157 lines
19 KiB
XML
157 lines
19 KiB
XML
<document id="17C66FCAA464160D6919D79CF2EA8FB4" ID-DOI="10.11646/zootaxa.5228.5.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="7543172" ID-ZooBank="88FE61B0-65C6-4E22-8E18-B34BA534302C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1673942899316" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2023" docId="942F8787FFC1636A529D1B4ACA6BBCB0" docLanguage="en" docName="zootaxa.5228.5.1.pdf" docOrigin="Zootaxa 5228 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ochthebius (Aulacochthebius) nigeriensis Perkins 2023, new species" docType="treatment" docVersion="3" lastPageNumber="537" masterDocId="6816FFFFFFE5634E520A1A48C90CB874" masterDocTitle="Taxonomic revision of African water beetles in the genus Ochthebius Leach (1815) subgenus Aulacochthebius Kuwert (1887) (Coleoptera: Hydraenidae)" masterLastPageNumber="546" masterPageNumber="501" pageNumber="537" updateTime="1699447117083" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:titleInfo id="8FC9A5BB7F646F06EE5249AE88779A66">
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<mods:title id="71D16E44E62773FE91ADAFFDCE1F8C60">Taxonomic revision of African water beetles in the genus Ochthebius Leach (1815) subgenus Aulacochthebius Kuwert (1887) (Coleoptera: Hydraenidae)</mods:title>
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<mods:name id="44FE2AE1E416858B87056263B9119DF3" type="personal">
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<mods:roleTerm id="647B10146DC9807B449F4E868E37D6F4">Author</mods:roleTerm>
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<mods:namePart id="2ABF54851A2039E82D57C775495F9B6A">Perkins, Philip D.</mods:namePart>
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<mods:typeOfResource id="4CAB6D2F9AFBF7C564C2970655329101">text</mods:typeOfResource>
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<mods:title id="7369EC4F6855E050ED8CD0841A7334F5">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="A0A547B9F8BA966DE3EAF50EAF7A99A4">
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<mods:date id="671BF6301D5AB3C3544983A147DB3D5F">2023</mods:date>
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<mods:number id="85C16ED2397C6E9FDD8A0832BA89E3E6">2023-01-17</mods:number>
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<mods:detail id="50F28FE258B66C379EC4DBF47E749B15" type="volume">
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<mods:number id="6B35774674585BC51F93DDB1E6B39730">5228</mods:number>
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<mods:number id="F2A1299B251D6E6ADC092FACA01941AF">5</mods:number>
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<mods:start id="C9D8CD35E3D2DF62FC530C5B7D4733E4">501</mods:start>
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<mods:classification id="BD4900EE4DE2E657ACE09A3BCE5A75E0">journal article</mods:classification>
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<mods:identifier id="2B0B212B6DFA110AA6315C41DDB21934" type="DOI">10.11646/zootaxa.5228.5.1</mods:identifier>
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<mods:identifier id="2D9BF81046F073F4185639C5104AB4FA" type="ISSN">1175-5326</mods:identifier>
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<treatment id="942F8787FFC1636A529D1B4ACA6BBCB0" ID-DOI="http://doi.org/10.5281/zenodo.7543229" ID-Zenodo-Dep="7543229" LSID="urn:lsid:plazi:treatment:942F8787FFC1636A529D1B4ACA6BBCB0" httpUri="http://treatment.plazi.org/id/942F8787FFC1636A529D1B4ACA6BBCB0" lastPageNumber="537" pageId="36" pageNumber="537">
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<subSubSection id="549C651AFFC1636A529D1B4ACBC8B968" box="[151,708,258,285]" pageId="36" pageNumber="537" type="nomenclature">
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<paragraph id="1C393691FFC1636A529D1B4ACBC8B968" blockId="36.[151,708,258,320]" box="[151,708,258,285]" pageId="36" pageNumber="537">
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<heading id="477181FDFFC1636A529D1B4ACBC8B968" bold="true" box="[151,708,258,285]" fontSize="11" level="1" pageId="36" pageNumber="537" reason="1">
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<emphasis id="2EF2EA83FFC1636A529D1B4ACBC8B968" bold="true" box="[151,708,258,285]" pageId="36" pageNumber="537">
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<taxonomicName id="DB864D12FFC1636A529D1B4ACB2AB968" authority="Perkins, 2023" authorityName="Perkins" authorityYear="2023" box="[151,550,258,285]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="537" phylum="Arthropoda" rank="species" species="nigeriensis" status="sp. nov." subGenus="Aulacochthebius">
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<emphasis id="2EF2EA83FFC1636A529D1B4ACB2AB968" bold="true" box="[151,550,258,285]" italics="true" pageId="36" pageNumber="537">O. (Aulacochthebius) nigeriensis</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="35C157F8FFC1636A503E1B4ACBC8B968" box="[564,708,258,284]" pageId="36" pageNumber="537" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="549C651AFFC1636A529D1B6EC85FB934" box="[151,339,294,320]" pageId="36" pageNumber="537" type="description">
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||
<paragraph id="1C393691FFC1636A529D1B6EC85FB934" blockId="36.[151,708,258,320]" box="[151,339,294,320]" pageId="36" pageNumber="537">
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<figureCitation id="84BD2A14FFC1636A529D1B6EC9E9B934" box="[151,229,294,320]" captionStart="FIGURE 2" captionStartId="8.[151,250,1966,1990]" captionTargetBox="[213,1380,198,1924]" captionTargetId="figure-25@8.[177,1410,181,1942]" captionTargetPageId="8" captionText="FIGURE 2. Dorsal habitus of species in O. (A.) tenuipunctus Group." figureDoi="http://doi.org/10.5281/zenodo.7543176" httpUri="https://zenodo.org/record/7543176/files/figure.png" pageId="36" pageNumber="537">Figs. 2</figureCitation>
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,
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<figureCitation id="84BD2A14FFC1636A52FB1B6EC80DB934" box="[241,257,294,320]" captionStart="FIGURE 4" captionStartId="10.[151,250,1867,1891]" captionTargetBox="[170,1423,212,1791]" captionTargetId="figure-25@10.[151,1435,181,1844]" captionTargetPageId="10" captionText="FIGURE 4. Venter and aedeagi (ventral view, flagellum deleted) of species in O. (A.) continentalis Group and O. (A.) tenuipunctus Group. Not to same scale." figureDoi="http://doi.org/10.5281/zenodo.7543186" httpUri="https://zenodo.org/record/7543186/files/figure.png" pageId="36" pageNumber="537">4</figureCitation>
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,
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<figureCitation id="84BD2A14FFC1636A53071B6EC827B934" box="[269,299,294,320]" captionStart="FIGURE 19" captionStartId="33.[151,250,1944,1968]" captionTargetBox="[197,1392,196,1884]" captionTargetId="figure-15@33.[164,1424,181,1920]" captionTargetPageId="33" captionText="FIGURE 19. O. (A.) nigeriensis n. sp., holotype habitus and aedeagus." figureDoi="http://doi.org/10.5281/zenodo.7543241" httpUri="https://zenodo.org/record/7543241/files/figure.png" pageId="36" pageNumber="537">19</figureCitation>
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,
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<figureCitation id="84BD2A14FFC1636A533D1B6EC85FB934" box="[311,339,294,320]" captionStart="FIGURES 27–28" captionStartId="41.[151,264,1572,1596]" captionTargetBox="[151,1435,181,1549]" captionTargetId="figure-15@41.[151,1435,181,1549]" captionTargetPageId="41" captionText="FIGURES 27–28. Geographical distributions. 27. O. (A.) species in Kenya and Uganda. 28. O. (A.) strangulatus Régimbart, O. (A.) tenuipunctus Régimbart and O. (A.) nigeriensis n. sp." figureDoi="http://doi.org/10.5281/zenodo.7543257" httpUri="https://zenodo.org/record/7543257/files/figure.png" pageId="36" pageNumber="537">28</figureCitation>
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</paragraph>
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</subSubSection>
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||
<subSubSection id="549C651AFFC1636A529D1B26CAA2B9D8" pageId="36" pageNumber="537" type="materials_examined">
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<paragraph id="1C393691FFC1636A529D1B26CAA2B9D8" blockId="36.[151,1437,366,1221]" pageId="36" pageNumber="537">
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<emphasis id="2EF2EA83FFC1636A529D1B26C847B9FC" bold="true" box="[151,331,366,392]" pageId="36" pageNumber="537">Type Material.</emphasis>
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<materialsCitation id="ACEE3CCCFFC1636A535E1B26C878B9D8" collectingDate="1949-02-12" collectingDateMax="1949-02-13" collectingDateMin="1949-02-12" collectionCode="NHMUK" collectorName="B. Malkin" country="Nigeria" latitude="8.133333" location="Ogbomosho" longLatPrecision="1302" longitude="4.25" pageId="36" pageNumber="537" specimenCount="1" stateProvince="Oyo" typeStatus="holotype">
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<emphasis id="2EF2EA83FFC1636A535E1B26CB13B9FC" bold="true" box="[340,543,366,392]" pageId="36" pageNumber="537">
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<typeStatus id="C33D8833FFC1636A535E1B26C8CDB9FC" box="[340,449,366,392]" pageId="36" pageNumber="537" type="holotype">Holotype</typeStatus>
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(male):
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</emphasis>
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“
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<collectingCountry id="64917601FFC1636A50391B26CBA1B9FC" box="[563,685,366,392]" name="Nigeria" pageId="36" pageNumber="537">NIGERIA</collectingCountry>
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:
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<collectingRegion id="DE42F873FFC1636A50B01B26CA3DB9FD" box="[698,817,366,393]" country="Nigeria" name="Oyo" pageId="36" pageNumber="537">Oyo Prov.</collectingRegion>
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,
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<location id="1959604AFFC1636A51341B26CAC6B9FC" LSID="urn:lsid:plazi:treatment:942F8787FFC1636A529D1B4ACA6BBCB0:1959604AFFC1636A51341B26CAC6B9FC" box="[830,970,366,392]" country="Nigeria" latitude="8.133333" longLatPrecision="1302" longitude="4.25" name="Ogbomosho" pageId="36" pageNumber="537" stateProvince="Oyo">Ogbomosho</location>
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,
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<date id="68381051FFC1636A51DD1B26CD7DB9FC" box="[983,1137,366,392]" pageId="36" pageNumber="537" value="1949-02-12" valueMax="1949-02-13" valueMin="1949-02-12">
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||
<collectingDate id="787CE9B9FFC1636A51DD1B26CD7DB9FC" box="[983,1137,366,392]" pageId="36" pageNumber="537" value="1949-02-12" valueMax="1949-02-13" valueMin="1949-02-12">12-13.ii.1949</collectingDate>
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</date>
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.
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<collectorName id="B1735347FFC1636A56741B27CDFEB9FC" box="[1150,1266,366,393]" pageId="36" pageNumber="537">B. Malkin</collectorName>
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coll.”; [
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<geoCoordinate id="79B25056FFC1636A57581B38CC94B9FF" box="[1362,1432,368,395]" degrees="8" direction="north" minutes="8" orientation="latitude" pageId="36" pageNumber="537" precision="925" value="8.133333">8°8′N</geoCoordinate>
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,
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<geoCoordinate id="79B25056FFC1636A529D1BDCC9E9B9DB" box="[151,229,404,431]" degrees="4" direction="east" minutes="15" orientation="longitude" pageId="36" pageNumber="537" precision="925" value="4.25">4°15′E</geoCoordinate>
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] (
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<collectionCode id="7A97AE54FFC1636A52F61BDAC861B9D8" box="[252,365,402,428]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="36" pageNumber="537" type="Museum">NHMUK</collectionCode>
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)
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</materialsCitation>
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.
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<materialsCitation id="ACEE3CCCFFC1636A53741BDACAA7B9D8" box="[382,939,402,429]" collectingDate="1949-02-12" collectingDateMax="1949-02-13" collectingDateMin="1949-02-12" collectionCode="NHMUK" collectorName="B. Malkin" country="Nigeria" latitude="8.133333" location="Ogbomosho" longLatPrecision="1302" longitude="4.25" pageId="36" pageNumber="537" specimenCount="1" stateProvince="Oyo" typeStatus="paratype">
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<emphasis id="2EF2EA83FFC1636A53741BDAC8FAB9D8" bold="true" box="[382,502,402,428]" pageId="36" pageNumber="537">
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<typeStatus id="C33D8833FFC1636A53741BDAC8FAB9D8" box="[382,502,402,428]" pageId="36" pageNumber="537" type="paratype">Paratypes</typeStatus>
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</emphasis>
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: Same data as holotype (12
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<collectionCode id="7A97AE54FFC1636A51391BDACAAEB9D8" box="[819,930,402,428]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="36" pageNumber="537" type="Museum">NHMUK</collectionCode>
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)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="549C651AFFC1636A52CD1BFECDAABA6C" pageId="36" pageNumber="537" type="diagnosis">
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<paragraph id="1C393691FFC1636A52CD1BFECDAABA6C" blockId="36.[151,1437,366,1221]" pageId="36" pageNumber="537">
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<emphasis id="2EF2EA83FFC1636A52CD1BFEC8DCB9A4" bold="true" box="[199,464,438,464]" pageId="36" pageNumber="537">Differential diagnosis.</emphasis>
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Differentiated from other similar members of the
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<taxonomicName id="DB864D12FFC1636A561E1BFFCDF7B9A4" box="[1044,1275,439,465]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="537" phylum="Arthropoda" rank="species" species="tenuipunctus" subGenus="Aulacochthebius">
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<emphasis id="2EF2EA83FFC1636A561E1BFFCDF7B9A4" box="[1044,1275,439,465]" italics="true" pageId="36" pageNumber="537">O. (A.) tenuipunctus</emphasis>
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</taxonomicName>
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Group by the combination of the slightly smaller size, and by the aedeagus. Very similar in characters to
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<taxonomicName id="DB864D12FFC1636A56A81B93CC45B980" box="[1186,1353,474,501]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="537" phylum="Arthropoda" rank="species" species="alienus" subGenus="Aulacochthebius">
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<emphasis id="2EF2EA83FFC1636A56A81B93CC45B980" box="[1186,1353,474,501]" italics="true" pageId="36" pageNumber="537">O. (A.) alienus</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="DB864D12FFC1636A578B1B93C83FBA6C" authorityName="Perkins" authorityYear="2023" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="537" phylum="Arthropoda" rank="species" species="solitarius" subGenus="Aulacochthebius">
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<emphasis id="2EF2EA83FFC1636A578B1B93C83FBA6C" italics="true" pageId="36" pageNumber="537">O. (A.) solitarius</emphasis>
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</taxonomicName>
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; reliable determinations will require careful examination of the male genitalia.
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</paragraph>
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</subSubSection>
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<subSubSection id="549C651AFFC1636A52CD186ACB05BC09" pageId="36" pageNumber="537" type="description">
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<paragraph id="1C393691FFC1636A52CD186ACAC8BA85" blockId="36.[151,1437,366,1221]" pageId="36" pageNumber="537">
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<emphasis id="2EF2EA83FFC1636A52CD186AC85DBA48" bold="true" box="[199,337,546,572]" pageId="36" pageNumber="537">Description</emphasis>
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. Size:
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<typeStatus id="C33D8833FFC1636A53AE186ACB0ABA48" box="[420,518,546,572]" pageId="36" pageNumber="537" type="holotype">holotype</typeStatus>
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(length/width, mm): body 1.16/0.57 (length to elytral apices); head width 0.35; pronotum length 0.30, width at base 0.22, width at hyaline border 0.55, width of lateral sclerotized lobes 0.42 at anterior, 0.41 at posterior; elytra 0.76/0.57. Habitus as illustrated (
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<figureCitation id="84BD2A14FFC1636A51861822CAEEBAF0" box="[908,994,618,644]" captionStart="FIGURE 19" captionStartId="33.[151,250,1944,1968]" captionTargetBox="[197,1392,196,1884]" captionTargetId="figure-15@33.[164,1424,181,1920]" captionTargetPageId="33" captionText="FIGURE 19. O. (A.) nigeriensis n. sp., holotype habitus and aedeagus." figureDoi="http://doi.org/10.5281/zenodo.7543241" httpUri="https://zenodo.org/record/7543241/files/figure.png" pageId="36" pageNumber="537">Fig. 19</figureCitation>
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). Body longitudinally and transversely very convex. Dorsum brown to dark brown except hyaline borders of prothorax translucent. Eyes with moderately coarse, convex facets. Frons with deep impressions, borders of impressions rounded, smooth. Clypeus with very few, very shallow setigerous punctures. Labrum anterior margin arcuate.
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</paragraph>
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<paragraph id="1C393691FFC1636A52CD18B2CAF5BBBD" blockId="36.[151,1437,366,1221]" pageId="36" pageNumber="537">Pronotal reliefs very smooth, with very sparse and very short setae; anterior and posterior transverse impressions deep, anterior contiguous with shallow transverse impression immediately behind anterior margin of lateral lobe; median longitudinal impression obsolete, almost imperceptible, only present at anterior transverse impression. Pronotum anterior margin bisinuate. Pronotum lateral hyaline borders very wide, extended from posterior to anterior angles, outer margin strongly arcuate; sides of non-hyaline part with long conspicuous setae immediately above and resting on hyaline border; anterior and posterior hyaline borders very narrow.</paragraph>
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<paragraph id="1C393691FFC1636A52CD199ACA40BC40" blockId="36.[151,1437,366,1221]" pageId="36" pageNumber="537">Elytra slightly wider than hyaline part of pronotum, posteriorly acuminate, sides markedly arcuate; very smooth, non-punctate, with sparse, very short, widely separated serial or subserial setae; lateral margin with very dense, very short setae that extend outward from margin in dry specimens</paragraph>
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<paragraph id="1C393691FFC1636A529D1E76CB05BC09" blockId="36.[151,1437,366,1221]" pageId="36" pageNumber="537">
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Abdominal venter (
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<figureCitation id="84BD2A14FFC1636A537B1E76C8C9BC2C" box="[369,453,1086,1112]" captionStart="FIGURE 19" captionStartId="33.[151,250,1944,1968]" captionTargetBox="[197,1392,196,1884]" captionTargetId="figure-15@33.[164,1424,181,1920]" captionTargetPageId="33" captionText="FIGURE 19. O. (A.) nigeriensis n. sp., holotype habitus and aedeagus." figureDoi="http://doi.org/10.5281/zenodo.7543241" httpUri="https://zenodo.org/record/7543241/files/figure.png" pageId="36" pageNumber="537">Fig. 19</figureCitation>
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): first four ventrites bearing rather long hydrofuge pubescence; remaining ventrites glabrous, shining, with sparse setae.
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</paragraph>
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</subSubSection>
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<subSubSection id="549C651AFFC1636A52CD1ECECA57BCD4" box="[199,859,1158,1185]" pageId="36" pageNumber="537" type="etymology">
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<paragraph id="1C393691FFC1636A52CD1ECECA57BCD4" blockId="36.[151,1437,366,1221]" box="[199,859,1158,1185]" pageId="36" pageNumber="537">
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<emphasis id="2EF2EA83FFC1636A52CD1ECEC842BCD4" bold="true" box="[199,334,1158,1184]" pageId="36" pageNumber="537">Etymology.</emphasis>
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Named in reference to the known distribution.
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</paragraph>
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</subSubSection>
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<subSubSection id="549C651AFFC1636A52CD1EE2CA6BBCB0" box="[199,871,1194,1221]" pageId="36" pageNumber="537" type="distribution">
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||
<paragraph id="1C393691FFC1636A52CD1EE2CA6BBCB0" blockId="36.[151,1437,366,1221]" box="[199,871,1194,1221]" pageId="36" pageNumber="537">
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||
<emphasis id="2EF2EA83FFC1636A52CD1EE2C855BCB0" bold="true" box="[199,345,1194,1220]" pageId="36" pageNumber="537">Distribution</emphasis>
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. Currently known only from
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<collectingCountry id="64917601FFC1636A50A91EE2CBFBBCB0" box="[675,759,1194,1220]" name="Nigeria" pageId="36" pageNumber="537">Nigeria</collectingCountry>
|
||
(
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<figureCitation id="84BD2A14FFC1636A510C1EE2CA57BCB0" box="[774,859,1194,1220]" captionStart="FIGURES 27–28" captionStartId="41.[151,264,1572,1596]" captionTargetBox="[151,1435,181,1549]" captionTargetId="figure-15@41.[151,1435,181,1549]" captionTargetPageId="41" captionText="FIGURES 27–28. Geographical distributions. 27. O. (A.) species in Kenya and Uganda. 28. O. (A.) strangulatus Régimbart, O. (A.) tenuipunctus Régimbart and O. (A.) nigeriensis n. sp." figureDoi="http://doi.org/10.5281/zenodo.7543257" httpUri="https://zenodo.org/record/7543257/files/figure.png" pageId="36" pageNumber="537">Fig. 28</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |