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<document ID-CLB-Dataset="256667" ID-DOI="http://dx.doi.org/10.3897/phytokeys.225.99356" ID-GBIF-Dataset="9edaba47-012e-4c83-a606-8a760dc24de6" ID-PMC="PMC10209509" ID-Pensoft-Pub="1314-2003-225-115" ID-Pensoft-UUID="AB75E2E9609C5A7583F34B48EA61B976" ID-PubMed="37252326" ModsDocID="1314-2003-225-115" checkinTime="1682624511249" checkinUser="pensoft" docAuthor="Khamar, Hamid, Oualidi, Jalal El, Touhami, Amina Ouazzani &amp; Civeyrel, Laure" docDate="2023" docId="34E7283141BA563398E7304180014C15" docLanguage="en" docName="PhytoKeys 225: 115-152" docOrigin="PhytoKeys 225" docPubDate="2023-04-27" docSource="http://dx.doi.org/10.3897/phytokeys.225.99356" docTitle="Verbascum letourneuxii Asch. ex Asch. &amp; Schweinf., Ill. Fl. Egypt 2: 189 &amp; 114. 1887." docType="treatment" docVersion="5" id="AB75E2E9609C5A7583F34B48EA61B976" lastPageNumber="115" masterDocId="AB75E2E9609C5A7583F34B48EA61B976" masterDocTitle="Nomenclature and typification in Verbascum (Scrophulariaceae) from North Africa" masterLastPageNumber="152" masterPageNumber="115" pageNumber="115" updateTime="1732936468263" updateUser="ExternalLinkService">
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<mods:title id="A68B9359A17CFD48F69C4C2752847B06">Nomenclature and typification in Verbascum (Scrophulariaceae) from North Africa</mods:title>
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<mods:namePart id="B30DD359955965634CC5D845EAADF6A1">Khamar, Hamid</mods:namePart>
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<mods:affiliation id="F9912822D5C5A63D9934B28255F01C1B">Department of Botany and Plant Ecology, Scientific Institute, University Mohammed V in Rabat, B. P. 703, Rabat 10106, Morocco</mods:affiliation>
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<mods:namePart id="E0EC1CAD2DCEC17B8A0566A4FA915BB9">Oualidi, Jalal El</mods:namePart>
<mods:affiliation id="B79194C169F6975AFC3E66A8F470900E">Department of Botany and Plant Ecology, Scientific Institute, University Mohammed V in Rabat, B. P. 703, Rabat 10106, Morocco</mods:affiliation>
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<treatment id="34E7283141BA563398E7304180014C15" ID-GBIF-Taxon="209579297" LSID="urn:lsid:plazi:treatment:34E7283141BA563398E7304180014C15" httpUri="http://treatment.plazi.org/id/34E7283141BA563398E7304180014C15" lastPageNumber="115" pageId="0" pageNumber="115" scope_family="Scrophulariaceae">
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<taxonomicName id="C6080027E38DF0E000124C97DC1187EB" LSID="34E72831-41BA-5633-98E7-304180014C15" authority="Asch. ex Asch. &amp; Schweinf., Ill. Fl. Egypt 2: 189 &amp; 114. 1887." authorityName="Asch. ex Asch. &amp; Schweinf., Ill. Fl. Egypt 2: 189 &amp; 114. 1887." class="Dicotyledoneae" family="Scrophulariaceae" genus="Verbascum" higherTaxonomySource="treatment-meta" kingdom="Plantae" lsidName="Verbascum letourneuxii" order="Lamiales" pageId="0" pageNumber="115" rank="species" species="letourneuxii">Verbascum letourneuxii Asch. ex Asch. &amp; Schweinf., Ill. Fl. Egypt 2: 189 &amp; 114. 1887.</taxonomicName>
</paragraph>
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<subSubSection id="FA5251ED930A4FABFC03922D026EA257" pageId="0" pageNumber="115" type="reference_group">
<paragraph id="C1224CEEF0CCB91E9D21609DB2E9A2E7" pageId="0" pageNumber="115">
<taxonomicName id="C0E7BE228CAC6D9E4F19AE19A5B08B40" class="Dicotyledoneae" family="Scrophulariaceae" genus="Verbascum" higherTaxonomySource="treatment-meta" kingdom="Plantae" lsidName="Verbascum spinosum" order="Lamiales" pageId="0" pageNumber="115" rank="species" species="spinosum">Verbascum spinosum</taxonomicName>
=
<taxonomicName id="E070B5669E12414561E7CCF37186AE60" class="Dicotyledoneae" family="Scrophulariaceae" genus="Verbascum" higherTaxonomySource="treatment-meta" kingdom="Plantae" lsidName="Verbascum spinosum" order="Lamiales" pageId="0" pageNumber="115" rank="species" species="spinosum">Verbascum spinosum</taxonomicName>
Delile, Fl. AEg. Illustr.: 55. 1813; non L. Cent. II. plant.: 10. 1756. &amp; Amoen. Acad. IV: 307. 1759.
</paragraph>
<paragraph id="44344925BDB027130753E9BE0B3E5B20" pageId="0" pageNumber="115">
<taxonomicName id="39D72F3E3FF1467025F9C0B872516AAE" class="Dicotyledoneae" family="Scrophulariaceae" genus="Verbascum" higherTaxonomySource="treatment-meta" kingdom="Plantae" lsidName="Verbascum marniaricum" order="Lamiales" pageId="0" pageNumber="115" rank="species" species="marniaricum">Verbascum marniaricum</taxonomicName>
=
<taxonomicName id="4259DAB20686FEE7E30BCF874545FCC8" class="Dicotyledoneae" family="Scrophulariaceae" genus="Verbascum" higherTaxonomySource="treatment-meta" kingdom="Plantae" lsidName="Verbascum marniaricum" order="Lamiales" pageId="0" pageNumber="115" rank="species" species="marniaricum">Verbascum marniaricum</taxonomicName>
Letournex ap. Barbey Herboris, au Levant: 148. 1882, nom. nud.
</paragraph>
<paragraph id="038F5729774CB6D4B933FD3AF9B5DD05" pageId="0" pageNumber="115">
<taxonomicName id="F03C129F4F7EA10495567D9EC51FE116" class="Dicotyledoneae" family="Scrophulariaceae" genus="Verbascum" higherTaxonomySource="treatment-meta" kingdom="Plantae" lsidName="Verbascum tourneuxii" order="Lamiales" pageId="0" pageNumber="115" rank="species" species="tourneuxii">Verbascum tourneuxii</taxonomicName>
=
<taxonomicName id="743C61DC5C71C9BD86F71BDD1E1C23B5" class="Dicotyledoneae" family="Scrophulariaceae" genus="Verbascum" higherTaxonomySource="treatment-meta" kingdom="Plantae" lsidName="Verbascum tourneuxii" order="Lamiales" pageId="0" pageNumber="115" rank="species" species="tourneuxii">Verbascum tourneuxii</taxonomicName>
Aschers., ap. Barbey 1. C: 182, nom. nud.. Aschers., ap. Aschers. &amp; Schweinf. Illustr.
<normalizedToken id="66661B473A9E5619BC7F40EAFFA19F44" originalValue="lI">l'I</normalizedToken>
. d. Egypt. in
<normalizedToken id="FE5141D721F6A01E33680A7D99C848E6" originalValue="Mém">Mem</normalizedToken>
. Instit. Egypt., II: 114. 1887.
</paragraph>
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<paragraph id="CFE72EBE3AB74CA89B2EEB6E6E800F67" pageId="0" pageNumber="115">
<typeStatus id="4C5DACF7BC3ABF04A0A8D3BCE999DD6D">Type</typeStatus>
.
</paragraph>
<paragraph id="98A9DCC68D21A9C5E80475F0EA2A5E52" pageId="0" pageNumber="115">
[
<materialsCitation id="5CAED6B144A7E75E05D216883C4F168A" collectingDate="1879-04" collectorName="A. Letourneux" country="Egypt" county="Cyrenaicae" location="In" municipality="Matrouka in Marmorica" specimenCount="1" typeStatus="lectotype">
<collectingCountry id="EF3DF56EFE0EA14B6B307BED908A9A16" name="Egypt">Egypt</collectingCountry>
].
<location id="1438B6BF1529B68B890A28AFA660DA9D" LSID="urn:lsid:plazi:treatment:34E7283141BA563398E7304180014C15:1438B6BF1529B68B890A28AFA660DA9D" country="Egypt" county="Cyrenaicae" municipality="Matrouka in Marmorica" name="In">In</location>
apricis calcareo-argillosis prope
<location id="F4A860A3A5BB7B02D1CAAD54E51C7E21" LSID="urn:lsid:plazi:treatment:34E7283141BA563398E7304180014C15:F4A860A3A5BB7B02D1CAAD54E51C7E21" country="Egypt" county="Cyrenaicae" municipality="Matrouka in Marmorica" name="Oum Rakoumi">Oum Rakoumi</location>
et
<collectingMunicipality id="AEA0665CDB97030AAC4EDB4BF78E2B0F">Matrouka in Marmorica</collectingMunicipality>
ad limites
<collectingCounty id="1EEE8D9355AFD8747CECBA12EBCCA572">Cyrenaicae</collectingCounty>
,
<collectingDate id="31BABBF18F8AE7F92CE890C25817B792" value="1879-04">April 1879</collectingDate>
,
<emphasis id="F418DDD610BC68EBCBC9105CB80AF255" italics="true" pageId="0" pageNumber="115">
<collectorName id="1AC7BA6500AA43FFC31E34638BA98636">A. Letourneux</collectorName>
, s.n.
</emphasis>
(
<typeStatus id="DC2C5179342AAA58EDF8E252F5C58B95">lectotype</typeStatus>
, designated here: G [G00015113]!; isolectotype: W [W1889-0043225]!, G [G00015111, G00015112, G00015114]!, S [S10-27120]!, K [K000975868]!, P [P03417358, P03417357, P03417360, P03417361]!). [image of
<typeStatus id="3B5F9FDB727992684554716795949991">lectotype</typeStatus>
available at https://plants.jstor.org/stable/viewer/10.5555/al.ap.specimen.g00015113]
</materialsCitation>
.
</paragraph>
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<subSubSection id="733929BB645CE70FDA6815F3152ECFE1" pageId="0" pageNumber="115" type="notes">
<paragraph id="8EA361C1E490CD7A3A87B32DFBBB9262" pageId="0" pageNumber="115">Notes.</paragraph>
<paragraph id="5B2FFF33461E2B7D9A893D14A2690E13" pageId="0" pageNumber="115">
Ascherson (in
<bibRefCitation id="4545882948F7E655C0261397CCB321AC" author="Ascherson, P" journalOrPublisher="PhD Thesis, Colorado State University, United States" pageId="0" pageNumber="115" refId="B5" refString="Ascherson, P, Schweinfurth, G, 1887. Illustration de la Flore d'Egypte. Memoires de l'Institut Egyptien [preprint] 2(1): 189, 114." title="Illustration de la Flore d'Egypte. Memoires de l'Institut Egyptien [preprint] 2 (1): 189, 114." year="1887">Ascherson and Schweinfurth 1887</bibRefCitation>
) described
<taxonomicName id="87290CBF03C774A51164371EC9B8F6BA" class="Dicotyledoneae" family="Scrophulariaceae" genus="Verbascum" kingdom="Plantae" lsidName="Verbascum letourneuxii" order="Lamiales" pageId="0" pageNumber="115" rank="species" species="letourneuxii">
<emphasis id="836C595B972C00776D99AD2D666DE1B9" italics="true" pageId="0" pageNumber="115">Verbascum letourneuxii</emphasis>
</taxonomicName>
on the basis of specimens separately collected by Ehrenberg and A. Letourneux from the Alexandria region in the north of Egypt. Within the protologue
<bibRefCitation id="C87AE3D43497E9AFB068E68BF8B0233F" author="Ascherson, P" journalOrPublisher="PhD Thesis, Colorado State University, United States" pageId="0" pageNumber="115" refId="B5" refString="Ascherson, P, Schweinfurth, G, 1887. Illustration de la Flore d'Egypte. Memoires de l'Institut Egyptien [preprint] 2(1): 189, 114." title="Illustration de la Flore d'Egypte. Memoires de l'Institut Egyptien [preprint] 2 (1): 189, 114." year="1887">Ascherson and Schweinfurth (1887)</bibRefCitation>
confess that the specimen brought by Ehrenberg only constitutes a skeleton of a plant in fruit, while specimens collected by Letourneux, from a few kilometers from the collection locality of Ehrenberg, are well-developed specimens. However, the authors did not indicate the name of the herbarium housing the type specimen. According to
<bibRefCitation id="601D34922197A0D7373A34C106A44A9F" DOI="https://doi.org/10.5962/bhl.title.48631" author="Stafleu, FA" journalOrPublisher="Phytotaxa" pageId="0" pageNumber="115" refId="B106" refString="Stafleu, FA, Cowan, RS, 1976-1988. Taxonomic literature: A selective guide to botanical publications and collections with dates, commentaries and types, edition. 2 (Vol. 1-7). Bohn, Scheltema &amp; Holkema, Utrecht. https://doi.org/10.5962/bhl.title.48631" title="Taxonomic literature: A selective guide to botanical publications and collections with dates, commentaries and types, edition. 2 (Vol. 1 - 7). Bohn, Scheltema &amp; Holkema, Utrecht." url="https://doi.org/10.5962/bhl.title.48631" year="1976 - 1988">Stafleu and Cowan (1976-1988)</bibRefCitation>
, the original material of Ehrenberg, Letourneux, Ascherson and Schweinfurth were kept at Herbs B, C, G, K, L, LE, P, S, and W. Nevertheless, the Berlin (B) herbarium was bombed during World War II on the night of 1-2 March 1943 (
<bibRefCitation id="E5764DCF449E689A2747F63503939D97" author="Hiepko, P" journalOrPublisher="Englera" pageId="0" pageNumber="115" pagination="219 - 252" refId="B47" refString="Hiepko, P, 1987. The collections of the Botanical Museum Berlin-Dahlem (B) and their history. Englera 7: 219 - 252" title="The collections of the Botanical Museum Berlin-Dahlem (B) and their history." volume="7" year="1987">Hiepko 1987</bibRefCitation>
); hence, a good part of the
<normalizedToken id="34EB3293144EDCF9DC546CB1FF7BCC33" originalValue="authors">authors'</normalizedToken>
collection has been lost (
<bibRefCitation id="5993DAD0B59C9A0526F6F1303A0F9281" DOI="https://doi.org/10.5962/bhl.title.48631" author="Stafleu, FA" journalOrPublisher="Phytotaxa" pageId="0" pageNumber="115" refId="B106" refString="Stafleu, FA, Cowan, RS, 1976-1988. Taxonomic literature: A selective guide to botanical publications and collections with dates, commentaries and types, edition. 2 (Vol. 1-7). Bohn, Scheltema &amp; Holkema, Utrecht. https://doi.org/10.5962/bhl.title.48631" title="Taxonomic literature: A selective guide to botanical publications and collections with dates, commentaries and types, edition. 2 (Vol. 1 - 7). Bohn, Scheltema &amp; Holkema, Utrecht." url="https://doi.org/10.5962/bhl.title.48631" year="1976 - 1988">Stafleu and Cowan 1976-1988</bibRefCitation>
).
</paragraph>
<paragraph id="DC6273AC4E604B96298A9BB942577C45" pageId="0" pageNumber="115">
Based on the type specification given in the protologue (locality, collector, and collection date), eleven sheets were traced in different herbaria belonging to the Letourneux collections: W (W1889-0043225), G (G00015111, G00015112, G00015113, G00015114), S (S10-27120), K (K000975868), and P (P03417358, P03417357, P03417360, P03417361). Since the type has not been specified, all of the specimens are to be recognized as syntypes according to Art. 9.6 of the ICN (
<bibRefCitation id="CA4CEB301FB99F9D8AF1C4CFC31FB9A1" DOI="https://doi.org/10.12705/Code.2018" author="Turland, NJ" journalOrPublisher="Phytotaxa" pageId="0" pageNumber="115" refId="B108" refString="Turland, NJ, Wiersema, JH, Barrie, FR, Greuter, W, Hawksworth, DL, Herendeen, PS, Knapp, S, Kusber, WH, Li, DZ, Marhold, K, May, TW, McNeill, J, Monro, AM, Prado, J, Price, MJ, Smith, GF [Eds], 2018. International Code of Nomenclature for algae, fungi, and plants (Shenzhen Code) adopted by the Nineteenth International Botanical Congress Shenzhen, China, July 2017. Regnum Vegetabile 159. Koeltz Botanical Books, Glashuetten. https://doi.org/10.12705/Code.2018" title="International Code of Nomenclature for algae, fungi, and plants (Shenzhen Code) adopted by the Nineteenth International Botanical Congress Shenzhen, China, July 2017. Regnum Vegetabile 159. Koeltz Botanical Books, Glashuetten." url="https://doi.org/10.12705/Code.2018" year="2018">Turland et al. 2018</bibRefCitation>
). The sheet G00015113 preserved at Herb. G is here designated as lectotype for the name
<taxonomicName id="7D1F78D655FF36D4A29594B0104F32B3" class="Magnoliopsida" family="Scrophulariaceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="V. letourneuxii" order="Lamiales" pageId="0" pageNumber="115" phylum="Tracheophyta" rank="species" species="letourneuxii">
<emphasis id="68765881AC19785BF9B4B5E9BA7CCEC2" italics="true" pageId="0" pageNumber="115">V. letourneuxii</emphasis>
</taxonomicName>
, since it is the specimen that shows the best quality of preservation of the important diagnostic features.
</paragraph>
</subSubSection>
</treatment>
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