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<document id="7EFE077F024170D870E4020A76906BD5" ID-CLB-Dataset="45522" ID-DOI="10.11646/zootaxa.3814.4.3" ID-GBIF-Dataset="b1c0a4d9-3b72-496e-846b-c05dd2f6d96a" ID-ISSN="1175-5326" ID-Zenodo-Dep="229706" ID-ZooBank="670FF8E1-49C6-4D36-8981-1242C18360E3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460435152864" checkinUser="plazi" docAuthor="Silva, Guilherme Liberato Da, Cunha, Uemerson Silva Da, Rocha, Matheus Dos Santos, Panou, Eleni N. &amp; Ferla, Noeli Juarez" docDate="2014" docId="03FAB66161123A56D3F5FA16FA3FF878" docLanguage="en" docName="zt03814p511.pdf" docOrigin="Zootaxa 3814 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tydeus Koch 1835" docType="treatment" docVersion="12" lastPageNumber="501" masterDocId="FFC3CE1961143A50D362FFA4FFB7FFFB" masterDocTitle="Tydeid and triophtydeid mites (Acari: Tydeoidea) associated with grapevine (Vitaceae: Vitis spp.) in Brazil, with the descriptions of species of Prelorryia (André, 1980) and Tydeus Koch, 1835" masterLastPageNumber="511" masterPageNumber="495" pageNumber="501" updateTime="1698332017546" updateUser="ExternalLinkService">
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<mods:title id="55BA336810C4E69B8FDE32D2FE202550">Tydeid and triophtydeid mites (Acari: Tydeoidea) associated with grapevine (Vitaceae: Vitis spp.) in Brazil, with the descriptions of species of Prelorryia (André, 1980) and Tydeus Koch, 1835</mods:title>
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<mods:namePart id="60DE078840F77DB7B5EF92B69DC05D40">Silva, Guilherme Liberato Da</mods:namePart>
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<mods:namePart id="26EAB2FFD601AB905F16C0EEEAA63592">Cunha, Uemerson Silva Da</mods:namePart>
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<mods:namePart id="A9DDE4E51991D834737B15D68C12CA7C">Rocha, Matheus Dos Santos</mods:namePart>
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<mods:namePart id="A3DA3402B54FDD715A7CAD04057230FB">Panou, Eleni N.</mods:namePart>
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<mods:namePart id="50007FA29842F0FE372AC13E486AA11B">Ferla, Noeli Juarez</mods:namePart>
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<mods:date id="B32B09C2A5598F2EDAB1AF6B400BFDF0">2014</mods:date>
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<heading id="D0A4B01B61123A56D3F5FA16FE37FA37" bold="true" box="[151,384,1458,1484]" fontSize="11" level="1" pageId="6" pageNumber="501" reason="1">
<taxonomicName id="4C537CF461123A56D3F5FA16FE37FA37" authority="Koch, 1835" authorityName="Koch" authorityYear="1835" box="[151,384,1458,1484]" class="Arachnida" family="Tydeidae" genus="Tydeus" kingdom="Animalia" order="Prostigmata" pageId="6" pageNumber="501" phylum="Arthropoda" rank="genus">
<emphasis id="B927DB6561123A56D3F5FA16FE37FA37" bold="true" box="[151,384,1458,1484]" pageId="6" pageNumber="501">
<emphasis id="B927DB6561123A56D3F5FA16FF5BFA37" bold="true" box="[151,236,1458,1484]" italics="true" pageId="6" pageNumber="501">Tydeus</emphasis>
<bibRefCitation id="EFC27A8661123A56D391FA16FE37FA37" author="Koch" box="[243,384,1458,1484]" pageId="6" pageNumber="501" refString="Koch, C. L. (1835) Deutschlands Crustaceen, Myriapoden und Arachniden 4. Herrich-Schaffer, Regensburg, pp. 11 - 12." type="journal article" year="1835">Koch, 1835</bibRefCitation>
</emphasis>
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<taxonomicName id="4C537CF461123A56D3F5FA5DFECEF9F4" authority="Koch, 1835:11" authorityName="Koch" authorityPageNumber="11" authorityYear="1835" box="[151,377,1529,1551]" class="Arachnida" family="Tydeidae" genus="Tydeus" kingdom="Animalia" order="Prostigmata" pageId="6" pageNumber="501" phylum="Arthropoda" rank="genus">
<emphasis id="B927DB6561123A56D3F5FA5DFF6AF9F4" box="[151,221,1529,1551]" italics="true" pageId="6" pageNumber="501">Tydeus</emphasis>
<treatmentCitation id="0AF2216661123A56D381FA5DFECEF9F4" author="Koch" box="[227,377,1529,1551]" page="11" pageId="6" pageNumber="501" year="1835">
<bibRefCitation id="EFC27A8661123A56D381FA5DFEEAF9F4" author="Koch" box="[227,349,1529,1551]" pageId="6" pageNumber="501" refString="Koch, C. L. (1835) Deutschlands Crustaceen, Myriapoden und Arachniden 4. Herrich-Schaffer, Regensburg, pp. 11 - 12." type="journal article" year="1835">Koch, 1835</bibRefCitation>
:11
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12;
<treatmentCitation id="0AF2216661123A56D2CEFA5DFD3CF9F4" author="Oudemans" box="[428,651,1529,1551]" page="922" pageId="6" pageNumber="501" year="1937">
<bibRefCitation id="EFC27A8661123A56D2CEFA5DFDECF9F4" author="Oudemans" box="[428,603,1529,1551]" pageId="6" pageNumber="501" refString="Oudemans, A. C. (1937) Kritisch Historisch Overzicht der Acarologie. Derde Gedeelte Band C. E. J. Brill, Leiden, pp. 799 - 1348." type="book chapter" year="1937">Oudemans, 1937</bibRefCitation>
: 922
</treatmentCitation>
(in part);
<treatmentCitation id="0AF2216661123A56D193FA5DFC53F9F4" author="Kazmierski" box="[753,996,1529,1551]" page="292" pageId="6" pageNumber="501" year="1989">
<bibRefCitation id="EFC27A8661123A56D193FA5DFC02F9F4" author="Kazmierski" box="[753,949,1529,1551]" pageId="6" pageNumber="501" refString="Kazmierski, A. (1989 b) Revision of the genera Tydeus Koch sensu Andre, Homeotydeus Andre and Orthotydeus Andre with description of a new genus and four new species of Tydeinae (Acari: Actinedida: Tydeidae). Mitteilungen Hamburgisches Zoologisches Museum und Institut, 86, 289 - 314." type="journal article" year="1989" yearSuffix="b">Kaźmierski, 1989b</bibRefCitation>
: 292
</treatmentCitation>
(in part).
<taxonomicName id="4C537CF461123A56D3F5F9B3FE0AF9D6" authority="Thor, 1933: 54" authorityName="Thor" authorityPageNumber="54" authorityYear="1933" box="[151,445,1559,1581]" class="Arachnida" family="Tydeidae" genus="Brachytydeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="6" pageNumber="501" phylum="Arthropoda" rank="genus">
<emphasis id="B927DB6561123A56D3F5F9B3FE96F9D6" box="[151,289,1559,1581]" italics="true" pageId="6" pageNumber="501">Brachytydeus</emphasis>
<treatmentCitation id="0AF2216661123A56D245F9B3FE0AF9D6" author="Thor" box="[295,445,1559,1581]" page="54" pageId="6" pageNumber="501" year="1933">
<bibRefCitation id="EFC27A8661123A56D245F9B3FE2DF9D6" author="Thor" box="[295,410,1559,1581]" pageId="6" pageNumber="501" refString="Thor, S. (1933) Acarina. Tydeidae, Erynetidae. Das Tierreich, 60, 1 - 82." type="journal article" year="1933">Thor, 1933</bibRefCitation>
: 54
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</taxonomicName>
(in part).
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</paragraph>
<paragraph id="8BEC077761123A56D3F5F992FDF9F9B7" blockId="6.[151,1092,1529,1642]" box="[151,590,1590,1612]" pageId="6" pageNumber="501">
<emphasis id="B927DB6561123A56D3F5F992FEBEF9B7" box="[151,265,1590,1612]" italics="true" pageId="6" pageNumber="501">Calotydeus</emphasis>
<bibRefCitation id="EFC27A8661123A56D26DF992FE09F9B7" author="Oudemans" box="[271,446,1590,1612]" pageId="6" pageNumber="501" refString="Oudemans, A. C. (1937) Kritisch Historisch Overzicht der Acarologie. Derde Gedeelte Band C. E. J. Brill, Leiden, pp. 799 - 1348." type="book chapter" year="1937">Oudemans, 1937</bibRefCitation>
: 923 (in part).
</paragraph>
<paragraph id="8BEC077761123A56D3F5F9F0FE7BF991" blockId="6.[151,1092,1529,1642]" box="[151,460,1620,1642]" pageId="6" pageNumber="501">
<treatmentCitationGroup id="AB43205961123A56D3F5F9F0FE7BF991" box="[151,460,1620,1642]" pageId="6" pageNumber="501">
<taxonomicName id="4C537CF461123A56D3F5F9F0FE7BF991" authority="Andre, 1980: 127" authorityName="Andre" authorityPageNumber="127" authorityYear="1980" box="[151,460,1620,1642]" class="Arachnida" family="Tydeidae" genus="Orthotydeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="6" pageNumber="501" phylum="Arthropoda" rank="genus">
<emphasis id="B927DB6561123A56D3F5F9F0FEA3F991" box="[151,276,1620,1642]" italics="true" pageId="6" pageNumber="501">Orthotydeus</emphasis>
<treatmentCitation id="0AF2216661123A56D278F9F0FE7BF991" author="Andre" box="[282,460,1620,1642]" page="127" pageId="6" pageNumber="501" year="1980">
<bibRefCitation id="EFC27A8661123A56D278F9F0FE2BF991" author="Andre" box="[282,412,1620,1642]" pageId="6" pageNumber="501" refString="Andre, H. M. (1980) A generic revision of the family Tydeidae (Acari: Actinedida). IV. Generic descriptions, keys and conclusion. Bulletin et Annales de la Societe royale Belge d´Entomologie, 116, 103 - 168." type="journal article" year="1980">André, 1980</bibRefCitation>
: 127
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</paragraph>
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<typeStatus id="54E8B9D561123A56D3F5F93CFF7EF956" box="[151,201,1688,1709]" pageId="6" pageNumber="501">Type</typeStatus>
species:
<taxonomicName id="4C537CF461123A56D244F933FCEEF956" authority="Oudemans, 1928 sensu Andre 2005" authorityName="Oudemans, 1928 sensu Andre" authorityYear="2005" box="[294,857,1687,1710]" class="Arachnida" family="Tydeidae" genus="Tydeus" kingdom="Animalia" order="Prostigmata" pageId="6" pageNumber="501" phylum="Arthropoda" rank="species" species="spathulatus">
<emphasis id="B927DB6561123A56D244F933FE50F956" box="[294,487,1687,1709]" italics="true" pageId="6" pageNumber="501">Tydeus spathulatus</emphasis>
<bibRefCitation id="EFC27A8661123A56D28FF933FD2EF956" author="Oudemans" box="[493,665,1687,1709]" pageId="6" pageNumber="501" refString="Oudemans, A. C. (1928) Acarologische Aantekeningen 94. Entomologische Berrichten, 7, 374 - 382." type="journal article" year="1928">Oudemans, 1928</bibRefCitation>
<emphasis id="B927DB6561123A56D1FDF93DFD60F955" box="[671,727,1689,1710]" italics="true" pageId="6" pageNumber="501">sensu</emphasis>
<bibRefCitation id="EFC27A8661123A56D1BFF933FCEEF956" author="Andre" box="[733,857,1687,1709]" pageId="6" pageNumber="501" refString="Andre, H. M. (2005) In search of the true Tydeus (Acari: Tydeidae). Journal of Natural History, 39, 975 - 1001. http: // dx. doi. org / 10.1080 / 00222930400002838" type="journal article" year="2005">André 2005</bibRefCitation>
</taxonomicName>
.
</paragraph>
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<subSubSection id="C34954FC61123A56D3F5F97EFA3FF878" pageId="6" pageNumber="501" type="discussion">
<paragraph id="8BEC077761123A56D3F5F97EFA3FF878" blockId="6.[151,1436,1754,1923]" pageId="6" pageNumber="501">
<emphasis id="B927DB6561123A56D3F5F97EFEBCF90A" bold="true" box="[151,267,1754,1779]" pageId="6" pageNumber="501">Remarks.</emphasis>
A few
<taxonomicName id="4C537CF461123A56D205F97EFE03F908" box="[359,436,1754,1779]" class="Arachnida" family="Tydeidae" genus="Tydeus" kingdom="Animalia" order="Prostigmata" pageId="6" pageNumber="501" phylum="Arthropoda" rank="genus">
<emphasis id="B927DB6561123A56D205F97EFE03F908" box="[359,436,1754,1779]" italics="true" pageId="6" pageNumber="501">Tydeus</emphasis>
</taxonomicName>
-like species have clavate or spatulate setae at their posterior idiosomal margin. These were placed in
<taxonomicName id="4C537CF461123A56D22FF95AFE60F8EC" box="[333,471,1790,1815]" class="Arachnida" family="Tydeidae" genus="Orthotydeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="6" pageNumber="501" phylum="Arthropoda" rank="genus">
<emphasis id="B927DB6561123A56D22FF95AFE60F8EC" box="[333,471,1790,1815]" italics="true" pageId="6" pageNumber="501">Orthotydeus</emphasis>
</taxonomicName>
, as proposed by
<bibRefCitation id="EFC27A8661123A56D1C7F95AFCF4F8EC" author="Andre" box="[677,835,1790,1815]" pageId="6" pageNumber="501" refString="Andre, H. M. (1980) A generic revision of the family Tydeidae (Acari: Actinedida). IV. Generic descriptions, keys and conclusion. Bulletin et Annales de la Societe royale Belge d´Entomologie, 116, 103 - 168." type="journal article" year="1980">André (1980)</bibRefCitation>
. However,
<bibRefCitation id="EFC27A8661123A56D0A8F95AFB19F8EC" author="Kazmierski" box="[970,1198,1790,1815]" pageId="6" pageNumber="501" refString="Kazmierski, A. (1989 b) Revision of the genera Tydeus Koch sensu Andre, Homeotydeus Andre and Orthotydeus Andre with description of a new genus and four new species of Tydeinae (Acari: Actinedida: Tydeidae). Mitteilungen Hamburgisches Zoologisches Museum und Institut, 86, 289 - 314." type="journal article" year="1989" yearSuffix="b">Kaźmierski (1989b)</bibRefCitation>
redefined
<taxonomicName id="4C537CF461123A56D64CF95AFACCF8EC" box="[1326,1403,1790,1815]" class="Arachnida" family="Tydeidae" genus="Tydeus" kingdom="Animalia" order="Prostigmata" pageId="6" pageNumber="501" phylum="Arthropoda" rank="genus">
<emphasis id="B927DB6561123A56D64CF95AFACCF8EC" box="[1326,1403,1790,1815]" italics="true" pageId="6" pageNumber="501">Tydeus</emphasis>
</taxonomicName>
to include species with such modified setae, and subsequently
<taxonomicName id="4C537CF461123A56D05DF886FC7DF8C0" box="[831,970,1826,1851]" class="Arachnida" family="Tydeidae" genus="Orthotydeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="6" pageNumber="501" phylum="Arthropoda" rank="genus">
<emphasis id="B927DB6561123A56D05DF886FC7DF8C0" box="[831,970,1826,1851]" italics="true" pageId="6" pageNumber="501">Orthotydeus</emphasis>
</taxonomicName>
was synonymized with
<taxonomicName id="4C537CF461123A56D7BEF886FA9EF8C0" box="[1244,1321,1826,1851]" class="Arachnida" family="Tydeidae" genus="Tydeus" kingdom="Animalia" order="Prostigmata" pageId="6" pageNumber="501" phylum="Arthropoda" rank="genus">
<emphasis id="B927DB6561123A56D7BEF886FA9EF8C0" box="[1244,1321,1826,1851]" italics="true" pageId="6" pageNumber="501">Tydeus</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFC27A8661123A56D634F886FF53F8A4" author="Andre" pageId="6" pageNumber="501" refString="Andre, H. M. (2005) In search of the true Tydeus (Acari: Tydeidae). Journal of Natural History, 39, 975 - 1001. http: // dx. doi. org / 10.1080 / 00222930400002838" type="journal article" year="2005">André (2005)</bibRefCitation>
. This genus is characterized by having four pairs of aggenital and four or six pairs of genital setae; epimeral formula 3-1-4-2; leg setal counts: Leg I: 8 (1ω)-4-3-3-1; Leg II: 6-2-2-2-0; Leg III: 5-2-1-1-1; Leg IV: 5-2-1-1-0.
</paragraph>
</subSubSection>
</treatment>
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