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<document id="4D27B685723AFE809F42A018C0E0F210" ID-DOI="10.5281/zenodo.187806" ID-GBIF-Dataset="9a495f32-3eb8-4b35-bd46-adca3b5db2d2" ID-ISSN="1175-5326" ID-Zenodo-Dep="187806" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461074216081" checkinUser="plazi" docAuthor="Machado, Angelo B. M." docDate="2009" docId="1312774BFFD1FFE3FF04FF70FCA9FBE8" docLanguage="en" docName="zt02108p036.pdf" docOrigin="Zootaxa 2108" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Denticulobasis Machado, 2009, gen. nov." docType="treatment" docVersion="12" lastPageNumber="9" masterDocId="EF2B0F33FFD9FFEBFF93FFE7FFA7FFA6" masterDocTitle="Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae)." masterLastPageNumber="36" masterPageNumber="1" pageNumber="9" updateTime="1698585892014" updateUser="plazi">
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<mods:title id="B78EF757F288F48D71AA454E6235AD4D">Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae).</mods:title>
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<mods:name id="95E76BCFDC1F7B1FFCA64DF2ED55BA30" type="personal">
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<mods:namePart id="290EE17179F99A479EF7BD12DADC2989">Machado, Angelo B. M.</mods:namePart>
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<mods:typeOfResource id="791E5CC2ECD5DF717F7372A1BB1AE994">text</mods:typeOfResource>
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<mods:part id="AF057C2DC64C7C05E4A47E38C9540960">
<mods:date id="44D04F9B2FFE2AA594CA1C37EB1D3DAE">2009</mods:date>
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<mods:identifier id="090677BFCEFED2E1BAF6A01DF1C73249" type="DOI">10.5281/zenodo.187806</mods:identifier>
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<treatment id="1312774BFFD1FFE3FF04FF70FCA9FBE8" ID-DOI="http://doi.org/10.5281/zenodo.4391351" ID-GBIF-Taxon="119617246" ID-Zenodo-Dep="4391351" LSID="urn:lsid:plazi:treatment:1312774BFFD1FFE3FF04FF70FCA9FBE8" httpUri="http://treatment.plazi.org/id/1312774BFFD1FFE3FF04FF70FCA9FBE8" lastPageNumber="9" pageId="8" pageNumber="9">
<subSubSection id="D3A195D6FFD1FFE3FF04FF70FE18FF17" box="[151,447,151,177]" pageId="8" pageNumber="9" type="nomenclature">
<paragraph id="9B04C65DFFD1FFE3FF04FF70FE18FF17" blockId="8.[151,447,151,177]" box="[151,447,151,177]" pageId="8" pageNumber="9">
<heading id="C04C7131FFD1FFE3FF04FF70FE18FF17" bold="true" box="[151,447,151,177]" fontSize="11" level="1" pageId="8" pageNumber="9" reason="1">
<emphasis id="A9CF1A4FFFD1FFE3FF04FF70FE18FF17" bold="true" box="[151,447,151,177]" pageId="8" pageNumber="9">
<taxonomicName id="5CBBBDDEFFD1FFE3FF04FF70FEEDFF17" ID-CoL="4396" box="[151,330,151,177]" class="Insecta" family="Coenagrionidae" genus="Denticulobasis" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="A9CF1A4FFFD1FFE3FF04FF70FEEDFF17" bold="true" box="[151,330,151,177]" italics="true" pageId="8" pageNumber="9">Denticulobasis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B2FCA734FFD1FFE3FEC2FF70FE18FF17" box="[337,447,151,177]" pageId="8" pageNumber="9" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="D3A195D6FFD1FFE3FF04FF07FC4DFF5D" box="[151,1002,224,251]" pageId="8" pageNumber="9" type="materials_examined">
<paragraph id="9B04C65DFFD1FFE3FF04FF07FC4DFF5D" blockId="8.[151,1438,224,1102]" box="[151,1002,224,251]" pageId="8" pageNumber="9">
<emphasis id="A9CF1A4FFFD1FFE3FF04FF07FE9CFF5C" bold="true" box="[151,315,224,250]" pageId="8" pageNumber="9">
<typeStatus id="440078FFFFD1FFE3FF04FF07FF73FF5C" box="[151,212,224,250]" pageId="8" pageNumber="9">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="5CBBBDDEFFD1FFE3FED0FF06FDC0FF5C" box="[323,615,225,250]" class="Insecta" family="Coenagrionidae" genus="Denticulobasis" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="garrisoni" status="sp. nov.">
<emphasis id="A9CF1A4FFFD1FFE3FED0FF06FDC0FF5C" box="[323,615,225,250]" italics="true" pageId="8" pageNumber="9">Denticulobasis garrisoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B2FCA734FFD1FFE3FDFDFF07FD6CFF5D" box="[622,715,224,251]" pageId="8" pageNumber="9" rank="species">
<emphasis id="A9CF1A4FFFD1FFE3FDFDFF07FD63FF5C" bold="true" box="[622,708,224,250]" pageId="8" pageNumber="9">sp. nov</emphasis>
.
</taxonomicNameLabel>
, by present designation.
</paragraph>
</subSubSection>
<subSubSection id="D3A195D6FFD1FFE3FF55FEE0FCD2FEC9" pageId="8" pageNumber="9" type="etymology">
<paragraph id="9B04C65DFFD1FFE3FF55FEE0FCD2FEC9" blockId="8.[151,1438,224,1102]" pageId="8" pageNumber="9">
<emphasis id="A9CF1A4FFFD1FFE3FF55FEE0FEF0FE87" bold="true" box="[198,343,263,289]" pageId="8" pageNumber="9">Etymology:</emphasis>
From the Latin
<emphasis id="A9CF1A4FFFD1FFE3FD8BFEEFFD37FE87" box="[536,656,264,289]" italics="true" pageId="8" pageNumber="9">denticulus</emphasis>
meaning diminutive of tooth and
<emphasis id="A9CF1A4FFFD1FFE3FBD2FEEFFBDAFE87" box="[1089,1149,264,289]" italics="true" pageId="8" pageNumber="9">basis</emphasis>
, a Greek feminine noun for pedestal or foundation; a common suffix used in
<taxonomicName id="5CBBBDDEFFD1FFE3FC90FEC9FC67FEEE" box="[771,960,302,328]" class="Insecta" family="Coenagrionidae" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="9" phylum="Arthropoda" rank="family">Coenagrionidae</taxonomicName>
. The name
<taxonomicName id="5CBBBDDEFFD1FFE3FBDBFEC8FB5FFEEE" box="[1096,1272,303,328]" class="Insecta" family="Coenagrionidae" genus="Denticulobasis" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="A9CF1A4FFFD1FFE3FBDBFEC8FB5FFEEE" box="[1096,1272,303,328]" italics="true" pageId="8" pageNumber="9">Denticulobasis</emphasis>
</taxonomicName>
is an allusion to the presence of a plate with denticles on female sternum S8.
</paragraph>
</subSubSection>
<subSubSection id="D3A195D6FFD1FFE3FF55FE9CFCA9FBE8" pageId="8" pageNumber="9" type="distribution">
<paragraph id="9B04C65DFFD1FFE3FF55FE9CFD0DFE30" blockId="8.[151,1438,224,1102]" box="[198,682,379,406]" pageId="8" pageNumber="9">
<emphasis id="A9CF1A4FFFD1FFE3FF55FE9CFECEFE33" bold="true" box="[198,361,379,405]" pageId="8" pageNumber="9">Distribution:</emphasis>
<collectingCountry id="E3AC86CDFFD1FFE3FEE3FE9BFE1FFE30" box="[368,440,380,406]" name="Brazil" pageId="8" pageNumber="9">Brazil</collectingCountry>
and
<collectingCountry id="E3AC86CDFFD1FFE3FE62FE9BFD80FE30" box="[497,551,380,406]" name="Peru" pageId="8" pageNumber="9">Peru</collectingCountry>
(
<figureCitation id="0380DAD8FFD1FFE3FDA5FE9BFD3AFE30" box="[566,669,380,406]" captionStart="FIGURE 101" captionStartId="32.[151,255,1622,1646]" captionText="FIGURE 101. Distribution map of Denticulobasis and Tuberculobasis." pageId="8" pageNumber="9">Fig. 101</figureCitation>
).
</paragraph>
<paragraph id="9B04C65DFFD1FFE3FF55FE45FEDCFDD8" blockId="8.[151,1438,224,1102]" pageId="8" pageNumber="9">
<emphasis id="A9CF1A4FFFD1FFE3FF55FE45FE58FE1A" bold="true" box="[198,511,418,444]" pageId="8" pageNumber="9">Generic characterization</emphasis>
. Medium size damselflies (HW 18.219.2 mm). Top of head brown with blue postocular spots. Thorax dominantly yellow, with three longitudinal pale blue stripes: antehumeral, metepisternal, and metepimeral. S17 dorsally brown, laterally yellow, S810 dorso-laterally yellow. Frons rounded.
<collectingCountry id="E3AC86CDFFD1FFE3FE96FDF1FEE7FD96" box="[261,320,534,560]" name="India" pageId="8" pageNumber="9">Hind</collectingCountry>
prothoracic lobe in males with very small lateral lobes, median lobe well-developed, projected caudally and two-lipped (
<figureCitation id="0380DAD8FFD1FFE3FE5AFDDAFD90FDF1" box="[457,567,573,599]" captionStart="FIGURES 4 11" captionStartId="6.[151,273,1897,1921]" captionTargetBox="[257,1330,190,1837]" captionTargetId="figure@6.[211,1375,151,1877]" captionTargetPageId="6" captionText="FIGURES 4 11. Denticulobasis, male thorax in dorsal and lateral view: D. dunklei, holotype (4, 5); D. garrisoni, holotype (6, 7); female thorax in dorsal view: D. ariken, holotype (8); D. dunklei, allotype (9); and D. garrisoni, allotype (10); female S 8 10 in lateral view, D. garrisoni, allotype (11)." httpUri="https://zenodo.org/record/187810/files/figure.png" pageId="8" pageNumber="9">Figs 46</figureCitation>
); females with well-developed lateral and median lobes (
<figureCitation id="0380DAD8FFD1FFE3FB76FDDAFAC6FDF1" box="[1253,1377,573,599]" captionStart="FIGURES 4 11" captionStartId="6.[151,273,1897,1921]" captionTargetBox="[257,1330,190,1837]" captionTargetId="figure@6.[211,1375,151,1877]" captionTargetPageId="6" captionText="FIGURES 4 11. Denticulobasis, male thorax in dorsal and lateral view: D. dunklei, holotype (4, 5); D. garrisoni, holotype (6, 7); female thorax in dorsal view: D. ariken, holotype (8); D. dunklei, allotype (9); and D. garrisoni, allotype (10); female S 8 10 in lateral view, D. garrisoni, allotype (11)." httpUri="https://zenodo.org/record/187810/files/figure.png" pageId="8" pageNumber="9">Figs 810</figureCitation>
), the latter single-lipped.
</paragraph>
<paragraph id="9B04C65DFFD1FFE3FF55FD6DFCA9FBE8" blockId="8.[151,1438,224,1102]" pageId="8" pageNumber="9">
Male mesepisternum with an antero-medial horn with a basal smooth portion and a distal portion ridged as a screw. Female mesepisternum with an antero-medial mammiliform tubercle. Metafemoral spines on distal 1/2 as long as width of femur (
<figureCitation id="0380DAD8FFD1FFE3FDBFFD3FFDDBFD54" box="[556,636,728,754]" captionStart="FIGURE 3" captionStartId="5.[151,255,1018,1042]" captionTargetBox="[434,1146,187,1017]" captionTargetId="figure@5.[415,1171,144,1052]" captionTargetPageId="5" captionText="FIGURE 3. Denticulobasis garrisoni, male, holotype: (a) metafemur; (b) metatibia; (c) tarsus; (d) tarsal claw; (e) wings [3 e from a wing scan by R. W. Garrison]." httpUri="https://zenodo.org/record/187809/files/figure.png" pageId="8" pageNumber="9">Fig. 3</figureCitation>
a); metatibial spurs shorter than intervening spaces (
<figureCitation id="0380DAD8FFD1FFE3FAB8FD3FFADEFD54" box="[1323,1401,728,754]" captionStart="FIGURE 3" captionStartId="5.[151,255,1018,1042]" captionTargetBox="[434,1146,187,1017]" captionTargetId="figure@5.[415,1171,144,1052]" captionTargetPageId="5" captionText="FIGURE 3. Denticulobasis garrisoni, male, holotype: (a) metafemur; (b) metatibia; (c) tarsus; (d) tarsal claw; (e) wings [3 e from a wing scan by R. W. Garrison]." httpUri="https://zenodo.org/record/187809/files/figure.png" pageId="8" pageNumber="9">Fig. 3</figureCitation>
b). Supplementary tooth of tarsal claw well-developed (
<figureCitation id="0380DAD8FFD1FFE3FD69FD19FCE3FCBE" box="[762,836,766,792]" captionStart="FIGURE 3" captionStartId="5.[151,255,1018,1042]" captionTargetBox="[434,1146,187,1017]" captionTargetId="figure@5.[415,1171,144,1052]" captionTargetPageId="5" captionText="FIGURE 3. Denticulobasis garrisoni, male, holotype: (a) metafemur; (b) metatibia; (c) tarsus; (d) tarsal claw; (e) wings [3 e from a wing scan by R. W. Garrison]." httpUri="https://zenodo.org/record/187809/files/figure.png" pageId="8" pageNumber="9">Figs 3</figureCitation>
c, d). Wings petiolated distal to Ac for a distance as long as or twice as long as Ac length (
<figureCitation id="0380DAD8FFD1FFE3FDCDFCC2FD03FC99" box="[606,676,805,831]" captionStart="FIGURE 3" captionStartId="5.[151,255,1018,1042]" captionTargetBox="[434,1146,187,1017]" captionTargetId="figure@5.[415,1171,144,1052]" captionTargetPageId="5" captionText="FIGURE 3. Denticulobasis garrisoni, male, holotype: (a) metafemur; (b) metatibia; (c) tarsus; (d) tarsal claw; (e) wings [3 e from a wing scan by R. W. Garrison]." httpUri="https://zenodo.org/record/187809/files/figure.png" pageId="8" pageNumber="9">Fig. 3</figureCitation>
e). R
<quantity id="5C436BB8FFD1FFE3FD4CFCC2FCACFC99" box="[735,779,805,831]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="8" pageNumber="9" unit="in" value="3.0">3 in</quantity>
FW and HW originating closer to Px 6 (
<figureCitation id="0380DAD8FFD1FFE3FB63FCC2FA91FC99" box="[1264,1334,805,831]" captionStart="FIGURE 3" captionStartId="5.[151,255,1018,1042]" captionTargetBox="[434,1146,187,1017]" captionTargetId="figure@5.[415,1171,144,1052]" captionTargetPageId="5" captionText="FIGURE 3. Denticulobasis garrisoni, male, holotype: (a) metafemur; (b) metatibia; (c) tarsus; (d) tarsal claw; (e) wings [3 e from a wing scan by R. W. Garrison]." httpUri="https://zenodo.org/record/187809/files/figure.png" pageId="8" pageNumber="9">Fig. 3</figureCitation>
e). Penis (
<figureCitation id="0380DAD8FFD1FFE3FF0CFCABFF48FCC0" box="[159,239,844,870]" captionStart="FIGURE 1" captionStartId="4.[151,255,1006,1030]" captionTargetBox="[562,1032,498,981]" captionTargetId="figure@4.[541,1045,465,982]" captionTargetPageId="4" captionText="FIGURE 1. Denticulobasis garrisoni, paratype: penis with distal segment raised [by R. W. Garrison]." httpUri="https://zenodo.org/record/187807/files/figure.png" pageId="8" pageNumber="9">Figs. 1</figureCitation>
) distal segment expanded laterally along distal 1/3 forming a large lateral terminal lobe and with two lateral lobes on each side. No terminal fold. Internal fold with a filiform process directed disto-laterally (
<figureCitation id="0380DAD8FFD1FFE3FAFEFC95FF02FC15" captionStart="FIGURE 1" captionStartId="4.[151,255,1006,1030]" captionTargetBox="[562,1032,498,981]" captionTargetId="figure@4.[541,1045,465,982]" captionTargetPageId="4" captionText="FIGURE 1. Denticulobasis garrisoni, paratype: penis with distal segment raised [by R. W. Garrison]." httpUri="https://zenodo.org/record/187807/files/figure.png" pageId="8" pageNumber="9">Fig. 1</figureCitation>
). Male cercus with a dorso-lateral membranous depression at base (
<figureCitation id="0380DAD8FFD1FFE3FC72FC7EFBD4FC15" box="[993,1139,921,947]" captionStart="FIGURES 12 17" captionStartId="7.[151,269,1977,2001]" captionTargetBox="[186,1411,190,1976]" captionTargetId="figure@7.[176,1420,183,2013]" captionTargetPageId="7" captionText="FIGURES 12 17. Denticulobasis, male anal appendages in dorsal, lateral, and ventro-posterior view in holotypes of D." httpUri="https://zenodo.org/record/187811/files/figure.png" pageId="8" pageNumber="9">Figs. 1213</figureCitation>
) and no ventral process. Cercus in lateral view subequal to S10 (excluding distal lobe). Female sternum S8 with a denticulated vulvar plate (Figs. 18, 20, 22). Ovipositor extending to about level of tip of cercus (
<figureCitation id="0380DAD8FFD1FFE3FB84FC01FBCAFBA6" box="[1047,1133,998,1024]" captionStart="FIGURES 4 11" captionStartId="6.[151,273,1897,1921]" captionTargetBox="[257,1330,190,1837]" captionTargetId="figure@6.[211,1375,151,1877]" captionTargetPageId="6" captionText="FIGURES 4 11. Denticulobasis, male thorax in dorsal and lateral view: D. dunklei, holotype (4, 5); D. garrisoni, holotype (6, 7); female thorax in dorsal view: D. ariken, holotype (8); D. dunklei, allotype (9); and D. garrisoni, allotype (10); female S 8 10 in lateral view, D. garrisoni, allotype (11)." httpUri="https://zenodo.org/record/187810/files/figure.png" pageId="8" pageNumber="9">Fig. 11</figureCitation>
). Ventral border of lateral valve of ovipositor with one row of denticles each one subrectangular, directed ventro-posteriorly and associated with a robust short truncated seta (
<figureCitation id="0380DAD8FFD1FFE3FD39FBD3FD56FBE8" box="[682,753,1076,1102]" captionStart="FIGURE 2" captionStartId="4.[151,255,1888,1912]" captionTargetBox="[479,1102,1096,1850]" captionTargetId="figure@4.[472,1115,1084,1865]" captionTargetPageId="4" captionText="FIGURE 2. Ovipositors: denticulated part of ventral border of lateral valve. SEM: (a) Denticulobasis dunklei in lateral view; (b) T. williamsoni ventro-lateral view showing borders of two valves; (c) L. vacillans in lateral view." httpUri="https://zenodo.org/record/187808/files/figure.png" pageId="8" pageNumber="9">Fig. 2</figureCitation>
a).
</paragraph>
</subSubSection>
</treatment>
</document>