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<document id="30210B6F289F74B5449E7A2966E52F36" ID-DOI="10.1080/00222933.2012.759285" ID-ISSN="1464-5262" ID-Zenodo-Dep="5197572" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628882145230" checkinUser="felipe" docAuthor="Park, Kyu-Tek, Bae, Yang-Seop, Kim, Sora &amp; Heppner, John B." docDate="2013" docId="BD468B24D061303A54984EB6FC25FF20" docLanguage="en" docName="JNatHist.47.35-36.2289-2304.pdf" docOrigin="Journal of Natural History 47 (35 - 36)" docSource="http://dx.doi.org/10.1080/00222933.2012.759285" docStyle="DocumentStyle:210803DF447D5DCB84349979EDD3507F.6:JNatHist.2013-2015.journal_article.2cover" docStyleId="210803DF447D5DCB84349979EDD3507F" docStyleName="JNatHist.2013-2015.journal_article.2cover" docStyleVersion="6" docTitle="Torodora occidentalis Park &amp; Bae &amp; Kim &amp; Heppner 2013, sp. nov." docType="treatment" docVersion="4" lastPageNumber="2300" masterDocId="417FF35CD06D303755764E11FFFEFFC7" masterDocTitle="Genus Torodora Meyrick in Cambodia (Lepidoptera: Lecithoceridae: Torodorinae), with descriptions of three new species" masterLastPageNumber="2304" masterPageNumber="2289" pageNumber="2299" updateTime="1699054287737" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="155EEE9EF95E8D2546E771D475556F31">Genus Torodora Meyrick in Cambodia (Lepidoptera: Lecithoceridae: Torodorinae), with descriptions of three new species</mods:title>
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<mods:namePart id="1D3EC04101ED142C113129190934953C">Park, Kyu-Tek</mods:namePart>
<mods:affiliation id="A8DDA14F502401F4BBE43EED490E5BC9">The Korean Academy of Science and Technology, Seungnam, Korea; &amp; McGuire Center for Lepidoptera and Biodiversity, Florida Museum of Natural History, University of Florida, Gainesville, FL, USA</mods:affiliation>
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<mods:namePart id="49C65FBEE1121F974904A9619838C1DB">Bae, Yang-Seop</mods:namePart>
<mods:affiliation id="33CB52997A6F81D2430AF68335259BAF">Division of Life Sciences, College of Natural Sciences, University of Incheon, Incheon, Korea;</mods:affiliation>
</mods:name>
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<mods:namePart id="BE4BB7D9166542DA43D7258C14A82079">Kim, Sora</mods:namePart>
<mods:affiliation id="56C4DE88DE9A6683BE6C10B58D6A0022">Division of Entomology, School of Agricultural Biotechnology, Research Institute for Agriculture and Life Sciences, Seoul National University, Seoul, Korea;</mods:affiliation>
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<mods:namePart id="998B861C83E6CF5E4CE1ED902390DA26">Heppner, John B.</mods:namePart>
<mods:affiliation id="5AC5C10AF547750DE22FAEF9648F80F4">McGuire Center for Lepidoptera and Biodiversity, Florida Museum of Natural History, University of Florida, Gainesville, FL, USA</mods:affiliation>
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<mods:title id="7CA2712E0B1CCBD608A364D258949C0E">Journal of Natural History</mods:title>
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<mods:part id="3005D77EDD9BEE82873ADB00EC8137A5">
<mods:date id="8723D6FD3B2A6549A9FF1A3BEF442179">2013</mods:date>
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<mods:number id="F661F010F6887F1AFB63F8F3C6824667">2013-08-06</mods:number>
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<mods:number id="1AC6C0291EAC2CBA8E40868ED15C4BDE">47</mods:number>
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<mods:number id="9EB5E8C8ED2DF822CA5F539E9614C48E">35 - 36</mods:number>
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<mods:start id="BF932996AB4AB1F38D209EAFE3653C67">2289</mods:start>
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<treatment id="BD468B24D061303A54984EB6FC25FF20" ID-DOI="http://doi.org/10.5281/zenodo.5197566" ID-Zenodo-Dep="5197566" LSID="urn:lsid:zoobank.org:act:BC81BF8B-A61E-414B-A157-A09305192A85" httpUri="http://treatment.plazi.org/id/BD468B24D061303A54984EB6FC25FF20" lastPageId="13" lastPageNumber="2300" pageId="12" pageNumber="2299">
<subSubSection id="7DF569B9D061303B54984EB6FC87FF78" box="[494,889,167,191]" pageId="12" pageNumber="2299" type="nomenclature">
<paragraph id="35503A32D061303B54984EB6FC87FF78" blockId="12.[494,889,167,223]" box="[494,889,167,191]" pageId="12" pageNumber="2299">
<heading id="6E188D5ED061303B54984EB6FC87FF78" box="[494,889,167,191]" centered="true" fontSize="10" level="2" pageId="12" pageNumber="2299" reason="2">
<taxonomicName id="F2EF41B1D061303B54984EB6FCE7FF78" ID-CoL="9FNT5" authority="Park" authorityName="Park &amp; Bae &amp; Kim &amp; Heppner" authorityYear="2013" box="[494,793,167,191]" class="Insecta" family="Lecithoceridae" genus="Torodora" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="2299" phylum="Arthropoda" rank="species" species="occidentalis" status="sp. nov.">
<emphasis id="079BE620D061303B54984EB6FD25FF78" bold="true" box="[494,731,167,191]" italics="true" pageId="12" pageNumber="2299">Torodora occidentalis</emphasis>
Park
</taxonomicName>
,
<taxonomicNameLabel id="1CA85B5BD061303B56534EB9FC87FF78" box="[805,889,168,191]" pageId="12" pageNumber="2299" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="7DF569B9D061303B574D4ED6FCD5FF18" box="[571,811,199,223]" pageId="12" pageNumber="2299" type="description">
<paragraph id="35503A32D061303B574D4ED6FCD5FF18" blockId="12.[494,889,167,223]" box="[571,811,199,223]" pageId="12" pageNumber="2299">
(
<figureCitation id="ADD426B7D061303B57354ED6FD22FF18" box="[579,732,199,223]" captionStart="Figure 2" captionStartId="6.[211,278,1409,1431]" captionTargetBox="[213,1172,167,1376]" captionTargetId="figure-5@6.[212,1172,167,1376]" captionTargetPageId="6" captionText="Figure 2. Adults of Torodora spp. (AC) Torodora parotidosa (Wu); (B) head part in dorsal view; (C) labial palpus; (DF) T. osamensis Park, sp. nov.; (E) head part in dorsal view; (F) labial palpus; (GI) T. occidentalis Park, sp. nov.; (H) head part in dorsal view; (I) labial palpus; (JL) T. aritai Park, sp. nov.; (K) head part in dorsal view; (L) labial palpus." figureDoi="http://doi.org/10.5281/zenodo.5197576" httpUri="https://zenodo.org/record/5197576/files/figure.png" pageId="12" pageNumber="2299">Figures 2GI</figureCitation>
;
<figureCitation id="ADD426B7D061303B579E4ED6FCDDFF18" box="[744,803,199,223]" captionStart="Figure 3" captionStartId="8.[211,278,1613,1635]" captionTargetBox="[213,1172,165,1580]" captionTargetId="figure-5@8.[212,1172,162,1580]" captionTargetPageId="8" captionText="Figure 3. Male genitalia of Torodora spp. (AC) Torodora parotidosa (Wu); (B) aedeagus; (C) abdominal segments; (DF) T. osamensis Park, sp. nov.; (E) aedeagus; (F) abdominal segments; (GI) T. occidentalis Park, sp. nov.; (H) aedeagus; (I) abdominal segments; (JL) T. aritai Park, sp. nov.; (K) aedeagus; (L) abdominal segments." figureDoi="http://doi.org/10.5281/zenodo.5197580" httpUri="https://zenodo.org/record/5197580/files/figure.png" pageId="12" pageNumber="2299">3GI</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="7DF569B9D061303B55A54F3FFE14FE4A" pageId="12" pageNumber="2299" type="materials_examined">
<paragraph id="35503A32D061303B55A54F3FFEF7FE82" blockId="12.[211,1172,302,397]" box="[211,265,302,325]" pageId="12" pageNumber="2299">
<heading id="6E188D5ED061303B55A54F3FFEF7FE82" box="[211,265,302,325]" fontSize="10" level="3" pageId="12" pageNumber="2299" reason="5">
<emphasis id="079BE620D061303B55A54F3FFEF7FE82" box="[211,265,302,325]" italics="true" pageId="12" pageNumber="2299">Type</emphasis>
</heading>
</paragraph>
<paragraph id="35503A32D061303B55A54F44FE14FE4A" blockId="12.[211,1172,302,397]" pageId="12" pageNumber="2299">
<materialsCitation id="8587306FD061303B55A54F44FE14FE4A" collectingDate="2010-05-03" country="Cambodia" location="Pursat" pageId="12" pageNumber="2299" specimenCount="1" specimenCount-male="1" stateProvince="Pursat" typeStatus="holotype">
<emphasis id="079BE620D061303B55A54F44FEC4FEAA" box="[211,314,341,365]" italics="true" pageId="12" pageNumber="2299">
<typeStatus id="EA548490D061303B55A54F44FEC4FEAA" box="[211,314,341,365]" pageId="12" pageNumber="2299" type="holotype">Holotype</typeStatus>
</emphasis>
.
<specimenCount id="23E9F1BBD061303B543C4F44FEA1FEAA" box="[330,351,341,365]" pageId="12" pageNumber="2299" type="male"></specimenCount>
,
<collectingCountry id="4DF87AA2D061303B54194F44FE17FEAA" box="[367,489,341,365]" name="Cambodia" pageId="12" pageNumber="2299">Cambodia</collectingCountry>
, Prov.
<collectingRegion id="F72BF4D0D061303B57414F47FD7EFEAA" box="[567,640,342,365]" country="Cambodia" name="Pursat" pageId="12" pageNumber="2299">Pursat</collectingRegion>
(Osam),
<date id="41511CF2D061303B579D4F44FC8CFEAA" box="[747,882,341,366]" pageId="12" pageNumber="2299" value="2010-05-03">
<collectingDate id="5115E51AD061303B579D4F44FC8CFEAA" box="[747,882,341,366]" pageId="12" pageNumber="2299" value="2010-05-03">3 May 2010</collectingDate>
</date>
(Bae, Chae and Vi), gen. slide no. CIS-6066
<emphasis id="079BE620D061303B54D74F64FE53FE4A" bold="true" box="[417,429,373,397]" italics="true" pageId="12" pageNumber="2299">/</emphasis>
Park.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="7DF569B9D061303B55A54FCAFC2FFD7E" pageId="12" pageNumber="2299" type="diagnosis">
<paragraph id="35503A32D061303B55A54FCAFEC1FE35" blockId="12.[211,1172,475,697]" box="[211,319,475,498]" pageId="12" pageNumber="2299">
<heading id="6E188D5ED061303B55A54FCAFEC1FE35" box="[211,319,475,498]" fontSize="10" level="3" pageId="12" pageNumber="2299" reason="5">
<emphasis id="079BE620D061303B55A54FCAFEC1FE35" box="[211,319,475,498]" italics="true" pageId="12" pageNumber="2299">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="35503A32D061303B55A54C13FC2FFD7E" blockId="12.[211,1172,475,697]" pageId="12" pageNumber="2299">
This species is superficially similar to
<taxonomicName id="F2EF41B1D061303B570E4C12FCEEFDDD" authority="Park" authorityName="Park" authorityYear="2002" box="[632,784,514,538]" class="Insecta" family="Lecithoceridae" genus="Torodora" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="2299" phylum="Arthropoda" rank="species" species="aritai">
<emphasis id="079BE620D061303B570E4C12FD2FFDDD" box="[632,721,515,538]" italics="true" pageId="12" pageNumber="2299">T. aritai</emphasis>
Park
</taxonomicName>
, but the basal part of the forewing is darker than that of the latter, and the distance between R
<subScript id="A96B3877D061303B561F4C3AFC8CFDFB" attach="left" box="[873,882,555,572]" fontSize="7" pageId="12" pageNumber="2299">1</subScript>
and R
<subScript id="A96B3877D061303B56C94C3AFC36FDFB" attach="left" box="[959,968,555,572]" fontSize="7" pageId="12" pageNumber="2299">2</subScript>
is about 1.5 times that between R
<subScript id="A96B3877D061303B540B4C5AFE78FD9B" attach="left" box="[381,390,587,604]" fontSize="7" pageId="12" pageNumber="2299">2</subScript>
and R
<subScript id="A96B3877D061303B54A54C5AFE22FD9B" attach="left" box="[467,476,587,604]" fontSize="7" pageId="12" pageNumber="2299">3</subScript>
, whereas in
<taxonomicName id="F2EF41B1D061303B57104C52FD41FD9D" authorityName="Park" authorityYear="2002" box="[614,703,579,602]" class="Insecta" family="Lecithoceridae" genus="Torodora" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="2299" phylum="Arthropoda" rank="species" species="aritai">
<emphasis id="079BE620D061303B57104C52FD41FD9D" box="[614,703,579,602]" italics="true" pageId="12" pageNumber="2299">T. aritai</emphasis>
</taxonomicName>
, the distance of R
<subScript id="A96B3877D061303B56FC4C5AFC6DFD9B" attach="left" box="[906,915,587,604]" fontSize="7" pageId="12" pageNumber="2299">1</subScript>
R
<subScript id="A96B3877D061303B56C14C5AFC3EFD9B" attach="left" box="[951,960,587,604]" fontSize="7" pageId="12" pageNumber="2299">2</subScript>
is about twice that of R
<subScript id="A96B3877D061303B547E4C7AFEEFFDBB" attach="left" box="[264,273,619,636]" fontSize="7" pageId="12" pageNumber="2299">2</subScript>
R
<subScript id="A96B3877D061303B54424C7AFEC3FDBB" attach="left" box="[308,317,619,636]" fontSize="7" pageId="12" pageNumber="2299">3</subScript>
, It can be easily distinguished by the male genital structure as follows: the valva more elongate, without conic particles on ventral surface, with a crescent sclerite separating basal part and cucullus, and the aedeagus nearly straight.
</paragraph>
</subSubSection>
<subSubSection id="7DF569B9D061303B55A54D19FBF4F9CB" pageId="12" pageNumber="2299" type="description">
<paragraph id="35503A32D061303B55A54D19FEACFCD8" blockId="12.[211,1172,776,1318]" box="[211,338,776,799]" pageId="12" pageNumber="2299">
<heading id="6E188D5ED061303B55A54D19FEACFCD8" box="[211,338,776,799]" fontSize="10" level="3" pageId="12" pageNumber="2299" reason="5">
<emphasis id="079BE620D061303B55A54D19FEACFCD8" box="[211,338,776,799]" italics="true" pageId="12" pageNumber="2299">Description</emphasis>
</heading>
</paragraph>
<paragraph id="35503A32D061303B55A54D3EFB6AFAE2" blockId="12.[211,1172,776,1318]" pageId="12" pageNumber="2299">
<emphasis id="079BE620D061303B55A54D3EFEEFFC80" box="[211,273,815,839]" italics="true" pageId="12" pageNumber="2299">Adult</emphasis>
(
<figureCitation id="ADD426B7D061303B54544D3EFE4EFC80" box="[290,432,815,839]" captionStart="Figure 2" captionStartId="6.[211,278,1409,1431]" captionTargetBox="[213,1172,167,1376]" captionTargetId="figure-5@6.[212,1172,167,1376]" captionTargetPageId="6" captionText="Figure 2. Adults of Torodora spp. (AC) Torodora parotidosa (Wu); (B) head part in dorsal view; (C) labial palpus; (DF) T. osamensis Park, sp. nov.; (E) head part in dorsal view; (F) labial palpus; (GI) T. occidentalis Park, sp. nov.; (H) head part in dorsal view; (I) labial palpus; (JL) T. aritai Park, sp. nov.; (K) head part in dorsal view; (L) labial palpus." figureDoi="http://doi.org/10.5281/zenodo.5197576" httpUri="https://zenodo.org/record/5197576/files/figure.png" pageId="12" pageNumber="2299">Figure 2GI</figureCitation>
). Wingspan
<quantity id="F21797D7D061303B57354D3EFD6BFC80" box="[579,661,815,839]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="12" pageNumber="2299" unit="mm" value="17.0">17 mm</quantity>
. Head, tegula, and thorax dark fuscous dorsally. Basal segment of antenna orange white, densely speckled with dark fuscous scales dorsally; flagellum orange white, with distinct brownish annulations beyond four-fifths of length. Second segment of labial palpus thickened, dark brown in basal two-thirds length on outer surface with orange-white apex, orange white on inner surface; third segment slender, as long as second segment, dark brown on ventral surface. Forewing evenly covered with dark brown scales throughout, with blackish reniform stigma at end of cell; costa gently arched before quarter of length, then nearly straight medially, slightly arched beyond three-quarters of length, with a small triangular orange-white costal patch at three-quarters of length; apex obtuse; termen oblique, sinuate, with blackish scales along margin; fringe the same colour throughout. Venation with R
<subScript id="A96B3877D061303B51FF4A66FB6CFB4F" attach="left" box="[1161,1170,1143,1160]" fontSize="7" pageId="12" pageNumber="2299">1</subScript>
arising before middle of cell; R
<subScript id="A96B3877D061303B57594A86FDC6FB6F" attach="left" box="[559,568,1175,1192]" fontSize="7" pageId="12" pageNumber="2299">1</subScript>
arising before middle; R
<subScript id="A96B3877D061303B56274A86FCA4FB6F" attach="left" box="[849,858,1175,1192]" fontSize="7" pageId="12" pageNumber="2299">2</subScript>
close to R
<subScript id="A96B3877D061303B56A24A86FC23FB6F" attach="left" box="[980,989,1175,1192]" fontSize="7" pageId="12" pageNumber="2299">3</subScript>
at base; R
<subScript id="A96B3877D061303B51214A86FB9EFB6F" attach="left" box="[1111,1120,1175,1192]" fontSize="7" pageId="12" pageNumber="2299">3</subScript>
and R
<subScript id="A96B3877D061303B559E4AA6FF0FFB0F" attach="left" box="[232,241,1207,1224]" fontSize="7" pageId="12" pageNumber="2299">4</subScript>
stalked for half their length; R
<subScript id="A96B3877D061303B572A4AA6FD9BFB0F" attach="left" box="[604,613,1207,1224]" fontSize="7" pageId="12" pageNumber="2299">4</subScript>
and R
<subScript id="A96B3877D061303B57CE4AA6FD3FFB0F" attach="left" box="[696,705,1207,1224]" fontSize="7" pageId="12" pageNumber="2299">5</subScript>
stalked beyond two-thirds of length; R
<subScript id="A96B3877D061303B51FF4AA6FB6CFB0F" attach="left" box="[1161,1170,1207,1224]" fontSize="7" pageId="12" pageNumber="2299">5</subScript>
to termen; M
<subScript id="A96B3877D061303B541F4AC6FE8CFB2F" attach="left" box="[361,370,1239,1256]" fontSize="7" pageId="12" pageNumber="2299">2</subScript>
close to M
<subScript id="A96B3877D061303B54844AC6FE05FB2F" attach="left" box="[498,507,1239,1256]" fontSize="7" pageId="12" pageNumber="2299">3</subScript>
; M
<subScript id="A96B3877D061303B57534AC6FDD0FB2F" attach="left" box="[549,558,1239,1256]" fontSize="7" pageId="12" pageNumber="2299">3</subScript>
and CuA
<subScript id="A96B3877D061303B57E84AC6FD59FB2F" attach="left" box="[670,679,1239,1256]" fontSize="7" pageId="12" pageNumber="2299">1</subScript>
stalked for half length. Hindwing brownish grey, broader than forewing; venation with Rs and M
<subScript id="A96B3877D061303B564D4AE6FCBAFACF" attach="left" box="[827,836,1271,1288]" fontSize="7" pageId="12" pageNumber="2299">1</subScript>
short-stalked; M
<subScript id="A96B3877D061303B517D4AE6FBEAFACF" attach="left" box="[1035,1044,1271,1288]" fontSize="7" pageId="12" pageNumber="2299">3</subScript>
and CuA
<subScript id="A96B3877D061303B51FF4AE6FB6CFACF" attach="left" box="[1161,1170,1271,1288]" fontSize="7" pageId="12" pageNumber="2299">1</subScript>
also short-stalked. Hind tibia with dark-brown scales dorsally, with orange-grey apex.
</paragraph>
<paragraph id="35503A32D061303B55A54B44FBF4F9CB" blockId="12.[211,1172,1365,1548]" pageId="12" pageNumber="2299">
<emphasis id="079BE620D061303B55A54B44FE8CFAAA" box="[211,370,1365,1389]" italics="true" pageId="12" pageNumber="2299">Male genitalia</emphasis>
(
<figureCitation id="ADD426B7D061303B54F44B44FDF3FAAA" box="[386,525,1365,1389]" captionStart="Figure 3" captionStartId="8.[211,278,1613,1635]" captionTargetBox="[213,1172,165,1580]" captionTargetId="figure-5@8.[212,1172,162,1580]" captionTargetPageId="8" captionText="Figure 3. Male genitalia of Torodora spp. (AC) Torodora parotidosa (Wu); (B) aedeagus; (C) abdominal segments; (DF) T. osamensis Park, sp. nov.; (E) aedeagus; (F) abdominal segments; (GI) T. occidentalis Park, sp. nov.; (H) aedeagus; (I) abdominal segments; (JL) T. aritai Park, sp. nov.; (K) aedeagus; (L) abdominal segments." figureDoi="http://doi.org/10.5281/zenodo.5197580" httpUri="https://zenodo.org/record/5197580/files/figure.png" pageId="12" pageNumber="2299">Figure 3GI</figureCitation>
). Uncus heavily sclerotized, stout, gently bent downward, with round apex. Gnathos relatively short, strongly bent median process. Valva broad basally; costa gently concave beyond one-fifth; ventral margin with slightly expanded median lobe; cucullus elongate, slightly upturned before apex; apex more or less acute. Juxta with semiovate free flap in distal part. Saccus rounded anteriorly. Aedeagus nearly straight, as long as valva, with more than 10 short spines in vesica.
</paragraph>
</subSubSection>
<subSubSection id="7DF569B9D061303B55A5484BFE0DF95D" box="[211,499,1626,1690]" pageId="12" pageNumber="2299" type="distribution">
<paragraph id="35503A32D061303B55A5484BFEA6F9B5" blockId="12.[211,499,1626,1690]" box="[211,344,1626,1650]" pageId="12" pageNumber="2299">
<heading id="6E188D5ED061303B55A5484BFEA6F9B5" box="[211,344,1626,1650]" fontSize="10" level="3" pageId="12" pageNumber="2299" reason="5">
<emphasis id="079BE620D061303B55A5484BFEA6F9B5" box="[211,344,1626,1650]" italics="true" pageId="12" pageNumber="2299">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="35503A32D061303B55A54893FE0DF95D" blockId="12.[211,499,1626,1690]" box="[211,499,1666,1690]" pageId="12" pageNumber="2299">
<collectingCountry id="4DF87AA2D061303B55A54893FEB4F95D" box="[211,330,1666,1690]" name="Cambodia" pageId="12" pageNumber="2299">Cambodia</collectingCountry>
(
<collectingRegion id="F72BF4D0D061303B542E4893FE19F95D" box="[344,487,1666,1690]" country="Cambodia" name="Pursat" pageId="12" pageNumber="2299">Pursat Prov.</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="7DF569B9D060303A55F94EB9FC25FF20" box="[143,987,168,231]" pageId="13" pageNumber="2300" type="etymology">
<paragraph id="35503A32D060303A55F94EB9FEF9FF07" blockId="13.[143,987,168,231]" box="[143,263,168,192]" pageId="13" pageNumber="2300">
<heading id="6E188D5ED060303A55F94EB9FEF9FF07" box="[143,263,168,192]" fontSize="10" level="3" pageId="13" pageNumber="2300" reason="5">
<emphasis id="079BE620D060303A55F94EB9FEF9FF07" box="[143,263,168,192]" italics="true" pageId="13" pageNumber="2300">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="35503A32D060303A55F94EDEFC25FF20" blockId="13.[143,987,168,231]" box="[143,987,207,231]" pageId="13" pageNumber="2300">
The species name is derived from the Latin,
<emphasis id="079BE620D060303A570B4EDEFD12FF20" box="[637,748,207,231]" italics="true" pageId="13" pageNumber="2300">occidental</emphasis>
, meaning “western”.
</paragraph>
</subSubSection>
</treatment>
</document>