166 lines
18 KiB
XML
166 lines
18 KiB
XML
<document id="517BF93C804624F2E27AC9A1D04AC496" ID-DOI="10.1206/3961.1" ID-ISSN="0003-0082" ID-Zenodo-Dep="5370399" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630268866948" checkinUser="carolina" docAuthor="Sinclair, Bradley J. & Grimaldi, David A." docDate="2020" docId="03B94C5BFFC23C4867EB37C0FF2B7617" docLanguage="en" docName="N3961.pdf" docOrigin="American Museum Novitates 2020 (3961)" docSource="https://bioone.org/journals/american-museum-novitates/volume-2020/issue-3961/3961.1/Cretaceous-Diversity-of-the-Relict-Genus-Alavesia-Waters-and-Arillo/10.1206/3961.1.full" docStyle="DocumentStyle:26C3580A17D4BBE58BD8D8AC6AE09AC4.6:AmMusNovit.2010-.journal_article" docStyleId="26C3580A17D4BBE58BD8D8AC6AE09AC4" docStyleName="AmMusNovit.2010-.journal_article" docStyleVersion="6" docTitle="Alavesia brevipennae Sinclair & Grimaldi 2020, new species" docType="treatment" docVersion="4" lastPageNumber="15" masterDocId="FF803423FFCC3C46662D3721FC617321" masterDocTitle="Cretaceous diversity of the relict genus Alavesia Waters and Arillo (Diptera: Empidoidea: Atelestidae)" masterLastPageNumber="40" masterPageNumber="1" pageNumber="15" updateTime="1699205599378" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:mods id="4F873BB6577E93007CA24825954D4219" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="0ECABDAF2A8034A24D28FD79165D02C8">
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<mods:title id="013EFB4037392FC31CE4EFE840BE47EA">Cretaceous diversity of the relict genus Alavesia Waters and Arillo (Diptera: Empidoidea: Atelestidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="17BB9B3A3A051D7EDA275C2D5B1ADEB9" type="personal">
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<mods:role id="2F6809FBF3FB8002B39E137C41D5CB8D">
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<mods:roleTerm id="F37282548951127A7A3A9CB4F9AF5D02">Author</mods:roleTerm>
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<mods:namePart id="2ED6CEAE3B14558307719400C6956FCE">Sinclair, Bradley J.</mods:namePart>
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</mods:name>
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<mods:name id="872960F25EC6C8ECF6811C7C07E35E0F" type="personal">
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<mods:role id="FEA5FBCA7DAED9CB787D57C05661CF4E">
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<mods:roleTerm id="68E30C4534E0AA1A6C51BF47C3C6607F">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="3316FF08C6457D692D05EC1EA68AE5B8">Grimaldi, David A.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="7DA850764A31C46BB9E7D1591054D96A">text</mods:typeOfResource>
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<mods:title id="D94ADF924F222569648BB355D54467FF">American Museum Novitates</mods:title>
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<mods:part id="CE559790BE8BC46E63D9E3B1DC9E4622">
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<mods:date id="3A0BC684B0F5DF34D940EA22FF55F28B">2020</mods:date>
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<mods:detail id="D5FC1BA1CE4D8A5443054E243222153F" type="pubDate">
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<mods:number id="AEAC58BCAB3B0BA276782D9F2904717E">2020-10-06</mods:number>
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<mods:detail id="45C27479E481F9A9BEC9E065F5DA6B36" type="volume">
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<mods:number id="9FA709A96D0F8B03467342DBAB470A38">2020</mods:number>
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</mods:detail>
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<mods:detail id="B718D5E70EB615DC334256E6BA76FD88" type="issue">
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<mods:number id="E65D958D54E9C6210661FA742DCA4B7A">3961</mods:number>
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</mods:detail>
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<mods:start id="4F78BFAE667EF37A5758EED936A4D056">1</mods:start>
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<mods:location id="C264521A3DD663A2519998535684B880">
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<mods:url id="D6CEF5E221B4B1086E63422071A5D47C">https://bioone.org/journals/american-museum-novitates/volume-2020/issue-3961/3961.1/Cretaceous-Diversity-of-the-Relict-Genus-Alavesia-Waters-and-Arillo/10.1206/3961.1.full</mods:url>
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</mods:location>
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<mods:classification id="7AE2C7E73498E5C6B72BAFCCF65116FF">journal article</mods:classification>
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<mods:identifier id="644450962FD461360AE4F37CBF8C98B6" type="DOI">10.1206/3961.1</mods:identifier>
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<mods:identifier id="01A34AF7AE87955442C8195F0B39B84B" type="ISSN">0003-0082</mods:identifier>
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<mods:identifier id="94E72535FE68CD00B16F7316E6B7E91E" type="Zenodo-Dep">5370399</mods:identifier>
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<treatment id="03B94C5BFFC23C4867EB37C0FF2B7617" ID-DOI="http://doi.org/10.5281/zenodo.5462031" ID-Zenodo-Dep="5462031" LSID="urn:lsid:plazi:treatment:03B94C5BFFC23C4867EB37C0FF2B7617" httpUri="http://treatment.plazi.org/id/03B94C5BFFC23C4867EB37C0FF2B7617" lastPageNumber="15" pageId="14" pageNumber="15">
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<subSubSection id="C30AAEC6FFC23C4867EB37C0FF0373DA" box="[454,866,225,252]" pageId="14" pageNumber="15" type="nomenclature">
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<paragraph id="8BAFFD4DFFC23C4867EB37C0FF0373DA" blockId="14.[454,866,225,252]" box="[454,866,225,252]" pageId="14" pageNumber="15">
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<heading id="D0E74A21FFC23C4867EB37C0FF0373DA" box="[454,866,225,252]" centered="true" fontSize="11" level="2" pageId="14" pageNumber="15" reason="2">
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<emphasis id="B964215FFFC23C4867EB37C0FEAE73DA" bold="true" box="[454,719,225,252]" pageId="14" pageNumber="15">
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<taxonomicName id="4C1086CEFFC23C4867EB37C0FEA973DD" ID-CoL="89WYF" authority="Sinclair & Grimaldi, 2020" authorityName="Sinclair & Grimaldi" authorityYear="2020" box="[454,712,225,252]" class="Insecta" family="Atelestidae" genus="Alavesia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="brevipennae" status="sp. nov.">
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<emphasis id="B964215FFFC23C4867EB37C0FEA973DD" bold="true" box="[454,712,225,252]" italics="true" pageId="14" pageNumber="15">Alavesia brevipennae</emphasis>
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</taxonomicName>
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,
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</emphasis>
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<taxonomicNameLabel id="A2579C24FFC23C4864FA37C3FF0373DA" box="[727,866,225,252]" pageId="14" pageNumber="15" rank="species">new species</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30AAEC6FFC23C4867DA3638FF507212" box="[503,817,281,307]" pageId="14" pageNumber="15" type="description">
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<paragraph id="8BAFFD4DFFC23C4867DA3638FF507212" blockId="14.[503,817,281,307]" box="[503,817,281,307]" pageId="14" pageNumber="15">
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<heading id="D0E74A21FFC23C4867DA3638FF507212" box="[503,817,281,307]" centered="true" fontSize="36" level="3" pageId="14" pageNumber="15" reason="7">
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<figureCitation id="132BE1C8FFC23C4867DA3638FE177212" box="[503,630,281,307]" captionStart="FIGURE" captionStartId="6.[108,197,1285,1306]" captionTargetBox="[200,1125,232,1256]" captionTargetId="figure-108@6.[108,1212,224,1262]" captionTargetPageId="6" captionText="FIGURE. 1. Antennae of Burmese amber Alavesia species, lateral views. All are males except N, O, P. Some antennae were reversed in direction for optimal comparison; longicornuta and longistylata are slightly reconstructed to account for curvature of the stylus. All are to the same scale. Specimen numbers: A. USNM PAL 726867. B. AMNH Bu-494. C. AMNH Bu-496. D. USNM PAL 726868. E. AMNH Bu-KL2-7. F. AMNH Bu-1476. G. JZC Bu-556 (AMNH). H. USNM PAL 726872. K. USNM PAL 726869. L. AMNH Bu-KL7-18. M. JZC Bu-282. N. Undetermined species. O. A. zigrasi. P. A. zigrasi." figureDoi="http://doi.org/10.5281/zenodo.5370401" httpUri="https://zenodo.org/record/5370401/files/figure.png" pageId="14" pageNumber="15">Figures 1B</figureCitation>
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,
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<figureCitation id="132BE1C8FFC23C4864AE3638FEC57212" box="[643,676,281,307]" captionStart="FIGURE 2" captionStartId="7.[108,194,1456,1477]" captionTargetBox="[121,1197,229,1434]" captionTargetId="figure-45@7.[108,1212,224,1437]" captionTargetPageId="7" captionText="FIGURE 2. Wings of six species of Alavesia in Burmese amber. All are males, to the same scale. Specimens: A. USNM PAL 726867. B. AMNH Bu-494. C. AMNH Bu-496. D. USNM PAL 726868. E. AMNH Bu-1476. F. JZC Bu-556." figureDoi="http://doi.org/10.5281/zenodo.5370403" httpUri="https://zenodo.org/record/5370403/files/figure.png" pageId="14" pageNumber="15">2B</figureCitation>
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,
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<figureCitation id="132BE1C8FFC23C48649C3638FEB47212" box="[689,725,281,307]" captionStart="FIGURE 8" captionStartId="17.[108,194,1252,1273]" captionTargetBox="[108,1212,224,1227]" captionTargetId="figure-95@17.[108,1212,224,1229]" captionTargetPageId="17" captionText="FIGURE 8. Heads and antennae of Burmese amber Alavesia. A. A. lanceolata, head and antennae, dorsolateral view, holotype, USNM 726868. B. A. pankowskiorum, head and antennae, lateral view, paratype, AMNH Bu-KL7-19. C. A. magna, left antenna, lateral view, holotype, AMNH Bu-KL2-7. D. A. magna, left antenna detail, lateral view, holotype, AMNH Bu-KL2-7. E. A. brevipennae, lateral view, AMNH Bu-494. Note the longer microtrichia on the postpedicel. F. A. longicornuta, holotype, stylus tip, lateral view, JZC Bu-556." figureDoi="http://doi.org/10.5281/zenodo.5370415" httpUri="https://zenodo.org/record/5370415/files/figure.png" pageId="14" pageNumber="15">8D</figureCitation>
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,
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<figureCitation id="132BE1C8FFC23C4864CF3638FF507212" box="[738,817,281,307]" captionStart="FIGURE 12" captionStartId="21.[108,194,1223,1244]" captionTargetBox="[108,1212,224,1202]" captionTargetId="figure-51@21.[625,1212,705,1208]" captionTargetPageId="21" captionText="FIGURE 12. Alavesia spp. in Burmese amber. A. A. angusta, USNM PAL 726867. B. Drawing of A. brevipennae, AMNH Bu-494. C. Photomicrograph of A. brevipennae, AMNH Bu-494. D. A. myanmarensis, AMNH Bu-496." figureDoi="http://doi.org/10.5281/zenodo.5370423" httpUri="https://zenodo.org/record/5370423/files/figure.png" pageId="14" pageNumber="15">12B, C</figureCitation>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30AAEC6FFC23C48668E3670FEB772FD" pageId="14" pageNumber="15" type="diagnosis">
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<paragraph id="8BAFFD4DFFC23C48668E3670FEB772FD" blockId="14.[108,1213,337,1335]" pageId="14" pageNumber="15">
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DIAGNOSIS: Tiny species (fig. 12B, C), wing length
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<quantity id="4CE850A8FFC23C4864DA3670FF00724D" box="[759,865,337,364]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" pageId="14" pageNumber="15" unit="mm" value="0.75">0.75 mm</quantity>
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; postpedicel 2.3
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<emphasis id="B964215FFFC23C4862303670F84F724A" bold="true" box="[1053,1070,337,363]" pageId="14" pageNumber="15">×</emphasis>
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longer than aristalike stylus, slightly curved; wing shortened, rounded, and broad (L/
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<geoCoordinate id="EE249B8AFFC23C4865983656F86D72B0" box="[949,1036,374,401]" degrees="1.75" direction="west" orientation="longitude" pageId="14" pageNumber="15" precision="555" value="-1.75">W 1.75</geoCoordinate>
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); M
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<subScript id="1794FF08FFC23C48621436A5F83672B4" attach="left" box="[1081,1111,388,405]" fontSize="7" pageId="14" pageNumber="15">1+2</subScript>
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strongly arched, this vein and M
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<subScript id="1794FF08FFC23C4867AC368BFDEB729A" attach="left" box="[385,394,426,443]" fontSize="7" pageId="14" pageNumber="15">4</subScript>
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very divergent; CuA arched apically, CuA+CuP weakened, not reaching wing margin, M
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<subScript id="1794FF08FFC23C48677436EEFD0372C1" attach="left" box="[345,354,463,480]" fontSize="7" pageId="14" pageNumber="15">4</subScript>
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closer to CuA+CuP than M
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<subScript id="1794FF08FFC23C48649C36EEFEAE72C1" attach="left" box="[689,719,463,480]" fontSize="7" pageId="14" pageNumber="15">1+2</subScript>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C30AAEC6FFC23C48668936C6FF1877E7" pageId="14" pageNumber="15" type="description">
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<paragraph id="8BAFFD4DFFC23C48668936C6F8C97150" blockId="14.[108,1213,337,1335]" pageId="14" pageNumber="15">
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DESCRIPTION:
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<emphasis id="B964215FFFC23C48676136C6FDEA7120" box="[332,395,487,513]" italics="true" pageId="14" pageNumber="15">Male.</emphasis>
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<emphasis id="B964215FFFC23C4867BF36C7FE757121" bold="true" box="[402,532,486,512]" pageId="14" pageNumber="15">Coloration</emphasis>
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not preserved.
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<emphasis id="B964215FFFC23C4864EA36C6FF6C7120" bold="true" box="[711,781,487,513]" pageId="14" pageNumber="15">Head:</emphasis>
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Scape and pedicel quadrate, subequal in size and length. Postpedicel elongate, straplike, slightly curved, gradually tapered apically, 2.3
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<emphasis id="B964215FFFC23C486286352DF8DD7107" bold="true" box="[1195,1212,524,550]" pageId="14" pageNumber="15">×</emphasis>
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longer than stylus (fig. 1B). Stylus with two basal articles, short (fig. 8D), without apical flaglike expansion; length 0.32
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<emphasis id="B964215FFFC23C4867453576FD187150" bold="true" box="[360,377,599,625]" pageId="14" pageNumber="15">×</emphasis>
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that of postpedicel. Mouthparts very short; palpus short, not projecting.
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</paragraph>
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<paragraph id="8BAFFD4DFFC23C48668B355DFEE271C0" blockId="14.[108,1213,337,1335]" pageId="14" pageNumber="15">
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<emphasis id="B964215FFFC23C48668B355DFD6571B7" bold="true" box="[166,260,636,662]" pageId="14" pageNumber="15">Thorax:</emphasis>
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Chaetotaxy thickened, in numerous rows (not clearly visible): 2 anterior and 1 posterior postpronotal setae; 4 notopleural setae of variable lengths, 1 postsutural supraalar seta; 1 postalar seta; 2 pairs of scutellar setae.
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</paragraph>
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<paragraph id="8BAFFD4DFFC23C48668E35CDF83470BA" blockId="14.[108,1213,337,1335]" pageId="14" pageNumber="15">
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<emphasis id="B964215FFFC23C48668E35CDFC807027" bold="true" box="[163,225,748,774]" pageId="14" pageNumber="15">Legs:</emphasis>
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Most setae thickened. Forecoxa with short erect anterior setae; anteroapical setae longer and stouter. Forefemur with anteroventral and posteroventral row of short setae. Mid and hind femora with anteroventral row of short setae. Mid and hind coxae with 2–3 stout lateral setae on apical half. Tibiae with biserial row of short, stout ventral setae; apex of tibiae with long, apical ventral seta; hind tibia with several dorsal setae, length about half width of tibia.
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</paragraph>
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<paragraph id="8BAFFD4DFFC23C48668E3487FE15775A" blockId="14.[108,1213,337,1335]" pageId="14" pageNumber="15">
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<emphasis id="B964215FFFC23C48668E3487FC8770E1" bold="true" box="[163,230,934,960]" pageId="14" pageNumber="15">Wing</emphasis>
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(wing length
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<quantity id="4CE850A8FFC23C4867A73487FD9570E0" box="[394,500,934,961]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" pageId="14" pageNumber="15" unit="mm" value="0.75">0.75 mm</quantity>
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) (fig. 2B): shortened and rounded; costa terminating slightly beyond M
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<subScript id="1794FF08FFC23C4866C934FBFD6370CA" attach="left" box="[228,258,986,1003]" fontSize="7" pageId="14" pageNumber="15">1+2</subScript>
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; Sc nearly reaching costal margin; R
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<subScript id="1794FF08FFC23C48648034FBFEAA70CA" attach="left" box="[685,715,986,1003]" fontSize="7" pageId="14" pageNumber="15">4+5</subScript>
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closely associated with M
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<subScript id="1794FF08FFC23C4865D134FBF87B70CA" attach="left" box="[1020,1050,986,1003]" fontSize="7" pageId="14" pageNumber="15">1+2</subScript>
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; M
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<subScript id="1794FF08FFC23C48626F34FBF80170CA" attach="left" box="[1090,1120,986,1003]" fontSize="7" pageId="14" pageNumber="15">1+2</subScript>
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arched, strongly divergent from R
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<subScript id="1794FF08FFC23C4867B634DEFDD87731" attach="left" box="[411,441,1023,1040]" fontSize="7" pageId="14" pageNumber="15">4+5</subScript>
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distally; cell dm very short, crossvein r-m nearly opposite apex of cell bm; M
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<subScript id="1794FF08FFC23C4866C13305FC947714" attach="left" box="[236,245,1060,1077]" fontSize="7" pageId="14" pageNumber="15">4</subScript>
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straight to wing margin; apex of M
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<subScript id="1794FF08FFC23C4864B23305FEC97714" attach="left" box="[671,680,1060,1077]" fontSize="7" pageId="14" pageNumber="15">4</subScript>
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closer to CuA+CuP than M
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<subScript id="1794FF08FFC23C4865D73305F8797714" attach="left" box="[1018,1048,1060,1077]" fontSize="7" pageId="14" pageNumber="15">1+2</subScript>
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; CuA arched; apex of cell cua convex, length of cell cua subequal to cell bm; CuA+CuP mostly straight, strongly curved subapically to wing margin.
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</paragraph>
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<paragraph id="8BAFFD4DFFC23C48668933A7FF1877E7" blockId="14.[108,1213,337,1335]" pageId="14" pageNumber="15">
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<emphasis id="B964215FFFC23C48668933A7FD457781" bold="true" box="[164,292,1158,1184]" pageId="14" pageNumber="15">Abdomen:</emphasis>
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Setae short, without distinct posteromarginal setae. Segments 1–5 broad. Terminalia and remaining segments damaged and partially obscured.
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</paragraph>
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</subSubSection>
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<subSubSection id="C30AAEC6FFC23C48668933F3FF0477CA" box="[164,869,1233,1260]" pageId="14" pageNumber="15" type="materials_examined">
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<paragraph id="8BAFFD4DFFC23C48668933F3FF0477CA" blockId="14.[108,1213,337,1335]" box="[164,869,1233,1260]" pageId="14" pageNumber="15">
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<materialsCitation id="3B78F710FFC23C48668933F3FF0477CA" box="[164,869,1233,1260]" collectionCode="TYPE, MATERIAL, AMNH" pageId="14" pageNumber="15" specimenCount="1" typeStatus="holotype">
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<collectionCode id="ED016588FFC23C48668933F3FD0077CD" box="[164,353,1234,1260]" pageId="14" pageNumber="15">TYPE MATERIAL</collectionCode>
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:
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<typeStatus id="54AB43EFFFC23C48674133F0FDBA77CA" box="[364,475,1233,1259]" pageId="14" pageNumber="15" type="holotype">Holotype</typeStatus>
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, male,
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<collectionCode id="ED016588FFC23C48640233F3FEEB77CD" box="[559,650,1234,1260]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="14" pageNumber="15">AMNH</collectionCode>
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Bu-494 (
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<collectionCode id="ED016588FFC23C4864DA33F0FF3977CA" box="[759,856,1233,1259]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="14" pageNumber="15">AMNH</collectionCode>
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||
).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C30AAEC6FFC23C48668F33D6FF2B7617" pageId="14" pageNumber="15" type="etymology">
|
||
<paragraph id="8BAFFD4DFFC23C48668F33D6FF2B7617" blockId="14.[108,1213,337,1335]" pageId="14" pageNumber="15">
|
||
ETYMOLOGY: The specific epithet is from the Latin
|
||
<emphasis id="B964215FFFC23C4864C133D6FF537631" box="[748,818,1271,1296]" italics="true" pageId="14" pageNumber="15">penna</emphasis>
|
||
(wing, feather) and
|
||
<emphasis id="B964215FFFC23C48623133D6F83E7630" box="[1052,1119,1271,1297]" italics="true" pageId="14" pageNumber="15">brevis</emphasis>
|
||
(short), referring to the shortened wing that characterizes this species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |