-eee9465760f0" ID-ISSN="1175-5326" ID-Zenodo-Dep="260651" ID-ZooBank="BFF1365C-78F6-4B7B-8F70-57B28C3A1138" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1474872590518" checkinUser="plazi" docAuthor="Solodovnikov, Alexey &amp; Hansen, Aslak Kappel" docDate="2016" docId="156487CBE8621C6BFF55F9A4FAC2DD3C" docLanguage="en" docName="zootaxa.4170.3.3.pdf" docOrigin="Zootaxa 4170 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Quedius roma Solodovnikov et Hansen, sp. n." docType="treatment" docVersion="9" lastPageNumber="487" masterDocId="E95DFFB3E86A1C67FFC2FF93FFA5DF01" masterDocTitle="Review of subterranean Quedius, with description of the first hypogean species from the Russian Far East (Coleoptera: Staphylinidae: Staphylinini)" masterLastPageNumber="490" masterPageNumber="475" pageNumber="483" updateTime="1698387186525" updateUser="plazi">
<mods:mods id="DB723B93C98C721100638C495238543D" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="0DE75A29F476D35D72E85321FB16C6D5">
<mods:title id="702096DC39AC49502F3ADADDD1CE587F">Review of subterranean Quedius, with description of the first hypogean species from the Russian Far East (Coleoptera: Staphylinidae: Staphylinini)</mods:title>
</mods:titleInfo>
<mods:name id="B7AD8AD4994590128A900A18E25B25D4" type="personal">
<mods:role id="3C4568E69B13C2D40E93D4430A50DFE1">
<mods:roleTerm id="A51E2625115988585187ADC6A8591781">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DA5F38F724B3D0F2D069C843C34B65DC">Solodovnikov, Alexey</mods:namePart>
</mods:name>
<mods:name id="9EB51C146EB93DCC34A6699DFCEE03D7" type="personal">
<mods:role id="FA353F25502ABF784832B08B3FEEFCB1">
<mods:roleTerm id="CD17D2F8BEA6F8059C2BE1D6DB91CA12">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="75FBEB717D4873208F5CE986AB31D250">Hansen, Aslak Kappel</mods:namePart>
</mods:name>
<mods:typeOfResource id="74ACC5BC129FD505F3381395FA8DF3A7">text</mods:typeOfResource>
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<mods:title id="FB32B507E6996A40B4B52C9ABA629153">Zootaxa</mods:title>
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<mods:part id="94915AE27024395F60012F5B3F07CEF5">
<mods:date id="BB2A66D3A25A9B0734ACD5D7BEC0C0B2">2016</mods:date>
<mods:detail id="F319362DC71B337661F4891289C75C07" type="volume">
<mods:number id="D416FA8AC469E67EC2E2B22A64100462">4170</mods:number>
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<mods:detail id="198477B6CEF3561F6ABBDDCCD08E6458" type="issue">
<mods:number id="6C145F04C71CA7FF138EA2967E215316">3</mods:number>
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<mods:start id="EB883B57E398CC96FABD9F2BC4E4A3DF">475</mods:start>
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<mods:classification id="13D7E688CFD05A741F1DCE3E15B7BECC">journal article</mods:classification>
<mods:identifier id="6D83875F928BB6DD339BBD38D68372D1" type="DOI">10.11646/zootaxa.4170.3.3</mods:identifier>
<mods:identifier id="8135EEC22FACFD8D351D1F6AE72AC0F5" type="GBIF-Dataset">e9fcf827-dcd7-4575-ae01-eee9465760f0</mods:identifier>
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<mods:identifier id="848F190DEE732A4D7F5FE351743BF4E9" type="Zenodo-Dep">260651</mods:identifier>
<mods:identifier id="14336923D57EEBEF2AFA5E3D6927C28C" type="ZooBank">BFF1365C-78F6-4B7B-8F70-57B28C3A1138</mods:identifier>
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<treatment id="156487CBE8621C6BFF55F9A4FAC2DD3C" ID-DOI="http://doi.org/10.5281/zenodo.5674848" ID-GBIF-Taxon="124552847" ID-Zenodo-Dep="5674848" LSID="urn:lsid:plazi:treatment:156487CBE8621C6BFF55F9A4FAC2DD3C" httpUri="http://treatment.plazi.org/id/156487CBE8621C6BFF55F9A4FAC2DD3C" lastPageId="12" lastPageNumber="487" pageId="8" pageNumber="483">
<subSubSection id="D5D76556E8621C6FFF55F9A4FE84D972" pageId="8" pageNumber="483" type="nomenclature">
<paragraph id="9D7236DDE8621C6FFF55F9A4FD6DD953" blockId="8.[151,712,1591,1651]" box="[151,712,1591,1618]" pageId="8" pageNumber="483">
<heading id="C63A81B1E8621C6FFF55F9A4FD6DD953" bold="true" box="[151,712,1591,1618]" fontSize="11" level="1" pageId="8" pageNumber="483" reason="1">
<emphasis id="AFB9EACFE8621C6FFF55F9A4FD6DD953" bold="true" box="[151,712,1591,1618]" pageId="8" pageNumber="483">
<taxonomicName id="5ACD4D5EE8621C6FFF55F9A4FDDCD953" ID-CoL="6WPY4" authority="Solodovnikov et Hansen" authorityName="Solodovnikov et Hansen" box="[151,633,1591,1618]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="483" phylum="Arthropoda" rank="species" species="roma" status="sp. nov.">
<emphasis id="AFB9EACFE8621C6FFF55F9A4FEE7D950" bold="true" box="[151,322,1591,1617]" italics="true" pageId="8" pageNumber="483">Quedius roma</emphasis>
Solodovnikov et Hansen
</taxonomicName>
,
<taxonomicNameLabel id="B48A57B4E8621C6FFD44F9ABFD6DD953" box="[646,712,1592,1618]" pageId="8" pageNumber="483" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="9D7236DDE8621C6FFF55F9C9FE84D972" blockId="8.[151,712,1591,1651]" box="[151,289,1626,1651]" pageId="8" pageNumber="483">
<figureCitation id="05F62A58E8621C6FFF55F9C9FF79D972" box="[151,220,1626,1651]" captionStart="FIGURE 3" captionStartId="9.[151,250,1345,1367]" captionText="FIGURE 3. Quedius roma sp. n., habitus. Scale bar 5 mm." pageId="8" pageNumber="483">Figs 3</figureCitation>
and
<figureCitation id="05F62A58E8621C6FFED1F9C8FE84D972" box="[275,289,1627,1651]" captionStart="FIGURE 4" captionStartId="10.[151,250,1916,1938]" captionTargetBox="[240,1354,185,1893]" captionTargetId="figure@10.[222,1354,185,1893]" captionTargetPageId="10" captionText="FIGURE 4. Quedius roma sp. n., details of morphology: head dorsally (A); pronotum dorsally (B); male sternite VIII (C); male sternite IX (D); male tergite X (E); aedeagus, parameral side (F); aedeagus, laterally (G); paramere, side facing median lobe (H); female tergite X (I); d—dorsal puncture, f—frontal puncture, s—sublateral puncture, v—vertical puncture. Scale bars 1 mm." httpUri="https://zenodo.org/record/260654/files/figure.png" pageId="8" pageNumber="483">4</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="D5D76556E8621C6FFF55F931FDE9D826" pageId="8" pageNumber="483" type="materials_examined">
<paragraph id="9D7236DDE8621C6FFF55F931FDE9D826" blockId="8.[151,1436,1698,1975]" pageId="8" pageNumber="483">
<materialsCitation id="2DA53C80E8621C6FFF55F931FD24D802" ID-GBIF-Occurrence="1315044601" collectingDate="2015-05-26" collectionCode="ZMUC" collectorName="A. Hansen &amp; M. Justesen &amp; A. Solodovnikov" country="Russia" county="river Ko" elevation="750" latitude="47.0716" location="Material" longLatPrecision="6" longitude="136.4572" municipality="Sikhote-Alin" pageId="8" pageNumber="483" specimenCount="1" typeStatus="holotype">
<emphasis id="AFB9EACFE8621C6FFF55F931FE26D9BA" bold="true" box="[151,387,1698,1723]" pageId="8" pageNumber="483">
<location id="98126006E8621C6FFF55F931FF5AD9BA" LSID="urn:lsid:plazi:treatment:156487CBE8621C6BFF55F9A4FAC2DD3C:98126006E8621C6FFF55F931FF5AD9BA" box="[151,255,1698,1723]" country="Russia" county="river Ko" latitude="47.0716" longLatPrecision="6" longitude="136.4572" municipality="Sikhote-Alin" name="Material" pageId="8" pageNumber="483">Material</location>
examined.
</emphasis>
<typeStatus id="4276887FE8621C6FFE4EF931FE5CD9BA" box="[396,505,1698,1723]" pageId="8" pageNumber="483" type="holotype">Holotype</typeStatus>
: male,
<collectingCountry id="E5DA764DE8621C6FFD8EF931FD3CD9BA" box="[588,665,1698,1723]" name="Russia" pageId="8" pageNumber="483">Russia</collectingCountry>
,
<location id="98126006E8621C6FFD65F931FCDCD9BA" LSID="urn:lsid:plazi:treatment:156487CBE8621C6BFF55F9A4FAC2DD3C:98126006E8621C6FFD65F931FCDCD9BA" box="[679,889,1698,1723]" country="Russia" county="river Ko" latitude="47.0716" longLatPrecision="6" longitude="136.4572" municipality="Sikhote-Alin" name="Khabarovskij Kraj" pageId="8" pageNumber="483">Khabarovskij Kraj</location>
(southern),
<collectingMunicipality id="7D16ACA7E8621C6FFBC4F931FB32D9BA" box="[1030,1175,1698,1723]" pageId="8" pageNumber="483">Sikhote-Alin</collectingMunicipality>
(Central) range, upper course of
<collectingCounty id="74134E51E8621C6FFEC4F955FEC2D9DF" box="[262,359,1734,1759]" pageId="8" pageNumber="483">river Ko</collectingCounty>
,
<geoCoordinate id="F8F9501AE8621C6FFEB1F955FE55D9DE" box="[371,496,1734,1759]" direction="north" orientation="latitude" pageId="8" pageNumber="483" precision="5" value="47.0716">47,0716°N</geoCoordinate>
,
<geoCoordinate id="F8F9501AE8621C6FFE3EF955FD21D9DE" box="[508,644,1734,1759]" direction="east" orientation="longitude" pageId="8" pageNumber="483" precision="5" value="136.4572">136,4572°E</geoCoordinate>
,
<quantity id="5A359B38E8621C6FFD4DF955FD7DD9DF" box="[655,728,1734,1759]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="8" pageNumber="483" unit="m" value="750.0">
<elevation id="16E0D1EEE8621C6FFD4DF955FD7DD9DF" box="[655,728,1734,1759]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="8" pageNumber="483" unit="m" value="750.0">750 m</elevation>
</quantity>
, fir-birch forest,
<date id="E973101DE8621C6FFC55F955FBA9D9DE" box="[919,1036,1734,1759]" pageId="8" pageNumber="483" value="2015-05-26">
<collectingDate id="F937E9F5E8621C6FFC55F955FBA9D9DE" box="[919,1036,1734,1759]" pageId="8" pageNumber="483" value="2015-05-26">26.V.2015</collectingDate>
</date>
, talus [rock slides], leg.
<collectorName id="3038530BE8621C6FFADDF954FA3DD9DF" box="[1311,1432,1735,1759]" pageId="8" pageNumber="483">A. Hansen</collectorName>
,
<collectorName id="3038530BE8621C6FFF55F978FEBAD803" box="[151,287,1771,1794]" pageId="8" pageNumber="483">M. Justesen</collectorName>
&amp;
<collectorName id="3038530BE8621C6FFE8EF978FDAAD802" box="[332,527,1770,1795]" pageId="8" pageNumber="483">A. Solodovnikov</collectorName>
(
<collectionCode id="FBDCAE18E8621C6FFDE0F978FDDCD802" box="[546,633,1771,1795]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="8" pageNumber="483">ZMUC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="2DA53C80E8621C6FFD53F978FDEDD826" ID-GBIF-Occurrence="1315044600" collectingDate="2015-05-26" collectionCode="ZMUC, ISEA" collectorName="A. Hansen &amp; M. Justesen &amp; A. Solodovnikov" country="Russia" county="river Ko" elevation="750" latitude="47.0716" location="Material" longLatPrecision="6" longitude="136.4572" municipality="Sikhote-Alin" pageId="8" pageNumber="483" specimenCount="12" specimenCount-female="10" specimenCount-male="2" typeStatus="paratype">
<typeStatus id="4276887FE8621C6FFD53F978FCA4D802" box="[657,769,1771,1795]" pageId="8" pageNumber="483" type="paratype">paratypes</typeStatus>
:
<specimenCount id="8BCBFD54E8621C6FFCCDF978FCC4D802" box="[783,865,1770,1795]" pageId="8" pageNumber="483" type="male">1 male</specimenCount>
,
<specimenCount id="8BCBFD54E8621C6FFCB3F979FC41D802" box="[881,996,1770,1795]" pageId="8" pageNumber="483" type="female">5 females</specimenCount>
, same data as in
<typeStatus id="4276887FE8621C6FFB74F979FABDD802" box="[1206,1304,1770,1795]" pageId="8" pageNumber="483" type="holotype">holotype</typeStatus>
(
<specimenCount id="8BCBFD54E8621C6FFAECF978FADBD802" box="[1326,1406,1770,1795]" pageId="8" pageNumber="483" type="male">1 male</specimenCount>
,
<specimenCount id="8BCBFD54E8621C6FFA4CF978FF4BD826" pageId="8" pageNumber="483" type="female">2 females</specimenCount>
at
<collectionCode id="FBDCAE18E8621C6FFED2F89CFEC1D826" box="[272,356,1807,1831]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="8" pageNumber="483">ZMUC</collectionCode>
,
<specimenCount id="8BCBFD54E8621C6FFEADF89CFE7ED826" box="[367,475,1806,1831]" pageId="8" pageNumber="483" type="female">3 females</specimenCount>
at
<collectionCode id="FBDCAE18E8621C6FFE3FF89CFDE5D826" box="[509,576,1807,1831]" pageId="8" pageNumber="483">ISEA</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="D5D76556E8621C6CFF05F8A1FEA3D8DD" lastPageId="11" lastPageNumber="486" pageId="8" pageNumber="483" type="description">
<paragraph id="9D7236DDE8621C6FFF05F8A1FC88D8B7" blockId="8.[151,1436,1698,1975]" pageId="8" pageNumber="483">
<emphasis id="AFB9EACFE8621C6FFF05F8A1FEFCD84A" bold="true" box="[199,345,1842,1867]" pageId="8" pageNumber="483">Description.</emphasis>
Measurements (
<specimenCount id="8BCBFD54E8621C6FFDD8F8A0FD0ED84A" box="[538,683,1842,1867]" pageId="8" pageNumber="483" type="generic">7 specimens</specimenCount>
, min-max, in mm): head length (from base of labrum to nuchal ridge) 1.2–1.4, head width (maximal) 1.3–1.4, pronotum length (along midline) 1.7–1.9, pronotum width (maximal) 1.9–2.0, elytral length (from shoulder to apical margin) 1.6–1.9, elytral width (maximal, when elytra closed along suture) 2–2.2; total body length 9.0–
<quantity id="5A359B38E8621C6FFD06F80DFC8CD8B7" box="[708,809,1950,1975]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" pageId="8" pageNumber="483" unit="mm" value="11.5">11.5 mm</quantity>
.
</paragraph>
<caption id="C9B26655E8631C6EFF55FAD2FD4ADA56" box="[151,751,1345,1367]" pageId="9" pageNumber="484">
<paragraph id="9D7236DDE8631C6EFF55FAD2FD4ADA56" blockId="9.[151,751,1345,1367]" box="[151,751,1345,1367]" pageId="9" pageNumber="484">
<emphasis id="AFB9EACFE8631C6EFF55FAD2FEB7DA57" bold="true" box="[151,274,1345,1367]" pageId="9" pageNumber="484">FIGURE 3.</emphasis>
<taxonomicName id="5ACD4D5EE8631C6EFEDBFAD2FE0DDA56" box="[281,424,1345,1367]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="484" phylum="Arthropoda" rank="species" species="roma" status="sp. nov.">
<emphasis id="AFB9EACFE8631C6EFEDBFAD2FE0DDA56" box="[281,424,1345,1367]" italics="true" pageId="9" pageNumber="484">Quedius roma</emphasis>
</taxonomicName>
<emphasis id="AFB9EACFE8631C6EFE6CFAD1FE42DA56" bold="true" box="[430,487,1346,1367]" pageId="9" pageNumber="484">
<taxonomicNameLabel id="B48A57B4E8631C6EFE6CFAD1FE42DA56" box="[430,487,1346,1367]" pageId="9" pageNumber="484" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, habitus. Scale bar 5 mm.
</paragraph>
</caption>
<paragraph id="9D7236DDE8631C6EFF05FA13FA21DABA" blockId="9.[151,1436,1406,2007]" pageId="9" pageNumber="484">
Brown to dark brown, medium large species, distinctly flattened dorso-ventrally, with very small eyes, relatively narrow head, anteriorly strongly narrowed pronotum, shortened elytra and lacking hind wings (
<figureCitation id="05F62A58E8631C6EFAF0FA31FAD0DABA" box="[1330,1397,1442,1467]" captionStart="FIGURE 3" captionStartId="9.[151,250,1345,1367]" captionText="FIGURE 3. Quedius roma sp. n., habitus. Scale bar 5 mm." pageId="9" pageNumber="484">Fig. 3</figureCitation>
).
</paragraph>
<paragraph id="9D7236DDE8631C6EFF05FA55FCDAD9DA" blockId="9.[151,1436,1406,2007]" pageId="9" pageNumber="484">
Head about as long as wide, without notable temporal angles; surface with distinct fingerprint-like microsculpture and hardly visible micropunctation; eyes small, in dorsal view hardly protruding over contour of head, tempora ca. 2.5 times as long as eyes. ‘Taxonomic’ setae on disk of head capsule represented by pair of anterior and posterior frontal setae, and two pairs of vertical setae (‘f’ and ‘v’ in
<figureCitation id="05F62A58E8631C6EFBF0F9A1FBD0D94A" box="[1074,1141,1586,1611]" captionStart="FIGURE 4" captionStartId="10.[151,250,1916,1938]" captionTargetBox="[240,1354,185,1893]" captionTargetId="figure@10.[222,1354,185,1893]" captionTargetPageId="10" captionText="FIGURE 4. Quedius roma sp. n., details of morphology: head dorsally (A); pronotum dorsally (B); male sternite VIII (C); male sternite IX (D); male tergite X (E); aedeagus, parameral side (F); aedeagus, laterally (G); paramere, side facing median lobe (H); female tergite X (I); d—dorsal puncture, f—frontal puncture, s—sublateral puncture, v—vertical puncture. Scale bars 1 mm." httpUri="https://zenodo.org/record/260654/files/figure.png" pageId="9" pageNumber="484">Fig. 4</figureCitation>
A, respectively); posterior frontal setae situated at about same distance from eye margin and nuchal ridge, or closer to the latter. Infraorbital ridges fully developed reaching base of mandibles anteriorly. Antennae moderately long; their second and third antennomeres combined distinctly longer than first; fourth and fifth antennomeres, each, distinctly longer than wide; sixth to tenth antennomeres, each, slightly longer than wide.
</paragraph>
<paragraph id="9D7236DDE8631C6EFF05F97BFCCED8D6" blockId="9.[151,1436,1406,2007]" pageId="9" pageNumber="484">
Pronotum about as wide as long, widest at about its middle with lateral sides very strongly narrowing anteriad, and weakly-posteriad; latero-anterior areas strongly deflexed downward so pronotal anterior angles not visible in dorsal view; lateral portions in posterior half of pronotal length distinctly explanate, without iridescence; basal margin nearly straight, posterior angles distinct. ‘Taxonomic’ setae on disc of pronotum consisting of two punctures in dorsal row on disk (not counting smaller puncture near anterior margin) (‘d’ in
<figureCitation id="05F62A58E8631C6EFB67F8E5FB4DD88E" box="[1189,1256,1910,1935]" captionStart="FIGURE 4" captionStartId="10.[151,250,1916,1938]" captionTargetBox="[240,1354,185,1893]" captionTargetId="figure@10.[222,1354,185,1893]" captionTargetPageId="10" captionText="FIGURE 4. Quedius roma sp. n., details of morphology: head dorsally (A); pronotum dorsally (B); male sternite VIII (C); male sternite IX (D); male tergite X (E); aedeagus, parameral side (F); aedeagus, laterally (G); paramere, side facing median lobe (H); female tergite X (I); d—dorsal puncture, f—frontal puncture, s—sublateral puncture, v—vertical puncture. Scale bars 1 mm." httpUri="https://zenodo.org/record/260654/files/figure.png" pageId="9" pageNumber="484">Fig. 4</figureCitation>
B), and one-two punctures in sublateral row (of them posterior, when present, situated before or at level of large lateral puncture) (‘s’ in
<figureCitation id="05F62A58E8631C6EFF23F82DFE80D8D6" box="[225,293,1982,2007]" captionStart="FIGURE 4" captionStartId="10.[151,250,1916,1938]" captionTargetBox="[240,1354,185,1893]" captionTargetId="figure@10.[222,1354,185,1893]" captionTargetPageId="10" captionText="FIGURE 4. Quedius roma sp. n., details of morphology: head dorsally (A); pronotum dorsally (B); male sternite VIII (C); male sternite IX (D); male tergite X (E); aedeagus, parameral side (F); aedeagus, laterally (G); paramere, side facing median lobe (H); female tergite X (I); d—dorsal puncture, f—frontal puncture, s—sublateral puncture, v—vertical puncture. Scale bars 1 mm." httpUri="https://zenodo.org/record/260654/files/figure.png" pageId="9" pageNumber="484">Fig. 4</figureCitation>
B). Micropunctation and microsclupture as on head.
</paragraph>
<caption id="C9B26655E8601C6DFF55F8EFFF77D8EF" httpUri="https://zenodo.org/record/260654/files/figure.png" pageId="10" pageNumber="485" targetBox="[240,1354,185,1893]" targetPageId="10">
<paragraph id="9D7236DDE8601C6DFF55F8EFFF77D8EF" blockId="10.[151,1436,1916,2030]" pageId="10" pageNumber="485">
<emphasis id="AFB9EACFE8601C6DFF55F8EFFEB0D890" bold="true" box="[151,277,1916,1938]" pageId="10" pageNumber="485">FIGURE 4.</emphasis>
<taxonomicName id="5ACD4D5EE8601C6DFEDFF8EFFE0AD893" box="[285,431,1916,1938]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="485" phylum="Arthropoda" rank="species" species="roma" status="sp. nov.">
<emphasis id="AFB9EACFE8601C6DFEDFF8EFFE0AD893" box="[285,431,1916,1938]" italics="true" pageId="10" pageNumber="485">Quedius roma</emphasis>
</taxonomicName>
<emphasis id="AFB9EACFE8601C6DFE7AF8EEFE56D893" bold="true" box="[440,499,1917,1938]" pageId="10" pageNumber="485">
<taxonomicNameLabel id="B48A57B4E8601C6DFE7AF8EEFE56D893" box="[440,499,1917,1938]" pageId="10" pageNumber="485" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, details of morphology: head dorsally (A); pronotum dorsally (B); male sternite VIII (C); male sternite IX (D); male tergite X (E); aedeagus, parameral side (F); aedeagus, laterally (G); paramere, side facing median lobe (H); female tergite X (I); d—dorsal puncture, f—frontal puncture, s—sublateral puncture, v—vertical puncture. Scale bars 1mm.
</paragraph>
</caption>
<paragraph id="9D7236DDE8611C6CFF05FF04FC03DFB1" blockId="11.[151,1437,151,2013]" box="[199,934,151,176]" pageId="11" pageNumber="486">Mesoscutellum impunctate, with faint transversal microsculpture.</paragraph>
<paragraph id="9D7236DDE8611C6CFF05FF2EFC74DFF9" blockId="11.[151,1437,151,2013]" pageId="11" pageNumber="486">Legs rather long, posterior distinctly longer than anterior and middle; anterior tarsi enlarged in both sexes; middle and especially posterior femora with dark iridescent inner surface.</paragraph>
<paragraph id="9D7236DDE8611C6CFF05FE97FE58DE64" blockId="11.[151,1437,151,2013]" pageId="11" pageNumber="486">Elytra about as long as pronotum, moderately densely punctate, with setiferous punctures sometimes nearly contiguous horizontally but separated by interspaces larger than puncture diameter vertically; interspaces glossy, with very slight microsculpture.</paragraph>
<paragraph id="9D7236DDE8611C6CFF05FEFCFDFDDDF1" blockId="11.[151,1437,151,2013]" pageId="11" pageNumber="486">
Abdominal tergites sparsely punctate, with distinct irregular microscuplture and slight iridescence; tergite VII without apical seam of palisade fringe.
<emphasis id="AFB9EACFE8611C6CFD9EFE07FD3ADEAC" box="[604,671,404,429]" italics="true" pageId="11" pageNumber="486">Male:</emphasis>
Medial emargination of sternite VIII almost at right angle and not rounded (
<figureCitation id="05F62A58E8611C6CFEC6FE24FEEFDED1" box="[260,330,439,464]" captionStart="FIGURE 4" captionStartId="10.[151,250,1916,1938]" captionTargetBox="[240,1354,185,1893]" captionTargetId="figure@10.[222,1354,185,1893]" captionTargetPageId="10" captionText="FIGURE 4. Quedius roma sp. n., details of morphology: head dorsally (A); pronotum dorsally (B); male sternite VIII (C); male sternite IX (D); male tergite X (E); aedeagus, parameral side (F); aedeagus, laterally (G); paramere, side facing median lobe (H); female tergite X (I); d—dorsal puncture, f—frontal puncture, s—sublateral puncture, v—vertical puncture. Scale bars 1 mm." httpUri="https://zenodo.org/record/260654/files/figure.png" pageId="11" pageNumber="486">Fig. 4</figureCitation>
C); sternite IX with asymmetrical, apically pointed glabrous basal portion, and medially sparsely setose symmetrical apical portion with faint apical emargination (
<figureCitation id="05F62A58E8611C6CFCB0FE4FFC10DEF5" box="[882,949,476,501]" captionStart="FIGURE 4" captionStartId="10.[151,250,1916,1938]" captionTargetBox="[240,1354,185,1893]" captionTargetId="figure@10.[222,1354,185,1893]" captionTargetPageId="10" captionText="FIGURE 4. Quedius roma sp. n., details of morphology: head dorsally (A); pronotum dorsally (B); male sternite VIII (C); male sternite IX (D); male tergite X (E); aedeagus, parameral side (F); aedeagus, laterally (G); paramere, side facing median lobe (H); female tergite X (I); d—dorsal puncture, f—frontal puncture, s—sublateral puncture, v—vertical puncture. Scale bars 1 mm." httpUri="https://zenodo.org/record/260654/files/figure.png" pageId="11" pageNumber="486">Fig. 4</figureCitation>
D); male tergite X apically broadly rounded (
<figureCitation id="05F62A58E8611C6CFF5DFE6CFF4CDD19" box="[159,233,511,536]" captionStart="FIGURE 4" captionStartId="10.[151,250,1916,1938]" captionTargetBox="[240,1354,185,1893]" captionTargetId="figure@10.[222,1354,185,1893]" captionTargetPageId="10" captionText="FIGURE 4. Quedius roma sp. n., details of morphology: head dorsally (A); pronotum dorsally (B); male sternite VIII (C); male sternite IX (D); male tergite X (E); aedeagus, parameral side (F); aedeagus, laterally (G); paramere, side facing median lobe (H); female tergite X (I); d—dorsal puncture, f—frontal puncture, s—sublateral puncture, v—vertical puncture. Scale bars 1 mm." httpUri="https://zenodo.org/record/260654/files/figure.png" pageId="11" pageNumber="486">Fig. 4</figureCitation>
E). Aedeagus only slightly asymmetrical; median lobe (in parameral and anteparameral view) rather parallel-sided along most of its length, with obtuse apex (
<figureCitation id="05F62A58E8611C6CFCDDFDB7FCC6DD3D" box="[799,867,548,573]" captionStart="FIGURE 4" captionStartId="10.[151,250,1916,1938]" captionTargetBox="[240,1354,185,1893]" captionTargetId="figure@10.[222,1354,185,1893]" captionTargetPageId="10" captionText="FIGURE 4. Quedius roma sp. n., details of morphology: head dorsally (A); pronotum dorsally (B); male sternite VIII (C); male sternite IX (D); male tergite X (E); aedeagus, parameral side (F); aedeagus, laterally (G); paramere, side facing median lobe (H); female tergite X (I); d—dorsal puncture, f—frontal puncture, s—sublateral puncture, v—vertical puncture. Scale bars 1 mm." httpUri="https://zenodo.org/record/260654/files/figure.png" pageId="11" pageNumber="486">Fig. 4</figureCitation>
F), with short blunt apical carina on anteparameral side more basally followed by shallow invagination (
<figureCitation id="05F62A58E8611C6CFCC0FDD4FCECDD61" box="[770,841,583,608]" captionStart="FIGURE 4" captionStartId="10.[151,250,1916,1938]" captionTargetBox="[240,1354,185,1893]" captionTargetId="figure@10.[222,1354,185,1893]" captionTargetPageId="10" captionText="FIGURE 4. Quedius roma sp. n., details of morphology: head dorsally (A); pronotum dorsally (B); male sternite VIII (C); male sternite IX (D); male tergite X (E); aedeagus, parameral side (F); aedeagus, laterally (G); paramere, side facing median lobe (H); female tergite X (I); d—dorsal puncture, f—frontal puncture, s—sublateral puncture, v—vertical puncture. Scale bars 1 mm." httpUri="https://zenodo.org/record/260654/files/figure.png" pageId="11" pageNumber="486">Fig. 4</figureCitation>
G); paramere (in parameral view) about as wide as median lobe, apically obtuse, with slight median emargination, with 8–10 sensory peg setae on its underside densely clustered in two lateral groups close to anterior margin, with two pairs of apical and two pairs of apicolateral setae (
<figureCitation id="05F62A58E8611C6CFEEFFD27FED5DDCD" box="[301,368,692,717]" captionStart="FIGURE 4" captionStartId="10.[151,250,1916,1938]" captionTargetBox="[240,1354,185,1893]" captionTargetId="figure@10.[222,1354,185,1893]" captionTargetPageId="10" captionText="FIGURE 4. Quedius roma sp. n., details of morphology: head dorsally (A); pronotum dorsally (B); male sternite VIII (C); male sternite IX (D); male tergite X (E); aedeagus, parameral side (F); aedeagus, laterally (G); paramere, side facing median lobe (H); female tergite X (I); d—dorsal puncture, f—frontal puncture, s—sublateral puncture, v—vertical puncture. Scale bars 1 mm." httpUri="https://zenodo.org/record/260654/files/figure.png" pageId="11" pageNumber="486">Fig. 4</figureCitation>
H).
<emphasis id="AFB9EACFE8611C6CFE5EFD27FE5CDDCC" box="[412,505,692,717]" italics="true" pageId="11" pageNumber="486">Female:</emphasis>
tergite X with stronger sclerotized apical lobe and soft, membranous lateral lobes, apically with sparse long setae (
<figureCitation id="05F62A58E8611C6CFE3EFD44FDE4DDF1" box="[508,577,727,752]" captionStart="FIGURE 4" captionStartId="10.[151,250,1916,1938]" captionTargetBox="[240,1354,185,1893]" captionTargetId="figure@10.[222,1354,185,1893]" captionTargetPageId="10" captionText="FIGURE 4. Quedius roma sp. n., details of morphology: head dorsally (A); pronotum dorsally (B); male sternite VIII (C); male sternite IX (D); male tergite X (E); aedeagus, parameral side (F); aedeagus, laterally (G); paramere, side facing median lobe (H); female tergite X (I); d—dorsal puncture, f—frontal puncture, s—sublateral puncture, v—vertical puncture. Scale bars 1 mm." httpUri="https://zenodo.org/record/260654/files/figure.png" pageId="11" pageNumber="486">
Fig.
<date id="E973101DE8611C6CFDF3FD4BFDE4DDF1" box="[561,577,728,752]" pageId="11" pageNumber="486">4</date>
</figureCitation>
I).
</paragraph>
<paragraph id="9D7236DDE8611C6CFF05FD6FFC06DC39" blockId="11.[151,1437,151,2013]" pageId="11" pageNumber="486">
<emphasis id="AFB9EACFE8611C6CFF05FD6FFE03DC14" box="[199,422,764,789]" italics="true" pageId="11" pageNumber="486">Sexual dimorphism</emphasis>
. Without dissection of genitalia males can be separated from females by medially emarginated sternite VIII (not emarginated, concave in females) only.
</paragraph>
<paragraph id="9D7236DDE8611C6CFF05FCD7FD0BDCA5" blockId="11.[151,1437,151,2013]" pageId="11" pageNumber="486">
<emphasis id="AFB9EACFE8611C6CFF05FCD7FE9EDC5C" box="[199,315,836,861]" italics="true" pageId="11" pageNumber="486">Variability</emphasis>
. The species slightly varies in size, and distinctly in the body coloration: from darker, nearly blackish to paler, brown specimens. Basal setiferous pores in dorsal and sublateral rows of pronotum may be absent in some specimens, sometimes only in one row.
</paragraph>
<paragraph id="9D7236DDE8611C6CFF05FC3CFC02D950" blockId="11.[151,1437,151,2013]" pageId="11" pageNumber="486">
<emphasis id="AFB9EACFE8611C6CFF05FC3CFD35DCC9" bold="true" box="[199,656,943,968]" pageId="11" pageNumber="486">Systematic position and comparison.</emphasis>
Based on two vertical setae in each pair on head, and laterally explanate pronotum, the new species can be safely placed to the subgenus
<taxonomicName id="5ACD4D5EE8611C6CFC37FC46FB26DCED" box="[1013,1155,981,1004]" class="Insecta" family="Staphylinidae" genus="Microsaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="486" phylum="Arthropoda" rank="genus">
<emphasis id="AFB9EACFE8611C6CFC37FC46FB26DCED" box="[1013,1155,981,1004]" italics="true" pageId="11" pageNumber="486">Microsaurus</emphasis>
</taxonomicName>
. Within the Russian Far East fauna,
<taxonomicName id="5ACD4D5EE8611C6CFEDFFC6BFEDFDB11" box="[285,378,1016,1040]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="486" phylum="Arthropoda" rank="species" species="roma">
<emphasis id="AFB9EACFE8611C6CFEDFFC6BFEDFDB11" box="[285,378,1016,1040]" italics="true" pageId="11" pageNumber="486">Q. roma</emphasis>
</taxonomicName>
is the only
<taxonomicName id="5ACD4D5EE8611C6CFDC0FC6AFD35DB11" box="[514,656,1017,1040]" class="Insecta" family="Staphylinidae" genus="Microsaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="486" phylum="Arthropoda" rank="genus">
<emphasis id="AFB9EACFE8611C6CFDC0FC6AFD35DB11" box="[514,656,1017,1040]" italics="true" pageId="11" pageNumber="486">Microsaurus</emphasis>
</taxonomicName>
species with pronounced morphological modifications for hypogean biology and therefore it is very distinct. Based on glabrous, non-punctate scutellum, taxonomic chaetotaxy of head and pronotum, and characteristic structure of the slightly asymmetrical aedeagus (obtuse median lobe having blunt short median carina followed by an invagination; broad, apically only slightly incised paramere with 8–10 sensory peg setae clustered in two groups near its apex), the new species is affiliated with the core of the
<taxonomicName id="5ACD4D5EE8611C6CFB10FB14FAF7DBA1" box="[1234,1362,1159,1184]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="486" phylum="Arthropoda" rank="species" species="przewalskii">
<emphasis id="AFB9EACFE8611C6CFB10FB14FAF7DBA1" box="[1234,1362,1159,1184]" italics="true" pageId="11" pageNumber="486">przewalskii</emphasis>
</taxonomicName>
-group (as defined in
<bibRefCitation id="F95C4B2CE8611C6CFEFAFB3FFE79DBC5" author="Smetana" box="[312,476,1196,1220]" pageId="11" pageNumber="486" refString="Smetana, A. (2001) Contributions to the knowledge of the Quediina (Coleoptera, Staphylinidae, Staphylinini) of China. Part 20. Genus Quedius Stephens, 1829. Subgenus Microsaurus Dejean, 1833. Section 12. Elytra (Tokyo), 29 (1), 193 - 216." type="journal article" year="2001">Smetana 2001</bibRefCitation>
). However, relatively more narrow head, more strongly deflexed downwards anterior angles of pronotum, more sparse punctation of abdomen (mostly dense in
<taxonomicName id="5ACD4D5EE8611C6CFC2EFB5CFBC9DBE9" box="[1004,1132,1231,1256]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="486" phylum="Arthropoda" rank="species" species="przewalskii">
<emphasis id="AFB9EACFE8611C6CFC2EFB5CFBC9DBE9" box="[1004,1132,1231,1256]" italics="true" pageId="11" pageNumber="486">przewalskii</emphasis>
</taxonomicName>
-group) and more elongate subapical segments of antennae easily distinguish
<taxonomicName id="5ACD4D5EE8611C6CFD0BFB66FC80DA0D" box="[713,805,1269,1292]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="486" phylum="Arthropoda" rank="species" species="roma">
<emphasis id="AFB9EACFE8611C6CFD0BFB66FC80DA0D" box="[713,805,1269,1292]" italics="true" pageId="11" pageNumber="486">Q. roma</emphasis>
</taxonomicName>
from any other species of the
<taxonomicName id="5ACD4D5EE8611C6CFBBBFB67FB5CDA0C" box="[1145,1273,1268,1293]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="486" phylum="Arthropoda" rank="species" species="przewalskii">
<emphasis id="AFB9EACFE8611C6CFBBBFB67FB5CDA0C" box="[1145,1273,1268,1293]" italics="true" pageId="11" pageNumber="486">przewalskii</emphasis>
</taxonomicName>
-group even in its broadest sense (as in
<bibRefCitation id="F95C4B2CE8611C6CFE79FA8BFDC7DA31" author="Smetana" box="[443,610,1304,1328]" pageId="11" pageNumber="486" refString="Smetana, A. (2014) Contributions to the knowledge of the Quediina (Coleoptera, Staphylinidae, Staphylinini) of China. Part 44. Genus Quedius Stephens, 1829. Subgenus Microsaurus Dejean, 1833. Section 22. Folia Heyrovskyana, series A, 21 (1 - 4), 1 - 39." type="journal article" year="2014">Smetana 2014</bibRefCitation>
, 2015). Although the aedeagus of the new species looks similar to
<taxonomicName id="5ACD4D5EE8611C6CFA43FA8BFEB3DA54" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="486" phylum="Arthropoda" rank="species" species="przewalskii">
<emphasis id="AFB9EACFE8611C6CFA43FA8BFEB3DA54" italics="true" pageId="11" pageNumber="486">Q. przewalskii</emphasis>
</taxonomicName>
and similar species, its paramere is relatively broader and entirely covers the contours of the median lobe in parameral view. Also the new species has lesser number of sensory peg setae that are situated distinctly denser and more apically on the parameral underside. Even though in habitus and chaetotaxy of head and pronotum
<taxonomicName id="5ACD4D5EE8611C6CFF55FA3BFF51DAC1" box="[151,244,1448,1472]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="486" phylum="Arthropoda" rank="species" species="roma">
<emphasis id="AFB9EACFE8611C6CFF55FA3BFF51DAC1" box="[151,244,1448,1472]" italics="true" pageId="11" pageNumber="486">Q. roma</emphasis>
</taxonomicName>
resembles
<taxonomicName id="5ACD4D5EE8611C6CFEB6FA3BFE5ADAC1" box="[372,511,1447,1472]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="486" phylum="Arthropoda" rank="species" species="mutilatus">
<emphasis id="AFB9EACFE8611C6CFEB6FA3BFE5ADAC1" box="[372,511,1447,1472]" italics="true" pageId="11" pageNumber="486">Q. mutilatus</emphasis>
</taxonomicName>
and
<taxonomicName id="5ACD4D5EE8611C6CFDFAFA3BFD04DAC1" box="[568,673,1447,1472]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="486" phylum="Arthropoda" rank="species" species="kalabi">
<emphasis id="AFB9EACFE8611C6CFDFAFA3BFD04DAC1" box="[568,673,1447,1472]" italics="true" pageId="11" pageNumber="486">Q. kalabi</emphasis>
</taxonomicName>
, it differs from them in glabrous scutellum, relatively more narrow head, and more anterior position of posterior frontal puncture (in
<taxonomicName id="5ACD4D5EE8611C6CFC5BFA5EFB82DAE4" box="[921,1063,1484,1509]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="486" phylum="Arthropoda" rank="species" species="mutilatus">
<emphasis id="AFB9EACFE8611C6CFC5BFA5EFB82DAE4" box="[921,1063,1484,1509]" italics="true" pageId="11" pageNumber="486">Q. mutilatus</emphasis>
</taxonomicName>
and
<taxonomicName id="5ACD4D5EE8611C6CFBA4FA5EFB77DAE4" box="[1126,1234,1484,1509]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="486" phylum="Arthropoda" rank="species" species="kalabi">
<emphasis id="AFB9EACFE8611C6CFBA4FA5EFB77DAE4" box="[1126,1234,1484,1509]" italics="true" pageId="11" pageNumber="486">Q. kalabi</emphasis>
</taxonomicName>
located closer to nuchal ridge than to postero-median margin of eye). Compared to the former species,
<taxonomicName id="5ACD4D5EE8611C6CFBAAFA63FB63D909" box="[1128,1222,1520,1544]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="486" phylum="Arthropoda" rank="species" species="roma">
<emphasis id="AFB9EACFE8611C6CFBAAFA63FB63D909" box="[1128,1222,1520,1544]" italics="true" pageId="11" pageNumber="486">Q. roma</emphasis>
</taxonomicName>
has quite different structure of the aedeagus.
<taxonomicName id="5ACD4D5EE8611C6CFE09F987FDC8D92D" box="[459,621,1556,1581]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="486" phylum="Arthropoda" rank="species" species="roma">
<emphasis id="AFB9EACFE8611C6CFE09F987FDC8D92D" box="[459,621,1556,1581]" italics="true" pageId="11" pageNumber="486">Quedius roma</emphasis>
</taxonomicName>
differs from
<taxonomicName id="5ACD4D5EE8611C6CFCCEF986FC2FD92D" box="[780,906,1556,1580]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="486" phylum="Arthropoda" rank="species" species="fonteius">
<emphasis id="AFB9EACFE8611C6CFCCEF986FC2FD92D" box="[780,906,1556,1580]" italics="true" pageId="11" pageNumber="486">Q. fonteius</emphasis>
</taxonomicName>
in habitus and in lack of additional puncture between posterior frontal puncture and postero-median margin of eye.
</paragraph>
<paragraph id="9D7236DDE8611C6CFF05F9CFFEA3D8DD" blockId="11.[151,1437,151,2013]" pageId="11" pageNumber="486">
Unlike hypogean
<taxonomicName id="5ACD4D5EE8611C6CFE4DF9CEFDB8D975" box="[399,541,1629,1652]" class="Insecta" family="Staphylinidae" genus="Microsaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="486" phylum="Arthropoda" rank="genus">
<emphasis id="AFB9EACFE8611C6CFE4DF9CEFDB8D975" box="[399,541,1629,1652]" italics="true" pageId="11" pageNumber="486">Microsaurus</emphasis>
</taxonomicName>
from
<collectingCountry id="E5DA764DE8611C6CFDA0F9CEFD07D975" box="[610,674,1629,1652]" name="Japan" pageId="11" pageNumber="486">Japan</collectingCountry>
or
<collectingCountry id="E5DA764DE8611C6CFD0AF9CFFCBFD974" box="[712,794,1628,1653]" name="Taiwan" pageId="11" pageNumber="486">Taiwan</collectingCountry>
(referred as members of
<emphasis id="AFB9EACFE8611C6CFBF5F9CFFB13D974" box="[1079,1206,1628,1653]" italics="true" pageId="11" pageNumber="486">abnormalis</emphasis>
-group)
<taxonomicName id="5ACD4D5EE8611C6CFAD0F9CEFACAD975" box="[1298,1391,1629,1652]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="486" phylum="Arthropoda" rank="species" species="roma">
<emphasis id="AFB9EACFE8611C6CFAD0F9CEFACAD975" box="[1298,1391,1629,1652]" italics="true" pageId="11" pageNumber="486">Q. roma</emphasis>
</taxonomicName>
has glabrous non-punctate scutellum and different
<typeStatus id="4276887FE8611C6CFD6BF912FD7CD998" box="[681,729,1665,1689]" pageId="11" pageNumber="486">type</typeStatus>
of the aedeagus (in the
<emphasis id="AFB9EACFE8611C6CFC2CF9ECFBC8D999" box="[1006,1133,1663,1688]" italics="true" pageId="11" pageNumber="486">abnormalis</emphasis>
-group aedeagus is usually symmetrical; with apically broad deeply bilobed paramere having numerous sensory peg setae; and with two carinae near apex on the anteparameral side). From the Chinese members of
<emphasis id="AFB9EACFE8611C6CFBDAF954FB04D9E1" box="[1048,1185,1735,1760]" italics="true" pageId="11" pageNumber="486">kiangsiensis</emphasis>
- and
<emphasis id="AFB9EACFE8611C6CFB25F95AFAE3D9E1" box="[1255,1350,1737,1760]" italics="true" pageId="11" pageNumber="486">mnemon</emphasis>
-groups collected in caves,
<taxonomicName id="5ACD4D5EE8611C6CFEA8F97EFE62D805" box="[362,455,1773,1796]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="486" phylum="Arthropoda" rank="species" species="roma">
<emphasis id="AFB9EACFE8611C6CFEA8F97EFE62D805" box="[362,455,1773,1796]" italics="true" pageId="11" pageNumber="486">Q. roma</emphasis>
</taxonomicName>
differs in its brachypterous habitus (shorter elytra, lacking palisade fringe on abdominal tergite VII), brownish coloration, and in very different shape of the aedeagus. From the Turkish hypogean endemic
<emphasis id="AFB9EACFE8611C6CFF55F8A6FE3CD84D" box="[151,409,1844,1869]" italics="true" pageId="11" pageNumber="486">
<taxonomicName id="5ACD4D5EE8611C6CFF55F8A6FE94D84C" box="[151,305,1844,1869]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="486" phylum="Arthropoda" rank="species" species="weiratheri">Q. weiratheri</taxonomicName>
,
<taxonomicName id="5ACD4D5EE8611C6CFEFFF8A6FE3CD84D" box="[317,409,1845,1868]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="486" phylum="Arthropoda" rank="species" species="roma">Q. roma</taxonomicName>
</emphasis>
differs both in habitus (in particular more narrow head with temples not diverging towards more distinct posterior angles) and very different shape of the aedeagus. Likewise,
<taxonomicName id="5ACD4D5EE8611C6CFBF4F8CBFB37D871" box="[1078,1170,1880,1904]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="486" phylum="Arthropoda" rank="species" species="roma">
<emphasis id="AFB9EACFE8611C6CFBF4F8CBFB37D871" box="[1078,1170,1880,1904]" italics="true" pageId="11" pageNumber="486">Q. roma</emphasis>
</taxonomicName>
appears very distinct in comparison with the North American
<taxonomicName id="5ACD4D5EE8611C6CFD8BF8EEFD6CD895" box="[585,713,1916,1940]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="486" phylum="Arthropoda" rank="species" species="infernus">
<emphasis id="AFB9EACFE8611C6CFD8BF8EEFD6CD895" box="[585,713,1916,1940]" italics="true" pageId="11" pageNumber="486">Q. infernus</emphasis>
</taxonomicName>
, in particular in its clearly brachypterous habitus and aedeagus having median lobe with differently shaped apex, relatively broader paramere, and other arrangement of sensory peg setae.
</paragraph>
</subSubSection>
<subSubSection id="D5D76556E8661C6BFF05FF04FC8CDED1" pageId="12" pageNumber="487" type="distribution">
<paragraph id="9D7236DDE8661C6BFF05FF04FC8CDED1" blockId="12.[151,1437,151,573]" pageId="12" pageNumber="487">
<emphasis id="AFB9EACFE8661C6BFF05FF04FE5DDFB1" bold="true" box="[199,504,151,176]" pageId="12" pageNumber="487">Distribution and ecology.</emphasis>
The new species is known only from the
<typeStatus id="4276887FE8661C6BFC23FF0BFBB4DFB1" box="[993,1041,152,176]" pageId="12" pageNumber="487">type</typeStatus>
locality at Mt. Ko. All specimens were hand collected at one talus field (
<figureCitation id="05F62A58E8661C6BFD85FF2FFD2CDFD5" box="[583,649,188,213]" captionStart="FIGURE 1" captionStartId="1.[151,250,1831,1853]" captionTargetBox="[151,1436,844,1807]" captionTargetId="figure@1.[151,1436,844,1808]" captionTargetPageId="1" captionText="FIGURE 1. Talus-associated habitat of Quedius roma sp. n. at the type locality at Mt. Ko." httpUri="https://zenodo.org/record/260652/files/figure.png" pageId="12" pageNumber="487">Fig. 1</figureCitation>
). When such talus is formed on a slope, continuous erosion transports smaller rocks and debris to its lower levels, filling its internal cavities. Our specimens were collected under smaller rocks of the upper levels of the talus that we could access and manually lift. We were collecting there from evening dusk to about midnight with spotlights.
<taxonomicName id="5ACD4D5EE8661C6BFD9BFEB4FD13DE41" box="[601,694,295,320]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="487" phylum="Arthropoda" rank="genus">
<emphasis id="AFB9EACFE8661C6BFD9BFEB4FD13DE41" box="[601,694,295,320]" italics="true" pageId="12" pageNumber="487">Quedius</emphasis>
</taxonomicName>
were easily spotted in the layer of wet humus accumulated under those rocks by their shiny appearance. Unlike various carabids, weevils, spiders and opilionids that were actively crawling on the boulders in these night hours, all
<taxonomicName id="5ACD4D5EE8661C6BFD0BFEE3FC83DE89" box="[713,806,368,392]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="487" phylum="Arthropoda" rank="species" species="roma">
<emphasis id="AFB9EACFE8661C6BFD0BFEE3FC83DE89" box="[713,806,368,392]" italics="true" pageId="12" pageNumber="487">Q. roma</emphasis>
</taxonomicName>
specimens were collected from the humus under rocks. Internal crevices between rocks were often filled with ice and snow that was still abundant and formed large patches in the shadowy spots in the forest at this elevation.
</paragraph>
</subSubSection>
<subSubSection id="D5D76556E8661C6BFF05FE4FFAC2DD3C" pageId="12" pageNumber="487" type="etymology">
<paragraph id="9D7236DDE8661C6BFF05FE4FFAC2DD3C" blockId="12.[151,1437,151,573]" pageId="12" pageNumber="487">
<emphasis id="AFB9EACFE8661C6BFF05FE4FFEEBDEF4" bold="true" box="[199,334,476,501]" pageId="12" pageNumber="487">Etymology.</emphasis>
The new species is dedicated to our friend and colleague Roman (
<taxonomicName id="5ACD4D5EE8661C6BFB93FE4FFB32DEF5" box="[1105,1175,476,500]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="487" phylum="Arthropoda" rank="species" species="roma">Roma</taxonomicName>
) Dudko who collected several specimens that formed the
<typeStatus id="4276887FE8661C6BFDF4FD93FDC3DD19" box="[566,614,512,536]" pageId="12" pageNumber="487">type</typeStatus>
series, and introduced us to talus, a very interesting and still poorly explored habitat for diverse beetles in Siberia and Russian Far East. The species name is a noun in apposition.
</paragraph>
</subSubSection>
</treatment>
</document>
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