treatments-xml/data/2E/BA/8E/2EBA8EB5F4A818324C85FCE46395A1ED.xml

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<mods:title id="93C853FE60B67421E2617B2397E41C22">Playing hard to get: two new species of subterranean Trechini beetles (Coleoptera, Carabidae, Trechinae) from the Dinaric Karst</mods:title>
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<mods:namePart id="AC5C9749EB5D05F2B56BCAA6EA9332E8">Lohaj, Roman</mods:namePart>
<mods:affiliation id="C39EDD5E79B1A8157DF0F1131572FC8C">Zahradna 61 A, SK- 900 91 Limbach, Slovakia</mods:affiliation>
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<mods:namePart id="5E8972DD5309163D98C10AE00061A53E">Delic, Teo</mods:namePart>
<mods:affiliation id="BE5A584D212CC3CAFBECF0A2DCF4EAFE">SubBioLab, Department of Biology, Biotechnical Faculty, University of Ljubljana, Jamnikarjeva 101, SI- 1000 Ljubljana, Slovenia</mods:affiliation>
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<treatment id="2EBA8EB5F4A818324C85FCE46395A1ED" ID-GBIF-Taxon="182223588" LSID="urn:lsid:zoobank.org:act:BCB33301-A988-465D-A0F5-82060EAD16B0" httpUri="http://treatment.plazi.org/id/2EBA8EB5F4A818324C85FCE46395A1ED" lastPageNumber="1" pageId="0" pageNumber="1" scope_family="Carabidae" scope_order="Coleoptera" scope_subFamily="Trechinae" scope_tribe="Trechini">
<subSubSection id="E4CA517408152864136B4C0F5B7D7C6C" pageId="0" pageNumber="1" type="nomenclature">
<paragraph id="D66982A3227DD9F16A98B19EE8CD7903" pageId="0" pageNumber="1">
<taxonomicName id="AF37F2E4DA07BE1A4D6B14C0C3829BCD" ID-CoL="8LXQL" LSID="http://zoobank.org/BCB33301-A988-465D-A0F5-82060EAD16B0" authority="Lohaj &amp; Delić, 2019" authorityName="Lohaj &amp; Delić" authorityYear="2019" class="Insecta" family="Carabidae" genus="Derossiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Derossiella lukici" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="lukici" status="sp. n.">Derossiella lukici</taxonomicName>
<taxonomicNameLabel id="1917063DFF786FDE48862BAC33988785" pageId="0" pageNumber="1">sp. n.</taxonomicNameLabel>
<figureCitation id="982A9220FEDB4E41FD666AA551D39640" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Geographical distribution of the Dinaric aphaenopsoid trechines of the genera Derossiella and Adriaphenops. Source of data: http: // subbio. net / db /" figureDoi="10.3897/dez.66.31754.figure2" httpUri="https://binary.pensoft.net/fig/258009" pageId="0" pageNumber="1">Figs 2</figureCitation>
<figureCitation id="317082336FA197A8AE357C5B4818B926" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Derossiella lukici sp. n. in its habitat in the cave Biokovka, Golubinjak, Mt Biokovo (Photo courtesy of P. Bregovic)." figureDoi="10.3897/dez.66.31754.figure3" httpUri="https://binary.pensoft.net/fig/258010" pageId="0" pageNumber="1">, 3</figureCitation>
<figureCitation id="AF718DE8D3DE0978449FE5514021874C" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Habitus of Derosiella species. Derossiella lukici sp. n. (a) and Derossiella nonveilleri (b). Chetotaxy is presented as white points (photo courtesy of Dusan Beno)." figureDoi="10.3897/dez.66.31754.figure4" httpUri="https://binary.pensoft.net/fig/258011" pageId="0" pageNumber="1">, 4</figureCitation>
<figureCitation id="57DC4F46539994751E11649E1D1810DB" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male and female genitalia of Derossiella representatives: aedeagus of D. lukici sp. n., dorsal view (a); and lateral view (b); female genitalia gonocoxite 1 and 2 (basal and apical segments of gonostylus) copulatory piece of D. lukici sp. n. (c); and D. nonveilleri, lateral view of male aedeagus with parameres detached (d) (illustration by Fedor Ciampor)." figureDoi="10.3897/dez.66.31754.figure5" httpUri="https://binary.pensoft.net/fig/258012" pageId="0" pageNumber="1">, 5</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="BA6C641033BD41E18A40B03B434EDA05" pageId="0" pageNumber="1" type="type series">
<paragraph id="E352215A5BABBAA76581043631C7CD3B" pageId="0" pageNumber="1">Type series.</paragraph>
<paragraph id="F5D7C562CA690A6B2CD4234D9CBA4239" pageId="0" pageNumber="1">
Holotype male labelled: &quot;Croatia, Mt Biokovo, Golubinjak, Biokovka, -300 m, 24.6.2017, T. 
<normalizedToken id="2ABF048A23FC51E2793D0373B7684954" originalValue="Delić">Delic</normalizedToken>
lgt.&quot; (white label, printed) / &quot;DNA extraction RL-07&quot; (orange label, printed) / &quot;HOLOTYPUS 
<taxonomicName id="058756B81B5AE2B1837F64BE7E21618B" authorityName="Lohaj &amp; Delić" authorityYear="2019" class="Insecta" family="Carabidae" genus="Derossiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Derossiella lukici" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="lukici">
<emphasis id="582A02AC3AAFD994119F0A86BA71AC89" italics="true" pageId="0" pageNumber="1">Derossiella lukici</emphasis>
</taxonomicName>
sp. n. Lohaj &amp; 
<normalizedToken id="5E903FD9ED354B44C03113D59C7A1BA4" originalValue="Delić">Delic</normalizedToken>
des. 2018&quot; (red label, printed), (CNHM, voucher code 600: ZAG; ZEC2, 4194 Coll. 
<normalizedToken id="0B9F190BB3FF6BDD9FF2430683A8F020" originalValue="Jalžić">Jalzic</normalizedToken>
). Paratypes: one female (right posterior tarsus missing) labelled: &quot;Croatia, Mt Biokovo, Golubinjak, Biokovka, 2.9.2007, M. 
<normalizedToken id="3D5416F8ED9137FB1D12261299AD0E01" originalValue="Lukić">Lukic</normalizedToken>
lgt.&quot; (white label, printed) / &quot;PARATYPUS 
<taxonomicName id="CEAA4AB431DD5FDA183317FE416E2943" authorityName="Lohaj &amp; Delić" authorityYear="2019" class="Insecta" family="Carabidae" genus="Derossiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Derossiella lukici" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="lukici">
<emphasis id="C5A7C85BD63D672A36E29D4301EE9D4C" italics="true" pageId="0" pageNumber="1">Derossiella lukici</emphasis>
</taxonomicName>
sp. n. Lohaj &amp; 
<normalizedToken id="5E723CC50EFB8D75B79CDF42A3D7F431" originalValue="Delić">Delic</normalizedToken>
des. 2018&quot; (red label, printed), (CNHM, voucher code 600: ZAG; ZEC2, 4195 Coll. 
<normalizedToken id="D586378885C687754AB22A864187DBA2" originalValue="Jalžić">Jalzic</normalizedToken>
)., one female (last three antennomeres of right antenna missing) labelled: &quot;Croatia, Mt Biokovo, Lokva, Pretnerova, -120 m, 19.05.2015, E. Premate lgt.&quot; (white label, printed) / &quot;PARATYPUS 
<taxonomicName id="E7250DBF75B1F493E28A43845834E5D5" authorityName="Lohaj &amp; Delić" authorityYear="2019" class="Insecta" family="Carabidae" genus="Derossiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Derossiella lukici" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="lukici">
<emphasis id="F462E682E7FF9096022C346F4BA22212" italics="true" pageId="0" pageNumber="1">Derossiella lukici</emphasis>
</taxonomicName>
sp. n. Lohaj &amp; 
<normalizedToken id="8CEDA96011A96482D1FAF61E1E4D98CE" originalValue="Delić">Delic</normalizedToken>
des. 2018&quot; (red label, printed), (CRL).
</paragraph>
</subSubSection>
<subSubSection id="A40149E3D5681EFFCCC18248CB0FCBF6" pageId="0" pageNumber="1" type="diagnosis">
<paragraph id="D3D36241994363B6E45E00C1652E5F5C" pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph id="8ECA2802F291679308FD68FBFCECA77F" pageId="0" pageNumber="1">
Medium-sized aphaenopsoid trechine with morphological features fully matching generic description proposed by 
<bibRefCitation id="7DC1208F6B244294934C1DB0E7364953" author="Queinnec, E" editor="Pavicevic, D" journalOrPublisher="Institute for Nature Conservation of Serbia, Belgrade" pageId="0" pageNumber="1" pagination="157 - 176" refId="B44" refString="Queinnec, E, 2008. Two new genera and species of 'aphaenopsid' cave beetles from the Balkan Peninsula and considerations about the evolutionary trends of the so-called &quot;hyper-specialised&quot; trechines. In: Pavicevic, D, Perreau, M, Eds., Advances in the Studies of the Fauna of the Balkan Peninsula. Institute for Nature Conservation of Serbia, Belgrade: 157 - 176" title="Two new genera and species of ' aphaenopsid' cave beetles from the Balkan Peninsula and considerations about the evolutionary trends of the so-called &quot; hyper-specialised &quot; trechines." volumeTitle="Advances in the Studies of the Fauna of the Balkan Peninsula." year="2008">
<normalizedToken id="47CD02B56247DEB2050A7CA3D8EE543C" originalValue="Quéinnec">Queinnec</normalizedToken>
(2008)
</bibRefCitation>
. Head long, parallel-sided, with complete, deep frontal furrows reaching neck constriction and two pairs of supraorbital setae, posterior setae doubled. Eyes absent, mentum with simple tooth. Aphaenopsoid habitus with very narrow head and pronotum; elytra oblong-oval, 2.6 times wider than head and pronotum. Hind angles of pronotum without setae. Surface completely glabrous, striae absent, elytral chaetotaxy with macrochetae and microchetae. Cuticle depigmented, reddish-yellow, body strongly flattened dorso-ventrally, with very long and slender legs and antennae (Figs 
<figureCitation id="43F97B1B9428D0ABEA29939881C2593D" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Derossiella lukici sp. n. in its habitat in the cave Biokovka, Golubinjak, Mt Biokovo (Photo courtesy of P. Bregovic)." figureDoi="10.3897/dez.66.31754.figure3" httpUri="https://binary.pensoft.net/fig/258010" pageId="0" pageNumber="1">3</figureCitation>
, 
<figureCitation id="A59093D63DC4486134CE1C38C7B3F1A4" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Habitus of Derosiella species. Derossiella lukici sp. n. (a) and Derossiella nonveilleri (b). Chetotaxy is presented as white points (photo courtesy of Dusan Beno)." figureDoi="10.3897/dez.66.31754.figure4" httpUri="https://binary.pensoft.net/fig/258011" pageId="0" pageNumber="1">4</figureCitation>
). Closely related to the type species of the genus, 
<taxonomicName id="1EC77F998654AE96B23C356931512A17" authorityName="Queinnec" authorityYear="2008" class="Insecta" family="Carabidae" genus="Derossiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Derossiella nonveilleri" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="nonveilleri">
<emphasis id="F6EE26D9E5134613FCBB2D7CB23B33E7" italics="true" pageId="0" pageNumber="1">Derossiella nonveilleri</emphasis>
</taxonomicName>
, from which it differs by paired posterior supraorbital setiferous punctures, differently shaped elytra with higher number of macro- and microchetae, narrower pronotum, as well as by the differently shaped aedeagus.
</paragraph>
<caption id="34D742D071194540520F7125C75ADE3B" ID-Zenodo-Dep="2551937" doi="10.3897/dez.66.31754.figure3" httpUri="https://binary.pensoft.net/fig/258010" pageId="0" pageNumber="1" start="Figure 3" startId="F3">
<paragraph id="135912F94325AE641BD2A9DD94875C8E" pageId="0" pageNumber="1">
<emphasis id="A1E7AF7C179FCFAFC5D7A30CA042BD99" bold="true" pageId="0" pageNumber="1">Figure 3.</emphasis>
<taxonomicName id="6EE76E1511D3239B6B4C6C3C3B14DE6F" authorityName="Lohaj &amp; Delić" authorityYear="2019" class="Insecta" family="Carabidae" genus="Derossiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Derossiella lukici" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="lukici">
<emphasis id="7158C0EDD7B3CE97709A1D214629B657" italics="true" pageId="0" pageNumber="1">Derossiella lukici</emphasis>
</taxonomicName>
sp. n. in its habitat in the cave Biokovka, Golubinjak, Mt Biokovo (Photo courtesy of P. 
<normalizedToken id="89F511EBD5F3C2D79EFA8A3A1F2BB58A" originalValue="Bregović">Bregovic</normalizedToken>
).
</paragraph>
</caption>
<caption id="42CF329CE50E9509E2F44555ED4C9174" ID-Zenodo-Dep="2551939" doi="10.3897/dez.66.31754.figure4" httpUri="https://binary.pensoft.net/fig/258011" pageId="0" pageNumber="1" start="Figure 4" startId="F4">
<paragraph id="B31FF08B48295B05BC95DD6139D20AC3" pageId="0" pageNumber="1">
<emphasis id="6303986AC59ABEFDB3AE846FFF0F6F9D" bold="true" pageId="0" pageNumber="1">Figure 4.</emphasis>
Habitus of 
<taxonomicName id="81516986E41FA6F34A8862C92B3FBF61" genus="Derosiella" lsidName="Derosiella" pageId="0" pageNumber="1" rank="genus">
<emphasis id="2421B194D6D098DA97F4DBF397AE045F" italics="true" pageId="0" pageNumber="1">Derosiella</emphasis>
</taxonomicName>
species. 
<taxonomicName id="2A22F03723368D3AA2355FDD8C39AB23" authorityName="Lohaj &amp; Delić" authorityYear="2019" class="Insecta" family="Carabidae" genus="Derossiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Derossiella lukici" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="lukici">
<emphasis id="1B1F79D8FEF072B8BBC4A3D3E47D02B0" italics="true" pageId="0" pageNumber="1">Derossiella lukici</emphasis>
</taxonomicName>
sp. n. 
<emphasis id="A327A89B9690859D3672DF3A8122E73E" bold="true" pageId="0" pageNumber="1">(a)</emphasis>
and 
<taxonomicName id="90D49FEE886463B157B50D3D39052988" authorityName="Queinnec" authorityYear="2008" class="Insecta" family="Carabidae" genus="Derossiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Derossiella nonveilleri" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="nonveilleri">
<emphasis id="91D84774425F27FC840062D149ECBBB8" italics="true" pageId="0" pageNumber="1">Derossiella nonveilleri</emphasis>
</taxonomicName>
<emphasis id="62C11AF86DD32D2951B26D13B26DC7C9" bold="true" pageId="0" pageNumber="1">(b)</emphasis>
. Chetotaxy is presented as white points (photo courtesy of 
<normalizedToken id="3F739BBAC08A6BF935176D63C22B25CA" originalValue="Dušan">Dusan</normalizedToken>
<normalizedToken id="9008836E291FE7E47D12070AA6670F9D" originalValue="Beňo">Beno</normalizedToken>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="89B376AE1E2AEF751E616A282358DD8B" pageId="0" pageNumber="1" type="description">
<paragraph id="178DD5A88B810746A1F12B09BF8AB51A" pageId="0" pageNumber="1">Description.</paragraph>
<paragraph id="1D590B6106D3C5DF1CDA1EAAEDF57BA7" pageId="0" pageNumber="1">L: 5.5 mm (PT)-6.0 mm (HT), TL: 4.8 mm (PT)-5.4 mm (HT). Head relatively large, nearly parallel-sided, with maximum width behind middle, distinctly longer than wide (index HL/HW 1.35 (PT)-1.42 (HT), slightly wider than pronotum, sparsely pubescent. Frontal furrows deep, complete, reaching neck constriction, slightly divergent posteriorly. Anterior pair of supraorbital setae situated before middle of head length, posterior supraorbital setae paired, two setae on each side of head situated close to neck constriction. Neck constriction distinct; genae gently convex. Clypeus and labrum with three pairs of setae, outer pairs longer. Antennae length 4.9 mm (HT)-4.3 mm (PT), scape as long as pedicel, almost as long as terminal antennomere.</paragraph>
<paragraph id="99B868999764C4BF287AF4C40E7D19F8" pageId="0" pageNumber="1">Pronotum elongated, slightly longer and narrower than head, with maximum width in anterior third, index PL/PW 1.75 (PT)-1.83 (HT), only slightly narrowed anteriorly, posterior part distinctly narrower than anterior. Surface glabrous, median furrow distinct, visible in whole pronotal length. Propleura visible from dorsal aspect in basal two-thirds. Anterior angles of pronotum not protruding, posterior angles obtuse. Lateral furrows well developed, deep, with one pair of anterolateral setae, situated in the apical fifth of pronotal length.</paragraph>
<paragraph id="9DFE3584F6E08CD2700C954C68291A2A" pageId="0" pageNumber="1">
Elytra subovate elongate, distinctly longer than wide, index EL/EW 1.74 (PT)-1.85 (HT), with maximum width in posterior third; elytral surface glabrous, without pubescence; striae absent. Stria 3 with 4-5 (3-4 discal and one preapical) macrochetae and 3 or 4 microchetae situated between macrochetae, stria 5 with 4 or 5 microchetae (Fig. 
<figureCitation id="2E7D00EC1FFA7977AF8DCDE0C394F30C" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Habitus of Derosiella species. Derossiella lukici sp. n. (a) and Derossiella nonveilleri (b). Chetotaxy is presented as white points (photo courtesy of Dusan Beno)." figureDoi="10.3897/dez.66.31754.figure4" httpUri="https://binary.pensoft.net/fig/258011" pageId="0" pageNumber="1">4</figureCitation>
). Humeral group of umbilicate pores not aggregated, first anterior pore of humeral group isolated and situated at level of the first discal seta.
</paragraph>
<paragraph id="2FAC791DE594E22D7388AC9245BF5788" pageId="0" pageNumber="1">Legs long, slender, densely pubescent. First two tarsomeres of male protarsi distinctly dilated and protracted at their internal margins. Tarsal claws very long and slender, without traces of denticulation on their internal sides.</paragraph>
<paragraph id="762392A506B35704DCB142CBC1A1BDD6" pageId="0" pageNumber="1">
Male genitalia. Aedeagus (Fig. 
<figureCitation id="1B9E782CB9B465373ABF761F611A4F78" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male and female genitalia of Derossiella representatives: aedeagus of D. lukici sp. n., dorsal view (a); and lateral view (b); female genitalia gonocoxite 1 and 2 (basal and apical segments of gonostylus) copulatory piece of D. lukici sp. n. (c); and D. nonveilleri, lateral view of male aedeagus with parameres detached (d) (illustration by Fedor Ciampor)." figureDoi="10.3897/dez.66.31754.figure5" httpUri="https://binary.pensoft.net/fig/258012" pageId="0" pageNumber="1">5</figureCitation>
) 0.58 mm long, relatively robust, regularly wide, lacking apical constriction, laterally flattened. Parameres relatively long and slender, longer than half of the length of aedeagus. Apex very short, tip obtuse. Each paramere at apex with three setae, two long, and one short.
</paragraph>
<paragraph id="8F701DF1B79CEA72AA1098433833D82E" pageId="0" pageNumber="1">
Female genitalia as in Figure 
<figureCitation id="75225AB5666576B7FFDEBA920FBD1767" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male and female genitalia of Derossiella representatives: aedeagus of D. lukici sp. n., dorsal view (a); and lateral view (b); female genitalia gonocoxite 1 and 2 (basal and apical segments of gonostylus) copulatory piece of D. lukici sp. n. (c); and D. nonveilleri, lateral view of male aedeagus with parameres detached (d) (illustration by Fedor Ciampor)." figureDoi="10.3897/dez.66.31754.figure5" httpUri="https://binary.pensoft.net/fig/258012" pageId="0" pageNumber="1">5</figureCitation>
.
</paragraph>
<caption id="E7F07BF7372C6356507B61CAF43F767A" ID-Zenodo-Dep="2551941" doi="10.3897/dez.66.31754.figure5" httpUri="https://binary.pensoft.net/fig/258012" pageId="0" pageNumber="1" start="Figure 5" startId="F5">
<paragraph id="482A1724DFAE9DE6392314C476BB0CE1" pageId="0" pageNumber="1">
<emphasis id="A391066B92BF1B09D793C8EF0589BA5F" bold="true" pageId="0" pageNumber="1">Figure 5.</emphasis>
Male and female genitalia of 
<taxonomicName id="3FE6B42C12176B514F6B7BE445B5407B" authorityName="Queinnec" authorityYear="2008" class="Insecta" family="Carabidae" genus="Derossiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Derossiella" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis id="190E11CF5C11355DF0FCC4AD6E9FE626" italics="true" pageId="0" pageNumber="1">Derossiella</emphasis>
</taxonomicName>
representatives: aedeagus of 
<taxonomicName id="F7C263D5AABB0C87A1300AB996BE0052" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="D. lukici" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="lukici">
<emphasis id="63F3771FA4ADF319DB83A08E18EBA241" italics="true" pageId="0" pageNumber="1">D. lukici</emphasis>
</taxonomicName>
sp. n., dorsal view 
<emphasis id="1A4807A790A507699070CC27F00814B2" bold="true" pageId="0" pageNumber="1">(a)</emphasis>
; and lateral view 
<emphasis id="037EE1062327CE45180E1E0BDB7772D8" bold="true" pageId="0" pageNumber="1">(b)</emphasis>
; female genitalia gonocoxite 1 and 2 (basal and apical segments of gonostylus) copulatory piece of 
<taxonomicName id="E9DEB0C150B2E0AC2E29AF7706E6C0CA" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="D. lukici" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="lukici">
<emphasis id="A2C64502B958F292AAB7D4F762C32934" italics="true" pageId="0" pageNumber="1">D. lukici</emphasis>
</taxonomicName>
sp. n. 
<emphasis id="249731BC325BA76935C2B79BF26B185A" bold="true" pageId="0" pageNumber="1">(c)</emphasis>
; and 
<taxonomicName id="EDCE368B007FEE755D2D5AA103A135C1" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="D. nonveilleri" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="nonveilleri">
<emphasis id="29452B33E7550D0F641148EF6972D193" italics="true" pageId="0" pageNumber="1">D. nonveilleri</emphasis>
</taxonomicName>
, lateral view of male aedeagus with parameres detached 
<emphasis id="4D96344003E440B5F948B75BEE2D733A" bold="true" pageId="0" pageNumber="1">(d)</emphasis>
(illustration by Fedor 
<normalizedToken id="535E707B07BF65997100EE9B62164B38" originalValue="Čiampor">Ciampor</normalizedToken>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="85B4766B5DB765E6543BC90E0F5A368F" pageId="0" pageNumber="1" type="etymology">
<paragraph id="D881C01D84241F569E9966273D6244D6" pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph id="E851FA91D22CAE09DFB834EDB015AAD1" pageId="0" pageNumber="1">
Patronymic, dedicated to our dear friend Marko 
<normalizedToken id="06D2A855A44BF342AFE606EE0C1C0DDE" originalValue="Lukić">Lukic</normalizedToken>
(Zagreb, Croatia), enthusiastic speleologist and speleobiologist, taxonomic specialist on subterranean 
<taxonomicName id="544004241ADE5C8BBE82968E06604B2C" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Collembola" pageId="0" pageNumber="1" phylum="Arthropoda" rank="order">Collembola</taxonomicName>
, and collector of the first specimen of the new species.
</paragraph>
</subSubSection>
<subSubSection id="CAEA53263973F968855C6FD4EF260D2B" pageId="0" pageNumber="1" type="differential diagnosis">
<paragraph id="A7D8EC979BC3E750830C919F8AD7ADE2" pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
<paragraph id="DF46FE78576840F86966903603F7B361" pageId="0" pageNumber="1">
<taxonomicName id="41E2E939F984C5912DEA2FFC5A2C55B1" authorityName="Lohaj &amp; Delić" authorityYear="2019" class="Insecta" family="Carabidae" genus="Derossiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Derossiella lukici" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="lukici">
<emphasis id="5AF028208A746DEA1546C49C7E965608" italics="true" pageId="0" pageNumber="1">Derossiella lukici</emphasis>
</taxonomicName>
sp. n. is closely related to the type species of the genus, 
<taxonomicName id="332D1F511533F8E112DA17D31D06CAE4" authorityName="Queinnec" authorityYear="2008" class="Insecta" family="Carabidae" genus="Derossiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Derossiella nonveilleri" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="nonveilleri">
<emphasis id="566E07D3573EFF97DF85399664F23129" italics="true" pageId="0" pageNumber="1">Derossiella nonveilleri</emphasis>
</taxonomicName>
. However, these two species can be easily recognized using the following key:
</paragraph>
<paragraph id="A946546B5A2BADA547AF0C6AB167857D" pageId="0" pageNumber="1">
<table id="091E9A83C4C2C9C0467EF5B10EFE2CBC" inLine="true" pageId="0" pageNumber="1">
<tr id="F59017022C79CF9F3CC67C02192C0EDB" pageId="0" pageNumber="1">
<td id="B31A6CA582671EB67B06DBDBAD1D66E8" colspan="1" pageId="0" pageNumber="1" rowspan="1">1(2)</td>
<td id="0F89D7044B247F0F7CFD6ABB15713561" colspan="1" pageId="0" pageNumber="1" rowspan="1">
Head with 2 posterior supraorbital setiferous punctures on each side, which are very close to each other. Elytra with maximum width in the posterior third. Putative stria 3 with 4 or 5 macrochetae and 4-6 microchetae, putative stria 5 with 3 or 4 microchetae (Fig. 
<figureCitation id="B6E91F1D67F3A43C6E76B7B760BA5957" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Habitus of Derosiella species. Derossiella lukici sp. n. (a) and Derossiella nonveilleri (b). Chetotaxy is presented as white points (photo courtesy of Dusan Beno)." figureDoi="10.3897/dez.66.31754.figure4" httpUri="https://binary.pensoft.net/fig/258011" pageId="0" pageNumber="1">4</figureCitation>
). Pronotum narrower, index PL/PW: 1.75-1.83, anterior angles not protruding. Aedeagus (Fig. 
<figureCitation id="442960695F2FBC0ACFC1332049E97537" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male and female genitalia of Derossiella representatives: aedeagus of D. lukici sp. n., dorsal view (a); and lateral view (b); female genitalia gonocoxite 1 and 2 (basal and apical segments of gonostylus) copulatory piece of D. lukici sp. n. (c); and D. nonveilleri, lateral view of male aedeagus with parameres detached (d) (illustration by Fedor Ciampor)." figureDoi="10.3897/dez.66.31754.figure5" httpUri="https://binary.pensoft.net/fig/258012" pageId="0" pageNumber="1">5</figureCitation>
) shorter, more robust. L: 5.5-6.0 mm. Croatia, Mt Biokovo
</td>
<td id="5765A931091F6BFDC35269C11AC99BD6" colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName id="C97CD8A22C0AF5109B878F5A4996BD78" authorityName="Lohaj &amp; Delić" authorityYear="2019" class="Insecta" family="Carabidae" genus="Derossiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Derossiella lukici" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="lukici">
<emphasis id="41B64435E238543C875F7D8D523964E7" italics="true" pageId="0" pageNumber="1">Derossiella lukici</emphasis>
</taxonomicName>
sp. n.
</td>
</tr>
<tr id="F9532DAEF00F725EBFA2C9E52B093209" pageId="0" pageNumber="1">
<td id="6EB2E05C0A3E145EC0B779D3D1A426ED" colspan="1" pageId="0" pageNumber="1" rowspan="1">2(1)</td>
<td id="47E0B403DE220E38F35284CB095FAD98" colspan="1" pageId="0" pageNumber="1" rowspan="1">
Head with only 2 posterior supraorbital setiferous puncture on each side. Elytra with maximum width in middle. Putative stria 3 with 3 macrochetae and 2 microchetae between each macrochetae, putative stria 5 with 1 microcheta in basal fourth (see Fig. 
<figureCitation id="91E6DB71BBF7C0E1B07ABF348E42C058" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Habitus of Derosiella species. Derossiella lukici sp. n. (a) and Derossiella nonveilleri (b). Chetotaxy is presented as white points (photo courtesy of Dusan Beno)." figureDoi="10.3897/dez.66.31754.figure4" httpUri="https://binary.pensoft.net/fig/258011" pageId="0" pageNumber="1">4</figureCitation>
). Pronotum wider, index PL/PW: 1.43-1.44, anterior angles slightly protruding. Aedeagus (Fig. 
<figureCitation id="FE63F145052E7E4FE5CECD4BA9D21FE4" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male and female genitalia of Derossiella representatives: aedeagus of D. lukici sp. n., dorsal view (a); and lateral view (b); female genitalia gonocoxite 1 and 2 (basal and apical segments of gonostylus) copulatory piece of D. lukici sp. n. (c); and D. nonveilleri, lateral view of male aedeagus with parameres detached (d) (illustration by Fedor Ciampor)." figureDoi="10.3897/dez.66.31754.figure5" httpUri="https://binary.pensoft.net/fig/258012" pageId="0" pageNumber="1">5</figureCitation>
) longer and slenderer. L: 5.4-5.8 mm. Croatia, Mt Mosor
</td>
<td id="A3B1D61B0E27410C4C096F718991F496" colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName id="67B44D5468A8D7602EECDA94845269A9" authorityName="Queinnec" authorityYear="2008" class="Insecta" family="Carabidae" genus="Derossiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Derossiella nonveilleri" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="nonveilleri">
<emphasis id="774390A8BB36AC770578CAD478530A83" italics="true" pageId="0" pageNumber="1">Derossiella nonveilleri</emphasis>
</taxonomicName>
<normalizedToken id="4C3A71F8FD4DB29340F5533A469ABDF9" originalValue="Quéinnec">Queinnec</normalizedToken>
, 2008
</td>
</tr>
<tr id="571BC89AA346B5E1DB0A6129AE78CF99" pageId="0" pageNumber="1">
<td id="318FB07EF000E08A122365E2742229E7" colspan="1" isEmpty="true" pageId="0" pageNumber="1" rowspan="1">-</td>
</tr>
</table>
</paragraph>
</subSubSection>
<subSubSection id="3F5238A8D01BC5773F87EF5AA78D6D6D" pageId="0" pageNumber="1" type="distribution">
<paragraph id="A43A479299A1050FD1C37E3598BF9FCB" pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph id="C0991C0DB3647B3C3AE12684711423C3" pageId="0" pageNumber="1">
So far this species is known from the two pits on Mt Biokovo, the type locality, Biokovka, Golubinjak and Pretnerova jama, Lokva. All three specimens were found in deeper parts of the caves, attaining depths of 120 to 300 m. They were all found walking on the 
<normalizedToken id="EF1858B39E91FD3EDC00F03E66F6E501" originalValue="“moonmilk”">&quot;moonmilk&quot;</normalizedToken>
, a white, pastelite material consisting of microbiologically transformed microcrystalline calcites with high water content (60-90%), and near the cave hygropetric.
</paragraph>
<paragraph id="DABEA62A0A6CBC2BBFD91095810CE440" pageId="0" pageNumber="1">Associated subterranean coleopteran fauna observed in the pits:</paragraph>
<paragraph id="106FEFBD6981DA1BE4E7BA06B1055ABE" pageId="0" pageNumber="1">1 Biokovka, Golubinjak, Biokovo, Croatia:</paragraph>
<paragraph id="F83D4276C9103A322BD23BA30D403ED1" pageId="0" pageNumber="1">
<taxonomicName id="A6F6B0F673DF27C8A97904204FE0E9B8" authorityName="Latreille" authorityYear="1802" class="Insecta" family="Carabidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="family">Carabidae</taxonomicName>
: 
<taxonomicName id="20626E944CE4BEA02CFBF81BA544F280" authorityName="Bonelli" authorityYear="1810" lsidName="" pageId="0" pageNumber="1" rank="subFamily" subFamily="Trechinae">Trechinae</taxonomicName>
</paragraph>
<paragraph id="8BE113970F005DE1728B2CE145EF4C31" pageId="0" pageNumber="1">
<taxonomicName id="91111FDC3FE235353CFC051094298A97" baseAuthorityName="L. Miller" baseAuthorityYear="1861" class="Insecta" family="Carabidae" genus="Neotrechus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Neotrechus dalmatinus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="dalmatinus">
<emphasis id="A819EC2C0E980F35B7B8FF40C76D195A" italics="true" pageId="0" pageNumber="1">Neotrechus dalmatinus</emphasis>
</taxonomicName>
(Miller, 1861)
</paragraph>
<paragraph id="85CEA186ED9961FC81DB4BA019C93147" pageId="0" pageNumber="1">
<taxonomicName id="476715BDA51C096913974B23C11FD0A1" authorityName="Fleming" authorityYear="1821" class="Insecta" family="Leiodidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="family">Leiodidae</taxonomicName>
: 
<taxonomicName id="422550CD2FD2C2333779A1A0710E4614" authorityName="Kirby" authorityYear="1837" lsidName="" pageId="0" pageNumber="1" rank="subFamily" subFamily="Cholevinae">Cholevinae</taxonomicName>
</paragraph>
<paragraph id="A2435AE6F971B1457CFE5B03727A22B4" pageId="0" pageNumber="1">
<taxonomicName id="C539FE419ED4B003B2FF1228BAE18F17" class="Insecta" family="Leiodidae" genus="Leptomeson" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Leptomeson biokovensis" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="biokovensis">
<emphasis id="C4EB604E1E3C8D5B2747BAF4E73D11CB" italics="true" pageId="0" pageNumber="1">Leptomeson biokovensis</emphasis>
</taxonomicName>
Giachino, 
<normalizedToken id="E9EE0AF8C58BCC57DA0EB11A3785A5BB" originalValue="Bregović">Bregovic</normalizedToken>
&amp; 
<normalizedToken id="A9A5A136AFC076B7B8FFA26A713AB27F" originalValue="Jalžić">Jalzic</normalizedToken>
, 2012
</paragraph>
<paragraph id="3C0487910B3B8B8729D2DAEED83A8D57" pageId="0" pageNumber="1">
<taxonomicName id="05FCB304FFC65C50AD0C9149C84F92A5" class="Insecta" family="Leiodidae" genus="Radziella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Radziella styx" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="styx">
<emphasis id="FA36FB4467A2B2C172752039B63CF347" italics="true" pageId="0" pageNumber="1">Radziella styx</emphasis>
</taxonomicName>
Casale &amp; 
<normalizedToken id="D935D01B75BA83C0ADB2A19EB5F0BC82" originalValue="Jalžić">Jalzic</normalizedToken>
, 1988
</paragraph>
<paragraph id="00D122AD4ECF0C44683CD976CF1F4BDF" pageId="0" pageNumber="1">
<taxonomicName id="05EB12943E0DBB4F904749DE4B08B832" class="Insecta" family="Leiodidae" genus="Speoplanes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Speoplanes giganteus subsp. biocovensis" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="subSpecies" species="giganteus" subSpecies="biocovensis">
<emphasis id="7B470D3F42D3B34F859CBE506F8CC4D6" italics="true" pageId="0" pageNumber="1">Speoplanes giganteus biocovensis</emphasis>
</taxonomicName>
<normalizedToken id="5405E8AEE34CE122183249B88642BA5F" originalValue="Müller">Mueller</normalizedToken>
, 1934
</paragraph>
<paragraph id="69D6568854A93E1C63C0BCE0DA0AE9F1" pageId="0" pageNumber="1">2 Pretnerova jama, Lokva, Biokovo, Croatia:</paragraph>
<paragraph id="B77994353BA0FA5D91EDEEB0DE37BD3A" pageId="0" pageNumber="1">
<taxonomicName id="53E91C5245FB451D3CFC4A5A8D78D256" authorityName="Latreille" authorityYear="1802" class="Insecta" family="Carabidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="family">Carabidae</taxonomicName>
: 
<taxonomicName id="FB18D1666A95A45E164B37FA29461483" authorityName="Bonelli" authorityYear="1810" lsidName="" pageId="0" pageNumber="1" rank="subFamily" subFamily="Trechinae">Trechinae</taxonomicName>
</paragraph>
<paragraph id="6551841604CF460FD0339F21979D1879" pageId="0" pageNumber="1">
<taxonomicName id="2E2ADED3D89F321702B77B760B9C1806" authorityName="Monguzzi" authorityYear="1993" class="Insecta" family="Carabidae" genus="Dalmataphaenops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Dalmataphaenops chiarae" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="chiarae">
<emphasis id="B3D6972960240F5B0F8F26CA13824209" italics="true" pageId="0" pageNumber="1">Dalmataphaenops chiarae</emphasis>
</taxonomicName>
Monguzzi, 1993
</paragraph>
<paragraph id="8BE4DA10CAFA2C17CB8F19FB35DAC146" pageId="0" pageNumber="1">
<taxonomicName id="8E1876407AA7897659D43FB2BE88FE25" authorityName="Latreille" authorityYear="1802" class="Insecta" family="Carabidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="family">Carabidae</taxonomicName>
: 
<taxonomicName id="DB11432807F12AF28DB38C70D9D0A35B" authorityName="Laporte de Castelnau" authorityYear="1834" lsidName="" pageId="0" pageNumber="1" rank="tribe" tribe="Sphodrini">Sphodrini</taxonomicName>
</paragraph>
<paragraph id="7B73E3F274DBF3F39E8A34D0817F7ACE" pageId="0" pageNumber="1">
<taxonomicName id="36880A87962D1F5B070E42624AE38F68" baseAuthorityName="Schaum" baseAuthorityYear="1858" class="Insecta" family="Carabidae" genus="Laemostenus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Laemostenus cavicola" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="cavicola">
<emphasis id="706A1CEDD07EA952A09D0574E06B3F51" italics="true" pageId="0" pageNumber="1">Laemostenus cavicola</emphasis>
</taxonomicName>
(Schaum, 1858)
</paragraph>
<paragraph id="1DB7828C7F90F47FB5B8C943F397B861" pageId="0" pageNumber="1">
<taxonomicName id="1488F36A2E1BE6D67B42C813DA95B5D1" authorityName="Fleming" authorityYear="1821" class="Insecta" family="Leiodidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="family">Leiodidae</taxonomicName>
: 
<taxonomicName id="4CFC0CAD0AA6B634163291F5BA992240" authorityName="Kirby" authorityYear="1837" lsidName="" pageId="0" pageNumber="1" rank="subFamily" subFamily="Cholevinae">Cholevinae</taxonomicName>
</paragraph>
<paragraph id="BE61880BA9C56DB751C9CCDF16AD8535" pageId="0" pageNumber="1">
<taxonomicName id="2269BFCA45829D4173FFC3FDEA0214D2" class="Insecta" family="Leiodidae" genus="Leptomeson" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Leptomeson biokovensis" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="biokovensis">
<emphasis id="4C516BD803C0207884E01B6011C8B51F" italics="true" pageId="0" pageNumber="1">Leptomeson biokovensis</emphasis>
</taxonomicName>
Giachino, 
<normalizedToken id="79B4901E541F4367CDB3B1CB68083158" originalValue="Bregović">Bregovic</normalizedToken>
&amp; 
<normalizedToken id="023CC84BB9D7DA711483DCFA34DBE20B" originalValue="Jalžić">Jalzic</normalizedToken>
, 2012
</paragraph>
<paragraph id="A1FDDD5B4881773C8DD39362A184375F" pageId="0" pageNumber="1">
<taxonomicName id="160645B0117B65E626350D8FD3BA5F15" class="Insecta" family="Leiodidae" genus="Radziella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Radziella styx" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="styx">
<emphasis id="0CA6466BA098DB32A43E053EB9C4F1F9" italics="true" pageId="0" pageNumber="1">Radziella styx</emphasis>
</taxonomicName>
Casale &amp; 
<normalizedToken id="9D9C394D9743CC47DE3446D25247D690" originalValue="Jalžić">Jalzic</normalizedToken>
, 1988
</paragraph>
<paragraph id="8B0C7399338B2573FC514E94FCFB1AEA" pageId="0" pageNumber="1">
<taxonomicName id="BB70BF913B000036CFA585BC19CF8EAC" class="Insecta" family="Leiodidae" genus="Speoplanes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Speoplanes giganteus subsp. biocovensis" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="subSpecies" species="giganteus" subSpecies="biocovensis">
<emphasis id="25F0F7F7A5B4A801DFFC8B871F91EC2B" italics="true" pageId="0" pageNumber="1">Speoplanes giganteus biocovensis</emphasis>
</taxonomicName>
<normalizedToken id="1E88AB8DEA6719B94499AA2E6788D541" originalValue="Müller">Mueller</normalizedToken>
, 1934
</paragraph>
<paragraph id="3DA0656FFE3A7A2BFCF53A0DFB65F266" pageId="0" pageNumber="1">
<taxonomicName id="33FFCBE61538492DADBF87EE150FC8BC" class="Insecta" family="Staphylinidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="family">Staphylinidae</taxonomicName>
: 
<taxonomicName id="4D6806A589EE2C0EF1819EA65D90FD64" authorityName="Latreille" authorityYear="1802" lsidName="" pageId="0" pageNumber="1" rank="subFamily" subFamily="Pselaphinae">Pselaphinae</taxonomicName>
</paragraph>
<paragraph id="0843BE02906342F3150304B85D548BF9" pageId="0" pageNumber="1">
<taxonomicName id="3C442B2A735AF956A54205C712E8A674" authorityName="Latreille" authorityYear="1802" lsidName="" pageId="0" pageNumber="1" rank="subFamily" subFamily="Pselaphinae">Pselaphinae</taxonomicName>
gen.
</paragraph>
</subSubSection>
</treatment>
</document>