185 lines
20 KiB
XML
185 lines
20 KiB
XML
<document id="BAF52DFC99FBC94B883F6E26C3F4198B" ID-DOI="10.11646/zootaxa.3949.3.3" ID-GBIF-Dataset="25ece91c-6566-4122-a406-236f147b58b1" ID-ISSN="1175-5326" ID-Zenodo-Dep="238900" ID-ZooBank="CBEF804D-5FBF-48D2-8CCD-036B70FE6ECD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461046281391" checkinUser="plazi" docAuthor="Arias, Andrés & Paxton, Hannelore" docDate="2015" docId="03BA8797FFBAFFA0FF2BFD37FE3DC546" docLanguage="en" docName="zt03949p369.pdf" docOrigin="Zootaxa 3949 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Onuphis eremita Audouin & Milne Edwards 1883" docType="treatment" docVersion="15" lastPageNumber="354" masterDocId="FF83FFEFFFB3FFA9FFBCFFE0FFA7C254" masterDocTitle="Onuphis and Aponuphis (Annelida: Onuphidae) from southwestern Europe, with the description of a new species" masterLastPageNumber="369" masterPageNumber="345" pageNumber="354" updateTime="1698577207772" updateUser="plazi">
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<mods:title id="D9EFE6A23EBCACFC76805925492DD435">Onuphis and Aponuphis (Annelida: Onuphidae) from southwestern Europe, with the description of a new species</mods:title>
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<mods:namePart id="DB704A4C8A2CC18E18381F8A54929C11">Arias, Andrés</mods:namePart>
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<mods:namePart id="4587A9479D7FE00FB0B373AF00E2277E">Paxton, Hannelore</mods:namePart>
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<mods:typeOfResource id="68BB504D26FBC7F2497A7DC9657BD60B">text</mods:typeOfResource>
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<mods:title id="73E3B499F56F3949E6A6C9FAB3A2CF75">Zootaxa</mods:title>
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<mods:date id="829771BB48E09EC72E7EE85FB7FFE289">2015</mods:date>
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<mods:number id="85BB06A0233DB43066B589A32BB0231D">3949</mods:number>
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<treatment id="03BA8797FFBAFFA0FF2BFD37FE3DC546" ID-DOI="http://doi.org/10.5281/zenodo.3513478" ID-GBIF-Taxon="119610572" ID-Zenodo-Dep="3513478" LSID="urn:lsid:plazi:treatment:03BA8797FFBAFFA0FF2BFD37FE3DC546" httpUri="http://treatment.plazi.org/id/03BA8797FFBAFFA0FF2BFD37FE3DC546" lastPageNumber="354" pageId="9" pageNumber="354">
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<subSubSection id="C309650AFFBAFFA0FF2BFD37FCA0C0A6" box="[151,775,727,754]" pageId="9" pageNumber="354" type="nomenclature">
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<paragraph id="8BAC3681FFBAFFA0FF2BFD37FCA0C0A6" blockId="9.[151,775,727,754]" box="[151,775,727,754]" pageId="9" pageNumber="354">
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<heading id="D0E481EDFFBAFFA0FF2BFD37FCA0C0A6" bold="true" box="[151,775,727,754]" fontSize="11" level="1" pageId="9" pageNumber="354" reason="1">
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<taxonomicName id="4C134D02FFBAFFA0FF2BFD37FCA0C0A6" ID-CoL="49QJ7" authority="Audouin & Milne Edwards, 1883" authorityName="Audouin & Milne Edwards" authorityYear="1883" box="[151,775,727,754]" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="9" pageNumber="354" phylum="Annelida" rank="species" species="eremita">
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<emphasis id="B967EA93FFBAFFA0FF2BFD37FCA0C0A6" bold="true" box="[151,775,727,754]" pageId="9" pageNumber="354">
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<emphasis id="B967EA93FFBAFFA0FF2BFD37FEC6C0A5" bold="true" box="[151,353,727,753]" italics="true" pageId="9" pageNumber="354">Onuphis eremita</emphasis>
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Audouin & Milne Edwards, 1883
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C309650AFFBAFFA0FF2BFCFEFB87C1C4" pageId="9" pageNumber="354" type="reference_group">
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<paragraph id="8BAC3681FFBAFFA0FF2BFCFEFE81C107" blockId="9.[151,1436,798,912]" pageId="9" pageNumber="354">
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<treatmentCitationGroup id="AB0311AFFFBAFFA0FF2BFCFEFE81C107" pageId="9" pageNumber="354">
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<taxonomicName id="4C134D02FFBAFFA0FF2BFCFEFD73C160" ID-CoL="49QJ7" authority="Audouin & Milne Edwards, 1833: 226" authorityName="Audouin & Milne Edwards" authorityPageNumber="226" authorityYear="1833" box="[151,724,798,821]" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="9" pageNumber="354" phylum="Annelida" rank="species" species="eremita">
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<emphasis id="B967EA93FFBAFFA0FF2BFCFEFEE7C161" box="[151,320,798,821]" italics="true" pageId="9" pageNumber="354">Onuphis eremita</emphasis>
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<treatmentCitation id="0AB21090FFBAFFA0FEFBFCFEFD73C160" author="Audouin" box="[327,724,798,820]" page="226" pageId="9" pageNumber="354" year="1833">
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<bibRefCitation id="EF824B70FFBAFFA0FEFBFCFEFD06C160" author="Audouin" box="[327,673,798,820]" pageId="9" pageNumber="354" refString="Audouin, J. V. & Milne Edwards, H. (1833) Classification des Annelides et description de celles qui habitent les cotes de la France. Annales des sciences naturelles, 29, 195 - 269." type="journal article" year="1833">Audouin & Milne Edwards, 1833</bibRefCitation>
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: 226
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</treatmentCitation>
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</taxonomicName>
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; Redondo & San Martín, 1997: 227;
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<treatmentCitation id="0AB21090FFBAFFA0FBECFCFEFAC4C160" ID-DOI="http://doi.org/10.5281/zenodo.5217764" ID-Zenodo-Dep="5217764" author="Arias" box="[1104,1379,798,820]" httpUri="http://treatment.plazi.org/id/03CA87FEFF9F9C09FF0206ACFCE3FE23" page="148" pageId="9" pageNumber="354" year="2014">
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<bibRefCitation id="EF824B70FFBAFFA0FBECFCFEFA94C160" author="Arias" box="[1104,1331,798,820]" pageId="9" pageNumber="354" refString="Arias, A. & Paxton, H. (2014) Hidden diversity within the polychaete Onuphis eremita sensu lato (Annelida: Onuphidae) - redescription of O. eremita Audouin & Milne-Edwards, 1833 and reinstatement of Onuphis pancerii Claparede, 1868. Zootaxa, 3861 (2), 145 - 169." type="journal article" year="2014">Arias & Paxton, 2014</bibRefCitation>
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: 148
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</treatmentCitation>
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–158, figs. 1–8.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BAC3681FFBAFFA0FF2BFCBCFCC7C126" blockId="9.[151,1436,798,912]" box="[151,864,860,882]" pageId="9" pageNumber="354">
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<treatmentCitationGroup id="AB0311AFFFBAFFA0FF2BFCBCFCC7C126" box="[151,864,860,882]" pageId="9" pageNumber="354">
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<taxonomicName id="4C134D02FFBAFFA0FF2BFCBCFE54C126" ID-CoL="6SLW2" authority="Castelli, 1982: 45" authorityName="Castelli" authorityPageNumber="45" authorityYear="1982" box="[151,499,860,882]" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="9" pageNumber="354" phylum="Annelida" rank="species" species="falesia">
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<emphasis id="B967EA93FFBAFFA0FF2BFCBCFE90C126" box="[151,311,860,882]" italics="true" pageId="9" pageNumber="354">Onuphis falesia</emphasis>
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<treatmentCitation id="0AB21090FFBAFFA0FE81FCBCFE54C126" author="Castelli" box="[317,499,860,882]" page="45" pageId="9" pageNumber="354" year="1982">
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<bibRefCitation id="EF824B70FFBAFFA0FE81FCBCFE6AC126" author="Castelli" box="[317,461,860,882]" pageId="9" pageNumber="354" refString="Castelli, A. (1982) Onuphis falesia, a new species of Onuphinae (Polychaeta, Eunicidae). Bollettino di Zoologia, 49, 45 - 49. http: // dx. doi. org / 10.1080 / 11250008209439371" type="journal article" year="1982">Castelli, 1982</bibRefCitation>
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: 45
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</treatmentCitation>
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</taxonomicName>
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, fig. 1–11 (Gulf of Follonica,
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<collectingCountry id="F3047611FFBAFFA0FC9EFCBCFCF2C126" box="[802,853,860,882]" name="Italy" pageId="9" pageNumber="354">Italy</collectingCountry>
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).
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BAC3681FFBAFFA0FF2BFC9AFB87C1C4" blockId="9.[151,1436,798,912]" box="[151,1056,890,912]" pageId="9" pageNumber="354">
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<treatmentCitationGroup id="AB0311AFFFBAFFA0FF2BFC9AFB87C1C4" box="[151,1056,890,912]" pageId="9" pageNumber="354">
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<taxonomicName id="4C134D02FFBAFFA0FF2BFC9AFE35C1C4" ID-CoL="7HLPT" box="[151,402,890,912]" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="9" pageNumber="354" phylum="Annelida" rank="subSpecies" species="eremita" subSpecies="oculata">
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<emphasis id="B967EA93FFBAFFA0FF2BFC9AFE35C1C4" box="[151,402,890,912]" italics="true" pageId="9" pageNumber="354">Onuphis eremita oculata</emphasis>
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</taxonomicName>
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.—
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<treatmentCitation id="0AB21090FFBAFFA0FE0CFC9AFDCFC1C4" author="Cinar" box="[432,616,890,912]" httpUri="http://treatment.plazi.org/id/03BF3042C867FFFF8EA39AB3FE7C278B" page="2297" pageId="9" pageNumber="354" year="2009">
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<bibRefCitation id="EF824B70FFBAFFA0FE0CFC9AFD8CC1C4" author="Cinar" box="[432,555,890,912]" pageId="9" pageNumber="354" refString="Cinar, M. E. (2009) Alien polychaete species (Annelida: Polychaeta) on the southern coast of Turkey (Levantine Sea, eastern Mediterranean), with 13 new records for the Mediterranean Sea. Journal of Natural History, 43, 2283 - 2328. http: // dx. doi. org / 10.1080 / 00222930903094654" type="journal article" year="2009">Çinar, 2009</bibRefCitation>
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:
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<date id="FFAD1041FFBAFFA0FD89FC9AFDCFC1C4" box="[565,616,890,912]" pageId="9" pageNumber="354" value="2297" valueMax="2299">2297</date>
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</treatmentCitation>
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–2299, fig. 4 (
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<collectingCountry id="F3047611FFBAFFA0FD4AFC9AFCE3C1C4" box="[758,836,890,912]" name="Turkey" pageId="9" pageNumber="354">Turkey</collectingCountry>
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). Not
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<bibRefCitation id="EF824B70FFBAFFA0FC3DFC9BFBBAC1C4" author="Hartman" box="[897,1053,890,912]" pageId="9" pageNumber="354" refString="Hartman, O. (1951) The littoral marine annelids of the Gulf of Mexico. Publications of the Institute of Marine Sciences, 2, 7 - 124." type="journal article" year="1951">Hartman, 1951</bibRefCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C309650AFFBAFFA0FF2BFC5DFC07C632" pageId="9" pageNumber="354" type="materials_examined">
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||
<paragraph id="8BAC3681FFBAFFA0FF2BFC5DFCD7C1AE" blockId="9.[151,1437,957,1810]" pageId="9" pageNumber="354">
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<emphasis id="B967EA93FFBAFFA0FF2BFC5DFE25C182" bold="true" box="[151,386,957,982]" pageId="9" pageNumber="354">Material examined.</emphasis>
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<typeStatus id="54A88823FFBAFFA0FE36FC5EFE66C182" box="[394,449,958,982]" pageId="9" pageNumber="354">Type</typeStatus>
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material.
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<typeStatus id="54A88823FFBAFFA0FD89FC5EFD33C182" box="[565,660,958,982]" pageId="9" pageNumber="354" type="neotype">Neotype</typeStatus>
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(
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<collectionCode id="ED02AE44FFBAFFA0FD18FC5EFD5BC182" LSID="urn:lsid:biocol.org:col:34988" box="[676,764,958,982]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="9" pageNumber="354">MNHN</collectionCode>
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<typeStatus id="54A88823FFBAFFA0FCB9FC5EFCECC181" box="[773,843,958,981]" pageId="9" pageNumber="354">
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<collectionCode id="ED02AE44FFBAFFA0FCB9FC5EFCECC181" box="[773,843,958,981]" pageId="9" pageNumber="354">TYPE</collectionCode>
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</typeStatus>
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1555) La Rochelle,
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<collectingCountry id="F3047611FFBAFFA0FB8AFC5EFB22C181" box="[1078,1157,958,981]" name="France" pageId="9" pageNumber="354">France</collectingCountry>
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, East Atlantic,
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<geoCoordinate id="EE275046FFBAFFA0FA88FC5DFA3FC182" box="[1332,1432,957,982]" direction="north" orientation="latitude" pageId="9" pageNumber="354" precision="925" value="46.15">46°09’N</geoCoordinate>
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,
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<geoCoordinate id="EE275046FFBAFFA0FF2BFC01FF5AC1AE" box="[151,253,993,1018]" direction="west" orientation="longitude" pageId="9" pageNumber="354" precision="925" value="-1.15">01°09’W</geoCoordinate>
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–
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<geoCoordinate id="EE275046FFBAFFA0FEB1FC01FEC8C1AE" box="[269,367,993,1018]" direction="north" orientation="latitude" pageId="9" pageNumber="354" precision="925" value="46.15">46°09’N</geoCoordinate>
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,
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<geoCoordinate id="EE275046FFBAFFA0FEC7FC02FE46C1AE" box="[379,481,994,1018]" direction="west" orientation="longitude" pageId="9" pageNumber="354" precision="925" value="-1.1666666">01°10’W</geoCoordinate>
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, intertidal sands, coll.
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<date id="FFAD1041FFBAFFA0FD61FC02FCCBC1AE" box="[733,876,994,1018]" pageId="9" pageNumber="354" value="2011-09-28">28 Sep 2011</date>
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.
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</paragraph>
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<paragraph id="8BAC3681FFBAFFA0FF7BFBE6FD56C616" blockId="9.[151,1437,957,1810]" pageId="9" pageNumber="354">
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Non-type material. For a detailed listing of additional material from
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<collectingCountry id="F3047611FFBAFFA0FC58FBE6FB94C649" box="[996,1075,1030,1053]" name="France" pageId="9" pageNumber="354">France</collectingCountry>
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, Cantabrian and Mediterranean coasts of
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<collectingCountry id="F3047611FFBAFFA0FEBDFBC9FEE6C616" box="[257,321,1065,1090]" name="Spain" pageId="9" pageNumber="354">Spain</collectingCountry>
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and
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<collectingCountry id="F3047611FFBAFFA0FECBFBC9FE1FC616" box="[375,440,1065,1090]" name="Malta" pageId="9" pageNumber="354">Malta</collectingCountry>
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see
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<bibRefCitation id="EF824B70FFBAFFA0FE56FBC9FD4AC616" author="Arias" box="[490,749,1065,1090]" pageId="9" pageNumber="354" refString="Arias, A. & Paxton, H. (2014) Hidden diversity within the polychaete Onuphis eremita sensu lato (Annelida: Onuphidae) - redescription of O. eremita Audouin & Milne-Edwards, 1833 and reinstatement of Onuphis pancerii Claparede, 1868. Zootaxa, 3861 (2), 145 - 169." type="journal article" year="2014">Arias & Paxton (2014)</bibRefCitation>
|
||
.
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||
</paragraph>
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||
<paragraph id="8BAC3681FFBAFFA0FF7BFBADFC07C632" blockId="9.[151,1437,957,1810]" box="[199,928,1101,1126]" pageId="9" pageNumber="354">
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||
<emphasis id="B967EA93FFBAFFA0FF7BFBADFEC2C632" bold="true" box="[199,357,1101,1126]" pageId="9" pageNumber="354">
|
||
<typeStatus id="54A88823FFBAFFA0FF7BFBADFEA5C632" box="[199,258,1101,1126]" pageId="9" pageNumber="354">Type</typeStatus>
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||
locality.
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||
</emphasis>
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La Rochelle,
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<collectingCountry id="F3047611FFBAFFA0FDBEFBAEFDF6C631" box="[514,593,1102,1125]" name="France" pageId="9" pageNumber="354">France</collectingCountry>
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||
, Bay of Biscay, East Atlantic.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C309650AFFBAFFA0FF7BFB91FC7DC79A" pageId="9" pageNumber="354" type="diagnosis">
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||
<paragraph id="8BAC3681FFBAFFA0FF7BFB91FC7DC79A" blockId="9.[151,1437,957,1810]" pageId="9" pageNumber="354">
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||
<emphasis id="B967EA93FFBAFFA0FF7BFB91FEE6C6DE" bold="true" box="[199,321,1137,1162]" pageId="9" pageNumber="354">Diagnosis.</emphasis>
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Prostomium anteriorly extended, one pair of small eyespots present at anterolateral end of prostomium. Palps reaching chaetiger 2–4, lateral antennae reaching chaetiger 7–14 and median antenna chaetiger 4–7. Ceratophores long and strongly ringed, palpal ceratophores with 16–20 rings, lateral antennae with 19–22 and median antenna with 13–19 rings. Subulate ventral cirri in first five to six chaetigers; distinct subulate postchaetal lobes in first 10–17 chaetigers. Small interramal papilla present at base of dorsal cirrus in chaetigers 4(5)–9. Strongly tridentate pseudocompound slender, long and robust, short appendaged hooks in first three chaetigers. Subacicular hooks from chaetiger 10. Flat, distally oblique pectinate chaetae with 10–13 teeth. Single branchial filaments from chaetiger 1 to 20–26, thereafter number increasing rapidly to maximum of three to five. Colour pattern consisting of two rows of brownish patches on dorsal anterior end or brown transverse stripes when lateral patches coalesce medially. Tube thin-walled and covered with sand grains.
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||
</paragraph>
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</subSubSection>
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||
<subSubSection id="C309650AFFBAFFA0FF7BFA39FC85C4F2" pageId="9" pageNumber="354" type="discussion">
|
||
<paragraph id="8BAC3681FFBAFFA0FF7BFA39FC85C4F2" blockId="9.[151,1437,957,1810]" pageId="9" pageNumber="354">
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||
<emphasis id="B967EA93FFBAFFA0FF7BFA39FE9CC7A6" bold="true" box="[199,315,1497,1522]" pageId="9" pageNumber="354">Remarks.</emphasis>
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The taxonomic identity of
|
||
<taxonomicName id="4C134D02FFBAFFA0FDD0FA3AFD44C7A5" box="[620,739,1498,1521]" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="9" pageNumber="354" phylum="Annelida" rank="species" species="eremita">
|
||
<emphasis id="B967EA93FFBAFFA0FDD0FA3AFD44C7A5" box="[620,739,1498,1521]" italics="true" pageId="9" pageNumber="354">O. eremita</emphasis>
|
||
</taxonomicName>
|
||
, the
|
||
<typeStatus id="54A88823FFBAFFA0FCA7FA3AFCECC7A6" box="[795,843,1498,1522]" pageId="9" pageNumber="354">type</typeStatus>
|
||
species of the genus, was extremely controversial. It was confounded by vague descriptions, redescriptions based on worldwide material and the establishment of dubious synonymies over the last century. However, a
|
||
<typeStatus id="54A88823FFBAFFA0FC9BF9C2FC27C46E" box="[807,896,1570,1594]" pageId="9" pageNumber="354" type="neotype">neotype</typeStatus>
|
||
was recently designated and the species was redescribed by
|
||
<bibRefCitation id="EF824B70FFBAFFA0FEFBF9A5FDF5C40A" author="Arias" box="[327,594,1605,1630]" pageId="9" pageNumber="354" refString="Arias, A. & Paxton, H. (2014) Hidden diversity within the polychaete Onuphis eremita sensu lato (Annelida: Onuphidae) - redescription of O. eremita Audouin & Milne-Edwards, 1833 and reinstatement of Onuphis pancerii Claparede, 1868. Zootaxa, 3861 (2), 145 - 169." type="journal article" year="2014">Arias & Paxton (2014)</bibRefCitation>
|
||
. These authors redefined its diagnostic features, restricted its distribution range to the European Atlantic and the Mediterranean Sea, making its conspecificity with the specimens reported from outside Europe and the Mediterranean very unlikely.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C309650AFFBAFFA0FF7BF951FE3DC546" pageId="9" pageNumber="354" type="distribution">
|
||
<paragraph id="8BAC3681FFBAFFA0FF7BF951FE3DC546" blockId="9.[151,1437,957,1810]" pageId="9" pageNumber="354">
|
||
<emphasis id="B967EA93FFBAFFA0FF7BF951FEC7C49E" bold="true" box="[199,352,1713,1738]" pageId="9" pageNumber="354">Distribution.</emphasis>
|
||
East Atlantic (Cantabrian Sea, Bay of Biscay) and western and central Mediterranean. However, it is most likely that the species is widely distributed along the Atlantic coasts of
|
||
<collectingCountry id="F3047611FFBAFFA0FB95F935FBCDC4BA" box="[1065,1130,1749,1774]" name="Georgia" pageId="9" pageNumber="354">Iberia</collectingCountry>
|
||
and the Levantine basin of the Mediterranean Sea.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |