treatments-xml/data/5B/50/E9/5B50E916FFD4396F4D2DFA7AFBABFE84.xml
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<mods:title id="6DF33990EE7AEAE8126221F666988841">A revision of Nearctic species of the genus Geostiba Thomson, 1858 (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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15.
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<emphasis id="E18D8412FFD439694D11FA7AFE76FA3A" bold="true" box="[308,414,1478,1504]" italics="true" pageId="74" pageNumber="75">Geostiba</emphasis>
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(
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.)
<taxonomicName id="14F92383FFD439694DDBFA7AFC92FA3A" authority="Gravenhorst, 1806" authorityName="Gravenhorst" authorityYear="1806" box="[510,890,1478,1504]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="circellaris">
<emphasis id="E18D8412FFD439694DDBFA7AFD9FFA3A" bold="true" box="[510,631,1478,1504]" italics="true" pageId="74" pageNumber="75">circellaris</emphasis>
(
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)
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(
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;
<figureCitation id="4BC24485FFD439694801FA7AFB66FA3A" box="[1060,1166,1478,1504]" captionStart="FIGURES 1 ­ 7" captionStartId="5.[264,383,1751,1775]" captionTargetBox="[367,1205,490,1733]" captionTargetId="figure@5.[365,1207,490,1736]" captionTargetPageId="5" captionText="FIGURES 1 ­ 7. Mouthparts of Geostiba circellaris (Gravenhorst) (1 ­ 2, Moscow Region, Russia), G. carteriensis Pace (3 ­ 4, Tennessee), G. appalachigena Gusarov, sp. n. (5 ­ 6, paratype from West Virginia) and G. pluvigena Gusarov, sp. n. (7, paratype from North Carolina). 1, 3 labrum; 2, 4 epipharynx; 5 left mandible, dorsal view; 6 left mandible, ventral view; 7 right mandible, dorsal view. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/155702/files/figure.png" pageId="74" pageNumber="75">
Figs.
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in
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</emphasis>
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</paragraph>
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<emphasis id="E18D8412FFD439694D2DF988FE02F991" box="[264,490,1588,1611]" italics="true" pageId="74" pageNumber="75">Aleochara circellaris</emphasis>
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: 155
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.
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</paragraph>
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<treatmentCitationGroup id="F3E97F2EFFD439694D2DF9E8FD54F951" pageId="74" pageNumber="75">
<taxonomicName id="14F92383FFD439694D2DF9E8FE80F9B1" ID-CoL="8VRPP" box="[264,360,1620,1643]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="genus">
<emphasis id="E18D8412FFD439694D2DF9E8FE80F9B1" box="[264,360,1620,1643]" italics="true" pageId="74" pageNumber="75">Geostiba</emphasis>
</taxonomicName>
(
<emphasis id="E18D8412FFD439694D5DF9E8FDC3F9B1" box="[376,555,1620,1643]" italics="true" pageId="74" pageNumber="75">
s. str.)
<taxonomicName id="14F92383FFD439694D9AF9E8FDC3F9B1" authority="Benick &amp; Lohse 1974: 112" authorityName="Benick &amp; Lohse" authorityPageNumber="112" authorityYear="1974" box="[447,555,1620,1643]" class="Insecta" family="Staphylinidae" genus="Aleochara" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="circellaris">circellaris</taxonomicName>
</emphasis>
:
<treatmentCitation id="52587E11FFD439694E65F9E8FC82F9B1" author="Benick" box="[576,874,1620,1643]" page="112" pageId="74" pageNumber="75" year="1974">
<bibRefCitation id="B76825F1FFD439694E65F9E8FCC7F9B1" author="Benick" box="[576,815,1620,1643]" lastPageId="74" lastPageNumber="87" pageId="74" pageNumber="86" refString="Benick, G. &amp; Lohse, G. A. (1974) 14. Tribus: Callicerini (Athetae). In: Freude, H., Harde, K. W. &amp; Lohse, G. A. (Eds.), Die Kafer Mitteleuropas. Band 5, Staphylinidae II (Hypocyphtinae und Aleocharinae). Pselaphidae. Krefeld: Goecke &amp; Evers Verlag, pp. 72 - 220." type="book chapter" year="1974">Benick &amp; Lohse 1974</bibRefCitation>
: 112
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.
<taxonomicName id="14F92383FFD439694D2DF9C8FD50F951" ID-CoL="3FSFC" authority="Muona 1984: 229" authorityName="Muona" authorityPageNumber="229" authorityYear="1984" box="[264,696,1652,1675]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="circellaris">
<emphasis id="E18D8412FFD439694D2DF9C8FE33F951" box="[264,475,1652,1675]" italics="true" pageId="74" pageNumber="75">Geostiba circellaris</emphasis>
:
<treatmentCitation id="52587E11FFD439694DD5F9C8FD50F951" author="Muona" box="[496,696,1652,1675]" page="229" pageId="74" pageNumber="75" year="1984">
<bibRefCitation id="B76825F1FFD439694DD5F9C8FD94F951" author="Muona" box="[496,636,1652,1675]" pageId="74" pageNumber="88" refString="Muona, J. (1984) Review of Palaearctic Aleocharinae also occurring in North America (Coleoptera: Staphylinidae). Entomologica scandinavica, 15, 227 - 231." type="journal article" year="1984">Muona 1984</bibRefCitation>
: 229
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.
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</paragraph>
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<taxonomicName id="14F92383FFD439694D2DF928FE80F971" ID-CoL="8VRPP" box="[264,360,1684,1707]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="genus">
<emphasis id="E18D8412FFD439694D2DF928FE80F971" box="[264,360,1684,1707]" italics="true" pageId="74" pageNumber="75">Geostiba</emphasis>
</taxonomicName>
(
<emphasis id="E18D8412FFD439694D5DF928FE43F971" box="[376,427,1684,1707]" italics="true" pageId="74" pageNumber="75">s. str</emphasis>
.)
<taxonomicName id="14F92383FFD439694DE4F928FCEFF971" ID-CoL="3FSFC" authority="Assing 2001: 139" authorityName="Assing" authorityPageNumber="139" authorityYear="2001" box="[449,775,1684,1707]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="circellaris">
<emphasis id="E18D8412FFD439694DE4F928FDC5F971" box="[449,557,1684,1707]" italics="true" pageId="74" pageNumber="75">circellaris</emphasis>
:
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: 139
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.
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</paragraph>
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<emphasis id="E18D8412FFD439694D2DF967FCECF92F" bold="true" box="[264,772,1755,1781]" pageId="74" pageNumber="75">
Material.
<collectingCountry id="ABEE1890FFD439694DBEF967FDF7F92F" box="[411,543,1755,1781]" name="Canada" pageId="74" pageNumber="75">CANADA</collectingCountry>
: Newfoundland:
</emphasis>
,, S Newfoundland, Rencontre West [
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] (Lindroth),
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(
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).
</paragraph>
</subSubSection>
<subSubSection id="9BE30B8BFFD439684D1DF897FE54FDAC" lastPageId="75" lastPageNumber="76" pageId="74" pageNumber="75" type="diagnosis">
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<emphasis id="E18D8412FFD439694D1DF897FE5FF89F" bold="true" box="[312,439,1835,1861]" pageId="74" pageNumber="75">Diagnosis.</emphasis>
<taxonomicName id="14F92383FFD439694DEEF890FD5EF89F" box="[459,694,1836,1861]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="circellaris">
<emphasis id="E18D8412FFD439694DEEF890FD5EF89F" box="[459,694,1836,1861]" italics="true" pageId="74" pageNumber="75">Geostiba circellaris</emphasis>
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can be distinguished from other Nearctic species of
<taxonomicName id="14F92383FFD439694D2DF8E8FE9AF8B7" box="[264,370,1876,1901]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="genus">
<emphasis id="E18D8412FFD439694D2DF8E8FE9AF8B7" box="[264,370,1876,1901]" italics="true" pageId="74" pageNumber="75">Geostiba</emphasis>
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by having large eyes (temple length to eye length ratio 1.6­1.9), pronotal pubescence of
<typeStatus id="0C42E6A2FFD539684D54FEA0FE4CFEEC" box="[369,420,284,310]" pageId="75" pageNumber="76">type</typeStatus>
VI, reduced wings, elytra shorter than pronotum (pronotum length to elytron length ratio 1.1), the presence of one medial carina on male abdominal tergum
<quantity id="1401F5E5FFD539684885FEF8FB24FE84" box="[1184,1228,324,350]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="75" pageNumber="76" unit="in" value="7.0">7 in</quantity>
front of posterior margin, the shape of the aedeagus and spermatheca (
<figureCitation id="4BC24485FFD539684FCCFED0FBBBFE5C" box="[1001,1107,364,390]" captionStart="FIGURES 1 ­ 7" captionStartId="5.[264,383,1751,1775]" captionTargetBox="[367,1205,490,1733]" captionTargetId="figure@5.[365,1207,490,1736]" captionTargetPageId="5" captionText="FIGURES 1 ­ 7. Mouthparts of Geostiba circellaris (Gravenhorst) (1 ­ 2, Moscow Region, Russia), G. carteriensis Pace (3 ­ 4, Tennessee), G. appalachigena Gusarov, sp. n. (5 ­ 6, paratype from West Virginia) and G. pluvigena Gusarov, sp. n. (7, paratype from North Carolina). 1, 3 labrum; 2, 4 epipharynx; 5 left mandible, dorsal view; 6 left mandible, ventral view; 7 right mandible, dorsal view. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/155702/files/figure.png" pageId="75" pageNumber="76">
Figs.
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in
<bibRefCitation id="B76825F1FFD53968485EFED0FACEFE5C" author="Assing" box="[1147,1318,364,390]" pageId="75" pageNumber="86" refString="Assing, V. (2001) A revision of the Turkish species of Geostiba Thomson 1858 and Tropimenelytron Pace 1983 (Coleoptera: Staphylinidae, Aleocharinae). Linzer biologische Beitrage, 33 (1), 137 - 185." type="journal article" year="2001">Assing (2001)</bibRefCitation>
,
<figureCitation id="4BC24485FFD539684D2DFE28FE79FE74" box="[264,401,404,430]" captionStart="FIGURES 21 ­ 25" captionStartId="10.[264,383,1697,1721]" captionTargetBox="[310,1273,423,1660]" captionTargetId="figure@10.[302,1285,409,1671]" captionTargetPageId="10" captionText="FIGURES 21 ­ 25. Partially everted internal sac of aedeagus of Geostiba circellaris (Gravenhorst) (21 ­ 23, St. Petersburg Region, Russia) and G. infirma (Weise) (24 ­ 25, Ivano­Frankovsk Region, Ukraine). 21, 24 ventral view; 22, 25 lateral view; 23 dorsal view. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/155705/files/figure.png" pageId="75" pageNumber="76">Figs. 21­23</figureCitation>
).
</paragraph>
<paragraph id="D3465800FFD539684D1DFE00FE54FDAC" blockId="75.[264,1324,284,1870]" pageId="75" pageNumber="76">
In North
<collectingCountry id="ABEE1890FFD539684D81FE00FDE1FE0C" box="[420,521,444,470]" name="United States of America" pageId="75" pageNumber="76">America</collectingCountry>
there is no native species closely related to
<taxonomicName id="14F92383FFD53968482AFE00FB42FE0F" box="[1039,1194,444,469]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="circellaris">
<emphasis id="E18D8412FFD53968482AFE00FB42FE0F" box="[1039,1194,444,469]" italics="true" pageId="75" pageNumber="76">G. circellaris</emphasis>
</taxonomicName>
.
<taxonomicName id="14F92383FFD5396848E4FE00FE97FE27" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="circellaris">
<emphasis id="E18D8412FFD5396848E4FE00FE97FE27" italics="true" pageId="75" pageNumber="76">Geostiba circellaris</emphasis>
</taxonomicName>
is superficially similar to
<taxonomicName id="14F92383FFD539684E9DFE58FCB5FE27" box="[696,861,484,509]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="impressula">
<emphasis id="E18D8412FFD539684E9DFE58FCB5FE27" box="[696,861,484,509]" italics="true" pageId="75" pageNumber="76">G. impressula</emphasis>
</taxonomicName>
in having pronotal pubescence of
<typeStatus id="0C42E6A2FFD5396848D2FE58FAC1FE24" box="[1271,1321,484,510]" pageId="75" pageNumber="76">type</typeStatus>
VI but can be distinguished from the latter species by having tergum 7 with one medial carina in males and a different shape of spermatheca in females (
<figureCitation id="4BC24485FFD53968482EFD88FBBAFD94" box="[1035,1106,564,590]" captionStart="FIGURES 1 ­ 7" captionStartId="5.[264,383,1751,1775]" captionTargetBox="[367,1205,490,1733]" captionTargetId="figure@5.[365,1207,490,1736]" captionTargetPageId="5" captionText="FIGURES 1 ­ 7. Mouthparts of Geostiba circellaris (Gravenhorst) (1 ­ 2, Moscow Region, Russia), G. carteriensis Pace (3 ­ 4, Tennessee), G. appalachigena Gusarov, sp. n. (5 ­ 6, paratype from West Virginia) and G. pluvigena Gusarov, sp. n. (7, paratype from North Carolina). 1, 3 labrum; 2, 4 epipharynx; 5 left mandible, dorsal view; 6 left mandible, ventral view; 7 right mandible, dorsal view. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/155702/files/figure.png" pageId="75" pageNumber="76">
Fig.
<quantity id="1401F5E5FFD539684866FD88FBBAFD94" box="[1091,1106,564,590]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="75" pageNumber="76" unit="in" value="4.0">4</quantity>
</figureCitation>
in
<bibRefCitation id="B76825F1FFD53968485CFD88FACCFD94" author="Assing" box="[1145,1316,564,590]" pageId="75" pageNumber="86" refString="Assing, V. (2001) A revision of the Turkish species of Geostiba Thomson 1858 and Tropimenelytron Pace 1983 (Coleoptera: Staphylinidae, Aleocharinae). Linzer biologische Beitrage, 33 (1), 137 - 185." type="journal article" year="2001">Assing (2001)</bibRefCitation>
,
<figureCitation id="4BC24485FFD539684D2DFDE0FE58FDAC" box="[264,432,604,630]" captionStart="FIGURES 316 ­ 320" captionStartId="71.[264,383,1481,1505]" captionTargetBox="[294,1301,904,1448]" captionTargetId="figure@71.[281,1305,889,1455]" captionTargetPageId="71" captionText="FIGURES 316 ­ 320. Genitalia of Geostiba impressula (Casey) (holotype (316) and specimens from Triangle Lake, Oregon (317 ­ 320 )). 316 ­ 317 spermatheca; 318 everted internal sac, ventral view; 319 everted internal sac, lateral view; 320 apex of left paramere, side facing median lobe. Scale bar 0.2 mm (316 ­ 317, 320), 0.4 mm (318 ­ 319)." httpUri="https://zenodo.org/record/155745/files/figure.png" pageId="75" pageNumber="76">Figs. 316­317</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="9BE30B8BFFD539684D1DFD3FFCC5F90C" pageId="75" pageNumber="76" type="description">
<paragraph id="D3465800FFD539684D1DFD3FFACDFD34" blockId="75.[264,1324,284,1870]" pageId="75" pageNumber="76">
<emphasis id="E18D8412FFD539684D1DFD3FFE38FD47" bold="true" box="[312,464,643,669]" pageId="75" pageNumber="76">Description.</emphasis>
Length
<quantity id="1401F5E5FFD539684E65FD38FD26FD44" box="[576,718,644,670]" metricMagnitude="-3" metricUnit="m" metricValue="2.55" metricValueMax="2.8" metricValueMin="2.3" pageId="75" pageNumber="76" unit="mm" value="2.55" valueMax="2.8" valueMin="2.3">2.3­2.8 mm</quantity>
. Body reddish brown to brownish black, in some specimens pronotum, elytra and abdominal segments 7­8 lighter, antennae reddish brown to dark brown, legs and mouthparts from brownish yellow to brown. Body parallel­sided.
</paragraph>
<paragraph id="D3465800FFD539684D1DFD40FB67FC54" blockId="75.[264,1324,284,1870]" pageId="75" pageNumber="76">Head as wide as long, surface on disk with fine isodiametric microsculpture, puncturation very fine, distance between punctures equal to 2­4 times their diameter. Temple length to eye length ratio 1.6­1.9. Antennal article 2 longer than article 3, article 4 subquadrate, articles 5­10 transverse, last article longer than 9 and 10 combined.</paragraph>
<paragraph id="D3465800FFD539684D1DFC20FC49FB7C" blockId="75.[264,1324,284,1870]" pageId="75" pageNumber="76">
Pronotum as wide as long, width
<quantity id="1401F5E5FFD539684EEAFC20FC93FC6C" box="[719,891,924,950]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" metricValueMax="5.3" metricValueMin="4.3" pageId="75" pageNumber="76" unit="mm" value="0.48" valueMax="0.53" valueMin="0.43">0.43­0.53 mm</quantity>
, wider than head (pronotal width to head width ratio 1.2); surface on disk with fine isodiametric microsculpture, and fine puncturation, distance between punctures equal to 1­2 times their diameter; pronotal pubescence of
<typeStatus id="0C42E6A2FFD539684DE6FBA8FE1EFBF4" box="[451,502,1044,1070]" pageId="75" pageNumber="76">type</typeStatus>
VI. Elytra measured from humeral angle shorter than pronotum (pronotal length to elytral length ratio 1.1), wider than long (1.4), with fine isodiametric microsculpture and fine asperate puncturation, punctures stronger than on pronotum, distance between punctures equals 1­2 times their diameter.
</paragraph>
<paragraph id="D3465800FFD539684D1DFB08FC2AFAC4" blockId="75.[264,1324,284,1870]" pageId="75" pageNumber="76">Abdominal terga with fine transverse microsculpture, with fine puncturation, puncturation becoming finer towards abdomen apex, on terga 3­5 distance between punctures equals 2­4 times their diameter. Tergum 7 with white edge.</paragraph>
<paragraph id="D3465800FFD539684D1DFA90FD29F9D4" blockId="75.[264,1324,284,1870]" pageId="75" pageNumber="76">In males pronotum with small medial tubercle at posterior margin (some females have very weak tubercle) and medial impression in front, each elytron with a tubercle near scutellum but distant from elytral suture. Male tergum 7 with one short medial carina in front of posterior margin. Male tergum 8 with two weak and short parallel carina in front of middle of convex posterior margin. Male sternum 8 with posterior margin convex, extending posteriorly beyond tergum.</paragraph>
<paragraph id="D3465800FFD539684D1DF9A0FD86F984" blockId="75.[264,1324,284,1870]" pageId="75" pageNumber="76">
Aedeagus as in
<figureCitation id="4BC24485FFD539684DD1F9A0FDB6F9EC" box="[500,606,1564,1590]" captionStart="FIGURES 1 ­ 7" captionStartId="5.[264,383,1751,1775]" captionTargetBox="[367,1205,490,1733]" captionTargetId="figure@5.[365,1207,490,1736]" captionTargetPageId="5" captionText="FIGURES 1 ­ 7. Mouthparts of Geostiba circellaris (Gravenhorst) (1 ­ 2, Moscow Region, Russia), G. carteriensis Pace (3 ­ 4, Tennessee), G. appalachigena Gusarov, sp. n. (5 ­ 6, paratype from West Virginia) and G. pluvigena Gusarov, sp. n. (7, paratype from North Carolina). 1, 3 labrum; 2, 4 epipharynx; 5 left mandible, dorsal view; 6 left mandible, ventral view; 7 right mandible, dorsal view. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/155702/files/figure.png" pageId="75" pageNumber="76">
Figs.
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</figureCitation>
in
<bibRefCitation id="B76825F1FFD539684EA3F9A0FCDAF9EC" author="Assing" box="[646,818,1564,1590]" pageId="75" pageNumber="86" refString="Assing, V. (2001) A revision of the Turkish species of Geostiba Thomson 1858 and Tropimenelytron Pace 1983 (Coleoptera: Staphylinidae, Aleocharinae). Linzer biologische Beitrage, 33 (1), 137 - 185." type="journal article" year="2001">Assing (2001)</bibRefCitation>
. Internal sac of aedeagus with one pair of large diverticula (
<figureCitation id="4BC24485FFD539684DF2F9F8FD89F984" box="[471,609,1604,1630]" captionStart="FIGURES 21 ­ 25" captionStartId="10.[264,383,1697,1721]" captionTargetBox="[310,1273,423,1660]" captionTargetId="figure@10.[302,1285,409,1671]" captionTargetPageId="10" captionText="FIGURES 21 ­ 25. Partially everted internal sac of aedeagus of Geostiba circellaris (Gravenhorst) (21 ­ 23, St. Petersburg Region, Russia) and G. infirma (Weise) (24 ­ 25, Ivano­Frankovsk Region, Ukraine). 21, 24 ventral view; 22, 25 lateral view; 23 dorsal view. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/155705/files/figure.png" pageId="75" pageNumber="76">Figs. 21­22</figureCitation>
).
</paragraph>
<paragraph id="D3465800FFD539684D1DF9D0FD0BF974" blockId="75.[264,1324,284,1870]" pageId="75" pageNumber="76">Posterior margin of female tergum 8 convex. Posterior margin of female sternum 8 distinctly convex, without emargination.</paragraph>
<paragraph id="D3465800FFD539684D1DF900FCC5F90C" blockId="75.[264,1324,284,1870]" box="[312,813,1724,1750]" pageId="75" pageNumber="76">
Spermatheca as in
<figureCitation id="4BC24485FFD539684E30F900FDB2F90C" box="[533,602,1724,1750]" captionStart="FIGURES 1 ­ 7" captionStartId="5.[264,383,1751,1775]" captionTargetBox="[367,1205,490,1733]" captionTargetId="figure@5.[365,1207,490,1736]" captionTargetPageId="5" captionText="FIGURES 1 ­ 7. Mouthparts of Geostiba circellaris (Gravenhorst) (1 ­ 2, Moscow Region, Russia), G. carteriensis Pace (3 ­ 4, Tennessee), G. appalachigena Gusarov, sp. n. (5 ­ 6, paratype from West Virginia) and G. pluvigena Gusarov, sp. n. (7, paratype from North Carolina). 1, 3 labrum; 2, 4 epipharynx; 5 left mandible, dorsal view; 6 left mandible, ventral view; 7 right mandible, dorsal view. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/155702/files/figure.png" pageId="75" pageNumber="76">
Fig.
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</figureCitation>
in
<bibRefCitation id="B76825F1FFD539684E5AF900FCC2F90C" author="Assing" box="[639,810,1724,1750]" pageId="75" pageNumber="86" refString="Assing, V. (2001) A revision of the Turkish species of Geostiba Thomson 1858 and Tropimenelytron Pace 1983 (Coleoptera: Staphylinidae, Aleocharinae). Linzer biologische Beitrage, 33 (1), 137 - 185." type="journal article" year="2001">Assing (2001)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="9BE30B8BFFD5396F4D1DF95FFBABFE84" lastPageId="76" lastPageNumber="77" pageId="75" pageNumber="76" type="distribution">
<paragraph id="D3465800FFD5396F4D1DF95FFBABFE84" blockId="75.[264,1324,284,1870]" lastBlockId="76.[264,1323,284,350]" lastPageId="76" lastPageNumber="77" pageId="75" pageNumber="76">
<emphasis id="E18D8412FFD539684D1DF95FFE30F927" bold="true" box="[312,472,1763,1789]" pageId="75" pageNumber="76">Distribution.</emphasis>
<taxonomicName id="14F92383FFD539684DC0F958FD25F927" box="[485,717,1764,1789]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="circellaris">
<emphasis id="E18D8412FFD539684DC0F958FD25F927" box="[485,717,1764,1789]" italics="true" pageId="75" pageNumber="76">Geostiba circellaris</emphasis>
</taxonomicName>
is a widespread Palaearctic species. In the Nearctic region, it is known only from Newfoundland (
<bibRefCitation id="B76825F1FFD539684F06F8B0FC55F8FC" author="Muona" box="[803,957,1804,1830]" pageId="75" pageNumber="88" refString="Muona, J. (1984) Review of Palaearctic Aleocharinae also occurring in North America (Coleoptera: Staphylinidae). Entomologica scandinavica, 15, 227 - 231." type="journal article" year="1984">Muona 1984</bibRefCitation>
), where it was introduced from Europe. Contrary to the opinion expressed by
<bibRefCitation id="B76825F1FFD539684F47F888FB59F894" author="Lohse" box="[866,1201,1844,1870]" pageId="75" pageNumber="88" refString="Lohse, G. A. &amp; Smetana, A. (1988) Four new species of Geostiba Thomson from the Appalachian mountains of North Carolina, with a key to North American species and synonymic notes (Coleoptera: Staphylinidae: Aleocharinae). The Coleopterists Bulletin, 42 (3), 269 - 278." type="journal article" year="1988">Lohse and Smetana (1988)</bibRefCitation>
<taxonomicName id="14F92383FFD5396F48E4F888FDDAFEEC" authority="Casey, 1906" authorityName="Casey" authorityYear="1906" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" lastPageId="76" lastPageNumber="77" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="impressula">
<emphasis id="E18D8412FFD5396F48E4F888FE60FEEF" italics="true" lastPageId="76" lastPageNumber="77" pageId="75" pageNumber="76">Geostiba impressula</emphasis>
(
<bibRefCitation id="B76825F1FFD2396F4DBDFEA0FDC2FEEC" author="Casey" box="[408,554,284,310]" pageId="76" pageNumber="87" refString="Casey, T. L. (1906) Observations on the staphylinid groups Aleocharinae and Xantholinini, chiefly of America. Transactions of the Academy of Science of St. Louis, 16, 125 - 434." type="journal article" year="1906">Casey, 1906</bibRefCitation>
)
</taxonomicName>
is not a synonym of
<taxonomicName id="14F92383FFD2396F4F0CFEA0FC2CFEEF" box="[809,964,284,309]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="circellaris">
<emphasis id="E18D8412FFD2396F4F0CFEA0FC2CFEEF" box="[809,964,284,309]" italics="true" pageId="76" pageNumber="77">G. circellaris</emphasis>
</taxonomicName>
(see discussion for
<taxonomicName id="14F92383FFD2396F488BFEA0FED0FE87" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="impressula">
<emphasis id="E18D8412FFD2396F488BFEA0FED0FE87" italics="true" pageId="76" pageNumber="77">G. impressula</emphasis>
</taxonomicName>
).
<taxonomicName id="14F92383FFD2396F4D7CFEF8FDA9FE87" box="[345,577,324,349]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="circellaris">
<emphasis id="E18D8412FFD2396F4D7CFEF8FDA9FE87" box="[345,577,324,349]" italics="true" pageId="76" pageNumber="77">Geostiba circellaris</emphasis>
</taxonomicName>
is not known from western North
<collectingCountry id="ABEE1890FFD2396F4FF3FEF8FBD7FE84" box="[982,1087,324,350]" name="United States of America" pageId="76" pageNumber="77">America</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>