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<document id="9DBB5043A3AF18F095A4EB3E068296BC" ID-CLB-Dataset="21918" ID-DOI="10.11646/zootaxa.4780.1.1" ID-GBIF-Dataset="af0a7e60-adc6-4896-8544-63725501fa13" ID-ISSN="1175-5326" ID-Zenodo-Dep="3856493" ID-ZooBank="D6CC1683-8BF0-4ABF-ABFE-3EC63E66AE5C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1590138264658" checkinUser="plazi" docAuthor="Suárez, William" docDate="2020" docId="039EF96AFFE3224DED83FBB6FE81FCA0" docLanguage="en" docName="zootaxa.4780.1.1.pdf" docOrigin="Zootaxa 4780 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Gigantohierax itchei Suárez 2020, sp. nov." docType="treatment" docVersion="7" lastPageNumber="27" masterDocId="FFA78112FFFB2257ED14FFF7FFB9FFB8" masterDocTitle="The fossil avifauna of the tar seeps Las Breas de San Felipe, Matanzas, Cuba" masterLastPageNumber="53" masterPageNumber="1" pageNumber="25" updateTime="1698840883311" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="D444236F633691BF9558AFCB92507079">The fossil avifauna of the tar seeps Las Breas de San Felipe, Matanzas, Cuba</mods:title>
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<mods:namePart id="8E988408BEFAE736120F3D94B153872D">Suárez, William</mods:namePart>
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<mods:date id="2029830999462E9210072F1A9CCB1975">2020</mods:date>
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<mods:number id="65051DF0F1B62F6DB5D5427E98A941B3">2020-05-22</mods:number>
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<treatment id="039EF96AFFE3224DED83FBB6FE81FCA0" ID-DOI="http://doi.org/10.5281/zenodo.3856810" ID-GBIF-Taxon="164248776" ID-Zenodo-Dep="3856810" LSID="urn:lsid:plazi:treatment:039EF96AFFE3224DED83FBB6FE81FCA0" httpUri="http://treatment.plazi.org/id/039EF96AFFE3224DED83FBB6FE81FCA0" lastPageId="26" lastPageNumber="27" pageId="24" pageNumber="25">
<subSubSection id="C32D1BF7FFE3224FED83FBB6FE5DFBC7" pageId="24" pageNumber="25" type="nomenclature">
<paragraph id="8B88487CFFE3224FED83FBB6FDBCFBE3" blockId="24.[151,517,1089,1188]" box="[151,517,1089,1116]" pageId="24" pageNumber="25">
<heading id="D0C0FF10FFE3224FED83FBB6FDBCFBE3" bold="true" box="[151,517,1089,1116]" fontSize="11" level="1" pageId="24" pageNumber="25" reason="1">
<emphasis id="B943946EFFE3224FED83FBB6FDBCFBE3" bold="true" box="[151,517,1089,1116]" pageId="24" pageNumber="25">
<taxonomicName id="4C3733FFFFE3224FEDB2FBB6FE19FBE4" authority="Suárez, 2020" authorityName="Suárez" authorityYear="2020" box="[166,416,1089,1116]" class="Aves" family="Accipitridae" genus="Gigantohierax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Accipitriformes" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="itchei" status="sp. nov.">
<emphasis id="B943946EFFE3224FEDB2FBB6FE19FBE4" bold="true" box="[166,416,1089,1116]" italics="true" pageId="24" pageNumber="25">Gigantohierax itchei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2702915FFE3224FECB3FBB6FDBCFBE3" box="[423,517,1089,1115]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B88487CFFE3224FED83FB92FE5DFBC7" blockId="24.[151,517,1089,1188]" box="[151,484,1124,1151]" pageId="24" pageNumber="25">Itches Eagle; Águila de Itche</paragraph>
</subSubSection>
<subSubSection id="C32D1BF7FFE3224FED83FB7EFE17FB1C" box="[151,430,1161,1188]" pageId="24" pageNumber="25" type="description">
<paragraph id="8B88487CFFE3224FED83FB7EFE17FB1C" blockId="24.[151,517,1089,1188]" box="[151,430,1161,1188]" pageId="24" pageNumber="25">
(
<figureCitation id="130C54F9FFE3224FED8BFB7EFEB6FB1B" box="[159,271,1161,1187]" captionStart="FIGURE 11" captionStartId="22.[151,250,1826,1851]" captionTargetBox="[342,1245,189,1800]" captionTargetId="figure@22.[337,1251,181,1801]" captionTargetPageId="22" captionText="FIGURE 11. Gigantohierax suarezi: Left humerus (MNHNCu 75.4722) in anterior/cranial (A) and posterior/caudal (B) views; left carpometacarpus (MNHNCu 75.4723) in dorsal (C) view; right tarsometatarsus (MNHNCu 75.4728) in distal (D), anterior (E), medial (F) and posterior (G) views; phalanx I, left digit I, in dorsal (H) view; ungual phalanx, left digit I (MNHNCu 75.4731) in lateral (I) view. Gigantohierax itchei sp. nov.: Right femur (Paratype, MNHNCu 75.4725) in anterior (J), posterior (K) and lateral (L) views; right tarsometatarsus (Holotype, MNHNCu 75.4869) in anterior (M), medial (N), posterior (O) and distal (P) views. Scale = 2 cm." figureDoi="http://doi.org/10.5281/zenodo.3856513" httpUri="https://zenodo.org/record/3856513/files/figure.png" pageId="24" pageNumber="25">Figure 11</figureCitation>
: JP;
<tableCitation id="C6B57DC7FFE3224FEC45FB7EFE1CFB1C" box="[337,421,1161,1188]" captionStart="TABLE 6" captionStartId="25.[151,239,658,682]" captionText="TABLE 6. Measurements (mm) in the tarsometatarsus of Gigantohierax itchei sp. nov., compared to those of other antil-" pageId="24" pageNumber="25">Table 6</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C32D1BF7FFE3224FED83FB26FE16FA8B" pageId="24" pageNumber="25" type="materials_examined">
<paragraph id="8B88487CFFE3224FED83FB26FE16FA8B" blockId="24.[151,1436,1232,2015]" pageId="24" pageNumber="25">
<materialsCitation id="3B5F4221FFE3224FED83FB26FE16FA8B" ID-GBIF-Occurrence="2622603311" collectionCode="IV, II" county="San Felipe" location="Distal" municipality="Paleontologia" pageId="24" pageNumber="25" specimenCount="1" typeStatus="holotype">
<emphasis id="B943946EFFE3224FED83FB26FEB2FB53" bold="true" box="[151,267,1233,1259]" pageId="24" pageNumber="25">
<typeStatus id="548CF6DEFFE3224FED83FB26FEBFFB53" box="[151,262,1233,1259]" pageId="24" pageNumber="25" type="holotype">Holotype</typeStatus>
.
</emphasis>
<location id="8EE81EA7FFE3224FEC02FB26FEE0FB53" LSID="urn:lsid:plazi:treatment:039EF96AFFE3224DED83FBB6FE81FCA0:8EE81EA7FFE3224FEC02FB26FEE0FB53" box="[278,345,1233,1259]" county="San Felipe" municipality="Paleontologia" name="Distal" pageId="24" pageNumber="25">Distal</location>
third of right tarsometatarsus without trochlea metatarsi
<collectionCode id="ED26D0B9FFE3224FEEE6FB26FBB6FB53" box="[1010,1039,1233,1259]" pageId="24" pageNumber="25">IV</collectionCode>
, MNHNCu 75.4869. Collected by members of the
<location id="8EE81EA7FFE3224FEC59FB02FDC2FAB7" LSID="urn:lsid:plazi:treatment:039EF96AFFE3224DED83FBB6FE81FCA0:8EE81EA7FFE3224FEC59FB02FDC2FAB7" box="[333,635,1269,1295]" county="San Felipe" municipality="Paleontologia" name="Departamento de Geologia" pageId="24" pageNumber="25">Departamento de Geología</location>
y
<collectingMunicipality id="6BECD206FFE3224FEF83FB02FC8DFAB7" box="[663,820,1269,1295]" pageId="24" pageNumber="25">Paleontología</collectingMunicipality>
, MNHNCu, in
<collectingCounty id="62E930F0FFE3224FEEF4FB02FBEEFAB7" box="[992,1111,1269,1295]" pageId="24" pageNumber="25">San Felipe</collectingCounty>
<collectionCode id="ED26D0B9FFE3224FE94AFB02FBCAFAB7" box="[1118,1139,1269,1295]" pageId="24" pageNumber="25">II</collectionCode>
, during field expedition to the type locality in 1998.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32D1BF7FFE3224FEDD3FACAFC69F997" pageId="24" pageNumber="25" type="description">
<paragraph id="8B88487CFFE3224FEDD3FACAFD16FA18" blockId="24.[151,1436,1232,2015]" pageId="24" pageNumber="25">
<emphasis id="B943946EFFE3224FEDD3FACAFDEEFAEF" bold="true" box="[199,599,1340,1367]" pageId="24" pageNumber="25">
Measurements (mm) of
<typeStatus id="548CF6DEFFE3224FECFDFACAFDEBFAEF" box="[489,594,1341,1367]" pageId="24" pageNumber="25" type="holotype">holotype</typeStatus>
.
</emphasis>
Total length as preserved: 69.5; least width and depth of shaft: 12.09.0; distal width from the wing of trochlea metatarsi II to the outer rim of trochlea metatarsi III: 23.5; width and depth of trochlea metatarsi III: 8.410.1 (see
<tableCitation id="C6B57DC7FFE3224FEF58FA72FD19FA18" box="[588,672,1413,1440]" captionStart="TABLE 6" captionStartId="25.[151,239,658,682]" captionText="TABLE 6. Measurements (mm) in the tarsometatarsus of Gigantohierax itchei sp. nov., compared to those of other antil-" pageId="24" pageNumber="25">Table 6</tableCitation>
).
</paragraph>
<paragraph id="8B88487CFFE3224FEDD3FA5EFA9EFA7B" blockId="24.[151,1436,1232,2015]" box="[199,1319,1448,1475]" pageId="24" pageNumber="25">
<materialsCitation id="3B5F4221FFE3224FEDD3FA5EFA9AFA7B" ID-GBIF-Occurrence="2622603302" box="[199,1315,1448,1475]" collectionCode="II" pageId="24" pageNumber="25" specimenCount="1" typeStatus="paratype">
<emphasis id="B943946EFFE3224FEDD3FA5EFE04FA7B" bold="true" box="[199,445,1448,1475]" pageId="24" pageNumber="25">
<typeStatus id="548CF6DEFFE3224FEDD3FA5EFE8DFA7B" box="[199,308,1449,1475]" pageId="24" pageNumber="25" type="paratype">Paratype</typeStatus>
(topotype).
</emphasis>
San Felipe
<collectionCode id="ED26D0B9FFE3224FEF56FA5EFDE1FA7B" box="[578,600,1449,1475]" pageId="24" pageNumber="25">II</collectionCode>
: Proximal end of fragmentary right femur (MNHNCu 75.4725)
</materialsCitation>
.
</paragraph>
<paragraph id="8B88487CFFE3224FEDD3FA3AFC69F997" blockId="24.[151,1436,1232,2015]" pageId="24" pageNumber="25">
<emphasis id="B943946EFFE3224FEDD3FA3AFDE3FA5F" bold="true" box="[199,602,1484,1511]" pageId="24" pageNumber="25">
Measurements (mm) of
<typeStatus id="548CF6DEFFE3224FECFCFA3AFDECFA5F" box="[488,597,1485,1511]" pageId="24" pageNumber="25" type="paratype">paratype</typeStatus>
.
</emphasis>
Femur (
<taxonomicName id="4C3733FFFFE3224FEFA9FA3AFC88FA5F" box="[701,817,1485,1511]" class="Aves" family="Accipitridae" genus="Gigantohierax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Accipitriformes" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="suarezi">
<emphasis id="B943946EFFE3224FEFA9FA3AFC88FA5F" box="[701,817,1485,1511]" italics="true" pageId="24" pageNumber="25">G. suarezi</emphasis>
</taxonomicName>
in parentheses): depth of neck: 13.7 (19.0); width and depth at level of the distal end of the trochanteric ridge: 18.5 (23.9)12.2 (16.2); length from distal point of the pneumatic foramen to the distal end of the trochanteric ridge: 19.5 (15.5).
</paragraph>
</subSubSection>
<subSubSection id="C32D1BF7FFE3224FEDD3F9CEFCCBF9CF" pageId="24" pageNumber="25" type="etymology">
<paragraph id="8B88487CFFE3224FEDD3F9CEFCCBF9CF" blockId="24.[151,1436,1232,2015]" pageId="24" pageNumber="25">
<emphasis id="B943946EFFE3224FEDD3F9CEFEF7F9EB" bold="true" box="[199,334,1593,1619]" pageId="24" pageNumber="25">Etymology.</emphasis>
After my dear friend Irving “Itche” Himel, Toronto,
<collectingCountry id="F32008ECFFE3224FEE8BF9CEFC4FF9EB" box="[927,1014,1593,1619]" name="Canada" pageId="24" pageNumber="25">Canada</collectingCountry>
, for his sincere friendship and unconditional support during my paleontological studies in his country.
</paragraph>
</subSubSection>
<subSubSection id="C32D1BF7FFE3224FEDD3F977FCD7F923" box="[199,878,1664,1691]" pageId="24" pageNumber="25" type="diagnosis">
<paragraph id="8B88487CFFE3224FEDD3F977FCD7F923" blockId="24.[151,1436,1232,2015]" box="[199,878,1664,1691]" pageId="24" pageNumber="25">
<emphasis id="B943946EFFE3224FEDD3F977FEFBF923" bold="true" box="[199,322,1664,1691]" pageId="24" pageNumber="25">Diagnosis.</emphasis>
The smallest species in the genus
<taxonomicName id="4C3733FFFFE3224FEFD7F976FCDFF923" authorityName="Arredondo &amp; Arredondo" authorityYear="1999" box="[707,870,1665,1691]" class="Aves" family="Accipitridae" genus="Gigantohierax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Accipitriformes" pageId="24" pageNumber="25" phylum="Chordata" rank="genus">
<emphasis id="B943946EFFE3224FEFD7F976FCDFF923" box="[707,870,1665,1691]" italics="true" pageId="24" pageNumber="25">Gigantohierax</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C32D1BF7FFE3224DEDD3F953FE85FE33" lastPageId="26" lastPageNumber="27" pageId="24" pageNumber="25" type="description">
<paragraph id="8B88487CFFE3224EEDD3F953FCC8FDDB" blockId="24.[151,1436,1232,2015]" lastBlockId="25.[151,1437,153,611]" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25">
<emphasis id="B943946EFFE3224FEDD3F953FEE0F907" bold="true" box="[199,345,1700,1727]" pageId="24" pageNumber="25">Description.</emphasis>
Specimens of
<taxonomicName id="4C3733FFFFE3224FECEAF952FD5FF907" authority="Suárez, 2020" authorityName="Suárez" authorityYear="2020" box="[510,742,1701,1727]" class="Aves" family="Accipitridae" genus="Gigantohierax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Accipitriformes" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="itchei" status="sp. nov.">
<emphasis id="B943946EFFE3224FECEAF952FD5FF907" box="[510,742,1701,1727]" italics="true" pageId="24" pageNumber="25">Gigantohierax itchei</emphasis>
</taxonomicName>
<emphasis id="B943946EFFE3224FEFF9F952FCFFF907" bold="true" box="[749,838,1701,1727]" pageId="24" pageNumber="25">
<taxonomicNameLabel id="A2702915FFE3224FEFF9F952FCFFF907" box="[749,838,1701,1727]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
are ~29% smaller than the equivalent elements in the skeleton of
<taxonomicName id="4C3733FFFFE3224FEC0FF93EFE37F95B" box="[283,398,1737,1763]" class="Aves" family="Accipitridae" genus="Gigantohierax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Accipitriformes" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="suarezi">
<emphasis id="B943946EFFE3224FEC0FF93EFE96F95B" box="[283,303,1737,1763]" italics="true" pageId="24" pageNumber="25">G</emphasis>
.
<emphasis id="B943946EFFE3224FEC2AF93DFE37F95B" box="[318,398,1738,1763]" italics="true" pageId="24" pageNumber="25">suarezi</emphasis>
</taxonomicName>
(mean 17.63%, range 9.2726.84% in between-species of large
<taxonomicName id="4C3733FFFFE3224FE971F93EFB4BF95B" authorityName="Vieillot" authorityYear="1816" box="[1125,1266,1737,1763]" class="Aves" family="Accipitridae" kingdom="Animalia" order="Accipitriformes" pageId="24" pageNumber="25" phylum="Chordata" rank="family">Accipitridae</taxonomicName>
, see
<bibRefCitation id="EFA6358DFFE3224FE83DF93EFF6AF8BF" author="Holdaway, R. N." pageId="24" pageNumber="25" pagination="39 - 47" refId="ref27120" refString="Holdaway, R. N. (1990) Harpagornis assimilis Haast, 1874, a synonym of Harpagornis moorei Haast, 1872 (Aves: Accipitridae). New Zealand Natural Sciences, 17, 39 - 47." type="journal article" year="1990">Holdaway 1990</bibRefCitation>
). The femur (
<figureCitation id="130C54F9FFE3224FEC7FF91AFE0EF8BF" box="[363,439,1773,1799]" captionStart="FIGURE 11" captionStartId="22.[151,250,1826,1851]" captionTargetBox="[342,1245,189,1800]" captionTargetId="figure@22.[337,1251,181,1801]" captionTargetPageId="22" captionText="FIGURE 11. Gigantohierax suarezi: Left humerus (MNHNCu 75.4722) in anterior/cranial (A) and posterior/caudal (B) views; left carpometacarpus (MNHNCu 75.4723) in dorsal (C) view; right tarsometatarsus (MNHNCu 75.4728) in distal (D), anterior (E), medial (F) and posterior (G) views; phalanx I, left digit I, in dorsal (H) view; ungual phalanx, left digit I (MNHNCu 75.4731) in lateral (I) view. Gigantohierax itchei sp. nov.: Right femur (Paratype, MNHNCu 75.4725) in anterior (J), posterior (K) and lateral (L) views; right tarsometatarsus (Holotype, MNHNCu 75.4869) in anterior (M), medial (N), posterior (O) and distal (P) views. Scale = 2 cm." figureDoi="http://doi.org/10.5281/zenodo.3856513" httpUri="https://zenodo.org/record/3856513/files/figure.png" pageId="24" pageNumber="25">Fig. 11</figureCitation>
JL) in anterior view has a large pneumatic depression (= proximal pneumatic foramen), located proximomediad to the distal pneumatic foramen (similar, greater pneumaticity and presence of perforations in the basal wall by some very small foramina in
<taxonomicName id="4C3733FFFFE3224FEE08F8C2FC28F8F7" box="[796,913,1845,1871]" class="Aves" family="Accipitridae" genus="Gigantohierax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Accipitriformes" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="suarezi">
<emphasis id="B943946EFFE3224FEE08F8C2FC89F8F7" box="[796,816,1845,1871]" italics="true" pageId="24" pageNumber="25">G</emphasis>
.
<emphasis id="B943946EFFE3224FEE55F8C1FC28F8F7" box="[833,913,1846,1871]" italics="true" pageId="24" pageNumber="25">suarezi</emphasis>
</taxonomicName>
); small distal pneumatic foramen, apparently semitriangular, according to the preserved portions (larger, semitriangular in
<taxonomicName id="4C3733FFFFE3224FEEE8F8AEFBD6F8CB" box="[1020,1135,1881,1907]" class="Aves" family="Accipitridae" genus="Gigantohierax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Accipitriformes" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="suarezi">
<emphasis id="B943946EFFE3224FEEE8F8AEFBA9F8CB" box="[1020,1040,1881,1907]" italics="true" pageId="24" pageNumber="25">G</emphasis>
.
<emphasis id="B943946EFFE3224FE90BF8ADFBD6F8CB" box="[1055,1135,1882,1907]" italics="true" pageId="24" pageNumber="25">suarezi</emphasis>
</taxonomicName>
); the distance between the distal margin of the distal pneumatic foramen and the distal end of the trochanteric ridge is very large (smaller in
<taxonomicName id="4C3733FFFFE3224FED83F856FEB2F803" box="[151,267,1953,1979]" class="Aves" family="Accipitridae" genus="Gigantohierax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Accipitriformes" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="suarezi">
<emphasis id="B943946EFFE3224FED83F856FF12F803" box="[151,171,1953,1979]" italics="true" pageId="24" pageNumber="25">G</emphasis>
.
<emphasis id="B943946EFFE3224FEDAFF855FEB2F803" box="[187,267,1954,1979]" italics="true" pageId="24" pageNumber="25">suarezi</emphasis>
</taxonomicName>
). In posterior view, the proximal surface is irregular, being inflated (or bulky) in its center, as result of the high development of the obturator ridge (flattened and pneumatic surface, with little or no development of the obturator ridge in
<taxonomicName id="4C3733FFFFE2224EEC76FF6EFE6CFF0B" box="[354,469,153,179]" class="Aves" family="Accipitridae" genus="Gigantohierax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Accipitriformes" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="suarezi">
<emphasis id="B943946EFFE2224EEC76FF6EFECFFF0B" box="[354,374,153,179]" italics="true" pageId="25" pageNumber="26">G</emphasis>
.
<emphasis id="B943946EFFE2224EEC91FF6DFE6CFF0B" box="[389,469,154,179]" italics="true" pageId="25" pageNumber="26">suarezi</emphasis>
</taxonomicName>
). The tarsometatarsus (
<figureCitation id="130C54F9FFE2224EEFC8FF6EFC91FF0B" box="[732,808,153,179]" captionStart="FIGURE 11" captionStartId="22.[151,250,1826,1851]" captionTargetBox="[342,1245,189,1800]" captionTargetId="figure@22.[337,1251,181,1801]" captionTargetPageId="22" captionText="FIGURE 11. Gigantohierax suarezi: Left humerus (MNHNCu 75.4722) in anterior/cranial (A) and posterior/caudal (B) views; left carpometacarpus (MNHNCu 75.4723) in dorsal (C) view; right tarsometatarsus (MNHNCu 75.4728) in distal (D), anterior (E), medial (F) and posterior (G) views; phalanx I, left digit I, in dorsal (H) view; ungual phalanx, left digit I (MNHNCu 75.4731) in lateral (I) view. Gigantohierax itchei sp. nov.: Right femur (Paratype, MNHNCu 75.4725) in anterior (J), posterior (K) and lateral (L) views; right tarsometatarsus (Holotype, MNHNCu 75.4869) in anterior (M), medial (N), posterior (O) and distal (P) views. Scale = 2 cm." figureDoi="http://doi.org/10.5281/zenodo.3856513" httpUri="https://zenodo.org/record/3856513/files/figure.png" pageId="25" pageNumber="26">Fig. 11</figureCitation>
MP) has a shaft proportionally thinner (anterior and/or posterior view), less flattened or compressed (lateral and/or medial view) than in the largest species (more flattened and compressed, being wide in
<taxonomicName id="4C3733FFFFE2224EECE8FF16FDD7FF43" box="[508,622,225,251]" class="Aves" family="Accipitridae" genus="Gigantohierax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Accipitriformes" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="suarezi">
<emphasis id="B943946EFFE2224EECE8FF16FDA9FF43" box="[508,528,225,251]" italics="true" pageId="25" pageNumber="26">G</emphasis>
.
<emphasis id="B943946EFFE2224EEF0AFF15FDD7FF43" box="[542,622,226,251]" italics="true" pageId="25" pageNumber="26">suarezi</emphasis>
</taxonomicName>
). In anterior view, it is deeply excavated by a distinctive, very deep and narrow anterior metatarsal groove, which extends well distad (groove much less excavated or extended distad, with anterior surface of the shaft more flattened or convex in
<taxonomicName id="4C3733FFFFE2224EEE0BFEDEFC2AFEFB" box="[799,915,297,323]" class="Aves" family="Accipitridae" genus="Gigantohierax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Accipitriformes" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="suarezi">
<emphasis id="B943946EFFE2224EEE0BFEDEFC8AFEFB" box="[799,819,297,323]" italics="true" pageId="25" pageNumber="26">G</emphasis>
.
<emphasis id="B943946EFFE2224EEE57FEDDFC2AFEFB" box="[835,915,298,323]" italics="true" pageId="25" pageNumber="26">suarezi</emphasis>
</taxonomicName>
). The distal foramen is large (proportionately smaller in
<taxonomicName id="4C3733FFFFE2224EEC1BFEBAFE3BFEDF" box="[271,386,333,359]" class="Aves" family="Accipitridae" genus="Gigantohierax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Accipitriformes" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="suarezi">
<emphasis id="B943946EFFE2224EEC1BFEBAFE9AFEDF" box="[271,291,333,359]" italics="true" pageId="25" pageNumber="26">G</emphasis>
.
<emphasis id="B943946EFFE2224EEC26FEB9FE3BFEDF" box="[306,386,334,359]" italics="true" pageId="25" pageNumber="26">suarezi</emphasis>
</taxonomicName>
), trochlea metatarsi III with very wide and grooved articular surface (relatively narrow in
<taxonomicName id="4C3733FFFFE2224EE895FEBAFF5EFE33" class="Aves" family="Accipitridae" genus="Gigantohierax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Accipitriformes" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="suarezi">
<emphasis id="B943946EFFE2224EE895FEBAFA2CFEDF" box="[1409,1429,333,359]" italics="true" pageId="25" pageNumber="26">G</emphasis>
.
<emphasis id="B943946EFFE2224EED83FE85FF5EFE33" box="[151,231,370,395]" italics="true" pageId="25" pageNumber="26">suarezi</emphasis>
</taxonomicName>
). In posterior view, the posterior metatarsal groove is slightly deeper, but less extended distally (superficial, ending more distad in
<taxonomicName id="4C3733FFFFE2224EEC89FE62FDABFE17" box="[413,530,405,431]" class="Aves" family="Accipitridae" genus="Gigantohierax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Accipitriformes" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="suarezi">
<emphasis id="B943946EFFE2224EEC89FE62FE08FE17" box="[413,433,405,431]" italics="true" pageId="25" pageNumber="26">G</emphasis>
.
<emphasis id="B943946EFFE2224EECD6FE61FDABFE17" box="[450,530,406,431]" italics="true" pageId="25" pageNumber="26">suarezi</emphasis>
</taxonomicName>
); crista plantaris medialis better defined (rounded, less defined in
<taxonomicName id="4C3733FFFFE2224EE802FE62FA32FE17" box="[1302,1419,405,431]" class="Aves" family="Accipitridae" genus="Gigantohierax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Accipitriformes" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="suarezi">
<emphasis id="B943946EFFE2224EE802FE62FA93FE17" box="[1302,1322,405,431]" italics="true" pageId="25" pageNumber="26">G</emphasis>
.
<emphasis id="B943946EFFE2224EE82FFE61FA32FE17" box="[1339,1419,406,431]" italics="true" pageId="25" pageNumber="26">suarezi</emphasis>
</taxonomicName>
); larger distal foramen (relatively smaller in
<taxonomicName id="4C3733FFFFE2224EEF61FE4EFD5EFE6B" box="[629,743,441,467]" class="Aves" family="Accipitridae" genus="Gigantohierax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Accipitriformes" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="suarezi">
<emphasis id="B943946EFFE2224EEF61FE4EFD30FE6B" box="[629,649,441,467]" italics="true" pageId="25" pageNumber="26">G</emphasis>
.
<emphasis id="B943946EFFE2224EEF83FE4DFD5EFE6B" box="[663,743,442,467]" italics="true" pageId="25" pageNumber="26">suarezi</emphasis>
</taxonomicName>
); fossa supratrochlearis plantaris shallow (deep and expanded in
<taxonomicName id="4C3733FFFFE2224EEDA0FE2AFE9FFE4F" box="[180,294,477,503]" class="Aves" family="Accipitridae" genus="Gigantohierax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Accipitriformes" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="suarezi">
<emphasis id="B943946EFFE2224EEDA0FE2AFF71FE4F" box="[180,200,477,503]" italics="true" pageId="25" pageNumber="26">G</emphasis>
.
<emphasis id="B943946EFFE2224EEDC2FE29FE9FFE4F" box="[214,294,478,503]" italics="true" pageId="25" pageNumber="26">suarezi</emphasis>
</taxonomicName>
); broadly furrowed trochlea metatarsi III (narrow groove in
<taxonomicName id="4C3733FFFFE2224EEEDCFE2AFB83FE4F" box="[968,1082,477,503]" class="Aves" family="Accipitridae" genus="Gigantohierax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Accipitriformes" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="suarezi">
<emphasis id="B943946EFFE2224EEEDCFE2AFC65FE4F" box="[968,988,477,503]" italics="true" pageId="25" pageNumber="26">G</emphasis>
.
<emphasis id="B943946EFFE2224EEEFEFE29FB83FE4F" box="[1002,1082,478,503]" italics="true" pageId="25" pageNumber="26">suarezi</emphasis>
</taxonomicName>
) with very narrow and laterally oriented inner ring (much wider and mediad in
<taxonomicName id="4C3733FFFFE2224EEF8BFDF6FCA9FDA3" box="[671,784,513,539]" class="Aves" family="Accipitridae" genus="Gigantohierax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Accipitriformes" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="suarezi">
<emphasis id="B943946EFFE2224EEF8BFDF6FD0AFDA3" box="[671,691,513,539]" italics="true" pageId="25" pageNumber="26">G</emphasis>
.
<emphasis id="B943946EFFE2224EEFD4FDF5FCA9FDA3" box="[704,784,514,539]" italics="true" pageId="25" pageNumber="26">suarezi</emphasis>
</taxonomicName>
); internal side of this last trochlea very excavated medially (less excavated, more superficial in
<taxonomicName id="4C3733FFFFE2224EEF3EFDD2FD25FD87" box="[554,668,549,575]" class="Aves" family="Accipitridae" genus="Gigantohierax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Accipitriformes" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="suarezi">
<emphasis id="B943946EFFE2224EEF3EFDD2FD87FD87" box="[554,574,549,575]" italics="true" pageId="25" pageNumber="26">G</emphasis>
.
<emphasis id="B943946EFFE2224EEF58FDD1FD25FD87" box="[588,668,550,575]" italics="true" pageId="25" pageNumber="26">suarezi</emphasis>
</taxonomicName>
). In cross section, the shaft is squarer, furrowed by the deep anterior metatarsal groove (more triangular, less excavated in
<taxonomicName id="4C3733FFFFE2224EEFFAFDBEFCD9FDDB" box="[750,864,585,611]" class="Aves" family="Accipitridae" genus="Gigantohierax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Accipitriformes" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="suarezi">
<emphasis id="B943946EFFE2224EEFFAFDBEFCBBFDDB" box="[750,770,585,611]" italics="true" pageId="25" pageNumber="26">G</emphasis>
.
<emphasis id="B943946EFFE2224EEE04FDBDFCD9FDDB" box="[784,864,586,611]" italics="true" pageId="25" pageNumber="26">suarezi</emphasis>
</taxonomicName>
).
</paragraph>
<caption id="DF4818F4FFE2224EED83FD65FA25FD13" box="[151,1436,658,683]" pageId="25" pageNumber="26">
<paragraph id="8B88487CFFE2224EED83FD65FA25FD13" blockId="25.[151,1436,658,755]" box="[151,1436,658,683]" pageId="25" pageNumber="26">
<emphasis id="B943946EFFE2224EED83FD65FEB0FD13" bold="true" box="[151,265,658,683]" pageId="25" pageNumber="26">TABLE 6.</emphasis>
Measurements (mm) in the tarsometatarsus of
<taxonomicName id="4C3733FFFFE2224EEFE9FD64FC63FD13" authority="Suárez, 2020" authorityName="Suárez" authorityYear="2020" box="[765,986,659,683]" class="Aves" family="Accipitridae" genus="Gigantohierax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Accipitriformes" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="itchei" status="sp. nov.">
<emphasis id="B943946EFFE2224EEFE9FD64FC63FD13" box="[765,986,659,683]" italics="true" pageId="25" pageNumber="26">Gigantohierax itchei</emphasis>
</taxonomicName>
<emphasis id="B943946EFFE2224EEEF4FD64FB8CFD13" bold="true" box="[992,1077,659,683]" pageId="25" pageNumber="26">
<taxonomicNameLabel id="A2702915FFE2224EEEF4FD64FB8CFD13" box="[992,1077,659,683]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, compared to those of other antil-
</paragraph>
</caption>
<tableNote id="76D149F2FFE2224EED83FD40FD4EFD4B" pageId="25" pageNumber="26" targetBox="[163,1423,772,1343]" targetPageId="25">
<paragraph id="8B88487CFFE2224EED83FD40FA25FD77" blockId="25.[151,1436,658,755]" box="[151,1436,695,719]" pageId="25" pageNumber="26">
lean extinct accipitrids in the genera
<taxonomicName id="4C3733FFFFE2224EEF36FD40FCACFD76" authority="Wetmore" authorityName="Wetmore" authorityYear="1937" box="[546,789,695,719]" class="Aves" family="Accipitridae" genus="Titanohierax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Accipitriformes" pageId="25" pageNumber="26" phylum="Chordata" rank="genus">
<emphasis id="B943946EFFE2224EEF36FD40FD13FD77" box="[546,682,695,719]" italics="true" pageId="25" pageNumber="26">Titanohierax</emphasis>
Wetmore
</taxonomicName>
, “
<taxonomicName id="4C3733FFFFE2224EEE38FD40FC1EFD77" authority="Campbell" authorityName="Campbell" box="[812,935,695,719]" class="Aves" family="Accipitridae" genus="Amplibuteo" higherTaxonomySource="GBIF" kingdom="Animalia" order="Accipitriformes" pageId="25" pageNumber="26" phylum="Chordata" rank="genus">
<emphasis id="B943946EFFE2224EEE38FD40FC1EFD77" box="[812,935,695,719]" italics="true" pageId="25" pageNumber="26">Amplibuteo</emphasis>
</taxonomicName>
” Campbell,
<taxonomicName id="4C3733FFFFE2224EE93BFD40FABBFD76" authority="Lesson" authorityName="Lesson" authorityYear="1830" box="[1071,1282,695,719]" class="Aves" family="Accipitridae" genus="Buteogallus" kingdom="Animalia" order="Accipitriformes" pageId="25" pageNumber="26" phylum="Chordata" rank="genus">
<emphasis id="B943946EFFE2224EE93BFD40FB17FD77" box="[1071,1198,695,719]" italics="true" pageId="25" pageNumber="26">Buteogallus</emphasis>
Lesson
</taxonomicName>
and the living
</paragraph>
<paragraph id="8B88487CFFE2224EED83FD2CFD4EFD4B" blockId="25.[151,1436,658,755]" box="[151,759,731,755]" pageId="25" pageNumber="26">
<taxonomicName id="4C3733FFFFE2224EED83FD2CFEDBFD4B" authority="(Linnaeus)" baseAuthorityName="Linnaeus" box="[151,354,731,755]" class="Aves" family="Accipitridae" genus="Harpia" kingdom="Animalia" order="Accipitriformes" pageId="25" pageNumber="26" phylum="Chordata" rank="genus">
<emphasis id="B943946EFFE2224EED83FD2CFF5DFD4A" box="[151,228,731,754]" italics="true" pageId="25" pageNumber="26">Harpia</emphasis>
(Linnaeus)
</taxonomicName>
. Sequence: range (mean) sample size.
</paragraph>
</tableNote>
<paragraph id="8B88487CFFE2224EEDB7FCE1FA37FA95" pageId="25" pageNumber="26">
<table id="F937BADCFFE2DDA8EDB7FCF3FA36FA87" box="[163,1423,772,1343]" gridcols="7" gridrows="7" pageId="25" pageNumber="26">
<tr id="35074A3EFFE2DDA8EDB7FCF3FA36FC85" box="[163,1423,772,829]" gridrow="0" pageId="25" pageNumber="26">
<th id="76D62342FFE2DDA8EDB7FCF3FEC4FC85" box="[163,381,772,829]" gridcol="0" gridrow="0" pageId="25" pageNumber="26">CHARACTER</th>
<th id="76D62342FFE2DDA8EC8FFCF3FE54FC85" box="[411,493,772,829]" gridcol="1" gridrow="0" pageId="25" pageNumber="26">
<taxonomicName id="4C3733FFFFE2224EEC8FFCF3FE54FCA1" authority="Suárez, 2020" authorityName="Suárez" authorityYear="2020" box="[411,493,772,793]" class="Aves" family="Accipitridae" genus="Gigantohierax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Accipitriformes" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="itchei" status="sp. nov.">
<emphasis id="B943946EFFE2224EEC8FFCF3FE54FCA1" box="[411,493,772,793]" italics="true" pageId="25" pageNumber="26">G. itchei</emphasis>
</taxonomicName>
<emphasis id="B943946EFFE2224EEC8FFCDFFE5EFC85" bold="true" box="[411,487,808,829]" pageId="25" pageNumber="26">
<taxonomicNameLabel id="A2702915FFE2224EEC8FFCDFFE5EFC85" box="[411,487,808,829]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</th>
<th id="76D62342FFE2DDA8EF13FCF3FDCBFC85" box="[519,626,772,829]" gridcol="2" gridrow="0" pageId="25" pageNumber="26">
<taxonomicName id="4C3733FFFFE2224EEF13FCE1FDD1FC93" box="[519,616,790,811]" class="Aves" family="Accipitridae" genus="Gigantohierax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Accipitriformes" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="suarezi">
<emphasis id="B943946EFFE2224EEF13FCE1FDD1FC93" box="[519,616,790,811]" italics="true" pageId="25" pageNumber="26">G. suarezi</emphasis>
</taxonomicName>
</th>
<th id="76D62342FFE2DDA8EF83FCF3FC94FC85" box="[663,813,772,829]" gridcol="3" gridrow="0" pageId="25" pageNumber="26">
<taxonomicName id="4C3733FFFFE2224EEF83FCE1FD46FC93" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[663,767,790,811]" class="Aves" family="Accipitridae" genus="Harpia" kingdom="Animalia" order="Accipitriformes" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="harpyja">
<emphasis id="B943946EFFE2224EEF83FCE1FD46FC93" box="[663,767,790,811]" italics="true" pageId="25" pageNumber="26">H. harpyja</emphasis>
</taxonomicName>
</th>
<th id="76D62342FFE2DDA8EE5EFCF3FC6FFC85" box="[842,982,772,829]" gridcol="4" gridrow="0" pageId="25" pageNumber="26">
<taxonomicName id="4C3733FFFFE2224EEE5EFCE1FC6FFC93" authorityName="Wetmore" authorityYear="1937" box="[842,982,790,811]" class="Aves" family="Accipitridae" genus="Titanohierax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Accipitriformes" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="gloveralleni">
<emphasis id="B943946EFFE2224EEE5EFCE1FC6FFC93" box="[842,982,790,811]" italics="true" pageId="25" pageNumber="26">T. gloveralleni</emphasis>
</taxonomicName>
</th>
<th id="76D62342FFE2DDA8EEE4FCF3FB0EFC85" box="[1008,1207,772,829]" gridcol="5" gridrow="0" pageId="25" pageNumber="26">
<emphasis id="B943946EFFE2224EEEEFFCE1FBB3FC93" box="[1019,1034,790,811]" italics="true" pageId="25" pageNumber="26">A</emphasis>
.”
<emphasis id="B943946EFFE2224EE934FCE1FB32FC93" box="[1056,1163,790,811]" italics="true" pageId="25" pageNumber="26">woodwardi</emphasis>
</th>
<th id="76D62342FFE2DDA8E9C6FCF3FA36FC85" box="[1234,1423,772,829]" gridcol="6" gridrow="0" pageId="25" pageNumber="26">
<taxonomicName id="4C3733FFFFE2224EE9C6FCE1FA8DFC93" box="[1234,1332,790,811]" class="Aves" family="Accipitridae" genus="Buteogallus" kingdom="Animalia" order="Accipitriformes" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="borrasi">
<emphasis id="B943946EFFE2224EE9C6FCE1FA8DFC93" box="[1234,1332,790,811]" italics="true" pageId="25" pageNumber="26">B. borrasi</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="35074A3EFFE2DDA8EDB7FCAEFA36FCD6" box="[163,1423,857,878]" gridrow="1" pageId="25" pageNumber="26">
<th id="76D62342FFE2DDA8EDB7FCAEFA36FCD6" box="[163,1423,857,878]" colspan="7" colspanRight="6" gridcol="0" gridrow="1" pageId="25" pageNumber="26">TARSOMETATARSUS</th>
</tr>
<tr id="35074A3EFFE2DDA8EDB7FC7EFA36FC52" box="[163,1423,905,1002]" gridrow="2" pageId="25" pageNumber="26">
<th id="76D62342FFE2DDA8EDB7FC7EFEC4FC52" box="[163,381,905,1002]" gridcol="0" gridrow="2" pageId="25" pageNumber="26">Width above proxi- mal end of metatarsal facet</th>
<td id="76D62342FFE2DDA8EC8FFC7EFE54FC52" box="[411,493,905,1002]" gridcol="1" gridrow="2" pageId="25" pageNumber="26">13</td>
<td id="76D62342FFE2DDA8EF13FC7EFDCBFC52" box="[519,626,905,1002]" gridcol="2" gridrow="2" pageId="25" pageNumber="26">19.1 21.1 (20.0) 4</td>
<td id="76D62342FFE2DDA8EF83FC7EFC94FC52" box="[663,813,905,1002]" gridcol="3" gridrow="2" pageId="25" pageNumber="26">12.414.7 (13.4) 4</td>
<td id="76D62342FFE2DDA8EE5EFC7EFC6FFC52" box="[842,982,905,1002]" gridcol="4" gridrow="2" pageId="25" pageNumber="26">13.1</td>
<td id="76D62342FFE2DDA8EEE4FC7EFB0EFC52" box="[1008,1207,905,1002]" gridcol="5" gridrow="2" pageId="25" pageNumber="26">10.112.4 (11.0) 20</td>
<td id="76D62342FFE2DDA8E9C6FC7EFA36FC52" box="[1234,1423,905,1002]" gridcol="6" gridrow="2" pageId="25" pageNumber="26">10.411.4 (10.8) 7</td>
</tr>
<tr id="35074A3EFFE2DDA8EDB7FC0EFA36FBE2" box="[163,1423,1017,1114]" gridrow="3" pageId="25" pageNumber="26">
<th id="76D62342FFE2DDA8EDB7FC0EFEC4FBE2" box="[163,381,1017,1114]" gridcol="0" gridrow="3" pageId="25" pageNumber="26">Depth above proxi- mal end of metatarsal facet</th>
<td id="76D62342FFE2DDA8EC8FFC0EFE54FBE2" box="[411,493,1017,1114]" gridcol="1" gridrow="3" pageId="25" pageNumber="26">8.8</td>
<td id="76D62342FFE2DDA8EF13FC0EFDCBFBE2" box="[519,626,1017,1114]" gridcol="2" gridrow="3" pageId="25" pageNumber="26">11.913.7 (12.8) 5</td>
<td id="76D62342FFE2DDA8EF83FC0EFC94FBE2" box="[663,813,1017,1114]" gridcol="3" gridrow="3" pageId="25" pageNumber="26">15.017.6 (15.8) 4</td>
<td id="76D62342FFE2DDA8EE5EFC0EFC6FFBE2" box="[842,982,1017,1114]" gridcol="4" gridrow="3" pageId="25" pageNumber="26">9.5</td>
<td id="76D62342FFE2DDA8EEE4FC0EFB0EFBE2" box="[1008,1207,1017,1114]" gridcol="5" gridrow="3" pageId="25" pageNumber="26">7.710.0 (8.7) 28</td>
<td id="76D62342FFE2DDA8E9C6FC0EFA36FBE2" box="[1234,1423,1017,1114]" gridcol="6" gridrow="3" pageId="25" pageNumber="26">7.79.0 (8.5) 7</td>
</tr>
<tr id="35074A3EFFE2DDA8EDB7FB9DFA36FB1E" box="[163,1423,1130,1190]" gridrow="4" pageId="25" pageNumber="26">
<th id="76D62342FFE2DDA8EDB7FB9DFEC4FB1E" box="[163,381,1130,1190]" gridcol="0" gridrow="4" pageId="25" pageNumber="26">Depth trochlea II</th>
<td id="76D62342FFE2DDA8EC8FFB9DFE54FB1E" box="[411,493,1130,1190]" gridcol="1" gridrow="4" pageId="25" pageNumber="26">8.8</td>
<td id="76D62342FFE2DDA8EF13FB9DFDCBFB1E" box="[519,626,1130,1190]" gridcol="2" gridrow="4" pageId="25" pageNumber="26">19.8</td>
<td id="76D62342FFE2DDA8EF83FB9DFC94FB1E" box="[663,813,1130,1190]" gridcol="3" gridrow="4" pageId="25" pageNumber="26">17.019.7 (17.8) 4</td>
<td id="76D62342FFE2DDA8EE5EFB9DFC6FFB1E" box="[842,982,1130,1190]" gridcol="4" gridrow="4" pageId="25" pageNumber="26">-</td>
<td id="76D62342FFE2DDA8EEE4FB9DFB0EFB1E" box="[1008,1207,1130,1190]" gridcol="5" gridrow="4" pageId="25" pageNumber="26">12.515.5(14.0) 27</td>
<td id="76D62342FFE2DDA8E9C6FB9DFA36FB1E" box="[1234,1423,1130,1190]" gridcol="6" gridrow="4" pageId="25" pageNumber="26">12.513.4 (12.9) 4</td>
</tr>
<tr id="35074A3EFFE2DDA8EDB7FB42FA36FB4A" box="[163,1423,1205,1266]" gridrow="5" pageId="25" pageNumber="26">
<th id="76D62342FFE2DDA8EDB7FB42FEC4FB4A" box="[163,381,1205,1266]" gridcol="0" gridrow="5" pageId="25" pageNumber="26">Width trochlea III</th>
<td id="76D62342FFE2DDA8EC8FFB42FE54FB4A" box="[411,493,1205,1266]" gridcol="1" gridrow="5" pageId="25" pageNumber="26">8.4</td>
<td id="76D62342FFE2DDA8EF13FB42FDCBFB4A" box="[519,626,1205,1266]" gridcol="2" gridrow="5" pageId="25" pageNumber="26">11.512.6 (11.9) 3</td>
<td id="76D62342FFE2DDA8EF83FB42FC94FB4A" box="[663,813,1205,1266]" gridcol="3" gridrow="5" pageId="25" pageNumber="26">7.99.4 (8.4) 4</td>
<td id="76D62342FFE2DDA8EE5EFB42FC6FFB4A" box="[842,982,1205,1266]" gridcol="4" gridrow="5" pageId="25" pageNumber="26">7.6</td>
<td id="76D62342FFE2DDA8EEE4FB42FB0EFB4A" box="[1008,1207,1205,1266]" gridcol="5" gridrow="5" pageId="25" pageNumber="26">6.88.4 (7.4) 28</td>
<td id="76D62342FFE2DDA8E9C6FB42FA36FB4A" box="[1234,1423,1205,1266]" gridcol="6" gridrow="5" pageId="25" pageNumber="26">6.47.4 (6.9) 5</td>
</tr>
<tr id="35074A3EFFE2DDA8EDB7FAF5FA36FA87" box="[163,1423,1282,1343]" gridrow="6" pageId="25" pageNumber="26">
<th id="76D62342FFE2DDA8EDB7FAF5FEC4FA87" box="[163,381,1282,1343]" gridcol="0" gridrow="6" pageId="25" pageNumber="26">Depth trochlea III</th>
<td id="76D62342FFE2DDA8EC8FFAF5FE54FA87" box="[411,493,1282,1343]" gridcol="1" gridrow="6" pageId="25" pageNumber="26">10.1</td>
<td id="76D62342FFE2DDA8EF13FAF5FDCBFA87" box="[519,626,1282,1343]" gridcol="2" gridrow="6" pageId="25" pageNumber="26">14.815.7 (15.2) 3</td>
<td id="76D62342FFE2DDA8EF83FAF5FC94FA87" box="[663,813,1282,1343]" gridcol="3" gridrow="6" pageId="25" pageNumber="26">11.813.3 (12.5) 4</td>
<td id="76D62342FFE2DDA8EE5EFAF5FC6FFA87" box="[842,982,1282,1343]" gridcol="4" gridrow="6" pageId="25" pageNumber="26">11</td>
<td id="76D62342FFE2DDA8EEE4FAF5FB0EFA87" box="[1008,1207,1282,1343]" gridcol="5" gridrow="6" pageId="25" pageNumber="26">9.812.1 (10.6) 27</td>
<td id="76D62342FFE2DDA8E9C6FAF5FA36FA87" box="[1234,1423,1282,1343]" gridcol="6" gridrow="6" pageId="25" pageNumber="26">10.212.4 (11.8) 6</td>
</tr>
</table>
</paragraph>
<paragraph id="8B88487CFFE2224DEDD3FA61FE85FE33" blockId="25.[151,1437,1430,2033]" lastBlockId="26.[151,1437,153,792]" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26">
<emphasis id="B943946EFFE2224EEDD3FA61FBF2FA09" bold="true" box="[199,1099,1430,1457]" pageId="25" pageNumber="26">Comparisons with other Antillean and continental related fossil accipitrids.</emphasis>
<taxonomicName id="4C3733FFFFE2224EE946FA60FA82FA09" authority="Suárez, 2020" authorityName="Suárez" authorityYear="2020" box="[1106,1339,1431,1457]" class="Aves" family="Accipitridae" genus="Gigantohierax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Accipitriformes" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="itchei" status="sp. nov.">
<emphasis id="B943946EFFE2224EE946FA60FA82FA09" box="[1106,1339,1431,1457]" italics="true" pageId="25" pageNumber="26">Gigantohierax itchei</emphasis>
</taxonomicName>
<emphasis id="B943946EFFE2224EE856FA60FA25FA09" bold="true" box="[1346,1436,1431,1457]" pageId="25" pageNumber="26">
<taxonomicNameLabel id="A2702915FFE2224EE856FA60FA25FA09" box="[1346,1436,1431,1457]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is a larger species compared with
<taxonomicName id="4C3733FFFFE2224EEF08FA4CFD45FA6D" box="[540,764,1467,1493]" class="Aves" family="Accipitridae" genus="Buteogallus" kingdom="Animalia" order="Accipitriformes" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="borrasi">
<emphasis id="B943946EFFE2224EEF08FA4CFD45FA6D" box="[540,764,1467,1493]" italics="true" pageId="25" pageNumber="26">Buteogallus borrasi</emphasis>
</taxonomicName>
, “
<taxonomicName id="4C3733FFFFE2224EEE0DFA4CFC23FA6D" box="[793,922,1467,1493]" class="Aves" family="Accipitridae" genus="Amplibuteo" higherTaxonomySource="GBIF" kingdom="Animalia" order="Accipitriformes" pageId="25" pageNumber="26" phylum="Chordata" rank="genus">
<emphasis id="B943946EFFE2224EEE0DFA4CFC23FA6D" box="[793,922,1467,1493]" italics="true" pageId="25" pageNumber="26">Amplibuteo</emphasis>
</taxonomicName>
<emphasis id="B943946EFFE2224EEEBBFA4CFB95FA6D" box="[943,1068,1467,1493]" italics="true" pageId="25" pageNumber="26">woodwardi</emphasis>
, or “
<emphasis id="B943946EFFE2224EE97CFA4CFBC0FA6D" box="[1128,1145,1467,1493]" italics="true" pageId="25" pageNumber="26">A</emphasis>
<emphasis id="B943946EFFE2224EE99AFA4CFB54FA6D" box="[1166,1261,1467,1493]" italics="true" pageId="25" pageNumber="26">hibbardi</emphasis>
, being close in size to
<taxonomicName id="4C3733FFFFE2224EEDF8FA28FDB5FA41" authorityName="Wetmore" authorityYear="1937" box="[236,524,1503,1529]" class="Aves" family="Accipitridae" genus="Titanohierax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Accipitriformes" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="gloveralleni">
<emphasis id="B943946EFFE2224EEDF8FA28FDB5FA41" box="[236,524,1503,1529]" italics="true" pageId="25" pageNumber="26">Titanohierax gloveralleni</emphasis>
</taxonomicName>
of
<collectingCountry id="F32008ECFFE2224EEF2CFA28FD72FA41" box="[568,715,1503,1529]" name="Bahamas" pageId="25" pageNumber="26">the Bahamas</collectingCountry>
(
<tableCitation id="C6B57DC7FFE2224EEFC8FA28FC8FFA42" box="[732,822,1503,1530]" captionStart="TABLE 6" captionStartId="25.[151,239,658,682]" captionText="TABLE 6. Measurements (mm) in the tarsometatarsus of Gigantohierax itchei sp. nov., compared to those of other antil-" pageId="25" pageNumber="26">Table 6</tableCitation>
).
<taxonomicName id="4C3733FFFFE2224EEE5AFA28FB80FA41" authority="Suárez, 2020" authorityName="Suárez" authorityYear="2020" box="[846,1081,1503,1529]" class="Aves" family="Accipitridae" genus="Gigantohierax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Accipitriformes" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="itchei" status="sp. nov.">
<emphasis id="B943946EFFE2224EEE5AFA28FB80FA41" box="[846,1081,1503,1529]" italics="true" pageId="25" pageNumber="26">Gigantohierax itchei</emphasis>
</taxonomicName>
<emphasis id="B943946EFFE2224EE957FA28FB19FA41" bold="true" box="[1091,1184,1503,1529]" pageId="25" pageNumber="26">
<taxonomicNameLabel id="A2702915FFE2224EE957FA28FB19FA41" box="[1091,1184,1503,1529]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from all these species by having a wider femur (unknown element in
<taxonomicName id="4C3733FFFFE2224EEE07F9F3FC00F9A5" authorityName="Wetmore" authorityYear="1937" box="[787,953,1539,1565]" class="Aves" family="Accipitridae" genus="Titanohierax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Accipitriformes" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="gloveralleni">
<emphasis id="B943946EFFE2224EEE07F9F3FC9AF9A5" box="[787,803,1540,1565]" italics="true" pageId="25" pageNumber="26">T</emphasis>
.
<emphasis id="B943946EFFE2224EEE26F9F4FC00F9A5" box="[818,953,1539,1565]" italics="true" pageId="25" pageNumber="26">gloveralleni</emphasis>
</taxonomicName>
), flaring greatly (anterior and/or posterior view) proximad; presence of distinctive concavity formed by the most proximal pneumatic foramen; second (more distad) pneumatic foramen small and semitriangular in shape. In posterior view, the proximal surface is flattened and expanded, not rounded. In lateral view, this element is greatly compressed in anteroposterior direction, being almost flat and the trochanteric ridge is narrow and long distad. Tarsometatarsus (
<figureCitation id="130C54F9FFE2224EE91BF964FBE2F915" box="[1039,1115,1683,1709]" captionStart="FIGURE 11" captionStartId="22.[151,250,1826,1851]" captionTargetBox="[342,1245,189,1800]" captionTargetId="figure@22.[337,1251,181,1801]" captionTargetPageId="22" captionText="FIGURE 11. Gigantohierax suarezi: Left humerus (MNHNCu 75.4722) in anterior/cranial (A) and posterior/caudal (B) views; left carpometacarpus (MNHNCu 75.4723) in dorsal (C) view; right tarsometatarsus (MNHNCu 75.4728) in distal (D), anterior (E), medial (F) and posterior (G) views; phalanx I, left digit I, in dorsal (H) view; ungual phalanx, left digit I (MNHNCu 75.4731) in lateral (I) view. Gigantohierax itchei sp. nov.: Right femur (Paratype, MNHNCu 75.4725) in anterior (J), posterior (K) and lateral (L) views; right tarsometatarsus (Holotype, MNHNCu 75.4869) in anterior (M), medial (N), posterior (O) and distal (P) views. Scale = 2 cm." figureDoi="http://doi.org/10.5281/zenodo.3856513" httpUri="https://zenodo.org/record/3856513/files/figure.png" pageId="25" pageNumber="26">Fig. 11</figureCitation>
MP) also more compressed anteroposteriorly, being wide and flat, especially at the distal end; very deep anterior metatarsal groove that extends well down (distad) on shaft; distal foramen large; surface between this last foramen and the external border very wide and rounded (very thin and acute in
<taxonomicName id="4C3733FFFFE2224EEF76F8F7FCBFF8A1" authorityName="Wetmore" authorityYear="1937" box="[610,774,1791,1817]" class="Aves" family="Accipitridae" genus="Titanohierax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Accipitriformes" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="gloveralleni">
<emphasis id="B943946EFFE2224EEF76F8F7FDCBF8A1" box="[610,626,1792,1817]" italics="true" pageId="25" pageNumber="26">T</emphasis>
.
<emphasis id="B943946EFFE2224EEF6BF908FCBFF8A1" box="[639,774,1791,1817]" italics="true" pageId="25" pageNumber="26">gloveralleni</emphasis>
</taxonomicName>
; similar in the other species); trochleae flaring abruptly and distally on shaft; trochlea metatarsi III short, wide, and with a distinct wide groove (longer, thinner, with thin groove in
<taxonomicName id="4C3733FFFFE2224EEDA0F8BFFEE0F8D9" authorityName="Wetmore" authorityYear="1937" box="[180,345,1863,1889]" class="Aves" family="Accipitridae" genus="Titanohierax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Accipitriformes" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="gloveralleni">
<emphasis id="B943946EFFE2224EEDA0F8BFFF7DF8D9" box="[180,196,1864,1889]" italics="true" pageId="25" pageNumber="26">T</emphasis>
.
<emphasis id="B943946EFFE2224EEDC6F8B0FEE0F8D9" box="[210,345,1863,1889]" italics="true" pageId="25" pageNumber="26">gloveralleni</emphasis>
</taxonomicName>
; proportionately similar in the remaining species compared); in lateral view, the lateral border or surface is thinner (by the great anteroposterior compression) and rounded, especially at the distal end (wide, concave because it is deeply grooved, less rounded distally in
<taxonomicName id="4C3733FFFFE2224EEE31F867FC2CF811" box="[805,917,1935,1961]" class="Aves" family="Accipitridae" genus="Buteogallus" kingdom="Animalia" order="Accipitriformes" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="borrasi">
<emphasis id="B943946EFFE2224EEE31F867FC8FF811" box="[805,822,1936,1961]" italics="true" pageId="25" pageNumber="26">B</emphasis>
.
<emphasis id="B943946EFFE2224EEE57F878FC2CF811" box="[835,917,1935,1961]" italics="true" pageId="25" pageNumber="26">borrasi</emphasis>
</taxonomicName>
; very wide and flat even distally, only slightly grooved in
<taxonomicName id="4C3733FFFFE2224EEC0EF843FE79F875" authorityName="Wetmore" authorityYear="1937" box="[282,448,1971,1997]" class="Aves" family="Accipitridae" genus="Titanohierax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Accipitriformes" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="gloveralleni">
<emphasis id="B943946EFFE2224EEC0EF843FE93F875" box="[282,298,1972,1997]" italics="true" pageId="25" pageNumber="26">T</emphasis>
.
<emphasis id="B943946EFFE2224EEC2DF844FE79F875" box="[313,448,1971,1997]" italics="true" pageId="25" pageNumber="26">gloveralleni</emphasis>
</taxonomicName>
; intermediate in characters in “
<emphasis id="B943946EFFE2224EEE33F844FC81F875" box="[807,824,1971,1997]" italics="true" pageId="25" pageNumber="26">A</emphasis>
”.
<emphasis id="B943946EFFE2224EEE40F844FC68F875" box="[852,977,1971,1997]" italics="true" pageId="25" pageNumber="26">woodwardi</emphasis>
and “
<emphasis id="B943946EFFE2224EE903F844FB91F875" box="[1047,1064,1971,1997]" italics="true" pageId="25" pageNumber="26">A</emphasis>
”.
<emphasis id="B943946EFFE2224EE950F844FB1AF875" box="[1092,1187,1971,1997]" italics="true" pageId="25" pageNumber="26">hibbardi</emphasis>
); internal (inner) side forming a thin, acute or sharp border, specially above the level of the metatarsal facet. In posterior view, the poste- rior metatarsal groove is shallow, crista plantaris medialis and crista plantaris lateralis with poor development; metatarsal facet higher (proximal) on shaft; distal foramen ovoid and much larger; trochlea metatarsi III wide, widely grooved, with short articular surface and distinct long neck at base (wide and also widely grooved, long surface without neck in
<taxonomicName id="4C3733FFFFE1224DEC5FFEF1FE05FEA7" box="[331,444,261,287]" class="Aves" family="Accipitridae" genus="Buteogallus" kingdom="Animalia" order="Accipitriformes" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="borrasi">
<emphasis id="B943946EFFE1224DEC5FFEF1FEE5FEA7" box="[331,348,262,287]" italics="true" pageId="26" pageNumber="27">B</emphasis>
.
<emphasis id="B943946EFFE1224DEC7EFEF2FE05FEA7" box="[362,444,261,287]" italics="true" pageId="26" pageNumber="27">borrasi</emphasis>
</taxonomicName>
; much thinner and less grooved, much longer or extended posterior surface without neck in
<taxonomicName id="4C3733FFFFE1224DEDA7FEDDFEEEFEFB" authorityName="Wetmore" authorityYear="1937" box="[179,343,297,323]" class="Aves" family="Accipitridae" genus="Titanohierax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Accipitriformes" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="gloveralleni">
<emphasis id="B943946EFFE1224DEDA7FEDDFF7AFEFB" box="[179,195,298,323]" italics="true" pageId="26" pageNumber="27">T</emphasis>
.
<emphasis id="B943946EFFE1224DEDC4FEDEFEEEFEFB" box="[208,343,297,323]" italics="true" pageId="26" pageNumber="27">gloveralleni</emphasis>
</taxonomicName>
; intermediate in characters in “
<emphasis id="B943946EFFE1224DEFA1FEDEFD7FFEFB" box="[693,710,297,323]" italics="true" pageId="26" pageNumber="27">A</emphasis>
”.
<emphasis id="B943946EFFE1224DEFF4FEDEFCE4FEFB" box="[736,861,297,323]" italics="true" pageId="26" pageNumber="27">woodwardi</emphasis>
and “
<emphasis id="B943946EFFE1224DEE8AFEDEFC16FEFB" box="[926,943,297,323]" italics="true" pageId="26" pageNumber="27">A</emphasis>
”.
<emphasis id="B943946EFFE1224DEEDDFEDEFB91FEFB" box="[969,1064,297,323]" italics="true" pageId="26" pageNumber="27">hibbardi</emphasis>
); in distal view, the later trochlea is wide but not deep, being squared in morphology (much thinner, very deep, rectangular in
<taxonomicName id="4C3733FFFFE1224DE9B1FEB9FAF3FEDF" authorityName="Wetmore" authorityYear="1937" box="[1189,1354,333,359]" class="Aves" family="Accipitridae" genus="Titanohierax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Accipitriformes" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="gloveralleni">
<emphasis id="B943946EFFE1224DE9B1FEB9FB0CFEDF" box="[1189,1205,334,359]" italics="true" pageId="26" pageNumber="27">T</emphasis>
.
<emphasis id="B943946EFFE1224DE9D7FEBAFAF3FEDF" box="[1219,1354,333,359]" italics="true" pageId="26" pageNumber="27">gloveralleni</emphasis>
</taxonomicName>
); outer rim very wide.
</paragraph>
</subSubSection>
<subSubSection id="C32D1BF7FFE1224DEDD3FE63FE81FCA0" pageId="26" pageNumber="27" type="discussion">
<paragraph id="8B88487CFFE1224DEDD3FE63FE81FCA0" blockId="26.[151,1437,153,792]" pageId="26" pageNumber="27">
<emphasis id="B943946EFFE1224DEDD3FE63FEF6FE17" bold="true" box="[199,335,404,431]" pageId="26" pageNumber="27">Comments.</emphasis>
In the initial stages of the process of identification of the specimens described here as
<taxonomicName id="4C3733FFFFE1224DE823FE62FE9CFE6B" authority="Suárez, 2020" authorityName="Suárez" authorityYear="2020" class="Aves" family="Accipitridae" genus="Gigantohierax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Accipitriformes" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="itchei" status="sp. nov.">
<emphasis id="B943946EFFE1224DE823FE62FE9CFE6B" italics="true" pageId="26" pageNumber="27">Gigantohierax itchei</emphasis>
</taxonomicName>
<emphasis id="B943946EFFE1224DEC3AFE4EFE30FE6B" bold="true" box="[302,393,441,467]" pageId="26" pageNumber="27">
<taxonomicNameLabel id="A2702915FFE1224DEC3AFE4EFE30FE6B" box="[302,393,441,467]" pageId="26" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
the author considered as the first hypothesis the logical possibility that the
<typeStatus id="548CF6DEFFE1224DE9FDFE4EFAF2FE6B" box="[1257,1355,441,467]" pageId="26" pageNumber="27" type="holotype">holotype</typeStatus>
of this species, due to its similar size, could represent
<taxonomicName id="4C3733FFFFE1224DEFBEFE2AFC71FE4F" authorityName="Wetmore" authorityYear="1937" box="[682,968,477,503]" class="Aves" family="Accipitridae" genus="Titanohierax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Accipitriformes" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="gloveralleni">
<emphasis id="B943946EFFE1224DEFBEFE2AFC71FE4F" box="[682,968,477,503]" italics="true" pageId="26" pageNumber="27">Titanohierax gloveralleni</emphasis>
</taxonomicName>
, an extinct and large buteonine hawk unknown in
<collectingCountry id="F32008ECFFE1224DEC1EFDF6FEFCFDA3" box="[266,325,513,539]" name="Cuba" pageId="26" pageNumber="27">Cuba</collectingCountry>
so far, and described from the nearby
<collectingCountry id="F32008ECFFE1224DEFEAFDF6FCDCFDA3" box="[766,869,513,539]" name="Bahamas" pageId="26" pageNumber="27">Bahamas</collectingCountry>
(
<bibRefCitation id="EFA6358DFFE1224DEE61FDF6FB99FDA3" author="Wetmore, A." box="[885,1056,513,539]" pageId="26" pageNumber="27" pagination="427 - 441" refId="ref32224" refString="Wetmore, A. (1937) Bird remains from cave deposits on Great Exuma Island in the Bahamas. Bulletin of the Museum of Comparative Zoology, 80, 427 - 441." type="journal article" year="1937">Wetmore 1937</bibRefCitation>
;
<bibRefCitation id="EFA6358DFFE1224DE939FDF6FAF7FDA3" author="Olson, S. L. &amp; Hilgartner, W. B." box="[1069,1358,513,539]" pageId="26" pageNumber="27" pagination="22 - 56" refId="ref29528" refString="Olson, S. L. &amp; Hilgartner, W. B. (1982) Fossil and subfossil birds from the Bahamas. In: Olson, S. L. (Ed.), Fossil vertebrates from the Bahamas, Smithsonian Contributions to Paleobiology, 48, pp. 22 - 56. https: // doi. org / 10.5479 / si. 00810266.48.1" type="journal article" year="1982">Olson &amp; Hilgartner 1982</bibRefCitation>
). After direct comparisons with the type series of the Bahamian taxon, this hypothesis was quickly dismissed.
<taxonomicName id="4C3733FFFFE1224DE819FDD2FA25FD87" authorityName="Wetmore" authorityYear="1937" box="[1293,1436,549,575]" class="Aves" family="Accipitridae" genus="Titanohierax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Accipitriformes" pageId="26" pageNumber="27" phylum="Chordata" rank="genus">
<emphasis id="B943946EFFE1224DE819FDD2FA25FD87" box="[1293,1436,549,575]" italics="true" pageId="26" pageNumber="27">Titanohierax</emphasis>
</taxonomicName>
is closer to the genus
<taxonomicName id="4C3733FFFFE1224DEC87FDBEFD9DFDDB" authorityName="Kaup" authorityYear="1844" box="[403,548,585,611]" class="Aves" family="Accipitridae" genus="Geranoaetus" kingdom="Animalia" order="Accipitriformes" pageId="26" pageNumber="27" phylum="Chordata" rank="genus">
<emphasis id="B943946EFFE1224DEC87FDBEFD9DFDDB" box="[403,548,585,611]" italics="true" pageId="26" pageNumber="27">Geranoaetus</emphasis>
</taxonomicName>
(and
<taxonomicName id="4C3733FFFFE1224DEF7CFDBDFCA6FDDC" authority="Lacepede" authorityName="Lacepede" box="[616,799,585,612]" class="Aves" family="Accipitridae" genus="Buteo" kingdom="Animalia" order="Accipitriformes" pageId="26" pageNumber="27" phylum="Chordata" rank="genus">
<emphasis id="B943946EFFE1224DEF7CFDBDFD10FDDB" box="[616,681,586,611]" italics="true" pageId="26" pageNumber="27">Buteo</emphasis>
Lacépède
</taxonomicName>
) than to
<taxonomicName id="4C3733FFFFE1224DEE9CFDBEFBB7FDDB" authorityName="Lesson" authorityYear="1830" box="[904,1038,585,611]" class="Aves" family="Accipitridae" genus="Buteogallus" kingdom="Animalia" order="Accipitriformes" pageId="26" pageNumber="27" phylum="Chordata" rank="genus">
<emphasis id="B943946EFFE1224DEE9CFDBEFBB7FDDB" box="[904,1038,585,611]" italics="true" pageId="26" pageNumber="27">Buteogallus</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFA6358DFFE1224DE90BFDBEFAFBFDDB" author="Olson, S. L. &amp; Hilgartner, W. B." box="[1055,1346,585,611]" pageId="26" pageNumber="27" pagination="22 - 56" refId="ref29528" refString="Olson, S. L. &amp; Hilgartner, W. B. (1982) Fossil and subfossil birds from the Bahamas. In: Olson, S. L. (Ed.), Fossil vertebrates from the Bahamas, Smithsonian Contributions to Paleobiology, 48, pp. 22 - 56. https: // doi. org / 10.5479 / si. 00810266.48.1" type="journal article" year="1982">Olson &amp; Hilgartner 1982</bibRefCitation>
;
<bibRefCitation id="EFA6358DFFE1224DE844FDBEFE85FD3F" author="Suarez, W. &amp; Olson, S. L." pageId="26" pageNumber="27" pagination="288 - 298" refId="ref31261" refString="Suarez, W. &amp; Olson, S. L. (2007) The Cuban fossil eagle Aquila borrasi Arredondo: A scaled-up version of the Great Black- Hawk Buteogallus urubitinga (Gmelin). Journal of Raptor Research, 41, 288 - 298. https: // doi. org / 10.3356 / 0892 - 1016 (2007) 41 [288: TCFEAB] 2.0. CO; 2" type="journal article" year="2007">Suárez &amp; Olson 2007</bibRefCitation>
), or
<emphasis id="B943946EFFE1224DEC7BFD9AFD92FD3F" box="[367,555,621,647]" italics="true" pageId="26" pageNumber="27">Gigantohierax—</emphasis>
which shares some characters with
<taxonomicName id="4C3733FFFFE1224DEEA8FD9AFBFBFD3F" authorityName="Lesson" authorityYear="1830" box="[956,1090,621,647]" class="Aves" family="Accipitridae" genus="Buteogallus" kingdom="Animalia" order="Accipitriformes" pageId="26" pageNumber="27" phylum="Chordata" rank="genus">
<emphasis id="B943946EFFE1224DEEA8FD9AFBFBFD3F" box="[956,1090,621,647]" italics="true" pageId="26" pageNumber="27">Buteogallus</emphasis>
</taxonomicName>
(see descriptions above)—, as seen in the anatomy of its tarsometatarsus, especially on both sides of the shaft and in the trochleae. After comparisons and explorations on the osteology of both species described in the extinct genus “
<taxonomicName id="4C3733FFFFE1224DE972FD42FB5EFD77" authority="Campbell" authorityName="Campbell" box="[1126,1255,693,719]" class="Aves" family="Accipitridae" genus="Amplibuteo" higherTaxonomySource="GBIF" kingdom="Animalia" order="Accipitriformes" pageId="26" pageNumber="27" phylum="Chordata" rank="genus">
<emphasis id="B943946EFFE1224DE972FD42FB5EFD77" box="[1126,1255,693,719]" italics="true" pageId="26" pageNumber="27">Amplibuteo</emphasis>
</taxonomicName>
” Campbell (see
<bibRefCitation id="EFA6358DFFE1224DED83FD2EFEFEFD4B" author="Campbell, K. E. Jr." box="[151,327,729,755]" pageId="26" pageNumber="27" pagination="1 - 203" refId="ref25456" refString="Campbell, K. E. Jr. (1979) The non-passerine Pleistocene avifauna of the Talara Tar Seeps, northwestern Peru. Royal Ontario Museum Life Sciences Contributions, 118, 1 - 203. https: // doi. org / 10.5962 / bhl. title. 52133" type="journal article" year="1979">Campbell 1979</bibRefCitation>
), it seems that those must be more accurately placed under
<taxonomicName id="4C3733FFFFE1224DEEF5FD2EFBDEFD4B" authorityName="Lesson" authorityYear="1830" box="[993,1127,729,755]" class="Aves" family="Accipitridae" genus="Buteogallus" kingdom="Animalia" order="Accipitriformes" pageId="26" pageNumber="27" phylum="Chordata" rank="genus">
<emphasis id="B943946EFFE1224DEEF5FD2EFBDEFD4B" box="[993,1127,729,755]" italics="true" pageId="26" pageNumber="27">Buteogallus</emphasis>
</taxonomicName>
(see
<bibRefCitation id="EFA6358DFFE1224DE9B6FD2EFA9EFD4B" author="Olson, S. L." box="[1186,1319,729,755]" pageId="26" pageNumber="27" pagination="110 - 114" refId="ref29357" refString="Olson, S. L. (2007) The '' walking eagle' ' Wetmoregyps daggetti Miller: a scaled-up version of the Savanna Hawk (Buteogallus meridionalis). Ornithological Monographs, 63, 110 - 114. https: // doi. org / 10.1642 / 0078 - 6594 (2007) 63 [110: TWEWDM] 2.0. CO; 2" type="journal article" year="2007">Olson 2007</bibRefCitation>
;
<bibRefCitation id="EFA6358DFFE1224DE827FD2EFE95FCA0" author="Suarez, W. &amp; Olson, S. L." pageId="26" pageNumber="27" pagination="250 - 254" refId="ref31383" refString="Suarez, W. &amp; Olson, S. L. (2009 b) The generic position of Miraquila terrestris Campbell: Another addition to the buteogalline radiation from the Pleistocene of Peru. Journal of Raptor Research, 43, 250 - 254. https: // doi. org / 10.3356 / JRR- 08 - 85.1" type="journal article" year="2009">Suárez &amp; Olson 2009b</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>