911 lines
119 KiB
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911 lines
119 KiB
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<document ID-CLB-Dataset="129873" ID-DOI="http://dx.doi.org/10.3897/zookeys.1119.85400" ID-GBIF-Dataset="418bd521-e4b9-4ee6-981f-a6f471eab287" ID-PMC="PMC9848625" ID-Pensoft-Pub="1313-2970-1119-1" ID-Pensoft-UUID="F158A7C5261D52B69288A62F3C777CAF" ID-PubMed="36762355" ID-ZooBank="A3BE91C6B79344E1A886A803BF104D8B" ModsDocID="1313-2970-1119-1" checkinTime="1661532673136" checkinUser="pensoft" docAuthor="Jirapatrasilp, Parin, Sutcharit, Chirasak & Panha, Somsak" docDate="2022" docId="38446D96E8785C33B44B278839E24F7B" docLanguage="en" docName="ZooKeys 1119: 1-115" docOrigin="ZooKeys 1119" docPubDate="2022-08-25" docSource="http://dx.doi.org/10.3897/zookeys.1119.85400" docTitle="Pupina artata Benson 1856" docType="treatment" docVersion="6" id="F158A7C5261D52B69288A62F3C777CAF" lastPageNumber="1" masterDocId="F158A7C5261D52B69288A62F3C777CAF" masterDocTitle="Annotated checklist of the operculated land snails from Thailand (Mollusca, Gastropoda, Caenogastropoda): the family Pupinidae, with descriptions of several new species and subspecies, and notes on classification of Pupina Vignard, 1829 and Pupinella Gray, 1850 from mainland Southeast Asia" masterLastPageNumber="115" masterPageNumber="1" pageNumber="1" updateTime="1732923707409" updateUser="ExternalLinkService">
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<mods:title id="D061255EEB9A2D0A9B1EA124CCC16512">Annotated checklist of the operculated land snails from Thailand (Mollusca, Gastropoda, Caenogastropoda): the family Pupinidae, with descriptions of several new species and subspecies, and notes on classification of Pupina Vignard, 1829 and Pupinella Gray, 1850 from mainland Southeast Asia</mods:title>
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<mods:namePart id="EDD6AB6701EFD38D545CE3531FCE0D5B">Jirapatrasilp, Parin</mods:namePart>
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<mods:affiliation id="900A92BC954B8E83509B80DF71E2DF22">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:namePart id="C5C40A0B67413F0CCC938D90C639BB34">Sutcharit, Chirasak</mods:namePart>
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<mods:affiliation id="BF8BFBDD23A0049BDCEDEA706796C520">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:nameIdentifier id="D91D6C1C641DED4B258C39D92670DF87" type="email">jirasak4@yahoo.com</mods:nameIdentifier>
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<mods:namePart id="3B8EA909A256BB315A7888132553FD16">Panha, Somsak</mods:namePart>
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<mods:affiliation id="E3612C6D466E999416E4E3DE437B51A3">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand & Academy of Science, The Royal Society of Thailand, Bangkok 10300, Thailand</mods:affiliation>
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<mods:nameIdentifier id="D23452212110819111571A98F06822F1" type="email">somsak.pan@chula.ac.th</mods:nameIdentifier>
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<mods:date id="86CA3EC77BC65D8014245D0AD73B4CB2">2022</mods:date>
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<mods:number id="7AC0211428454AF26D80DF3368752194">1119</mods:number>
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<mods:identifier id="D0BEFA73FAB4987D6890087207F61551" type="DOI">http://dx.doi.org/10.3897/zookeys.1119.85400</mods:identifier>
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<treatment id="38446D96E8785C33B44B278839E24F7B" ID-GBIF-Taxon="199839023" LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B" httpUri="http://treatment.plazi.org/id/38446D96E8785C33B44B278839E24F7B" lastPageNumber="1" pageId="0" pageNumber="1" scope_class="Gastropoda" scope_family="Pupinidae" scope_phylum="Mollusca" scope_subClass="Caenogastropoda" scope_subFamily="Pupininae" scope_superFamily="Cyclophoroidea">
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<subSubSection id="C6F938344E6C38AC07168623547CD52D" pageId="0" pageNumber="1" type="nomenclature">
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<paragraph id="27E08A8DF8182BEBFFCEC8AAFB996705" pageId="0" pageNumber="1">
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<taxonomicName id="504F7C9A198E7AC78CD1D034B83124A0" ID-CoL="7SWXB" LSID="38446D96-E878-5C33-B44B-278839E24F7B" authority="Benson, 1856" authorityName="Benson" authorityYear="1856" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina artata" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="artata">Pupina artata Benson, 1856</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="32B0B87D7DCE8B217CB83EEC774D01FB" pageId="0" pageNumber="1" type="description">
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<paragraph id="8CD00B20EB78AF4B63405F11F6B6130A" pageId="0" pageNumber="1">
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<figureCitation id="A013F50416F81CCAC5BEEC0F1D6DF275" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Live specimens of A, B Pollicaria mouhoti mouhoti: specimens A CUMZ 12166 and B CUMZ 12175 from Wang Daeng Cave, Phitsanulok C Pollicaria mouhoti monochroma, paratype CUMZ 1562 from Tam Pha Bing Temple, Loei D Tortulosa tortuosa, specimen CUMZ 12155 from Tham Suea Temple, Krabi E-H Pupina artata: specimens of E CUMZ 12006 from Pha Daeng Cave, Mae Hong Son F CUMZ 12008 from Tham Nam Pha Pha Ngam Temple, Lampang, and G, H CUMZ 12029 from Khao Tham Raet Temple, Chachoengsao showing the brown (G) and grey (H) shell morphs; All not to scale." figureDoi="10.3897/zookeys.1119.85400.figure8" httpUri="https://binary.pensoft.net/fig/735369" pageId="0" pageNumber="1">Figs 8E-H</figureCitation>
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<figureCitation id="35BE4688AC5F71DE08128B8A8287A2D3" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Umbilical, columellar and parietal views of A Pupinella mansuyi, specimen CUMZ 12148 from Pha Chu, Nan B Pupina artata from the Pupina artata species group, specimen CUMZ 12003 from Ban Ping Khong, Chiang Mai C Pupina godwinausteni sp. nov. from the Pupina arula species group, holotype CUMZ 12090 / 1 D Pupina aureola from the Pupina aureola species group, specimen CUMZ 12130 from Sra Morakot, Krabi." figureDoi="10.3897/zookeys.1119.85400.figure10" httpUri="https://binary.pensoft.net/fig/735371" pageId="0" pageNumber="1">, 10B</figureCitation>
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<figureCitation id="0C98C32C6C811F4C3352352D1F544CC1" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. External shell morphology of three mainland Southeast Asian Pupina species groups A Pupina artata from the Pupina artata species group, CUMZ 12003 from Ban Ping Khong, Chiang Mai B Pupina peguensis from the Pupina arula species group, CUMZ 12094 from Khao Tham Phra Temple, Chiang Rai C Pupina siamensis from the Pupina arula species group, CUMZ 12052 from Sri Thong Cave, Sra Keo, and D Pupina tchehelensis from the Pupina aureola species group, CUMZ 12136 from limestone mountain, Phang Nga. Red frames focus on the parietal tooth and posterior canal; blue frames focus on the curvature of the apertural lip when observed from lateral view." figureDoi="10.3897/zookeys.1119.85400.figure18" httpUri="https://binary.pensoft.net/fig/735379" pageId="0" pageNumber="1">, 18A</figureCitation>
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<figureCitation id="1F06A7FD7038827A3B2839676A447323" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Shells of Pupina artata species group from mainland Southeast Asia A-M Pupina artata A syntype UMZC I. 102960. A and specimens B CUMZ 12011 C CUMZ 12023 D CUMZ 12038 E CUMZ 12026 F CUMZ 12015 G CUMZ 12009 H CUMZ 12001 I CUMZ 12003 J NHMUK 1906.4.4.28 K CUMZ 12037 L CUMZ 12031, and M CUMZ 12018 N, O Pupina pallens N lectotype SMF 109951 and O specimen CUMZ 12042 P-R Pupina limitanea P, Q syntypes NHMUK 1903.7.1.2967 and R specimen CUMZ 12044 S Pupina billeti, holotype MNHN-IM- 2000 - 35841 T Pupina verneaui, syntype MNHN-IM- 2000 - 35843 U, V Pupina hungerfordiana, specimens NHMUK 91.3.14.686 - 7 W, X Pupina bensoni sp. nov. W holotype CUMZ 12045 / 1 and X paratype CUMZ 12046 / 1. Photo: H. Taylor, NHM (A, P, Q), P. Maestrati, MNHN (S, T)." figureDoi="10.3897/zookeys.1119.85400.figure21" httpUri="https://binary.pensoft.net/fig/735382" pageId="0" pageNumber="1">, 21A-M</figureCitation>
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<figureCitation id="F7F8C0074D1C7B6D96897DC1EB032802" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Pupina artata A syntype UMZC I. 102960. A from Moulmein B specimen NHMUK 1906.4.4.28 from Moulmein C specimen CUMZ 12001 from Khao Tham Phra Temple, Chiang Rai D specimen CUMZ 12003 from Ban Ping Khong, Chiang Mai E specimen CUMZ 12009 from Phu Sang Waterfall, Phayao, and F specimen CUMZ 12011 from Thep Sathaporn Temple, Nakhon Sawan. Photo: H. Taylor, NHM (A)." figureDoi="10.3897/zookeys.1119.85400.figure22" httpUri="https://binary.pensoft.net/fig/735383" pageId="0" pageNumber="1">, 22</figureCitation>
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<figureCitation id="E6D573A1FFC6803B4DAD0F410051B34D" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Pupina artata: specimens A CUMZ 12015 from Khao Wong Phrommachan Temple, Uthai Thani B CUMZ 12018 from Tham Khao Thalu Temple, Ratchaburi C CUMZ 12023 from Tham Khiriwong Temple, Prachub Kirikhan D CUMZ 12026 from Tham Khao Cha Ang On Temple, Chonburi E CUMZ 12031 from Tham Khao Chakan Temple, Sa Kaeo, and F CUMZ 12037 from Wua Ta Lap Island, Surat Thani." figureDoi="10.3897/zookeys.1119.85400.figure23" httpUri="https://binary.pensoft.net/fig/735384" pageId="0" pageNumber="1">, 23</figureCitation>
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<figureCitation id="743E25700BA65984D853354B4451E464" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. A Pupina artata, specimen CUMZ 12038 from Tham Suea Temple, Krabi B, C Pupina pallens B lectotype SMF 109951 and C specimen CUMZ 12042 from Suan Wiwek Bureau of Monks, Prachuap Khiri Khan D-F Pupina limitanea D, E syntypes NHMUK 1903.7.1.2967 from East of Burma & Siam and F specimen CUMZ 12044 from Mae Lana junction, Mae Hong Son G Pupina bensoni sp. nov., holotype CUMZ 12045 / 1. Photo: H. Taylor, NHM (D, E)." figureDoi="10.3897/zookeys.1119.85400.figure24" httpUri="https://binary.pensoft.net/fig/735385" pageId="0" pageNumber="1">, 24A</figureCitation>
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<figureCitation id="98F0FEC115094DEAE7334FDC16BDF277" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Live specimens of A-C Pupina artata: specimens A CUMZ 12020 from Buri Ratchawanaram Temple, Ratchaburi B CUMZ 12022 from Golden Dragon Cave, Ratchaburi, and C CUMZ 12037 from Wua Ta Lap Island, Surat Thani D, E Pupina bensoni sp. nov. D paratype CUMZ 12045 / 2 from Khao Wong Cave, Uthai Thani and E specimen CUMZ 12047 from Tham Namthip Bureau of Monks, Uthai Thani F, G Pupina peguensis: specimens F CUMZ 12050 from Chai Thong Wararam Temple, Nakhon Sawan and G CUMZ 12051 from Tham Saeng Wiset Bureau of Monks, Nakhon Sawan H Pupina siamensis, specimen CUMZ 12069 from Khao Chi Chan Buddha Image, Chonburi. All not to scale." figureDoi="10.3897/zookeys.1119.85400.figure25" httpUri="https://binary.pensoft.net/fig/735386" pageId="0" pageNumber="1">, 25A-C</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="652F1138498280D7F4BD941A97A77230" pageId="0" pageNumber="1" type="reference_group">
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<paragraph id="5C81C009A1E3CF0E22684AA648B69137" pageId="0" pageNumber="1">
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<taxonomicName id="C6A8B71B685E81C279458B2D9DFD7A86" authorityName="Benson" authorityYear="1856" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina artata" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="artata">Pupina artata</taxonomicName>
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Benson, 1856: 230. Type locality: Moulmein [Mawlamyine, Mawlamyine Township, Mawlamyine District, Mon State, Myanmar].
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<bibRefCitation id="7AE74440418A88D015E089D190431EAC" author="Theobald, W" journalOrPublisher="Journal of the Asiatic Society of Bengal" pageId="0" pageNumber="1" pagination="245 - 254" refId="B167" refString="Theobald, W, 1858. Notes on the distribution of some of the land and freshwater shells of India. Journal of the Asiatic Society of Bengal 26 (1857): 245 - 254" title="Notes on the distribution of some of the land and freshwater shells of India." volume="26" year="1858">Theobald 1858</bibRefCitation>
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[1857]: 247, 248. Pfeiffer 1860: 142, pl. 37, figs 10-12. Sowerby I 1866:
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<taxonomicName id="9C4B9DA223E803BC152EEAAAF901733A" authorityName="L. Pfeiffer" authorityYear="1853" class="Gastropoda" family="Pupinidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="family">Pupinidae</taxonomicName>
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, pl. 3 (pl. 265),
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<taxonomicName id="D3F4E144A953A168F2C53720CBB48B25" authorityName="Vignard" authorityYear="1829" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Pupina</taxonomicName>
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, figs 1, 2. Hanley and Theobald 1870: 4, pl. 7, fig. 5.
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<bibRefCitation id="EAE7D50694B335B84140758FF8174968" author="Stoliczka, F" journalOrPublisher="Journal of the Asiatic Society of Bengal" pageId="0" pageNumber="1" pagination="143 - 259" refId="B150" refString="Stoliczka, F, 1871. Notes on terrestrial Mollusca from the neighbourhood of Moulmein (Tenasserim Provinces), with descriptions of new species. Journal of the Asiatic Society of Bengal 40: 143 - 259" title="Notes on terrestrial Mollusca from the neighbourhood of Moulmein (Tenasserim Provinces), with descriptions of new species." volume="40" year="1871">Stoliczka 1871</bibRefCitation>
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: 151, 152.
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<bibRefCitation id="9E90D5E39EB2B30242D2B9C39C2C4ED5" DOI="https://doi.org/10.5962/bhl.title.11957" author="Nevill, G" journalOrPublisher="Indian Museum, Calcutta" pageId="0" pageNumber="1" refId="B104" refString="Nevill, G, 1878. Hand List of Mollusca in the Indian Museum, Calcutta. Part 1. Gastropoda. Pulmonata and Prosobranchia-Neurobranchia. Indian Museum, Calcutta, DOI: https://doi.org/10.5962/bhl.title.11957" title="Hand List of Mollusca in the Indian Museum, Calcutta. Part 1. Gastropoda. Pulmonata and Prosobranchia-Neurobranchia." url="https://doi.org/10.5962/bhl.title.11957" year="1878">Nevill 1878</bibRefCitation>
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: 299, 300, Ava [Mandalay Region, Myanmar]; Moulmein; Buket Pondong [Gunung Pondok, Perak State, Malaysia].
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<bibRefCitation id="3A11EA4AF18A1A5E28743FD94D2B0485" author="Reeve, L" journalOrPublisher="Journal of Systematic Palaeontology" pageId="0" pageNumber="1" refId="B139" refString="Reeve, L, 1878. Conchologia Iconica: Illustrations of the shells of molluscous animals. Volume 20, Monograph of the genus Pupinidae, pls 1-10. Lovell Reeve & Co, London, [without pagination]." title="Conchologia Iconica: Illustrations of the shells of molluscous animals. Volume 20, Monograph of the genus Pupinidae, pls 1 - 10. Lovell Reeve & Co, London, [without pagination]." year="1878">Reeve 1878</bibRefCitation>
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:
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<taxonomicName id="668C2E2E35AAB373E8157B231E373B55" authorityName="L. Pfeiffer" authorityYear="1853" class="Gastropoda" family="Pupinidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="family">Pupinidae</taxonomicName>
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, pl. 1, sp. 3.
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<bibRefCitation id="09239822DE6B50B20C5DBDC75DBC792D" author="Crosse, H" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="1" pagination="336 - 340" refId="B26" refString="Crosse, H, 1879. Faune malacologique de Perak (Indo-Chine). Journal de Conchyliologie 27: 336 - 340" title="Faune malacologique de Perak (Indo-Chine)." volume="27" year="1879">Crosse 1879</bibRefCitation>
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: 340. de Morgan 1885: 413, Boukit Pondong,
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<normalizedToken id="E7C5C9CD4CE3081ECD329A746A49CA02" originalValue="Pérak">Perak</normalizedToken>
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; Java [doubtful]; Moulmein; Lahat, Ipoh,
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<normalizedToken id="35EA92A2D4D9A2F8F44EBA1826C69CCD" originalValue="Gôping">Goping</normalizedToken>
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, Kinta [Perak State, Malaysia].
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<bibRefCitation id="B712CF3C7932AFBFE2F738F336A0BDF9" author="von Moellendorff, OF" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="1" pagination="146 - 156" refId="B192" refString="von Moellendorff, OF, 1894. On a collection of land-shells from the Samui Islands, Gulf of Siam. Proceedings of the Zoological Society of London 62: 146 - 156" title="On a collection of land-shells from the Samui Islands, Gulf of Siam." volume="62" year="1894">
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von
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<normalizedToken id="8AA76ABA74CED1245C021CE014C791F5" originalValue="Möllendorff">Moellendorff</normalizedToken>
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1894
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</bibRefCitation>
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: 155, the Samui Islands, Gulf of Siam [Samui Island, Surat Thani Province, Thailand].
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<bibRefCitation id="9A155B4D3746B2F3A16F8A81D75D3511" author="Godwin-Austen, HH" journalOrPublisher="Journal of Malacology" pageId="0" pageNumber="1" refId="B51" refString="Godwin-Austen, HH, 1897. Land and freshwater mollusca of India, including South Arabia, Baluchistan, Afghanistan, Kashmir, Nepal, Burma, Pegu, Tenasserim, Malaya Peninsula, Ceylon and other islands of the Indian Ocean. Vol. 2, Part 7: 1-46." title="Land and freshwater mollusca of India, including South Arabia, Baluchistan, Afghanistan, Kashmir, Nepal, Burma, Pegu, Tenasserim, Malaya Peninsula, Ceylon and other islands of the Indian Ocean. Vol. 2, Part 7: 1 - 46." year="1897">Godwin-Austen 1897</bibRefCitation>
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: 38, 39, pl. 69, fig. 6, 6a, b.
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<bibRefCitation id="0AE1FC41555A0EC005A80666F72E47C2" author="Fischer, H" editor="Leroux, E" journalOrPublisher="E. Leroux, Paris" pageId="0" pageNumber="1" pagination="390 - 450" refId="B45" refString="Fischer, H, Dautzenberg, P, 1904. Catalogue des mollusques terrestres et fluviatiles de ľIndo-Chine orientale cites jusqu' a ce jour. In: Leroux, E, Ed., Mission Pavie Indo-Chine 1879-1895, Etudes diverses III Recherches sur l'histoire naturelle de l'Indo-Chine orientale. E. Leroux, Paris: 390 - 450" title="Catalogue des mollusques terrestres et fluviatiles de ľIndo-Chine orientale cites jusqu' a ce jour." volumeTitle="Mission Pavie Indo-Chine 1879 - 1895, Etudes diverses III Recherches sur l'histoire naturelle de l'Indo-Chine orientale." year="1904">Fischer and Dautzenberg 1904</bibRefCitation>
|
|
: 431, Ile Samui, golfe de Siam [Samui Island, Surat Thani Province, Thailand].
|
|
<bibRefCitation id="CB8F4954FE52127719243AD2E03F9AAB" author="van Benthem Jutting, WSS" journalOrPublisher="Basteria" pageId="0" pageNumber="1" pagination="10 - 20" refId="B181" refString="van Benthem Jutting, WSS, 1960. Some notes on land and freshwater Mollusca of Malaya. Basteria 24: 10 - 20" title="Some notes on land and freshwater Mollusca of Malaya." volume="24" year="1960">van Benthem Jutting 1960</bibRefCitation>
|
|
: 12, a hill near the hot springs, near Tandjong Rambutan, N.E. of Ipoh, Perak.
|
|
<bibRefCitation id="276226AE972DB7DA3D7A4A4199F5A9E2" author="Solem, A" journalOrPublisher="Spolia Zoologica Musei Hauniensis" pageId="0" pageNumber="1" pagination="1 - 110" refId="B146" refString="Solem, A, 1966. Some non-marine mollusks from Thailand, with notes on classification of the Helicarionidae. Spolia Zoologica Musei Hauniensis 24: 1 - 110" title="Some non-marine mollusks from Thailand, with notes on classification of the Helicarionidae." volume="24" year="1966">Solem 1966</bibRefCitation>
|
|
: 12, Chieng Dao, Doi Sutep [Chiang Dao District and Doi Suthep Mountain, Chiang Mai Province, Thailand].
|
|
<bibRefCitation id="1FE19AEFB8C5432566F3FE338A7B402B" author="Davison, GWH" journalOrPublisher="Malayan Nature Journal" pageId="0" pageNumber="1" pagination="233 - 248" refId="B31" refString="Davison, GWH, 1995. The terrestrial molluscan fauna of Temengor Forest Reserve, Hulu Perak, Malaysia. In: Belum Expedition, Temengor, 1993-1994. Malayan Nature Journal 48: 233 - 248" title="The terrestrial molluscan fauna of Temengor Forest Reserve, Hulu Perak, Malaysia. In: Belum Expedition, Temengor, 1993 - 1994." volume="48" year="1995">Davison 1995</bibRefCitation>
|
|
: 236, 237, limestone island C, Temengor dam, Perak, Malaysia.
|
|
<bibRefCitation id="0142F012830E794944EB831229B02917" author="Chan, S-Y" journalOrPublisher="The Papustyla" pageId="0" pageNumber="1" pagination="1 - 2" refId="B18" refString="Chan, S-Y, 1998b. A brief collecting trip to Perak, West Malaysia, part two. The Papustyla 12: 1 - 2" title="A brief collecting trip to Perak, West Malaysia, part two." volume="12" year="1998 b">Chan 1998b</bibRefCitation>
|
|
: 2, Ipoh, Perak.
|
|
<bibRefCitation id="3D5047681C7AA857B0CA7E17BAF7D83A" author="Maassen, WJM" journalOrPublisher="Proceedings of the United States National Museum" pageId="0" pageNumber="1" refId="B87" refString="Maassen, WJM, 2001. A preliminary checklist of the non-marine molluscs of West-Malaysia, A handlist. De Kreukel (Supplement): 1-155." title="A preliminary checklist of the non-marine molluscs of West-Malaysia, A handlist. De Kreukel (Supplement): 1 - 155." year="2001">Maassen 2001</bibRefCitation>
|
|
: 39, 40.
|
|
<bibRefCitation id="59604C91FFAE7C763DB0B646867321CB" author="Bavay, A" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="1" refId="B8" refString="BEDO (2017) Land Snails: Checklist of Molluscan Biodiversity in Thailand.BEDO, Bangkok, 300 pp. [in Thai]" year="1904">BEDO 2017</bibRefCitation>
|
|
: 87.
|
|
<bibRefCitation id="10F47FA730B2E1BA2C3BD9C5B5BC8B59" author="Sutcharit, C" journalOrPublisher="Zino Publishing, Bangkok" pageId="0" pageNumber="1" refId="B156" refString="Sutcharit, C, Tongkerd, P, Panha, S, 2018. Land snails: The invaluable bio-resources for the Kingdom of Thailand. Zino Publishing, Bangkok" title="Land snails: The invaluable bio-resources for the Kingdom of Thailand." year="2018">Sutcharit et al. 2018</bibRefCitation>
|
|
: fig. 5-13d.
|
|
</paragraph>
|
|
<paragraph id="8C17A311924501D256E80139CB1A2D7B" pageId="0" pageNumber="1">
|
|
<taxonomicName id="ADAD35B35B05C25EE50472DFA5C09994" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina artata subsp. var. var. blanfordiana" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="variety" species="artata" subSpecies="var." variety="blanfordiana">Pupina artata var. blanfordiana</taxonomicName>
|
|
Nevill, 1878: 300. Type locality: Thyet Myo [Thayetmyo, Magway Region, Myanmar]; Akoutong [Akauk Taung, Pyay District, Bago Region, Myanmar]; Kamah Hill, Tongoop, & c., Arakan [Toungup, Thandwe District, Rakhine State, Myanmar]; Prome [Pyay, Bago Region, Myanmar].
|
|
</paragraph>
|
|
<paragraph id="4D192853C05B57BACBC5FEEA27700871" pageId="0" pageNumber="1">
|
|
<taxonomicName id="9E6A90DC774795232315444A028FDC81" authorityName="Benson" authorityYear="1860" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina peguensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peguensis">Pupina peguensis</taxonomicName>
|
|
[non Benson]-
|
|
<bibRefCitation id="C9EF3B93F41A80E3DCDB5AEEBDAAF2D3" author="Godwin-Austen, HH" journalOrPublisher="Journal of Malacology" pageId="0" pageNumber="1" refId="B51" refString="Godwin-Austen, HH, 1897. Land and freshwater mollusca of India, including South Arabia, Baluchistan, Afghanistan, Kashmir, Nepal, Burma, Pegu, Tenasserim, Malaya Peninsula, Ceylon and other islands of the Indian Ocean. Vol. 2, Part 7: 1-46." title="Land and freshwater mollusca of India, including South Arabia, Baluchistan, Afghanistan, Kashmir, Nepal, Burma, Pegu, Tenasserim, Malaya Peninsula, Ceylon and other islands of the Indian Ocean. Vol. 2, Part 7: 1 - 46." year="1897">Godwin-Austen 1897</bibRefCitation>
|
|
: 40, pl. 69, fig. 3, 3a-d, Kama on the right bank of the Irrawaddy, Pegu [Kamma Township, Thayet District, Magway Region, Myanmar].
|
|
<bibRefCitation id="B73B1E0A5DF1EDF2C042349E032EB247" author="Bavay, A" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="1" refId="B8" refString="BEDO (2017) Land Snails: Checklist of Molluscan Biodiversity in Thailand.BEDO, Bangkok, 300 pp. [in Thai]" year="1904">BEDO 2017</bibRefCitation>
|
|
: 93.
|
|
</paragraph>
|
|
<paragraph id="CE034DD6BCD5871EF4EAF0356392C1C8" pageId="0" pageNumber="1">
|
|
<taxonomicName id="71F3671F8E068951318B54FC1C39B7A7" authorityName="Benson" authorityYear="1856" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina (Tylotoechus) artata" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="artata" subGenus="Tylotoechus">Pupina (Tylotoechus) artata</taxonomicName>
|
|
-
|
|
<bibRefCitation id="F1E6A1C6A80A6794904163F6CD4F0ACE" author="Kobelt, W" journalOrPublisher="Friedlaender und Sohn, Berlin" pageId="0" pageNumber="1" refId="B76" refString="Kobelt, W, 1902. Cyclophoridae. Das Tierreich. R. Friedlaender und Sohn, Berlin" title="Cyclophoridae. Das Tierreich. R." year="1902">Kobelt 1902</bibRefCitation>
|
|
: 306, 307.
|
|
<bibRefCitation id="12BDCF774D6F248A2F23DA29475F811D" author="Gude, GK" journalOrPublisher="Taylor and Francis, London" pageId="0" pageNumber="1" refId="B60" refString="Gude, GK, 1921. The Fauna of British India including Ceylon and Burma. Mollusca-- III. Land Operculates. Taylor and Francis, London" title="The Fauna of British India including Ceylon and Burma. Mollusca- - III. Land Operculates." year="1921">Gude 1921</bibRefCitation>
|
|
: 193.
|
|
<bibRefCitation id="D25FBD423C8B780534C453D95405C67B" author="Laidlaw, FF" journalOrPublisher="Journal of the Malaysian Branch of the Royal Asiatic Society" pageId="0" pageNumber="1" pagination="25 - 37" refId="B82" refString="Laidlaw, FF, 1928. A list of the land and fresh-water mollusca of the Malay Peninsula with notes. Journal of the Malaysian Branch of the Royal Asiatic Society 6: 25 - 37" title="A list of the land and fresh-water mollusca of the Malay Peninsula with notes." volume="6" year="1928">Laidlaw 1928</bibRefCitation>
|
|
: 33.
|
|
</paragraph>
|
|
<paragraph id="F76C6350BBC00E584E010C8D61E4D7D5" pageId="0" pageNumber="1">
|
|
<taxonomicName id="E8511A72B3670B611EC6F7B8701FEDC7" authorityName="Benson" authorityYear="1856" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina (Pupina) artata" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="artata" subGenus="Pupina">Pupina (Pupina) artata</taxonomicName>
|
|
-
|
|
<bibRefCitation id="DD3803BF7970A03C73657291EA1DE123" author="Hemmen, J" journalOrPublisher="Schriften zur Malakozoologie" pageId="0" pageNumber="1" pagination="35 - 70" refId="B66" refString="Hemmen, J, Hemmen, C, 2001. Aktualisierte Liste der terrestrischen Gastropoden Thailands. Schriften zur Malakozoologie 18: 35 - 70" title="Aktualisierte Liste der terrestrischen Gastropoden Thailands." volume="18" year="2001">Hemmen and Hemmen 2001</bibRefCitation>
|
|
: 39.
|
|
</paragraph>
|
|
<paragraph id="45F9187A501FFA895B43B7B78B9D3B2B" pageId="0" pageNumber="1">
|
|
<taxonomicName id="13F2AD8465B09599D08FE110F67AE934" authorityName="Theobald" authorityYear="1864" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina blanfordi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="blanfordi">Pupina blanfordi</taxonomicName>
|
|
[non Theobald]-
|
|
<bibRefCitation id="4E469811C91541BDD5DDC126E7B5EA03" author="Bavay, A" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="1" refId="B8" refString="BEDO (2017) Land Snails: Checklist of Molluscan Biodiversity in Thailand.BEDO, Bangkok, 300 pp. [in Thai]" year="1904">BEDO 2017</bibRefCitation>
|
|
: 89.
|
|
</paragraph>
|
|
<paragraph id="CEA925568641130571E1733568DD921F" pageId="0" pageNumber="1">
|
|
<taxonomicName id="5BF2779F94BC14DFDE16D7018A544804" authorityName="Godwin-Austen" authorityYear="1897" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina limitanea" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="limitanea">Pupina limitanea</taxonomicName>
|
|
[non Godwin-Austen]-
|
|
<bibRefCitation id="DE7543AB80CC64255E8E3AA70E2A13FF" author="Bavay, A" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="1" refId="B8" refString="BEDO (2017) Land Snails: Checklist of Molluscan Biodiversity in Thailand.BEDO, Bangkok, 300 pp. [in Thai]" year="1904">BEDO 2017</bibRefCitation>
|
|
: 90.
|
|
<bibRefCitation id="37D17042266FD6D8F856656147A11766" author="Sutcharit, C" journalOrPublisher="Zino Publishing, Bangkok" pageId="0" pageNumber="1" refId="B156" refString="Sutcharit, C, Tongkerd, P, Panha, S, 2018. Land snails: The invaluable bio-resources for the Kingdom of Thailand. Zino Publishing, Bangkok" title="Land snails: The invaluable bio-resources for the Kingdom of Thailand." year="2018">Sutcharit et al. 2018</bibRefCitation>
|
|
: fig. 5-13g.
|
|
</paragraph>
|
|
<paragraph id="FD9E45769D222C27FA2DFFF9F96D5CBC" pageId="0" pageNumber="1">
|
|
<taxonomicName id="0FA51F022EB5198AACF10A95345419C1" authorityName="Vignard" authorityYear="1829" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Pupina</taxonomicName>
|
|
sp.-
|
|
<bibRefCitation id="0BF3220A6CDE2729FCAD6857CFE27084" author="Sutcharit, C" journalOrPublisher="Zino Publishing, Bangkok" pageId="0" pageNumber="1" refId="B156" refString="Sutcharit, C, Tongkerd, P, Panha, S, 2018. Land snails: The invaluable bio-resources for the Kingdom of Thailand. Zino Publishing, Bangkok" title="Land snails: The invaluable bio-resources for the Kingdom of Thailand." year="2018">Sutcharit et al. 2018</bibRefCitation>
|
|
: fig. 5-11a.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="802B84ED427020A4D75A080FABA880B0" pageId="0" pageNumber="1" type="type material">
|
|
<paragraph id="90972B3AE857491B987A04F95F324BE3" pageId="0" pageNumber="1">Type material examined.</paragraph>
|
|
<paragraph id="57D83194A19852602EAF800958B4BF4A" pageId="0" pageNumber="1">
|
|
<emphasis id="F0B41E8972D0E39D7B1A3D5B227C8CAD" bold="true" italics="true" pageId="0" pageNumber="1">Syntype</emphasis>
|
|
UMZC I.102960.A (1 shell; Figs
|
|
<figureCitation id="EA47D68C479F335E3466E418E4A4CB37" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Shells of Pupina artata species group from mainland Southeast Asia A-M Pupina artata A syntype UMZC I. 102960. A and specimens B CUMZ 12011 C CUMZ 12023 D CUMZ 12038 E CUMZ 12026 F CUMZ 12015 G CUMZ 12009 H CUMZ 12001 I CUMZ 12003 J NHMUK 1906.4.4.28 K CUMZ 12037 L CUMZ 12031, and M CUMZ 12018 N, O Pupina pallens N lectotype SMF 109951 and O specimen CUMZ 12042 P-R Pupina limitanea P, Q syntypes NHMUK 1903.7.1.2967 and R specimen CUMZ 12044 S Pupina billeti, holotype MNHN-IM- 2000 - 35841 T Pupina verneaui, syntype MNHN-IM- 2000 - 35843 U, V Pupina hungerfordiana, specimens NHMUK 91.3.14.686 - 7 W, X Pupina bensoni sp. nov. W holotype CUMZ 12045 / 1 and X paratype CUMZ 12046 / 1. Photo: H. Taylor, NHM (A, P, Q), P. Maestrati, MNHN (S, T)." figureDoi="10.3897/zookeys.1119.85400.figure21" httpUri="https://binary.pensoft.net/fig/735382" pageId="0" pageNumber="1">21A</figureCitation>
|
|
,
|
|
<figureCitation id="75571A76832620686720EE1F78D4E768" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Pupina artata A syntype UMZC I. 102960. A from Moulmein B specimen NHMUK 1906.4.4.28 from Moulmein C specimen CUMZ 12001 from Khao Tham Phra Temple, Chiang Rai D specimen CUMZ 12003 from Ban Ping Khong, Chiang Mai E specimen CUMZ 12009 from Phu Sang Waterfall, Phayao, and F specimen CUMZ 12011 from Thep Sathaporn Temple, Nakhon Sawan. Photo: H. Taylor, NHM (A)." figureDoi="10.3897/zookeys.1119.85400.figure22" httpUri="https://binary.pensoft.net/fig/735383" pageId="0" pageNumber="1">22A</figureCitation>
|
|
) from the R. McAndrew collection, labelled "Bens. col., Moulmein".
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="775850FB8CDADCB2CF6844F853842623" pageId="0" pageNumber="1" type="materials_examined">
|
|
<paragraph id="CCF4C8615B587A130758DC9463991C1A" pageId="0" pageNumber="1">Other material examined.</paragraph>
|
|
<paragraph id="81B54C3E8B23FD79B04825A546309A14" pageId="0" pageNumber="1">
|
|
<materialsCitation id="4CC77995CE327E16F29D22FAF7DCD8D0" collectingDate="2008-01-09" collectorName="Figs, Moulmein" country="Myanmar" county="Chiang Rai Province" location="Khao Tham Phra Temple" municipality="Mueang Chiang Rai District" specimenCount="1">
|
|
NHMUK 1906.4.4.28 (6 shells;
|
|
<collectorName id="1F4373C6618BCB203AC325DB8F85DFE3">Figs</collectorName>
|
|
<figureCitation id="B8B4C45E495D287F191D73ED02FA2EA5" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Shells of Pupina artata species group from mainland Southeast Asia A-M Pupina artata A syntype UMZC I. 102960. A and specimens B CUMZ 12011 C CUMZ 12023 D CUMZ 12038 E CUMZ 12026 F CUMZ 12015 G CUMZ 12009 H CUMZ 12001 I CUMZ 12003 J NHMUK 1906.4.4.28 K CUMZ 12037 L CUMZ 12031, and M CUMZ 12018 N, O Pupina pallens N lectotype SMF 109951 and O specimen CUMZ 12042 P-R Pupina limitanea P, Q syntypes NHMUK 1903.7.1.2967 and R specimen CUMZ 12044 S Pupina billeti, holotype MNHN-IM- 2000 - 35841 T Pupina verneaui, syntype MNHN-IM- 2000 - 35843 U, V Pupina hungerfordiana, specimens NHMUK 91.3.14.686 - 7 W, X Pupina bensoni sp. nov. W holotype CUMZ 12045 / 1 and X paratype CUMZ 12046 / 1. Photo: H. Taylor, NHM (A, P, Q), P. Maestrati, MNHN (S, T)." figureDoi="10.3897/zookeys.1119.85400.figure21" httpUri="https://binary.pensoft.net/fig/735382" pageId="0" pageNumber="1">21J</figureCitation>
|
|
,
|
|
<figureCitation id="464A9EF9B127B29A070B9CA1562AE33E" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Pupina artata A syntype UMZC I. 102960. A from Moulmein B specimen NHMUK 1906.4.4.28 from Moulmein C specimen CUMZ 12001 from Khao Tham Phra Temple, Chiang Rai D specimen CUMZ 12003 from Ban Ping Khong, Chiang Mai E specimen CUMZ 12009 from Phu Sang Waterfall, Phayao, and F specimen CUMZ 12011 from Thep Sathaporn Temple, Nakhon Sawan. Photo: H. Taylor, NHM (A)." figureDoi="10.3897/zookeys.1119.85400.figure22" httpUri="https://binary.pensoft.net/fig/735383" pageId="0" pageNumber="1">22B</figureCitation>
|
|
) from
|
|
<collectorName id="883B709F329D042534C51AD6BBA01A1F">Moulmein</collectorName>
|
|
,
|
|
<collectingCountry id="E4D86EBC9719B19DAF9AD67E977FF705" name="Myanmar">Myanmar</collectingCountry>
|
|
. CUMZ 12001 (7 shells; Figs
|
|
<figureCitation id="5FFB3BE1645FEBF3DF9D6BC4D36D8CBB" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Shells of Pupina artata species group from mainland Southeast Asia A-M Pupina artata A syntype UMZC I. 102960. A and specimens B CUMZ 12011 C CUMZ 12023 D CUMZ 12038 E CUMZ 12026 F CUMZ 12015 G CUMZ 12009 H CUMZ 12001 I CUMZ 12003 J NHMUK 1906.4.4.28 K CUMZ 12037 L CUMZ 12031, and M CUMZ 12018 N, O Pupina pallens N lectotype SMF 109951 and O specimen CUMZ 12042 P-R Pupina limitanea P, Q syntypes NHMUK 1903.7.1.2967 and R specimen CUMZ 12044 S Pupina billeti, holotype MNHN-IM- 2000 - 35841 T Pupina verneaui, syntype MNHN-IM- 2000 - 35843 U, V Pupina hungerfordiana, specimens NHMUK 91.3.14.686 - 7 W, X Pupina bensoni sp. nov. W holotype CUMZ 12045 / 1 and X paratype CUMZ 12046 / 1. Photo: H. Taylor, NHM (A, P, Q), P. Maestrati, MNHN (S, T)." figureDoi="10.3897/zookeys.1119.85400.figure21" httpUri="https://binary.pensoft.net/fig/735382" pageId="0" pageNumber="1">21H</figureCitation>
|
|
,
|
|
<figureCitation id="7506D2A8B7D5422CB429D784708A29DC" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Pupina artata A syntype UMZC I. 102960. A from Moulmein B specimen NHMUK 1906.4.4.28 from Moulmein C specimen CUMZ 12001 from Khao Tham Phra Temple, Chiang Rai D specimen CUMZ 12003 from Ban Ping Khong, Chiang Mai E specimen CUMZ 12009 from Phu Sang Waterfall, Phayao, and F specimen CUMZ 12011 from Thep Sathaporn Temple, Nakhon Sawan. Photo: H. Taylor, NHM (A)." figureDoi="10.3897/zookeys.1119.85400.figure22" httpUri="https://binary.pensoft.net/fig/735383" pageId="0" pageNumber="1">22C</figureCitation>
|
|
) from
|
|
<location id="F3458A76246BD6F4E33E000B48596955" LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:F3458A76246BD6F4E33E000B48596955" country="Myanmar" county="Chiang Rai Province" municipality="Mueang Chiang Rai District" name="Khao Tham Phra Temple">Khao Tham Phra Temple</location>
|
|
,
|
|
<collectingMunicipality id="D58D2D849D1B2FDB5D68A97B2052A3A2">Mueang Chiang Rai District</collectingMunicipality>
|
|
,
|
|
<collectingCounty id="E281309BA224E63DA87C1D05DA5917FB">Chiang Rai Province</collectingCounty>
|
|
,
|
|
<collectingDate id="8D4A887A3DAFD493829BD2CB8E308F23" value="2008-01-09">9 Jan. 2008</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="D95EDB07D6139A800822D9154AA48736" collectingDate="2015-10-23" country="Myanmar" county="Chiang Rai Province" location="Luang Cave" municipality="Mae Sai District" specimenCount="1">
|
|
CUMZ 12002 (1 shell) from
|
|
<location id="75DF4B534B3BCA50C1E226E6EEC83D4A" LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:75DF4B534B3BCA50C1E226E6EEC83D4A" country="Myanmar" county="Chiang Rai Province" municipality="Mae Sai District" name="Luang Cave">Luang Cave</location>
|
|
,
|
|
<collectingMunicipality id="7AD067340F30334806F546A5B60EBF49">Mae Sai District</collectingMunicipality>
|
|
,
|
|
<collectingCounty id="E58B8CCA7DE1A516AD94927BA9F47274">Chiang Rai Province</collectingCounty>
|
|
,
|
|
<collectingDate id="3428FFEA8509446362992D9F074BCD8E" value="2015-10-23">23 Oct. 2015</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="C7E864169477F9703FA3135203FC5575" collectingDate="2008-10-08" collectorName="Figs" country="Myanmar" county="Chiang Mai Province" location="Ban Ping Khong" municipality="Chiang Dao District" specimenCount="1">
|
|
CUMZ 12003 (21 shells;
|
|
<collectorName id="4A3C99D291780B5789C21F852E9CC2B7">Figs</collectorName>
|
|
<figureCitation id="576DD9EA02382585D7FFE8EC71A1F435" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Umbilical, columellar and parietal views of A Pupinella mansuyi, specimen CUMZ 12148 from Pha Chu, Nan B Pupina artata from the Pupina artata species group, specimen CUMZ 12003 from Ban Ping Khong, Chiang Mai C Pupina godwinausteni sp. nov. from the Pupina arula species group, holotype CUMZ 12090 / 1 D Pupina aureola from the Pupina aureola species group, specimen CUMZ 12130 from Sra Morakot, Krabi." figureDoi="10.3897/zookeys.1119.85400.figure10" httpUri="https://binary.pensoft.net/fig/735371" pageId="0" pageNumber="1">10B</figureCitation>
|
|
,
|
|
<figureCitation id="5A56CA2B7C455B3EBBF5297C4421FA8F" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. External shell morphology of three mainland Southeast Asian Pupina species groups A Pupina artata from the Pupina artata species group, CUMZ 12003 from Ban Ping Khong, Chiang Mai B Pupina peguensis from the Pupina arula species group, CUMZ 12094 from Khao Tham Phra Temple, Chiang Rai C Pupina siamensis from the Pupina arula species group, CUMZ 12052 from Sri Thong Cave, Sra Keo, and D Pupina tchehelensis from the Pupina aureola species group, CUMZ 12136 from limestone mountain, Phang Nga. Red frames focus on the parietal tooth and posterior canal; blue frames focus on the curvature of the apertural lip when observed from lateral view." figureDoi="10.3897/zookeys.1119.85400.figure18" httpUri="https://binary.pensoft.net/fig/735379" pageId="0" pageNumber="1">18A</figureCitation>
|
|
,
|
|
<figureCitation id="F0123300A80EE016C31F057B90F555C5" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Shells of Pupina artata species group from mainland Southeast Asia A-M Pupina artata A syntype UMZC I. 102960. A and specimens B CUMZ 12011 C CUMZ 12023 D CUMZ 12038 E CUMZ 12026 F CUMZ 12015 G CUMZ 12009 H CUMZ 12001 I CUMZ 12003 J NHMUK 1906.4.4.28 K CUMZ 12037 L CUMZ 12031, and M CUMZ 12018 N, O Pupina pallens N lectotype SMF 109951 and O specimen CUMZ 12042 P-R Pupina limitanea P, Q syntypes NHMUK 1903.7.1.2967 and R specimen CUMZ 12044 S Pupina billeti, holotype MNHN-IM- 2000 - 35841 T Pupina verneaui, syntype MNHN-IM- 2000 - 35843 U, V Pupina hungerfordiana, specimens NHMUK 91.3.14.686 - 7 W, X Pupina bensoni sp. nov. W holotype CUMZ 12045 / 1 and X paratype CUMZ 12046 / 1. Photo: H. Taylor, NHM (A, P, Q), P. Maestrati, MNHN (S, T)." figureDoi="10.3897/zookeys.1119.85400.figure21" httpUri="https://binary.pensoft.net/fig/735382" pageId="0" pageNumber="1">21I</figureCitation>
|
|
,
|
|
<figureCitation id="9F387D6C64511C49A5DCFB15B1824B98" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Pupina artata A syntype UMZC I. 102960. A from Moulmein B specimen NHMUK 1906.4.4.28 from Moulmein C specimen CUMZ 12001 from Khao Tham Phra Temple, Chiang Rai D specimen CUMZ 12003 from Ban Ping Khong, Chiang Mai E specimen CUMZ 12009 from Phu Sang Waterfall, Phayao, and F specimen CUMZ 12011 from Thep Sathaporn Temple, Nakhon Sawan. Photo: H. Taylor, NHM (A)." figureDoi="10.3897/zookeys.1119.85400.figure22" httpUri="https://binary.pensoft.net/fig/735383" pageId="0" pageNumber="1">22D</figureCitation>
|
|
) from
|
|
<location id="F926A1F50087751CCDF235571B4F42B5" LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:F926A1F50087751CCDF235571B4F42B5" country="Myanmar" county="Chiang Mai Province" municipality="Chiang Dao District" name="Ban Ping Khong">Ban Ping Khong</location>
|
|
,
|
|
<collectingMunicipality id="4D3CD7C9E1CE339DCCEE13BE56971BDE">Chiang Dao District</collectingMunicipality>
|
|
,
|
|
<collectingCounty id="87A91FFC81AFBC17055159C91C43418E">Chiang Mai Province</collectingCounty>
|
|
,
|
|
<collectingDate id="0F4FAB165D1252189B781CBB8314E78C" value="2008-10-08">8 Oct. 2008</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="BD40561D64970EDA42AA1C75B9914F99" collectingDate="2012-11-21" collectorName="Ban Ping Khong" country="Myanmar" location="Chiang Dao District" municipality="Chiang Mai Province" specimenCount="1">
|
|
CUMZ 12193 (4 shells) from
|
|
<collectorName id="A807FCEB809CB350E8102D63EF437408">Ban Ping Khong</collectorName>
|
|
,
|
|
<location id="2E305503310D178ADC931B14CC875208" LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:2E305503310D178ADC931B14CC875208" country="Myanmar" municipality="Chiang Mai Province" name="Chiang Dao District">Chiang Dao District</location>
|
|
,
|
|
<collectingMunicipality id="97FBB2BCB2C293E6FE4245C2EF4BF21C">Chiang Mai Province</collectingMunicipality>
|
|
,
|
|
<collectingDate id="B44D38E9FAFFE20FB41AA54D0E8A24C9" value="2012-11-21">21 Nov. 2012</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="810DFF9D62ADAA09F5D28D82E7DFE4A3" collectingDate="2015-10-25" country="Myanmar" county="Chiang Mai Province" location="Chiang Dao Cave" municipality="Chiang Dao District" specimenCount="1">
|
|
CUMZ 12190 (3 shells) from
|
|
<location id="BA4500FD7862A675F0A4E6735A73BF34" LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:BA4500FD7862A675F0A4E6735A73BF34" country="Myanmar" county="Chiang Mai Province" municipality="Chiang Dao District" name="Chiang Dao Cave">Chiang Dao Cave</location>
|
|
,
|
|
<collectingMunicipality id="CA167335BBB4687AF0624C7165EF9DEC">Chiang Dao District</collectingMunicipality>
|
|
,
|
|
<collectingCounty id="07C83C90E40F76A6F8CA8661C242B28F">Chiang Mai Province</collectingCounty>
|
|
,
|
|
<collectingDate id="86E0ABA0E8B58E2AD5043E9988D5DD4C" value="2015-10-25">25 Oct. 2015</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="297040213CACFECA81791BA0A576CA88" collectingDate="2017-10-08" country="Myanmar" county="Chiang Mai Province" location="Bua Tong Cave" municipality="Mae Tang District" specimenCount="4">
|
|
CUMZ 12004 (
|
|
<specimenCount id="C2D6328648848F7AE779729A1D3DFD41" type="generic">4 specimens</specimenCount>
|
|
in ethanol) from
|
|
<location id="8382F5FE09BCAD476EB39D11BFA179EB" LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:8382F5FE09BCAD476EB39D11BFA179EB" country="Myanmar" county="Chiang Mai Province" municipality="Mae Tang District" name="Bua Tong Cave">Bua Tong Cave</location>
|
|
,
|
|
<collectingMunicipality id="5F6416F890582AAF42CBD126294D4948">Mae Tang District</collectingMunicipality>
|
|
,
|
|
<collectingCounty id="493AB13C49805059E6C69CE2B118B448">Chiang Mai Province</collectingCounty>
|
|
,
|
|
<collectingDate id="EC10C8F73B4B49F55E1BB15FFEF2CA0C" value="2017-10-08">8 Oct. 2017</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="E222168EE29A8702C9C200016A1C9426" collectingDate="2015-10-24" collectorName="Doi Ang Khang" country="Myanmar" location="Fang District" municipality="Chiang Mai Province" specimenCount="1">
|
|
CUMZ 12168 (4 shells) from
|
|
<collectorName id="A29CB3EFB05A4983C1E453ABBA39C969">Doi Ang Khang</collectorName>
|
|
,
|
|
<location id="F3BF56D495AEF4540AC419134DDB906B" LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:F3BF56D495AEF4540AC419134DDB906B" country="Myanmar" municipality="Chiang Mai Province" name="Fang District">Fang District</location>
|
|
,
|
|
<collectingMunicipality id="D26CF29603BAE8B10D9691862717B7EA">Chiang Mai Province</collectingMunicipality>
|
|
,
|
|
<collectingDate id="B95B52538C555410BBF355FFC71DA88E" value="2015-10-24">24 Oct. 2015</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="3F22033547F7D0630B85A8776A18471B" collectingDate="2015-01-18" country="Myanmar" county="Mae Hong Son Province" location="Pha Daeng Cave" municipality="Mueang Mae Hong Son District" specimenCount="1">
|
|
CUMZ 12005 (43 shells) from
|
|
<location id="5BDE773E37E6BE65EEFD1ED90F38E9D4" LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:5BDE773E37E6BE65EEFD1ED90F38E9D4" country="Myanmar" county="Mae Hong Son Province" municipality="Mueang Mae Hong Son District" name="Pha Daeng Cave">Pha Daeng Cave</location>
|
|
,
|
|
<collectingMunicipality id="BD78E6685006A245ED49B35E367AD0A0">Mueang Mae Hong Son District</collectingMunicipality>
|
|
,
|
|
<collectingCounty id="0D93B2CFAFD3C058FAD8464AD0B37160">Mae Hong Son Province</collectingCounty>
|
|
,
|
|
<collectingDate id="89412E7662A682D73F908430CB62DD91" value="2015-01-18">18 Jan. 2015</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="500903694F1B4047AD14DF6820F9599C" collectingDate="2020-12-03" collectorName="Fig." country="Myanmar" county="Mae Hong Son Province" location="Pha Daeng Cave" municipality="Mueang Mae Hong Son District" specimenCount="1">
|
|
CUMZ 12006 (17 shells and
|
|
<specimenCount id="DB2220F82F6047A41697C61C649EDDF3" type="generic">1 specimen</specimenCount>
|
|
in ethanol;
|
|
<collectorName id="E2DBAB2C32C507524DBFBC7C007F2118">Fig.</collectorName>
|
|
<figureCitation id="DB9362B1A8335075C18A9E8FBE61FD03" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Live specimens of A, B Pollicaria mouhoti mouhoti: specimens A CUMZ 12166 and B CUMZ 12175 from Wang Daeng Cave, Phitsanulok C Pollicaria mouhoti monochroma, paratype CUMZ 1562 from Tam Pha Bing Temple, Loei D Tortulosa tortuosa, specimen CUMZ 12155 from Tham Suea Temple, Krabi E-H Pupina artata: specimens of E CUMZ 12006 from Pha Daeng Cave, Mae Hong Son F CUMZ 12008 from Tham Nam Pha Pha Ngam Temple, Lampang, and G, H CUMZ 12029 from Khao Tham Raet Temple, Chachoengsao showing the brown (G) and grey (H) shell morphs; All not to scale." figureDoi="10.3897/zookeys.1119.85400.figure8" httpUri="https://binary.pensoft.net/fig/735369" pageId="0" pageNumber="1">8E</figureCitation>
|
|
) from
|
|
<location id="ADF92299F57A4FCD2A10A8FA06EE05F5" LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:ADF92299F57A4FCD2A10A8FA06EE05F5" country="Myanmar" county="Mae Hong Son Province" municipality="Mueang Mae Hong Son District" name="Pha Daeng Cave">Pha Daeng Cave</location>
|
|
,
|
|
<collectingMunicipality id="D67E5CECE146001ABD5CA0B5C0CDC74F">Mueang Mae Hong Son District</collectingMunicipality>
|
|
,
|
|
<collectingCounty id="014FE1B92BBADB965B47FD62B5FEC98C">Mae Hong Son Province</collectingCounty>
|
|
,
|
|
<collectingDate id="CB9872D8C6B6F89F7CA7CF5C35F9654E" value="2020-12-03">3 Dec. 2020</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="5698308D867755F992DEE254EA8CA2A6" collectingDate="2008-01-07" collectorName="Tham Nam Pha Pha Ngam" country="Myanmar" location="Mae Phrik District" municipality="Lampang Province" specimenCount="1">
|
|
CUMZ 12007 (7 shells) from
|
|
<collectorName id="B4073CF0AB16CADACC46D38862851553">Tham Nam Pha Pha Ngam</collectorName>
|
|
Temple,
|
|
<location id="BE11E04038071036A8ED9000EC34D3A4" LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:BE11E04038071036A8ED9000EC34D3A4" country="Myanmar" municipality="Lampang Province" name="Mae Phrik District">Mae Phrik District</location>
|
|
,
|
|
<collectingMunicipality id="6D624CC154338A42ED53298B2B241AC5">Lampang Province</collectingMunicipality>
|
|
,
|
|
<collectingDate id="FD7246BE75D0D1F45BBA3AEA9D5509CF" value="2008-01-07">7 Jan. 2008</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="182D98BC9718F9E37152EE2CF3EDF13E" collectingDate="2020-10-08" collectorName="Fig." country="Myanmar" county="Lampang Province" location="Tham Nam Pha Pha Ngam" municipality="Mae Phrik District" specimenCount="4">
|
|
CUMZ 12008 (3 shells and
|
|
<specimenCount id="44E66A557BD60964427D6A55E2DC4218" type="generic">4 specimens</specimenCount>
|
|
in ethanol;
|
|
<collectorName id="9A29688BF4515C88097B1346C65EA216">Fig.</collectorName>
|
|
<figureCitation id="6CDD12F962ECD13E6474743BA20D9002" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Live specimens of A, B Pollicaria mouhoti mouhoti: specimens A CUMZ 12166 and B CUMZ 12175 from Wang Daeng Cave, Phitsanulok C Pollicaria mouhoti monochroma, paratype CUMZ 1562 from Tam Pha Bing Temple, Loei D Tortulosa tortuosa, specimen CUMZ 12155 from Tham Suea Temple, Krabi E-H Pupina artata: specimens of E CUMZ 12006 from Pha Daeng Cave, Mae Hong Son F CUMZ 12008 from Tham Nam Pha Pha Ngam Temple, Lampang, and G, H CUMZ 12029 from Khao Tham Raet Temple, Chachoengsao showing the brown (G) and grey (H) shell morphs; All not to scale." figureDoi="10.3897/zookeys.1119.85400.figure8" httpUri="https://binary.pensoft.net/fig/735369" pageId="0" pageNumber="1">8F</figureCitation>
|
|
) from
|
|
<location id="01D2E8299BF09483B4CD25152962D53F" LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:01D2E8299BF09483B4CD25152962D53F" country="Myanmar" county="Lampang Province" municipality="Mae Phrik District" name="Tham Nam Pha Pha Ngam">Tham Nam Pha Pha Ngam</location>
|
|
Temple,
|
|
<collectingMunicipality id="3F2E002EF01F2781B4007F4A427BA94D">Mae Phrik District</collectingMunicipality>
|
|
,
|
|
<collectingCounty id="EAA4273C858FF411920E072F4CFD88A3">Lampang Province</collectingCounty>
|
|
,
|
|
<collectingDate id="7181EE7E86B16EA69EBAEA19E11F8A51" value="2020-10-08">8 Oct. 2020</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="3F1DBB8C22EEF66DA69CA10A61D5318F" collectingDate="2008-10-24" collectorName="Figs" country="Myanmar" location="Phu Sang District" municipality="Phayao Province" specimenCount="1">
|
|
CUMZ 12009 (12 shells;
|
|
<collectorName id="12BB8D5BF5EB482206087A3165E5A4E7">Figs</collectorName>
|
|
<figureCitation id="928343912227DDA0BEE360D5F4D81EE6" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Shells of Pupina artata species group from mainland Southeast Asia A-M Pupina artata A syntype UMZC I. 102960. A and specimens B CUMZ 12011 C CUMZ 12023 D CUMZ 12038 E CUMZ 12026 F CUMZ 12015 G CUMZ 12009 H CUMZ 12001 I CUMZ 12003 J NHMUK 1906.4.4.28 K CUMZ 12037 L CUMZ 12031, and M CUMZ 12018 N, O Pupina pallens N lectotype SMF 109951 and O specimen CUMZ 12042 P-R Pupina limitanea P, Q syntypes NHMUK 1903.7.1.2967 and R specimen CUMZ 12044 S Pupina billeti, holotype MNHN-IM- 2000 - 35841 T Pupina verneaui, syntype MNHN-IM- 2000 - 35843 U, V Pupina hungerfordiana, specimens NHMUK 91.3.14.686 - 7 W, X Pupina bensoni sp. nov. W holotype CUMZ 12045 / 1 and X paratype CUMZ 12046 / 1. Photo: H. Taylor, NHM (A, P, Q), P. Maestrati, MNHN (S, T)." figureDoi="10.3897/zookeys.1119.85400.figure21" httpUri="https://binary.pensoft.net/fig/735382" pageId="0" pageNumber="1">21G</figureCitation>
|
|
,
|
|
<figureCitation id="00E2E0A7F1209A3EA360F98CEAE819A0" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Pupina artata A syntype UMZC I. 102960. A from Moulmein B specimen NHMUK 1906.4.4.28 from Moulmein C specimen CUMZ 12001 from Khao Tham Phra Temple, Chiang Rai D specimen CUMZ 12003 from Ban Ping Khong, Chiang Mai E specimen CUMZ 12009 from Phu Sang Waterfall, Phayao, and F specimen CUMZ 12011 from Thep Sathaporn Temple, Nakhon Sawan. Photo: H. Taylor, NHM (A)." figureDoi="10.3897/zookeys.1119.85400.figure22" httpUri="https://binary.pensoft.net/fig/735383" pageId="0" pageNumber="1">22E</figureCitation>
|
|
) from Phu Sang Waterfall,
|
|
<location id="4A671163378ACAF960EC3DC94C790046" LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:4A671163378ACAF960EC3DC94C790046" country="Myanmar" municipality="Phayao Province" name="Phu Sang District">Phu Sang District</location>
|
|
,
|
|
<collectingMunicipality id="FFD8E9D3C99E9088E7CE02707BE389B2">Phayao Province</collectingMunicipality>
|
|
,
|
|
<collectingDate id="773EE9BC59518CA70FF1EF2A247418CD" value="2008-10-24">24 Oct. 2008</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="262A319E58A1CD83201B312A44994110" collectingDate="2012-11-19" collectorName="Phu Sang Waterfall" country="Myanmar" location="Phu Sang District" municipality="Phayao Province" specimenCount="1">
|
|
CUMZ 12010 (5 shells) from
|
|
<collectorName id="ABC8EA6C869689EC6FF70834BDD6BA04">Phu Sang Waterfall</collectorName>
|
|
,
|
|
<location id="AE7A4C53D5517D19EBC953BE71C43A13" LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:AE7A4C53D5517D19EBC953BE71C43A13" country="Myanmar" municipality="Phayao Province" name="Phu Sang District">Phu Sang District</location>
|
|
,
|
|
<collectingMunicipality id="5CF2229E81562BDED8916ADB073A3C39">Phayao Province</collectingMunicipality>
|
|
,
|
|
<collectingDate id="7C46F3715AE889F7F45DC5FDB51C5508" value="2012-11-19">19 Nov. 2012</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="B59669DCB1FEBC18B099E128D579D515" collectingDate="2008-07-17" collectorName="Figs" country="Myanmar" county="Nakhon Sawan Province" location="Thep Sathaporn Temple" municipality="Banphot Phisai District" specimenCount="7">
|
|
CUMZ 12011 (12 shells and
|
|
<specimenCount id="C3A0104109657563B880D27FC0760DF7" type="generic">7 specimens</specimenCount>
|
|
in ethanol;
|
|
<collectorName id="D611318CB93858E5A68C3FD795595DCF">Figs</collectorName>
|
|
<figureCitation id="29C6EABD59D8022DA9894CDC48182F98" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Shells of Pupina artata species group from mainland Southeast Asia A-M Pupina artata A syntype UMZC I. 102960. A and specimens B CUMZ 12011 C CUMZ 12023 D CUMZ 12038 E CUMZ 12026 F CUMZ 12015 G CUMZ 12009 H CUMZ 12001 I CUMZ 12003 J NHMUK 1906.4.4.28 K CUMZ 12037 L CUMZ 12031, and M CUMZ 12018 N, O Pupina pallens N lectotype SMF 109951 and O specimen CUMZ 12042 P-R Pupina limitanea P, Q syntypes NHMUK 1903.7.1.2967 and R specimen CUMZ 12044 S Pupina billeti, holotype MNHN-IM- 2000 - 35841 T Pupina verneaui, syntype MNHN-IM- 2000 - 35843 U, V Pupina hungerfordiana, specimens NHMUK 91.3.14.686 - 7 W, X Pupina bensoni sp. nov. W holotype CUMZ 12045 / 1 and X paratype CUMZ 12046 / 1. Photo: H. Taylor, NHM (A, P, Q), P. Maestrati, MNHN (S, T)." figureDoi="10.3897/zookeys.1119.85400.figure21" httpUri="https://binary.pensoft.net/fig/735382" pageId="0" pageNumber="1">21B</figureCitation>
|
|
,
|
|
<figureCitation id="FF181C3E90CB6BF6980AD5E920EA4B00" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Pupina artata A syntype UMZC I. 102960. A from Moulmein B specimen NHMUK 1906.4.4.28 from Moulmein C specimen CUMZ 12001 from Khao Tham Phra Temple, Chiang Rai D specimen CUMZ 12003 from Ban Ping Khong, Chiang Mai E specimen CUMZ 12009 from Phu Sang Waterfall, Phayao, and F specimen CUMZ 12011 from Thep Sathaporn Temple, Nakhon Sawan. Photo: H. Taylor, NHM (A)." figureDoi="10.3897/zookeys.1119.85400.figure22" httpUri="https://binary.pensoft.net/fig/735383" pageId="0" pageNumber="1">22F</figureCitation>
|
|
) from
|
|
<location id="B492B2CF91C5E75245D5CB450C8E1799" LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:B492B2CF91C5E75245D5CB450C8E1799" country="Myanmar" county="Nakhon Sawan Province" municipality="Banphot Phisai District" name="Thep Sathaporn Temple">Thep Sathaporn Temple</location>
|
|
,
|
|
<collectingMunicipality id="9831AD9EF3C6681CFBF773FC3AF4FE36">Banphot Phisai District</collectingMunicipality>
|
|
,
|
|
<collectingCounty id="CD5EED830950064B8756CD5DBF0F55F3">Nakhon Sawan Province</collectingCounty>
|
|
,
|
|
<collectingDate id="F8A6DF0B7E71C925A3D3B6663AA2B6D0" value="2008-07-17">17 July 2008</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="AE8313AE2948E31005B349E28434216E" collectingDate="2009-07-08" collectorName="Khao Chuak Charoentham Temple" country="Myanmar" location="Ban Rai District" municipality="Uthai Thani Province" specimenCount="1">
|
|
CUMZ 12012 (17 shells) from
|
|
<collectorName id="104B4F121701E9B15EBD643D0FCAC7B7">Khao Chuak Charoentham Temple</collectorName>
|
|
,
|
|
<location id="85400E052CA5D5D88CAC4AF6F3D456D0" LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:85400E052CA5D5D88CAC4AF6F3D456D0" country="Myanmar" municipality="Uthai Thani Province" name="Ban Rai District">Ban Rai District</location>
|
|
,
|
|
<collectingMunicipality id="857AFC5E4011011E3F43D5801DDD11D5">Uthai Thani Province</collectingMunicipality>
|
|
,
|
|
<collectingDate id="941CC1FF493E621F5A84D2D43B772B4C" value="2009-07-08">8 July 2009</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="DEA4C18BB0F73B621C010A828F013EED" collectingDate="2016-08-27" collectorName="Khao Chuak Charoentham Temple" country="Myanmar" location="Ban Rai District" municipality="Uthai Thani Province" specimenCount="1">
|
|
CUMZ 12013 (3 shells) from
|
|
<collectorName id="CD090E93143FDE52A6DE7E9826465AFC">Khao Chuak Charoentham Temple</collectorName>
|
|
,
|
|
<location id="EB6774EBC5F9D94926C93F51C6460E7D" LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:EB6774EBC5F9D94926C93F51C6460E7D" country="Myanmar" municipality="Uthai Thani Province" name="Ban Rai District">Ban Rai District</location>
|
|
,
|
|
<collectingMunicipality id="95C28E80F695D52F2C95BA940B903FF1">Uthai Thani Province</collectingMunicipality>
|
|
,
|
|
<collectingDate id="F7789597FA61FF6270BB8819E17F836F" value="2016-08-27">27 Aug. 2016</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="3B718B94E0AA3175AAD62F8C3A64AC03" collectingDate="2020-12-05" collectorName="Khao Chuak Charoentham Temple" country="Myanmar" location="Ban Rai District" municipality="Uthai Thani Province" specimenCount="1">
|
|
CUMZ 12014 (1 shell) from
|
|
<collectorName id="9F3B5E9C548D91943BF27E753703C1C5">Khao Chuak Charoentham Temple</collectorName>
|
|
,
|
|
<location id="E885D15EB10AF982DE2D45561BFB2C27" LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:E885D15EB10AF982DE2D45561BFB2C27" country="Myanmar" municipality="Uthai Thani Province" name="Ban Rai District">Ban Rai District</location>
|
|
,
|
|
<collectingMunicipality id="8DAC08DDE895C60B2D999141212FA30F">Uthai Thani Province</collectingMunicipality>
|
|
,
|
|
<collectingDate id="2A9B80D9A2F605B726F0BB60EACD340C" value="2020-12-05">5 Dec. 2020</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="D15EFE36995D81F081B92A004176A3CA" collectingDate="2009-07-08" collectorName="Figs" country="Myanmar" location="Ban Rai District" municipality="Uthai Thani Province" specimenCount="1">
|
|
CUMZ 12015 (14 shells;
|
|
<collectorName id="F5F62035A490CC22A69533FD774B2D45">Figs</collectorName>
|
|
<figureCitation id="84014FED9B1997D933A02C232AF6A0B8" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Shells of Pupina artata species group from mainland Southeast Asia A-M Pupina artata A syntype UMZC I. 102960. A and specimens B CUMZ 12011 C CUMZ 12023 D CUMZ 12038 E CUMZ 12026 F CUMZ 12015 G CUMZ 12009 H CUMZ 12001 I CUMZ 12003 J NHMUK 1906.4.4.28 K CUMZ 12037 L CUMZ 12031, and M CUMZ 12018 N, O Pupina pallens N lectotype SMF 109951 and O specimen CUMZ 12042 P-R Pupina limitanea P, Q syntypes NHMUK 1903.7.1.2967 and R specimen CUMZ 12044 S Pupina billeti, holotype MNHN-IM- 2000 - 35841 T Pupina verneaui, syntype MNHN-IM- 2000 - 35843 U, V Pupina hungerfordiana, specimens NHMUK 91.3.14.686 - 7 W, X Pupina bensoni sp. nov. W holotype CUMZ 12045 / 1 and X paratype CUMZ 12046 / 1. Photo: H. Taylor, NHM (A, P, Q), P. Maestrati, MNHN (S, T)." figureDoi="10.3897/zookeys.1119.85400.figure21" httpUri="https://binary.pensoft.net/fig/735382" pageId="0" pageNumber="1">21F</figureCitation>
|
|
,
|
|
<figureCitation id="0DEA4520AE081A96B231A943827E5A36" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Pupina artata: specimens A CUMZ 12015 from Khao Wong Phrommachan Temple, Uthai Thani B CUMZ 12018 from Tham Khao Thalu Temple, Ratchaburi C CUMZ 12023 from Tham Khiriwong Temple, Prachub Kirikhan D CUMZ 12026 from Tham Khao Cha Ang On Temple, Chonburi E CUMZ 12031 from Tham Khao Chakan Temple, Sa Kaeo, and F CUMZ 12037 from Wua Ta Lap Island, Surat Thani." figureDoi="10.3897/zookeys.1119.85400.figure23" httpUri="https://binary.pensoft.net/fig/735384" pageId="0" pageNumber="1">23A</figureCitation>
|
|
) from Khao Wong Phrommachan Temple,
|
|
<location id="2F1B4E38622CC3B8C0F856B2B4B10889" LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:2F1B4E38622CC3B8C0F856B2B4B10889" country="Myanmar" municipality="Uthai Thani Province" name="Ban Rai District">Ban Rai District</location>
|
|
,
|
|
<collectingMunicipality id="2A4A15AF3AF8546796025C38C9020938">Uthai Thani Province</collectingMunicipality>
|
|
,
|
|
<collectingDate id="34D846C4FC9AEB181D8F43B0DE55D20F" value="2009-07-08">8 July 2009</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="C8C86D04DA2B38873AEC7FE0C29C7222" collectingDate="2020-12-05" collectorName="Tham Prathat Mueang Thep Temple" country="Myanmar" location="Ban Rai District" municipality="Uthai Thani Province" specimenCount="1">
|
|
CUMZ 12016 (5 shells) from
|
|
<collectorName id="06C23D63A248BA1C802E894F2662BE1B">Tham Prathat Mueang Thep Temple</collectorName>
|
|
,
|
|
<location id="92E29E54A9FE629C337BB66A119EC333" LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:92E29E54A9FE629C337BB66A119EC333" country="Myanmar" municipality="Uthai Thani Province" name="Ban Rai District">Ban Rai District</location>
|
|
,
|
|
<collectingMunicipality id="920072E068847ACB460D6EC9459F68B4">Uthai Thani Province</collectingMunicipality>
|
|
,
|
|
<collectingDate id="F2A3E9586A90EF6261042482EFDEA7D5" value="2020-12-05">5 Dec. 2020</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="7F9CD62B1E217F85F79877ACBFCD5DCA" collectingDate="2006-12-10" country="Myanmar" county="Kanchanaburi Province" location="Krasae Cave" municipality="Sai Yok District" specimenCount="1">
|
|
CUMZ 12017 (13 shells) from
|
|
<location id="2279B5C2C902810A8233C90779CF9586" LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:2279B5C2C902810A8233C90779CF9586" country="Myanmar" county="Kanchanaburi Province" municipality="Sai Yok District" name="Krasae Cave">Krasae Cave</location>
|
|
,
|
|
<collectingMunicipality id="D4AD4C19660F982AA440F561E65D96F5">Sai Yok District</collectingMunicipality>
|
|
,
|
|
<collectingCounty id="A0EC29F10E7984AA57BAF64694956168">Kanchanaburi Province</collectingCounty>
|
|
,
|
|
<collectingDate id="B8D26E13D9703FF5ABE84F3D7985BABC" value="2006-12-10">10 Dec. 2006</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="DBE8DF4B62EF45131388E0AB44D73B84" collectingDate="2020-08-19" collectorName="Tham Charoentham Temple" country="Myanmar" location="Mueang Kanchanaburi District" municipality="Kanchanaburi Province" specimenCount="1">
|
|
CUMZ 12173 (14 shells) from
|
|
<collectorName id="EF8A5CFF3ADBF45660CB3816249FA04D">Tham Charoentham Temple</collectorName>
|
|
,
|
|
<location id="538EED740A16EE79E004C506B562D599" LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:538EED740A16EE79E004C506B562D599" country="Myanmar" municipality="Kanchanaburi Province" name="Mueang Kanchanaburi District">Mueang Kanchanaburi District</location>
|
|
,
|
|
<collectingMunicipality id="6DC3D13351F9112B87D3A292C670B371">Kanchanaburi Province</collectingMunicipality>
|
|
,
|
|
<collectingDate id="6AE2D4D12051ED682273D9B5948F9A42" value="2020-08-19">19 Aug. 2020</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="88D61893678A997FDF71C20869473FC3" collectingDate="2015-06-30" collectorName="Ban Tapoepu-Wakruko" country="Myanmar" location="Umphang District" municipality="Tak Province" specimenCount="1">
|
|
CUMZ 12192 (1 shell) from
|
|
<collectorName id="AB16C1BFF09A8E786C3F7E023F20CCF2">Ban Tapoepu-Wakruko</collectorName>
|
|
,
|
|
<location id="3687A1ADD22DB26ECEC82DDA4146B532" LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:3687A1ADD22DB26ECEC82DDA4146B532" country="Myanmar" municipality="Tak Province" name="Umphang District">Umphang District</location>
|
|
,
|
|
<collectingMunicipality id="841F42760E671943E983B6441134A5D0">Tak Province</collectingMunicipality>
|
|
,
|
|
<collectingDate id="1F04BE551DD500E9538187EE26615C6F" value="2015-06-30">30 June 2015</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="C4CA9F5BC8F7100B7A422B25AEE02E66" collectingDate="2006-12-09" collectorName="Figs" country="Myanmar" county="Ratchaburi Province" location="Tham Khao Thalu Temple" municipality="Chom Bueang District" specimenCount="10">
|
|
CUMZ 12018 (68 shells and
|
|
<specimenCount id="ED9A85389C0A1FAEC351E52762E830E8" type="generic">10 specimens</specimenCount>
|
|
in ethanol;
|
|
<collectorName id="4FACC7ED17B3A32145D46EC727E1D0CA">Figs</collectorName>
|
|
<figureCitation id="0C2A4AF61E425E311A65AFF51614E8FA" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Shells of Pupina artata species group from mainland Southeast Asia A-M Pupina artata A syntype UMZC I. 102960. A and specimens B CUMZ 12011 C CUMZ 12023 D CUMZ 12038 E CUMZ 12026 F CUMZ 12015 G CUMZ 12009 H CUMZ 12001 I CUMZ 12003 J NHMUK 1906.4.4.28 K CUMZ 12037 L CUMZ 12031, and M CUMZ 12018 N, O Pupina pallens N lectotype SMF 109951 and O specimen CUMZ 12042 P-R Pupina limitanea P, Q syntypes NHMUK 1903.7.1.2967 and R specimen CUMZ 12044 S Pupina billeti, holotype MNHN-IM- 2000 - 35841 T Pupina verneaui, syntype MNHN-IM- 2000 - 35843 U, V Pupina hungerfordiana, specimens NHMUK 91.3.14.686 - 7 W, X Pupina bensoni sp. nov. W holotype CUMZ 12045 / 1 and X paratype CUMZ 12046 / 1. Photo: H. Taylor, NHM (A, P, Q), P. Maestrati, MNHN (S, T)." figureDoi="10.3897/zookeys.1119.85400.figure21" httpUri="https://binary.pensoft.net/fig/735382" pageId="0" pageNumber="1">21M</figureCitation>
|
|
,
|
|
<figureCitation id="8A4B8FFBF44F607DC94875BF5C77A590" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Pupina artata: specimens A CUMZ 12015 from Khao Wong Phrommachan Temple, Uthai Thani B CUMZ 12018 from Tham Khao Thalu Temple, Ratchaburi C CUMZ 12023 from Tham Khiriwong Temple, Prachub Kirikhan D CUMZ 12026 from Tham Khao Cha Ang On Temple, Chonburi E CUMZ 12031 from Tham Khao Chakan Temple, Sa Kaeo, and F CUMZ 12037 from Wua Ta Lap Island, Surat Thani." figureDoi="10.3897/zookeys.1119.85400.figure23" httpUri="https://binary.pensoft.net/fig/735384" pageId="0" pageNumber="1">23B</figureCitation>
|
|
) from
|
|
<location id="C7389ABB35BFA1589C509FDF77769369" LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:C7389ABB35BFA1589C509FDF77769369" country="Myanmar" county="Ratchaburi Province" municipality="Chom Bueang District" name="Tham Khao Thalu Temple">Tham Khao Thalu Temple</location>
|
|
,
|
|
<collectingMunicipality id="DE73D7A66C39AEF494AE6F430597DA76">Chom Bueang District</collectingMunicipality>
|
|
,
|
|
<collectingCounty id="75936D807DB524FCFDE239ED42192A03">Ratchaburi Province</collectingCounty>
|
|
,
|
|
<collectingDate id="47E2F3D80A856CFABCC33FC5A9DC3D4E" value="2006-12-09">9 Dec. 2006</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="C4996DA962DA59F7D2F697A4B0CB9A78" collectingDate="2009-12-09" collectorName="Tham Khao Thalu Temple" country="Myanmar" location="Chom Bueang District" municipality="Ratchaburi Province" specimenCount="1">
|
|
CUMZ 12019 (21 shells) from
|
|
<collectorName id="1357A7FDF21020DE3FB28B40EC19E585">Tham Khao Thalu Temple</collectorName>
|
|
,
|
|
<location id="6F38B9E2E6D9A1C4D861B8CAEA87A70C" LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:6F38B9E2E6D9A1C4D861B8CAEA87A70C" country="Myanmar" municipality="Ratchaburi Province" name="Chom Bueang District">Chom Bueang District</location>
|
|
,
|
|
<collectingMunicipality id="EDF1F538F78957D9B257CA8438AAAEC4">Ratchaburi Province</collectingMunicipality>
|
|
,
|
|
<collectingDate id="A15DFD5A61598B9B69087D47187771B0" value="2009-12-09">9 Dec. 2009</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="751D205CC085C9DCB5E05A0BB2F58211" collectingDate="2017-05-08" collectorName="Fig." country="Myanmar" county="Ratchaburi Province" location="Buri Ratchawanaram Temple" municipality="Pak Tho District" specimenCount="6">
|
|
CUMZ 12020 (4 shells and
|
|
<specimenCount id="4C910637258A1211EB623EDB94728257" type="generic">6 specimens</specimenCount>
|
|
in ethanol;
|
|
<collectorName id="7FDEC480BA935EC2FED2BF9FEA32B620">Fig.</collectorName>
|
|
<figureCitation id="66224FE7A310CD6F30DA32FF9415F627" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Live specimens of A-C Pupina artata: specimens A CUMZ 12020 from Buri Ratchawanaram Temple, Ratchaburi B CUMZ 12022 from Golden Dragon Cave, Ratchaburi, and C CUMZ 12037 from Wua Ta Lap Island, Surat Thani D, E Pupina bensoni sp. nov. D paratype CUMZ 12045 / 2 from Khao Wong Cave, Uthai Thani and E specimen CUMZ 12047 from Tham Namthip Bureau of Monks, Uthai Thani F, G Pupina peguensis: specimens F CUMZ 12050 from Chai Thong Wararam Temple, Nakhon Sawan and G CUMZ 12051 from Tham Saeng Wiset Bureau of Monks, Nakhon Sawan H Pupina siamensis, specimen CUMZ 12069 from Khao Chi Chan Buddha Image, Chonburi. All not to scale." figureDoi="10.3897/zookeys.1119.85400.figure25" httpUri="https://binary.pensoft.net/fig/735386" pageId="0" pageNumber="1">25A</figureCitation>
|
|
) from
|
|
<location id="05F52406279442C30BF7702955CCBC50" LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:05F52406279442C30BF7702955CCBC50" country="Myanmar" county="Ratchaburi Province" municipality="Pak Tho District" name="Buri Ratchawanaram Temple">Buri Ratchawanaram Temple</location>
|
|
,
|
|
<collectingMunicipality id="7F464EE0BA345E5C3DF24426A85EB3E5">Pak Tho District</collectingMunicipality>
|
|
,
|
|
<collectingCounty id="C6408F374C7FD45FEBE6B8E87E5B53C6">Ratchaburi Province</collectingCounty>
|
|
,
|
|
<collectingDate id="CAB738A14E735A17809E26CB99C5DF8A" value="2017-05-08">8 May 2017</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="918FC1E57A9F6B87896D47FF642A34B0" collectingDate="2020-08-18" country="Myanmar" county="Ratchaburi Province" location="Buri Ratchawanaram Temple" municipality="Pak Tho District" specimenCount="1">
|
|
CUMZ 12021 (20 shells and
|
|
<specimenCount id="2892C6530EF1E57D2AC8103FC8C98C5D" type="generic">1 specimen</specimenCount>
|
|
in ethanol) from
|
|
<location id="923BA5CE72B828BA62D735B8A3D3AFD5" LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:923BA5CE72B828BA62D735B8A3D3AFD5" country="Myanmar" county="Ratchaburi Province" municipality="Pak Tho District" name="Buri Ratchawanaram Temple">Buri Ratchawanaram Temple</location>
|
|
,
|
|
<collectingMunicipality id="C6F7090899FEC081B9E5012BC726A7E4">Pak Tho District</collectingMunicipality>
|
|
,
|
|
<collectingCounty id="D9164FC74A425FEA746940A1BD91ADE9">Ratchaburi Province</collectingCounty>
|
|
,
|
|
<collectingDate id="8DEE6F26CB52A901CD3B30210B5A989E" value="2020-08-18">18 Aug. 2020</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="9A7633C414A2E2E507D5738010F51CC2" collectingDate="2019-08-18" collectorName="Fig." country="Myanmar" county="Ratchaburi Province" location="Golden Dragon Cave" municipality="Pak Tho District" specimenCount="5">
|
|
CUMZ 12022 (
|
|
<specimenCount id="CDC8E74D66A06E13F6DBD3EDD9B23576" type="generic">5 specimens</specimenCount>
|
|
in ethanol;
|
|
<collectorName id="33BD78ECB35A135827E0CBDDF3F0DE7F">Fig.</collectorName>
|
|
<figureCitation id="6040DD7B6C501177A3AA64078EDBDB2D" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Live specimens of A-C Pupina artata: specimens A CUMZ 12020 from Buri Ratchawanaram Temple, Ratchaburi B CUMZ 12022 from Golden Dragon Cave, Ratchaburi, and C CUMZ 12037 from Wua Ta Lap Island, Surat Thani D, E Pupina bensoni sp. nov. D paratype CUMZ 12045 / 2 from Khao Wong Cave, Uthai Thani and E specimen CUMZ 12047 from Tham Namthip Bureau of Monks, Uthai Thani F, G Pupina peguensis: specimens F CUMZ 12050 from Chai Thong Wararam Temple, Nakhon Sawan and G CUMZ 12051 from Tham Saeng Wiset Bureau of Monks, Nakhon Sawan H Pupina siamensis, specimen CUMZ 12069 from Khao Chi Chan Buddha Image, Chonburi. All not to scale." figureDoi="10.3897/zookeys.1119.85400.figure25" httpUri="https://binary.pensoft.net/fig/735386" pageId="0" pageNumber="1">25B</figureCitation>
|
|
) from
|
|
<location id="18E4645F1C44CE967CAC46ADC96CE929" LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:18E4645F1C44CE967CAC46ADC96CE929" country="Myanmar" county="Ratchaburi Province" municipality="Pak Tho District" name="Golden Dragon Cave">Golden Dragon Cave</location>
|
|
,
|
|
<collectingMunicipality id="6B01BED5E2D473DFEA0C6301DCF2CFEA">Pak Tho District</collectingMunicipality>
|
|
,
|
|
<collectingCounty id="2C17D5BA7963E2687EE86607C9A7181E">Ratchaburi Province</collectingCounty>
|
|
,
|
|
<collectingDate id="5135BD4D6F08051843FD546B062B263D" value="2019-08-18">18 Aug. 2019</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="3617253B4CBC52B6D43814A4985EB72D" collectingDate="2007-04-21" collectorName="Figs" country="Myanmar" location="Bang Saphan District" municipality="Prachub Kirikhan Province" specimenCount="1">
|
|
CUMZ 12023 (17 shells;
|
|
<collectorName id="4357450B3745226FE2490B0E69C4A0B5">Figs</collectorName>
|
|
<figureCitation id="C2679296B6AE8669AAC456B1F83C6A8E" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Shells of Pupina artata species group from mainland Southeast Asia A-M Pupina artata A syntype UMZC I. 102960. A and specimens B CUMZ 12011 C CUMZ 12023 D CUMZ 12038 E CUMZ 12026 F CUMZ 12015 G CUMZ 12009 H CUMZ 12001 I CUMZ 12003 J NHMUK 1906.4.4.28 K CUMZ 12037 L CUMZ 12031, and M CUMZ 12018 N, O Pupina pallens N lectotype SMF 109951 and O specimen CUMZ 12042 P-R Pupina limitanea P, Q syntypes NHMUK 1903.7.1.2967 and R specimen CUMZ 12044 S Pupina billeti, holotype MNHN-IM- 2000 - 35841 T Pupina verneaui, syntype MNHN-IM- 2000 - 35843 U, V Pupina hungerfordiana, specimens NHMUK 91.3.14.686 - 7 W, X Pupina bensoni sp. nov. W holotype CUMZ 12045 / 1 and X paratype CUMZ 12046 / 1. Photo: H. Taylor, NHM (A, P, Q), P. Maestrati, MNHN (S, T)." figureDoi="10.3897/zookeys.1119.85400.figure21" httpUri="https://binary.pensoft.net/fig/735382" pageId="0" pageNumber="1">21C</figureCitation>
|
|
,
|
|
<figureCitation id="55000505F770EBABF4A697F5169E747A" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Pupina artata: specimens A CUMZ 12015 from Khao Wong Phrommachan Temple, Uthai Thani B CUMZ 12018 from Tham Khao Thalu Temple, Ratchaburi C CUMZ 12023 from Tham Khiriwong Temple, Prachub Kirikhan D CUMZ 12026 from Tham Khao Cha Ang On Temple, Chonburi E CUMZ 12031 from Tham Khao Chakan Temple, Sa Kaeo, and F CUMZ 12037 from Wua Ta Lap Island, Surat Thani." figureDoi="10.3897/zookeys.1119.85400.figure23" httpUri="https://binary.pensoft.net/fig/735384" pageId="0" pageNumber="1">23C</figureCitation>
|
|
) from Tham Khiriwong Temple,
|
|
<location id="47CF34F5E00BC3137795A7547A77F428" LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:47CF34F5E00BC3137795A7547A77F428" country="Myanmar" municipality="Prachub Kirikhan Province" name="Bang Saphan District">Bang Saphan District</location>
|
|
,
|
|
<collectingMunicipality id="A3EBE97F7CE704793BE8020966C596DA">Prachub Kirikhan Province</collectingMunicipality>
|
|
,
|
|
<collectingDate id="14D0B9AC8044341DA15ACE6BC1D9EF60" value="2007-04-21">21 Apr. 2007</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="E89B21ECD7E422144315E7ADAEF0BDF9" collectingDate="2019-07-29" collectorName="Tham Khiriwong Temple" country="Myanmar" location="Bang Saphan District" municipality="Prachub Kirikhan Province" specimenCount="1">
|
|
CUMZ 12024 (147 shells) from
|
|
<collectorName id="62B3BAA682B376FA832BAC13128684BA">Tham Khiriwong Temple</collectorName>
|
|
,
|
|
<location id="5D0A27A4465E4E6234BAC370B9F1CB4B" LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:5D0A27A4465E4E6234BAC370B9F1CB4B" country="Myanmar" municipality="Prachub Kirikhan Province" name="Bang Saphan District">Bang Saphan District</location>
|
|
,
|
|
<collectingMunicipality id="4A659C5270DFC68088BAF6415D5FEF7B">Prachub Kirikhan Province</collectingMunicipality>
|
|
,
|
|
<collectingDate id="ACC42F4A062FBEDA04245F5E199D445D" value="2019-07-29">29 July 2019</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="DF7B2750332596050F302B8F5FBD4D92" collectingDate="2020-08-24" collectorName="Tham Thep Nimit Temple" country="Myanmar" location="Pak Chong District" municipality="Nakhon Ratchasima Province" specimenCount="1">
|
|
CUMZ 12169 (10 shells) from
|
|
<collectorName id="6DB6ABCF7AE59C83907849D5BC2A98DF">Tham Thep Nimit Temple</collectorName>
|
|
,
|
|
<location id="3482F7EF37CFCEF70560A24194D8CE45" LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:3482F7EF37CFCEF70560A24194D8CE45" country="Myanmar" municipality="Nakhon Ratchasima Province" name="Pak Chong District">Pak Chong District</location>
|
|
,
|
|
<collectingMunicipality id="235AB9141EC28C4CF37D4755BBFAE9AB">Nakhon Ratchasima Province</collectingMunicipality>
|
|
,
|
|
<collectingDate id="9B1E6A0B6C7327DA3DEF24B401A5E5F2" value="2020-08-24">24 Aug. 2020</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="C0D65C348A68DAA5FD3A1D2946369626" collectingDate="2006-03-13" country="Myanmar" county="Chonburi Province" location="Tham Khao Cha Ang On Temple" municipality="Bo Thong District" specimenCount="1">
|
|
CUMZ 12025 (28 shells) from
|
|
<location id="D83DDE694FDE763CD586AC0D9E694C10" LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:D83DDE694FDE763CD586AC0D9E694C10" country="Myanmar" county="Chonburi Province" municipality="Bo Thong District" name="Tham Khao Cha Ang On Temple">Tham Khao Cha Ang On Temple</location>
|
|
,
|
|
<collectingMunicipality id="C0880AB24ADC268CA1D88DBDEEC7C3B7">Bo Thong District</collectingMunicipality>
|
|
,
|
|
<collectingCounty id="5D884624EBA22FE9190218A0917A89BC">Chonburi Province</collectingCounty>
|
|
,
|
|
<collectingDate id="0CC208B4CDE41C964076B537EF242F76" value="2006-03-13">13 Mar. 2006</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="0CB1F9BEB45145A1A1DB43D7AF1C7D1F" collectingDate="2006-08-17" collectorName="Figs" country="Myanmar" county="Chonburi Province" location="Tham Khao Cha Ang On Temple" municipality="Bo Thong District" specimenCount="23">
|
|
CUMZ 12026 (28 shells and
|
|
<specimenCount id="BB39AEACD257236B333B909149A10D52" type="generic">23 specimens</specimenCount>
|
|
in ethanol;
|
|
<collectorName id="97CF19F68895ABDF5FAA1F815577B621">Figs</collectorName>
|
|
<figureCitation id="C1DB7ABED9FCF0B564FEF55F448946CD" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Shells of Pupina artata species group from mainland Southeast Asia A-M Pupina artata A syntype UMZC I. 102960. A and specimens B CUMZ 12011 C CUMZ 12023 D CUMZ 12038 E CUMZ 12026 F CUMZ 12015 G CUMZ 12009 H CUMZ 12001 I CUMZ 12003 J NHMUK 1906.4.4.28 K CUMZ 12037 L CUMZ 12031, and M CUMZ 12018 N, O Pupina pallens N lectotype SMF 109951 and O specimen CUMZ 12042 P-R Pupina limitanea P, Q syntypes NHMUK 1903.7.1.2967 and R specimen CUMZ 12044 S Pupina billeti, holotype MNHN-IM- 2000 - 35841 T Pupina verneaui, syntype MNHN-IM- 2000 - 35843 U, V Pupina hungerfordiana, specimens NHMUK 91.3.14.686 - 7 W, X Pupina bensoni sp. nov. W holotype CUMZ 12045 / 1 and X paratype CUMZ 12046 / 1. Photo: H. Taylor, NHM (A, P, Q), P. Maestrati, MNHN (S, T)." figureDoi="10.3897/zookeys.1119.85400.figure21" httpUri="https://binary.pensoft.net/fig/735382" pageId="0" pageNumber="1">21E</figureCitation>
|
|
,
|
|
<figureCitation id="4F9B7C6EFE11B7272E5AAE6EC157468B" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Pupina artata: specimens A CUMZ 12015 from Khao Wong Phrommachan Temple, Uthai Thani B CUMZ 12018 from Tham Khao Thalu Temple, Ratchaburi C CUMZ 12023 from Tham Khiriwong Temple, Prachub Kirikhan D CUMZ 12026 from Tham Khao Cha Ang On Temple, Chonburi E CUMZ 12031 from Tham Khao Chakan Temple, Sa Kaeo, and F CUMZ 12037 from Wua Ta Lap Island, Surat Thani." figureDoi="10.3897/zookeys.1119.85400.figure23" httpUri="https://binary.pensoft.net/fig/735384" pageId="0" pageNumber="1">23D</figureCitation>
|
|
) from
|
|
<location id="AED7EE91FDB4C53A8EFB9CAFABA20BBD" LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:AED7EE91FDB4C53A8EFB9CAFABA20BBD" country="Myanmar" county="Chonburi Province" municipality="Bo Thong District" name="Tham Khao Cha Ang On Temple">Tham Khao Cha Ang On Temple</location>
|
|
,
|
|
<collectingMunicipality id="C7A7F1AD06C9943D6FE0A272598E0798">Bo Thong District</collectingMunicipality>
|
|
,
|
|
<collectingCounty id="FA717DB28A30108B06B299A317E231B2">Chonburi Province</collectingCounty>
|
|
,
|
|
<collectingDate id="0002844D20AF2515A8E6BE731074665E" value="2006-08-17">17 Aug. 2006</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="A367319DE9F3520896FF2420A9D46CC0" collectingDate="2008-05-09" country="Myanmar" location="Bo Thong District" municipality="Chonburi Province" specimenCount="1">
|
|
CUMZ 12028 (34 shells) from
|
|
<location id="046BF8165D0DD2ADEE3B2D3727BCC743" LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:046BF8165D0DD2ADEE3B2D3727BCC743" country="Myanmar" municipality="Chonburi Province" name="Bo Thong District">Bo Thong District</location>
|
|
,
|
|
<collectingMunicipality id="705760B798ECDD82C21089CE2DC7F8DC">Chonburi Province</collectingMunicipality>
|
|
,
|
|
<collectingDate id="897920B54DF81450EB12E370B5F71370" value="2008-05-09">9 May 2008</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="71AFE158305D13B96E6CF59916EA725D" collectingDate="2020-09-19" collectorName="Phromawat Temple" country="Myanmar" location="Si Racha District" municipality="Chonburi Province" specimenCount="2">
|
|
CUMZ 12027 (
|
|
<specimenCount id="2DA82305CC23254B3B0C9C38F5CEDACB" type="generic">2 specimens</specimenCount>
|
|
in ethanol) from
|
|
<collectorName id="1D705853D21DAE1E01B5A88F0CE1210D">Phromawat Temple</collectorName>
|
|
,
|
|
<location id="89C438925D5D5ED3E71334F50D8C8A74" LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:89C438925D5D5ED3E71334F50D8C8A74" country="Myanmar" municipality="Chonburi Province" name="Si Racha District">Si Racha District</location>
|
|
,
|
|
<collectingMunicipality id="2A26F4665A7DC064A42B3D4F5A83964E">Chonburi Province</collectingMunicipality>
|
|
,
|
|
<collectingDate id="80D8C6B4B653C1C00800EE14F797B44F" value="2020-09-19">19 Sept. 2020</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="18052EF6615BA6E69912C949EB8AB5C1" collectingDate="2010-10-23" collectorName="Tham Khao Loi Temple" country="Myanmar" location="Khao Chamao District" municipality="Rayong Province" specimenCount="1">
|
|
CUMZ 12174 (1 shell) from
|
|
<collectorName id="673ADB916936EAB35D09460DBE2B56F2">Tham Khao Loi Temple</collectorName>
|
|
,
|
|
<location id="8C5408AEC44EEC57FC6B882C97BDFBDE" LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:8C5408AEC44EEC57FC6B882C97BDFBDE" country="Myanmar" municipality="Rayong Province" name="Khao Chamao District">Khao Chamao District</location>
|
|
,
|
|
<collectingMunicipality id="12253501FEB8EEEAB2A11B4A3262BF29">Rayong Province</collectingMunicipality>
|
|
,
|
|
<collectingDate id="9CAF5BEFC38862F80C577CBA762A8639" value="2010-10-23">23 Oct. 2010</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="A05EBF6C5FE46E784B9D262A353C46A9" collectingDate="2012-05-21" collectorName="Fig." country="Myanmar" county="Chachoengsao Province" location="Khao Tham Raet Temple" municipality="Tha Takiap District" specimenCount="10">
|
|
CUMZ 12029 (85 shells and
|
|
<specimenCount id="E7174168002AFAE12E8DDDC32468EB74" type="generic">10 specimens</specimenCount>
|
|
in ethanol;
|
|
<collectorName id="3467E9EAC55D5C15878C8C45039A5E8B">Fig.</collectorName>
|
|
<figureCitation id="169D919BFD01D7E42820222B043EE0FF" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Live specimens of A, B Pollicaria mouhoti mouhoti: specimens A CUMZ 12166 and B CUMZ 12175 from Wang Daeng Cave, Phitsanulok C Pollicaria mouhoti monochroma, paratype CUMZ 1562 from Tam Pha Bing Temple, Loei D Tortulosa tortuosa, specimen CUMZ 12155 from Tham Suea Temple, Krabi E-H Pupina artata: specimens of E CUMZ 12006 from Pha Daeng Cave, Mae Hong Son F CUMZ 12008 from Tham Nam Pha Pha Ngam Temple, Lampang, and G, H CUMZ 12029 from Khao Tham Raet Temple, Chachoengsao showing the brown (G) and grey (H) shell morphs; All not to scale." figureDoi="10.3897/zookeys.1119.85400.figure8" httpUri="https://binary.pensoft.net/fig/735369" pageId="0" pageNumber="1">8G, H</figureCitation>
|
|
) from
|
|
<location id="AF158BCED7B44D87E9D1E965D9A98C52" LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:AF158BCED7B44D87E9D1E965D9A98C52" country="Myanmar" county="Chachoengsao Province" municipality="Tha Takiap District" name="Khao Tham Raet Temple">Khao Tham Raet Temple</location>
|
|
,
|
|
<collectingMunicipality id="D4D286BAE05AD8718E9D9D70ADE92EEF">Tha Takiap District</collectingMunicipality>
|
|
,
|
|
<collectingCounty id="ED8AF92CF6469339468E8B31A226B7F5">Chachoengsao Province</collectingCounty>
|
|
,
|
|
<collectingDate id="2962897AEF773231CC9535D012FCA517" value="2012-05-21">21 May 2012</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="FA7413804C10CA5022318191E42F377D" collectingDate="2018-03-01" collectorName="Khao Tham Raet Temple" country="Myanmar" location="Tha Takiap District" municipality="Chachoengsao Province" specimenCount="1">
|
|
CUMZ 12030 (3 shells) from
|
|
<collectorName id="F18295AF56540A74A5A6BB4DAB8755B0">Khao Tham Raet Temple</collectorName>
|
|
,
|
|
<location id="424F18465441B8527D56966F908844B9" LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:424F18465441B8527D56966F908844B9" country="Myanmar" municipality="Chachoengsao Province" name="Tha Takiap District">Tha Takiap District</location>
|
|
,
|
|
<collectingMunicipality id="F86D98D299567819310316212B181120">Chachoengsao Province</collectingMunicipality>
|
|
,
|
|
<collectingDate id="14C485E7172AED23762633949B513C55" value="2018-03-01">1 Mar. 2018</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="0414BC16798C4CA5E65D218C05D1FE9E" collectingDate="2000-04-07" collectorName="Figs" country="Myanmar" location="Khao Chakan District" municipality="Sa Kaeo Province" specimenCount="1">
|
|
CUMZ 12031 (43 shells;
|
|
<collectorName id="B97537E7864D5D62DE988EE652DF2919">Figs</collectorName>
|
|
<figureCitation id="6D45A5BE6578A514C462BD946FE22868" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Shells of Pupina artata species group from mainland Southeast Asia A-M Pupina artata A syntype UMZC I. 102960. A and specimens B CUMZ 12011 C CUMZ 12023 D CUMZ 12038 E CUMZ 12026 F CUMZ 12015 G CUMZ 12009 H CUMZ 12001 I CUMZ 12003 J NHMUK 1906.4.4.28 K CUMZ 12037 L CUMZ 12031, and M CUMZ 12018 N, O Pupina pallens N lectotype SMF 109951 and O specimen CUMZ 12042 P-R Pupina limitanea P, Q syntypes NHMUK 1903.7.1.2967 and R specimen CUMZ 12044 S Pupina billeti, holotype MNHN-IM- 2000 - 35841 T Pupina verneaui, syntype MNHN-IM- 2000 - 35843 U, V Pupina hungerfordiana, specimens NHMUK 91.3.14.686 - 7 W, X Pupina bensoni sp. nov. W holotype CUMZ 12045 / 1 and X paratype CUMZ 12046 / 1. Photo: H. Taylor, NHM (A, P, Q), P. Maestrati, MNHN (S, T)." figureDoi="10.3897/zookeys.1119.85400.figure21" httpUri="https://binary.pensoft.net/fig/735382" pageId="0" pageNumber="1">21L</figureCitation>
|
|
,
|
|
<figureCitation id="4EF8520D049918E922782FA91FE24A58" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Pupina artata: specimens A CUMZ 12015 from Khao Wong Phrommachan Temple, Uthai Thani B CUMZ 12018 from Tham Khao Thalu Temple, Ratchaburi C CUMZ 12023 from Tham Khiriwong Temple, Prachub Kirikhan D CUMZ 12026 from Tham Khao Cha Ang On Temple, Chonburi E CUMZ 12031 from Tham Khao Chakan Temple, Sa Kaeo, and F CUMZ 12037 from Wua Ta Lap Island, Surat Thani." figureDoi="10.3897/zookeys.1119.85400.figure23" httpUri="https://binary.pensoft.net/fig/735384" pageId="0" pageNumber="1">23E</figureCitation>
|
|
) from Tham Khao Chakan Temple,
|
|
<location id="D02B1FBCF5464D182E0BB6676F37BD42" LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:D02B1FBCF5464D182E0BB6676F37BD42" country="Myanmar" municipality="Sa Kaeo Province" name="Khao Chakan District">Khao Chakan District</location>
|
|
,
|
|
<collectingMunicipality id="6CD69733A7BEB4855BE150BE73668CF6">Sa Kaeo Province</collectingMunicipality>
|
|
,
|
|
<collectingDate id="FD16642408551D9295A7CA3CB0AB1106" value="2000-04-07">7 Apr. 2000</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="5830831E82E95C0D63A7E795E49121EA" collectingDate="2018-02-25" collectorName="Tham Khao Chakan Temple" country="Myanmar" location="Khao Chakan District" municipality="Sa Kaeo Province" specimenCount="7">
|
|
CUMZ 12032 (
|
|
<specimenCount id="E8A7471A9899F1249C5D59EE06CD8F58" type="generic">7 specimens</specimenCount>
|
|
in ethanol) from
|
|
<collectorName id="89AAF12463F340E4BC118039F31AE734">Tham Khao Chakan Temple</collectorName>
|
|
,
|
|
<location id="EC32765B241B884683E80B23103FDB18" LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:EC32765B241B884683E80B23103FDB18" country="Myanmar" municipality="Sa Kaeo Province" name="Khao Chakan District">Khao Chakan District</location>
|
|
,
|
|
<collectingMunicipality id="21F50501C1572AC122B60B6B3EA74CF8">Sa Kaeo Province</collectingMunicipality>
|
|
,
|
|
<collectingDate id="4F639C74F6D00B6F24090320AA123F52" value="2018-02-25">25 Feb. 2018</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="FCAC5A8913459DE2A84F3A227502426D" collectingDate="2008-11-02" collectorName="Tham Khao Maka Temple" country="Myanmar" location="Mueang Sa Kaeo District" municipality="Sa Kaeo Province" specimenCount="1">
|
|
CUMZ 12033 (2 shells) from
|
|
<collectorName id="23DE045C33591132D26F642A3D246789">Tham Khao Maka Temple</collectorName>
|
|
,
|
|
<location id="810B7821D50D150C529009752F02AAAD" LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:810B7821D50D150C529009752F02AAAD" country="Myanmar" municipality="Sa Kaeo Province" name="Mueang Sa Kaeo District">Mueang Sa Kaeo District</location>
|
|
,
|
|
<collectingMunicipality id="8F0B72BB1DD08B06F0EA9EF5740EABD8">Sa Kaeo Province</collectingMunicipality>
|
|
,
|
|
<collectingDate id="46750156EEE4AE73055127ACC5B79F38" value="2008-11-02">2 Nov. 2008</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="4C26DF06E71B7B116913C3DF93BEB840" collectingDate="2018-05-21" collectorName="Khao Pha Pheung Temple" country="Myanmar" location="Klong Had District" municipality="Sra Keo Province" specimenCount="1">
|
|
CUMZ 12034 (2 shells) from
|
|
<collectorName id="22CDE24DE5FCE0971D23321FB88B761B">Khao Pha Pheung Temple</collectorName>
|
|
,
|
|
<location id="C29BB84C9CDABA8AC50EDD26B9CD5A44" LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:C29BB84C9CDABA8AC50EDD26B9CD5A44" country="Myanmar" municipality="Sra Keo Province" name="Klong Had District">Klong Had District</location>
|
|
,
|
|
<collectingMunicipality id="1703B7F1CD34C936A134EFEC6752E9B3">Sra Keo Province</collectingMunicipality>
|
|
,
|
|
<collectingDate id="C8D016346858F38A25277F43C5B1E71E" value="2018-05-21">21 May 2018</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="2629806914418F93802207B4DAB21114" collectingDate="2007-03-04" collectorName="Na Mueang Waterfall" country="Myanmar" location="Ko Samui District" municipality="Surat Thani Province" specimenCount="1">
|
|
CUMZ 12035 (
|
|
<specimenCount id="11C2201CE4554B0F6F0471601AF91DCF" type="generic">1 specimen</specimenCount>
|
|
in ethanol) from
|
|
<collectorName id="9D8ACBF5ED2DD5076A24D48B5B96C8C3">Na Mueang Waterfall</collectorName>
|
|
,
|
|
<location id="456FA03FBD025C6BA868454C22FCFFE9" LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:456FA03FBD025C6BA868454C22FCFFE9" country="Myanmar" municipality="Surat Thani Province" name="Ko Samui District">Ko Samui District</location>
|
|
,
|
|
<collectingMunicipality id="ABF5607F67E97A65723792126A8374F2">Surat Thani Province</collectingMunicipality>
|
|
,
|
|
<collectingDate id="B984CA20D5A730850683F34BFDA24CB8" value="2007-03-04">4 Mar. 2007</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="858CEDF9E067DB10A887D8DD3FCDEE02" collectingDate="2007-03-05" country="Myanmar" county="Surat Thani Province" location="Wua Ta Lap Island" municipality="Ko Samui District" specimenCount="1">
|
|
CUMZ 12036 (10 shells) from
|
|
<location id="22F7E8E43133B334F33DA398FB233FF5" LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:22F7E8E43133B334F33DA398FB233FF5" country="Myanmar" county="Surat Thani Province" municipality="Ko Samui District" name="Wua Ta Lap Island">Wua Ta Lap Island</location>
|
|
,
|
|
<collectingMunicipality id="89A96A61B756B2F00C1523085A0BA1EA">Ko Samui District</collectingMunicipality>
|
|
,
|
|
<collectingCounty id="44E48034BD95FFFFB052ACC801207CF9">Surat Thani Province</collectingCounty>
|
|
,
|
|
<collectingDate id="4CBCED0069D5AEDA2D0CA505EA8D1741" value="2007-03-05">5 Mar. 2007</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="19F6391A4C96176AA8B49EA84225EF76" collectingDate="2009-06-06" collectorName="Figs" country="Myanmar" county="Surat Thani Province" location="Wua Ta Lap Island" municipality="Ko Samui District" specimenCount="2">
|
|
CUMZ 12037 (1 shell and
|
|
<specimenCount id="3CE3E7250C1F4CB9C0A895AA528BDA8B" type="generic">2 specimens</specimenCount>
|
|
in ethanol;
|
|
<collectorName id="514CF898FC5C21BDEB243D8B666850F3">Figs</collectorName>
|
|
<figureCitation id="EE2C4BF0227A1F8DBC60F0298CF9F029" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Shells of Pupina artata species group from mainland Southeast Asia A-M Pupina artata A syntype UMZC I. 102960. A and specimens B CUMZ 12011 C CUMZ 12023 D CUMZ 12038 E CUMZ 12026 F CUMZ 12015 G CUMZ 12009 H CUMZ 12001 I CUMZ 12003 J NHMUK 1906.4.4.28 K CUMZ 12037 L CUMZ 12031, and M CUMZ 12018 N, O Pupina pallens N lectotype SMF 109951 and O specimen CUMZ 12042 P-R Pupina limitanea P, Q syntypes NHMUK 1903.7.1.2967 and R specimen CUMZ 12044 S Pupina billeti, holotype MNHN-IM- 2000 - 35841 T Pupina verneaui, syntype MNHN-IM- 2000 - 35843 U, V Pupina hungerfordiana, specimens NHMUK 91.3.14.686 - 7 W, X Pupina bensoni sp. nov. W holotype CUMZ 12045 / 1 and X paratype CUMZ 12046 / 1. Photo: H. Taylor, NHM (A, P, Q), P. Maestrati, MNHN (S, T)." figureDoi="10.3897/zookeys.1119.85400.figure21" httpUri="https://binary.pensoft.net/fig/735382" pageId="0" pageNumber="1">21K</figureCitation>
|
|
,
|
|
<figureCitation id="7762843B4F51B09E165DF654EFB38419" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Pupina artata: specimens A CUMZ 12015 from Khao Wong Phrommachan Temple, Uthai Thani B CUMZ 12018 from Tham Khao Thalu Temple, Ratchaburi C CUMZ 12023 from Tham Khiriwong Temple, Prachub Kirikhan D CUMZ 12026 from Tham Khao Cha Ang On Temple, Chonburi E CUMZ 12031 from Tham Khao Chakan Temple, Sa Kaeo, and F CUMZ 12037 from Wua Ta Lap Island, Surat Thani." figureDoi="10.3897/zookeys.1119.85400.figure23" httpUri="https://binary.pensoft.net/fig/735384" pageId="0" pageNumber="1">23F</figureCitation>
|
|
,
|
|
<figureCitation id="47E317BEFDCF7B13A95315489CC54809" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Live specimens of A-C Pupina artata: specimens A CUMZ 12020 from Buri Ratchawanaram Temple, Ratchaburi B CUMZ 12022 from Golden Dragon Cave, Ratchaburi, and C CUMZ 12037 from Wua Ta Lap Island, Surat Thani D, E Pupina bensoni sp. nov. D paratype CUMZ 12045 / 2 from Khao Wong Cave, Uthai Thani and E specimen CUMZ 12047 from Tham Namthip Bureau of Monks, Uthai Thani F, G Pupina peguensis: specimens F CUMZ 12050 from Chai Thong Wararam Temple, Nakhon Sawan and G CUMZ 12051 from Tham Saeng Wiset Bureau of Monks, Nakhon Sawan H Pupina siamensis, specimen CUMZ 12069 from Khao Chi Chan Buddha Image, Chonburi. All not to scale." figureDoi="10.3897/zookeys.1119.85400.figure25" httpUri="https://binary.pensoft.net/fig/735386" pageId="0" pageNumber="1">25C</figureCitation>
|
|
) from
|
|
<location id="7F331EE4AF9DC1C59C0918F27452E889" LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:7F331EE4AF9DC1C59C0918F27452E889" country="Myanmar" county="Surat Thani Province" municipality="Ko Samui District" name="Wua Ta Lap Island">Wua Ta Lap Island</location>
|
|
,
|
|
<collectingMunicipality id="6516297A79A6DCB10E0B4157A7F71625">Ko Samui District</collectingMunicipality>
|
|
,
|
|
<collectingCounty id="7AFB8A40B8502D41857A388FC5D31999">Surat Thani Province</collectingCounty>
|
|
,
|
|
<collectingDate id="564C51E156B0CC734017A0FFA92E5010" value="2009-06-06">6 June 2009</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="66C23CF7037FB03F17FCEBAD2612986F" collectingDate="2006-10-06" collectorName="Figs" country="Myanmar" location="Mueang Krabi District" municipality="Krabi Province" specimenCount="1">
|
|
CUMZ 12038 (4 shells;
|
|
<collectorName id="1E26449499819B34C12E29BEF88889E2">Figs</collectorName>
|
|
<figureCitation id="80970DBF6E7CBC73F9CFD3BF118F24ED" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Shells of Pupina artata species group from mainland Southeast Asia A-M Pupina artata A syntype UMZC I. 102960. A and specimens B CUMZ 12011 C CUMZ 12023 D CUMZ 12038 E CUMZ 12026 F CUMZ 12015 G CUMZ 12009 H CUMZ 12001 I CUMZ 12003 J NHMUK 1906.4.4.28 K CUMZ 12037 L CUMZ 12031, and M CUMZ 12018 N, O Pupina pallens N lectotype SMF 109951 and O specimen CUMZ 12042 P-R Pupina limitanea P, Q syntypes NHMUK 1903.7.1.2967 and R specimen CUMZ 12044 S Pupina billeti, holotype MNHN-IM- 2000 - 35841 T Pupina verneaui, syntype MNHN-IM- 2000 - 35843 U, V Pupina hungerfordiana, specimens NHMUK 91.3.14.686 - 7 W, X Pupina bensoni sp. nov. W holotype CUMZ 12045 / 1 and X paratype CUMZ 12046 / 1. Photo: H. Taylor, NHM (A, P, Q), P. Maestrati, MNHN (S, T)." figureDoi="10.3897/zookeys.1119.85400.figure21" httpUri="https://binary.pensoft.net/fig/735382" pageId="0" pageNumber="1">21D</figureCitation>
|
|
,
|
|
<figureCitation id="EA60056A5F1F3AB46EC5BD530A5B1C53" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. A Pupina artata, specimen CUMZ 12038 from Tham Suea Temple, Krabi B, C Pupina pallens B lectotype SMF 109951 and C specimen CUMZ 12042 from Suan Wiwek Bureau of Monks, Prachuap Khiri Khan D-F Pupina limitanea D, E syntypes NHMUK 1903.7.1.2967 from East of Burma & Siam and F specimen CUMZ 12044 from Mae Lana junction, Mae Hong Son G Pupina bensoni sp. nov., holotype CUMZ 12045 / 1. Photo: H. Taylor, NHM (D, E)." figureDoi="10.3897/zookeys.1119.85400.figure24" httpUri="https://binary.pensoft.net/fig/735385" pageId="0" pageNumber="1">24A</figureCitation>
|
|
) from Tham Suea Temple,
|
|
<location id="F1E617E913559D5635B6CC7F38A6E8CE" LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:F1E617E913559D5635B6CC7F38A6E8CE" country="Myanmar" municipality="Krabi Province" name="Mueang Krabi District">Mueang Krabi District</location>
|
|
,
|
|
<collectingMunicipality id="2F644289636E2C807A0049FC0FA2583A">Krabi Province</collectingMunicipality>
|
|
,
|
|
<collectingDate id="55BB756D74904FCC2846C431028F80FF" value="2006-10-06">6 Oct. 2006</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="36B63E2687A27493CC17588EA3D6545D" collectingDate="2015-08-31" collectorName="Khao Noi Phothiyan Temple" country="Myanmar" location="Mueang Satul District" municipality="Satul Province" specimenCount="1">
|
|
CUMZ 12039 (4 shells) from
|
|
<collectorName id="FBC2DC1AE0421B8FF6652CD323544332">Khao Noi Phothiyan Temple</collectorName>
|
|
,
|
|
<location id="C8F64BF6B66466976826ECBB7AC02F35" LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:C8F64BF6B66466976826ECBB7AC02F35" country="Myanmar" municipality="Satul Province" name="Mueang Satul District">Mueang Satul District</location>
|
|
,
|
|
<collectingMunicipality id="1B1E2B68A31951C934B5F75F17FE5450">Satul Province</collectingMunicipality>
|
|
,
|
|
<collectingDate id="3878ABA09BD432E6B237971351B23A39" value="2015-08-31">31 Aug. 2015</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="E9506181559B4232C519C29B77CC1551" collectingDate="2007-01-23" collectorName="Khao Rup Chang" country="Myanmar" location="Mueang Songkhla District" municipality="Songkhla Province" specimenCount="1">
|
|
CUMZ 12040 (1 shell) from
|
|
<collectorName id="C94DF859DCC8FEDFD9712FE7567B0CF9">Khao Rup Chang</collectorName>
|
|
,
|
|
<location id="F603591020FB75FEFD9BBF0B77EBA8AD" LSID="urn:lsid:plazi:treatment:38446D96E8785C33B44B278839E24F7B:F603591020FB75FEFD9BBF0B77EBA8AD" country="Myanmar" municipality="Songkhla Province" name="Mueang Songkhla District">Mueang Songkhla District</location>
|
|
,
|
|
<collectingMunicipality id="88C955FE4F1BFD5ECDACB5C4EDADAEDF">Songkhla Province</collectingMunicipality>
|
|
,
|
|
<collectingDate id="68464C47F7B256C871D45B453893EB58" value="2007-01-23">23 Jan. 2007</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="96BB5F5388743E3C046D1B87C74E4FAE" pageId="0" pageNumber="1" type="diagnosis">
|
|
<paragraph id="8496248E33C4D7C568C9BD319835A334" pageId="0" pageNumber="1">Diagnosis.</paragraph>
|
|
<paragraph id="D6A2E901191CB42768C0F316E2911C86" pageId="0" pageNumber="1">Shell ovate; last whorl ca. three quarters of shell height. Apertural lip slightly thickened, not expanded. Both parietal and columellar teeth fin-shaped and slightly thickened; parietal tooth covering posterior canal; columellar tooth next to slit-like anterior canal.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="D5952AEE463E3E4CD5D430C718983F71" pageId="0" pageNumber="1" type="differential diagnosis">
|
|
<paragraph id="0E163DDA0586D5D135DED1188305D57A" pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
|
|
<paragraph id="574CA3529B596F0C9F6355B952F3BF06" pageId="0" pageNumber="1">
|
|
<taxonomicName id="4FD5C162AA0CF82EDE78D76F3B33B6A2" authorityName="Benson" authorityYear="1856" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina artata" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="artata">
|
|
<emphasis id="7449D03037DC07881E5819C72C1DA756" italics="true" pageId="0" pageNumber="1">Pupina artata</emphasis>
|
|
</taxonomicName>
|
|
is most similar to
|
|
<taxonomicName id="98900BB93D0089DA7491AC97B5047960" class="Gastropoda" family="Pupinidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. pallens" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="pallens">
|
|
<emphasis id="7B7BEFD37915020C540A9AA0E749532E" italics="true" pageId="0" pageNumber="1">P. pallens</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="0458C7201F1A0CFD5A52A74060224424" class="Gastropoda" family="Pupinidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. limitanea" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="limitanea">
|
|
<emphasis id="AD63A4859C1F1159BBCF4C3619B805D8" italics="true" pageId="0" pageNumber="1">P. limitanea</emphasis>
|
|
</taxonomicName>
|
|
in shell shape, but different from
|
|
<taxonomicName id="D7976833D82EA0F0B49A586FC2C9BC08" class="Gastropoda" family="Pupinidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. pallens" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="pallens">
|
|
<emphasis id="CFDD4BEAD45EEC8D833D36F81F411681" italics="true" pageId="0" pageNumber="1">P. pallens</emphasis>
|
|
</taxonomicName>
|
|
in that the basal position of the apertural lip is not widened, and different from
|
|
<taxonomicName id="064ED92C3F2F613CCC4D1A83E1480B64" class="Gastropoda" family="Pupinidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. limitanea" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="limitanea">
|
|
<emphasis id="57F476A479478479EB9EB442109CC014" italics="true" pageId="0" pageNumber="1">P. limitanea</emphasis>
|
|
</taxonomicName>
|
|
by a longer last whorl, and parietal and columellar teeth and apertural lip less thickened.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="D95BFB4183703CEBF7117E6BC7076CA2" pageId="0" pageNumber="1" type="distribution">
|
|
<paragraph id="58CA3919CE7BC51D84AF51854CEA7776" pageId="0" pageNumber="1">Distribution.</paragraph>
|
|
<paragraph id="F806AE6AED4DDA15EEB909B6570EA060" pageId="0" pageNumber="1">
|
|
Peninsular Malaysia, Myanmar (
|
|
<bibRefCitation id="3BFB12A7DF4C3A7ADE219040505BDB90" author="Laidlaw, FF" journalOrPublisher="Journal of the Malaysian Branch of the Royal Asiatic Society" pageId="0" pageNumber="1" pagination="25 - 37" refId="B82" refString="Laidlaw, FF, 1928. A list of the land and fresh-water mollusca of the Malay Peninsula with notes. Journal of the Malaysian Branch of the Royal Asiatic Society 6: 25 - 37" title="A list of the land and fresh-water mollusca of the Malay Peninsula with notes." volume="6" year="1928">Laidlaw 1928</bibRefCitation>
|
|
;
|
|
<bibRefCitation id="BAE76A2C5695EEE9AA9D31DAFCF659BF" author="Solem, A" journalOrPublisher="Spolia Zoologica Musei Hauniensis" pageId="0" pageNumber="1" pagination="1 - 110" refId="B146" refString="Solem, A, 1966. Some non-marine mollusks from Thailand, with notes on classification of the Helicarionidae. Spolia Zoologica Musei Hauniensis 24: 1 - 110" title="Some non-marine mollusks from Thailand, with notes on classification of the Helicarionidae." volume="24" year="1966">Solem 1966</bibRefCitation>
|
|
), and throughout Thailand except in the northeastern region.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="75621C09E51DE80006BB3C78EFE06F03" pageId="0" pageNumber="1" type="remarks">
|
|
<paragraph id="16E46E2E356530D3273D530F007CA7AF" pageId="0" pageNumber="1">Remarks.</paragraph>
|
|
<paragraph id="8DC7B15D004449829B72A607C039D904" pageId="0" pageNumber="1">
|
|
The type specimen of
|
|
<taxonomicName id="4F2ECC4475223B082184D7AF162991F4" class="Gastropoda" family="Pupinidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. artata subsp. blanfordiana" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="artata" subSpecies="blanfordiana">
|
|
<emphasis id="B00962A458A433F76551BF1A0BB915E6" italics="true" pageId="0" pageNumber="1">P. artata blanfordiana</emphasis>
|
|
</taxonomicName>
|
|
could not be located, so the validity of this subspecies is still unknown. The specimen identified as
|
|
<taxonomicName id="94C38C8B1DD2C14CD94D1A93B948F8D8" class="Gastropoda" family="Pupinidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. peguensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peguensis">
|
|
<emphasis id="79C30C779ED91F91DF87501C337F31FF" italics="true" pageId="0" pageNumber="1">P. peguensis</emphasis>
|
|
</taxonomicName>
|
|
and figured in
|
|
<bibRefCitation id="E1EDDAEAB926EAAFD10B87A4E63FCC43" author="Godwin-Austen, HH" journalOrPublisher="Journal of Malacology" pageId="0" pageNumber="1" refId="B51" refString="Godwin-Austen, HH, 1897. Land and freshwater mollusca of India, including South Arabia, Baluchistan, Afghanistan, Kashmir, Nepal, Burma, Pegu, Tenasserim, Malaya Peninsula, Ceylon and other islands of the Indian Ocean. Vol. 2, Part 7: 1-46." title="Land and freshwater mollusca of India, including South Arabia, Baluchistan, Afghanistan, Kashmir, Nepal, Burma, Pegu, Tenasserim, Malaya Peninsula, Ceylon and other islands of the Indian Ocean. Vol. 2, Part 7: 1 - 46." year="1897">Godwin-Austen (1897</bibRefCitation>
|
|
: pl. 69, fig. 3, 3a-d) from Kama on the right bank of the Irrawaddy River, Pegu is different from the holotype of
|
|
<taxonomicName id="154C6092DD86B38D8F387DD9DC18F81F" class="Gastropoda" family="Pupinidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. peguensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="peguensis">
|
|
<emphasis id="E60F568CDA8AE31F57295E076BE31BAB" italics="true" pageId="0" pageNumber="1">P. peguensis</emphasis>
|
|
</taxonomicName>
|
|
(see
|
|
<bibRefCitation id="0E66EDC1F6D81CD6E87A6F2D7D1C9D13" author="Tripathy, B" journalOrPublisher="Records of the Zoological Survey of India" pageId="0" pageNumber="1" pagination="508 - 510" refId="B177" refString="Tripathy, B, Sajan, S, 2019. Pupina peguensis Benson, 1860 (Mollusca: Pupinidae): Tracing a long lost molluscan type. Records of the Zoological Survey of India 119: 508 - 510" title="Pupina peguensis Benson, 1860 (Mollusca: Pupinidae): Tracing a long lost molluscan type." volume="119" year="2019">Tripathy and Sajan 2019</bibRefCitation>
|
|
), but similar to the type specimen of
|
|
<taxonomicName id="FD318473C7B2C09F4514C97272C0EFA7" class="Gastropoda" family="Pupinidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. artata" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="artata">
|
|
<emphasis id="44DBB7FED64B94746EF6956D395BA813" italics="true" pageId="0" pageNumber="1">P. artata</emphasis>
|
|
</taxonomicName>
|
|
. Thus, this specimen is herein identified as
|
|
<taxonomicName id="7E81801B2CE1B6D62B21FF4AD41436AC" class="Gastropoda" family="Pupinidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. artata" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="artata">
|
|
<emphasis id="076B9B9E0704A0E71ED97BB9718D3F95" italics="true" pageId="0" pageNumber="1">P. artata</emphasis>
|
|
</taxonomicName>
|
|
.
|
|
</paragraph>
|
|
<paragraph id="C1EE2CEBD383E59521169667F6E9378B" pageId="0" pageNumber="1">
|
|
The specimen of
|
|
<taxonomicName id="295EADCC5BB1494ED18BC5A9624D0140" class="Gastropoda" family="Pupinidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. artata" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="artata">
|
|
<emphasis id="19824ADF42DA07E318C12ADAF1738209" italics="true" pageId="0" pageNumber="1">P. artata</emphasis>
|
|
</taxonomicName>
|
|
figured in
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<bibRefCitation id="87410A304802BFEB4DFB178BBED28144" author="Maassen, WJM" editor="Falkner, M" journalOrPublisher="ConchBooks, Hackenheim" pageId="0" pageNumber="1" pagination="277 - 290" refId="B88" refString="Maassen, WJM, 2002. Remarks on the Pupinidae of Sumatra (Indonesia) with description of two new species (Gastropoda, Prosobranchia: Pupinidae). In: Falkner, M, Groh, K, Speight, MCD, Eds., Collectanea Malacologica Festschrift fuer Gerhard Falkner. ConchBooks, Hackenheim: 277 - 290" title="Remarks on the Pupinidae of Sumatra (Indonesia) with description of two new species (Gastropoda, Prosobranchia: Pupinidae)." volumeTitle="Collectanea Malacologica Festschrift fuer Gerhard Falkner." year="2002">Maassen (2002</bibRefCitation>
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: text-fig. 3) from Sumatra should constitute a different species as it is different from the syntype figured here in having a smaller, sharper parietal tooth revealing the posterior canal and an ear-lobe-shaped columellar tooth covering the anterior canal. Thus, those specimens should belong to the
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<taxonomicName id="10DB953F9B01761E735CEB2BD1395703" class="Gastropoda" family="Pupinidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. arula" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="arula">
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<emphasis id="10A02D68A0178765ED8D3E99EDFB938B" italics="true" pageId="0" pageNumber="1">P. arula</emphasis>
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</taxonomicName>
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species group instead (see below).
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</paragraph>
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<paragraph id="325E4580CE631B2FF1C2E9C8EDE3C815" pageId="0" pageNumber="1">
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All specimens from Thailand with a slightly thickened, fin-shaped parietal tooth covering the posterior canal are herein identified as
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<taxonomicName id="F4533E3F54A87A099BA3BEAEAF4FE8FD" class="Gastropoda" family="Pupinidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. artata" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="artata">
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<emphasis id="ADD813F03D375357344D448D07B49152" italics="true" pageId="0" pageNumber="1">P. artata</emphasis>
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</taxonomicName>
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. However, these specimens exhibit a variable shell size (smaller with shell height 5.4 mm, shell width 3.5 mm, to larger with shell height 8.4 mm; shell width 5.9 mm; Fig.
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<figureCitation id="EF0F77B4AEF3F3DAD30CA6462018E072" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Shells of Pupina artata species group from mainland Southeast Asia A-M Pupina artata A syntype UMZC I. 102960. A and specimens B CUMZ 12011 C CUMZ 12023 D CUMZ 12038 E CUMZ 12026 F CUMZ 12015 G CUMZ 12009 H CUMZ 12001 I CUMZ 12003 J NHMUK 1906.4.4.28 K CUMZ 12037 L CUMZ 12031, and M CUMZ 12018 N, O Pupina pallens N lectotype SMF 109951 and O specimen CUMZ 12042 P-R Pupina limitanea P, Q syntypes NHMUK 1903.7.1.2967 and R specimen CUMZ 12044 S Pupina billeti, holotype MNHN-IM- 2000 - 35841 T Pupina verneaui, syntype MNHN-IM- 2000 - 35843 U, V Pupina hungerfordiana, specimens NHMUK 91.3.14.686 - 7 W, X Pupina bensoni sp. nov. W holotype CUMZ 12045 / 1 and X paratype CUMZ 12046 / 1. Photo: H. Taylor, NHM (A, P, Q), P. Maestrati, MNHN (S, T)." figureDoi="10.3897/zookeys.1119.85400.figure21" httpUri="https://binary.pensoft.net/fig/735382" pageId="0" pageNumber="1">21A-M</figureCitation>
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). The shell shape is also variable from ovate which is similar to the syntype (Fig.
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<figureCitation id="3CA08CA22012992BEFDD0C0766816387" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Shells of Pupina artata species group from mainland Southeast Asia A-M Pupina artata A syntype UMZC I. 102960. A and specimens B CUMZ 12011 C CUMZ 12023 D CUMZ 12038 E CUMZ 12026 F CUMZ 12015 G CUMZ 12009 H CUMZ 12001 I CUMZ 12003 J NHMUK 1906.4.4.28 K CUMZ 12037 L CUMZ 12031, and M CUMZ 12018 N, O Pupina pallens N lectotype SMF 109951 and O specimen CUMZ 12042 P-R Pupina limitanea P, Q syntypes NHMUK 1903.7.1.2967 and R specimen CUMZ 12044 S Pupina billeti, holotype MNHN-IM- 2000 - 35841 T Pupina verneaui, syntype MNHN-IM- 2000 - 35843 U, V Pupina hungerfordiana, specimens NHMUK 91.3.14.686 - 7 W, X Pupina bensoni sp. nov. W holotype CUMZ 12045 / 1 and X paratype CUMZ 12046 / 1. Photo: H. Taylor, NHM (A, P, Q), P. Maestrati, MNHN (S, T)." figureDoi="10.3897/zookeys.1119.85400.figure21" httpUri="https://binary.pensoft.net/fig/735382" pageId="0" pageNumber="1">21A</figureCitation>
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), to more globose (Fig.
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<figureCitation id="21871A225157C8360FBE8C745E186028" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Shells of Pupina artata species group from mainland Southeast Asia A-M Pupina artata A syntype UMZC I. 102960. A and specimens B CUMZ 12011 C CUMZ 12023 D CUMZ 12038 E CUMZ 12026 F CUMZ 12015 G CUMZ 12009 H CUMZ 12001 I CUMZ 12003 J NHMUK 1906.4.4.28 K CUMZ 12037 L CUMZ 12031, and M CUMZ 12018 N, O Pupina pallens N lectotype SMF 109951 and O specimen CUMZ 12042 P-R Pupina limitanea P, Q syntypes NHMUK 1903.7.1.2967 and R specimen CUMZ 12044 S Pupina billeti, holotype MNHN-IM- 2000 - 35841 T Pupina verneaui, syntype MNHN-IM- 2000 - 35843 U, V Pupina hungerfordiana, specimens NHMUK 91.3.14.686 - 7 W, X Pupina bensoni sp. nov. W holotype CUMZ 12045 / 1 and X paratype CUMZ 12046 / 1. Photo: H. Taylor, NHM (A, P, Q), P. Maestrati, MNHN (S, T)." figureDoi="10.3897/zookeys.1119.85400.figure21" httpUri="https://binary.pensoft.net/fig/735382" pageId="0" pageNumber="1">21F</figureCitation>
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) or more elongate (Fig.
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<figureCitation id="4ED6EC20B9426D1CF9A7984F9A0FCF8A" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Shells of Pupina artata species group from mainland Southeast Asia A-M Pupina artata A syntype UMZC I. 102960. A and specimens B CUMZ 12011 C CUMZ 12023 D CUMZ 12038 E CUMZ 12026 F CUMZ 12015 G CUMZ 12009 H CUMZ 12001 I CUMZ 12003 J NHMUK 1906.4.4.28 K CUMZ 12037 L CUMZ 12031, and M CUMZ 12018 N, O Pupina pallens N lectotype SMF 109951 and O specimen CUMZ 12042 P-R Pupina limitanea P, Q syntypes NHMUK 1903.7.1.2967 and R specimen CUMZ 12044 S Pupina billeti, holotype MNHN-IM- 2000 - 35841 T Pupina verneaui, syntype MNHN-IM- 2000 - 35843 U, V Pupina hungerfordiana, specimens NHMUK 91.3.14.686 - 7 W, X Pupina bensoni sp. nov. W holotype CUMZ 12045 / 1 and X paratype CUMZ 12046 / 1. Photo: H. Taylor, NHM (A, P, Q), P. Maestrati, MNHN (S, T)." figureDoi="10.3897/zookeys.1119.85400.figure21" httpUri="https://binary.pensoft.net/fig/735382" pageId="0" pageNumber="1">21L</figureCitation>
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). In addition, these specimens exhibit a variation in length, outer curvature and thickness of the parietal tooth, and body colour. There is also a case of different shell colour morphs (brown and grey) within the same population (Fig.
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<figureCitation id="FF4A18818D7CA50A946B48CB39126668" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Live specimens of A, B Pollicaria mouhoti mouhoti: specimens A CUMZ 12166 and B CUMZ 12175 from Wang Daeng Cave, Phitsanulok C Pollicaria mouhoti monochroma, paratype CUMZ 1562 from Tam Pha Bing Temple, Loei D Tortulosa tortuosa, specimen CUMZ 12155 from Tham Suea Temple, Krabi E-H Pupina artata: specimens of E CUMZ 12006 from Pha Daeng Cave, Mae Hong Son F CUMZ 12008 from Tham Nam Pha Pha Ngam Temple, Lampang, and G, H CUMZ 12029 from Khao Tham Raet Temple, Chachoengsao showing the brown (G) and grey (H) shell morphs; All not to scale." figureDoi="10.3897/zookeys.1119.85400.figure8" httpUri="https://binary.pensoft.net/fig/735369" pageId="0" pageNumber="1">8G, H</figureCitation>
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). Therefore, DNA data is needed to reveal the extent of genetic differentiation or cryptic diversity within the
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<taxonomicName id="AC3D18C167E1968A17E989CF8C80E721" class="Gastropoda" family="Pupinidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. artata" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="artata">
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<emphasis id="D87CD4585C3D4A54190E16E580C432CB" italics="true" pageId="0" pageNumber="1">P. artata</emphasis>
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</taxonomicName>
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morphotype.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |