716 lines
85 KiB
XML
716 lines
85 KiB
XML
<document id="A0CA6CEA69FA97AD3FC20E36DF5A37D5" ID-CLB-Dataset="27107" ID-DOI="10.11646/zootaxa.4586.2.3" ID-GBIF-Dataset="99c5e021-c0f5-40d3-b1fe-65bfb0715bf1" ID-ISSN="1175-5326" ID-Zenodo-Dep="2644466" ID-ZooBank="576F091A-E066-487A-91C9-A55DE721A6F4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1555539506244" checkinUser="plazi" docAuthor="Luu, Vinh Quang, Ziegler, Thomas, Ha, Nghia Van, Le, Minh Duc & Hoang, Tuoi Thi" docDate="2019" docId="AC0AE132FF874B1392FEF96DEEE6FBE1" docLanguage="en" docName="zootaxa.4586.2.3.pdf" docOrigin="Zootaxa 4586 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lycodon namdongensis Luu & Ziegler & Ha & Le & Hoang 2019, sp. nov." docType="treatment" docVersion="9" lastPageNumber="271" masterDocId="5033994AFF814B199269FFE1ED57FF9E" masterDocTitle="A new species of Lycodon Boie, 1826 (Serpentes: Colubridae) from Thanh Hoa Province, Vietnam" masterLastPageNumber="277" masterPageNumber="261" pageNumber="267" updateTime="1698722235096" updateUser="ExternalLinkService">
|
||
<mods:mods id="97609036F85D68B8FF9661E4107DCA10" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="EE2B7B84592D6F322C7F5AC66BD8BE45">
|
||
<mods:title id="D3B2CAE6439C4C920BBE2DCE5FBCCD3B">A new species of Lycodon Boie, 1826 (Serpentes: Colubridae) from Thanh Hoa Province, Vietnam</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="0DA97FBEDAF1916D431E5B01CC81F2E7" type="personal">
|
||
<mods:role id="015466D913A4C7B5B20652569557B096">
|
||
<mods:roleTerm id="DE235B7425CF76E38A226AE292338B02">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="6F299F991ACB16388D0448E7D22A89B0">Luu, Vinh Quang</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="435F6C6BC0CC7579F3311D7BEED11526" type="personal">
|
||
<mods:role id="2FD3CF4CF4E200ECC87100CEEA85F1C4">
|
||
<mods:roleTerm id="AA36E2F340AD698078640653B4AF3D62">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="1D7175E63EA2A647DDB0904AF85F45F0">Ziegler, Thomas</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="252839E514C6F8CA6546D66B4AD8C7BE" type="personal">
|
||
<mods:role id="9057AA3A9B2499D4BE8D89C028E6A230">
|
||
<mods:roleTerm id="2673F1087069F8B52086FDDF731E59F6">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="6DB835F354790328632446144259D01A">Ha, Nghia Van</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="1C92EADA9DDAC04A0B674333FF066D77" type="personal">
|
||
<mods:role id="7341DD6A844BA561F00BB5219F3BA264">
|
||
<mods:roleTerm id="76330EC047E56F554BD20142D08302AC">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="E4AA946DF19AF81DE9A485619BE07628">Le, Minh Duc</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="D2A7109286FD7384962DFCCCC387EE7F" type="personal">
|
||
<mods:role id="003DA0DB1089BB0029A5B3A7B33ED2F0">
|
||
<mods:roleTerm id="63BC7E95BEF7D48D4BDF02B15D67DB28">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="DF51EC2FD13EACB88BDDD40FC8246283">Hoang, Tuoi Thi</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="C61776D19A5510548385BFC85F18553F">text</mods:typeOfResource>
|
||
<mods:relatedItem id="A7B304E92ADA4E868722A50857B9B97A" type="host">
|
||
<mods:titleInfo id="D4F3A4CFB0269017CFE1835E97357E0F">
|
||
<mods:title id="330A90E08F82DD4B2775BD829254A995">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="3E6468B9A179FF9DDA7ADD5890E73C2E">
|
||
<mods:date id="912C316FB67DE91E1E63674297A616D5">2019</mods:date>
|
||
<mods:detail id="DD0048E2BB640F6A26F148F7FC93035C" type="pubDate">
|
||
<mods:number id="0A671A966965345BDE8DF660533A885B">2019-04-17</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="1C080EAC6AA36AF694045548EF6A8CA2" type="volume">
|
||
<mods:number id="86BEDDE9416FC341F76E2EDAB646EE0B">4586</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="EAFA2FD70DC6EBEB6F54FC2A56BE17B4" type="issue">
|
||
<mods:number id="7F1B2A7A81C1450E2651895A4BD353F4">2</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="05A2652FF2F3536230CB19073C3A0EA1" unit="page">
|
||
<mods:start id="DD7582957BE53FA3D178681A643A7C77">261</mods:start>
|
||
<mods:end id="95938FA78E40122532C93026B7E124B5">277</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="70E628692B46A6A6E27F8CEA54EC9A39">journal article</mods:classification>
|
||
<mods:identifier id="721F4CCA35A0E8FBFBF6F7A7ACEC49B4" type="CLB-Dataset">27107</mods:identifier>
|
||
<mods:identifier id="DABEC295F4C7C72B34EC615C899DA2D2" type="DOI">10.11646/zootaxa.4586.2.3</mods:identifier>
|
||
<mods:identifier id="83BEDBA1EF6069F2EFD2531EAE720A49" type="GBIF-Dataset">99c5e021-c0f5-40d3-b1fe-65bfb0715bf1</mods:identifier>
|
||
<mods:identifier id="197F7995B76FB0E70DF4E21563B7F348" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="0FE632EC203AA3AB7AF4B2DBA8EC36BC" type="Zenodo-Dep">2644466</mods:identifier>
|
||
<mods:identifier id="16D38F149B2685DC52773E4325233F46" type="ZooBank">576F091A-E066-487A-91C9-A55DE721A6F4</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="AC0AE132FF874B1392FEF96DEEE6FBE1" ID-DOI="http://doi.org/10.5281/zenodo.5587198" ID-GBIF-Taxon="156788471" ID-Zenodo-Dep="5587198" LSID="urn:lsid:plazi:treatment:AC0AE132FF874B1392FEF96DEEE6FBE1" httpUri="http://treatment.plazi.org/id/AC0AE132FF874B1392FEF96DEEE6FBE1" lastPageId="10" lastPageNumber="271" pageId="6" pageNumber="267">
|
||
<subSubSection id="6CB903AFFF874B1F92FEF96DEF4FF938" box="[151,536,1676,1702]" pageId="6" pageNumber="267" type="nomenclature">
|
||
<paragraph id="241C5024FF874B1F92FEF96DEF4FF938" blockId="6.[151,536,1676,1735]" box="[151,536,1676,1702]" pageId="6" pageNumber="267">
|
||
<heading id="7F54E748FF874B1F92FEF96DEF4FF938" bold="true" box="[151,536,1676,1702]" fontSize="11" level="1" pageId="6" pageNumber="267" reason="1">
|
||
<emphasis id="16D78C36FF874B1F92FEF96DEF4FF938" bold="true" box="[151,536,1676,1702]" pageId="6" pageNumber="267">
|
||
<taxonomicName id="E3A32BA7FF874B1F92FEF96DECE4F938" authority="Luu & Ziegler & Ha & Le & Hoang, 2019" authorityName="Luu & Ziegler & Ha & Le & Hoang" authorityYear="2019" box="[151,435,1676,1702]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="267" phylum="Chordata" rank="species" species="namdongensis" status="sp. nov.">
|
||
<emphasis id="16D78C36FF874B1F92FEF96DECE4F938" bold="true" box="[151,435,1676,1702]" italics="true" pageId="6" pageNumber="267">Lycodon namdongensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="0DE4314DFF874B1F93D3F96DEF4FF938" box="[442,536,1676,1702]" pageId="6" pageNumber="267" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="6CB903AFFF874B1F92FEF94FEC55F959" box="[151,258,1710,1735]" pageId="6" pageNumber="267" type="description">
|
||
<paragraph id="241C5024FF874B1F92FEF94FEC55F959" blockId="6.[151,536,1676,1735]" box="[151,258,1710,1735]" pageId="6" pageNumber="267">
|
||
<figureCitation id="BC984CA1FF874B1F92FEF94FEDB2F959" box="[151,229,1710,1735]" captionStart="FIGURE 2" captionStartId="7.[151,250,825,847]" captionTargetBox="[151,1435,265,804]" captionTargetId="figure@7.[151,1435,265,804]" captionTargetPageId="7" captionText="FIGURE 2. Dorsal view of the holotype of Lycodon namdongensis sp. nov. (VNUF R.2017.23) in life. Photo: N. V. Ha" httpUri="https://zenodo.org/record/2644470/files/figure.png" pageId="6" pageNumber="267">Figs. 2</figureCitation>
|
||
¯4
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="6CB903AFFF874B1F92FEF917E9CDF8C9" pageId="6" pageNumber="267" type="materials_examined">
|
||
<paragraph id="241C5024FF874B1F92FEF917E9CDF8C9" blockId="6.[151,1436,1782,2024]" pageId="6" pageNumber="267">
|
||
<materialsCitation id="94CB5A79FF874B1F92FEF917E9CDF8C9" ID-GBIF-Occurrence="2236308345" collectingDate="2017-06-13" collectionCode="VNUF" collectorName="Nghia Van Ha & Oanh Van Lo & Ngoan Van Ha" country="Vietnam" elevation="616" latitude="20.304966" location="Nam Dong Nature Reserve" longLatPrecision="1" longitude="104.91293" municipality="Quan Son District" pageId="6" pageNumber="267" specimenCount="1" stateProvince="Thanh Hoa" typeStatus="holotype">
|
||
<emphasis id="16D78C36FF874B1F92FEF917EC5AF891" bold="true" box="[151,269,1782,1807]" pageId="6" pageNumber="267">
|
||
<typeStatus id="FB18EE86FF874B1F92FEF917EC5FF891" box="[151,264,1782,1807]" pageId="6" pageNumber="267" type="holotype">Holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
<collectionCode id="42B2C8E1FF874B1F937DF919EC37F891" box="[276,352,1784,1807]" pageId="6" pageNumber="267">VNUF</collectionCode>
|
||
R.2017.23 (field number: TT 17.23), adult male, collected on
|
||
<date id="501D76E4FF874B1F9672F916E9E7F891" box="[1051,1200,1783,1807]" pageId="6" pageNumber="267" value="2017-06-13">
|
||
<collectingDate id="40598F0CFF874B1F9672F916E9E7F891" box="[1051,1200,1783,1807]" pageId="6" pageNumber="267" value="2017-06-13">13 June 2017</collectingDate>
|
||
</date>
|
||
by Vinh Quang Luu,
|
||
<collectorName id="895635F2FF874B1F92FEF8FAEC60F8AD" box="[151,311,1819,1844]" pageId="6" pageNumber="267">Nghia Van Ha</collectorName>
|
||
,
|
||
<collectorName id="895635F2FF874B1F932AF8FAEC8DF8AD" box="[323,474,1819,1844]" pageId="6" pageNumber="267">Oanh Van Lo</collectorName>
|
||
, and
|
||
<collectorName id="895635F2FF874B1F907DF8FDEFEEF8AD" box="[532,697,1820,1844]" pageId="6" pageNumber="267">Ngoan Van Ha</collectorName>
|
||
in the karst forest,
|
||
<location id="217C06FFFF874B1F91E6F8FDE996F8AD" LSID="urn:lsid:plazi:treatment:AC0AE132FF874B1392FEF96DEEE6FBE1:217C06FFFF874B1F91E6F8FDE996F8AD" box="[911,1217,1820,1844]" country="Vietnam" latitude="20.304966" longLatPrecision="1" longitude="104.91293" municipality="Quan Son District" name="Nam Dong Nature Reserve" pageId="6" pageNumber="267" stateProvince="Thanh Hoa">Nam Dong Nature Reserve</location>
|
||
,
|
||
<collectingMunicipality id="C478CA5EFF874B1F96A2F8FAE8CFF8AA" box="[1227,1432,1819,1844]" pageId="6" pageNumber="267">Quan Son District</collectingMunicipality>
|
||
,
|
||
<collectingRegion id="E6679EC6FF874B1F92FEF8DFECD6F8C9" box="[151,385,1854,1879]" country="Vietnam" name="Thanh Hoa" pageId="6" pageNumber="267">Thanh Hoa Province</collectingRegion>
|
||
,
|
||
<collectingCountry id="5CB410B4FF874B1F93E4F8DFECBBF8C9" box="[397,492,1854,1879]" name="Vietnam" pageId="6" pageNumber="267">Vietnam</collectingCountry>
|
||
(
|
||
<geoCoordinate id="419736E3FF874B1F9392F8DEEFC7F8C9" box="[507,656,1855,1879]" direction="north" orientation="latitude" pageId="6" pageNumber="267" precision="1" value="20.304966">20°18.298’N</geoCoordinate>
|
||
;
|
||
<geoCoordinate id="419736E3FF874B1F90F5F8DFEE6CF8C9" box="[668,827,1854,1879]" direction="east" orientation="longitude" pageId="6" pageNumber="267" precision="1" value="104.91293">104°54.776’E</geoCoordinate>
|
||
, at an elevation of
|
||
<elevation id="AF8EB717FF874B1F9679F8DFE9C7F8C9" box="[1040,1168,1854,1879]" metricMagnitude="2" metricUnit="m" metricValue="6.16" pageId="6" pageNumber="267" unit="m" value="616.0">
|
||
<quantity id="E35BFDC1FF874B1F9679F8DFE900F8C9" box="[1040,1111,1854,1879]" metricMagnitude="2" metricUnit="m" metricValue="6.16" pageId="6" pageNumber="267" unit="m" value="616.0">616 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
)
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="6CB903AFFF874B1E92AEF882EE2BFF4B" lastPageId="7" lastPageNumber="268" pageId="6" pageNumber="267" type="diagnosis">
|
||
<paragraph id="241C5024FF874B1E92AEF882EE2BFF4B" blockId="6.[151,1436,1782,2024]" lastBlockId="7.[151,1437,151,213]" lastPageId="7" lastPageNumber="268" pageId="6" pageNumber="267">
|
||
<emphasis id="16D78C36FF874B1F92AEF882EC14F8E2" bold="true" box="[199,323,1891,1916]" pageId="6" pageNumber="267">Diagnosis:</emphasis>
|
||
<taxonomicName id="E3A32BA7FF874B1F9322F882EF05F8E5" authority="Luu & Ziegler & Ha & Le & Hoang, 2019" authorityName="Luu & Ziegler & Ha & Le & Hoang" authorityYear="2019" box="[331,594,1891,1915]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="267" phylum="Chordata" rank="species" species="namdongensis" status="sp. nov.">
|
||
<emphasis id="16D78C36FF874B1F9322F882EF05F8E5" box="[331,594,1891,1915]" italics="true" pageId="6" pageNumber="267">Lycodon namdongensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="16D78C36FF874B1F9030F882EFF9F8E2" bold="true" box="[601,686,1891,1916]" pageId="6" pageNumber="267">
|
||
<taxonomicNameLabel id="0DE4314DFF874B1F9030F882EFF9F8E2" box="[601,686,1891,1916]" pageId="6" pageNumber="267" rank="species">sp. nov</taxonomicNameLabel>
|
||
</emphasis>
|
||
is characterized by the following morphological characters: dorsal scales in 17¯17¯15 rows, all smooth; supralabials 8; infralabials 10; one large loreal on each side, separated from the eye; cloacal plate single; ventral scales 218 (plus two preventral scales); subcaudals 85; a total length of
|
||
<quantity id="E35BFDC1FF874B1F971BF84AED94F879" metricMagnitude="-1" metricUnit="m" metricValue="7.2299999999999995" pageId="6" pageNumber="267" unit="mm" value="723.0">723 mm</quantity>
|
||
in the single male; tail / total length ratio 0.205; dorsal head pattern whitish grey; dorsal surface of body with 23 narrowed greyish cream bands; dorsal surface of tail with 14 cream bands; ventral surface of body mostly cream with a few dark spots posteriorly; ventral surface of tail dark grey.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="70DC00ACFF864B1E92FEFCD8E802FCD1" box="[151,1365,825,847]" httpUri="https://zenodo.org/record/2644470/files/figure.png" pageId="7" pageNumber="268" startId="7.[151,250,825,847]" targetBox="[151,1435,265,804]" targetPageId="7">
|
||
<paragraph id="241C5024FF864B1E92FEFCD8E802FCD1" blockId="7.[151,1365,825,847]" box="[151,1365,825,847]" pageId="7" pageNumber="268">
|
||
<emphasis id="16D78C36FF864B1E92FEFCD8EC45FCD0" bold="true" box="[151,274,825,847]" pageId="7" pageNumber="268">FIGURE 2.</emphasis>
|
||
Dorsal view of the holotype of
|
||
<taxonomicName id="E3A32BA7FF864B1E903CFCD8EE14FCD1" authority="Luu & Ziegler & Ha & Le & Hoang, 2019" authorityName="Luu & Ziegler & Ha & Le & Hoang" authorityYear="2019" box="[597,835,825,847]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="268" phylum="Chordata" rank="species" species="namdongensis" status="sp. nov.">
|
||
<emphasis id="16D78C36FF864B1E903CFCD8EE14FCD1" box="[597,835,825,847]" italics="true" pageId="7" pageNumber="268">Lycodon namdongensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="16D78C36FF864B1E9120FCDBEECEFCD1" bold="true" box="[841,921,826,847]" pageId="7" pageNumber="268">
|
||
<taxonomicNameLabel id="0DE4314DFF864B1E9120FCDBEECEFCD1" box="[841,921,826,847]" pageId="7" pageNumber="268" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(VNUF R.2017.23) in life. Photo: N. V. Ha
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="70DC00ACFF864B1E92FEF9BCEF2AF90F" httpUri="https://zenodo.org/record/2644472/files/figure.png" pageId="7" pageNumber="268" startId="7.[151,250,1629,1651]" targetBox="[151,1435,904,1607]" targetPageId="7">
|
||
<paragraph id="241C5024FF864B1E92FEF9BCEF2AF90F" blockId="7.[151,1436,1628,1681]" pageId="7" pageNumber="268">
|
||
<emphasis id="16D78C36FF864B1E92FEF9BCEC44F9EC" bold="true" box="[151,275,1629,1651]" pageId="7" pageNumber="268">FIGURE 3.</emphasis>
|
||
Lateral (A, B), dorsal (C), and ventral (D) views of the head of the preserved holotype of
|
||
<taxonomicName id="E3A32BA7FF864B1E96C7F9BDE8CBF9EC" authority="Luu & Ziegler & Ha & Le & Hoang, 2019" authorityName="Luu & Ziegler & Ha & Le & Hoang" authorityYear="2019" box="[1198,1436,1628,1650]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="268" phylum="Chordata" rank="species" species="namdongensis" status="sp. nov.">
|
||
<emphasis id="16D78C36FF864B1E96C7F9BDE8CBF9EC" box="[1198,1436,1628,1650]" italics="true" pageId="7" pageNumber="268">Lycodon namdongensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="16D78C36FF864B1E92FEF99DEDB0F90F" bold="true" box="[151,231,1660,1681]" pageId="7" pageNumber="268">
|
||
<taxonomicNameLabel id="0DE4314DFF864B1E92FEF99DEDB0F90F" box="[151,231,1660,1681]" pageId="7" pageNumber="268" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(VNUF R.2017.23). Photos: V. Q. Luu.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="6CB903AFFF864B1392AEF95FEF90FCA6" lastPageId="10" lastPageNumber="271" pageId="7" pageNumber="268" type="description">
|
||
<paragraph id="241C5024FF864B1192AEF95FEE47FE19" blockId="7.[151,1437,1726,2003]" lastBlockId="8.[151,1436,151,392]" lastPageId="8" lastPageNumber="269" pageId="7" pageNumber="268">
|
||
<emphasis id="16D78C36FF864B1E92AEF95FEF4DF949" bold="true" box="[199,538,1726,1751]" pageId="7" pageNumber="268">
|
||
Description of the
|
||
<typeStatus id="FB18EE86FF864B1E93C3F95FEF42F949" box="[426,533,1726,1751]" pageId="7" pageNumber="268" type="holotype">holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
Head elongate (HL
|
||
<quantity id="E35BFDC1FF864B1E9161F95EEE25F949" box="[776,882,1727,1751]" metricMagnitude="-2" metricUnit="m" metricValue="2.04" pageId="7" pageNumber="268" unit="mm" value="20.4">20.4 mm</quantity>
|
||
), moderately distinct from the neck, longer than wide (HW/HL ratio 0.54), depressed (HH/HL ratio 0.40), narrow anteriorly (IN/IO ratio 0.68); snout elongate (SnL/HL ratio 0.33); nostril lateral, oval shaped, located in the middle of the nasal; eye large (ED/HL ratio 0.13), pupils vertically elliptic; rostral triangular, much broader than high, hardly visible from above; nasal divided into two scales by a vertical ridge along posterior edge of nostril; two square internasals, as wide as long, bordered by two large, subpentagonal prefrontals posteriorly; frontal single, enlarged, pentagonal, narrow posteriorly; parietals longer than wide, in contact with each other medially, with upper anterior and posterior temporals, paraparietal laterally and three nuchal scales posteriorly; loreal 1/1 (on the left side the normal sized loreal posteriorly is followed by an additional, very tiny one), elongate, not entering orbit; supralabials 8/8, first and second in contact with nasal, only fifth entering orbit on left side, and third to fifth entering orbit on right side, sixth largest; infralabials 10/10, first pair in broad contact with each other, first to fifth in contact with anterior chin shields; anterior and posterior pairs of chin shields elongate, of about the same size and shape, separated by a medial groove, anterior pair slightly larger than the posterior; preoculars 3/1 (the lower two, tiny preoculars on the left side represent separated upper parts of supralabials 3¯4); postoculars 2/2; anterior temporals 2/2, posterior temporals 2/ 2, upper anterior ones distinctly thinner than lower ones.
|
||
</paragraph>
|
||
<caption id="70DC00ACFF894B1192FEF993ECC9F938" httpUri="https://zenodo.org/record/2644474/files/figure.png" pageId="8" pageNumber="269" startId="8.[151,250,1650,1672]" targetBox="[292,1262,455,1597]" targetPageId="8">
|
||
<paragraph id="241C5024FF894B1192FEF993ECC9F938" blockId="8.[151,1437,1650,1702]" pageId="8" pageNumber="269">
|
||
<emphasis id="16D78C36FF894B1192FEF993EC42F919" bold="true" box="[151,277,1650,1672]" pageId="8" pageNumber="269">FIGURE 4.</emphasis>
|
||
Dorsal (A) and lateral head (B) views of the holotype of
|
||
<taxonomicName id="E3A32BA7FF894B119111F993E93FF916" authority="Luu & Ziegler & Ha & Le & Hoang, 2019" authorityName="Luu & Ziegler & Ha & Le & Hoang" authorityYear="2019" box="[888,1128,1650,1672]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="269" phylum="Chordata" rank="species" species="namdongensis" status="sp. nov.">
|
||
<emphasis id="16D78C36FF894B119111F993E93FF916" box="[888,1128,1650,1672]" italics="true" pageId="8" pageNumber="269">Lycodon namdongensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="16D78C36FF894B119618F993E993F919" bold="true" box="[1137,1220,1650,1671]" pageId="8" pageNumber="269">
|
||
<taxonomicNameLabel id="0DE4314DFF894B119618F993E993F919" box="[1137,1220,1650,1671]" pageId="8" pageNumber="269" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(VNUF R.2017.23). Drawings by T. T. Hoang.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="241C5024FF894B1192AEF90DE867F8EF" blockId="8.[151,1437,1772,2013]" pageId="8" pageNumber="269">Left maxilla arched, with an angulous apex, distinctly bent inwards anteriorly. A total of 12 maxillary teeth or teeth alveola, with the following formula: four small anterior teeth, with the last two ones being somewhat enlarged + two strongly enlarged teeth, thick, and not much curved + a wide gap, of about the length of the largest teeth before + three small teeth + a small gap + three posterior teeth, with the first two of them being enlarged.</paragraph>
|
||
<paragraph id="241C5024FF894B1192AEF89DEE07F843" blockId="8.[151,1437,1772,2013]" pageId="8" pageNumber="269">
|
||
Body elongate, SVL
|
||
<quantity id="E35BFDC1FF894B1193D1F89DEF4EF80A" box="[440,537,1916,1941]" metricMagnitude="-1" metricUnit="m" metricValue="5.75" pageId="8" pageNumber="269" unit="mm" value="575.0">575 mm</quantity>
|
||
; TaL
|
||
<quantity id="E35BFDC1FF894B119032F89CEFEAF80A" box="[603,701,1917,1941]" metricMagnitude="-1" metricUnit="m" metricValue="1.48" pageId="8" pageNumber="269" unit="mm" value="148.0">148 mm</quantity>
|
||
; TL
|
||
<quantity id="E35BFDC1FF894B11909CF89CEE01F80A" box="[757,854,1917,1941]" metricMagnitude="-1" metricUnit="m" metricValue="7.2299999999999995" pageId="8" pageNumber="269" unit="mm" value="723.0">723 mm</quantity>
|
||
; ratio of tail length/total length 0.205; ventrals 218 (plus two preventrals); subcaudals 85, divided, weakly notched laterally; cloacal single; DSR 17¯17¯15; all dorsal scale rows smooth; the vertebral scales not enlarged (
|
||
<tableCitation id="6921659FFF894B119082F825EE68F843" box="[747,831,1988,2013]" captionStart="TABLE 1" captionStartId="1.[147,231,574,596]" captionTargetPageId="1" captionText="TABLE 1. Taxonomic review of Lycodon members from Vietnam (data obtained from Boulenger 1893; Smith 1943; Deuve 1970; Hu et al. 1975; Ziegler et al. 2007; Nguyen et al. 2009; Vogel & David 2010; Geissler et al. 2011; Orlov et al. 2011; Neang et al. 2014; Luu et al. 2013a,b; Jestrzemski et al. 2013; Phan et al. 2014 Grismer et al. 2014; Ziegler et al. 2014; Gawor et al. 2016; Ngo & Duong 2016; Nguyen et al. 2016; Do et al. 2016; Vassilieva et al. 2016; Pham et al. 2017; Do et al. 2017; Luu et al. 2018). 1—maximum snout-vent length (in mm); 2—maximum tail length; 3—maximum of total length; 4—number of supralabials; 5—number of infralabials; 6—loreal touching orbit; 7— number of preoculars; 8—number of postoculars; 9—number of loreals; 10—dorsal scale row fomula; 11—number of ventral scales; 12—number of subcaudals; 13—number of cloacal (1 or 2); 14—head pattern; 15—belly banded; 16—dorsal body pattern; 17—number of body bands/blotches; 18—number of tail bands/blotches; 19—color of light body bands/blotches; 20— ground coloration on dorsum; 21— dorsal scales keeled/smooth; 22—belly pattern; 23— vertebral scale row enlarged; 24— first body band position (at VEN); 25—first body band width; 26—ventral tail banded; 27—maxillary teeth, anterior=A, posterior=P; 28—temporals (A+P); 29—nuchal band; 30—distribution. “1” = presence of character state; “0” = absence of character state; “̅” = missing data; “juv.” = juvenile; “*” = this study." httpUri="http://table.plazi.org/id/70DC00ACFF804B1892FAFDDFE85AFCA6" pageId="8" pageNumber="269" tableUuid="70DC00ACFF804B1892FAFDDFE85AFCA6">Table 1</tableCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="241C5024FF884B1092AEFF76ECD5FE16" blockId="9.[151,1436,151,2013]" pageId="9" pageNumber="270">
|
||
<emphasis id="16D78C36FF884B1092AEFF76ECFCFF2E" bold="true" box="[199,427,151,176]" pageId="9" pageNumber="270">Colouration in life:</emphasis>
|
||
Head pattern whitish grey; dorsal surface of the head dark in the anterior half and slightly cream posteriorly; ventral side of the head cream, dark-grey anteriorly; dorsal surface of body dark grey with 23 narrowed greyish cream dorsal bands, each band with dark flank spots; first body band starting at ventral scale 25, one vertebral scale covered by this band, and slightly extended laterally to a width of 2–4 (mostly 2) scales; ventral surface mostly cream with a few small dark spots posteriorly; dorsal surface of tail with 14 cream bands, ventral surface of tail dark grey with irregular blotches corresponding to light crossbands, however, these bands do not form complete rings.
|
||
</paragraph>
|
||
<paragraph id="241C5024FF884B1092AEFE75EF83FE4E" blockId="9.[151,1436,151,2013]" pageId="9" pageNumber="270">
|
||
<emphasis id="16D78C36FF884B1092AEFE75EC07FE33" bold="true" box="[199,336,404,429]" pageId="9" pageNumber="270">Hemipenis:</emphasis>
|
||
The left hemipenis is only in part everted, hemipenis cylindrical at base, lower part smooth, upper part densely covered with spines, slightly widened.
|
||
</paragraph>
|
||
<paragraph id="241C5024FF884B1392AEFE3DEC73FE4E" blockId="9.[151,1436,151,2013]" lastBlockId="10.[151,1436,151,1151]" lastPageId="10" lastPageNumber="271" pageId="9" pageNumber="270">
|
||
<emphasis id="16D78C36FF884B1092AEFE3DEC38FE6B" bold="true" box="[199,367,476,501]" pageId="9" pageNumber="270">Comparisons:</emphasis>
|
||
<taxonomicName id="E3A32BA7FF884B10931CFE3DEF2BFE6A" authority="Luu & Ziegler & Ha & Le & Hoang, 2019" authorityName="Luu & Ziegler & Ha & Le & Hoang" authorityYear="2019" box="[373,636,476,500]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="270" phylum="Chordata" rank="species" species="namdongensis" status="sp. nov.">
|
||
<emphasis id="16D78C36FF884B10931CFE3DEF2BFE6A" box="[373,636,476,500]" italics="true" pageId="9" pageNumber="270">Lycodon namdongensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="16D78C36FF884B1090EBFE3DEF8BFE6B" bold="true" box="[642,732,476,501]" pageId="9" pageNumber="270">
|
||
<taxonomicNameLabel id="0DE4314DFF884B1090EBFE3DEF8BFE6B" box="[642,732,476,501]" pageId="9" pageNumber="270" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be distinguished from its closest relative
|
||
<taxonomicName id="E3A32BA7FF884B1096B1FE3CE82EFE6A" baseAuthorityName="Pope" baseAuthorityYear="1928" box="[1240,1401,476,500]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="270" phylum="Chordata" rank="species" species="futsingensis">
|
||
<emphasis id="16D78C36FF884B1096B1FE3CE82EFE6A" box="[1240,1401,476,500]" italics="true" pageId="9" pageNumber="270">L. futsingensis</emphasis>
|
||
</taxonomicName>
|
||
by having more ventral scales (218
|
||
<emphasis id="16D78C36FF884B109077FDE0EF31FD86" box="[542,614,513,536]" italics="true" pageId="9" pageNumber="270">versus</emphasis>
|
||
193–208), ventral pattern mostly cream with a few small dark spots posteriorly (
|
||
<emphasis id="16D78C36FF884B109348FDC4EC3EFDA2" box="[289,361,549,572]" italics="true" pageId="9" pageNumber="270">versus</emphasis>
|
||
speckled posteriorly), first body band starting at ventral scale 25 (
|
||
<emphasis id="16D78C36FF884B10963AFDC4E9CCFDA2" box="[1107,1179,549,572]" italics="true" pageId="9" pageNumber="270">versus</emphasis>
|
||
13–21), and first body band covered by one vertebral scale (
|
||
<emphasis id="16D78C36FF884B109022FDA8EFC4FDFE" box="[587,659,585,608]" italics="true" pageId="9" pageNumber="270">versus</emphasis>
|
||
5–10) (
|
||
<bibRefCitation id="40322DD5FF884B109086FDA6EEECFDFE" author="Vogel, G. & David, P. & Pauwels, O. S. G. & Sumontha, M. & Norval, G. & Hendrix, R. & Vu, N. T. & Ziegler, T." box="[751,955,583,608]" pageId="9" pageNumber="270" pagination="131" refId="ref10038" refString="Vogel, G., David, P., Pauwels, O. S. G., Sumontha, M., Norval, G., Hendrix, R., Vu, N. T. & Ziegler, T. (2009) A revision of Lycodon ruhstrati (Fischer 1886) auctorum (Squamata Colubridae), with the description of a new species from Thailand and a new subspecies from the Asian mainland. Tropical Zoology, 22, 131 ‾ 182." type="journal article" year="2009">
|
||
Vogel
|
||
<emphasis id="16D78C36FF884B109152FDA8EE39FDFE" box="[827,878,584,608]" italics="true" pageId="9" pageNumber="270">et al</emphasis>
|
||
. 2009
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="40322DD5FF884B1091A3FDA9E9D9FDFE" author="Luu, V. Q. & Nguyen, T. Q. & Calame, T. & Hoang, T. T. & Southichack, S. & Bonkowski, M. & Ziegler, T." box="[970,1166,584,608]" pageId="9" pageNumber="270" refId="ref8782" refString="Luu, V. Q., Nguyen, T. Q., Calame, T., Hoang, T. T., Southichack, S., Bonkowski, M. & Ziegler, T. (2013 a) New country records of reptiles from Laos. Biodiversity Data Journal, 1, 1 ‾ 14. https: // doi. org / 10.3897 / BDJ. 1. e 1015" type="journal volume" year="2013">
|
||
Luu
|
||
<emphasis id="16D78C36FF884B109668FDA8E962FDFE" box="[1025,1077,584,608]" italics="true" pageId="9" pageNumber="270">et al</emphasis>
|
||
. 2013a
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="40322DD5FF884B1096F5FDA9E827FDFE" author="Neang, T. & Hartmann, T. & Hun, S. & Souter, N. J. & Furey, N. M." box="[1180,1392,584,608]" pageId="9" pageNumber="270" pagination="1" refId="ref8967" refString="Neang, T., Hartmann, T., Hun, S., Souter, N. J. & Furey, N. M. (2014) A new species of wolf snake (Colubridae: Lycodon Fitzinger, 1826) from Phnom Samkos Wildlife Sanctuary, Cardamom Mountains, southwest Cambodia. Zootaxa, 3814 (1), 068 ‾ 080. https: // doi. org / 10.11646 / zootaxa. 3814.1.3" type="journal article" year="2014">
|
||
Neang
|
||
<emphasis id="16D78C36FF884B109686FDA8E874FDFE" box="[1263,1315,584,608]" italics="true" pageId="9" pageNumber="270">et al</emphasis>
|
||
. 2014
|
||
</bibRefCitation>
|
||
), in concert with an uncorrected
|
||
<emphasis id="16D78C36FF884B109382FD8CECAEFD1A" box="[491,505,621,644]" italics="true" pageId="9" pageNumber="270">p</emphasis>
|
||
-distance of 6.5%; from
|
||
<taxonomicName id="E3A32BA7FF884B109176FD8CEE87FD1A" baseAuthorityName="Pope" baseAuthorityYear="1928" box="[799,976,620,644]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="270" phylum="Chordata" rank="species" species="flavozonatus">
|
||
<emphasis id="16D78C36FF884B109176FD8CEE87FD1A" box="[799,976,620,644]" italics="true" pageId="9" pageNumber="270">L. flavozonatus</emphasis>
|
||
</taxonomicName>
|
||
by having cloacal plate single (
|
||
<emphasis id="16D78C36FF884B10973DFD8CE8CBFD1A" box="[1364,1436,621,644]" italics="true" pageId="9" pageNumber="270">versus</emphasis>
|
||
divided), dorsal scale rows being completely smooth (
|
||
<emphasis id="16D78C36FF884B109172FD70EE34FD36" box="[795,867,657,680]" italics="true" pageId="9" pageNumber="270">versus</emphasis>
|
||
10–12 keeled dorsal scale rows at midbody), a distinctly lower body band count (23
|
||
<emphasis id="16D78C36FF884B109051FD54EFD7FD52" box="[568,640,693,716]" italics="true" pageId="9" pageNumber="270">versus</emphasis>
|
||
51–78), belly pattern mostly cream with a few small dark spots (
|
||
<emphasis id="16D78C36FF884B10973DFD54E8CBFD52" box="[1364,1436,693,716]" italics="true" pageId="9" pageNumber="270">versus</emphasis>
|
||
yellow with large black spots) (
|
||
<bibRefCitation id="40322DD5FF884B109392FD36EFA8FD6E" author="Orlov, N. & Ryabov, S. A." box="[507,767,727,752]" pageId="9" pageNumber="270" pagination="181" refId="ref9353" refString="Orlov, N. & Ryabov, S. A. (2004) Revalidization and change of taxonomic status of Dinodon rufozonatus meridionale Bourret, 1935 (Serpentes: Colubridae: Colubrinae). Russian Journal of Herpetology, 11 (3), 181 ‾ 197." type="journal article" year="2004">Orlov & Ryabov 2004</bibRefCitation>
|
||
); from
|
||
<taxonomicName id="E3A32BA7FF884B109138FD38EEABFD6E" box="[849,1020,728,752]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="270" phylum="Chordata" rank="species" species="meridionalis">
|
||
<emphasis id="16D78C36FF884B109138FD38EEABFD6E" box="[849,1020,728,752]" italics="true" pageId="9" pageNumber="270">L. meridionalis</emphasis>
|
||
</taxonomicName>
|
||
by having whitish grey head pattern (
|
||
<emphasis id="16D78C36FF884B1092C9FD1CEDBFFC8A" box="[160,232,765,788]" italics="true" pageId="9" pageNumber="270">versus</emphasis>
|
||
with yellow-black marble markings), fewer ventral scales (218
|
||
<emphasis id="16D78C36FF884B1091ACFD1CE95AFC8A" box="[965,1037,765,788]" italics="true" pageId="9" pageNumber="270">versus</emphasis>
|
||
234–246), dorsal scale rows being completely smooth (
|
||
<emphasis id="16D78C36FF884B109316FCC0EC90FCA6" box="[383,455,801,824]" italics="true" pageId="9" pageNumber="270">versus</emphasis>
|
||
distinctly keeled), dorsal pattern banded (
|
||
<emphasis id="16D78C36FF884B1091CDFCC0EEBCFCA6" box="[932,1003,801,824]" italics="true" pageId="9" pageNumber="270">versus</emphasis>
|
||
blotched), fewer subcaudal scales (86
|
||
<emphasis id="16D78C36FF884B1092FEFCA4ED88FCC2" box="[151,223,837,860]" italics="true" pageId="9" pageNumber="270">versus</emphasis>
|
||
98–106), ventral surface mostly cream with a few dark spots posteriorly (
|
||
<emphasis id="16D78C36FF884B10962BFCA4E9DDFCC2" box="[1090,1162,837,860]" italics="true" pageId="9" pageNumber="270">versus</emphasis>
|
||
yellow with dark spots posteriorly), and fewer tail bands (14
|
||
<emphasis id="16D78C36FF884B109025FC88EFC3FC1E" box="[588,660,873,896]" italics="true" pageId="9" pageNumber="270">versus</emphasis>
|
||
25–35) (
|
||
<bibRefCitation id="40322DD5FF884B109094FC89EECFFC1E" author="Bourret, R." box="[765,920,871,896]" pageId="9" pageNumber="270" pagination="17" refId="ref7759" refString="Bourret, R. (1935) Notes herpetologiques sur l'Indochine francaise. IX. Les Serpents de Chapa. Bulletin general de l'Instruction publique, 14 (1934 ‾ 1935), 7 (Mars), 239 ‾ 251 (Reprint: 5 ‾ 17)." type="journal article" year="1935">Bourret 1935</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="40322DD5FF884B1091CEFC86E9FBFC1E" author="Orlov, N. & Ryabov, S. A." box="[935,1196,871,896]" pageId="9" pageNumber="270" pagination="181" refId="ref9353" refString="Orlov, N. & Ryabov, S. A. (2004) Revalidization and change of taxonomic status of Dinodon rufozonatus meridionale Bourret, 1935 (Serpentes: Colubridae: Colubrinae). Russian Journal of Herpetology, 11 (3), 181 ‾ 197." type="journal article" year="2004">Orlov & Ryabov 2004</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="40322DD5FF884B1096D0FC89E8C7FC1E" author="Gawor, A. & Pham, C. T. & Nguyen, T. Q. & Nguyen, T. T. & Schmitz, A. & Ziegler, T." box="[1209,1424,871,896]" pageId="9" pageNumber="270" pagination="23 - 41" refId="ref8174" refString="Gawor, A., Pham, C. T., Nguyen, T. Q., Nguyen, T. T., Schmitz, A. & Ziegler, T. (2016) The herpetofauna of the Bai Tu Long National Park, northeastern Vietnam. Salamandra, 52 (1), 23 - 41." type="journal article" year="2016">
|
||
Gawor
|
||
<emphasis id="16D78C36FF884B109766FC88E816FC1E" box="[1295,1345,872,896]" italics="true" pageId="9" pageNumber="270">et al</emphasis>
|
||
. 2016
|
||
</bibRefCitation>
|
||
); from
|
||
<taxonomicName id="E3A32BA7FF884B1092BDFC6CEC6CFC3A" authorityName="Luu, Bonkowski, Nguyen, Le, Calame & Ziegler" authorityYear="2018" box="[212,315,908,932]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="270" phylum="Chordata" rank="species" species="banksi">
|
||
<emphasis id="16D78C36FF884B1092BDFC6CEC6CFC3A" box="[212,315,908,932]" italics="true" pageId="9" pageNumber="270">L. banksi</emphasis>
|
||
</taxonomicName>
|
||
by having a loreal not entering the orbit (
|
||
<emphasis id="16D78C36FF884B109167FC6CEE01FC3A" box="[782,854,909,932]" italics="true" pageId="9" pageNumber="270">versus</emphasis>
|
||
entering the orbit), fewer ventral scales (218
|
||
<emphasis id="16D78C36FF884B10973DFC6CE8CBFC3A" box="[1364,1436,909,932]" italics="true" pageId="9" pageNumber="270">versus</emphasis>
|
||
241), dorsal scale rows being completely smooth (
|
||
<emphasis id="16D78C36FF884B1090B0FC50EE77FC56" box="[729,800,945,968]" italics="true" pageId="9" pageNumber="270">versus</emphasis>
|
||
feebly keeled), dorsal pattern banded (
|
||
<emphasis id="16D78C36FF884B10968BFC50E87DFC56" box="[1250,1322,945,968]" italics="true" pageId="9" pageNumber="270">versus</emphasis>
|
||
blotched) (
|
||
<bibRefCitation id="40322DD5FF884B1092F7FC35EC0FFC72" author="Luu, V. Q. & Bonkowski, M. & Nguyen, T. Q. & Le, M. D. & Calame, T. & Ziegler, T." box="[158,344,980,1004]" pageId="9" pageNumber="270" pagination="263" refId="ref8713" refString="Luu, V. Q., Bonkowski, M., Nguyen, T. Q., Le, M. D., Calame, T. & Ziegler, T. (2018) A new species of Lycodon Boie, 1826 (Serpentes: Colubridae) from central Laos. Revue Suisse de Zoologie, 125 (2), 263 ‾ 267." type="journal article" year="2018">
|
||
Luu
|
||
<emphasis id="16D78C36FF884B1092BEFC34EC5EFC72" box="[215,265,980,1004]" italics="true" pageId="9" pageNumber="270">et al</emphasis>
|
||
. 2018
|
||
</bibRefCitation>
|
||
); from
|
||
<taxonomicName id="E3A32BA7FF884B1093C4FC34EF06FC72" authorityName="Cantor" authorityYear="1842" box="[429,593,980,1004]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="270" phylum="Chordata" rank="species" species="rufozonatus">
|
||
<emphasis id="16D78C36FF884B1093C4FC34EF06FC72" box="[429,593,980,1004]" italics="true" pageId="9" pageNumber="270">L. rufozonatus</emphasis>
|
||
</taxonomicName>
|
||
by having a distinctly lower body band count (23
|
||
<emphasis id="16D78C36FF884B1096F6FC34E9B0FC72" box="[1183,1255,981,1004]" italics="true" pageId="9" pageNumber="270">versus</emphasis>
|
||
44–52), having fewer dorsal scale rows at neck (17
|
||
<emphasis id="16D78C36FF884B10905CFC18EF2BFB8E" box="[565,636,1017,1040]" italics="true" pageId="9" pageNumber="270">versus</emphasis>
|
||
19), whitish grey dorsal head pattern (
|
||
<emphasis id="16D78C36FF884B109653FC18E9D6FB8E" box="[1082,1153,1017,1040]" italics="true" pageId="9" pageNumber="270">versus</emphasis>
|
||
dark brown with yellow borders), and body bands cream with a few small dark spots laterally (
|
||
<emphasis id="16D78C36FF884B109661FBFCE918FBAA" box="[1032,1103,1053,1076]" italics="true" pageId="9" pageNumber="270">versus</emphasis>
|
||
yellow or cream, speckled posteriorly) (
|
||
<bibRefCitation id="40322DD5FF884B109340FBDEECB5FBC6" author="Boulenger, G. A." box="[297,482,1087,1112]" pageId="9" pageNumber="270" refId="ref7724" refString="Boulenger, G. A. (1893) Catalogue of the snakes in the British Museum (Natural History). Fol. I. Taylor and Francis, London, 448 pp." type="book" year="1893">Boulenger 1893</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="40322DD5FF884B109387FBA1EF2CFBC6" author="Deuve, J." box="[494,635,1088,1112]" pageId="9" pageNumber="270" pagination="1 - 251" refId="ref7914" refString="Deuve, J. (1970) Serpents du Laos. Memoires de l'ORSTOM, Paris, 39, 1 - 251." type="journal article" year="1970">Deuve 1970</bibRefCitation>
|
||
); from
|
||
<taxonomicName id="E3A32BA7FF884B1090A7FBA0EEDEFBC9" baseAuthorityName="Cope" baseAuthorityYear="1860" box="[718,905,1088,1112]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="270" phylum="Chordata" rank="species" species="semicarinatus">
|
||
<emphasis id="16D78C36FF884B1090A7FBA0EEDEFBC9" box="[718,905,1088,1112]" italics="true" pageId="9" pageNumber="270">L. semicarinatus</emphasis>
|
||
</taxonomicName>
|
||
by dorsal scale rows being completely smooth (
|
||
<emphasis id="16D78C36FF884B1092C9FB84EDBFFBE2" box="[160,232,1125,1148]" italics="true" pageId="9" pageNumber="270">versus</emphasis>
|
||
keeled along anterior half), whitish grey dorsal head pattern (
|
||
<emphasis id="16D78C36FF884B1091DBFB84EEAEFBE2" box="[946,1017,1125,1148]" italics="true" pageId="9" pageNumber="270">versus</emphasis>
|
||
black), first and second labials dark grey (
|
||
<emphasis id="16D78C36FF884B1092B2FB68EC74FB3E" box="[219,291,1161,1184]" italics="true" pageId="9" pageNumber="270">versus</emphasis>
|
||
yellow with black sutures), and belly pattern mostly cream with a few small dark spots posteriorly (
|
||
<emphasis id="16D78C36FF884B1092C9FB4CEDBFFB5A" box="[160,232,1197,1220]" italics="true" pageId="9" pageNumber="270">versus</emphasis>
|
||
yellow) (
|
||
<bibRefCitation id="40322DD5FF884B109337FB4DEF49FB5A" author="Boulenger, G. A." box="[350,542,1196,1221]" pageId="9" pageNumber="270" refId="ref7724" refString="Boulenger, G. A. (1893) Catalogue of the snakes in the British Museum (Natural History). Fol. I. Taylor and Francis, London, 448 pp." type="book" year="1893">Boulenger 1893</bibRefCitation>
|
||
); from
|
||
<taxonomicName id="E3A32BA7FF884B109012FB4CEE47FB5A" baseAuthorityName="Boie" baseAuthorityYear="1827" box="[635,784,1197,1220]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="270" phylum="Chordata" rank="species" species="capucinus">
|
||
<emphasis id="16D78C36FF884B109012FB4CEE47FB5A" box="[635,784,1197,1220]" italics="true" pageId="9" pageNumber="270">L. capucinus</emphasis>
|
||
</taxonomicName>
|
||
by having more ventrals (218
|
||
<emphasis id="16D78C36FF884B1096E3FB4CE986FB5A" box="[1162,1233,1197,1220]" italics="true" pageId="9" pageNumber="270">versus</emphasis>
|
||
182–211), fewer supralabials (8
|
||
<emphasis id="16D78C36FF884B10932DFB30ECDBFB76" box="[324,396,1233,1256]" italics="true" pageId="9" pageNumber="270">versus</emphasis>
|
||
9–10), banded dorsal pattern (
|
||
<emphasis id="16D78C36FF884B109083FB30EE65FB76" box="[746,818,1233,1256]" italics="true" pageId="9" pageNumber="270">versus</emphasis>
|
||
reticulated), cloacal single (
|
||
<emphasis id="16D78C36FF884B10961CFB30E9EBFB76" box="[1141,1212,1233,1256]" italics="true" pageId="9" pageNumber="270">versus</emphasis>
|
||
divided), and more subcaudal scales (86
|
||
<emphasis id="16D78C36FF884B1093E3FB14EC85FA92" box="[394,466,1269,1292]" italics="true" pageId="9" pageNumber="270">versus</emphasis>
|
||
59–74) (
|
||
<bibRefCitation id="40322DD5FF884B109052FB15EE58FA92" author="Neang, T. & Hartmann, T. & Hun, S. & Souter, N. J. & Furey, N. M." box="[571,783,1268,1292]" pageId="9" pageNumber="270" pagination="1" refId="ref8967" refString="Neang, T., Hartmann, T., Hun, S., Souter, N. J. & Furey, N. M. (2014) A new species of wolf snake (Colubridae: Lycodon Fitzinger, 1826) from Phnom Samkos Wildlife Sanctuary, Cardamom Mountains, southwest Cambodia. Zootaxa, 3814 (1), 068 ‾ 080. https: // doi. org / 10.11646 / zootaxa. 3814.1.3" type="journal article" year="2014">
|
||
Neang
|
||
<emphasis id="16D78C36FF884B1090E6FB14EF95FA92" box="[655,706,1268,1292]" italics="true" pageId="9" pageNumber="270">et al</emphasis>
|
||
. 2014
|
||
</bibRefCitation>
|
||
); from
|
||
<taxonomicName id="E3A32BA7FF884B10910DFB14E964FA92" box="[868,1075,1268,1292]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="270" phylum="Chordata" rank="species" species="cardamomensis">
|
||
<emphasis id="16D78C36FF884B10910DFB14E964FA92" box="[868,1075,1268,1292]" italics="true" pageId="9" pageNumber="270">L. cardamomensis</emphasis>
|
||
</taxonomicName>
|
||
by having more light bands on dorsum (23
|
||
<emphasis id="16D78C36FF884B10934DFAF8EC3BFAAE" box="[292,364,1305,1328]" italics="true" pageId="9" pageNumber="270">versus</emphasis>
|
||
12–14), fewer dorsal scale rows at neck (17
|
||
<emphasis id="16D78C36FF884B109117FAF8EE92FAAE" box="[894,965,1305,1328]" italics="true" pageId="9" pageNumber="270">versus</emphasis>
|
||
19), dorsal scale rows being completely smooth (
|
||
<emphasis id="16D78C36FF884B109293FADCEC16FACA" box="[250,321,1341,1364]" italics="true" pageId="9" pageNumber="270">versus</emphasis>
|
||
keeled on midbody), and more tail bands (14
|
||
<emphasis id="16D78C36FF884B10912CFADCEEDAFACA" box="[837,909,1341,1364]" italics="true" pageId="9" pageNumber="270">versus</emphasis>
|
||
6–7) (
|
||
<bibRefCitation id="40322DD5FF884B1091BCFADDE9F6FACA" author="Neang, T. & Hartmann, T. & Hun, S. & Souter, N. J. & Furey, N. M." box="[981,1185,1340,1364]" pageId="9" pageNumber="270" pagination="1" refId="ref8967" refString="Neang, T., Hartmann, T., Hun, S., Souter, N. J. & Furey, N. M. (2014) A new species of wolf snake (Colubridae: Lycodon Fitzinger, 1826) from Phnom Samkos Wildlife Sanctuary, Cardamom Mountains, southwest Cambodia. Zootaxa, 3814 (1), 068 ‾ 080. https: // doi. org / 10.11646 / zootaxa. 3814.1.3" type="journal article" year="2014">
|
||
Neang
|
||
<emphasis id="16D78C36FF884B10964EFADCE90EFACA" box="[1063,1113,1340,1364]" italics="true" pageId="9" pageNumber="270">et al</emphasis>
|
||
. 2014
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="40322DD5FF884B1096C4FADCE806FACA" author="Do, D. T. & Ngo, C. D. & Ziegler, T. & Nguyen, T. Q." box="[1197,1361,1340,1364]" pageId="9" pageNumber="270" pagination="167 - 170" refId="ref8009" refString="Do, D. T., Ngo, C. D., Ziegler, T. & Nguyen, T. Q. (2017) First record of Lycodon cardamomensis Daltry et Wuster, 2002 (Squamata: Colubridae) from Vietnam. Russian Journal of Herpetology, 24 (2), 167 - 170." type="journal article" year="2017">
|
||
Do
|
||
<emphasis id="16D78C36FF884B1096BFFADCE85FFACA" box="[1238,1288,1340,1364]" italics="true" pageId="9" pageNumber="270">et al</emphasis>
|
||
. 2017
|
||
</bibRefCitation>
|
||
); from
|
||
<taxonomicName id="E3A32BA7FF884B1092FEFA80EDA8FAE6" authorityName="Vogel, Nguyen, Kingsada & Ziegler" authorityYear="2012" box="[151,255,1376,1400]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="270" phylum="Chordata" rank="species" species="davidi">
|
||
<emphasis id="16D78C36FF884B1092FEFA80EDA8FAE6" box="[151,255,1376,1400]" italics="true" pageId="9" pageNumber="270">L. davidi</emphasis>
|
||
</taxonomicName>
|
||
by having dorsal scale rows being completely smooth (
|
||
<emphasis id="16D78C36FF884B1091FDFA80EE8CFAE6" box="[916,987,1377,1400]" italics="true" pageId="9" pageNumber="270">versus</emphasis>
|
||
middorsal scale rows weakly keeled), slightly fewer ventrals (218
|
||
<emphasis id="16D78C36FF884B1093BDFA64EF4BFA02" box="[468,540,1413,1436]" italics="true" pageId="9" pageNumber="270">versus</emphasis>
|
||
224), body cross-bands greyish cream with dark spots laterally (
|
||
<emphasis id="16D78C36FF884B10969CFA64E86BFA02" box="[1269,1340,1413,1436]" italics="true" pageId="9" pageNumber="270">versus</emphasis>
|
||
pale tan brown) (
|
||
<bibRefCitation id="40322DD5FF884B10929EFA46ECE8FA5E" author="Vogel, G. & Nguyen, T. Q. & Kingsada, P. & Ziegler, T." box="[247,447,1447,1472]" pageId="9" pageNumber="270" pagination="344" refId="ref10187" refString="Vogel, G., Nguyen, T. Q., Kingsada, P. & Ziegler, T. (2012) A new species of the genus Lycodon Boie, 1826 from Laos (Squamata: Colubridae). North-Western Journal of Zoology, 8, 344 ‾ 352." type="journal article" year="2012">
|
||
Vogel
|
||
<emphasis id="16D78C36FF884B10932BFA48EC2CFA5E" box="[322,379,1448,1472]" italics="true" pageId="9" pageNumber="270">et al.</emphasis>
|
||
2012
|
||
</bibRefCitation>
|
||
); from
|
||
<taxonomicName id="E3A32BA7FF884B10907BFA48EFDBFA5E" box="[530,652,1448,1472]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="270" phylum="Chordata" rank="species" species="laoensis">
|
||
<emphasis id="16D78C36FF884B10907BFA48EFDBFA5E" box="[530,652,1448,1472]" italics="true" pageId="9" pageNumber="270">L. laoensis</emphasis>
|
||
</taxonomicName>
|
||
by having more ventrals (218
|
||
<emphasis id="16D78C36FF884B109180FA48E967FA5E" box="[1001,1072,1449,1472]" italics="true" pageId="9" pageNumber="270">versus</emphasis>
|
||
169–192), more subcaudals (85
|
||
<emphasis id="16D78C36FF884B1092FEFA2CED88FA7A" box="[151,223,1485,1508]" italics="true" pageId="9" pageNumber="270">versus</emphasis>
|
||
60–76), fewer supralabials (8
|
||
<emphasis id="16D78C36FF884B109053FA2CEFD6FA7A" box="[570,641,1485,1508]" italics="true" pageId="9" pageNumber="270">versus</emphasis>
|
||
9–10), anterior chinshields slightly longer than posterior ones (
|
||
<emphasis id="16D78C36FF884B10973DFA2CE8CCFA7A" box="[1364,1435,1485,1508]" italics="true" pageId="9" pageNumber="270">versus</emphasis>
|
||
anterior chinshields 2 to 3 times longer than posterior ones), cloacal single (
|
||
<emphasis id="16D78C36FF884B109187FA10E961F996" box="[1006,1078,1521,1544]" italics="true" pageId="9" pageNumber="270">versus</emphasis>
|
||
divided) (
|
||
<bibRefCitation id="40322DD5FF884B1096C0FA0EE8CFF996" author="Vogel, G. & David, P." box="[1193,1432,1519,1544]" pageId="9" pageNumber="270" pagination="289" refId="ref9990" refString="Vogel, G. & David, P. (2010) A new species of the genus Lycodon (Boie, 1826) from Yunnan Province, China (Serpentes: Colubridae). Bonn zoological Bulletin, 57 (2), 289 ‾ 296." type="journal article" year="2010">Vogel & David 2010</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="40322DD5FF884B1092FEF9F4EC33F9B2" author="Neang, T. & Hartmann, T. & Hun, S. & Souter, N. J. & Furey, N. M." box="[151,356,1556,1581]" pageId="9" pageNumber="270" pagination="1" refId="ref8967" refString="Neang, T., Hartmann, T., Hun, S., Souter, N. J. & Furey, N. M. (2014) A new species of wolf snake (Colubridae: Lycodon Fitzinger, 1826) from Phnom Samkos Wildlife Sanctuary, Cardamom Mountains, southwest Cambodia. Zootaxa, 3814 (1), 068 ‾ 080. https: // doi. org / 10.11646 / zootaxa. 3814.1.3" type="journal article" year="2014">
|
||
Neang
|
||
<emphasis id="16D78C36FF884B10928EF9F4EC4EF9B2" box="[231,281,1556,1580]" italics="true" pageId="9" pageNumber="270">et al</emphasis>
|
||
. 2014
|
||
</bibRefCitation>
|
||
); from
|
||
<taxonomicName id="E3A32BA7FF884B1093DCF9F4EF34F9B2" box="[437,611,1557,1580]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="270" phylum="Chordata" rank="species" species="rosozonatum">
|
||
<emphasis id="16D78C36FF884B1093DCF9F4EF34F9B2" box="[437,611,1557,1580]" italics="true" pageId="9" pageNumber="270">L. rosozonatum</emphasis>
|
||
</taxonomicName>
|
||
by having fewer dorsal scale rows at neck and at midbody (17–17
|
||
<emphasis id="16D78C36FF884B10973DF9F4E8CBF9B2" box="[1364,1436,1557,1580]" italics="true" pageId="9" pageNumber="270">versus</emphasis>
|
||
19–19, respectively), the vertebral scale row not enlarged (
|
||
<emphasis id="16D78C36FF884B109120F9D8EEC7F9CE" box="[841,912,1593,1616]" italics="true" pageId="9" pageNumber="270">versus</emphasis>
|
||
distinctly enlarged), dorsal scale rows being completely smooth (
|
||
<emphasis id="16D78C36FF884B1093EDF9BCEC9BF9EA" box="[388,460,1629,1652]" italics="true" pageId="9" pageNumber="270">versus</emphasis>
|
||
three to nine upper dorsal scale rows plus vertebral row weakly keeled), and fewer dorsal bands (23
|
||
<emphasis id="16D78C36FF884B10930DF960ECFBF906" box="[356,428,1665,1688]" italics="true" pageId="9" pageNumber="270">versus</emphasis>
|
||
28–35) (
|
||
<bibRefCitation id="40322DD5FF884B109073F961EF98F906" author="Hu, S. & Zhao, E. & Huang, Z." box="[538,719,1663,1688]" pageId="9" pageNumber="270" pagination="1 - 9" refId="ref8464" refString="Hu, S., Zhao, E. & Huang, Z. (1975) Three new species of reptiles from Hainan Island, Guangdong Province (In Chinese with English abstract). Acta zoologica Sinica, 21 (4), 379 - 384. [English translation of this paper was provided by Akihiro Koshikawa, 1982, Smithsonian herpetological Information Service, 53, 1 - 9]" type="journal article" year="1975">
|
||
Hu
|
||
<emphasis id="16D78C36FF884B109023F960EFD7F906" box="[586,640,1664,1688]" italics="true" pageId="9" pageNumber="270">et al</emphasis>
|
||
. 1975
|
||
</bibRefCitation>
|
||
); from
|
||
<taxonomicName id="E3A32BA7FF884B109140F961E903F906" box="[809,1108,1664,1688]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="270" phylum="Chordata" rank="subSpecies" species="ruhstrati" subSpecies="abditus">
|
||
<emphasis id="16D78C36FF884B109140F961E903F906" box="[809,1108,1664,1688]" italics="true" pageId="9" pageNumber="270">Lycodon ruhstrati abditus</emphasis>
|
||
</taxonomicName>
|
||
by dorsal scale rows being completely smooth (
|
||
<emphasis id="16D78C36FF884B1093E9F944EC90F922" box="[384,455,1701,1724]" italics="true" pageId="9" pageNumber="270">versus</emphasis>
|
||
three to five dorsal scale rows plus vertebral row distinctly keeled), fewer subcaudals (85
|
||
<emphasis id="16D78C36FF884B1092ADF928EC5BF97E" box="[196,268,1737,1760]" italics="true" pageId="9" pageNumber="270">versus</emphasis>
|
||
90–103), first body band starting at ventral scale 25 (
|
||
<emphasis id="16D78C36FF884B10911DF928EEEBF97E" box="[884,956,1737,1760]" italics="true" pageId="9" pageNumber="270">versus</emphasis>
|
||
12–17), and first body band covering one vertebral scale (
|
||
<emphasis id="16D78C36FF884B109327F90CECC2F89A" box="[334,405,1773,1796]" italics="true" pageId="9" pageNumber="270">versus</emphasis>
|
||
5–6) (
|
||
<bibRefCitation id="40322DD5FF884B10938BF90DEFFAF89A" author="Vogel, G. & David, P. & Pauwels, O. S. G. & Sumontha, M. & Norval, G. & Hendrix, R. & Vu, N. T. & Ziegler, T." box="[482,685,1772,1797]" pageId="9" pageNumber="270" pagination="131" refId="ref10038" refString="Vogel, G., David, P., Pauwels, O. S. G., Sumontha, M., Norval, G., Hendrix, R., Vu, N. T. & Ziegler, T. (2009) A revision of Lycodon ruhstrati (Fischer 1886) auctorum (Squamata Colubridae), with the description of a new species from Thailand and a new subspecies from the Asian mainland. Tropical Zoology, 22, 131 ‾ 182." type="journal article" year="2009">
|
||
Vogel
|
||
<emphasis id="16D78C36FF884B109047F90CEF35F89A" box="[558,610,1772,1796]" italics="true" pageId="9" pageNumber="270">et al</emphasis>
|
||
. 2009
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="40322DD5FF884B1090D2F90CEE29F89A" author="Luu, V. Q. & Nguyen, T. Q. & Calame, T. & Hoang, T. T. & Southichack, S. & Bonkowski, M. & Ziegler, T." box="[699,894,1772,1796]" pageId="9" pageNumber="270" refId="ref8782" refString="Luu, V. Q., Nguyen, T. Q., Calame, T., Hoang, T. T., Southichack, S., Bonkowski, M. & Ziegler, T. (2013 a) New country records of reptiles from Laos. Biodiversity Data Journal, 1, 1 ‾ 14. https: // doi. org / 10.3897 / BDJ. 1. e 1015" type="journal volume" year="2013">
|
||
Luu
|
||
<emphasis id="16D78C36FF884B109098F90CEE73F89A" box="[753,804,1772,1796]" italics="true" pageId="9" pageNumber="270">et al</emphasis>
|
||
. 2013a
|
||
</bibRefCitation>
|
||
); from
|
||
<taxonomicName id="E3A32BA7FF884B1091BAF90CE9D0F89A" authorityName="Neang, Hartmann, Hun, Souter & Furey" authorityYear="2014" box="[979,1159,1773,1796]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="270" phylum="Chordata" rank="species" species="zoosvictoriae">
|
||
<emphasis id="16D78C36FF884B1091BAF90CE9D0F89A" box="[979,1159,1773,1796]" italics="true" pageId="9" pageNumber="270">L. zoosvictoriae</emphasis>
|
||
</taxonomicName>
|
||
by having more ventral scales (218
|
||
<emphasis id="16D78C36FF884B10934BF8F0EC3DF8B6" box="[290,362,1809,1832]" italics="true" pageId="9" pageNumber="270">versus</emphasis>
|
||
213), dorsum banded (
|
||
<emphasis id="16D78C36FF884B109014F8F0EF92F8B6" box="[637,709,1809,1832]" italics="true" pageId="9" pageNumber="270">versus</emphasis>
|
||
blotched), dorsal scale rows being completely smooth (
|
||
<emphasis id="16D78C36FF884B10973DF8F0E8CBF8B6" box="[1364,1436,1809,1832]" italics="true" pageId="9" pageNumber="270">versus</emphasis>
|
||
weakly keeled), dorsal head whitish grey (
|
||
<emphasis id="16D78C36FF884B109015F8D4EF93F8D2" box="[636,708,1845,1868]" italics="true" pageId="9" pageNumber="270">versus</emphasis>
|
||
dark brown), dorsal tail banded (
|
||
<emphasis id="16D78C36FF884B10962DF8D4E9DBF8D2" box="[1092,1164,1845,1868]" italics="true" pageId="9" pageNumber="270">versus</emphasis>
|
||
blotched) (
|
||
<bibRefCitation id="40322DD5FF884B109766F8D5ED83F8EE" author="Neang, T. & Hartmann, T. & Hun, S. & Souter, N. J. & Furey, N. M." pageId="9" pageNumber="270" pagination="1" refId="ref8967" refString="Neang, T., Hartmann, T., Hun, S., Souter, N. J. & Furey, N. M. (2014) A new species of wolf snake (Colubridae: Lycodon Fitzinger, 1826) from Phnom Samkos Wildlife Sanctuary, Cardamom Mountains, southwest Cambodia. Zootaxa, 3814 (1), 068 ‾ 080. https: // doi. org / 10.11646 / zootaxa. 3814.1.3" type="journal article" year="2014">
|
||
Neang
|
||
<emphasis id="16D78C36FF884B10970BF8D4E8C2F8D2" box="[1378,1429,1844,1868]" italics="true" pageId="9" pageNumber="270">et al</emphasis>
|
||
. 2014
|
||
</bibRefCitation>
|
||
); from
|
||
<taxonomicName id="E3A32BA7FF884B109341F8B8ECB3F8EE" baseAuthorityName="Maki" baseAuthorityYear="1931" box="[296,484,1880,1904]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="270" phylum="Chordata" rank="species" species="multifasciatus">
|
||
<emphasis id="16D78C36FF884B109341F8B8ECB3F8EE" box="[296,484,1880,1904]" italics="true" pageId="9" pageNumber="270">L. multifasciatus</emphasis>
|
||
</taxonomicName>
|
||
by having fewer ventral scales (218
|
||
<emphasis id="16D78C36FF884B1091E7F8B8EE82F8EE" box="[910,981,1881,1904]" italics="true" pageId="9" pageNumber="270">versus</emphasis>
|
||
229–237), fewer subcaudals (85
|
||
<emphasis id="16D78C36FF884B10973DF8B8E8CCF8EE" box="[1364,1435,1881,1904]" italics="true" pageId="9" pageNumber="270">versus</emphasis>
|
||
<quantity id="E35BFDC1FF884B1092FEF89DEC43F80B" box="[151,276,1916,1941]" metricMagnitude="0" metricUnit="m" metricValue="2.8575" metricValueMax="3.0225999999999997" metricValueMin="2.6924" pageId="9" pageNumber="270" unit="in" value="112.5" valueMax="119.0" valueMin="106.0">106–119 in</quantity>
|
||
males), dorsal scale rows being completely smooth (
|
||
<emphasis id="16D78C36FF884B10910AF89CEEFCF80A" box="[867,939,1917,1940]" italics="true" pageId="9" pageNumber="270">versus</emphasis>
|
||
upper dorsal and vertebral rows keeled), tail bands (14
|
||
<emphasis id="16D78C36FF884B109365F840EC04F826" box="[268,339,1953,1976]" italics="true" pageId="9" pageNumber="270">versus</emphasis>
|
||
25–42), first body band starting at ventral scale 25 (
|
||
<emphasis id="16D78C36FF884B1091C8F840EEBFF826" box="[929,1000,1953,1976]" italics="true" pageId="9" pageNumber="270">versus</emphasis>
|
||
7–11), and first body band covered by one vertebral scale (
|
||
<emphasis id="16D78C36FF884B109315F824EC94F842" box="[380,451,1989,2012]" italics="true" pageId="9" pageNumber="270">versus</emphasis>
|
||
about 6) (
|
||
<bibRefCitation id="40322DD5FF884B109051F825EFEFF842" author="Maki, M." box="[568,696,1988,2013]" pageId="9" pageNumber="270" refId="ref8944" refString="Maki, M. (1931) A monograph of the snakes of Japan. Dai-ichi Bhobo, Tokyo, 247 pp." type="book" year="1931">Maki 1931</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="40322DD5FF884B1090ACF825EEDCF842" author="Vogel, G. & Nguyen, T. Q. & Kingsada, P. & Ziegler, T." box="[709,907,1988,2013]" pageId="9" pageNumber="270" pagination="344" refId="ref10187" refString="Vogel, G., Nguyen, T. Q., Kingsada, P. & Ziegler, T. (2012) A new species of the genus Lycodon Boie, 1826 from Laos (Squamata: Colubridae). North-Western Journal of Zoology, 8, 344 ‾ 352." type="journal article" year="2012">
|
||
Vogel
|
||
<emphasis id="16D78C36FF884B109164F824EE68F842" box="[781,831,1988,2012]" italics="true" pageId="9" pageNumber="270">et al</emphasis>
|
||
. 2012
|
||
</bibRefCitation>
|
||
); from
|
||
<taxonomicName id="E3A32BA7FF884B1091B4F824E9DAF842" authorityName="Vogel, David, Pauwels, Sumontha, Norval, Hendrix, Vu & Ziegler" authorityYear="2009" box="[989,1165,1988,2012]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="270" phylum="Chordata" rank="species" species="ophiophagus">
|
||
<emphasis id="16D78C36FF884B1091B4F824E9DAF842" box="[989,1165,1988,2012]" italics="true" pageId="9" pageNumber="270">L. ophiophagus</emphasis>
|
||
</taxonomicName>
|
||
by having more ventral scales (218
|
||
<emphasis id="16D78C36FF8B4B139376FF78EC30FF2E" box="[287,359,153,176]" italics="true" pageId="10" pageNumber="271">versus</emphasis>
|
||
211–212), fewer subcaudals (85
|
||
<emphasis id="16D78C36FF8B4B13908FFF78EE7AFF2E" box="[742,813,153,176]" italics="true" pageId="10" pageNumber="271">versus</emphasis>
|
||
87–90), and first body band covered by one vertebral scale (
|
||
<emphasis id="16D78C36FF8B4B139289FF5CEC7FFF4A" box="[224,296,189,212]" italics="true" pageId="10" pageNumber="271">versus</emphasis>
|
||
about 8) (
|
||
<bibRefCitation id="40322DD5FF8B4B1393F7FF5DEF31FF4A" author="Vogel, G. & David, P. & Pauwels, O. S. G. & Sumontha, M. & Norval, G. & Hendrix, R. & Vu, N. T. & Ziegler, T." box="[414,614,188,213]" pageId="10" pageNumber="271" pagination="131" refId="ref10038" refString="Vogel, G., David, P., Pauwels, O. S. G., Sumontha, M., Norval, G., Hendrix, R., Vu, N. T. & Ziegler, T. (2009) A revision of Lycodon ruhstrati (Fischer 1886) auctorum (Squamata Colubridae), with the description of a new species from Thailand and a new subspecies from the Asian mainland. Tropical Zoology, 22, 131 ‾ 182." type="journal article" year="2009">
|
||
Vogel
|
||
<emphasis id="16D78C36FF8B4B139380FF5CEF4DFF4A" box="[489,538,188,212]" italics="true" pageId="10" pageNumber="271">et al</emphasis>
|
||
. 2009
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="40322DD5FF8B4B13901AFF5DEE6DFF4A" author="Vogel, G. & Nguyen, T. Q. & Kingsada, P. & Ziegler, T." box="[627,826,188,213]" pageId="10" pageNumber="271" pagination="344" refId="ref10187" refString="Vogel, G., Nguyen, T. Q., Kingsada, P. & Ziegler, T. (2012) A new species of the genus Lycodon Boie, 1826 from Laos (Squamata: Colubridae). North-Western Journal of Zoology, 8, 344 ‾ 352." type="journal article" year="2012">
|
||
Vogel
|
||
<emphasis id="16D78C36FF8B4B1390D5FF5CEFB9FF4A" box="[700,750,188,212]" italics="true" pageId="10" pageNumber="271">et al</emphasis>
|
||
. 2012
|
||
</bibRefCitation>
|
||
); from
|
||
<taxonomicName id="E3A32BA7FF8B4B1391E4FF5CE919FF4A" box="[909,1102,188,212]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="271" phylum="Chordata" rank="species" species="paucifasciatus">
|
||
<emphasis id="16D78C36FF8B4B1391E4FF5CE919FF4A" box="[909,1102,188,212]" italics="true" pageId="10" pageNumber="271">L. paucifasciatus</emphasis>
|
||
</taxonomicName>
|
||
by having fewer dorsal scale rows at neck (17
|
||
<emphasis id="16D78C36FF8B4B13930AFF00ECFDFF66" box="[355,426,225,248]" italics="true" pageId="10" pageNumber="271">versus</emphasis>
|
||
19), dorsal scale rows being completely smooth (
|
||
<emphasis id="16D78C36FF8B4B139186FF00E960FF66" box="[1007,1079,225,248]" italics="true" pageId="10" pageNumber="271">versus</emphasis>
|
||
upper rows plus vertebral row keeled), fewer subcaudals (85
|
||
<emphasis id="16D78C36FF8B4B139387FEE4EF61FE82" box="[494,566,261,284]" italics="true" pageId="10" pageNumber="271">versus</emphasis>
|
||
<quantity id="E35BFDC1FF8B4B139054FEE5EF20FE83" box="[573,631,260,285]" metricMagnitude="0" metricUnit="m" metricValue="2.3368" pageId="10" pageNumber="271" unit="in" value="92.0">92 in</quantity>
|
||
males), first body band starting at ventral scale 25 (
|
||
<emphasis id="16D78C36FF8B4B1396ABFEE4E85EFE82" box="[1218,1289,261,284]" italics="true" pageId="10" pageNumber="271">versus</emphasis>
|
||
10–15), first body band covered by one vertebral scale (
|
||
<emphasis id="16D78C36FF8B4B139011FEC8EF97FEDE" box="[632,704,297,320]" italics="true" pageId="10" pageNumber="271">versus</emphasis>
|
||
9.5–11), and more light tail bands (14
|
||
<emphasis id="16D78C36FF8B4B13961AFEC8E9EDFEDE" box="[1139,1210,297,320]" italics="true" pageId="10" pageNumber="271">versus</emphasis>
|
||
8–11) (
|
||
<bibRefCitation id="40322DD5FF8B4B139779FEC6E8C0FEDE" author="Smith, M. A." box="[1296,1431,295,320]" pageId="10" pageNumber="271" refId="ref9742" refString="Smith, M. A. (1943) The fauna of British India, Ceylon and Burma, including the whole of the Indo-chinese subregion. Reptilia and Amphibia. Fol. III. Serpentes. Taylor & Francis, London, xii + 583 pp." type="book" year="1943">Smith 1943</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="40322DD5FF8B4B1392FEFEADEC0DFEFA" author="Vogel, G. & David, P. & Pauwels, O. S. G. & Sumontha, M. & Norval, G. & Hendrix, R. & Vu, N. T. & Ziegler, T." box="[151,346,332,357]" pageId="10" pageNumber="271" pagination="131" refId="ref10038" refString="Vogel, G., David, P., Pauwels, O. S. G., Sumontha, M., Norval, G., Hendrix, R., Vu, N. T. & Ziegler, T. (2009) A revision of Lycodon ruhstrati (Fischer 1886) auctorum (Squamata Colubridae), with the description of a new species from Thailand and a new subspecies from the Asian mainland. Tropical Zoology, 22, 131 ‾ 182." type="journal article" year="2009">
|
||
Vogel
|
||
<emphasis id="16D78C36FF8B4B1392B6FEACEC46FEFA" box="[223,273,332,356]" italics="true" pageId="10" pageNumber="271">et al</emphasis>
|
||
. 2009
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="40322DD5FF8B4B13930FFEADEF7DFEFA" author="Vogel, G. & Nguyen, T. Q. & Kingsada, P. & Ziegler, T." box="[358,554,332,357]" pageId="10" pageNumber="271" pagination="344" refId="ref10187" refString="Vogel, G., Nguyen, T. Q., Kingsada, P. & Ziegler, T. (2012) A new species of the genus Lycodon Boie, 1826 from Laos (Squamata: Colubridae). North-Western Journal of Zoology, 8, 344 ‾ 352." type="journal article" year="2012">
|
||
Vogel
|
||
<emphasis id="16D78C36FF8B4B1393C6FEACECB6FEFA" box="[431,481,332,356]" italics="true" pageId="10" pageNumber="271">et al</emphasis>
|
||
. 2012
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="40322DD5FF8B4B13905FFEACEE53FEFA" author="Neang, T. & Hartmann, T. & Hun, S. & Souter, N. J. & Furey, N. M." box="[566,772,332,357]" pageId="10" pageNumber="271" pagination="1" refId="ref8967" refString="Neang, T., Hartmann, T., Hun, S., Souter, N. J. & Furey, N. M. (2014) A new species of wolf snake (Colubridae: Lycodon Fitzinger, 1826) from Phnom Samkos Wildlife Sanctuary, Cardamom Mountains, southwest Cambodia. Zootaxa, 3814 (1), 068 ‾ 080. https: // doi. org / 10.11646 / zootaxa. 3814.1.3" type="journal article" year="2014">
|
||
Neang
|
||
<emphasis id="16D78C36FF8B4B1390EEFEACEFEFFEFA" box="[647,696,332,356]" italics="true" pageId="10" pageNumber="271">et al</emphasis>
|
||
. 2014
|
||
</bibRefCitation>
|
||
); from
|
||
<taxonomicName id="E3A32BA7FF8B4B13913FFEACEE81FEFA" authorityName="Vogel & David" authorityYear="2010" box="[854,982,333,356]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="271" phylum="Chordata" rank="species" species="synaptor">
|
||
<emphasis id="16D78C36FF8B4B13913FFEACEE81FEFA" box="[854,982,333,356]" italics="true" pageId="10" pageNumber="271">L. synaptor</emphasis>
|
||
</taxonomicName>
|
||
by having more ventral scale rows (218
|
||
<emphasis id="16D78C36FF8B4B1392FEFE90ED88FE16" box="[151,223,369,392]" italics="true" pageId="10" pageNumber="271">versus</emphasis>
|
||
201–203), fewer light bands on dorsum (23
|
||
<emphasis id="16D78C36FF8B4B1390B3FE90EE75FE16" box="[730,802,369,392]" italics="true" pageId="10" pageNumber="271">versus</emphasis>
|
||
30–31), more subcaudals (85
|
||
<emphasis id="16D78C36FF8B4B139610FE90E996FE16" box="[1145,1217,369,392]" italics="true" pageId="10" pageNumber="271">versus</emphasis>
|
||
68–69), and dorsal scale rows being completely smooth (
|
||
<emphasis id="16D78C36FF8B4B139056FE74EFD1FE32" box="[575,646,405,428]" italics="true" pageId="10" pageNumber="271">versus</emphasis>
|
||
6–7 dorsal scale rows keeled) (
|
||
<bibRefCitation id="40322DD5FF8B4B13918EFE75E984FE32" author="Vogel, G. & David, P." box="[999,1235,404,429]" pageId="10" pageNumber="271" pagination="289" refId="ref9990" refString="Vogel, G. & David, P. (2010) A new species of the genus Lycodon (Boie, 1826) from Yunnan Province, China (Serpentes: Colubridae). Bonn zoological Bulletin, 57 (2), 289 ‾ 296." type="journal article" year="2010">Vogel & David 2010</bibRefCitation>
|
||
) (for more details see
|
||
<tableCitation id="6921659FFF8B4B1392ABFE56EC44FE4E" box="[194,275,439,464]" captionStart="TABLE 1" captionStartId="1.[147,231,574,596]" captionTargetPageId="1" captionText="TABLE 1. Taxonomic review of Lycodon members from Vietnam (data obtained from Boulenger 1893; Smith 1943; Deuve 1970; Hu et al. 1975; Ziegler et al. 2007; Nguyen et al. 2009; Vogel & David 2010; Geissler et al. 2011; Orlov et al. 2011; Neang et al. 2014; Luu et al. 2013a,b; Jestrzemski et al. 2013; Phan et al. 2014 Grismer et al. 2014; Ziegler et al. 2014; Gawor et al. 2016; Ngo & Duong 2016; Nguyen et al. 2016; Do et al. 2016; Vassilieva et al. 2016; Pham et al. 2017; Do et al. 2017; Luu et al. 2018). 1—maximum snout-vent length (in mm); 2—maximum tail length; 3—maximum of total length; 4—number of supralabials; 5—number of infralabials; 6—loreal touching orbit; 7— number of preoculars; 8—number of postoculars; 9—number of loreals; 10—dorsal scale row fomula; 11—number of ventral scales; 12—number of subcaudals; 13—number of cloacal (1 or 2); 14—head pattern; 15—belly banded; 16—dorsal body pattern; 17—number of body bands/blotches; 18—number of tail bands/blotches; 19—color of light body bands/blotches; 20— ground coloration on dorsum; 21— dorsal scales keeled/smooth; 22—belly pattern; 23— vertebral scale row enlarged; 24— first body band position (at VEN); 25—first body band width; 26—ventral tail banded; 27—maxillary teeth, anterior=A, posterior=P; 28—temporals (A+P); 29—nuchal band; 30—distribution. “1” = presence of character state; “0” = absence of character state; “̅” = missing data; “juv.” = juvenile; “*” = this study." httpUri="http://table.plazi.org/id/70DC00ACFF804B1892FAFDDFE85AFCA6" pageId="10" pageNumber="271" tableUuid="70DC00ACFF804B1892FAFDDFE85AFCA6">Table 1</tableCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="241C5024FF8B4B1392AEFE3DEF90FCA6" blockId="10.[151,1436,151,1151]" pageId="10" pageNumber="271">
|
||
The new species has a loreal which is not in contact with the orbit and thus differs from the following species of the
|
||
<taxonomicName id="E3A32BA7FF8B4B13928FFDE1ECE2FD86" baseAuthorityName="Anderson" baseAuthorityYear="1879" box="[230,437,512,536]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="271" phylum="Chordata" rank="species" species="fasciatus">
|
||
<emphasis id="16D78C36FF8B4B13928FFDE1ECE2FD86" box="[230,437,512,536]" italics="true" pageId="10" pageNumber="271">Lycodon fasciatus</emphasis>
|
||
</taxonomicName>
|
||
group which have the loreal entering the orbit:
|
||
<taxonomicName id="E3A32BA7FF8B4B139186FDE0E989FD86" authority="Boulenger" authorityName="Boulenger" authorityYear="1900" box="[1007,1246,511,536]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="271" phylum="Chordata" rank="species" species="butleri">
|
||
<emphasis id="16D78C36FF8B4B139186FDE0E90CFD86" box="[1007,1115,512,536]" italics="true" pageId="10" pageNumber="271">L. butleri</emphasis>
|
||
Boulenger
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="E3A32BA7FF8B4B139684FDE0E824FD86" baseAuthorityName="Anderson" baseAuthorityYear="1879" box="[1261,1395,512,536]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="271" phylum="Chordata" rank="species" species="fasciatus">
|
||
<emphasis id="16D78C36FF8B4B139684FDE0E824FD86" box="[1261,1395,512,536]" italics="true" pageId="10" pageNumber="271">L. fasciatus</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="E3A32BA7FF8B4B1397ECFDE0EC5AFDA2" authorityName="Boie" authorityYear="1827" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="271" phylum="Chordata" rank="species" species="subcinctus">
|
||
<emphasis id="16D78C36FF8B4B1397ECFDE0EC5AFDA2" italics="true" pageId="10" pageNumber="271">L. subcinctus</emphasis>
|
||
</taxonomicName>
|
||
. In addition, the new species differs from
|
||
<taxonomicName id="E3A32BA7FF8B4B139091FDC4EE35FDA2" authorityName="Boulenger" authorityYear="1900" box="[760,866,548,572]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="271" phylum="Chordata" rank="species" species="butleri">
|
||
<emphasis id="16D78C36FF8B4B139091FDC4EE35FDA2" box="[760,866,548,572]" italics="true" pageId="10" pageNumber="271">L. butleri</emphasis>
|
||
</taxonomicName>
|
||
by having fewer dorsal bands (23
|
||
<emphasis id="16D78C36FF8B4B13969CFDC4E86AFDA2" box="[1269,1341,549,572]" italics="true" pageId="10" pageNumber="271">versus</emphasis>
|
||
28–37), fewer tail bands (14
|
||
<emphasis id="16D78C36FF8B4B1393EDFDA8EC9BFDFE" box="[388,460,585,608]" italics="true" pageId="10" pageNumber="271">versus</emphasis>
|
||
17–23), belly pattern mostly cream (
|
||
<emphasis id="16D78C36FF8B4B13911EFDA8EEE9FDFE" box="[887,958,585,608]" italics="true" pageId="10" pageNumber="271">versus</emphasis>
|
||
banded & spotted) (
|
||
<bibRefCitation id="40322DD5FF8B4B1396C3FDA6E8C7FDFE" author="Grismer, L. L. & Quah, E. S. H. & Anuar, S. & Muin, M. A. & Wood, P. L. Jr. & Aziza, S." box="[1194,1424,583,608]" pageId="10" pageNumber="271" pagination="1" refId="ref8295" refString="Grismer, L. L. Quah, E. S. H., Anuar, S., Muin, M. A., Wood, P. L. Jr. & Aziza, S. (2014) A diminutive new species of cavedwelling Wolf Snake (Colubridae: Lycodon Boie, 1826) from Peninsular Malaysia. Zootaxa, 3815 (1), 051 ‾ 067. https: // doi. org / 10.11646 / zootaxa. 3815.1.3" type="journal article" year="2014">
|
||
Grismer
|
||
<emphasis id="16D78C36FF8B4B139779FDA8E814FDFE" box="[1296,1347,584,608]" italics="true" pageId="10" pageNumber="271">et al</emphasis>
|
||
. 2014
|
||
</bibRefCitation>
|
||
); from
|
||
<taxonomicName id="E3A32BA7FF8B4B1392BDFD8CEC02FD1A" baseAuthorityName="Anderson" baseAuthorityYear="1879" box="[212,341,620,644]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="271" phylum="Chordata" rank="species" species="fasciatus">
|
||
<emphasis id="16D78C36FF8B4B1392BDFD8CEC02FD1A" box="[212,341,620,644]" italics="true" pageId="10" pageNumber="271">L. fasciatus</emphasis>
|
||
</taxonomicName>
|
||
by having dorsal scale rows being completely smooth (
|
||
<emphasis id="16D78C36FF8B4B1391ABFD8CE95EFD1A" box="[962,1033,621,644]" italics="true" pageId="10" pageNumber="271">versus</emphasis>
|
||
weakly keeled), dorsal head pattern whitish grey (
|
||
<emphasis id="16D78C36FF8B4B13935FFD70EC2AFD36" box="[310,381,657,680]" italics="true" pageId="10" pageNumber="271">versus</emphasis>
|
||
dark), first body band starting at ventral scale 25 (
|
||
<emphasis id="16D78C36FF8B4B1391AEFD70E958FD36" box="[967,1039,657,680]" italics="true" pageId="10" pageNumber="271">versus</emphasis>
|
||
5–18), first body band covered by one vertebral scale (
|
||
<emphasis id="16D78C36FF8B4B139311FD54ECE8FD52" box="[376,447,693,716]" italics="true" pageId="10" pageNumber="271">versus</emphasis>
|
||
3–12), ventral pattern mostly cream (
|
||
<emphasis id="16D78C36FF8B4B13910FFD54EEFAFD52" box="[870,941,693,716]" italics="true" pageId="10" pageNumber="271">versus</emphasis>
|
||
banded) (
|
||
<bibRefCitation id="40322DD5FF8B4B139675FD55E85EFD52" author="Vogel, G. & David, P." box="[1052,1289,692,717]" pageId="10" pageNumber="271" pagination="289" refId="ref9990" refString="Vogel, G. & David, P. (2010) A new species of the genus Lycodon (Boie, 1826) from Yunnan Province, China (Serpentes: Colubridae). Bonn zoological Bulletin, 57 (2), 289 ‾ 296." type="journal article" year="2010">Vogel & David 2010</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="40322DD5FF8B4B13977DFD54ED83FD6E" author="Neang, T. & Hartmann, T. & Hun, S. & Souter, N. J. & Furey, N. M." pageId="10" pageNumber="271" pagination="1" refId="ref8967" refString="Neang, T., Hartmann, T., Hun, S., Souter, N. J. & Furey, N. M. (2014) A new species of wolf snake (Colubridae: Lycodon Fitzinger, 1826) from Phnom Samkos Wildlife Sanctuary, Cardamom Mountains, southwest Cambodia. Zootaxa, 3814 (1), 068 ‾ 080. https: // doi. org / 10.11646 / zootaxa. 3814.1.3" type="journal article" year="2014">
|
||
Neang
|
||
<emphasis id="16D78C36FF8B4B13970DFD54E8C2FD52" box="[1380,1429,692,716]" italics="true" pageId="10" pageNumber="271">et al</emphasis>
|
||
. 2014
|
||
</bibRefCitation>
|
||
); from
|
||
<taxonomicName id="E3A32BA7FF8B4B139342FD38EC95FD6E" authorityName="Boie" authorityYear="1827" box="[299,450,728,752]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="271" phylum="Chordata" rank="species" species="subcinctus">
|
||
<emphasis id="16D78C36FF8B4B139342FD38EC95FD6E" box="[299,450,728,752]" italics="true" pageId="10" pageNumber="271">L. subcinctus</emphasis>
|
||
</taxonomicName>
|
||
by the presence of a preocular scale (
|
||
<emphasis id="16D78C36FF8B4B1391EBFD38EE9EFD6E" box="[898,969,729,752]" italics="true" pageId="10" pageNumber="271">versus</emphasis>
|
||
absent), having dorsal scale rows being completely smooth (
|
||
<emphasis id="16D78C36FF8B4B139314FD1CEC92FC8A" box="[381,453,765,788]" italics="true" pageId="10" pageNumber="271">versus</emphasis>
|
||
weakly keeled), single cloacal (
|
||
<emphasis id="16D78C36FF8B4B139147FD1CEE21FC8A" box="[814,886,765,788]" italics="true" pageId="10" pageNumber="271">versus</emphasis>
|
||
divided), and more body bands (23
|
||
<emphasis id="16D78C36FF8B4B139765FD1CE803FC8A" box="[1292,1364,765,788]" italics="true" pageId="10" pageNumber="271">versus</emphasis>
|
||
9–15) (
|
||
<bibRefCitation id="40322DD5FF8B4B1392F7FCC1EC3EFCA6" author="Neang, T. & Hartmann, T. & Hun, S. & Souter, N. J. & Furey, N. M." box="[158,361,800,824]" pageId="10" pageNumber="271" pagination="1" refId="ref8967" refString="Neang, T., Hartmann, T., Hun, S., Souter, N. J. & Furey, N. M. (2014) A new species of wolf snake (Colubridae: Lycodon Fitzinger, 1826) from Phnom Samkos Wildlife Sanctuary, Cardamom Mountains, southwest Cambodia. Zootaxa, 3814 (1), 068 ‾ 080. https: // doi. org / 10.11646 / zootaxa. 3814.1.3" type="journal article" year="2014">
|
||
Neang
|
||
<emphasis id="16D78C36FF8B4B139299FCC0EC77FCA6" box="[240,288,800,824]" italics="true" pageId="10" pageNumber="271">et al</emphasis>
|
||
. 2014
|
||
</bibRefCitation>
|
||
) (for more details see
|
||
<tableCitation id="6921659FFF8B4B13900DFCFEEFE2FCA6" box="[612,693,799,824]" captionStart="TABLE 1" captionStartId="1.[147,231,574,596]" captionTargetPageId="1" captionText="TABLE 1. Taxonomic review of Lycodon members from Vietnam (data obtained from Boulenger 1893; Smith 1943; Deuve 1970; Hu et al. 1975; Ziegler et al. 2007; Nguyen et al. 2009; Vogel & David 2010; Geissler et al. 2011; Orlov et al. 2011; Neang et al. 2014; Luu et al. 2013a,b; Jestrzemski et al. 2013; Phan et al. 2014 Grismer et al. 2014; Ziegler et al. 2014; Gawor et al. 2016; Ngo & Duong 2016; Nguyen et al. 2016; Do et al. 2016; Vassilieva et al. 2016; Pham et al. 2017; Do et al. 2017; Luu et al. 2018). 1—maximum snout-vent length (in mm); 2—maximum tail length; 3—maximum of total length; 4—number of supralabials; 5—number of infralabials; 6—loreal touching orbit; 7— number of preoculars; 8—number of postoculars; 9—number of loreals; 10—dorsal scale row fomula; 11—number of ventral scales; 12—number of subcaudals; 13—number of cloacal (1 or 2); 14—head pattern; 15—belly banded; 16—dorsal body pattern; 17—number of body bands/blotches; 18—number of tail bands/blotches; 19—color of light body bands/blotches; 20— ground coloration on dorsum; 21— dorsal scales keeled/smooth; 22—belly pattern; 23— vertebral scale row enlarged; 24— first body band position (at VEN); 25—first body band width; 26—ventral tail banded; 27—maxillary teeth, anterior=A, posterior=P; 28—temporals (A+P); 29—nuchal band; 30—distribution. “1” = presence of character state; “0” = absence of character state; “̅” = missing data; “juv.” = juvenile; “*” = this study." httpUri="http://table.plazi.org/id/70DC00ACFF804B1892FAFDDFE85AFCA6" pageId="10" pageNumber="271" tableUuid="70DC00ACFF804B1892FAFDDFE85AFCA6">Table 1</tableCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="6CB903AFFF8B4B1392AEFCA5ECCCFC3B" pageId="10" pageNumber="271" type="distribution">
|
||
<paragraph id="241C5024FF8B4B1392AEFCA5ECCCFC3B" blockId="10.[151,1436,151,1151]" pageId="10" pageNumber="271">
|
||
<emphasis id="16D78C36FF8B4B1392AEFCA5EC35FCC3" bold="true" box="[199,354,836,861]" pageId="10" pageNumber="271">Distribution:</emphasis>
|
||
<taxonomicName id="E3A32BA7FF8B4B139300FCA5EF38FCC2" authority="Luu & Ziegler & Ha & Le & Hoang, 2019" authorityName="Luu & Ziegler & Ha & Le & Hoang" authorityYear="2019" box="[361,623,836,860]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="271" phylum="Chordata" rank="species" species="namdongensis" status="sp. nov.">
|
||
<emphasis id="16D78C36FF8B4B139300FCA5EF38FCC2" box="[361,623,836,860]" italics="true" pageId="10" pageNumber="271">Lycodon namdongensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="16D78C36FF8B4B13901CFCA5EF99FCC3" bold="true" box="[629,718,836,861]" pageId="10" pageNumber="271">
|
||
<taxonomicNameLabel id="0DE4314DFF8B4B13901CFCA5EF99FCC3" box="[629,718,836,861]" pageId="10" pageNumber="271" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is currently known only from the
|
||
<typeStatus id="FB18EE86FF8B4B139622FCA4E92CFCC3" box="[1099,1147,837,861]" pageId="10" pageNumber="271">type</typeStatus>
|
||
locality in the Nam Dong NR,
|
||
<collectingRegion id="E6679EC6FF8B4B1392A5FC86ECEFFC1E" box="[204,440,871,896]" country="Vietnam" name="Thanh Hoa" pageId="10" pageNumber="271">Thanh Hoa Province</collectingRegion>
|
||
,
|
||
<collectingCountry id="5CB410B4FF8B4B1393ACFC86EF7FFC1E" box="[453,552,871,896]" name="Vietnam" pageId="10" pageNumber="271">Vietnam</collectingCountry>
|
||
. Distribution of the new species and other
|
||
<taxonomicName id="E3A32BA7FF8B4B139665FC89E93BFC1E" box="[1036,1132,872,896]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="271" phylum="Chordata" rank="genus">
|
||
<emphasis id="16D78C36FF8B4B139665FC89E93BFC1E" box="[1036,1132,872,896]" italics="true" pageId="10" pageNumber="271">Lycodon</emphasis>
|
||
</taxonomicName>
|
||
species from
|
||
<collectingCountry id="5CB410B4FF8B4B139763FC86E83EFC1E" box="[1290,1385,871,896]" name="Vietnam" pageId="10" pageNumber="271">Vietnam</collectingCountry>
|
||
was shown in
|
||
<figureCitation id="BC984CA1FF8B4B13936DFC6DEC07FC3B" box="[260,336,908,933]" captionStart="FIGURE 5" captionStartId="11.[151,250,1787,1809]" captionTargetBox="[170,1427,193,1760]" captionTargetId="figure@11.[152,1435,193,1766]" captionTargetPageId="11" captionText="FIGURE 5. Map showing the records of the Lycodon fasciatus group in Vietnam (Smith 1943; Nguyen et al. 2009; Geissler et al. 2011; Orlov et al. 2011; Jestrzemski et al. 2013; Phan et al. 2014; Do et al. 2016; Vassilieva et al. 2016)." httpUri="https://zenodo.org/record/2644476/files/figure.png" pageId="10" pageNumber="271">Figs. 5</figureCitation>
|
||
and
|
||
<figureCitation id="BC984CA1FF8B4B1393EFFC6DECC0FC3B" box="[390,407,908,933]" captionStart="FIGURE 6" captionStartId="12.[151,250,1792,1814]" captionTargetBox="[170,1427,197,1765]" captionTargetId="figure@12.[152,1435,193,1771]" captionTargetPageId="12" captionText="FIGURE 6. Map showing the type locality of Lycodon namdongensis sp. nov. in Thanh Hoa Province (yellow cross) and other records of the Lycodon ruhstrati group from in Vietnam (Boulenger 1893; Smith 1943; Deuve 1970; Hu et al. 1975; Ziegler et al. 2007; Nguyen et al. 2009; Vogel & David 2010; Geissler et al. 2011; Orlov et al. 2011; Neang et al. 2014; Luu et al. 2013a,b; Jestrzemski et al. 2013; Phan et al. 2014; Grismer et al. 2014; Ziegler et al. 2014; Gawor et al. 2016; Ngo & Duong 2016; Nguyen et al. 2016; Do et al. 2016; Vassilieva et al. 2016; Pham et al. 2017; Do et al. 2017; Luu et al. 2018)." httpUri="https://zenodo.org/record/2644478/files/figure.png" pageId="10" pageNumber="271">6</figureCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="6CB903AFFF8B4B1392AEFC4EEEE6FBE1" pageId="10" pageNumber="271" type="etymology">
|
||
<paragraph id="241C5024FF8B4B1392AEFC4EEFFBFB8E" blockId="10.[151,1436,151,1151]" pageId="10" pageNumber="271">
|
||
<emphasis id="16D78C36FF8B4B1392AEFC4EEC06FC56" bold="true" box="[199,337,943,968]" pageId="10" pageNumber="271">Etymology:</emphasis>
|
||
We name this species after its
|
||
<typeStatus id="FB18EE86FF8B4B1390BEFC51EE50FC56" box="[727,775,944,968]" pageId="10" pageNumber="271">type</typeStatus>
|
||
locality, Nam Dong NR, where the new
|
||
<taxonomicName id="E3A32BA7FF8B4B13976AFC51E834FC56" box="[1283,1379,944,968]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="271" phylum="Chordata" rank="genus">
|
||
<emphasis id="16D78C36FF8B4B13976AFC51E834FC56" box="[1283,1379,944,968]" italics="true" pageId="10" pageNumber="271">Lycodon</emphasis>
|
||
</taxonomicName>
|
||
was discovered and propose the following common names: Namdong Wolf Snake (English), Namdong Wolfszahnnatter (German), Rǻn khuyḗt nam động (Vietnamese).
|
||
</paragraph>
|
||
<paragraph id="241C5024FF8B4B1392AEFBFDEEE6FBE1" blockId="10.[151,1436,151,1151]" pageId="10" pageNumber="271">
|
||
<emphasis id="16D78C36FF8B4B1392AEFBFDECDEFBAB" bold="true" box="[199,393,1052,1077]" pageId="10" pageNumber="271">Natural history:</emphasis>
|
||
The
|
||
<typeStatus id="FB18EE86FF8B4B1393AAFBFDEF72FBAB" box="[451,549,1052,1077]" pageId="10" pageNumber="271" type="holotype">holotype</typeStatus>
|
||
was collected at 7:54 PM, while crawling on the forest path, at an elevation of
|
||
<quantity id="E35BFDC1FF8B4B1392FEFBDEEDB6FBC9" box="[151,225,1087,1112]" metricMagnitude="2" metricUnit="m" metricValue="6.16" pageId="10" pageNumber="271" unit="m" value="616.0">616 m</quantity>
|
||
a.s.l. The surrounding habitat was shrubs in secondary karst forest (
|
||
<figureCitation id="BC984CA1FF8B4B139192FBDEE915FBC6" box="[1019,1090,1087,1112]" captionStart="FIGURE 7" captionStartId="13.[151,250,732,754]" captionTargetBox="[151,1435,240,710]" captionTargetId="figure@13.[151,1435,193,710]" captionTargetPageId="13" captionText="FIGURE 7. Macrohabitat of Lycodon namdongensis sp. nov. at the type locality. Photo N. V. Ha" httpUri="https://zenodo.org/record/2644480/files/figure.png" pageId="10" pageNumber="271">Fig. 7</figureCitation>
|
||
). The humidity at the time of collection was approximately 66% and the air temperature was 28.4
|
||
<superScript id="D3D6FD6CFF8B4B1391E6FB82EEC1FBEF" attach="right" box="[911,918,1123,1137]" fontSize="6" pageId="10" pageNumber="271">o</superScript>
|
||
C.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |