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<document id="02940D966421ABB3A4B146BCF42F058C" ID-CLB-Dataset="31121" ID-DOI="10.11646/zootaxa.4365.4.4" ID-GBIF-Dataset="8718a968-18bd-49fa-aaa9-d8b56f49baa7" ID-ISSN="1175-5326" ID-Zenodo-Dep="1119233" ID-ZooBank="5414E5C8-8360-454F-84C5-E17E8BAF7CB3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1513752532573" checkinUser="plazi" docAuthor="Tan, Ming Kai &amp; Wahab, Rodzay Bin Haji Abdul" docDate="2017" docId="03AF5B280450C274FF2FFDD91BBFAA4E" docLanguage="en" docName="zootaxa.4365.4.4.pdf" docOrigin="Zootaxa 4365 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Endodrelanva nympha Tan &amp; Wahab 2017, new species" docType="treatment" docVersion="8" lastPageNumber="453" masterDocId="FF9623500459C279FFB8FFE81F37AE5E" masterDocTitle="New taxa and notes on crickets of the subfamily Landrevinae (Orthoptera: Gryllidae) from Brunei Darussalam, Borneo" masterLastPageNumber="454" masterPageNumber="440" pageNumber="449" updateTime="1698511074806" updateUser="ExternalLinkService">
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<mods:title id="3BE3E2592CBACB228B39218E08DCE03D">New taxa and notes on crickets of the subfamily Landrevinae (Orthoptera: Gryllidae) from Brunei Darussalam, Borneo</mods:title>
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<mods:namePart id="12135618C6C67F31F1204BCF251B80E5">Tan, Ming Kai</mods:namePart>
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<mods:namePart id="5499FF0529F32864147A04A55731C2B7">Wahab, Rodzay Bin Haji Abdul</mods:namePart>
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<mods:title id="3A86C8F57CDA9925D6D4B974A3FD4965">Zootaxa</mods:title>
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<mods:date id="B2BE094D896C797453F88C161C28ACDA">2017</mods:date>
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<mods:number id="C5F7FAA7281F934E0CDDF485592C27D7">2017-12-20</mods:number>
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<treatment id="03AF5B280450C274FF2FFDD91BBFAA4E" ID-DOI="http://doi.org/10.5281/zenodo.6020217" ID-GBIF-Taxon="139350619" ID-Zenodo-Dep="6020217" LSID="urn:lsid:plazi:treatment:03AF5B280450C274FF2FFDD91BBFAA4E" httpUri="http://treatment.plazi.org/id/03AF5B280450C274FF2FFDD91BBFAA4E" lastPageId="13" lastPageNumber="453" pageId="9" pageNumber="449">
<subSubSection id="C31CB9B50450C270FF2FFDD91E26AC32" pageId="9" pageNumber="449" type="nomenclature">
<paragraph id="8BB9EA3E0450C270FF2FFDD91D7EAC15" blockId="9.[151,1055,561,656]" box="[151,585,561,587]" pageId="9" pageNumber="449">
<heading id="D0F15D520450C270FF2FFDD91D7EAC15" bold="true" box="[151,585,561,587]" fontSize="11" level="1" pageId="9" pageNumber="449" reason="1">
<emphasis id="B972362C0450C270FF2FFDD91D7EAC15" bold="true" box="[151,585,561,587]" pageId="9" pageNumber="449">
<taxonomicName id="4C0691BD0450C270FF2FFDD91E9CAC15" authority="Tan &amp; Wahab, 2017" authorityName="Tan &amp; Wahab" authorityYear="2017" box="[151,427,561,587]" class="Insecta" family="Haglotettigoniidae" genus="Endodrelanva" kingdom="Animalia" order="Orthoptera" pageId="9" pageNumber="449" phylum="Arthropoda" rank="species" species="nympha" status="sp. nov.">
<emphasis id="B972362C0450C270FF2FFDD91E9CAC15" bold="true" box="[151,427,561,587]" italics="true" pageId="9" pageNumber="449">Endodrelanva nympha</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A2418B570450C270FE01FDD91D7EAC15" box="[441,585,561,587]" pageId="9" pageNumber="449" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BB9EA3E0450C270FF2FFDBC1E26AC32" blockId="9.[151,1055,561,656]" box="[151,273,596,621]" pageId="9" pageNumber="449">
(
<figureCitation id="133DF6BB0450C270FF27FDBC1FD9AC32" box="[159,238,596,621]" captionStart="FIGURE 7" captionStartId="9.[151,250,2012,2034]" captionTargetBox="[200,1392,1393,2003]" captionTargetId="figure@9.[176,1420,1393,2006]" captionTargetPageId="9" captionText="FIGURE 7. Endodrelanva nympha sp. nov. male habitus in dorsal view. Scale bar: 5 mm." httpUri="https://zenodo.org/record/1119249/files/figure.png" pageId="9" pageNumber="449">Figs. 7</figureCitation>
,
<figureCitation id="133DF6BB0450C270FF42FDBC1E3FAC32" box="[250,264,596,620]" captionStart="FIGURE 8" captionStartId="10.[151,250,1936,1958]" captionTargetBox="[180,1415,193,1895]" captionTargetId="figure@10.[171,1415,193,1910]" captionTargetPageId="10" captionText="FIGURE 8. Endodrelanva nympha sp. nov. male: body in profile view (A), tegmen in dorsal view (B), metanotal gland in dorsal view (C), abdominal apex in dorsal (D) and ventral (E) views, genitalia in dorsal (F), ventral (G) and profile (H) views, and posteromedial epiphallic lobule (I). Scale bars: 2 mm (A), 1 mm (BF)." httpUri="https://zenodo.org/record/1119251/files/figure.png" pageId="9" pageNumber="449">8</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31CB9B50450C270FF2FFD9F1B28ACCE" box="[151,1055,631,656]" pageId="9" pageNumber="449" type="description">
<paragraph id="8BB9EA3E0450C270FF2FFD9F1B28ACCE" blockId="9.[151,1055,561,656]" box="[151,1055,631,656]" pageId="9" pageNumber="449">
http://lsid.speciesfile.org/urn:lsid:
<taxonomicName id="4C0691BD0450C270FDB1FD9F1DBEACCE" box="[521,649,631,656]" class="Insecta" kingdom="Animalia" order="Orthoptera" pageId="9" pageNumber="449" phylum="Arthropoda" rank="order">Orthoptera</taxonomicName>
.speciesfile.org:TaxonName:500655
</paragraph>
</subSubSection>
<subSubSection id="C31CB9B50450C270FF2FFD571D6CAB62" pageId="9" pageNumber="449" type="materials_examined">
<paragraph id="8BB9EA3E0450C270FF2FFD571BA9AD7E" blockId="9.[151,1437,703,1341]" pageId="9" pageNumber="449">
<materialsCitation id="3B6EE0630450C270FF2FFD571BA9AD7E" ID-GBIF-Occurrence="1795862918" collectingDate="2017-07-29" collectionCode="IBER" collectorName="M. K. Tan" country="Brunei" county="Material" elevation="7" latitude="4.54728" location="Sungei Mata Ikan" longLatPrecision="1" longitude="115.15711" municipality="Kuala Belalong Field Studies Centre" pageId="9" pageNumber="449" specimenCount="1" stateProvince="Temburong" typeStatus="holotype">
<emphasis id="B972362C0450C270FF2FFD571D36AC86" bold="true" box="[151,513,703,728]" pageId="9" pageNumber="449">
<collectingCounty id="62D892B20450C270FF2FFD571FC8AC86" box="[151,255,703,728]" pageId="9" pageNumber="449">Material</collectingCounty>
examined.
<typeStatus id="54BD549C0450C270FE2CFD571D36AC86" box="[404,513,703,728]" pageId="9" pageNumber="449" type="holotype">Holotype</typeStatus>
</emphasis>
(male):
<emphasis id="B972362C0450C270FDD6FD571C6AAC86" bold="true" box="[622,861,703,728]" pageId="9" pageNumber="449">
<collectingCountry id="F311AAAE0450C270FDD6FD571C6AAC86" box="[622,861,703,728]" name="Brunei" pageId="9" pageNumber="449">Brunei Darussalam</collectingCountry>
</emphasis>
, Ulu
<collectingRegion id="49C224DC0450C270FC12FD571B19AC86" box="[938,1070,703,728]" country="Brunei" name="Temburong" pageId="9" pageNumber="449">Temburong</collectingRegion>
,
<collectingMunicipality id="6BDD70440450C270FB87FD571FD3ACA2" pageId="9" pageNumber="449">Kuala Belalong Field Studies Centre</collectingMunicipality>
, near
<location id="8ED9BCE50450C270FE8DFD0C1D3AACA3" LSID="urn:lsid:plazi:treatment:03AF5B280450C274FF2FFDD91BBFAA4E:8ED9BCE50450C270FE8DFD0C1D3AACA3" box="[309,525,740,765]" country="Brunei" county="Material" latitude="4.54728" longLatPrecision="1" longitude="115.15711" municipality="Kuala Belalong Field Studies Centre" name="Sungei Mata Ikan" pageId="9" pageNumber="449" stateProvince="Temburong">Sungei Mata Ikan</location>
, primary ridge dipterocarp forest, on tree branch near a female,
<geoCoordinate id="EE328CF90450C270FA9EFD0C1AAFACA3" box="[1318,1432,740,765]" direction="north" orientation="latitude" pageId="9" pageNumber="449" precision="1" value="4.54728">N4.54728</geoCoordinate>
,
<geoCoordinate id="EE328CF90450C270FF2FFCEF1E17AD7E" box="[151,288,775,800]" direction="east" orientation="longitude" pageId="9" pageNumber="449" precision="1" value="115.15711">E115.15711</geoCoordinate>
, 131.1 ±
<quantity id="4CFE47DB0450C270FE3FFCEF1EFEAD41" box="[391,457,775,800]" metricMagnitude="0" metricUnit="m" metricValue="6.8" pageId="9" pageNumber="449" unit="m" value="6.8">
<elevation id="002B0D0D0450C270FE3FFCEF1EFEAD41" box="[391,457,775,800]" metricMagnitude="0" metricUnit="m" metricValue="6.8" pageId="9" pageNumber="449" unit="m" value="6.8">6.8 m</elevation>
</quantity>
,
<date id="FFB8CCFE0450C270FE6DFCE01D5EAD7E" box="[469,617,775,800]" pageId="9" pageNumber="449" value="2017-07-29">
<collectingDate id="EFFC35160450C270FE6DFCE01D5EAD7E" box="[469,617,775,800]" pageId="9" pageNumber="449" value="2017-07-29">29 July 2017</collectingDate>
</date>
, 1915 hours, coll.
<collectorName id="26F38FE80450C270FC8FFCE01C9DAD41" box="[823,938,776,799]" pageId="9" pageNumber="449">M. K. Tan</collectorName>
(KB.17.115) (
<collectionCode id="ED1772FB0450C270FBF4FCE01BA5AD7E" box="[1100,1170,776,800]" pageId="9" pageNumber="449">IBER</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BB9EA3E0450C270FF7FFCC41B10ADEE" blockId="9.[151,1437,703,1341]" pageId="9" pageNumber="449">
<emphasis id="B972362C0450C270FF7FFCC41E4BAD1B" bold="true" box="[199,380,812,837]" pageId="9" pageNumber="449">Generic status.</emphasis>
We hypothesize that our species belongs to the genus
<taxonomicName id="4C0691BD0450C270FC57FCC41A7BAD1A" authority="Gorochov, 2000" authorityName="Gorochov" authorityYear="2000" box="[1007,1356,812,837]" class="Insecta" family="Haglotettigoniidae" genus="Endodrelanva" kingdom="Animalia" order="Orthoptera" pageId="9" pageNumber="449" phylum="Arthropoda" rank="genus">
<emphasis id="B972362C0450C270FC57FCC41BBAAD1A" box="[1007,1165,812,836]" italics="true" pageId="9" pageNumber="449">Endodrelanva</emphasis>
Gorochov, 2000
</taxonomicName>
on the basis of these characters: reduced stridulatory apparatus; dorsal field of tegmen longer (very slightly) than lateral field; roundly angular apex of tegmen; absence of outer tympanum; anterior epiphallic part raised (only slightly); and rachis distinctly produced beyond apices of posteromedial epiphallic lobules.
</paragraph>
<paragraph id="8BB9EA3E0450C270FF7FFC541FEAAA8E" blockId="9.[151,1437,703,1341]" pageId="9" pageNumber="449">
This peculiar species has characters of subgenera
<taxonomicName id="4C0691BD0450C270FD4BFC551CFFAD8A" authority="Otte, 1988" authorityName="Otte" authorityYear="1988" box="[755,968,956,980]" class="Insecta" family="Gryllidae" genus="Jorama" higherTaxonomySource="CoL" kingdom="Animalia" order="Orthoptera" pageId="9" pageNumber="449" phylum="Arthropoda" rank="genus">
<emphasis id="B972362C0450C270FD4BFC551C7FAD8A" box="[755,840,957,980]" italics="true" pageId="9" pageNumber="449">Jorama</emphasis>
<bibRefCitation id="EF9797CF0450C270FCF7FC541CFFAD8A" author="Otte" box="[847,968,956,980]" pageId="9" pageNumber="449" refString="Otte, D. (1988) Bark crickets of the Western Pacific Region (Gryllidae: Pteroplistinae). Proceedings of the Academy of Natural Sciences of Philadelphia, 140 (2), 281 - 334 .." type="journal article" year="1988">Otte, 1988</bibRefCitation>
</taxonomicName>
, and
<emphasis id="B972362C0450C270FBBAFC541BACAD8A" box="[1026,1179,956,980]" italics="true" pageId="9" pageNumber="449">Vietlandrevus</emphasis>
Gorochov, 1996 of the genus
<taxonomicName id="4C0691BD0450C270FF58FC081E4AADA6" authorityName="Kirby" authorityYear="1906" box="[224,381,992,1016]" class="Insecta" family="Haglotettigoniidae" genus="Duolandrevus" kingdom="Animalia" order="Orthoptera" pageId="9" pageNumber="449" phylum="Arthropoda" rank="genus">
<emphasis id="B972362C0450C270FF58FC081E4AADA6" box="[224,381,992,1016]" italics="true" pageId="9" pageNumber="449">Duolandrevus</emphasis>
</taxonomicName>
. It shares characters of subgenus
<taxonomicName id="4C0691BD0450C270FD44FC091C65ADA6" authorityName="Otte" authorityYear="1988" box="[764,850,993,1016]" class="Insecta" family="Gryllidae" genus="Jorama" higherTaxonomySource="CoL" kingdom="Animalia" order="Orthoptera" pageId="9" pageNumber="449" phylum="Arthropoda" rank="genus">
<emphasis id="B972362C0450C270FD44FC091C65ADA6" box="[764,850,993,1016]" italics="true" pageId="9" pageNumber="449">Jorama</emphasis>
</taxonomicName>
: metanotal gland relatively small; reduced venation of tegmen (with obsolete mirror), and closely spaced and plate-like apex of posterolateral epiphallic lobe. However, it differs from
<taxonomicName id="4C0691BD0450C270FE08FBC11D31AA1E" authorityName="Otte" authorityYear="1988" box="[432,518,1065,1088]" class="Insecta" family="Gryllidae" genus="Jorama" higherTaxonomySource="CoL" kingdom="Animalia" order="Orthoptera" pageId="9" pageNumber="449" phylum="Arthropoda" rank="genus">
<emphasis id="B972362C0450C270FE08FBC11D31AA1E" box="[432,518,1065,1088]" italics="true" pageId="9" pageNumber="449">Jorama</emphasis>
</taxonomicName>
by the absence of small denticles at apex of posterolateral epiphallic lobe. It also shares characters with species from the subgenus
<emphasis id="B972362C0450C270FD62FBA41C45AA3A" box="[730,882,1100,1124]" italics="true" pageId="9" pageNumber="449">Vietlandrevus</emphasis>
for the following characters: hind wing absent, tegmen with dorsal field longer than lateral field, male anal plate without strong setae at apex, and posterolateral lobes of epiphallus without denticles or tubercles. However, it differs by the presence of developed metanotal gland.
</paragraph>
<paragraph id="8BB9EA3E0450C270FF7FFB341D6CAB62" blockId="9.[151,1437,703,1341]" pageId="9" pageNumber="449">
This species also has characters of genera
<taxonomicName id="4C0691BD0450C270FD79FB351C94AAAA" authority="Otte, 1988" authorityName="Otte" authorityYear="1988" box="[705,931,1244,1268]" class="Insecta" family="Haglotettigoniidae" genus="Kotama" kingdom="Animalia" order="Orthoptera" pageId="9" pageNumber="449" phylum="Arthropoda" rank="genus">
<emphasis id="B972362C0450C270FD79FB351C2DAAAA" box="[705,794,1245,1268]" italics="true" pageId="9" pageNumber="449">Kotama</emphasis>
<bibRefCitation id="EF9797CF0450C270FC9DFB341C94AAAA" author="Otte" box="[805,931,1244,1268]" pageId="9" pageNumber="449" refString="Otte, D. (1988) Bark crickets of the Western Pacific Region (Gryllidae: Pteroplistinae). Proceedings of the Academy of Natural Sciences of Philadelphia, 140 (2), 281 - 334 .." type="journal article" year="1988">Otte, 1988</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C0691BD0450C270FC5FFB351BF4AAAA" authority="Otte, 1988" authorityName="Otte" authorityYear="1988" box="[999,1219,1244,1268]" class="Insecta" family="Haglotettigoniidae" genus="Repapa" kingdom="Animalia" order="Orthoptera" pageId="9" pageNumber="449" phylum="Arthropoda" rank="genus">
<emphasis id="B972362C0450C270FC5FFB351B0BAAAA" box="[999,1084,1245,1268]" italics="true" pageId="9" pageNumber="449">Repapa</emphasis>
<bibRefCitation id="EF9797CF0450C270FBF0FB341BF4AAAA" author="Otte" box="[1096,1219,1244,1268]" pageId="9" pageNumber="449" refString="Otte, D. (1988) Bark crickets of the Western Pacific Region (Gryllidae: Pteroplistinae). Proceedings of the Academy of Natural Sciences of Philadelphia, 140 (2), 281 - 334 .." type="journal article" year="1988">Otte, 1988</bibRefCitation>
</taxonomicName>
but differs by the combination of these characters: presence of tympana (absence in
<taxonomicName id="4C0691BD0450C270FC0FFAE91B27AB46" authorityName="Otte" authorityYear="1988" box="[951,1040,1281,1304]" class="Insecta" family="Haglotettigoniidae" genus="Kotama" kingdom="Animalia" order="Orthoptera" pageId="9" pageNumber="449" phylum="Arthropoda" rank="genus">
<emphasis id="B972362C0450C270FC0FFAE91B27AB46" box="[951,1040,1281,1304]" italics="true" pageId="9" pageNumber="449">Kotama</emphasis>
</taxonomicName>
), paired posteromedial epiphallic process (instead of unpaired in
<taxonomicName id="4C0691BD0450C270FE4DFACD1D7DAB62" authorityName="Otte" authorityYear="1988" box="[501,586,1317,1340]" class="Insecta" family="Haglotettigoniidae" genus="Repapa" kingdom="Animalia" order="Orthoptera" pageId="9" pageNumber="449" phylum="Arthropoda" rank="genus">
<emphasis id="B972362C0450C270FE4DFACD1D7DAB62" box="[501,586,1317,1340]" italics="true" pageId="9" pageNumber="449">Repapa</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<caption id="DF79BAB60450C270FF2FF8341B1EA9AC" box="[151,1065,2012,2034]" httpUri="https://zenodo.org/record/1119249/files/figure.png" pageId="9" pageNumber="449" targetBox="[200,1392,1393,2003]" targetPageId="9">
<paragraph id="8BB9EA3E0450C270FF2FF8341B1EA9AC" blockId="9.[151,1065,2012,2034]" box="[151,1065,2012,2034]" pageId="9" pageNumber="449">
<emphasis id="B972362C0450C270FF2FF8341E25A9AC" bold="true" box="[151,274,2012,2034]" pageId="9" pageNumber="449">FIGURE 7.</emphasis>
<taxonomicName id="4C0691BD0450C270FEA1F8341EC9A9AF" authority="Tan &amp; Wahab, 2017" authorityName="Tan &amp; Wahab" authorityYear="2017" box="[281,510,2012,2033]" class="Insecta" family="Haglotettigoniidae" genus="Endodrelanva" kingdom="Animalia" order="Orthoptera" pageId="9" pageNumber="449" phylum="Arthropoda" rank="species" species="nympha" status="sp. nov.">
<emphasis id="B972362C0450C270FEA1F8341EC9A9AF" box="[281,510,2012,2033]" italics="true" pageId="9" pageNumber="449">Endodrelanva nympha</emphasis>
</taxonomicName>
<emphasis id="B972362C0450C270FDBCF8351D62A9AC" bold="true" box="[516,597,2013,2034]" pageId="9" pageNumber="449">
<taxonomicNameLabel id="A2418B570450C270FDBCF8351D62A9AC" box="[516,597,2013,2034]" pageId="9" pageNumber="449" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
male habitus in dorsal view. Scale bar: 5 mm.
</paragraph>
</caption>
<caption id="DF79BAB60453C273FF2FF8781CA1A9BA" httpUri="https://zenodo.org/record/1119251/files/figure.png" pageId="10" pageNumber="450" targetBox="[180,1415,193,1895]" targetPageId="10">
<paragraph id="8BB9EA3E0453C273FF2FF8781CA1A9BA" blockId="10.[151,1436,1936,2020]" pageId="10" pageNumber="450">
<emphasis id="B972362C0453C273FF2FF8781E22A9FB" bold="true" box="[151,277,1936,1958]" pageId="10" pageNumber="450">FIGURE 8.</emphasis>
<taxonomicName id="4C0691BD0453C273FEA5F8791D32A9F8" authority="Tan &amp; Wahab, 2017" authorityName="Tan &amp; Wahab" authorityYear="2017" box="[285,517,1937,1958]" class="Insecta" family="Haglotettigoniidae" genus="Endodrelanva" kingdom="Animalia" order="Orthoptera" pageId="10" pageNumber="450" phylum="Arthropoda" rank="species" species="nympha" status="sp. nov.">
<emphasis id="B972362C0453C273FEA5F8791D32A9F8" box="[285,517,1937,1958]" italics="true" pageId="10" pageNumber="450">Endodrelanva nympha</emphasis>
</taxonomicName>
<emphasis id="B972362C0453C273FDB5F8791D57A9F8" bold="true" box="[525,608,1937,1958]" pageId="10" pageNumber="450">
<taxonomicNameLabel id="A2418B570453C273FDB5F8791D57A9F8" box="[525,608,1937,1958]" pageId="10" pageNumber="450" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
male: body in profile view (A), tegmen in dorsal view (B), metanotal gland in dorsal view (C), abdominal apex in dorsal (D) and ventral (E) views, genitalia in dorsal (F), ventral (G) and profile (H) views, and posteromedial epiphallic lobule (I). Scale bars: 2 mm (A), 1 mm (BF).
</paragraph>
</caption>
<caption id="DF79BAB60452C272FF2FFC4D1B48ADE5" box="[151,1151,933,955]" httpUri="https://zenodo.org/record/1119253/files/figure.png" pageId="11" pageNumber="451" targetBox="[151,1436,193,926]" targetPageId="11">
<paragraph id="8BB9EA3E0452C272FF2FFC4D1B48ADE5" blockId="11.[151,1151,933,955]" box="[151,1151,933,955]" pageId="11" pageNumber="451">
<emphasis id="B972362C0452C272FF2FFC4D1E25ADE4" bold="true" box="[151,274,933,955]" pageId="11" pageNumber="451">FIGURE 9.</emphasis>
<taxonomicName id="4C0691BD0452C272FEA1FC4D1EC9ADE4" authority="Tan &amp; Wahab, 2017" authorityName="Tan &amp; Wahab" authorityYear="2017" box="[281,510,933,954]" class="Insecta" family="Haglotettigoniidae" genus="Endodrelanva" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="451" phylum="Arthropoda" rank="species" species="nympha" status="sp. nov.">
<emphasis id="B972362C0452C272FEA1FC4D1EC9ADE4" box="[281,510,933,954]" italics="true" pageId="11" pageNumber="451">Endodrelanva nympha</emphasis>
</taxonomicName>
<emphasis id="B972362C0452C272FDBCFC4D1D62ADE4" bold="true" box="[516,597,933,954]" pageId="11" pageNumber="451">
<taxonomicNameLabel id="A2418B570452C272FDBCFC4D1D62ADE4" box="[516,597,933,954]" pageId="11" pageNumber="451" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
male in natural environment in Kuala Belalong forest.
</paragraph>
</caption>
<subSubSection id="C31CB9B50452C272FF7FFC0F1D1AA8D6" pageId="11" pageNumber="451" type="diagnosis">
<paragraph id="8BB9EA3E0452C272FF7FFC0F1AABAA32" blockId="11.[151,1437,999,2032]" pageId="11" pageNumber="451">
<emphasis id="B972362C0452C272FF7FFC0F1E76AA5E" bold="true" box="[199,321,999,1024]" pageId="11" pageNumber="451">Diagnosis.</emphasis>
The new species differs from all known species of
<taxonomicName id="4C0691BD0452C272FC03FC001B6EAA5E" authorityName="Gorochov" authorityYear="2000" box="[955,1113,1000,1024]" class="Insecta" family="Haglotettigoniidae" genus="Endodrelanva" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="451" phylum="Arthropoda" rank="genus">
<emphasis id="B972362C0452C272FC03FC001B6EAA5E" box="[955,1113,1000,1024]" italics="true" pageId="11" pageNumber="451">Endodrelanva</emphasis>
</taxonomicName>
by the combination of the following characters: dorsal field of tegmen only very slightly longer than lateral field; metanotal gland with a rectangular depression, with setose depression, anterior, and posterior margins; anterior epiphallic part only slightly raised; posterolateral epiphallic lobe plate-like at the apex with a dorsal projection and a shorter ventral projection.
</paragraph>
<paragraph id="8BB9EA3E0452C272FF7FFB9F1FDCAB7E" blockId="11.[151,1437,999,2032]" pageId="11" pageNumber="451">
<emphasis id="B972362C0452C272FF7FFB9F1D16AACE" bold="true" box="[199,545,1143,1168]" pageId="11" pageNumber="451">Comparison with congeners.</emphasis>
The new species is similar to
<taxonomicName id="4C0691BD0452C272FC3BFB911BE0AACE" authority="Gorochov, 2016" authorityName="Gorochov" authorityYear="2016" box="[899,1239,1143,1168]" class="Insecta" family="Haglotettigoniidae" genus="Endodrelanva" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="451" phylum="Arthropoda" rank="species" species="peculiaris">
<emphasis id="B972362C0452C272FC3BFB911B23AACE" box="[899,1044,1144,1168]" italics="true" pageId="11" pageNumber="451">E. peculiaris</emphasis>
<bibRefCitation id="EF9797CF0452C272FBA6FB9F1BE0AACE" author="Gorochov" box="[1054,1239,1143,1168]" pageId="11" pageNumber="451" refString="Gorochov, A. V. (2016) Taxonomic studies on the subfamily Landrevinae. Zoosystematica Rossica, 25 (1), 23 - 97" type="journal article" year="2016">Gorochov, 2016</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C0691BD0452C272FAAEFB911E7BAAEA" authority="Gorochov, 2016" authorityName="Gorochov" authorityYear="2016" class="Insecta" family="Haglotettigoniidae" genus="Endodrelanva" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="451" phylum="Arthropoda" rank="species" species="chopardi">
<emphasis id="B972362C0452C272FAAEFB911AABAACE" box="[1302,1436,1144,1168]" italics="true" pageId="11" pageNumber="451">E. chopardi</emphasis>
<bibRefCitation id="EF9797CF0452C272FF2FFB731E7BAAEA" author="Gorochov" box="[151,332,1179,1204]" pageId="11" pageNumber="451" refString="Gorochov, A. V. (2016) Taxonomic studies on the subfamily Landrevinae. Zoosystematica Rossica, 25 (1), 23 - 97" type="journal article" year="2016">Gorochov, 2016</bibRefCitation>
</taxonomicName>
from
<collectingRegion id="49C224DC0452C272FE2AFB731EE1AAEA" box="[402,470,1179,1204]" country="Malaysia" name="Sabah" pageId="11" pageNumber="451">Sabah</collectingRegion>
by the rectangular shape of metanotal gland; and posterolateral epiphallic lobe platelike at the apex with a dorsal and ventral projections (in profile view). It however differs from the two species by apex of dorsal tegminal field (not tapering acutely); anterior epiphallic part only slightly raised (instead of strongly raised).
</paragraph>
<paragraph id="8BB9EA3E0452C272FF7FFAC31D1AA8D6" blockId="11.[151,1437,999,2032]" pageId="11" pageNumber="451">
The new species also differs from
<taxonomicName id="4C0691BD0452C272FD1AFAC41B26AB1A" authority="(Chopard, 1931)" authorityName="Chopard" authorityYear="1931" baseAuthorityName="Chopard" baseAuthorityYear="1931" box="[674,1041,1323,1348]" class="Insecta" family="Haglotettigoniidae" genus="Endodrelanva" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="451" phylum="Arthropoda" rank="species" species="tomentosa">
<emphasis id="B972362C0452C272FD1AFAC41C74AB1D" box="[674,835,1324,1347]" italics="true" pageId="11" pageNumber="451">E. tomentosa</emphasis>
(Chopard, 1931)
</taxonomicName>
from Peninsular
<collectingCountry id="F311AAAE0452C272FAB8FAC31A5EAB1A" box="[1280,1385,1323,1348]" name="Malaysia" pageId="11" pageNumber="451">Malaysia</collectingCountry>
,
<taxonomicName id="4C0691BD0452C272FA3CFAC41ED4AB36" authority="Gorochov, 2016" authorityName="Gorochov" authorityYear="2016" class="Insecta" family="Haglotettigoniidae" genus="Endodrelanva" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="451" phylum="Arthropoda" rank="species" species="macrorachis">
<emphasis id="B972362C0452C272FA3CFAC41E13AB36" italics="true" pageId="11" pageNumber="451">E. macrorachis</emphasis>
<bibRefCitation id="EF9797CF0452C272FE92FAA71ED4AB36" author="Gorochov" box="[298,483,1359,1384]" pageId="11" pageNumber="451" refString="Gorochov, A. V. (2016) Taxonomic studies on the subfamily Landrevinae. Zoosystematica Rossica, 25 (1), 23 - 97" type="journal article" year="2016">Gorochov, 2016</bibRefCitation>
</taxonomicName>
from
<collectingRegion id="49C224DC0452C272FD97FAA71D41AB36" box="[559,630,1359,1384]" country="Malaysia" name="Sabah" pageId="11" pageNumber="451">Sabah</collectingRegion>
, and
<taxonomicName id="4C0691BD0452C272FD01FAB91B63AB36" authority="Tan &amp; Kamaruddin, 2016" authorityName="Tan &amp; Kamaruddin" authorityYear="2016" box="[697,1108,1359,1384]" class="Insecta" family="Haglotettigoniidae" genus="Endodrelanva" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="451" phylum="Arthropoda" rank="species" species="jimini">
<emphasis id="B972362C0452C272FD01FAB91C29AB39" box="[697,798,1360,1384]" italics="true" pageId="11" pageNumber="451">E. jimini</emphasis>
<bibRefCitation id="EF9797CF0452C272FC90FAB81B63AB36" author="Tan" box="[808,1108,1359,1384]" pageId="11" pageNumber="451" refString="Tan, M. K. &amp; Kamaruddin, K. N. (2016) New taxa and notes on some Landrevinae (Orthoptera: Gryllidae) from Malay Peninsula. Zootaxa, 4162 (3), 559 - 570. https: // doi. org / 10.11646 / zootaxa. 4162.3.9." type="journal article" year="2016">Tan &amp; Kamaruddin, 2016</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="F311AAAE0452C272FB19FAA71A24AB36" box="[1185,1299,1359,1384]" name="Singapore" pageId="11" pageNumber="451">Singapore</collectingCountry>
by apex of posterolateral epiphallic lobe (plate-like instead of falcate in profile); and shape of metanotal glands (posterior margin not concave). The new species is similar to
<taxonomicName id="4C0691BD0452C272FCA9FA701C8EABEE" authorityName="Saussure" authorityYear="1877" box="[785,953,1432,1456]" class="Insecta" family="Haglotettigoniidae" genus="Endolandrevus" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="451" phylum="Arthropoda" rank="genus">
<emphasis id="B972362C0452C272FCA9FA701C8EABEE" box="[785,953,1432,1456]" italics="true" pageId="11" pageNumber="451">Endolandrevus</emphasis>
</taxonomicName>
[perhaps
<taxonomicName id="4C0691BD0452C272FB8DFA701FD3AB8A" authority="sensu Gorochov (2016)" authorityName="Gorochov" authorityYear="2016" class="Insecta" family="Haglotettigoniidae" genus="Endodrelanva" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="451" phylum="Arthropoda" rank="genus">
<emphasis id="B972362C0452C272FB8DFA701BE4ABEE" box="[1077,1235,1432,1456]" italics="true" pageId="11" pageNumber="451">Endodrelanva</emphasis>
sensu
<bibRefCitation id="EF9797CF0452C272FA94FA7F1FD3AB8A" author="Gorochov" pageId="11" pageNumber="451" refString="Gorochov, A. V. (2016) Taxonomic studies on the subfamily Landrevinae. Zoosystematica Rossica, 25 (1), 23 - 97" type="journal article" year="2016">Gorochov (2016)</bibRefCitation>
</taxonomicName>
]
<emphasis id="B972362C0452C272FF4AFA541E52AB8A" box="[242,357,1468,1492]" italics="true" pageId="11" pageNumber="451">pubescens</emphasis>
Chopard, 1930 from
<collectingCountry id="F311AAAE0452C272FDECFA531D82AB8A" box="[596,693,1467,1492]" name="Malaysia" pageId="11" pageNumber="451">
<collectingRegion id="49C224DC0452C272FDECFA531D82AB8A" box="[596,693,1467,1492]" country="Malaysia" name="Sarawak" pageId="11" pageNumber="451">Sarawak</collectingRegion>
</collectingCountry>
by shape of male tegmen but differs by smaller size (BL = 14.9
<emphasis id="B972362C0452C272FAC5FA541AABAB8D" box="[1405,1436,1468,1491]" italics="true" pageId="11" pageNumber="451">vs.</emphasis>
<quantity id="4CFE47DB0452C272FF2FFA081FDEABA9" box="[151,233,1504,1528]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="11" pageNumber="451" unit="mm" value="18.0">18 mm</quantity>
; PL = 3.1
<emphasis id="B972362C0452C272FEE5FA091E4BABA6" box="[349,380,1505,1528]" italics="true" pageId="11" pageNumber="451">vs.</emphasis>
<quantity id="4CFE47DB0452C272FE3BFA371EEBABA9" box="[387,476,1503,1528]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="11" pageNumber="451" unit="mm" value="3.5">3.5 mm</quantity>
; HFL = 10.4
<emphasis id="B972362C0452C272FDCAFA091DA6ABA6" box="[626,657,1505,1528]" italics="true" pageId="11" pageNumber="451">vs.</emphasis>
<quantity id="4CFE47DB0452C272FD20FA081DDCABA6" box="[664,747,1504,1528]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="11" pageNumber="451" unit="mm" value="12.0">12 mm</quantity>
), shorter tegmen length (4.9
<emphasis id="B972362C0452C272FB95FA091B7CABA6" box="[1069,1099,1505,1528]" italics="true" pageId="11" pageNumber="451">vs.</emphasis>
<quantity id="4CFE47DB0452C272FBEAFA371BA0ABA6" box="[1106,1175,1503,1528]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="11" pageNumber="451" unit="mm" value="6.0">6 mm</quantity>
); hind tibiae with more subapical spurs; first chord of tegmen substraight (clearly not sinuated), chords closer to one another than with diagonal vein (chords and diagonal vein about equal distance to one another); lateral field of tegmen with four longitudinal veins (instead of six). Until the
<typeStatus id="54BD549C0452C272FD3BF9A31DD2A83A" box="[643,741,1611,1636]" pageId="11" pageNumber="451" type="holotype">holotype</typeStatus>
can be located and genitalia dissected, we cannot confirm that the two species are clearly different.
</paragraph>
</subSubSection>
<subSubSection id="C31CB9B50452C274FF7FF97B1A0AAD96" lastPageId="13" lastPageNumber="453" pageId="11" pageNumber="451" type="description">
<paragraph id="8BB9EA3E0452C272FF7FF97B1CBAA9AE" blockId="11.[151,1437,999,2032]" pageId="11" pageNumber="451">
<emphasis id="B972362C0452C272FF7FF97B1E6EA8F2" bold="true" box="[199,345,1683,1708]" pageId="11" pageNumber="451">Description.</emphasis>
Body dorso-ventrally compressed and fairly densely pubescent (
<figureCitation id="133DF6BB0452C272FB85F97B1BB4A8F2" box="[1085,1155,1683,1708]" captionStart="FIGURE 7" captionStartId="9.[151,250,2012,2034]" captionTargetBox="[200,1392,1393,2003]" captionTargetId="figure@9.[176,1420,1393,2006]" captionTargetPageId="9" captionText="FIGURE 7. Endodrelanva nympha sp. nov. male habitus in dorsal view. Scale bar: 5 mm." httpUri="https://zenodo.org/record/1119249/files/figure.png" pageId="11" pageNumber="451">Fig. 7</figureCitation>
). Head rostrum about as wide as scapus, with apex truncated (in dorsal view) (
<figureCitation id="133DF6BB0452C272FD48F95F1C03A88E" box="[752,820,1719,1744]" captionStart="FIGURE 7" captionStartId="9.[151,250,2012,2034]" captionTargetBox="[200,1392,1393,2003]" captionTargetId="figure@9.[176,1420,1393,2006]" captionTargetPageId="9" captionText="FIGURE 7. Endodrelanva nympha sp. nov. male habitus in dorsal view. Scale bar: 5 mm." httpUri="https://zenodo.org/record/1119249/files/figure.png" pageId="11" pageNumber="451">Fig. 7</figureCitation>
). Maxillary palpi with apical (fifth) segment triangular (longer than broad); distinctly larger, broader and longer than third and subapical (fourth) segments, with apex obtusely rounded; fourth and third segments subequal length (
<figureCitation id="133DF6BB0452C272FCD1F9171CF1A946" box="[873,966,1791,1816]" captionStart="FIGURE 8" captionStartId="10.[151,250,1936,1958]" captionTargetBox="[180,1415,193,1895]" captionTargetId="figure@10.[171,1415,193,1910]" captionTargetPageId="10" captionText="FIGURE 8. Endodrelanva nympha sp. nov. male: body in profile view (A), tegmen in dorsal view (B), metanotal gland in dorsal view (C), abdominal apex in dorsal (D) and ventral (E) views, genitalia in dorsal (F), ventral (G) and profile (H) views, and posteromedial epiphallic lobule (I). Scale bars: 2 mm (A), 1 mm (BF)." httpUri="https://zenodo.org/record/1119251/files/figure.png" pageId="11" pageNumber="451">Fig. 8A</figureCitation>
). Pronotum transverse, about as wide as long, longer but about as wide as head; anterior and posterior margins with strong hairs; disc with anterior and posterior margins straight (
<figureCitation id="133DF6BB0452C272FE7DF8AF1D3DA93E" box="[453,522,1863,1888]" captionStart="FIGURE 7" captionStartId="9.[151,250,2012,2034]" captionTargetBox="[200,1392,1393,2003]" captionTargetId="figure@9.[176,1420,1393,2006]" captionTargetPageId="9" captionText="FIGURE 7. Endodrelanva nympha sp. nov. male habitus in dorsal view. Scale bar: 5 mm." httpUri="https://zenodo.org/record/1119249/files/figure.png" pageId="11" pageNumber="451">Fig. 7</figureCitation>
). Pronotal lateral lobe longer than tall, anterior half taller than posterior half (
<figureCitation id="133DF6BB0452C272FAD6F8AF1F8BA9DA" captionStart="FIGURE 8" captionStartId="10.[151,250,1936,1958]" captionTargetBox="[180,1415,193,1895]" captionTargetId="figure@10.[171,1415,193,1910]" captionTargetPageId="10" captionText="FIGURE 8. Endodrelanva nympha sp. nov. male: body in profile view (A), tegmen in dorsal view (B), metanotal gland in dorsal view (C), abdominal apex in dorsal (D) and ventral (E) views, genitalia in dorsal (F), ventral (G) and profile (H) views, and posteromedial epiphallic lobule (I). Scale bars: 2 mm (A), 1 mm (BF)." httpUri="https://zenodo.org/record/1119251/files/figure.png" pageId="11" pageNumber="451">Fig. 8A</figureCitation>
). Fore tibia with small round inner tympanum but without outer tympanum. Hind tibia inner and outer margins with 5 stout articulated spurs (also known as movable spines) on each dorsal side; and about much smaller basal spines; inner margin with two long apical spurs, outer margin with one shorter apical spur (still longer than subapical spurs). Hind basitarsus with 4 inner and 4 outer denticles.
</paragraph>
<caption id="DF79BAB60455C275FF2FF8BA1A03A936" box="[151,1332,1874,1896]" httpUri="https://zenodo.org/record/1119255/files/figure.png" pageId="12" pageNumber="452" targetBox="[151,1436,193,1853]" targetPageId="12">
<paragraph id="8BB9EA3E0455C275FF2FF8BA1A03A936" blockId="12.[151,1332,1874,1896]" box="[151,1332,1874,1896]" pageId="12" pageNumber="452">
<emphasis id="B972362C0455C275FF2FF8BA1E28A939" bold="true" box="[151,287,1874,1896]" pageId="12" pageNumber="452">FIGURE 10.</emphasis>
Parasitic wasp
<taxonomicName id="4C0691BD0455C275FE05F8BB1EDAA936" box="[445,493,1875,1896]" class="Insecta" family="Crabronidae" genus="Liris" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="452" phylum="Arthropoda" rank="genus">
<emphasis id="B972362C0455C275FE05F8BB1EDAA936" box="[445,493,1875,1896]" italics="true" pageId="12" pageNumber="452">Liris</emphasis>
</taxonomicName>
paralyzing a
<taxonomicName id="4C0691BD0455C275FDCEF8BA1DC2A936" authorityName="Gorochov" authorityYear="1982" box="[630,757,1874,1896]" class="Insecta" family="Gryllidae" kingdom="Animalia" order="Orthoptera" pageId="12" pageNumber="452" phylum="Arthropoda" rank="subFamily" subFamily="Landrevinae">Landrevinae</taxonomicName>
nymph (A); habitus of wasp in profile. Scale bar: 5 mm.
</paragraph>
</caption>
<paragraph id="8BB9EA3E0455C274FF7FF8701B47ACF6" blockId="12.[151,1436,1940,2004]" lastBlockId="13.[151,1437,151,1040]" lastPageId="13" lastPageNumber="453" pageId="12" pageNumber="452">
Tegmen pubescent, extending to middle of 4th abdominal tergite, dorsal field longer than lateral field; eight harp veins (three anterior ones are small); anal area roundly obtuse (
<figureCitation id="133DF6BB0455C275FC1FF8531B37A98A" box="[935,1024,1979,2004]" captionStart="FIGURE 8" captionStartId="10.[151,250,1936,1958]" captionTargetBox="[180,1415,193,1895]" captionTargetId="figure@10.[171,1415,193,1910]" captionTargetPageId="10" captionText="FIGURE 8. Endodrelanva nympha sp. nov. male: body in profile view (A), tegmen in dorsal view (B), metanotal gland in dorsal view (C), abdominal apex in dorsal (D) and ventral (E) views, genitalia in dorsal (F), ventral (G) and profile (H) views, and posteromedial epiphallic lobule (I). Scale bars: 2 mm (A), 1 mm (BF)." httpUri="https://zenodo.org/record/1119251/files/figure.png" pageId="12" pageNumber="452">Fig. 8B</figureCitation>
); lateral field with four longitudinal veins (
<figureCitation id="133DF6BB0454C274FF5AFF7F1E0AAEEE" box="[226,317,151,176]" captionStart="FIGURE 8" captionStartId="10.[151,250,1936,1958]" captionTargetBox="[180,1415,193,1895]" captionTargetId="figure@10.[171,1415,193,1910]" captionTargetPageId="10" captionText="FIGURE 8. Endodrelanva nympha sp. nov. male: body in profile view (A), tegmen in dorsal view (B), metanotal gland in dorsal view (C), abdominal apex in dorsal (D) and ventral (E) views, genitalia in dorsal (F), ventral (G) and profile (H) views, and posteromedial epiphallic lobule (I). Scale bars: 2 mm (A), 1 mm (BF)." httpUri="https://zenodo.org/record/1119251/files/figure.png" pageId="13" pageNumber="453">Fig. 8A</figureCitation>
). Hind wings absent. Metanotal gland with a rectangular depression not separated by a ridge in the middle; depression, anterior and posterior margins with setae (
<figureCitation id="133DF6BB0454C274FCF6FF541C92AE8A" box="[846,933,188,213]" captionStart="FIGURE 8" captionStartId="10.[151,250,1936,1958]" captionTargetBox="[180,1415,193,1895]" captionTargetId="figure@10.[171,1415,193,1910]" captionTargetPageId="10" captionText="FIGURE 8. Endodrelanva nympha sp. nov. male: body in profile view (A), tegmen in dorsal view (B), metanotal gland in dorsal view (C), abdominal apex in dorsal (D) and ventral (E) views, genitalia in dorsal (F), ventral (G) and profile (H) views, and posteromedial epiphallic lobule (I). Scale bars: 2 mm (A), 1 mm (BF)." httpUri="https://zenodo.org/record/1119251/files/figure.png" pageId="13" pageNumber="453">Fig. 8C</figureCitation>
). Ninth abdominal tergite transverse, slightly convex along posterior margin (
<figureCitation id="133DF6BB0454C274FDB6FF371D5BAEA6" box="[526,620,223,248]" captionStart="FIGURE 8" captionStartId="10.[151,250,1936,1958]" captionTargetBox="[180,1415,193,1895]" captionTargetId="figure@10.[171,1415,193,1910]" captionTargetPageId="10" captionText="FIGURE 8. Endodrelanva nympha sp. nov. male: body in profile view (A), tegmen in dorsal view (B), metanotal gland in dorsal view (C), abdominal apex in dorsal (D) and ventral (E) views, genitalia in dorsal (F), ventral (G) and profile (H) views, and posteromedial epiphallic lobule (I). Scale bars: 2 mm (A), 1 mm (BF)." httpUri="https://zenodo.org/record/1119251/files/figure.png" pageId="13" pageNumber="453">Fig. 8D</figureCitation>
). Anal plate broadly rounded at basal half (wider than entire length), tapering to a slightly narrower apical half (and wider than length of apical half), with truncated apex (
<figureCitation id="133DF6BB0454C274FA8BFEEC1AB8AF42" box="[1331,1423,260,285]" captionStart="FIGURE 8" captionStartId="10.[151,250,1936,1958]" captionTargetBox="[180,1415,193,1895]" captionTargetId="figure@10.[171,1415,193,1910]" captionTargetPageId="10" captionText="FIGURE 8. Endodrelanva nympha sp. nov. male: body in profile view (A), tegmen in dorsal view (B), metanotal gland in dorsal view (C), abdominal apex in dorsal (D) and ventral (E) views, genitalia in dorsal (F), ventral (G) and profile (H) views, and posteromedial epiphallic lobule (I). Scale bars: 2 mm (A), 1 mm (BF)." httpUri="https://zenodo.org/record/1119251/files/figure.png" pageId="13" pageNumber="453">Fig. 8D</figureCitation>
). Subgenital plate triangular, about as long as wide, with subacute apex (
<figureCitation id="133DF6BB0454C274FC07FECF1B20AF1E" box="[959,1047,295,320]" captionStart="FIGURE 8" captionStartId="10.[151,250,1936,1958]" captionTargetBox="[180,1415,193,1895]" captionTargetId="figure@10.[171,1415,193,1910]" captionTargetPageId="10" captionText="FIGURE 8. Endodrelanva nympha sp. nov. male: body in profile view (A), tegmen in dorsal view (B), metanotal gland in dorsal view (C), abdominal apex in dorsal (D) and ventral (E) views, genitalia in dorsal (F), ventral (G) and profile (H) views, and posteromedial epiphallic lobule (I). Scale bars: 2 mm (A), 1 mm (BF)." httpUri="https://zenodo.org/record/1119251/files/figure.png" pageId="13" pageNumber="453">Fig. 8E</figureCitation>
). Male genitalia as shown in
<figureCitation id="133DF6BB0454C274FADDFECF1FF8AF3A" captionStart="FIGURE 8" captionStartId="10.[151,250,1936,1958]" captionTargetBox="[180,1415,193,1895]" captionTargetId="figure@10.[171,1415,193,1910]" captionTargetPageId="10" captionText="FIGURE 8. Endodrelanva nympha sp. nov. male: body in profile view (A), tegmen in dorsal view (B), metanotal gland in dorsal view (C), abdominal apex in dorsal (D) and ventral (E) views, genitalia in dorsal (F), ventral (G) and profile (H) views, and posteromedial epiphallic lobule (I). Scale bars: 2 mm (A), 1 mm (BF)." httpUri="https://zenodo.org/record/1119251/files/figure.png" pageId="13" pageNumber="453">Figs. 8FI</figureCitation>
. Male epiphallus narrowly notched, with transverse fold (when viewed dorsally) (
<figureCitation id="133DF6BB0454C274FB22FEA41BC3AF3A" box="[1178,1268,332,357]" captionStart="FIGURE 8" captionStartId="10.[151,250,1936,1958]" captionTargetBox="[180,1415,193,1895]" captionTargetId="figure@10.[171,1415,193,1910]" captionTargetPageId="10" captionText="FIGURE 8. Endodrelanva nympha sp. nov. male: body in profile view (A), tegmen in dorsal view (B), metanotal gland in dorsal view (C), abdominal apex in dorsal (D) and ventral (E) views, genitalia in dorsal (F), ventral (G) and profile (H) views, and posteromedial epiphallic lobule (I). Scale bars: 2 mm (A), 1 mm (BF)." httpUri="https://zenodo.org/record/1119251/files/figure.png" pageId="13" pageNumber="453">Fig. 8F</figureCitation>
); anterior part slightly curved dorso-anteriorly (when viewed laterally) (
<figureCitation id="133DF6BB0454C274FC99FE871C4BAFD6" box="[801,892,367,392]" captionStart="FIGURE 8" captionStartId="10.[151,250,1936,1958]" captionTargetBox="[180,1415,193,1895]" captionTargetId="figure@10.[171,1415,193,1910]" captionTargetPageId="10" captionText="FIGURE 8. Endodrelanva nympha sp. nov. male: body in profile view (A), tegmen in dorsal view (B), metanotal gland in dorsal view (C), abdominal apex in dorsal (D) and ventral (E) views, genitalia in dorsal (F), ventral (G) and profile (H) views, and posteromedial epiphallic lobule (I). Scale bars: 2 mm (A), 1 mm (BF)." httpUri="https://zenodo.org/record/1119251/files/figure.png" pageId="13" pageNumber="453">Fig. 8H</figureCitation>
). Posteromedial epiphallic lobule elongated and directed posteriorly, and with acute apex; very deeply and narrowly notched between the lobules (
<figureCitation id="133DF6BB0454C274FA82FE7C1AB9AFF2" box="[1338,1422,404,429]" captionStart="FIGURE 8" captionStartId="10.[151,250,1936,1958]" captionTargetBox="[180,1415,193,1895]" captionTargetId="figure@10.[171,1415,193,1910]" captionTargetPageId="10" captionText="FIGURE 8. Endodrelanva nympha sp. nov. male: body in profile view (A), tegmen in dorsal view (B), metanotal gland in dorsal view (C), abdominal apex in dorsal (D) and ventral (E) views, genitalia in dorsal (F), ventral (G) and profile (H) views, and posteromedial epiphallic lobule (I). Scale bars: 2 mm (A), 1 mm (BF)." httpUri="https://zenodo.org/record/1119251/files/figure.png" pageId="13" pageNumber="453">
Fig.
<date id="FFB8CCFE0454C274FACCFE7C1AB9AFF2" box="[1396,1422,404,428]" pageId="13" pageNumber="453">8I</date>
</figureCitation>
). Posterolateral epiphallic lobe plate-like at the apex (when viewed laterally); at the apex with a dorsal projection and a shorter ventral projection (when viewed laterally) (
<figureCitation id="133DF6BB0454C274FD4CFE341C67AFAA" box="[756,848,476,501]" captionStart="FIGURE 8" captionStartId="10.[151,250,1936,1958]" captionTargetBox="[180,1415,193,1895]" captionTargetId="figure@10.[171,1415,193,1910]" captionTargetPageId="10" captionText="FIGURE 8. Endodrelanva nympha sp. nov. male: body in profile view (A), tegmen in dorsal view (B), metanotal gland in dorsal view (C), abdominal apex in dorsal (D) and ventral (E) views, genitalia in dorsal (F), ventral (G) and profile (H) views, and posteromedial epiphallic lobule (I). Scale bars: 2 mm (A), 1 mm (BF)." httpUri="https://zenodo.org/record/1119251/files/figure.png" pageId="13" pageNumber="453">Fig. 8H</figureCitation>
). Rachis (or ectophallic fold) elongated, surpassing posterior of posteromedial epiphallic lobule and nearly reaching the posterior end of posterolateral epiphallic lobe (
<figureCitation id="133DF6BB0454C274FF27FDCC1E0EAC62" box="[159,313,548,573]" captionStart="FIGURE 8" captionStartId="10.[151,250,1936,1958]" captionTargetBox="[180,1415,193,1895]" captionTargetId="figure@10.[171,1415,193,1910]" captionTargetPageId="10" captionText="FIGURE 8. Endodrelanva nympha sp. nov. male: body in profile view (A), tegmen in dorsal view (B), metanotal gland in dorsal view (C), abdominal apex in dorsal (D) and ventral (E) views, genitalia in dorsal (F), ventral (G) and profile (H) views, and posteromedial epiphallic lobule (I). Scale bars: 2 mm (A), 1 mm (BF)." httpUri="https://zenodo.org/record/1119251/files/figure.png" pageId="13" pageNumber="453">Figs. 8F, 8G</figureCitation>
). Apodeme principale narrowly forked posteriorly, forming a Y-shape (when viewed dorsally); posterior apex narrows just anterior of the bulbous apex. Formula elongated, narrow anteriorly (when viewed dorsally) (
<figureCitation id="133DF6BB0454C274FEB1FD841E6AACDA" box="[265,349,620,645]" captionStart="FIGURE 8" captionStartId="10.[151,250,1936,1958]" captionTargetBox="[180,1415,193,1895]" captionTargetId="figure@10.[171,1415,193,1910]" captionTargetPageId="10" captionText="FIGURE 8. Endodrelanva nympha sp. nov. male: body in profile view (A), tegmen in dorsal view (B), metanotal gland in dorsal view (C), abdominal apex in dorsal (D) and ventral (E) views, genitalia in dorsal (F), ventral (G) and profile (H) views, and posteromedial epiphallic lobule (I). Scale bars: 2 mm (A), 1 mm (BF)." httpUri="https://zenodo.org/record/1119251/files/figure.png" pageId="13" pageNumber="453">Fig. 8F</figureCitation>
), flattened and raised (when viewed laterally) (
<figureCitation id="133DF6BB0454C274FCD0FD841CF5ACDA" box="[872,962,620,645]" captionStart="FIGURE 8" captionStartId="10.[151,250,1936,1958]" captionTargetBox="[180,1415,193,1895]" captionTargetId="figure@10.[171,1415,193,1910]" captionTargetPageId="10" captionText="FIGURE 8. Endodrelanva nympha sp. nov. male: body in profile view (A), tegmen in dorsal view (B), metanotal gland in dorsal view (C), abdominal apex in dorsal (D) and ventral (E) views, genitalia in dorsal (F), ventral (G) and profile (H) views, and posteromedial epiphallic lobule (I). Scale bars: 2 mm (A), 1 mm (BF)." httpUri="https://zenodo.org/record/1119251/files/figure.png" pageId="13" pageNumber="453">Fig. 8H</figureCitation>
). Ramus not fusing together at the anterior end, with flattened and slightly membranous interior plate near posterior third (
<figureCitation id="133DF6BB0454C274FBB6FD671B54ACF6" box="[1038,1123,655,680]" captionStart="FIGURE 8" captionStartId="10.[151,250,1936,1958]" captionTargetBox="[180,1415,193,1895]" captionTargetId="figure@10.[171,1415,193,1910]" captionTargetPageId="10" captionText="FIGURE 8. Endodrelanva nympha sp. nov. male: body in profile view (A), tegmen in dorsal view (B), metanotal gland in dorsal view (C), abdominal apex in dorsal (D) and ventral (E) views, genitalia in dorsal (F), ventral (G) and profile (H) views, and posteromedial epiphallic lobule (I). Scale bars: 2 mm (A), 1 mm (BF)." httpUri="https://zenodo.org/record/1119251/files/figure.png" pageId="13" pageNumber="453">Fig. 8F</figureCitation>
).
</paragraph>
<paragraph id="8BB9EA3E0454C274FF7FFD5C1E9BAC93" blockId="13.[151,1437,151,1040]" box="[199,428,692,717]" pageId="13" pageNumber="453">Female. Not known.</paragraph>
<paragraph id="8BB9EA3E0454C274FF7FFD3F1BBAADFB" blockId="13.[151,1437,151,1040]" pageId="13" pageNumber="453">Coloration. Generally red brown or brown. Head red brown, antenna scapus lighter brown. Gena, maxillary palpi and clypeus pale to yellow brown. Pronotum red brown. Tegmen red brown. Legs yellow brown. Hind femur mostly light brown, but with a pale band at apical third red brown near knee; hind tibia red brown; hind tarsus yellow brown; spines on hind tibia and tarsus brown with black tips. Thoracic segments pale yellow brown, meta- and meso-sterna brown with dark margins. Abdominal tergites and sternites dark red brown, male anal plate and subgenital plate dark brown. Abdominal sternites and male subgenital plate yellow brown.</paragraph>
<paragraph id="8BB9EA3E0454C274FF7FFC581A0AAD96" blockId="13.[151,1437,151,1040]" box="[199,1341,943,968]" pageId="13" pageNumber="453">Measurements (in mm). BL = 14.9; HL = 2.0; PL = 3.1; PW = 3.4; TL = 4.9; HFL = 10.4; HTL = 6.9.</paragraph>
</subSubSection>
<subSubSection id="C31CB9B50454C274FF7FFC3C1BBFAA4E" pageId="13" pageNumber="453" type="etymology">
<paragraph id="8BB9EA3E0454C274FF7FFC3C1BBFAA4E" blockId="13.[151,1437,151,1040]" pageId="13" pageNumber="453">
<emphasis id="B972362C0454C274FF7FFC3C1E79ADB3" bold="true" box="[199,334,980,1005]" pageId="13" pageNumber="453">Etymology.</emphasis>
The species can be reckoned to a wood nymph (mythological spirit inhabiting wood); in Latin,
<taxonomicName id="4C0691BD0454C274FF2FFC1F1FC6AA4E" authorityName="Tan &amp; Wahab" authorityYear="2017" box="[151,241,1015,1040]" class="Insecta" family="Haglotettigoniidae" genus="Endodrelanva" kingdom="Animalia" order="Orthoptera" pageId="13" pageNumber="453" phylum="Arthropoda" rank="species" species="nympha">nympha</taxonomicName>
= nymph (feminine). These crickets generally tend to hide among woods (
<figureCitation id="133DF6BB0454C274FB8EFC1F1B4DAA4E" box="[1078,1146,1015,1040]" captionStart="FIGURE 9" captionStartId="11.[151,250,933,955]" captionTargetBox="[151,1436,193,926]" captionTargetId="figure@11.[151,1436,193,926]" captionTargetPageId="11" captionText="FIGURE 9. Endodrelanva nympha sp. nov. male in natural environment in Kuala Belalong forest." httpUri="https://zenodo.org/record/1119253/files/figure.png" pageId="13" pageNumber="453">Fig. 9</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>