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<document id="CBCD1D46989679D6AB77F742A0BBBB52" ID-CLB-Dataset="177057" ID-DOI="10.5852/ejt.2022.845.1955" ID-GBIF-Dataset="93834279-6660-4c8b-90af-74cde9c0568f" ID-ISSN="2118-9773" ID-Zenodo-Dep="7258792" ID-ZooBank="CC9F793D-5AD6-4D7C-A25C-7B2C58CAB4C0" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1666888781492" checkinUser="felipe" docAuthor="Enghoff, Henrik" docDate="2022" docId="03AA87BFFFA2C973FDC6F92F8CB44ACC" docLanguage="en" docName="ejt-845_66-100.pdf" docOrigin="European Journal of Taxonomy 845" docSource="http://dx.doi.org/10.5852/ejt.2022.845.1955" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Cryptocorypha cactifer Enghoff 2022, sp. nov." docType="treatment" docUuid="F640EE01-ADBC-46DD-BF9B-0688B8A554E9" docUuidSource="ZooBank" docVersion="8" lastPageNumber="80" masterDocId="FF93FFC7FFA9C97DFF9AFFDE882C4D0E" masterDocTitle="A mountain of millipedes X: Species of Pyrgodesmidae and Cryptodesmidae in the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida)" masterLastPageNumber="100" masterPageNumber="66" pageNumber="77" updateTime="1699408525271" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="E4E08CF00D3534BDC7A44AB9D2B6938C" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="5116B7FB1ED68AACDA637CFBF3A26F1C">
<mods:title id="E932CE4A270EC953E0DC5F5E717A7E68">A mountain of millipedes X: Species of Pyrgodesmidae and Cryptodesmidae in the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida)</mods:title>
</mods:titleInfo>
<mods:name id="B1A0BDBA5A5F54783A7999673E80BFCE" type="personal">
<mods:role id="FE4269CE57F3427849EF4381F6B3EC38">
<mods:roleTerm id="54D7BD970C83B4DA04225965C933B13A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7E756FA1DE85960BA7D8064EBFDC7AD3">Enghoff, Henrik</mods:namePart>
<mods:nameIdentifier id="BD297DB79CB442E009AFF51CDDB5FCBE" type="LSID">9B9D901F-D6C8-4BCA-B11B-CF6EE85B16DC</mods:nameIdentifier>
<mods:affiliation id="20AF8A8BABC2155BC83D5033123352C3">Natural History Museum of Denmark, University of Copenhagen, Universitetsparken 15, DK- 2100 Copenhagen, Denmark.</mods:affiliation>
<mods:nameIdentifier id="BEE2C7ADBFA165F26A11909601F645B4" type="email">henghoff@snm.ku.dk</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="72579DD787D793A6215BA5C5B1D6FB66">text</mods:typeOfResource>
<mods:relatedItem id="BC48EC12260AF264C0182751B6786003" type="host">
<mods:titleInfo id="FFCD69D54FA940ACFA6521111C3BCCAA">
<mods:title id="EE4D7A0F9C119E52E325B22D3E28AFC6">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="37CBB58D77BCDA0AA8D20CD21F8606FB">
<mods:date id="7A7BB5BBD2CDD5130A13626961183B58">2022</mods:date>
<mods:detail id="90934E57945D7ABCFF25C7EF6BA71665" type="pubDate">
<mods:number id="187012B3761C6B097F2837F7181BF2AB">2022-10-24</mods:number>
</mods:detail>
<mods:detail id="D654924A8C0E8A865E316C5A61ACBB96" type="volume">
<mods:number id="9B71D72D0A5A47C94A03F65FBFC4CB8E">845</mods:number>
</mods:detail>
<mods:extent id="F558D345D2970D919DDDE9CD269A2F1C" unit="page">
<mods:start id="F3A10E4A7ACC6416449793F1B27B213D">66</mods:start>
<mods:end id="1D838C43E1069B350B069B6058226CC6">100</mods:end>
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<mods:location id="A724D2EAD166236D364532AE2EF13963">
<mods:url id="3F5455FE4959C0AE21BDE0BEF12242ED">http://dx.doi.org/10.5852/ejt.2022.845.1955</mods:url>
</mods:location>
<mods:classification id="F3290E8566A74B98C54E768021C2BA6E">journal article</mods:classification>
<mods:identifier id="F0D09BC011CDB468A026CAB3CA142680" type="CLB-Dataset">177057</mods:identifier>
<mods:identifier id="0FC57DBEEC8E783C979BA7158B037FDE" type="DOI">10.5852/ejt.2022.845.1955</mods:identifier>
<mods:identifier id="5DD2DC5D3B155188886498228D52AFFC" type="GBIF-Dataset">93834279-6660-4c8b-90af-74cde9c0568f</mods:identifier>
<mods:identifier id="B73ED4B6666735394759CBA9116DAEE6" type="ISSN">2118-9773</mods:identifier>
<mods:identifier id="8A78A18E7459D5DEC58EC745C4FFF2A0" type="Zenodo-Dep">7258792</mods:identifier>
<mods:identifier id="4BF074148C06B4DAF418593B66123EFF" type="ZooBank">CC9F793D-5AD6-4D7C-A25C-7B2C58CAB4C0</mods:identifier>
</mods:mods>
<treatment id="03AA87BFFFA2C973FDC6F92F8CB44ACC" ID-DOI="http://doi.org/10.5281/zenodo.7271435" ID-GBIF-Taxon="203721835" ID-Zenodo-Dep="7271435" LSID="urn:lsid:zoobank.org:act:F640EE01-ADBC-46DD-BF9B-0688B8A554E9" httpUri="http://treatment.plazi.org/id/03AA87BFFFA2C973FDC6F92F8CB44ACC" lastPageId="14" lastPageNumber="80" pageId="11" pageNumber="77">
<subSubSection id="C3196522FFA2C976FDC6F92F8BFB4A02" box="[604,983,1777,1804]" pageId="11" pageNumber="77" type="nomenclature">
<paragraph id="8BBC36A9FFA2C976FDC6F92F8BFB4A02" blockId="11.[358,1230,1777,1874]" box="[604,983,1777,1804]" pageId="11" pageNumber="77">
<heading id="D0F481C5FFA2C976FDC6F92F8BFB4A02" box="[604,983,1777,1804]" centered="true" fontSize="11" level="2" pageId="11" pageNumber="77" reason="2">
<taxonomicName id="4C034D2AFFA2C976FDC6F92F8B594A05" authority="Enghoff, 2022" authorityName="Enghoff" authorityYear="2022" box="[604,885,1777,1804]" class="Diplopoda" family="Pyrgodesmidae" genus="Cryptocorypha" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="77" phylum="Arthropoda" rank="species" species="cactifer" status="sp. nov.">
<emphasis id="B977EABBFFA2C976FDC6F92F8B594A05" bold="true" box="[604,885,1777,1804]" italics="true" pageId="11" pageNumber="77">Cryptocorypha cactifer</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24457C0FFA2C976FCE7F92C8BFB4A02" box="[893,983,1778,1804]" pageId="11" pageNumber="77" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3196522FFA2C976FEFCF8CB8B604A5C" pageId="11" pageNumber="77" type="description">
<paragraph id="8BBC36A9FFA2C976FEFCF8CB8CE24A3E" blockId="11.[358,1230,1777,1874]" box="[358,1230,1813,1840]" pageId="11" pageNumber="77">
<uri id="FF923AABFFA2C976FEFCF8CB8CE24A3E" box="[358,1230,1813,1840]" pageId="11" pageNumber="77">
urn:lsid:zoobank.org:act:
<uuid id="FFA50C7CFFA2C976FD18F8CB8CE24A3E" box="[642,1230,1813,1840]" pageId="11" pageNumber="77">F640EE01-ADBC-46DD-BF9B-0688B8A554E9</uuid>
</uri>
</paragraph>
<paragraph id="8BBC36A9FFA2C976FD7DF8E68B604A5C" blockId="11.[358,1230,1777,1874]" box="[743,844,1848,1874]" pageId="11" pageNumber="77">
<figureCitation id="13382A2CFFA2C976FD7DF8E68B604A5C" box="[743,844,1848,1874]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="12.[189,232,1928,1954]" captionStartId-1="13.[188,231,1892,1918]" captionTargetBox-0="[189,1398,268,1886]" captionTargetBox-1="[189,1398,383,1853]" captionTargetId-0="figure-11@12.[189,1398,268,1886]" captionTargetId-1="figure-48@13.[189,1398,383,1853]" captionTargetPageId-0="12" captionTargetPageId-1="13" captionText-0="Fig. 8. Cryptocorypha cactifer sp. nov., paratype (NHMD 621836), ♂, from Mito Mitatu. A. Head and rings 1–6, lateral view. B. Rings 12–18 and telson, dorsal view. C. Tergal seta. D. Head and collum, frontal view. Scale bars: A–B, D = 0.1 mm; C = 0.005 mm." captionText-1="Fig. 9. Cryptocorypha cactifer sp. nov., paratype (NHMD 621836), ♂, from Mito Mitatu, left gonopod. A. Lateral view. B. Basal part of telopodite, showing jagged?velum. C. (Sub)ventral view. D. Posterior view. E. Anterior view. Abbreviations: cx = coxa; ex = exomere; slm = solenomere; ve = velum? Scale bars: A, C–E = 0.01 mm; B = 0.005 mm." figureDoi-0="http://doi.org/10.5281/zenodo.7258814" figureDoi-1="http://doi.org/10.5281/zenodo.7258818" httpUri-0="https://zenodo.org/record/7258814/files/figure.png" httpUri-1="https://zenodo.org/record/7258818/files/figure.png" pageId="11" pageNumber="77">Figs 8–9</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3196522FFA2C973FF27F8A789FE4C48" lastPageId="14" lastPageNumber="80" pageId="11" pageNumber="77" type="diagnosis">
<paragraph id="8BBC36A9FFA2C976FF27F8A7891B4A9D" blockId="11.[189,311,1913,1939]" box="[189,311,1913,1939]" pageId="11" pageNumber="77">
<heading id="D0F481C5FFA2C976FF27F8A7891B4A9D" bold="true" box="[189,311,1913,1939]" fontSize="11" level="3" pageId="11" pageNumber="77" reason="3">
<emphasis id="B977EABBFFA2C976FF27F8A7891B4A9D" bold="true" box="[189,311,1913,1939]" pageId="11" pageNumber="77">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BBC36A9FFA2C976FF27F8768D5B4AE8" blockId="11.[185,1399,1960,2022]" pageId="11" pageNumber="77">
Identical with
<taxonomicName id="4C034D2AFFA2C976FEC1F8778A394ACC" authority="Enghoff, 2022" authorityName="Enghoff" authorityYear="2022" box="[347,533,1961,1987]" class="Diplopoda" family="Pyrgodesmidae" genus="Cryptocorypha" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="77" phylum="Arthropoda" rank="species" species="geminiramus" status="sp. nov.">
<emphasis id="B977EABBFFA2C976FEC1F8778A394ACC" box="[347,533,1961,1987]" italics="true" pageId="11" pageNumber="77">C. geminiramus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24457C0FFA2C976FD87F8778A5F4ACD" box="[541,627,1961,1987]" pageId="11" pageNumber="77" rank="species">sp. nov.</taxonomicNameLabel>
in almost all characters, but differs from the smaller
<taxonomicName id="4C034D2AFFA2C976FB23F8778D5F4ACC" authorityName="Enghoff" authorityYear="2022" box="[1209,1395,1961,1987]" class="Diplopoda" family="Pyrgodesmidae" genus="Cryptocorypha" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="77" phylum="Arthropoda" rank="species" species="geminiramus">
<emphasis id="B977EABBFFA2C976FB23F8778D5F4ACC" box="[1209,1395,1961,1987]" italics="true" pageId="11" pageNumber="77">C. geminiramus</emphasis>
</taxonomicName>
and from all other species of
<taxonomicName id="4C034D2AFFA2C976FDB8F8158AF94AEB" authorityName="Attems" authorityYear="1807" box="[546,725,1995,2021]" class="Diplopoda" family="Pyrgodesmidae" genus="Cryptocorypha" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="77" phylum="Arthropoda" rank="genus">
<emphasis id="B977EABBFFA2C976FDB8F8158AF94AEB" box="[546,725,1995,2021]" italics="true" pageId="11" pageNumber="77">Cryptocorypha</emphasis>
</taxonomicName>
by the presence of setae on podous body rings (
<figureCitation id="13382A2CFFA2C976FA84F8158D4B4AE8" box="[1310,1383,1995,2022]" captionStart="Fig" captionStartId="12.[189,232,1928,1954]" captionTargetBox="[189,1398,268,1886]" captionTargetId="figure-11@12.[189,1398,268,1886]" captionTargetPageId="12" captionText="Fig. 8. Cryptocorypha cactifer sp. nov., paratype (NHMD 621836), ♂, from Mito Mitatu. A. Head and rings 1–6, lateral view. B. Rings 12–18 and telson, dorsal view. C. Tergal seta. D. Head and collum, frontal view. Scale bars: A–B, D = 0.1 mm; C = 0.005 mm." figureDoi="http://doi.org/10.5281/zenodo.7258814" httpUri="https://zenodo.org/record/7258814/files/figure.png" pageId="11" pageNumber="77">Fig. 8</figureCitation>
).
</paragraph>
<caption id="DF7C6621FFA5C971FF27F8568B594AE7" ID-DOI="http://doi.org/10.5281/zenodo.7258814" ID-Zenodo-Dep="7258814" httpUri="https://zenodo.org/record/7258814/files/figure.png" pageId="12" pageNumber="78" startId="12.[189,232,1928,1954]" targetBox="[189,1398,268,1886]" targetPageId="12">
<paragraph id="8BBC36A9FFA5C971FF27F8568B594AE7" blockId="12.[189,1398,1928,2025]" pageId="12" pageNumber="78">
<emphasis id="B977EABBFFA5C971FF27F85689274AAC" bold="true" box="[189,267,1928,1954]" pageId="12" pageNumber="78">Fig. 8.</emphasis>
<taxonomicName id="4C034D2AFFA5C971FE89F8568A054AAC" authority="Enghoff, 2022" authorityName="Enghoff" authorityYear="2022" box="[275,553,1928,1954]" class="Diplopoda" family="Pyrgodesmidae" genus="Cryptocorypha" kingdom="Animalia" order="Polydesmida" pageId="12" pageNumber="78" phylum="Arthropoda" rank="species" species="cactifer" status="sp. nov.">
<emphasis id="B977EABBFFA5C971FE89F8568A054AAC" box="[275,553,1928,1954]" italics="true" pageId="12" pageNumber="78">Cryptocorypha cactifer</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24457C0FFA5C971FDAAF8568AA14AAC" box="[560,653,1928,1954]" pageId="12" pageNumber="78" rank="species">sp. nov.</taxonomicNameLabel>
, paratype (NHMD 621836), ♂, from Mito Mitatu.
<emphasis id="B977EABBFFA5C971FB79F8568CD44AAC" bold="true" box="[1251,1272,1928,1954]" pageId="12" pageNumber="78">A</emphasis>
. Head and rings 1–6, lateral view.
<emphasis id="B977EABBFFA5C971FE41F87589C04ACB" bold="true" box="[475,492,1963,1989]" pageId="12" pageNumber="78">B</emphasis>
. Rings 12–18 and telson, dorsal view.
<emphasis id="B977EABBFFA5C971FC5FF8758BCE4ACB" bold="true" box="[965,994,1963,1989]" pageId="12" pageNumber="78">C.</emphasis>
Tergal seta.
<emphasis id="B977EABBFFA5C971FBE4F8758CBD4ACB" bold="true" box="[1150,1169,1963,1989]" pageId="12" pageNumber="78">D</emphasis>
. Head and collum, frontal view. Scale bars: A–B, D = 0.1 mm; C = 0.005 mm.
</paragraph>
</caption>
<paragraph id="8BBC36A9FFA4C973FF27FED789FE4C48" blockId="13.[189,1399,265,327]" lastBlockId="14.[189,1399,265,326]" lastPageId="14" lastPageNumber="80" pageId="13" pageNumber="79">
Further differs from
<taxonomicName id="4C034D2AFFA4C970FE34FED78A474C2D" authorityName="Enghoff" authorityYear="2022" box="[430,619,265,291]" class="Diplopoda" family="Pyrgodesmidae" genus="Cryptocorypha" kingdom="Animalia" order="Polydesmida" pageId="13" pageNumber="79" phylum="Arthropoda" rank="species" species="geminiramus">
<emphasis id="B977EABBFFA4C970FE34FED78A474C2D" box="[430,619,265,291]" italics="true" pageId="13" pageNumber="79">C. geminiramus</emphasis>
</taxonomicName>
by the smooth part of head capsule extending well above antennal sockets (
<figureCitation id="13382A2CFFA4C970FEB8FEF289524C48" box="[290,382,300,326]" captionStart="Fig" captionStartId="12.[189,232,1928,1954]" captionTargetBox="[189,1398,268,1886]" captionTargetId="figure-11@12.[189,1398,268,1886]" captionTargetPageId="12" captionText="Fig. 8. Cryptocorypha cactifer sp. nov., paratype (NHMD 621836), ♂, from Mito Mitatu. A. Head and rings 1–6, lateral view. B. Rings 12–18 and telson, dorsal view. C. Tergal seta. D. Head and collum, frontal view. Scale bars: A–B, D = 0.1 mm; C = 0.005 mm." figureDoi="http://doi.org/10.5281/zenodo.7258814" httpUri="https://zenodo.org/record/7258814/files/figure.png" pageId="13" pageNumber="79">Fig. 8D</figureCitation>
, compare with
<figureCitation id="13382A2CFFA4C970FDB4FEF28AA64C48" box="[558,650,300,326]" captionStart="Fig" captionStartId="5.[188,231,1857,1883]" captionTargetBox="[192,1395,445,1819]" captionTargetId="figure-34@5.[190,1399,445,1820]" captionTargetPageId="5" captionText="Fig. 2. Cryptocorypha geminiramus sp. nov. A–E. Paratype (NHMD 621824), ♂, from Kidatu. F. Paratype (NHMD 621829), ♂, from Mitu Mitato. A. Head, frontal view. B. Head, (sub)posterior view. C. Head, sublateral view (left side). D. Bases of first and second pairs of legs. E. First and second pair of legs. F. Leg of second pair, tarsus. Abbreviations: fu = epibiotic fungus, possibly Basiodiobolus sp.; go = gonopore. Scale bars: A–B = 0.1 mm; C = 0.05 mm; D–F = 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.7258798" httpUri="https://zenodo.org/record/7258798/files/figure.png" pageId="13" pageNumber="79">Fig. 2A</figureCitation>
), by having the gonopod coxa (
<emphasis id="B977EABBFFA4C970FC6FFEF38C234C48" box="[1013,1039,301,326]" italics="true" pageId="13" pageNumber="79">cx</emphasis>
) more regularly hemispherical (
<figureCitation id="13382A2CFFA7C973FF5CFED7896B4C2D" box="[198,327,265,291]" captionStart="Fig" captionStartId="13.[188,231,1892,1918]" captionTargetBox="[189,1398,383,1853]" captionTargetId="figure-48@13.[189,1398,383,1853]" captionTargetPageId="13" captionText="Fig. 9. Cryptocorypha cactifer sp. nov., paratype (NHMD 621836), ♂, from Mito Mitatu, left gonopod. A. Lateral view. B. Basal part of telopodite, showing jagged?velum. C. (Sub)ventral view. D. Posterior view. E. Anterior view. Abbreviations: cx = coxa; ex = exomere; slm = solenomere; ve = velum? Scale bars: A, C–E = 0.01 mm; B = 0.005 mm." figureDoi="http://doi.org/10.5281/zenodo.7258818" httpUri="https://zenodo.org/record/7258818/files/figure.png" pageId="14" pageNumber="80">Fig. 9A, C</figureCitation>
) and by having a jagged semicircular ridge (velum?,
<emphasis id="B977EABBFFA7C973FC52FED48BCD4C2D" box="[968,993,266,291]" italics="true" pageId="14" pageNumber="80">ve</emphasis>
) on the basal part of the gonopod telopodite (
<figureCitation id="13382A2CFFA7C973FEDEFEF289E94C48" box="[324,453,300,326]" captionStart="Fig" captionStartId="13.[188,231,1892,1918]" captionTargetBox="[189,1398,383,1853]" captionTargetId="figure-48@13.[189,1398,383,1853]" captionTargetPageId="13" captionText="Fig. 9. Cryptocorypha cactifer sp. nov., paratype (NHMD 621836), ♂, from Mito Mitatu, left gonopod. A. Lateral view. B. Basal part of telopodite, showing jagged?velum. C. (Sub)ventral view. D. Posterior view. E. Anterior view. Abbreviations: cx = coxa; ex = exomere; slm = solenomere; ve = velum? Scale bars: A, C–E = 0.01 mm; B = 0.005 mm." figureDoi="http://doi.org/10.5281/zenodo.7258818" httpUri="https://zenodo.org/record/7258818/files/figure.png" pageId="14" pageNumber="80">Fig. 9B, D</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF7C6621FFA4C970FF26F8BA8AB14AE7" ID-DOI="http://doi.org/10.5281/zenodo.7258818" ID-Zenodo-Dep="7258818" httpUri="https://zenodo.org/record/7258818/files/figure.png" pageId="13" pageNumber="79" startId="13.[188,231,1892,1918]" targetBox="[189,1398,383,1853]" targetPageId="13">
<paragraph id="8BBC36A9FFA4C970FF26F8BA8AB14AE7" blockId="13.[188,1399,1892,2025]" pageId="13" pageNumber="79">
<emphasis id="B977EABBFFA4C970FF26F8BA89274A71" bold="true" box="[188,267,1892,1919]" pageId="13" pageNumber="79">Fig. 9.</emphasis>
<taxonomicName id="4C034D2AFFA4C970FE88F8BB8A0B4A71" authority="Enghoff, 2022" authorityName="Enghoff" authorityYear="2022" box="[274,551,1893,1919]" class="Diplopoda" family="Pyrgodesmidae" genus="Cryptocorypha" kingdom="Animalia" order="Polydesmida" pageId="13" pageNumber="79" phylum="Arthropoda" rank="species" species="cactifer" status="sp. nov.">
<emphasis id="B977EABBFFA4C970FE88F8BB8A0B4A71" box="[274,551,1893,1919]" italics="true" pageId="13" pageNumber="79">Cryptocorypha cactifer</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24457C0FFA4C970FDB7F8BB8AA64A71" box="[557,650,1893,1919]" pageId="13" pageNumber="79" rank="species">sp. nov.</taxonomicNameLabel>
, paratype (NHMD 621836), ♂, from Mito Mitatu, left gonopod.
<emphasis id="B977EABBFFA4C970FF27F85688FE4AAC" bold="true" box="[189,210,1928,1954]" pageId="13" pageNumber="79">A</emphasis>
. Lateral view.
<emphasis id="B977EABBFFA4C970FE1AF85689B84AAC" bold="true" box="[384,404,1928,1954]" pageId="13" pageNumber="79">B</emphasis>
. Basal part of telopodite, showing jagged?velum.
<emphasis id="B977EABBFFA4C970FC7CF8568BD74AAC" bold="true" box="[998,1019,1928,1954]" pageId="13" pageNumber="79">C</emphasis>
. (Sub)ventral view.
<emphasis id="B977EABBFFA4C970FB73F8568CD24AAC" bold="true" box="[1257,1278,1928,1954]" pageId="13" pageNumber="79">D</emphasis>
. Posterior view.
<emphasis id="B977EABBFFA4C970FE99F875893B4ACB" bold="true" box="[259,279,1963,1989]" pageId="13" pageNumber="79">E</emphasis>
. Anterior view. Abbreviations:
<emphasis id="B977EABBFFA4C970FD10F8728A884ACB" box="[650,676,1964,1989]" italics="true" pageId="13" pageNumber="79">cx</emphasis>
= coxa;
<emphasis id="B977EABBFFA4C970FC94F8728B044ACB" box="[782,808,1964,1989]" italics="true" pageId="13" pageNumber="79">ex</emphasis>
= exomere;
<emphasis id="B977EABBFFA4C970FC27F8758BCA4ACB" box="[957,998,1963,1989]" italics="true" pageId="13" pageNumber="79">slm</emphasis>
= solenomere;
<emphasis id="B977EABBFFA4C970FB04F8728C944ACB" box="[1182,1208,1964,1989]" italics="true" pageId="13" pageNumber="79">ve</emphasis>
= velum? Scale bars: A, C–E = 0.01 mm; B = 0.005 mm.
</paragraph>
</caption>
<subSubSection id="C3196522FFA7C973FF27FEB089BD4CD4" pageId="14" pageNumber="80" type="etymology">
<paragraph id="8BBC36A9FFA7C973FF27FEB089694C86" blockId="14.[189,325,366,392]" box="[189,325,366,392]" pageId="14" pageNumber="80">
<heading id="D0F481C5FFA7C973FF27FEB089694C86" bold="true" box="[189,325,366,392]" fontSize="11" level="3" pageId="14" pageNumber="80" reason="3">
<emphasis id="B977EABBFFA7C973FF27FEB089694C86" bold="true" box="[189,325,366,392]" pageId="14" pageNumber="80">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BBC36A9FFA7C973FF27FE4289BD4CD4" blockId="14.[189,1399,412,474]" pageId="14" pageNumber="80">The species epithet (noun in apposition) means ‘bearer of cacti’ and refers to the some cactus-shaped metazonital setae.</paragraph>
</subSubSection>
<subSubSection id="C3196522FFA7C973FF27FDDE8D714998" pageId="14" pageNumber="80" type="materials_examined">
<paragraph id="8BBC36A9FFA7C973FF27FDDE8B2C4F15" blockId="14.[189,768,512,539]" box="[189,768,512,539]" pageId="14" pageNumber="80">
<emphasis id="B977EABBFFA7C973FF27FDDE89814F14" bold="true" box="[189,429,512,538]" pageId="14" pageNumber="80">Material examined</emphasis>
(
<specimenCount id="9D05FD20FFA7C973FE24FDDF8A2E4F15" box="[446,514,513,539]" pageId="14" pageNumber="80" type="male">3 ♂♂</specimenCount>
,
<specimenCount id="9D05FD20FFA7C973FD94FDDF8A164F15" box="[526,570,513,539]" pageId="14" pageNumber="80" type="female">1 ♀</specimenCount>
,
<specimenCount id="9D05FD20FFA7C973FDD3FDDF8AEE4F15" box="[585,706,513,539]" pageId="14" pageNumber="80" type="subadult">3 subadult</specimenCount>
<specimenCount id="9D05FD20FFA7C973FD50FDDF8AD84F15" box="[714,756,513,539]" pageId="14" pageNumber="80" type="female">♀♀</specimenCount>
)
</paragraph>
<paragraph id="8BBC36A9FFA7C973FF70FDF18D5B4F9E" blockId="14.[189,1399,559,657]" pageId="14" pageNumber="80">
<materialsCitation id="3B6B3CF4FFA7C973FF70FDF18D5B4F9E" ID-GBIF-Occurrence="3948948308" collectingDate="2014-10-21" collectingMethod="pitfall trap" collectionCode="NHMD" collectorName="J. Malumbres-Olarte" country="Tanzania" county="Udzungwa Mountains Natl Park" elevation="1448" latitude="-7.6793056" location="Kidatu" longLatPrecision="1" longitude="36.918583" municipality="Udzungwa Mountains" pageId="14" pageNumber="80" specimenCode="NHMD 621833" specimenCount="1" specimenCount-male="1" stateProvince="Morogoro Region" typeStatus="holotype">
<emphasis id="B977EABBFFA7C973FF70FDF189714F47" bold="true" box="[234,349,559,585]" pageId="14" pageNumber="80">
<typeStatus id="54B8880BFFA7C973FF70FDF189714F47" box="[234,349,559,585]" pageId="14" pageNumber="80" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="F3147639FFA7C973FF27FD8D897D4F63" box="[189,337,595,621]" name="Tanzania" pageId="14" pageNumber="80">TANZANIA</collectingCountry>
•
<specimenCount id="9D05FD20FFA7C973FEE4FD8D89BB4F63" box="[382,407,595,621]" pageId="14" pageNumber="80" type="male">♂</specimenCount>
;
<collectingRegion id="49C7F84BFFA7C973FE34FD8D8AA34F62" box="[430,655,594,621]" country="Tanzania" name="Morogoro" pageId="14" pageNumber="80">Morogoro Region</collectingRegion>
,
<collectingCounty id="62DD4E25FFA7C973FD3EFD8C8C6D4F62" box="[676,1089,594,620]" pageId="14" pageNumber="80">Udzungwa Mountains Natl Park</collectingCounty>
,
<location id="8EDC6072FFA7C973FBCDFD8C8C874F62" LSID="urn:lsid:plazi:treatment:03AA87BFFFA2C973FDC6F92F8CB44ACC:8EDC6072FFA7C973FBCDFD8C8C874F62" box="[1111,1195,594,620]" country="Tanzania" county="Udzungwa Mountains Natl Park" latitude="-7.6793056" longLatPrecision="1" longitude="36.918583" municipality="Udzungwa Mountains" name="Kidatu" pageId="14" pageNumber="80" stateProvince="Morogoro Region">Kidatu</location>
;
<geoCoordinate id="EE37506EFFA7C973FB5BFD8F8D5D4F63" box="[1217,1393,593,621]" degrees="07" direction="south" minutes="40" orientation="latitude" pageId="14" pageNumber="80" precision="1" seconds="45.5" value="-7.6793056">07°40′45.5″ S</geoCoordinate>
,
<geoCoordinate id="EE37506EFFA7C973FF27FDAA89484F9E" box="[189,356,628,656]" degrees="36" direction="east" minutes="55" orientation="longitude" pageId="14" pageNumber="80" precision="1" seconds="06.9" value="36.918583">36°55′06.9″ E</geoCoordinate>
;
<elevation id="002ED19AFFA7C973FEF4FDA88A284F9E" box="[366,516,630,656]" metricMagnitude="3" metricUnit="m" metricValue="1.448" pageId="14" pageNumber="80" unit="m" value="1448.0">
<quantity id="4CFB9B4CFFA7C973FEF4FDA889EA4F9E" box="[366,454,630,656]" metricMagnitude="3" metricUnit="m" metricValue="1.448" pageId="14" pageNumber="80" unit="m" value="1448.0">1448 m</quantity>
a.s.l.
</elevation>
;
<date id="FFBD1069FFA7C973FD95FDA88A8B4F9E" box="[527,679,630,656]" pageId="14" pageNumber="80" value="2014-10-21">
<collectingDate id="EFF9E981FFA7C973FD95FDA88A8B4F9E" box="[527,679,630,656]" pageId="14" pageNumber="80" value="2014-10-21">21 Oct. 2014</collectingDate>
</date>
;
<collectorName id="26F6537FFFA7C973FD28FDA88B8F4F9E" box="[690,931,630,656]" pageId="14" pageNumber="80">J. Malumbres-Olarte</collectorName>
leg.;
<collectingMethod id="52424EBEFFA7C973FC7BFDA88C714F9E" box="[993,1117,630,656]" pageId="14" pageNumber="80">pitfall trap</collectingMethod>
, Plot 5;
<specimenCode id="DBA59ED2FFA7C973FB22FDA88D5F4F9E" box="[1208,1395,630,656]" collectionCode="NHMD" pageId="14" pageNumber="80">NHMD 621833</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BBC36A9FFA7C973FF70FD6989334900" blockId="14.[189,1399,695,1038]" pageId="14" pageNumber="80">
<materialsCitation id="3B6B3CF4FFA7C973FF70FD6989554E19" ID-GBIF-Occurrence="3948948325" collectingDate="2014-10-21" collectingMethod="pitfall traps" collectionCode="NHMD" collectorName="J. Malumbres-Olarte &amp; Plot" country="Tanzania" county="Udzungwa Mountains Natl Park" elevation="1448" latitude="-7.6793056" location="TANZANIA" longLatPrecision="1" longitude="36.918583" municipality="Udzungwa Mountains" pageId="14" pageNumber="80" specimenCode="NHMD 621834" specimenCount="3" specimenCount-female="1" specimenCount-subadult="2" stateProvince="Morogoro" typeStatus="paratype">
<emphasis id="B977EABBFFA7C973FF70FD6989444FDF" bold="true" box="[234,360,695,721]" pageId="14" pageNumber="80">
<typeStatus id="54B8880BFFA7C973FF70FD6989444FDF" box="[234,360,695,721]" pageId="14" pageNumber="80" type="paratype">Paratypes</typeStatus>
</emphasis>
<collectingCountry id="F3147639FFA7C973FF27FD05897D4FFB" box="[189,337,731,757]" name="Tanzania" pageId="14" pageNumber="80">TANZANIA</collectingCountry>
•
<specimenCount id="9D05FD20FFA7C973FEF0FD0489C84FFA" box="[362,484,730,756]" pageId="14" pageNumber="80" type="subadult">2 subadult</specimenCount>
<specimenCount id="9D05FD20FFA7C973FE76FD058A354FFB" box="[492,537,731,757]" pageId="14" pageNumber="80" type="female">♀♀</specimenCount>
; same collection data as for holotype; summed catch from 2
<collectingMethod id="52424EBEFFA7C973FB72FD048D5E4FFB" box="[1256,1394,730,757]" pageId="14" pageNumber="80">pitfall traps</collectingMethod>
;
<specimenCode id="DBA59ED2FFA7C973FF27FD2089554E19" box="[189,377,765,792]" collectionCode="NHMD" pageId="14" pageNumber="80">NHMD 621834</specimenCode>
</materialsCitation>
•
<materialsCitation id="3B6B3CF4FFA7C973FE03FD208A834E32" ID-GBIF-Occurrence="3948948363" collectingDate="2014-10-21" collectingMethod="pitfall trap" collectionCode="NHMD" collectorName="Plot" country="Tanzania" county="Udzungwa Mountains Natl Park" elevation="1482" latitude="-7.678361" location="Mountains Natl Park" longLatPrecision="1" longitude="36.918556" municipality="Udzungwa Mountains" pageId="14" pageNumber="80" specimenCode="NHMD 621835" specimenCount="1" specimenCount-female="1" stateProvince="Morogoro" typeStatus="paratype">
<specimenCount id="9D05FD20FFA7C973FE03FD2089E54E16" box="[409,457,766,792]" pageId="14" pageNumber="80" type="female">1 ♀</specimenCount>
; same collection data as for holotype;
<geoCoordinate id="EE37506EFFA7C973FC3FFD228C624E16" box="[933,1102,764,792]" degrees="07" direction="south" minutes="40" orientation="latitude" pageId="14" pageNumber="80" precision="1" seconds="42.1" value="-7.678361">07°40′42.1″ S</geoCoordinate>
,
<geoCoordinate id="EE37506EFFA7C973FBC5FD228D274E16" box="[1119,1291,764,792]" degrees="36" direction="east" minutes="55" orientation="longitude" pageId="14" pageNumber="80" precision="1" seconds="06.8" value="36.918556">36°55′06.8″ E</geoCoordinate>
;
<elevation id="002ED19AFFA7C973FA80FD2088DA4E35" metricMagnitude="3" metricUnit="m" metricValue="1.4819999999999998" pageId="14" pageNumber="80" unit="m" value="1482.0">
<quantity id="4CFB9B4CFFA7C973FA80FD208D5B4E16" box="[1306,1399,766,792]" metricMagnitude="3" metricUnit="m" metricValue="1.4819999999999998" pageId="14" pageNumber="80" unit="m" value="1482.0">1482 m</quantity>
a.s.l.
</elevation>
;
<collectingMethod id="52424EBEFFA7C973FE9CFCFF89A44E35" box="[262,392,801,827]" pageId="14" pageNumber="80">pitfall trap</collectingMethod>
, Plot 6;
<specimenCode id="DBA59ED2FFA7C973FE69FCFF8A834E32" box="[499,687,801,828]" collectionCode="NHMD" pageId="14" pageNumber="80">NHMD 621835</specimenCode>
</materialsCitation>
•
<materialsCitation id="3B6B3CF4FFA7C973FD55FCFF8D0B4E50" ID-GBIF-Occurrence="3948948322" collectingDate="2014-10-27" collectingMethod="pitfall trap" collectionCode="NHMD" collectorName="Plot" country="Tanzania" county="Udzungwa Mountains Natl Park" elevation="1552" latitude="-7.827722" location="Mito Mitatu" longLatPrecision="1" longitude="36.840553" municipality="Udzungwa Mountains" pageId="14" pageNumber="80" specimenCode="NHMD 621836" specimenCount="1" specimenCount-male="1" stateProvince="Morogoro" typeStatus="paratype">
<specimenCount id="9D05FD20FFA7C973FD55FCFF8B2D4E35" box="[719,769,801,827]" pageId="14" pageNumber="80" type="male">1 ♂</specimenCount>
; same collection data as for holotype;
<location id="8EDC6072FFA7C973FB46FCFF8D5E4E35" LSID="urn:lsid:plazi:treatment:03AA87BFFFA2C973FDC6F92F8CB44ACC:8EDC6072FFA7C973FB46FCFF8D5E4E35" box="[1244,1394,801,827]" country="Tanzania" county="Udzungwa Mountains Natl Park" latitude="-7.827722" longLatPrecision="1" longitude="36.840553" municipality="Udzungwa Mountains" name="Mito Mitatu" pageId="14" pageNumber="80" stateProvince="Morogoro">Mito Mitatu</location>
;
<geoCoordinate id="EE37506EFFA7C973FF27FC9C89484E50" box="[189,356,834,862]" degrees="07" direction="south" minutes="49" orientation="latitude" pageId="14" pageNumber="80" precision="1" seconds="39.8" value="-7.827722">07°49′39.8″ S</geoCoordinate>
,
<geoCoordinate id="EE37506EFFA7C973FEEBFC9C8A374E51" box="[369,539,834,863]" degrees="36" direction="east" minutes="50" orientation="longitude" pageId="14" pageNumber="80" precision="1" seconds="26.0" value="36.840553">36°50′26.0″ E</geoCoordinate>
;
<elevation id="002ED19AFFA7C973FDB2FC9A8AEF4E50" box="[552,707,836,863]" metricMagnitude="3" metricUnit="m" metricValue="1.552" pageId="14" pageNumber="80" unit="m" value="1552.0">
<quantity id="4CFB9B4CFFA7C973FDB2FC9A8AAE4E51" box="[552,642,836,863]" metricMagnitude="3" metricUnit="m" metricValue="1.552" pageId="14" pageNumber="80" unit="m" value="1552.0">1552 m</quantity>
a.s.l.
</elevation>
;
<date id="FFBD1069FFA7C973FD4BFC9A8B434E50" box="[721,879,836,862]" pageId="14" pageNumber="80" value="2014-10-27">
<collectingDate id="EFF9E981FFA7C973FD4BFC9A8B434E50" box="[721,879,836,862]" pageId="14" pageNumber="80" value="2014-10-27">27 Oct. 2014</collectingDate>
</date>
;
<collectingMethod id="52424EBEFFA7C973FCE6FC9A8BD74E51" box="[892,1019,836,863]" pageId="14" pageNumber="80">pitfall trap</collectingMethod>
, Plot 15;
<specimenCode id="DBA59ED2FFA7C973FBF7FC9B8D0B4E50" box="[1133,1319,836,863]" collectionCode="NHMD" pageId="14" pageNumber="80">NHMD 621836</specimenCode>
</materialsCitation>
•
<materialsCitation id="3B6B3CF4FFA7C973FADBFC9A89CA4EC9" ID-GBIF-Occurrence="3948948357" collectingDate="2017-12-27" collectingMethod="pitfall trap" collectionCode="R, T" collectorName="R. Lowassary &amp; T. Pape &amp; N. Scharff" country="Tanzania" elevation="1643" latitude="-7.816867" location="Udzungwa Mountains Natl Park" longLatPrecision="1" longitude="36.840015" municipality="Udzungwa Mountains" pageId="14" pageNumber="80" specimenCode="NHMD 621837" specimenCount="1" specimenCount-male="1" stateProvince="Morogoro Region" typeStatus="paratype">
<specimenCount id="9D05FD20FFA7C973FADBFC9A8D5D4E51" box="[1345,1393,836,863]" pageId="14" pageNumber="80" type="male">1 ♂</specimenCount>
;
<collectingRegion id="49C7F84BFFA7C973FF27FCB689B94E8F" box="[189,405,871,898]" country="Tanzania" name="Morogoro" pageId="14" pageNumber="80">Morogoro Region</collectingRegion>
,
<location id="8EDC6072FFA7C973FE39FCB98B044E8F" LSID="urn:lsid:plazi:treatment:03AA87BFFFA2C973FDC6F92F8CB44ACC:8EDC6072FFA7C973FE39FCB98B044E8F" box="[419,808,871,897]" country="Tanzania" latitude="-7.816867" longLatPrecision="1" longitude="36.840015" municipality="Udzungwa Mountains" name="Udzungwa Mountains Natl Park" pageId="14" pageNumber="80" stateProvince="Morogoro Region">Udzungwa Mountains Natl Park</location>
,
<locationDeviation id="9A0E2F8BFFA7C973FCACFCB98C364E8F" box="[822,1050,871,897]" pageId="14" pageNumber="80">above Mito Mitatu</locationDeviation>
;
<geoCoordinate id="EE37506EFFA7C973FBB2FCB88CEF4E8F" box="[1064,1219,870,898]" degrees="07" direction="south" minutes="49.012" orientation="latitude" pageId="14" pageNumber="80" precision="1" value="-7.816867">07°49.012′ S</geoCoordinate>
,
<geoCoordinate id="EE37506EFFA7C973FB49FCB88D5D4E8C" box="[1235,1393,870,898]" degrees="36" direction="east" minutes="50.401" orientation="longitude" pageId="14" pageNumber="80" precision="1" value="36.840015">36°50.401′ E</geoCoordinate>
;
<elevation id="002ED19AFFA7C973FF27FC5489794EAA" box="[189,341,906,933]" metricMagnitude="3" metricUnit="m" metricValue="1.643" pageId="14" pageNumber="80" unit="m" value="1643.0">
<quantity id="4CFB9B4CFFA7C973FF27FC54893A4EAB" box="[189,278,906,933]" metricMagnitude="3" metricUnit="m" metricValue="1.643" pageId="14" pageNumber="80" unit="m" value="1643.0">1643 m</quantity>
a.s.l.
</elevation>
;
<date id="FFBD1069FFA7C973FEF9FC548A2F4EAA" box="[355,515,906,933]" pageId="14" pageNumber="80" value="2017-12-27">
<collectingDate id="EFF9E981FFA7C973FEF9FC548A2F4EAA" box="[355,515,906,933]" pageId="14" pageNumber="80" value="2017-12-27">27 Dec. 2017</collectingDate>
</date>
;
<collectorName id="26F6537FFFA7C973FD95FC558A9D4EAB" box="[527,689,907,933]" pageId="14" pageNumber="80">R. Lowassary</collectorName>
,
<collectorName id="26F6537FFFA7C973FD26FC558B3F4EAB" box="[700,787,907,933]" pageId="14" pageNumber="80">T. Pape</collectorName>
and
<collectorName id="26F6537FFFA7C973FCD6FC558BE94EAA" box="[844,965,906,933]" pageId="14" pageNumber="80">N. Scharff</collectorName>
leg.;
<collectingMethod id="52424EBEFFA7C973FB9DFC548CAE4EAB" box="[1031,1154,906,933]" pageId="14" pageNumber="80">pitfall trap</collectingMethod>
nr. 5, Acc. No.2017- EN-002;
<specimenCode id="DBA59ED2FFA7C973FEB1FC7089CA4EC9" box="[299,486,941,968]" collectionCode="NHMD" pageId="14" pageNumber="80">NHMD 621837</specimenCode>
</materialsCitation>
•
<materialsCitation id="3B6B3CF4FFA7C973FD98FC7089374900" ID-GBIF-Occurrence="3948948307" collectedFrom="litter in montane rain forest" collectingDate="1928-09-29" collectionCode="NHMD" collectorName="N. Scharff" country="Tanzania" elevation="1825" location="Mwanihana Forest Reserve" municipality="Uzungwa Mountains" pageId="14" pageNumber="80" specimenCode="NHMD 621838" specimenCount="2" specimenCount-female="1" specimenCount-subadult="1" stateProvince="Morogoro Region" typeStatus="paratype">
<specimenCount id="9D05FD20FFA7C973FD98FC708A514EC9" box="[514,637,941,968]" pageId="14" pageNumber="80" type="subadult">1 subadult</specimenCount>
<specimenCount id="9D05FD20FFA7C973FD1CFC708AB14EC6" box="[646,669,942,968]" pageId="14" pageNumber="80" type="female">♀</specimenCount>
;
<collectingRegion id="49C7F84BFFA7C973FD37FC708BA94EC9" box="[685,901,941,968]" country="Tanzania" name="Morogoro" pageId="14" pageNumber="80">Morogoro Region</collectingRegion>
,
<collectingMunicipality id="6BD8ACD3FFA7C973FC08FC708CA14EC9" box="[914,1165,941,968]" pageId="14" pageNumber="80">Uzungwa Mountains</collectingMunicipality>
,
<location id="8EDC6072FFA7C973FB00FC7389334EE5" LSID="urn:lsid:plazi:treatment:03AA87BFFFA2C973FDC6F92F8CB44ACC:8EDC6072FFA7C973FB00FC7389334EE5" country="Tanzania" municipality="Uzungwa Mountains" name="Mwanihana Forest Reserve" pageId="14" pageNumber="80" stateProvince="Morogoro Region">Mwanihana Forest Reserve</location>
;
<elevation id="002ED19AFFA7C973FEB7FC0F8A3E4EE5" box="[301,530,977,1004]" metricMagnitude="3" metricUnit="m" metricValue="1.825" metricValueMax="1.85" metricValueMin="1.8" pageId="14" pageNumber="80" unit="m" value="1825.0" valueMax="1850.0" valueMin="1800.0">
<quantity id="4CFB9B4CFFA7C973FEB7FC0F89FD4EE5" box="[301,465,977,1004]" metricMagnitude="3" metricUnit="m" metricValue="1.825" metricValueMax="1.85" metricValueMin="1.8" pageId="14" pageNumber="80" unit="m" value="1825.0" valueMax="1850.0" valueMin="1800.0">1800–1850 m</quantity>
a.s.l.
</elevation>
;
<date id="FFBD1069FFA7C973FDBBFC0F8A864EE5" box="[545,682,977,1003]" pageId="14" pageNumber="80" value="1928-09-29">
<collectingDate id="EFF9E981FFA7C973FDBBFC0F8A864EE5" box="[545,682,977,1003]" pageId="14" pageNumber="80" value="1928-09-29">28–29 Sep.</collectingDate>
</date>
, 1984;
<collectorName id="26F6537FFFA7C973FC99FC0F8B534EE5" box="[771,895,977,1003]" pageId="14" pageNumber="80">N. Scharff</collectorName>
leg.;
<collectedFrom id="381F7554FFA7C973FC5EFC0F8D3C4EE5" box="[964,1296,977,1003]" pageId="14" pageNumber="80">litter in montane rain forest</collectedFrom>
;
<specimenCode id="DBA59ED2FFA7C973FA87FC0F89374900" collectionCode="NHMD" pageId="14" pageNumber="80">NHMD 621838</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BBC36A9FFA7C973FF70FBEB8A48495E" blockId="14.[189,1399,1077,1175]" box="[234,612,1077,1104]" pageId="14" pageNumber="80">
<emphasis id="B977EABBFFA7C973FF70FBEB8A48495E" bold="true" box="[234,612,1077,1104]" pageId="14" pageNumber="80">
Referred specimen, not a
<typeStatus id="54B8880BFFA7C973FDB4FBE88A48495E" box="[558,612,1078,1104]" pageId="14" pageNumber="80">type</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BBC36A9FFA7C973FF27FB878D714998" blockId="14.[189,1399,1077,1175]" pageId="14" pageNumber="80">
<materialsCitation id="3B6B3CF4FFA7C973FF27FB878D754998" ID-GBIF-Occurrence="3948948314" collectedFrom="montane forest, litter" collectingDate="1984-11-08" collectingDateMax="1984-11-13" collectingDateMin="1984-11-08" collectionCode="NHMD" collectorName="N. Scharff" country="Tanzania" county="Udzungwa Mountains" elevation="1625" location="Udzungwa Scarp Forest Reserve" municipality="Udzungwa Scarp Forest Reserve" pageId="14" pageNumber="80" specimenCode="NHMD 621839" specimenCount="1" specimenCount-female="1" stateProvince="Iringa Region">
<collectingCountry id="F3147639FFA7C973FF27FB87897D497D" box="[189,337,1113,1139]" name="Tanzania" pageId="14" pageNumber="80">TANZANIA</collectingCountry>
•
<specimenCount id="9D05FD20FFA7C973FEFDFB8789BF497D" box="[359,403,1113,1139]" pageId="14" pageNumber="80" type="female">1 ♀</specimenCount>
;
<collectingCounty id="62DD4E25FFA7C973FE3BFB878A8B497D" box="[417,679,1113,1139]" pageId="14" pageNumber="80">Udzungwa Mountains</collectingCounty>
,
<collectingRegion id="49C7F84BFFA7C973FD2BFB878B7A497D" box="[689,854,1113,1139]" country="Tanzania" name="Iringa" pageId="14" pageNumber="80">Iringa Region</collectingRegion>
,
<collectingMunicipality id="6BD8ACD3FFA7C973FCFAFB878CF3497D" box="[864,1247,1113,1139]" pageId="14" pageNumber="80">Udzungwa Scarp Forest Reserve</collectingMunicipality>
,
<locationDeviation id="9A0E2F8BFFA7C973FB71FB8789394998" pageId="14" pageNumber="80">above Chita Village</locationDeviation>
;
<elevation id="002ED19AFFA7C973FEBBFBA28A2F4998" box="[289,515,1148,1175]" metricMagnitude="3" metricUnit="m" metricValue="1.625" metricValueMax="1.65" metricValueMin="1.6" pageId="14" pageNumber="80" unit="m" value="1625.0" valueMax="1650.0" valueMin="1600.0">
<quantity id="4CFB9B4CFFA7C973FEBBFBA289EF4998" box="[289,451,1148,1175]" metricMagnitude="3" metricUnit="m" metricValue="1.625" metricValueMax="1.65" metricValueMin="1.6" pageId="14" pageNumber="80" unit="m" value="1625.0" valueMax="1650.0" valueMin="1600.0">1600–1650 m</quantity>
a.s.l.
</elevation>
;
<date id="FFBD1069FFA7C973FD8BFBA28AFC4998" box="[529,720,1148,1174]" pageId="14" pageNumber="80" value="1984-11-08" valueMax="1984-11-13" valueMin="1984-11-08">
<collectingDate id="EFF9E981FFA7C973FD8BFBA28AFC4998" box="[529,720,1148,1174]" pageId="14" pageNumber="80" value="1984-11-08" valueMax="1984-11-13" valueMin="1984-11-08">8–13 Nov. 1984</collectingDate>
</date>
;
<collectorName id="26F6537FFFA7C973FD46FBA28B7A4998" box="[732,854,1148,1174]" pageId="14" pageNumber="80">N. Scharff</collectorName>
leg.;
<collectedFrom id="381F7554FFA7C973FC02FBA28CBC4998" box="[920,1168,1148,1174]" pageId="14" pageNumber="80">montane forest, litter</collectedFrom>
;
<specimenCode id="DBA59ED2FFA7C973FB06FBA28D754998" box="[1180,1369,1148,1174]" collectionCode="NHMD" pageId="14" pageNumber="80">NHMD 621839</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3196522FFA7C973FF27FB638A914B30" pageId="14" pageNumber="80" type="description">
<paragraph id="8BBC36A9FFA7C973FF27FB63896349D9" blockId="14.[189,335,1213,1239]" box="[189,335,1213,1239]" pageId="14" pageNumber="80">
<heading id="D0F481C5FFA7C973FF27FB63896349D9" bold="true" box="[189,335,1213,1239]" fontSize="11" level="3" pageId="14" pageNumber="80" reason="3">
<emphasis id="B977EABBFFA7C973FF27FB63896349D9" bold="true" box="[189,335,1213,1239]" pageId="14" pageNumber="80">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BBC36A9FFA7C973FF27FB328A614842" blockId="14.[189,1399,1260,1357]" pageId="14" pageNumber="80">
SIZE. Males: Length
<quantity id="4CFB9B4CFFA7C973FE2AFB328A6E4808" box="[432,578,1260,1287]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" metricValueMax="5.5" metricValueMin="5.3" pageId="14" pageNumber="80" unit="mm" value="5.4" valueMax="5.5" valueMin="5.3">5.3–5.5 mm</quantity>
; maximal body width (across metazona)
<quantity id="4CFB9B4CFFA7C973FBB2FB328CFB4808" box="[1064,1239,1260,1286]" metricMagnitude="-4" metricUnit="m" metricValue="9.15" metricValueMax="9.2" metricValueMin="9.1" pageId="14" pageNumber="80" unit="mm" value="0.915" valueMax="0.92" valueMin="0.91">0.91–0.92 mm</quantity>
; 17 podous + 1 apodous ring + telson. Female: Length
<quantity id="4CFB9B4CFFA7C973FD3BFACE8AD34824" box="[673,767,1296,1322]" metricMagnitude="-3" metricUnit="m" metricValue="7.7" pageId="14" pageNumber="80" unit="mm" value="7.7">7.7 mm</quantity>
; maximal body width (across metazona)
<quantity id="4CFB9B4CFFA7C973FB78FAD18D624824" box="[1250,1358,1295,1322]" metricMagnitude="-3" metricUnit="m" metricValue="1.19" pageId="14" pageNumber="80" unit="mm" value="1.19">1.19 mm</quantity>
; 18 podous + 1 apodous ring + telson.
</paragraph>
<paragraph id="8BBC36A9FFA7C973FF27FAAA8A914B30" blockId="14.[188,1399,1396,1598]" pageId="14" pageNumber="80">
OTHER CHARACTERS. Virtually identical to the smaller
<taxonomicName id="4C034D2AFFA7C973FCCEFAAA8C3B4883" authority="Enghoff, 2022" authorityName="Enghoff" authorityYear="2022" box="[852,1047,1396,1422]" class="Diplopoda" family="Pyrgodesmidae" genus="Cryptocorypha" kingdom="Animalia" order="Polydesmida" pageId="14" pageNumber="80" phylum="Arthropoda" rank="species" species="geminiramus" status="sp. nov.">
<emphasis id="B977EABBFFA7C973FCCEFAAA8C3B4883" box="[852,1047,1396,1422]" italics="true" pageId="14" pageNumber="80">C. geminiramus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24457C0FFA7C973FBBEFAAA8CA94880" box="[1060,1157,1396,1422]" pageId="14" pageNumber="80" rank="species">sp. nov.</taxonomicNameLabel>
with the following exceptions: All level 1 tubercles and all lateral paratergal lobes on podous and apodous rings with a club-shaped, ornamented club-shaped, somewhat cactus-like seta (
<figureCitation id="13382A2CFFA7C973FC58FA648C3248DA" box="[962,1054,1466,1492]" captionStart="Fig" captionStartId="12.[189,232,1928,1954]" captionTargetBox="[189,1398,268,1886]" captionTargetId="figure-11@12.[189,1398,268,1886]" captionTargetPageId="12" captionText="Fig. 8. Cryptocorypha cactifer sp. nov., paratype (NHMD 621836), ♂, from Mito Mitatu. A. Head and rings 1–6, lateral view. B. Rings 12–18 and telson, dorsal view. C. Tergal seta. D. Head and collum, frontal view. Scale bars: A–B, D = 0.1 mm; C = 0.005 mm." figureDoi="http://doi.org/10.5281/zenodo.7258814" httpUri="https://zenodo.org/record/7258814/files/figure.png" pageId="14" pageNumber="80">Fig. 8C</figureCitation>
); smooth part of head capsule extending well above antennal sockets (
<figureCitation id="13382A2CFFA7C973FD3BFA038B2E48F9" box="[673,770,1501,1527]" captionStart="Fig" captionStartId="12.[189,232,1928,1954]" captionTargetBox="[189,1398,268,1886]" captionTargetId="figure-11@12.[189,1398,268,1886]" captionTargetPageId="12" captionText="Fig. 8. Cryptocorypha cactifer sp. nov., paratype (NHMD 621836), ♂, from Mito Mitatu. A. Head and rings 1–6, lateral view. B. Rings 12–18 and telson, dorsal view. C. Tergal seta. D. Head and collum, frontal view. Scale bars: A–B, D = 0.1 mm; C = 0.005 mm." figureDoi="http://doi.org/10.5281/zenodo.7258814" httpUri="https://zenodo.org/record/7258814/files/figure.png" pageId="14" pageNumber="80">Fig. 8D</figureCitation>
); gonopod coxa (
<emphasis id="B977EABBFFA7C973FC40FA008BD848F9" box="[986,1012,1502,1527]" italics="true" pageId="14" pageNumber="80">cx</emphasis>
) more regularly hemisphaerical (
<figureCitation id="13382A2CFFA7C973FF5CF9DE896F4B15" box="[198,323,1536,1563]" captionStart="Fig" captionStartId="13.[188,231,1892,1918]" captionTargetBox="[189,1398,383,1853]" captionTargetId="figure-48@13.[189,1398,383,1853]" captionTargetPageId="13" captionText="Fig. 9. Cryptocorypha cactifer sp. nov., paratype (NHMD 621836), ♂, from Mito Mitatu, left gonopod. A. Lateral view. B. Basal part of telopodite, showing jagged?velum. C. (Sub)ventral view. D. Posterior view. E. Anterior view. Abbreviations: cx = coxa; ex = exomere; slm = solenomere; ve = velum? Scale bars: A, C–E = 0.01 mm; B = 0.005 mm." figureDoi="http://doi.org/10.5281/zenodo.7258818" httpUri="https://zenodo.org/record/7258818/files/figure.png" pageId="14" pageNumber="80">Fig. 9A, C</figureCitation>
); basal part of gonopod telopodite with a jagged semicircular ridge (velum?,
<emphasis id="B977EABBFFA7C973FB20F9DF8CF84B14" box="[1210,1236,1537,1562]" italics="true" pageId="14" pageNumber="80">ve</emphasis>
), on the basal part of the gonopod telopodite (
<figureCitation id="13382A2CFFA7C973FDB5F9FA8A9C4B30" box="[559,688,1572,1598]" captionStart="Fig" captionStartId="13.[188,231,1892,1918]" captionTargetBox="[189,1398,383,1853]" captionTargetId="figure-48@13.[189,1398,383,1853]" captionTargetPageId="13" captionText="Fig. 9. Cryptocorypha cactifer sp. nov., paratype (NHMD 621836), ♂, from Mito Mitatu, left gonopod. A. Lateral view. B. Basal part of telopodite, showing jagged?velum. C. (Sub)ventral view. D. Posterior view. E. Anterior view. Abbreviations: cx = coxa; ex = exomere; slm = solenomere; ve = velum? Scale bars: A, C–E = 0.01 mm; B = 0.005 mm." figureDoi="http://doi.org/10.5281/zenodo.7258818" httpUri="https://zenodo.org/record/7258818/files/figure.png" pageId="14" pageNumber="80">Fig. 9B, D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3196522FFA7C973FF27F9BB8CB44ACC" pageId="14" pageNumber="80" type="discussion">
<paragraph id="8BBC36A9FFA7C973FF27F9BB891C4B71" blockId="14.[189,304,1637,1663]" box="[189,304,1637,1663]" pageId="14" pageNumber="80">
<heading id="D0F481C5FFA7C973FF27F9BB891C4B71" bold="true" box="[189,304,1637,1663]" fontSize="11" level="3" pageId="14" pageNumber="80" reason="3">
<emphasis id="B977EABBFFA7C973FF27F9BB891C4B71" bold="true" box="[189,304,1637,1663]" pageId="14" pageNumber="80">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BBC36A9FFA7C973FF27F94A8D544A19" blockId="14.[188,1400,1684,1816]" pageId="14" pageNumber="80">
Almost all specimens of
<taxonomicName id="4C034D2AFFA7C973FE47F94A8A764BA0" authority="Enghoff, 2022" authorityName="Enghoff" authorityYear="2022" box="[477,602,1684,1710]" class="Diplopoda" family="Pyrgodesmidae" genus="Cryptocorypha" kingdom="Animalia" order="Polydesmida" pageId="14" pageNumber="80" phylum="Arthropoda" rank="species" species="cactifer" status="sp. nov.">
<emphasis id="B977EABBFFA7C973FE47F94A8A764BA0" box="[477,602,1684,1710]" italics="true" pageId="14" pageNumber="80">C. cactifer</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24457C0FFA7C973FDFBF94A8A974BA0" box="[609,699,1684,1710]" pageId="14" pageNumber="80" rank="species">sp. nov.</taxonomicNameLabel>
were found in mixed samples with
<taxonomicName id="4C034D2AFFA7C973FBC3F94A8D3A4BA3" authority="Enghoff, 2022" authorityName="Enghoff" authorityYear="2022" box="[1113,1302,1684,1710]" class="Diplopoda" family="Pyrgodesmidae" genus="Cryptocorypha" kingdom="Animalia" order="Polydesmida" pageId="14" pageNumber="80" phylum="Arthropoda" rank="species" species="geminiramus" status="sp. nov.">
<emphasis id="B977EABBFFA7C973FBC3F94A8D3A4BA3" box="[1113,1302,1684,1710]" italics="true" pageId="14" pageNumber="80">C. geminiramus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24457C0FFA7C973FA87F94A8D5B4BA0" box="[1309,1399,1684,1710]" pageId="14" pageNumber="80" rank="species">sp. nov.</taxonomicNameLabel>
The single female from Udzungwa Scarp Forest Reserve fully matches the females from Udzungwa Moutains National Park, but in the light of the considerable geographical distance, combined with the absence of a male, the specimen from Udzungwa Scarp Forest Reserve is not included in the
<typeStatus id="54B8880BFFA7C973FB6FF9208D044A16" box="[1269,1320,1790,1816]" pageId="14" pageNumber="80">type</typeStatus>
series.
</paragraph>
<paragraph id="8BBC36A9FFA7C973FF27F8E18CB44ACC" blockId="14.[189,1400,1855,1987]" pageId="14" pageNumber="80">
If the Afrotropical alternative in the first couplet of the key provided by
<bibRefCitation id="EF924B58FFA7C973FBBFF8E18CDB4A57" author="Golovatch S. I." box="[1061,1271,1855,1881]" pageId="14" pageNumber="80" pagination="179 - 190" refId="ref14226" refString="Golovatch S. I. 2019. The millipede genus Cryptocorypha Attems, 1907 revisited, with descriptions of two new Oriental species. Arthropoda Selecta 28: 179 - 190." type="journal article" year="2019">Golovatch (2019)</bibRefCitation>
is chosen,
<taxonomicName id="4C034D2AFFA7C973FF27F8BC89114A72" authority="Enghoff, 2022" authorityName="Enghoff" authorityYear="2022" box="[189,317,1890,1916]" class="Diplopoda" family="Pyrgodesmidae" genus="Cryptocorypha" kingdom="Animalia" order="Polydesmida" pageId="14" pageNumber="80" phylum="Arthropoda" rank="species" species="cactifer" status="sp. nov.">
<emphasis id="B977EABBFFA7C973FF27F8BC89114A72" box="[189,317,1890,1916]" italics="true" pageId="14" pageNumber="80">C. cactifer</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24457C0FFA7C973FEDCF8BC898F4A72" box="[326,419,1890,1916]" pageId="14" pageNumber="80" rank="species">sp. nov.</taxonomicNameLabel>
runs to
<taxonomicName id="4C034D2AFFA7C973FD90F8BC8B454A72" authority="(Brolemann, 1920)" baseAuthorityName="Brolemann" baseAuthorityYear="1920" box="[522,873,1890,1916]" class="Diplopoda" family="Pyrgodesmidae" genus="Cryptocorypha" kingdom="Animalia" order="Polydesmida" pageId="14" pageNumber="80" phylum="Arthropoda" rank="species" species="diffusa">
<emphasis id="B977EABBFFA7C973FD90F8BC8A524A72" box="[522,638,1890,1916]" italics="true" pageId="14" pageNumber="80">C. diffusa</emphasis>
(Brolemann, 1920)
</taxonomicName>
from which it differs by several characters, especially by strongly different gonopods and hairy tergites. If the Asian alternative in the first couplet is chosen,
<taxonomicName id="4C034D2AFFA7C973FEA2F877899A4ACC" authority="Enghoff, 2022" authorityName="Enghoff" authorityYear="2022" box="[312,438,1960,1987]" class="Diplopoda" family="Pyrgodesmidae" genus="Cryptocorypha" kingdom="Animalia" order="Polydesmida" pageId="14" pageNumber="80" phylum="Arthropoda" rank="species" species="cactifer" status="sp. nov.">
<emphasis id="B977EABBFFA7C973FEA2F877899A4ACC" box="[312,438,1960,1987]" italics="true" pageId="14" pageNumber="80">C. cactifer</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24457C0FFA7C973FE27F8778A3B4ACD" box="[445,535,1961,1987]" pageId="14" pageNumber="80" rank="species">sp. nov.</taxonomicNameLabel>
runs to couplet 17(18) where it fits neither alternative.
</paragraph>
</subSubSection>
</treatment>
</document>
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