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<document id="B0EF59378B44FD1CDF7AED830BF73B48" ID-DOI="10.5281/zenodo.187806" ID-GBIF-Dataset="9a495f32-3eb8-4b35-bd46-adca3b5db2d2" ID-ISSN="1175-5326" ID-Zenodo-Dep="187806" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461074216081" checkinUser="plazi" docAuthor="Machado, Angelo B. M." docDate="2009" docId="1312774BFFC7FFF4FF04FBB8FA32FC36" docLanguage="en" docName="zt02108p036.pdf" docOrigin="Zootaxa 2108" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Tuberculobasis williamsoni Machado, 2009, sp. nov." docType="treatment" docVersion="12" lastPageNumber="32" masterDocId="EF2B0F33FFD9FFEBFF93FFE7FFA7FFA6" masterDocTitle="Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae)." masterLastPageNumber="36" masterPageNumber="1" pageNumber="31" updateTime="1698585892014" updateUser="plazi">
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<mods:title id="3AFB7083DA38EBBBF3383E331D629CA9">Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae).</mods:title>
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<mods:namePart id="43E4D767A432E7C08FBDBFED56B0FDAA">Machado, Angelo B. M.</mods:namePart>
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<mods:part id="FDD10CF192FB7644662BAD47D2650EEA">
<mods:date id="CE48AEA43B91D8207DBAE2E60B0F5148">2009</mods:date>
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<treatment id="1312774BFFC7FFF4FF04FBB8FA32FC36" ID-DOI="http://doi.org/10.5281/zenodo.4391385" ID-GBIF-Taxon="119617261" ID-Zenodo-Dep="4391385" LSID="urn:lsid:plazi:treatment:1312774BFFC7FFF4FF04FBB8FA32FC36" httpUri="http://treatment.plazi.org/id/1312774BFFC7FFF4FF04FBB8FA32FC36" lastPageId="31" lastPageNumber="32" pageId="30" pageNumber="31">
<subSubSection id="D3A195D6FFC7FFF5FF04FBB8FDE9FB06" pageId="30" pageNumber="31" type="nomenclature">
<paragraph id="9B04C65DFFC7FFF5FF04FBB8FDE0FBDF" blockId="30.[151,590,1119,1184]" box="[151,583,1119,1145]" pageId="30" pageNumber="31">
<heading id="C04C7131FFC7FFF5FF04FBB8FDE0FBDF" bold="true" box="[151,583,1119,1145]" fontSize="11" level="1" pageId="30" pageNumber="31" reason="1">
<emphasis id="A9CF1A4FFFC7FFF5FF04FBB8FDE0FBDF" bold="true" box="[151,583,1119,1145]" pageId="30" pageNumber="31">
<taxonomicName id="5CBBBDDEFFC7FFF5FF04FBB8FE44FBDF" ID-CoL="7D9XH" box="[151,483,1119,1145]" class="Insecta" family="Coenagrionidae" genus="Tuberculobasis" kingdom="Animalia" order="Odonata" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="williamsoni" status="sp. nov.">
<emphasis id="A9CF1A4FFFC7FFF5FF04FBB8FE44FBDF" bold="true" box="[151,483,1119,1145]" italics="true" pageId="30" pageNumber="31">Tuberculobasis williamsoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B2FCA734FFC7FFF5FE79FBB8FDE0FBDF" box="[490,583,1119,1145]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="9B04C65DFFC7FFF5FF04FB61FDE9FB06" blockId="30.[151,590,1119,1184]" box="[151,590,1158,1184]" pageId="30" pageNumber="31">
<figureCitation id="0380DAD8FFC7FFF5FF04FB61FEA2FB06" box="[151,261,1158,1184]" captionStart="FIGURE 2" captionStartId="4.[151,255,1888,1912]" captionTargetBox="[479,1102,1096,1850]" captionTargetId="figure@4.[472,1115,1084,1865]" captionTargetPageId="4" captionText="FIGURE 2. Ovipositors: denticulated part of ventral border of lateral valve. SEM: (a) Denticulobasis dunklei in lateral view; (b) T. williamsoni ventro-lateral view showing borders of two valves; (c) L. vacillans in lateral view." httpUri="https://zenodo.org/record/187808/files/figure.png" pageId="30" pageNumber="31">Figures 2</figureCitation>
b, 3334, 52, 7274, 93, 98.
</paragraph>
</subSubSection>
<subSubSection id="D3A195D6FFC7FFF5FF04FB34FB05FAB2" pageId="30" pageNumber="31" type="etymology">
<paragraph id="9B04C65DFFC7FFF5FF04FB34FB05FAB2" blockId="30.[151,1436,1235,2037]" pageId="30" pageNumber="31">
<emphasis id="A9CF1A4FFFC7FFF5FF04FB34FE8BFB4B" bold="true" box="[151,300,1235,1261]" pageId="30" pageNumber="31">Etymology:</emphasis>
Named after Edward Bruce Williamson, whose odonatological expeditions to
<collectingCountry id="E3AC86CDFFC7FFF5FB63FB34FAC0FB4B" box="[1264,1383,1235,1261]" name="Colombia" pageId="30" pageNumber="31">Colombia</collectingCountry>
and
<collectingCountry id="E3AC86CDFFC7FFF5FF04FB1DFEA8FAB2" box="[151,271,1274,1300]" name="Venezuela" pageId="30" pageNumber="31">Venezuela</collectingCountry>
yielded this species and an enormous number of valuable
<taxonomicName id="5CBBBDDEFFC7FFF5FC2FFB1DFBBEFAB2" box="[956,1049,1274,1300]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="30" pageNumber="36" phylum="Arthropoda" rank="order">odonata</taxonomicName>
specimens.
</paragraph>
</subSubSection>
<subSubSection id="D3A195D6FFC7FFF5FF55FAC7FAEFFA5A" pageId="30" pageNumber="31" type="materials_examined">
<paragraph id="9B04C65DFFC7FFF5FF55FAC7FAEFFA5A" blockId="30.[151,1436,1235,2037]" pageId="30" pageNumber="31">
<emphasis id="A9CF1A4FFFC7FFF5FF55FAC7FDB5FA9C" bold="true" box="[198,530,1312,1338]" pageId="30" pageNumber="31">
<typeStatus id="440078FFFFC7FFF5FF55FAC7FEA4FA9C" box="[198,259,1312,1338]" pageId="30" pageNumber="31">Type</typeStatus>
specimens:
<typeStatus id="440078FFFFC7FFF5FE33FAC7FDB5FA9C" box="[416,530,1312,1338]" pageId="30" pageNumber="31" type="holotype">Holotype</typeStatus>
</emphasis>
(3) and
<emphasis id="A9CF1A4FFFC7FFF5FD17FAC7FD40FA9C" bold="true" box="[644,743,1312,1338]" pageId="30" pageNumber="31">allotype</emphasis>
(Ƥ):
<collectingCountry id="E3AC86CDFFC7FFF5FCBEFAC7FC6BFA9C" box="[813,972,1312,1338]" name="Colombia" pageId="30" pageNumber="31">COLOMBIA</collectingCountry>
, Fundación, State of Magdalena,
<date id="EF05E09DFFC7FFF5FAF1FAC7FF73FAC7" pageId="30" pageNumber="31" value="1917-01-11">11-I- 1917</date>
, J.H. &amp; E.B. Williamson leg. (
<collectionCode id="FDAA5E98FFC7FFF5FDA4FAA0FD3EFAC7" box="[567,665,1351,1377]" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="30" pageNumber="31">UMMZ</collectionCode>
).
<emphasis id="A9CF1A4FFFC7FFF5FD3FFAA0FC94FAC7" bold="true" box="[684,819,1351,1377]" pageId="30" pageNumber="31">
<typeStatus id="440078FFFFC7FFF5FD3FFAA0FC89FAC7" box="[684,814,1351,1377]" pageId="30" pageNumber="31" type="paratype">Paratypes</typeStatus>
:
</emphasis>
3 3 and 5 Ƥ: same data as
<typeStatus id="440078FFFFC7FFF5FBE0FAA0FB7EFAC7" box="[1139,1241,1351,1377]" pageId="30" pageNumber="31" type="holotype">holotype</typeStatus>
(three
<typeStatus id="440078FFFFC7FFF5FAB8FAA0FA3BFAC7" box="[1323,1436,1351,1377]" pageId="30" pageNumber="31" type="paratype">paratypes</typeStatus>
were collected along a stream in a shaded area); 1 3, 2 Ƥ: El Banco, State of Magdalena,
<date id="EF05E09DFFC7FFF5FB54FA89FA98FA2E" box="[1223,1343,1390,1416]" pageId="30" pageNumber="31" value="1917-01-21">21-I-1917</date>
, J.H. &amp; E.B. Williamson leg.; 1 3, 2 Ƥ:
<collectingCountry id="E3AC86CDFFC7FFF5FD80FA73FD62FA08" box="[531,709,1428,1454]" name="Venezuela" pageId="30" pageNumber="31">VENEZUELA</collectingCountry>
, Zulia State, Encontrados,
<date id="EF05E09DFFC7FFF5FC6CFA73FB2EFA08" box="[1023,1161,1428,1454]" pageId="30" pageNumber="31" value="1920-04-20">20-IV-1920</date>
, J.M., E.B. Williamson &amp; D.H. Ditzleri leg; 1 3: San Felipe,
<date id="EF05E09DFFC7FFF5FDC8FA5CFD4FFA73" box="[603,744,1467,1493]" pageId="30" pageNumber="31" value="1920-03-20">20-III-1920</date>
, J.M., E.B. Williamson &amp; D.H. Ditzleri leg.. 3 3 and 7 Ƥ
<typeStatus id="440078FFFFC7FFF5FF04FA05FEA0FA5A" box="[151,263,1506,1532]" pageId="30" pageNumber="31" type="paratype">paratypes</typeStatus>
(
<collectionCode id="FDAA5E98FFC7FFF5FE85FA05FEDFFA5A" box="[278,376,1506,1532]" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="30" pageNumber="31">UMMZ</collectionCode>
), 1 3 and 1 Ƥ
<typeStatus id="440078FFFFC7FFF5FDB7FA05FD33FA5A" box="[548,660,1506,1532]" pageId="30" pageNumber="31" type="paratype">paratypes</typeStatus>
(
<collectionCode id="FDAA5E98FFC7FFF5FD37FA05FD5EFA5A" box="[676,761,1506,1532]" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="30" pageNumber="31">MNRJ</collectionCode>
); 1 3 and 1 Ƥ (A. B.M.M), 1 3 and 1 Ƥ (R.W.G.).
</paragraph>
</subSubSection>
<subSubSection id="D3A195D6FFC7FFF4FF55F9EFFB28FCBA" lastPageId="31" lastPageNumber="32" pageId="30" pageNumber="31" type="description">
<paragraph id="9B04C65DFFC7FFF5FF55F9EFFDBFF984" blockId="30.[151,1436,1235,2037]" box="[198,536,1544,1570]" pageId="30" pageNumber="31">
<emphasis id="A9CF1A4FFFC7FFF5FF55F9EFFEF0F984" bold="true" box="[198,343,1544,1570]" pageId="30" pageNumber="31">Description</emphasis>
. Male
<typeStatus id="440078FFFFC7FFF5FE3AF9EFFDB4F984" box="[425,531,1544,1570]" pageId="30" pageNumber="31" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="9B04C65DFFC7FFF5FF55F9C8FEFEF930" blockId="30.[151,1436,1235,2037]" pageId="30" pageNumber="31">
<emphasis id="A9CF1A4FFFC7FFF5FF55F9C8FEAEF9EF" bold="true" box="[198,265,1583,1609]" pageId="30" pageNumber="31">Head</emphasis>
. Labium yellowish white. Labrum yellow; genae grey, base of mandibles, anteclypeus, postclypeus, antefrons, and top of head brown. Postocular lobes light orange with grey postocular spots. Rear of head yellowish white.
</paragraph>
<paragraph id="9B04C65DFFC7FFF5FF55F944FF6AF827" blockId="30.[151,1436,1235,2037]" pageId="30" pageNumber="31">
<emphasis id="A9CF1A4FFFC7FFF5FF55F944FE8DF91B" bold="true" box="[198,298,1699,1725]" pageId="30" pageNumber="31">Thorax.</emphasis>
Prothorax: Anterior lobe grey, median and posterior lobes brown. Propleuron yellowish brown. Pterothorax (as
<figureCitation id="0380DAD8FFC7FFF5FEC1F92DFE12F942" box="[338,437,1738,1764]" captionStart="FIGURE 101" captionStartId="32.[151,255,1622,1646]" captionText="FIGURE 101. Distribution map of Denticulobasis and Tuberculobasis." pageId="30" pageNumber="31">Fig. 101</figureCitation>
b) brown with greyish stripes on mesepisternum and metepisternum. Legs yellowish brown, wings hyaline, venation and pterostigma brown. Px in FW 11 (10%), 12 (90%); in HW 9 (30%), 10 (40%), 11 (30%). R
<quantity id="5C436BB8FFC7FFF5FE16F8FDFE16F892" box="[389,433,1818,1844]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="30" pageNumber="31" unit="in" value="3.0">3 in</quantity>
FW arising near Px 5 (20%) and 6 (80%), in HW at Px 4 (10%), 5 (80%), 6 (10%). Petiolation distal to Ac by a distance equal (60%) or ½ (40%) of its length in FW, equal (30%) or ½ (70%) in HW.
</paragraph>
<paragraph id="9B04C65DFFC7FFF5FF55F869FB18F853" blockId="30.[151,1436,1235,2037]" pageId="30" pageNumber="31">
<emphasis id="A9CF1A4FFFC7FFF5FF55F869FEEFF80E" bold="true" box="[198,328,1934,1960]" pageId="30" pageNumber="31">Abdomen.</emphasis>
S12 brownish yellow with a lateral grey stripe. S2 with a square yellow dorsal spot at distal 1/4. S35 dorsally yellowish brown, laterally yellowish with brown distal rings; S6 brown, S710 yellowish brown. Cercus brownish yellow with dorso-lateral border dark brown; paraproct brownish.
</paragraph>
<paragraph id="9B04C65DFFC6FFF4FF55FF70FE5AFED5" blockId="31.[151,1437,151,912]" pageId="31" pageNumber="32">
<emphasis id="A9CF1A4FFFC6FFF4FF55FF70FE75FF17" bold="true" box="[198,466,151,177]" pageId="31" pageNumber="32">Structural characters</emphasis>
.
<collectingCountry id="E3AC86CDFFC6FFF4FE72FF7FFDBBFF14" box="[481,540,152,178]" name="India" pageId="31" pageNumber="32">Hind</collectingCountry>
lobe of prothorax (
<figureCitation id="0380DAD8FFC6FFF4FD6EFF7FFC24FF14" box="[765,899,152,178]" captionStart="FIGURES 25 34" captionStartId="18.[151,270,1973,1997]" captionTargetBox="[327,1275,181,1941]" captionTargetId="figure@18.[312,1277,161,1944]" captionTargetPageId="18" captionText="FIGURES 25 34. Tuberculobasis, male thorax in dorsal and lateral view in T. arara, holotype (25, 26); T. costalimai, holotype (27, 28); T. guarani, holotype (29, 30); T. inversa, holotype (31, 32); T. williamsoni, holotype (33, 34)." httpUri="https://zenodo.org/record/187813/files/figure.png" pageId="31" pageNumber="32">Figs 33, 34</figureCitation>
) with median lobe semicircular and projected caudally; lateral lobes very small. Borders of median lobe two-lipped; dorsal lip upraised. Mesepisternal tubercles high (1.58 mm) in dorsal view strongly divergent (
<figureCitation id="0380DAD8FFC6FFF4FCFBFF02FC66FF59" box="[872,961,229,255]" captionStart="FIGURES 25 34" captionStartId="18.[151,270,1973,1997]" captionTargetBox="[327,1275,181,1941]" captionTargetId="figure@18.[312,1277,161,1944]" captionTargetPageId="18" captionText="FIGURES 25 34. Tuberculobasis, male thorax in dorsal and lateral view in T. arara, holotype (25, 26); T. costalimai, holotype (27, 28); T. guarani, holotype (29, 30); T. inversa, holotype (31, 32); T. williamsoni, holotype (33, 34)." httpUri="https://zenodo.org/record/187813/files/figure.png" pageId="31" pageNumber="32">Fig. 33</figureCitation>
), conical, directed antero-laterally, with bases adjacent to mid-dorsal carina (
<figureCitation id="0380DAD8FFC6FFF4FDC1FEEBFD0AFE80" box="[594,685,268,294]" captionStart="FIGURES 25 34" captionStartId="18.[151,270,1973,1997]" captionTargetBox="[327,1275,181,1941]" captionTargetId="figure@18.[312,1277,161,1944]" captionTargetPageId="18" captionText="FIGURES 25 34. Tuberculobasis, male thorax in dorsal and lateral view in T. arara, holotype (25, 26); T. costalimai, holotype (27, 28); T. guarani, holotype (29, 30); T. inversa, holotype (31, 32); T. williamsoni, holotype (33, 34)." httpUri="https://zenodo.org/record/187813/files/figure.png" pageId="31" pageNumber="32">Fig. 33</figureCitation>
). Cercus 1/2 shorter than paraproct (
<figureCitation id="0380DAD8FFC6FFF4FBFBFEEBFB5BFE80" box="[1128,1276,268,294]" captionStart="FIGURES 69 77" captionStartId="25.[151,272,1670,1694]" captionTargetBox="[151,1426,187,1662]" captionTargetId="figure@25.[151,1436,138,1748]" captionTargetPageId="25" captionText="FIGURES 69 77. Tuberculobasis male, anal appendages in dorsal, lateral, and posterior views: T. macuxi, holotype (69 71); T. williamsoni, holotype (72 74); T. geijskesi, holotype (75 77)." httpUri="https://zenodo.org/record/187818/files/figure.png" pageId="31" pageNumber="32">Figs. 7273</figureCitation>
), in posterior view (
<figureCitation id="0380DAD8FFC6FFF4FF72FED5FE9FFEEA" box="[225,312,306,332]" captionStart="FIGURES 69 77" captionStartId="25.[151,272,1670,1694]" captionTargetBox="[151,1426,187,1662]" captionTargetId="figure@25.[151,1436,138,1748]" captionTargetPageId="25" captionText="FIGURES 69 77. Tuberculobasis male, anal appendages in dorsal, lateral, and posterior views: T. macuxi, holotype (69 71); T. williamsoni, holotype (72 74); T. geijskesi, holotype (75 77)." httpUri="https://zenodo.org/record/187818/files/figure.png" pageId="31" pageNumber="32">Fig. 74</figureCitation>
) with a dorso-lateral semilunar cavity limited by a denticulated dark brown border. Penis (
<figureCitation id="0380DAD8FFC6FFF4FAFFFED5FF12FED5" captionStart="FIGURES 87 94" captionStartId="29.[151,269,1856,1880]" captionTargetBox="[282,1311,194,1855]" captionTargetId="figure@29.[273,1313,117,1921]" captionTargetPageId="29" captionText="FIGURES 87 94. Penis in lateral and ectal views: D. dunklei, holotype (87); D. garrisoni, holotype (88); T. costalimai, paratype (89); T. geijskesi, holotype (90); T. mammilaris, lectotype (91); T. tirio, paratype (92); T. williamsoni, paratype (93); Leptobasis vacillans, Panamá (94) [91 by R. W. Garrison]." httpUri="https://zenodo.org/record/187820/files/figure.png" pageId="31" pageNumber="32">Fig. 93</figureCitation>
) as described for the genus.
</paragraph>
<paragraph id="9B04C65DFFC6FFF4FF55FE98FB37FE3C" blockId="31.[151,1437,151,912]" box="[198,1168,383,410]" pageId="31" pageNumber="32">
<emphasis id="A9CF1A4FFFC6FFF4FF55FE98FE0CFE3F" bold="true" box="[198,427,383,409]" pageId="31" pageNumber="32">Dimensions (mm).</emphasis>
Abdomen 32.034.4 (mean 32.6); HW 17.519.0 (mean 18.2).
</paragraph>
<paragraph id="9B04C65DFFC6FFF4FF55FE41FD8EFE66" blockId="31.[151,1437,151,912]" box="[198,553,422,448]" pageId="31" pageNumber="32">
<emphasis id="A9CF1A4FFFC6FFF4FF55FE41FEF9FE66" bold="true" box="[198,350,422,448]" pageId="31" pageNumber="32">Description.</emphasis>
Female allotype.
</paragraph>
<paragraph id="9B04C65DFFC6FFF4FF55FE2AFC0EFDFD" blockId="31.[151,1437,151,912]" pageId="31" pageNumber="32">
Color of head, thorax, and abdomen as described for male. Px in FW 12 (30%), 13 (60%), 14 (10%), in HW 10 (40%), 11 (30%), 12 (30%). R
<quantity id="5C436BB8FFC6FFF4FDC6FE13FD27FDA8" box="[597,640,500,526]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="31" pageNumber="32" unit="in" value="3.0">3 in</quantity>
FW originating near Px 6 (90%) or 5 (10%) in HW near Px 5 (80%) or 6 (20%). Petiolation in FW distal to Ac by a distance equal (14.2%) or ½ (71.5%) of its own length, in HW slightly distal (14.2%), equal (14.2%) or ½ (71.4%) its own length.
</paragraph>
<paragraph id="9B04C65DFFC6FFF4FF55FD80FF62FD50" blockId="31.[151,1437,151,912]" pageId="31" pageNumber="32">
<emphasis id="A9CF1A4FFFC6FFF4FF55FD80FE74FD27" bold="true" box="[198,467,615,641]" pageId="31" pageNumber="32">Structural characters</emphasis>
.
<collectingCountry id="E3AC86CDFFC6FFF4FE71FD8FFDBAFD24" box="[482,541,616,642]" name="India" pageId="31" pageNumber="32">Hind</collectingCountry>
lobe of prothorax (
<figureCitation id="0380DAD8FFC6FFF4FC90FD8FFCFEFD24" box="[771,857,616,642]" captionStart="FIGURES 47 54" captionStartId="20.[151,272,1755,1779]" captionTargetBox="[188,1374,193,1754]" captionTargetId="figure@20.[180,1408,117,1857]" captionTargetPageId="20" captionText="FIGURES 47 54. Tuberculobasis female thorax, in dorsal view: T. cardinalis, holotype (47); T. costalimai, allotype (48); T. inversa, Peru, Loreto (49); T. mammilaris, Venezuela (50); T. tirio, allotype (51); T. williamsoni, allotype (52); T. yanomami, paratype (53), and in dorso-lateral view: Tuberculobasis sp. 1, Manaus, Amazonas (54) [47 by N. von Ellenrieder; 50, 53 by J. De Marmels]." httpUri="https://zenodo.org/record/187815/files/figure.png" pageId="31" pageNumber="32">Fig. 52</figureCitation>
) slightly concave at middle with no definition of lateral lobes. Mesepisternal tubercles well-developed but low, connected with anterior margin of mesostigmal plate (
<figureCitation id="0380DAD8FFC6FFF4FF72FD52FE9CFD69" box="[225,315,693,719]" captionStart="FIGURES 47 54" captionStartId="20.[151,272,1755,1779]" captionTargetBox="[188,1374,193,1754]" captionTargetId="figure@20.[180,1408,117,1857]" captionTargetPageId="20" captionText="FIGURES 47 54. Tuberculobasis female thorax, in dorsal view: T. cardinalis, holotype (47); T. costalimai, allotype (48); T. inversa, Peru, Loreto (49); T. mammilaris, Venezuela (50); T. tirio, allotype (51); T. williamsoni, allotype (52); T. yanomami, paratype (53), and in dorso-lateral view: Tuberculobasis sp. 1, Manaus, Amazonas (54) [47 by N. von Ellenrieder; 50, 53 by J. De Marmels]." httpUri="https://zenodo.org/record/187815/files/figure.png" pageId="31" pageNumber="32">Fig. 52</figureCitation>
). Cercus conical (
<figureCitation id="0380DAD8FFC6FFF4FD82FD52FDCDFD69" box="[529,618,693,719]" captionStart="FIGURES 95 100" captionStartId="30.[151,274,937,961]" captionTargetBox="[292,1277,185,936]" captionTargetId="figure@30.[277,1310,164,947]" captionTargetPageId="30" captionText="FIGURES 95 100. Female S 8 10 in lateral view: T. cardinalis, holotype (95); T. mammilaris, Venezuela (96); T. costalimai, allotype (97); T. williamsoni, paratype (98); T. yanomami, paratype (99); Leptobasis vacillans, Venezuela (100) [95 by N. von Ellenrieder, 96, 99 by J. De Marmels]." httpUri="https://zenodo.org/record/187821/files/figure.png" pageId="31" pageNumber="32">Fig. 98</figureCitation>
). Ovipositor (excluding styli) not surpassing level of cercus tip (
<figureCitation id="0380DAD8FFC6FFF4FAFFFD52FF1FFD50" captionStart="FIGURES 95 100" captionStartId="30.[151,274,937,961]" captionTargetBox="[292,1277,185,936]" captionTargetId="figure@30.[277,1310,164,947]" captionTargetPageId="30" captionText="FIGURES 95 100. Female S 8 10 in lateral view: T. cardinalis, holotype (95); T. mammilaris, Venezuela (96); T. costalimai, allotype (97); T. williamsoni, paratype (98); T. yanomami, paratype (99); Leptobasis vacillans, Venezuela (100) [95 by N. von Ellenrieder, 96, 99 by J. De Marmels]." httpUri="https://zenodo.org/record/187821/files/figure.png" pageId="31" pageNumber="32">Fig. 98</figureCitation>
).
</paragraph>
<paragraph id="9B04C65DFFC6FFF4FF55FCE5FB28FCBA" blockId="31.[151,1437,151,912]" box="[198,1167,770,796]" pageId="31" pageNumber="32">
<emphasis id="A9CF1A4FFFC6FFF4FF55FCE5FE0DFCBA" bold="true" box="[198,426,770,796]" pageId="31" pageNumber="32">Dimensions (mm).</emphasis>
Abdomen 31.032.1 (mean 31.5); HW 18.920.4 (mean 19.8).
</paragraph>
</subSubSection>
<subSubSection id="D3A195D6FFC6FFF4FF55FCCFFA32FC36" pageId="31" pageNumber="32" type="discussion">
<paragraph id="9B04C65DFFC6FFF4FF55FCCFFA32FC36" blockId="31.[151,1437,151,912]" pageId="31" pageNumber="32">
<emphasis id="A9CF1A4FFFC6FFF4FF55FCCFFE9FFCE4" bold="true" box="[198,312,808,834]" pageId="31" pageNumber="32">Remarks</emphasis>
. By having male mesepisternal tubercles high with base adjacent to mid-dorsal carina, and male paraproct much longer than cercus,
<taxonomicName id="5CBBBDDEFFC6FFF4FDD6FCB7FD49FCCF" box="[581,750,848,873]" class="Insecta" family="Coenagrionidae" genus="Tuberculobasis" kingdom="Animalia" order="Odonata" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="williamsoni">
<emphasis id="A9CF1A4FFFC6FFF4FDD6FCB7FD49FCCF" box="[581,750,848,873]" italics="true" pageId="31" pageNumber="32">T. williamsoni</emphasis>
</taxonomicName>
belongs in the
<taxonomicName id="5CBBBDDEFFC6FFF4FC34FCB7FB84FCCF" box="[935,1059,848,873]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="costalimai">
<emphasis id="A9CF1A4FFFC6FFF4FC34FCB7FB84FCCF" box="[935,1059,848,873]" italics="true" pageId="31" pageNumber="32">costalimai</emphasis>
</taxonomicName>
species group, together with
<emphasis id="A9CF1A4FFFC6FFF4FA15FCB7FDA2FC36" italics="true" pageId="31" pageNumber="32">
<taxonomicName id="5CBBBDDEFFC6FFF4FA15FCB7FF7AFC36" class="Insecta" family="Coenagrionidae" genus="Tuberculobasis" kingdom="Animalia" order="Odonata" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="arara">T. arara</taxonomicName>
,
<taxonomicName id="5CBBBDDEFFC6FFF4FF7BFC90FE26FC36" box="[232,385,887,912]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="costalimai">T. costalimai</taxonomicName>
,
<taxonomicName id="5CBBBDDEFFC6FFF4FE1FFC90FDA2FC36" box="[396,517,887,912]" class="Insecta" family="Coenagrionidae" genus="Tuberculobasis" kingdom="Animalia" order="Odonata" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="guarani">T. guarani</taxonomicName>
</emphasis>
, and
<taxonomicName id="5CBBBDDEFFC6FFF4FDD7FC90FD10FC36" box="[580,695,887,912]" class="Insecta" family="Coenagrionidae" genus="Tuberculobasis" kingdom="Animalia" order="Odonata" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="inversa">
<emphasis id="A9CF1A4FFFC6FFF4FDD7FC90FD10FC36" box="[580,695,887,912]" italics="true" pageId="31" pageNumber="32">T. inversa</emphasis>
</taxonomicName>
. It can be separated from them by characters given in the keys.
</paragraph>
</subSubSection>
</treatment>
</document>